id sid tid token lemma pos 32505 1 1 UNIVERSITY UNIVERSITY NNP 32505 1 2 OF of IN 32505 1 3 KANSAS KANSAS NNP 32505 1 4 PUBLICATIONS PUBLICATIONS NNP 32505 1 5 MUSEUM MUSEUM NNP 32505 1 6 OF of IN 32505 1 7 NATURAL natural JJ 32505 1 8 HISTORY HISTORY NNP 32505 1 9 Volume volume NN 32505 1 10 18 18 CD 32505 1 11 , , , 32505 1 12 No no UH 32505 1 13 . . . 32505 2 1 6 6 CD 32505 2 2 , , , 32505 2 3 pp pp NNP 32505 2 4 . . . 32505 3 1 505 505 CD 32505 3 2 - - SYM 32505 3 3 545 545 CD 32505 3 4 , , , 32505 3 5 7 7 CD 32505 3 6 figs fig NNS 32505 3 7 . . . 32505 3 8 , , , 32505 3 9 4 4 CD 32505 3 10 pls pls UH 32505 3 11 . . . 32505 4 1 December December NNP 32505 4 2 2 2 CD 32505 4 3 , , , 32505 4 4 1969 1969 CD 32505 4 5 The the DT 32505 4 6 Systematics Systematics NNPS 32505 4 7 of of IN 32505 4 8 the the DT 32505 4 9 Frogs Frogs NNP 32505 4 10 of of IN 32505 4 11 the the DT 32505 4 12 _ _ NNP 32505 4 13 Hyla Hyla NNP 32505 4 14 rubra rubra NN 32505 4 15 _ _ NNP 32505 4 16 Group Group NNP 32505 4 17 in in IN 32505 4 18 Middle Middle NNP 32505 4 19 America America NNP 32505 4 20 BY by IN 32505 4 21 JUAN JUAN NNP 32505 4 22 R. R. NNP 32505 4 23 LEÓN LEÓN NNP 32505 4 24 University University NNP 32505 4 25 of of IN 32505 4 26 Kansas Kansas NNP 32505 4 27 Lawrence Lawrence NNP 32505 4 28 1969 1969 CD 32505 4 29 UNIVERSITY UNIVERSITY NNP 32505 4 30 OF of IN 32505 4 31 KANSAS KANSAS NNP 32505 4 32 PUBLICATIONS PUBLICATIONS NNP 32505 4 33 , , , 32505 4 34 MUSEUM MUSEUM NNP 32505 4 35 OF of IN 32505 4 36 NATURAL natural JJ 32505 4 37 HISTORY HISTORY NNP 32505 4 38 Editors Editors NNPS 32505 4 39 : : : 32505 4 40 Frank Frank NNP 32505 4 41 B. B. NNP 32505 5 1 Cross Cross NNP 32505 5 2 , , , 32505 5 3 Philip Philip NNP 32505 5 4 S. S. NNP 32505 5 5 Humphrey Humphrey NNP 32505 5 6 , , , 32505 5 7 Robert Robert NNP 32505 5 8 M. M. NNP 32505 5 9 Mengel Mengel NNP 32505 5 10 . . . 32505 6 1 Volume volume NN 32505 6 2 18 18 CD 32505 6 3 , , , 32505 6 4 No no UH 32505 6 5 . . . 32505 7 1 6 6 CD 32505 7 2 , , , 32505 7 3 pp pp NNP 32505 7 4 . . . 32505 8 1 505 505 CD 32505 8 2 - - SYM 32505 8 3 545 545 CD 32505 8 4 , , , 32505 8 5 7 7 CD 32505 8 6 figs fig NNS 32505 8 7 . . . 32505 8 8 , , , 32505 8 9 4 4 CD 32505 8 10 pls pls UH 32505 8 11 . . . 32505 9 1 Published publish VBN 32505 9 2 December December NNP 32505 9 3 2 2 CD 32505 9 4 , , , 32505 9 5 1969 1969 CD 32505 9 6 UNIVERSITY UNIVERSITY NNP 32505 9 7 OF of IN 32505 9 8 KANSAS KANSAS NNP 32505 9 9 Lawrence Lawrence NNP 32505 9 10 , , , 32505 9 11 Kansas Kansas NNP 32505 9 12 PRINTED PRINTED NNP 32505 9 13 BY by IN 32505 9 14 ROBERT ROBERT NNP 32505 9 15 R. R. NNP 32505 9 16 ( ( -LRB- 32505 9 17 BOB BOB NNP 32505 9 18 ) ) -RRB- 32505 9 19 SANDERS sander NNS 32505 9 20 , , , 32505 9 21 STATE STATE NNP 32505 9 22 PRINTER PRINTER NNP 32505 9 23 TOPEKA TOPEKA NNP 32505 9 24 , , , 32505 9 25 KANSAS KANSAS NNP 32505 9 26 1969 1969 CD 32505 9 27 The the DT 32505 9 28 Systematics Systematics NNPS 32505 9 29 of of IN 32505 9 30 the the DT 32505 9 31 Frogs Frogs NNP 32505 9 32 of of IN 32505 9 33 the the DT 32505 9 34 _ _ NNP 32505 9 35 Hyla Hyla NNP 32505 9 36 rubra rubra NN 32505 9 37 _ _ NNP 32505 9 38 Group Group NNP 32505 9 39 in in IN 32505 9 40 Middle Middle NNP 32505 9 41 America America NNP 32505 9 42 BY by IN 32505 9 43 JUAN JUAN NNP 32505 9 44 R. R. NNP 32505 9 45 LEÓN LEÓN VBD 32505 9 46 CONTENTS CONTENTS NNP 32505 9 47 PAGE PAGE NNP 32505 9 48 INTRODUCTION introduction NN 32505 9 49 508 508 CD 32505 9 50 Acknowledgments acknowledgment NNS 32505 9 51 508 508 CD 32505 9 52 Materials material NNS 32505 9 53 and and CC 32505 9 54 Methods method NNS 32505 9 55 509 509 CD 32505 9 56 THE the DT 32505 9 57 HYLA HYLA NNP 32505 9 58 RUBRA RUBRA NNP 32505 9 59 GROUP GROUP NNP 32505 9 60 509 509 CD 32505 9 61 Key Key NNP 32505 9 62 to to IN 32505 9 63 Species Species NNPS 32505 9 64 and and CC 32505 9 65 Subspecies Subspecies NNPS 32505 9 66 510 510 CD 32505 9 67 Key key JJ 32505 9 68 to to IN 32505 9 69 Known Known NNP 32505 9 70 Tadpoles Tadpoles NNP 32505 9 71 511 511 CD 32505 9 72 ACCOUNTS account NNS 32505 9 73 OF of IN 32505 9 74 SPECIES specie NNS 32505 9 75 AND and CC 32505 9 76 SUBSPECIES SUBSPECIES NNP 32505 9 77 511 511 CD 32505 9 78 _ _ NNP 32505 9 79 Hyla Hyla NNP 32505 9 80 boulengeri boulengeri NN 32505 9 81 _ _ NNP 32505 9 82 ( ( -LRB- 32505 9 83 Cope Cope NNP 32505 9 84 ) ) -RRB- 32505 9 85 511 511 CD 32505 9 86 _ _ NNP 32505 9 87 Hyla Hyla NNP 32505 9 88 foliamorta foliamorta NN 32505 9 89 _ _ NNP 32505 9 90 Fouquette Fouquette NNP 32505 9 91 520 520 CD 32505 9 92 _ _ NNP 32505 9 93 Hyla Hyla NNP 32505 9 94 rubra rubra NN 32505 9 95 _ _ NNP 32505 9 96 Laurenti Laurenti NNP 32505 9 97 524 524 CD 32505 9 98 _ _ NNP 32505 9 99 Hyla Hyla NNP 32505 9 100 elaeochroa elaeochroa NN 32505 9 101 _ _ NNP 32505 9 102 Cope Cope NNP 32505 9 103 525 525 CD 32505 9 104 _ _ NNP 32505 9 105 Hyla Hyla NNP 32505 9 106 staufferi staufferi NN 32505 9 107 _ _ NNP 32505 9 108 Cope Cope NNP 32505 9 109 532 532 CD 32505 9 110 _ _ NNP 32505 9 111 Hyla Hyla NNP 32505 9 112 staufferi staufferi NN 32505 9 113 staufferi staufferi NN 32505 9 114 _ _ NNP 32505 9 115 Cope Cope NNP 32505 9 116 537 537 CD 32505 9 117 _ _ NNP 32505 9 118 Hyla Hyla NNP 32505 9 119 staufferi staufferi NN 32505 9 120 altae altae NN 32505 9 121 _ _ NNP 32505 9 122 Dunn Dunn NNP 32505 9 123 540 540 CD 32505 9 124 EVOLUTIONARY EVOLUTIONARY NNP 32505 9 125 HISTORY history NN 32505 9 126 540 540 CD 32505 9 127 LITERATURE literature NN 32505 9 128 CITED cite VBD 32505 9 129 543 543 CD 32505 9 130 INTRODUCTION introduction NN 32505 9 131 The the DT 32505 9 132 tree tree NN 32505 9 133 frogs frog NNS 32505 9 134 of of IN 32505 9 135 the the DT 32505 9 136 _ _ NNP 32505 9 137 Hyla Hyla NNP 32505 9 138 rubra rubra NN 32505 9 139 _ _ NNP 32505 9 140 group group NN 32505 9 141 are be VBP 32505 9 142 abundant abundant JJ 32505 9 143 and and CC 32505 9 144 form form VBP 32505 9 145 a a DT 32505 9 146 conspicuous conspicuous JJ 32505 9 147 element element NN 32505 9 148 of of IN 32505 9 149 the the DT 32505 9 150 Neotropical neotropical JJ 32505 9 151 frog frog NN 32505 9 152 fauna fauna NNS 32505 9 153 . . . 32505 10 1 Representatives representative NNS 32505 10 2 of of IN 32505 10 3 the the DT 32505 10 4 group group NN 32505 10 5 occur occur VBP 32505 10 6 from from IN 32505 10 7 lowland lowland NNP 32505 10 8 México México NNP 32505 10 9 to to IN 32505 10 10 Argentina Argentina NNP 32505 10 11 ; ; : 32505 10 12 the the DT 32505 10 13 greatest great JJS 32505 10 14 diversity diversity NN 32505 10 15 is be VBZ 32505 10 16 reached reach VBN 32505 10 17 in in IN 32505 10 18 the the DT 32505 10 19 lowlands lowland NNS 32505 10 20 of of IN 32505 10 21 southeastern southeastern JJ 32505 10 22 Brazil Brazil NNP 32505 10 23 ( ( -LRB- 32505 10 24 Cochran Cochran NNP 32505 10 25 , , , 32505 10 26 1955 1955 CD 32505 10 27 ) ) -RRB- 32505 10 28 . . . 32505 11 1 The the DT 32505 11 2 group group NN 32505 11 3 apparently apparently RB 32505 11 4 originated originate VBD 32505 11 5 in in IN 32505 11 6 South South NNP 32505 11 7 America America NNP 32505 11 8 ; ; : 32505 11 9 the the DT 32505 11 10 endemic endemic JJ 32505 11 11 Central central JJ 32505 11 12 American american JJ 32505 11 13 species specie NNS 32505 11 14 evolved evolve VBN 32505 11 15 from from IN 32505 11 16 stocks stock NNS 32505 11 17 that that WDT 32505 11 18 invaded invade VBD 32505 11 19 Middle Middle NNP 32505 11 20 America America NNP 32505 11 21 after after IN 32505 11 22 the the DT 32505 11 23 closure closure NN 32505 11 24 of of IN 32505 11 25 the the DT 32505 11 26 Colombian Colombian NNP 32505 11 27 Portal Portal NNP 32505 11 28 in in IN 32505 11 29 the the DT 32505 11 30 late late JJ 32505 11 31 Pliocene Pliocene NNP 32505 11 32 . . . 32505 12 1 Dunn Dunn NNP 32505 12 2 ( ( -LRB- 32505 12 3 1933 1933 CD 32505 12 4 ) ) -RRB- 32505 12 5 partially partially RB 32505 12 6 defined define VBD 32505 12 7 the the DT 32505 12 8 _ _ NNP 32505 12 9 rubra rubra NNP 32505 12 10 _ _ NNP 32505 12 11 group group NN 32505 12 12 as as IN 32505 12 13 it -PRON- PRP 32505 12 14 occurs occur VBZ 32505 12 15 in in IN 32505 12 16 Central Central NNP 32505 12 17 America America NNP 32505 12 18 . . . 32505 13 1 Cope Cope NNP 32505 13 2 ( ( -LRB- 32505 13 3 1865 1865 CD 32505 13 4 , , , 32505 13 5 1876 1876 CD 32505 13 6 , , , 32505 13 7 1887 1887 CD 32505 13 8 ) ) -RRB- 32505 13 9 , , , 32505 13 10 Brocchi Brocchi NNP 32505 13 11 ( ( -LRB- 32505 13 12 1881 1881 CD 32505 13 13 ) ) -RRB- 32505 13 14 , , , 32505 13 15 Boulenger Boulenger NNP 32505 13 16 ( ( -LRB- 32505 13 17 1882 1882 CD 32505 13 18 ) ) -RRB- 32505 13 19 , , , 32505 13 20 Günther Günther NNP 32505 13 21 ( ( -LRB- 32505 13 22 1901 1901 CD 32505 13 23 ) ) -RRB- 32505 13 24 , , , 32505 13 25 Noble Noble NNP 32505 13 26 ( ( -LRB- 32505 13 27 1918 1918 CD 32505 13 28 ) ) -RRB- 32505 13 29 , , , 32505 13 30 Kellogg Kellogg NNP 32505 13 31 ( ( -LRB- 32505 13 32 1932 1932 CD 32505 13 33 ) ) -RRB- 32505 13 34 , , , 32505 13 35 Dunn Dunn NNP 32505 13 36 and and CC 32505 13 37 Emlen Emlen NNP 32505 13 38 ( ( -LRB- 32505 13 39 1932 1932 CD 32505 13 40 ) ) -RRB- 32505 13 41 , , , 32505 13 42 Stuart Stuart NNP 32505 13 43 ( ( -LRB- 32505 13 44 1935 1935 CD 32505 13 45 ) ) -RRB- 32505 13 46 , , , 32505 13 47 and and CC 32505 13 48 Gaige Gaige NNP 32505 13 49 ( ( -LRB- 32505 13 50 1936 1936 CD 32505 13 51 ) ) -RRB- 32505 13 52 dealt deal VBD 32505 13 53 with with IN 32505 13 54 the the DT 32505 13 55 Middle middle JJ 32505 13 56 American american JJ 32505 13 57 species specie NNS 32505 13 58 now now RB 32505 13 59 considered consider VBN 32505 13 60 to to TO 32505 13 61 make make VB 32505 13 62 up up RP 32505 13 63 the the DT 32505 13 64 _ _ NNP 32505 13 65 rubra rubra NNP 32505 13 66 _ _ NNP 32505 13 67 group group NN 32505 13 68 . . . 32505 14 1 More more RBR 32505 14 2 recently recently RB 32505 14 3 , , , 32505 14 4 Taylor Taylor NNP 32505 14 5 ( ( -LRB- 32505 14 6 1952 1952 CD 32505 14 7 , , , 32505 14 8 1958 1958 CD 32505 14 9 ) ) -RRB- 32505 14 10 , , , 32505 14 11 Fouquette Fouquette NNP 32505 14 12 ( ( -LRB- 32505 14 13 1958 1958 CD 32505 14 14 ) ) -RRB- 32505 14 15 , , , 32505 14 16 Starrett Starrett NNP 32505 14 17 ( ( -LRB- 32505 14 18 1960 1960 CD 32505 14 19 ) ) -RRB- 32505 14 20 , , , 32505 14 21 and and CC 32505 14 22 Duellman Duellman NNP 32505 14 23 ( ( -LRB- 32505 14 24 1960 1960 CD 32505 14 25 , , , 32505 14 26 1963 1963 CD 32505 14 27 , , , 32505 14 28 1966a 1966a CD 32505 14 29 ) ) -RRB- 32505 14 30 studied study VBN 32505 14 31 aspects aspect NNS 32505 14 32 of of IN 32505 14 33 the the DT 32505 14 34 taxonomy taxonomy NN 32505 14 35 and and CC 32505 14 36 biology biology NN 32505 14 37 of of IN 32505 14 38 the the DT 32505 14 39 species specie NNS 32505 14 40 of of IN 32505 14 41 this this DT 32505 14 42 group group NN 32505 14 43 . . . 32505 15 1 The the DT 32505 15 2 five five CD 32505 15 3 species specie NNS 32505 15 4 of of IN 32505 15 5 the the DT 32505 15 6 _ _ NNP 32505 15 7 rubra rubra NNP 32505 15 8 _ _ NNP 32505 15 9 group group NN 32505 15 10 in in IN 32505 15 11 Central Central NNP 32505 15 12 America America NNP 32505 15 13 have have VBP 32505 15 14 received receive VBN 32505 15 15 ten ten CD 32505 15 16 different different JJ 32505 15 17 names name NNS 32505 15 18 . . . 32505 16 1 One one CD 32505 16 2 species species NN 32505 16 3 , , , 32505 16 4 _ _ NNP 32505 16 5 Hyla Hyla NNP 32505 16 6 staufferi staufferi NN 32505 16 7 _ _ NNP 32505 16 8 , , , 32505 16 9 has have VBZ 32505 16 10 received receive VBN 32505 16 11 five five CD 32505 16 12 names name NNS 32505 16 13 ( ( -LRB- 32505 16 14 two two CD 32505 16 15 subspecies subspecie NNS 32505 16 16 are be VBP 32505 16 17 recognized recognize VBN 32505 16 18 herein herein NNP 32505 16 19 ) ) -RRB- 32505 16 20 . . . 32505 17 1 _ _ NNP 32505 17 2 Hyla Hyla NNP 32505 17 3 boulengeri boulengeri NN 32505 17 4 _ _ NNP 32505 17 5 was be VBD 32505 17 6 named name VBN 32505 17 7 in in IN 32505 17 8 the the DT 32505 17 9 genus genus NN 32505 17 10 _ _ NNP 32505 17 11 Scytopis Scytopis NNP 32505 17 12 _ _ NNP 32505 17 13 , , , 32505 17 14 but but CC 32505 17 15 the the DT 32505 17 16 type type NN 32505 17 17 species species NN 32505 17 18 of of IN 32505 17 19 _ _ NNP 32505 17 20 Scytopis Scytopis NNP 32505 17 21 _ _ NNP 32505 17 22 is be VBZ 32505 17 23 a a DT 32505 17 24 member member NN 32505 17 25 of of IN 32505 17 26 the the DT 32505 17 27 genus genus NNP 32505 17 28 _ _ NNP 32505 17 29 Phrynohyas Phrynohyas NNP 32505 17 30 _ _ NNP 32505 17 31 Fitzinger Fitzinger NNP 32505 17 32 , , , 32505 17 33 1843 1843 CD 32505 17 34 ( ( -LRB- 32505 17 35 Duellman Duellman NNP 32505 17 36 , , , 32505 17 37 1956 1956 CD 32505 17 38 . . . 32505 17 39 ) ) -RRB- 32505 18 1 Little Little NNP 32505 18 2 has have VBZ 32505 18 3 been be VBN 32505 18 4 published publish VBN 32505 18 5 concerning concern VBG 32505 18 6 the the DT 32505 18 7 ecology ecology NN 32505 18 8 , , , 32505 18 9 life life NN 32505 18 10 history history NN 32505 18 11 , , , 32505 18 12 osteology osteology NN 32505 18 13 , , , 32505 18 14 and and CC 32505 18 15 mating mating NN 32505 18 16 calls call NNS 32505 18 17 of of IN 32505 18 18 the the DT 32505 18 19 Middle middle JJ 32505 18 20 American american JJ 32505 18 21 species specie NNS 32505 18 22 of of IN 32505 18 23 this this DT 32505 18 24 group group NN 32505 18 25 . . . 32505 19 1 The the DT 32505 19 2 purpose purpose NN 32505 19 3 of of IN 32505 19 4 the the DT 32505 19 5 present present JJ 32505 19 6 report report NN 32505 19 7 is be VBZ 32505 19 8 to to TO 32505 19 9 describe describe VB 32505 19 10 the the DT 32505 19 11 species specie NNS 32505 19 12 occurring occur VBG 32505 19 13 in in IN 32505 19 14 Middle Middle NNP 32505 19 15 America America NNP 32505 19 16 and and CC 32505 19 17 to to TO 32505 19 18 comment comment VB 32505 19 19 on on IN 32505 19 20 their -PRON- PRP$ 32505 19 21 distributions distribution NNS 32505 19 22 , , , 32505 19 23 ecology ecology NN 32505 19 24 , , , 32505 19 25 cranial cranial JJ 32505 19 26 osteology osteology NN 32505 19 27 , , , 32505 19 28 and and CC 32505 19 29 mating mating NN 32505 19 30 calls call NNS 32505 19 31 , , , 32505 19 32 and and CC 32505 19 33 in in IN 32505 19 34 so so RB 32505 19 35 doing do VBG 32505 19 36 provide provide VB 32505 19 37 evidence evidence NN 32505 19 38 for for IN 32505 19 39 the the DT 32505 19 40 evolutionary evolutionary JJ 32505 19 41 history history NN 32505 19 42 of of IN 32505 19 43 the the DT 32505 19 44 species specie NNS 32505 19 45 inhabiting inhabit VBG 32505 19 46 Middle Middle NNP 32505 19 47 America America NNP 32505 19 48 . . . 32505 20 1 Acknowledgments acknowledgment NNS 32505 20 2 For for IN 32505 20 3 permission permission NN 32505 20 4 to to TO 32505 20 5 examine examine VB 32505 20 6 specimens specimen NNS 32505 20 7 in in IN 32505 20 8 their -PRON- PRP$ 32505 20 9 care care NN 32505 20 10 , , , 32505 20 11 I -PRON- PRP 32505 20 12 am be VBP 32505 20 13 grateful grateful JJ 32505 20 14 to to IN 32505 20 15 Drs Drs NNP 32505 20 16 . . . 32505 21 1 Richard Richard NNP 32505 21 2 G. G. NNP 32505 21 3 Zweifel Zweifel NNP 32505 21 4 , , , 32505 21 5 American American NNP 32505 21 6 Museum Museum NNP 32505 21 7 of of IN 32505 21 8 Natural Natural NNP 32505 21 9 History History NNP 32505 21 10 ( ( -LRB- 32505 21 11 AMNH AMNH NNP 32505 21 12 ) ) -RRB- 32505 21 13 ; ; : 32505 21 14 Robert Robert NNP 32505 21 15 F. F. NNP 32505 21 16 Inger Inger NNP 32505 21 17 , , , 32505 21 18 Field Field NNP 32505 21 19 Museum Museum NNP 32505 21 20 of of IN 32505 21 21 Natural Natural NNP 32505 21 22 History History NNP 32505 21 23 ( ( -LRB- 32505 21 24 FMNH FMNH NNP 32505 21 25 ) ) -RRB- 32505 21 26 ; ; : 32505 21 27 Ernest Ernest NNP 32505 21 28 E. E. NNP 32505 21 29 Williams Williams NNP 32505 21 30 , , , 32505 21 31 Museum Museum NNP 32505 21 32 of of IN 32505 21 33 Comparative Comparative NNP 32505 21 34 Zoology Zoology NNP 32505 21 35 ( ( -LRB- 32505 21 36 MCZ MCZ NNP 32505 21 37 ) ) -RRB- 32505 21 38 ; ; : 32505 21 39 Hobart Hobart NNP 32505 21 40 M. M. NNP 32505 21 41 Smith Smith NNP 32505 21 42 , , , 32505 21 43 University University NNP 32505 21 44 of of IN 32505 21 45 Illinois Illinois NNP 32505 21 46 Museum Museum NNP 32505 21 47 of of IN 32505 21 48 Natural Natural NNP 32505 21 49 History History NNP 32505 21 50 ( ( -LRB- 32505 21 51 UIMNH UIMNH NNP 32505 21 52 ) ) -RRB- 32505 21 53 ; ; : 32505 21 54 Charles Charles NNP 32505 21 55 F. F. NNP 32505 21 56 Walker Walker NNP 32505 21 57 , , , 32505 21 58 University University NNP 32505 21 59 of of IN 32505 21 60 Michigan Michigan NNP 32505 21 61 Museum Museum NNP 32505 21 62 of of IN 32505 21 63 Zoology Zoology NNP 32505 21 64 ( ( -LRB- 32505 21 65 UMMZ UMMZ NNP 32505 21 66 ) ) -RRB- 32505 21 67 ; ; : 32505 21 68 Jay Jay NNP 32505 21 69 M. M. NNP 32505 21 70 Savage Savage NNP 32505 21 71 , , , 32505 21 72 University University NNP 32505 21 73 of of IN 32505 21 74 Southern Southern NNP 32505 21 75 California California NNP 32505 21 76 ( ( -LRB- 32505 21 77 USC USC NNP 32505 21 78 ) ) -RRB- 32505 21 79 ; ; : 32505 21 80 James James NNP 32505 21 81 A. A. NNP 32505 21 82 Peters Peters NNP 32505 21 83 , , , 32505 21 84 United United NNP 32505 21 85 States States NNP 32505 21 86 National National NNP 32505 21 87 Museum Museum NNP 32505 21 88 ( ( -LRB- 32505 21 89 USNM USNM NNP 32505 21 90 ) ) -RRB- 32505 21 91 ; ; : 32505 21 92 Richard Richard NNP 32505 21 93 J. J. NNP 32505 21 94 Baldauf Baldauf NNP 32505 21 95 , , , 32505 21 96 Texas Texas NNP 32505 21 97 Cooperative Cooperative NNP 32505 21 98 Wildlife Wildlife NNP 32505 21 99 Collection Collection NNP 32505 21 100 ( ( -LRB- 32505 21 101 TCWC TCWC NNP 32505 21 102 ) ) -RRB- 32505 21 103 ; ; : 32505 21 104 and and CC 32505 21 105 W. W. NNP 32505 21 106 Frank Frank NNP 32505 21 107 Blair Blair NNP 32505 21 108 , , , 32505 21 109 Texas Texas NNP 32505 21 110 Natural Natural NNP 32505 21 111 History History NNP 32505 21 112 Collection Collection NNP 32505 21 113 ( ( -LRB- 32505 21 114 TNHC TNHC NNP 32505 21 115 ) ) -RRB- 32505 21 116 . . . 32505 22 1 KU KU NNP 32505 22 2 refers refer VBZ 32505 22 3 to to IN 32505 22 4 specimens specimen NNS 32505 22 5 in in IN 32505 22 6 the the DT 32505 22 7 Museum Museum NNP 32505 22 8 of of IN 32505 22 9 Natural Natural NNP 32505 22 10 History History NNP 32505 22 11 , , , 32505 22 12 University University NNP 32505 22 13 of of IN 32505 22 14 Kansas Kansas NNP 32505 22 15 . . . 32505 23 1 For for IN 32505 23 2 the the DT 32505 23 3 loan loan NN 32505 23 4 of of IN 32505 23 5 tape tape NN 32505 23 6 - - HYPH 32505 23 7 recordings recording NNS 32505 23 8 of of IN 32505 23 9 mating mating NN 32505 23 10 calls call NNS 32505 23 11 I -PRON- PRP 32505 23 12 thank thank VBP 32505 23 13 Drs Drs NNP 32505 23 14 . . . 32505 24 1 W. W. NNP 32505 24 2 Frank Frank NNP 32505 24 3 Blair Blair NNP 32505 24 4 , , , 32505 24 5 University University NNP 32505 24 6 of of IN 32505 24 7 Texas Texas NNP 32505 24 8 , , , 32505 24 9 and and CC 32505 24 10 Richard Richard NNP 32505 24 11 G. G. NNP 32505 24 12 Zweifel Zweifel NNP 32505 24 13 , , , 32505 24 14 American American NNP 32505 24 15 Museum Museum NNP 32505 24 16 of of IN 32505 24 17 Natural Natural NNP 32505 24 18 History History NNP 32505 24 19 . . . 32505 25 1 I -PRON- PRP 32505 25 2 am be VBP 32505 25 3 indebted indebted JJ 32505 25 4 to to IN 32505 25 5 the the DT 32505 25 6 Ford Ford NNP 32505 25 7 Foundation Foundation NNP 32505 25 8 - - HYPH 32505 25 9 Universidad Universidad NNP 32505 25 10 de de NNP 32505 25 11 Oriente Oriente NNP 32505 25 12 ( ( -LRB- 32505 25 13 Venezuela Venezuela NNP 32505 25 14 ) ) -RRB- 32505 25 15 Science Science NNP 32505 25 16 Project Project NNP 32505 25 17 for for IN 32505 25 18 a a DT 32505 25 19 scholarship scholarship NN 32505 25 20 which which WDT 32505 25 21 enabled enable VBD 32505 25 22 me -PRON- PRP 32505 25 23 to to TO 32505 25 24 study study VB 32505 25 25 for for IN 32505 25 26 two two CD 32505 25 27 years year NNS 32505 25 28 at at IN 32505 25 29 The the DT 32505 25 30 University University NNP 32505 25 31 of of IN 32505 25 32 Kansas Kansas NNP 32505 25 33 , , , 32505 25 34 foster foster JJ 32505 25 35 institution institution NN 32505 25 36 of of IN 32505 25 37 the the DT 32505 25 38 project project NN 32505 25 39 . . . 32505 26 1 I -PRON- PRP 32505 26 2 have have VBP 32505 26 3 benefited benefit VBN 32505 26 4 by by IN 32505 26 5 being be VBG 32505 26 6 able able JJ 32505 26 7 to to TO 32505 26 8 work work VB 32505 26 9 in in IN 32505 26 10 the the DT 32505 26 11 Museum Museum NNP 32505 26 12 of of IN 32505 26 13 Natural Natural NNP 32505 26 14 History History NNP 32505 26 15 at at IN 32505 26 16 The the DT 32505 26 17 University University NNP 32505 26 18 of of IN 32505 26 19 Kansas Kansas NNP 32505 26 20 and and CC 32505 26 21 I -PRON- PRP 32505 26 22 am be VBP 32505 26 23 grateful grateful JJ 32505 26 24 to to IN 32505 26 25 Dr. Dr. NNP 32505 26 26 E. E. NNP 32505 26 27 Raymond Raymond NNP 32505 26 28 Hall Hall NNP 32505 26 29 , , , 32505 26 30 Director Director NNP 32505 26 31 , , , 32505 26 32 for for IN 32505 26 33 providing provide VBG 32505 26 34 space space NN 32505 26 35 and and CC 32505 26 36 equipment equipment NN 32505 26 37 . . . 32505 27 1 I -PRON- PRP 32505 27 2 gratefully gratefully RB 32505 27 3 acknowledge acknowledge VBP 32505 27 4 the the DT 32505 27 5 assistance assistance NN 32505 27 6 and and CC 32505 27 7 advice advice NN 32505 27 8 of of IN 32505 27 9 Dr. Dr. NNP 32505 27 10 William William NNP 32505 27 11 E. E. NNP 32505 27 12 Duellman Duellman NNP 32505 27 13 , , , 32505 27 14 who who WP 32505 27 15 suggested suggest VBD 32505 27 16 and and CC 32505 27 17 directed direct VBD 32505 27 18 this this DT 32505 27 19 work work NN 32505 27 20 , , , 32505 27 21 made make VBN 32505 27 22 available available JJ 32505 27 23 specimens specimen NNS 32505 27 24 under under IN 32505 27 25 his -PRON- PRP$ 32505 27 26 care care NN 32505 27 27 and and CC 32505 27 28 gave give VBD 32505 27 29 much much JJ 32505 27 30 of of IN 32505 27 31 his -PRON- PRP$ 32505 27 32 time time NN 32505 27 33 in in IN 32505 27 34 reading read VBG 32505 27 35 the the DT 32505 27 36 manuscript manuscript NN 32505 27 37 and and CC 32505 27 38 suggesting suggest VBG 32505 27 39 improvements improvement NNS 32505 27 40 . . . 32505 28 1 I -PRON- PRP 32505 28 2 am be VBP 32505 28 3 grateful grateful JJ 32505 28 4 to to IN 32505 28 5 Dr. Dr. NNP 32505 28 6 Frank Frank NNP 32505 28 7 B. B. NNP 32505 29 1 Cross cross NN 32505 29 2 who who WP 32505 29 3 critically critically RB 32505 29 4 read read VBD 32505 29 5 the the DT 32505 29 6 manuscript manuscript NN 32505 29 7 and and CC 32505 29 8 made make VBD 32505 29 9 many many JJ 32505 29 10 editorial editorial JJ 32505 29 11 suggestions suggestion NNS 32505 29 12 . . . 32505 30 1 I -PRON- PRP 32505 30 2 am be VBP 32505 30 3 indebted indebted JJ 32505 30 4 to to IN 32505 30 5 Linda Linda NNP 32505 30 6 Trueb Trueb NNP 32505 30 7 for for IN 32505 30 8 assistance assistance NN 32505 30 9 with with IN 32505 30 10 the the DT 32505 30 11 osteological osteological JJ 32505 30 12 aspects aspect NNS 32505 30 13 of of IN 32505 30 14 this this DT 32505 30 15 study study NN 32505 30 16 ; ; : 32505 30 17 she -PRON- PRP 32505 30 18 helped help VBD 32505 30 19 to to TO 32505 30 20 clarify clarify VB 32505 30 21 many many JJ 32505 30 22 confusing confusing JJ 32505 30 23 points point NNS 32505 30 24 . . . 32505 31 1 I -PRON- PRP 32505 31 2 am be VBP 32505 31 3 grateful grateful JJ 32505 31 4 to to IN 32505 31 5 Charles Charles NNP 32505 31 6 W. W. NNP 32505 31 7 Myers Myers NNP 32505 31 8 for for IN 32505 31 9 making make VBG 32505 31 10 available available JJ 32505 31 11 his -PRON- PRP$ 32505 31 12 field field NN 32505 31 13 notes note NNS 32505 31 14 on on IN 32505 31 15 these these DT 32505 31 16 frogs frog NNS 32505 31 17 in in IN 32505 31 18 Panamá Panamá NNP 32505 31 19 , , , 32505 31 20 to to IN 32505 31 21 Arthur Arthur NNP 32505 31 22 C. C. NNP 32505 31 23 Echternacht Echternacht NNP 32505 31 24 for for IN 32505 31 25 reading read VBG 32505 31 26 part part NN 32505 31 27 of of IN 32505 31 28 the the DT 32505 31 29 manuscript manuscript NN 32505 31 30 , , , 32505 31 31 and and CC 32505 31 32 to to IN 32505 31 33 John John NNP 32505 31 34 D. D. NNP 32505 31 35 Lynch Lynch NNP 32505 31 36 for for IN 32505 31 37 many many JJ 32505 31 38 suggestions suggestion NNS 32505 31 39 and and CC 32505 31 40 helpful helpful JJ 32505 31 41 criticisms criticism NNS 32505 31 42 . . . 32505 32 1 The the DT 32505 32 2 illustrations illustration NNS 32505 32 3 were be VBD 32505 32 4 executed execute VBN 32505 32 5 by by IN 32505 32 6 David David NNP 32505 32 7 M. M. NNP 32505 32 8 Dennis Dennis NNP 32505 32 9 . . . 32505 33 1 Materials material NNS 32505 33 2 and and CC 32505 33 3 Methods method NNS 32505 33 4 For for IN 32505 33 5 the the DT 32505 33 6 purposes purpose NNS 32505 33 7 of of IN 32505 33 8 the the DT 32505 33 9 present present JJ 32505 33 10 study study NN 32505 33 11 I -PRON- PRP 32505 33 12 examined examine VBD 32505 33 13 1383 1383 CD 32505 33 14 preserved preserve VBN 32505 33 15 specimens specimen NNS 32505 33 16 , , , 32505 33 17 50 50 CD 32505 33 18 skeletons skeleton NNS 32505 33 19 , , , 32505 33 20 and and CC 32505 33 21 9 9 CD 32505 33 22 lots lot NNS 32505 33 23 of of IN 32505 33 24 tadpoles tadpole NNS 32505 33 25 . . . 32505 34 1 External external JJ 32505 34 2 characteristics characteristic NNS 32505 34 3 used use VBN 32505 34 4 in in IN 32505 34 5 the the DT 32505 34 6 analysis analysis NN 32505 34 7 of of IN 32505 34 8 variation variation NN 32505 34 9 are be VBP 32505 34 10 those those DT 32505 34 11 currently currently RB 32505 34 12 employed employ VBN 32505 34 13 in in IN 32505 34 14 the the DT 32505 34 15 study study NN 32505 34 16 of of IN 32505 34 17 anuran anuran NNP 32505 34 18 systematics systematic NNS 32505 34 19 . . . 32505 35 1 Twelve twelve CD 32505 35 2 measurements measurement NNS 32505 35 3 and and CC 32505 35 4 six six CD 32505 35 5 proportions proportion NNS 32505 35 6 were be VBD 32505 35 7 taken take VBN 32505 35 8 in in IN 32505 35 9 the the DT 32505 35 10 manner manner NN 32505 35 11 described describe VBN 32505 35 12 by by IN 32505 35 13 Duellman Duellman NNP 32505 35 14 ( ( -LRB- 32505 35 15 1956 1956 CD 32505 35 16 ) ) -RRB- 32505 35 17 . . . 32505 36 1 Only only RB 32505 36 2 the the DT 32505 36 3 most most RBS 32505 36 4 important important JJ 32505 36 5 references reference NNS 32505 36 6 are be VBP 32505 36 7 given give VBN 32505 36 8 in in IN 32505 36 9 the the DT 32505 36 10 synonymies synonymy NNS 32505 36 11 , , , 32505 36 12 except except IN 32505 36 13 those those DT 32505 36 14 of of IN 32505 36 15 the the DT 32505 36 16 two two CD 32505 36 17 subspecies subspecie NNS 32505 36 18 of of IN 32505 36 19 _ _ NNP 32505 36 20 Hyla Hyla NNP 32505 36 21 staufferi staufferi NN 32505 36 22 , , , 32505 36 23 _ _ NNP 32505 36 24 which which WDT 32505 36 25 are be VBP 32505 36 26 more more RBR 32505 36 27 nearly nearly RB 32505 36 28 complete complete JJ 32505 36 29 . . . 32505 37 1 The the DT 32505 37 2 taxonomic taxonomic JJ 32505 37 3 history history NN 32505 37 4 of of IN 32505 37 5 each each DT 32505 37 6 frog frog NN 32505 37 7 is be VBZ 32505 37 8 discussed discuss VBN 32505 37 9 under under IN 32505 37 10 _ _ NNP 32505 37 11 Remarks Remarks NNP 32505 37 12 _ _ NNP 32505 37 13 in in IN 32505 37 14 each each DT 32505 37 15 account account NN 32505 37 16 . . . 32505 38 1 The the DT 32505 38 2 cranial cranial JJ 32505 38 3 osteology osteology NN 32505 38 4 was be VBD 32505 38 5 studied study VBN 32505 38 6 by by IN 32505 38 7 using use VBG 32505 38 8 skeletons skeleton NNS 32505 38 9 and and CC 32505 38 10 cleared clear VBD 32505 38 11 and and CC 32505 38 12 stained stain VBN 32505 38 13 specimens specimen NNS 32505 38 14 of of IN 32505 38 15 all all DT 32505 38 16 species specie NNS 32505 38 17 . . . 32505 39 1 Developmental developmental JJ 32505 39 2 stages stage NNS 32505 39 3 of of IN 32505 39 4 tadpoles tadpole NNS 32505 39 5 were be VBD 32505 39 6 determined determine VBN 32505 39 7 from from IN 32505 39 8 Gosner Gosner NNP 32505 39 9 's 's POS 32505 39 10 ( ( -LRB- 32505 39 11 1960 1960 CD 32505 39 12 ) ) -RRB- 32505 39 13 table table NN 32505 39 14 . . . 32505 40 1 Personal personal JJ 32505 40 2 field field NN 32505 40 3 work work NN 32505 40 4 in in IN 32505 40 5 Central Central NNP 32505 40 6 America America NNP 32505 40 7 in in IN 32505 40 8 the the DT 32505 40 9 summer summer NN 32505 40 10 of of IN 32505 40 11 1966 1966 CD 32505 40 12 provided provide VBD 32505 40 13 an an DT 32505 40 14 opportunity opportunity NN 32505 40 15 to to TO 32505 40 16 make make VB 32505 40 17 observations observation NNS 32505 40 18 on on IN 32505 40 19 the the DT 32505 40 20 ecology ecology NN 32505 40 21 , , , 32505 40 22 calling call VBG 32505 40 23 sites site NNS 32505 40 24 , , , 32505 40 25 and and CC 32505 40 26 color color NN 32505 40 27 in in IN 32505 40 28 life life NN 32505 40 29 ; ; : 32505 40 30 these these DT 32505 40 31 data datum NNS 32505 40 32 were be VBD 32505 40 33 supplemented supplement VBN 32505 40 34 by by IN 32505 40 35 field field NN 32505 40 36 notes note NNS 32505 40 37 from from IN 32505 40 38 , , , 32505 40 39 and and CC 32505 40 40 discussions discussion NNS 32505 40 41 with with IN 32505 40 42 , , , 32505 40 43 Dr. Dr. NNP 32505 40 44 William William NNP 32505 40 45 E. E. NNP 32505 40 46 Duellman Duellman NNP 32505 40 47 and and CC 32505 40 48 Charles Charles NNP 32505 40 49 W. W. NNP 32505 40 50 Myers Myers NNP 32505 40 51 . . . 32505 41 1 The the DT 32505 41 2 mating mating NN 32505 41 3 calls call NNS 32505 41 4 of of IN 32505 41 5 the the DT 32505 41 6 frogs frog NNS 32505 41 7 were be VBD 32505 41 8 recorded record VBN 32505 41 9 in in IN 32505 41 10 the the DT 32505 41 11 field field NN 32505 41 12 on on IN 32505 41 13 Magnemite Magnemite NNP 32505 41 14 and and CC 32505 41 15 Uher Uher NNP 32505 41 16 Tape Tape NNP 32505 41 17 Recorders Recorders NNPS 32505 41 18 by by IN 32505 41 19 Dr. Dr. NNP 32505 41 20 Duellman Duellman NNP 32505 41 21 in in IN 32505 41 22 the the DT 32505 41 23 course course NN 32505 41 24 of of IN 32505 41 25 his -PRON- PRP$ 32505 41 26 work work NN 32505 41 27 on on IN 32505 41 28 the the DT 32505 41 29 hylid hylid JJ 32505 41 30 frogs frog NNS 32505 41 31 of of IN 32505 41 32 Middle Middle NNP 32505 41 33 America America NNP 32505 41 34 -- -- : 32505 41 35 supported support VBN 32505 41 36 by by IN 32505 41 37 grants grant NNS 32505 41 38 from from IN 32505 41 39 the the DT 32505 41 40 National National NNP 32505 41 41 Science Science NNP 32505 41 42 Foundation Foundation NNP 32505 41 43 ( ( -LRB- 32505 41 44 G-9827 g-9827 CD 32505 41 45 and and CC 32505 41 46 GB-1441 GB-1441 NNPS 32505 41 47 ) ) -RRB- 32505 41 48 . . . 32505 42 1 These these DT 32505 42 2 recordings recording NNS 32505 42 3 , , , 32505 42 4 plus plus CC 32505 42 5 those those DT 32505 42 6 borrowed borrow VBN 32505 42 7 from from IN 32505 42 8 other other JJ 32505 42 9 institutions institution NNS 32505 42 10 , , , 32505 42 11 provided provide VBN 32505 42 12 50 50 CD 32505 42 13 tapes tape NNS 32505 42 14 for for IN 32505 42 15 analysis analysis NN 32505 42 16 of of IN 32505 42 17 the the DT 32505 42 18 mating mating NN 32505 42 19 calls call NNS 32505 42 20 . . . 32505 43 1 The the DT 32505 43 2 calls call NNS 32505 43 3 were be VBD 32505 43 4 analyzed analyze VBN 32505 43 5 on on IN 32505 43 6 a a DT 32505 43 7 Vibralyzer Vibralyzer NNP 32505 43 8 ( ( -LRB- 32505 43 9 Kay Kay NNP 32505 43 10 Electric Electric NNP 32505 43 11 Company Company NNP 32505 43 12 ) ) -RRB- 32505 43 13 . . . 32505 44 1 THE the DT 32505 44 2 HYLA HYLA NNP 32505 44 3 RUBRA RUBRA NNP 32505 44 4 GROUP GROUP NNP 32505 44 5 _ _ NNP 32505 44 6 Definition._--The Definition._--The NNP 32505 44 7 species specie NNS 32505 44 8 forming form VBG 32505 44 9 the the DT 32505 44 10 group group NN 32505 44 11 are be VBP 32505 44 12 small small JJ 32505 44 13 to to TO 32505 44 14 moderate moderate JJ 32505 44 15 - - HYPH 32505 44 16 sized sized JJ 32505 44 17 tree tree NN 32505 44 18 frogs frog NNS 32505 44 19 ( ( -LRB- 32505 44 20 maximum maximum JJ 32505 44 21 snout snout NN 32505 44 22 - - HYPH 32505 44 23 vent vent NN 32505 44 24 length length NN 32505 44 25 of of IN 32505 44 26 males male NNS 32505 44 27 of of IN 32505 44 28 various various JJ 32505 44 29 species specie NNS 32505 44 30 20 20 CD 32505 44 31 - - SYM 32505 44 32 49 49 CD 32505 44 33 mm mm CD 32505 44 34 . . . 32505 45 1 ) ) -RRB- 32505 45 2 , , , 32505 45 3 distinguished distinguish VBN 32505 45 4 from from IN 32505 45 5 other other JJ 32505 45 6 groups group NNS 32505 45 7 in in IN 32505 45 8 the the DT 32505 45 9 genus genus NN 32505 45 10 _ _ NNP 32505 45 11 Hyla Hyla NNP 32505 45 12 _ _ NNP 32505 45 13 as as IN 32505 45 14 follows follow VBZ 32505 45 15 : : : 32505 45 16 Brown Brown NNP 32505 45 17 , , , 32505 45 18 grayish grayish NN 32505 45 19 brown brown NNP 32505 45 20 , , , 32505 45 21 or or CC 32505 45 22 yellowish yellowish JJ 32505 45 23 tan tan NN 32505 45 24 above above RB 32505 45 25 ; ; : 32505 45 26 thighs thigh NNS 32505 45 27 plain plain JJ 32505 45 28 , , , 32505 45 29 marbled marble VBN 32505 45 30 with with IN 32505 45 31 dark dark JJ 32505 45 32 brown brown NNP 32505 45 33 , , , 32505 45 34 or or CC 32505 45 35 having have VBG 32505 45 36 vertical vertical JJ 32505 45 37 bands band NNS 32505 45 38 ; ; : 32505 45 39 vocal vocal JJ 32505 45 40 sac sac NN 32505 45 41 single single JJ 32505 45 42 , , , 32505 45 43 median median JJ 32505 45 44 , , , 32505 45 45 subgular subgular JJ 32505 45 46 ; ; : 32505 45 47 snout snout NN 32505 45 48 flat flat JJ 32505 45 49 , , , 32505 45 50 protruding protrude VBG 32505 45 51 , , , 32505 45 52 rounded round VBD 32505 45 53 or or CC 32505 45 54 pointed point VBN 32505 45 55 ; ; : 32505 45 56 webbing web VBG 32505 45 57 between between IN 32505 45 58 fingers finger NNS 32505 45 59 reduced reduce VBN 32505 45 60 or or CC 32505 45 61 absent absent JJ 32505 45 62 ; ; : 32505 45 63 web web NN 32505 45 64 between between IN 32505 45 65 first first JJ 32505 45 66 and and CC 32505 45 67 second second JJ 32505 45 68 toes toe NNS 32505 45 69 reduced reduce VBN 32505 45 70 to to TO 32505 45 71 fringe fringe VB 32505 45 72 on on IN 32505 45 73 second second JJ 32505 45 74 toe toe NN 32505 45 75 , , , 32505 45 76 rest rest NN 32505 45 77 of of IN 32505 45 78 toes toe NNS 32505 45 79 about about IN 32505 45 80 half half RB 32505 45 81 webbed web VBN 32505 45 82 ; ; : 32505 45 83 tarsal tarsal JJ 32505 45 84 fold fold VB 32505 45 85 reduced reduced JJ 32505 45 86 or or CC 32505 45 87 absent absent JJ 32505 45 88 ; ; : 32505 45 89 shanks shank NNS 32505 45 90 robust robust JJ 32505 45 91 ; ; : 32505 45 92 inner inner JJ 32505 45 93 metatarsal metatarsal NN 32505 45 94 tubercle tubercle NN 32505 45 95 larger large JJR 32505 45 96 than than IN 32505 45 97 outer outer JJ 32505 45 98 ; ; , 32505 45 99 prevomerine prevomerine JJ 32505 45 100 teeth tooth NNS 32505 45 101 on on IN 32505 45 102 transverse transverse JJ 32505 45 103 ridges ridge NNS 32505 45 104 between between IN 32505 45 105 small small JJ 32505 45 106 to to IN 32505 45 107 large large JJ 32505 45 108 sized sized JJ 32505 45 109 choanae choanae NN 32505 45 110 ; ; : 32505 45 111 skull skull NNP 32505 45 112 generally generally RB 32505 45 113 longer long RBR 32505 45 114 than than IN 32505 45 115 wide wide JJ 32505 45 116 ; ; : 32505 45 117 nasals nasal NNS 32505 45 118 large large JJ 32505 45 119 ( ( -LRB- 32505 45 120 length length NN 32505 45 121 more more JJR 32505 45 122 than than IN 32505 45 123 40 40 CD 32505 45 124 per per NN 32505 45 125 cent cent NN 32505 45 126 total total JJ 32505 45 127 length length NN 32505 45 128 of of IN 32505 45 129 skull skull NN 32505 45 130 ) ) -RRB- 32505 45 131 and and CC 32505 45 132 having have VBG 32505 45 133 pointed point VBN 32505 45 134 maxillary maxillary JJ 32505 45 135 processes process NNS 32505 45 136 ; ; , 32505 45 137 maxillary maxillary JJ 32505 45 138 bearing bear VBG 32505 45 139 small small JJ 32505 45 140 ventromedial ventromedial JJ 32505 45 141 palatine palatine NN 32505 45 142 process process NN 32505 45 143 ; ; : 32505 45 144 quadratojugal quadratojugal NNP 32505 45 145 slender slender NN 32505 45 146 , , , 32505 45 147 always always RB 32505 45 148 joined join VBD 32505 45 149 to to IN 32505 45 150 maxillary maxillary RB 32505 45 151 by by IN 32505 45 152 bony bony NN 32505 45 153 suture suture NN 32505 45 154 ; ; , 32505 45 155 auditory auditory JJ 32505 45 156 region region NN 32505 45 157 of of IN 32505 45 158 proötic proötic JJ 32505 45 159 slender slender NN 32505 45 160 and and CC 32505 45 161 short short JJ 32505 45 162 ; ; : 32505 45 163 delicate delicate JJ 32505 45 164 spatulate spatulate JJ 32505 45 165 columella columella NN 32505 45 166 ventral ventral JJ 32505 45 167 to to TO 32505 45 168 crista crista NNP 32505 45 169 parotica parotica NNP 32505 45 170 , , , 32505 45 171 broad broad JJ 32505 45 172 basally basally RB 32505 45 173 , , , 32505 45 174 compressed compress VBN 32505 45 175 anterolaterally anterolaterally RB 32505 45 176 , , , 32505 45 177 slightly slightly RB 32505 45 178 rounded round VBN 32505 45 179 distally distally RB 32505 45 180 ; ; , 32505 45 181 anterior anterior JJ 32505 45 182 arm arm NN 32505 45 183 of of IN 32505 45 184 squamosal squamosal NN 32505 45 185 extending extend VBG 32505 45 186 about about IN 32505 45 187 half half JJ 32505 45 188 distance distance NN 32505 45 189 to to IN 32505 45 190 maxillary maxillary NNP 32505 45 191 ; ; : 32505 45 192 sphenethmoid sphenethmoid LS 32505 45 193 wider wide JJR 32505 45 194 than than IN 32505 45 195 long long JJ 32505 45 196 ; ; : 32505 45 197 frontoparietal frontoparietal JJ 32505 45 198 fontanelle fontanelle NN 32505 45 199 present present JJ 32505 45 200 or or CC 32505 45 201 absent absent JJ 32505 45 202 ; ; : 32505 45 203 prevomerine prevomerine NN 32505 45 204 , , , 32505 45 205 premaxillary premaxillary NN 32505 45 206 , , , 32505 45 207 and and CC 32505 45 208 maxillary maxillary JJ 32505 45 209 teeth tooth NNS 32505 45 210 present present JJ 32505 45 211 ; ; : 32505 45 212 prevomer prevomer NN 32505 45 213 with with IN 32505 45 214 two two CD 32505 45 215 lateral lateral JJ 32505 45 216 processes process NNS 32505 45 217 forming form VBG 32505 45 218 incomplete incomplete JJ 32505 45 219 bony bony NN 32505 45 220 margin margin NN 32505 45 221 to to IN 32505 45 222 internal internal JJ 32505 45 223 nares naris NNS 32505 45 224 ; ; : 32505 45 225 tadpoles tadpole NNS 32505 45 226 having have VBG 32505 45 227 pointed point VBN 32505 45 228 xiphicercal xiphicercal JJ 32505 45 229 tail tail NN 32505 45 230 , , , 32505 45 231 snout snout VBP 32505 45 232 short short JJ 32505 45 233 , , , 32505 45 234 rounded rounded JJ 32505 45 235 ; ; : 32505 45 236 2/3 2/3 CD 32505 45 237 tooth tooth NN 32505 45 238 rows row VBZ 32505 45 239 ; ; : 32505 45 240 dorsal dorsal NN 32505 45 241 fin fin NN 32505 45 242 deeper deep JJR 32505 45 243 than than IN 32505 45 244 ventral ventral JJ 32505 45 245 fin fin NN 32505 45 246 ; ; , 32505 45 247 sinistral sinistral JJ 32505 45 248 spiracle spiracle NN 32505 45 249 ; ; : 32505 45 250 short short JJ 32505 45 251 dextral dextral JJ 32505 45 252 anal anal JJ 32505 45 253 tube tube NN 32505 45 254 not not RB 32505 45 255 reaching reach VBG 32505 45 256 edge edge NN 32505 45 257 of of IN 32505 45 258 ventral ventral JJ 32505 45 259 fin fin NN 32505 45 260 ; ; : 32505 45 261 mating mating NN 32505 45 262 calls call NNS 32505 45 263 consisting consist VBG 32505 45 264 of of IN 32505 45 265 single single JJ 32505 45 266 long long JJ 32505 45 267 note note NN 32505 45 268 or or CC 32505 45 269 series series NN 32505 45 270 of of IN 32505 45 271 short short JJ 32505 45 272 notes note NNS 32505 45 273 . . . 32505 46 1 _ _ NNP 32505 46 2 Composition._--This composition._--this DT 32505 46 3 group group NN 32505 46 4 contains contain VBZ 32505 46 5 about about RB 32505 46 6 24 24 CD 32505 46 7 currently currently RB 32505 46 8 recognized recognize VBN 32505 46 9 species specie NNS 32505 46 10 , , , 32505 46 11 most most JJS 32505 46 12 of of IN 32505 46 13 which which WDT 32505 46 14 occur occur VBP 32505 46 15 in in IN 32505 46 16 Brazil Brazil NNP 32505 46 17 . . . 32505 47 1 Only only RB 32505 47 2 five five CD 32505 47 3 species--_boulengeri species--_boulengeri CD 32505 47 4 , , , 32505 47 5 _ _ NNP 32505 47 6 _ _ NNP 32505 47 7 elaeochroa elaeochroa NNP 32505 47 8 _ _ NNP 32505 47 9 , , , 32505 47 10 _ _ NNP 32505 47 11 foliamorta foliamorta NNP 32505 47 12 _ _ NNP 32505 47 13 , , , 32505 47 14 _ _ NNP 32505 47 15 rubra rubra NNP 32505 47 16 _ _ NNP 32505 47 17 , , , 32505 47 18 and and CC 32505 47 19 _ _ NNP 32505 47 20 staufferi staufferi NN 32505 47 21 _ _ NNP 32505 47 22 with with IN 32505 47 23 two two CD 32505 47 24 subspecies subspecie NNS 32505 47 25 -- -- : 32505 47 26 occur occur VBP 32505 47 27 in in IN 32505 47 28 Central Central NNP 32505 47 29 America America NNP 32505 47 30 . . . 32505 48 1 _ _ NNP 32505 48 2 Hyla Hyla NNP 32505 48 3 boulengeri boulengeri NN 32505 48 4 _ _ NNP 32505 48 5 and and CC 32505 48 6 _ _ NNP 32505 48 7 rubra rubra NNP 32505 48 8 _ _ NNP 32505 48 9 are be VBP 32505 48 10 widespread widespread JJ 32505 48 11 in in IN 32505 48 12 South South NNP 32505 48 13 America America NNP 32505 48 14 , , , 32505 48 15 and and CC 32505 48 16 _ _ NNP 32505 48 17 foliamorta foliamorta NNP 32505 48 18 _ _ NNP 32505 48 19 occurs occur VBZ 32505 48 20 in in IN 32505 48 21 Colombia Colombia NNP 32505 48 22 , , , 32505 48 23 whereas whereas IN 32505 48 24 the the DT 32505 48 25 other other JJ 32505 48 26 species specie NNS 32505 48 27 are be VBP 32505 48 28 known know VBN 32505 48 29 only only RB 32505 48 30 from from IN 32505 48 31 Middle Middle NNP 32505 48 32 America America NNP 32505 48 33 . . . 32505 49 1 _ _ NNP 32505 49 2 Distribution._--The Distribution._--The NNP 32505 49 3 species specie NNS 32505 49 4 of of IN 32505 49 5 the the DT 32505 49 6 _ _ NNP 32505 49 7 Hyla Hyla NNP 32505 49 8 rubra rubra NN 32505 49 9 _ _ NNP 32505 49 10 group group NN 32505 49 11 range range NN 32505 49 12 from from IN 32505 49 13 the the DT 32505 49 14 lowlands lowland NNS 32505 49 15 of of IN 32505 49 16 northern northern JJ 32505 49 17 Argentina Argentina NNP 32505 49 18 and and CC 32505 49 19 Bolivia Bolivia NNP 32505 49 20 to to IN 32505 49 21 southern southern JJ 32505 49 22 Tamaulipas Tamaulipas NNP 32505 49 23 and and CC 32505 49 24 Guerrero Guerrero NNP 32505 49 25 , , , 32505 49 26 México México NNP 32505 49 27 . . . 32505 50 1 _ _ NNP 32505 50 2 Comments._--In comments._--in NN 32505 50 3 Central Central NNP 32505 50 4 America America NNP 32505 50 5 two two CD 32505 50 6 subgroups subgroup NNS 32505 50 7 of of IN 32505 50 8 species specie NNS 32505 50 9 can can MD 32505 50 10 be be VB 32505 50 11 recognized recognize VBN 32505 50 12 . . . 32505 51 1 _ _ NNP 32505 51 2 Hyla Hyla NNP 32505 51 3 boulengeri boulengeri NN 32505 51 4 _ _ NNP 32505 51 5 and and CC 32505 51 6 _ _ NNP 32505 51 7 H. H. NNP 32505 52 1 foliamorta foliamorta NNP 32505 52 2 _ _ NNP 32505 52 3 are be VBP 32505 52 4 distinctive distinctive JJ 32505 52 5 in in IN 32505 52 6 the the DT 32505 52 7 large large JJ 32505 52 8 size size NN 32505 52 9 of of IN 32505 52 10 adults adult NNS 32505 52 11 ( ( -LRB- 32505 52 12 snout snout NN 32505 52 13 - - HYPH 32505 52 14 vent vent NN 32505 52 15 lengths length NNS 32505 52 16 41 41 CD 32505 52 17 - - SYM 32505 52 18 49 49 CD 32505 52 19 mm mm CD 32505 52 20 . . . 32505 53 1 ) ) -RRB- 32505 53 2 ; ; : 32505 53 3 both both DT 32505 53 4 have have VBP 32505 53 5 prominent prominent JJ 32505 53 6 bars bar NNS 32505 53 7 on on IN 32505 53 8 the the DT 32505 53 9 thighs thigh NNS 32505 53 10 , , , 32505 53 11 a a DT 32505 53 12 well well RB 32505 53 13 - - HYPH 32505 53 14 defined define VBN 32505 53 15 interorbital interorbital NNP 32505 53 16 triangular triangular NNP 32505 53 17 mark mark NNP 32505 53 18 , , , 32505 53 19 blotches blotch NNS 32505 53 20 or or CC 32505 53 21 spots spot NNS 32505 53 22 dorsally dorsally RB 32505 53 23 , , , 32505 53 24 and and CC 32505 53 25 large large JJ 32505 53 26 choanae choanae NN 32505 53 27 . . . 32505 54 1 _ _ NNP 32505 54 2 Hyla Hyla NNP 32505 54 3 elaeochroa elaeochroa NN 32505 54 4 , , , 32505 54 5 _ _ NNP 32505 54 6 _ _ NNP 32505 54 7 H. H. NNP 32505 55 1 rubra rubra NNP 32505 55 2 , , , 32505 55 3 _ _ NNP 32505 55 4 and and CC 32505 55 5 _ _ NNP 32505 55 6 H. H. NNP 32505 56 1 staufferi staufferi NNP 32505 56 2 _ _ NNP 32505 56 3 are be VBP 32505 56 4 smaller small JJR 32505 56 5 ( ( -LRB- 32505 56 6 snout snout NN 32505 56 7 - - HYPH 32505 56 8 vent vent NN 32505 56 9 lengths length NNS 32505 56 10 29 29 CD 32505 56 11 - - SYM 32505 56 12 40 40 CD 32505 56 13 mm mm CD 32505 56 14 . . . 32505 57 1 ) ) -RRB- 32505 57 2 ; ; : 32505 57 3 they -PRON- PRP 32505 57 4 have have VBP 32505 57 5 the the DT 32505 57 6 thighs thigh NNS 32505 57 7 weakly weakly RB 32505 57 8 barred bar VBN 32505 57 9 or or CC 32505 57 10 vermiculate vermiculate VBP 32505 57 11 anteriorly anteriorly RB 32505 57 12 and and CC 32505 57 13 posteriorly posteriorly RB 32505 57 14 or or CC 32505 57 15 unmarked unmarked JJ 32505 57 16 , , , 32505 57 17 an an DT 32505 57 18 ill ill RB 32505 57 19 - - HYPH 32505 57 20 defined define VBN 32505 57 21 interorbital interorbital NNP 32505 57 22 triangular triangular NNP 32505 57 23 mark mark NNP 32505 57 24 , , , 32505 57 25 linear linear JJ 32505 57 26 markings marking NNS 32505 57 27 dorsally dorsally RB 32505 57 28 , , , 32505 57 29 and and CC 32505 57 30 small small JJ 32505 57 31 choanae choanae NN 32505 57 32 . . . 32505 58 1 Key key JJ 32505 58 2 to to IN 32505 58 3 Species Species NNPS 32505 58 4 and and CC 32505 58 5 Subspecies Subspecies NNPS 32505 58 6 1 1 CD 32505 58 7 . . . 32505 59 1 Larger large JJR 32505 59 2 frogs frog NNS 32505 59 3 ( ( -LRB- 32505 59 4 males male NNS 32505 59 5 to to IN 32505 59 6 49 49 CD 32505 59 7 mm mm NNS 32505 59 8 . . . 32505 60 1 snout snout NN 32505 60 2 - - HYPH 32505 60 3 vent vent NN 32505 60 4 length length NN 32505 60 5 ) ) -RRB- 32505 60 6 ; ; : 32505 60 7 thighs thigh NNS 32505 60 8 strongly strongly RB 32505 60 9 barred bar VBD 32505 60 10 ; ; : 32505 60 11 supratympanic supratympanic JJ 32505 60 12 fold fold JJ 32505 60 13 black black JJ 32505 60 14 ; ; : 32505 60 15 dorsum dorsum NN 32505 60 16 blotched blotch VBN 32505 60 17 or or CC 32505 60 18 spotted spot VBD 32505 60 19 2 2 CD 32505 60 20 Smaller small JJR 32505 60 21 frogs frog NNS 32505 60 22 ( ( -LRB- 32505 60 23 males male NNS 32505 60 24 to to IN 32505 60 25 40 40 CD 32505 60 26 mm mm NNS 32505 60 27 . . . 32505 61 1 snout snout NN 32505 61 2 - - HYPH 32505 61 3 vent vent NN 32505 61 4 length length NN 32505 61 5 ) ) -RRB- 32505 61 6 ; ; : 32505 61 7 thighs thigh NNS 32505 61 8 weakly weakly RB 32505 61 9 barred bar VBD 32505 61 10 or or CC 32505 61 11 plain plain JJ 32505 61 12 ; ; : 32505 61 13 supratympanic supratympanic JJ 32505 61 14 fold fold JJ 32505 61 15 pale pale JJ 32505 61 16 brown brown JJ 32505 61 17 ; ; : 32505 61 18 dorsum dorsum NN 32505 61 19 usually usually RB 32505 61 20 having have VBG 32505 61 21 linear linear JJ 32505 61 22 pattern pattern NN 32505 61 23 3 3 CD 32505 61 24 2 2 CD 32505 61 25 . . . 32505 62 1 Dorsum dorsum NN 32505 62 2 tuberculate tuberculate NN 32505 62 3 ; ; : 32505 62 4 snout snout NN 32505 62 5 subacuminate subacuminate NN 32505 62 6 ; ; : 32505 62 7 vocal vocal JJ 32505 62 8 sac sac NN 32505 62 9 flecked fleck VBN 32505 62 10 with with IN 32505 62 11 brown brown JJ 32505 62 12 ; ; : 32505 62 13 tarsal tarsal JJ 32505 62 14 fold fold JJ 32505 62 15 rudimentary rudimentary NNP 32505 62 16 ; ; : 32505 62 17 web web NN 32505 62 18 absent absent NN 32505 62 19 between between IN 32505 62 20 fingers finger NNS 32505 62 21 ; ; : 32505 62 22 black black JJ 32505 62 23 spots spot NNS 32505 62 24 absent absent JJ 32505 62 25 in in IN 32505 62 26 scapular scapular JJ 32505 62 27 region region NN 32505 62 28 _ _ NNP 32505 62 29 H. H. NNP 32505 63 1 boulengeri boulengeri NNP 32505 63 2 _ _ NNP 32505 63 3 Dorsum Dorsum NNP 32505 63 4 smooth smooth JJ 32505 63 5 ; ; : 32505 63 6 snout snout NN 32505 63 7 pointed point VBD 32505 63 8 ; ; : 32505 63 9 vocal vocal JJ 32505 63 10 sac sac NN 32505 63 11 dark dark JJ 32505 63 12 gray gray NN 32505 63 13 ; ; : 32505 63 14 tarsal tarsal JJ 32505 63 15 fold fold VB 32505 63 16 absent absent JJ 32505 63 17 ; ; : 32505 63 18 trace trace NN 32505 63 19 of of IN 32505 63 20 web web NN 32505 63 21 between between IN 32505 63 22 fingers finger NNS 32505 63 23 ; ; : 32505 63 24 two two CD 32505 63 25 or or CC 32505 63 26 more more JJR 32505 63 27 elongate elongate VB 32505 63 28 dark dark JJ 32505 63 29 spots spot NNS 32505 63 30 in in IN 32505 63 31 scapular scapular JJ 32505 63 32 region region NN 32505 63 33 _ _ NNP 32505 63 34 H. H. NNP 32505 64 1 foliamorta foliamorta NNP 32505 64 2 _ _ NNP 32505 64 3 3 3 CD 32505 64 4 . . . 32505 65 1 Snout snout NN 32505 65 2 - - HYPH 32505 65 3 vent vent NN 32505 65 4 length length NN 32505 65 5 more more JJR 32505 65 6 than than IN 32505 65 7 30 30 CD 32505 65 8 mm mm NN 32505 65 9 . . . 32505 66 1 ; ; : 32505 66 2 tympanum tympanum NN 32505 66 3 2/3 2/3 CD 32505 66 4 to to IN 32505 66 5 3/4 3/4 CD 32505 66 6 diameter diameter NN 32505 66 7 of of IN 32505 66 8 eye eye NN 32505 66 9 ; ; : 32505 66 10 prevomerine prevomerine JJ 32505 66 11 elevations elevation NNS 32505 66 12 about about IN 32505 66 13 size size NN 32505 66 14 of of IN 32505 66 15 choanae choanae NN 32505 66 16 4 4 CD 32505 66 17 Snout Snout NNP 32505 66 18 - - HYPH 32505 66 19 vent vent NN 32505 66 20 length length NN 32505 66 21 less less JJR 32505 66 22 than than IN 32505 66 23 30 30 CD 32505 66 24 mm mm NN 32505 66 25 . . . 32505 67 1 ; ; : 32505 67 2 tympanum tympanum VB 32505 67 3 less less JJR 32505 67 4 than than IN 32505 67 5 1/2 1/2 CD 32505 67 6 diameter diameter NN 32505 67 7 of of IN 32505 67 8 eye eye NN 32505 67 9 ; ; : 32505 67 10 prevomerine prevomerine JJ 32505 67 11 elevations elevation NNS 32505 67 12 smaller small JJR 32505 67 13 than than IN 32505 67 14 choanae choanae NNP 32505 67 15 5 5 CD 32505 67 16 4 4 CD 32505 67 17 . . . 32505 68 1 Thighs thigh NNS 32505 68 2 mottled mottle VBD 32505 68 3 posteriorly posteriorly RB 32505 68 4 ; ; : 32505 68 5 discs disc NNS 32505 68 6 on on IN 32505 68 7 fingers finger NNS 32505 68 8 about about RB 32505 68 9 1/2 1/2 CD 32505 68 10 size size NN 32505 68 11 of of IN 32505 68 12 tympanum tympanum NN 32505 68 13 ; ; , 32505 68 14 faint faint JJ 32505 68 15 dark dark JJ 32505 68 16 line line NN 32505 68 17 from from IN 32505 68 18 nostril nostril NNP 32505 68 19 to to IN 32505 68 20 eye eye NN 32505 68 21 _ _ NNP 32505 68 22 H. H. NNP 32505 69 1 rubra rubra NNP 32505 69 2 _ _ NNP 32505 69 3 Thighs Thighs NNP 32505 69 4 faintly faintly RB 32505 69 5 barred bar VBN 32505 69 6 or or CC 32505 69 7 plain plain RB 32505 69 8 posteriorly posteriorly RB 32505 69 9 ; ; : 32505 69 10 discs disc NNS 32505 69 11 on on IN 32505 69 12 fingers finger NNS 32505 69 13 about about IN 32505 69 14 size size NN 32505 69 15 of of IN 32505 69 16 tympanum tympanum NN 32505 69 17 ; ; : 32505 69 18 distinct distinct JJ 32505 69 19 dark dark JJ 32505 69 20 line line NN 32505 69 21 from from IN 32505 69 22 nostril nostril NNP 32505 69 23 to to IN 32505 69 24 eye eye NN 32505 69 25 _ _ NNP 32505 69 26 H. H. NNP 32505 70 1 elaeochroa elaeochroa NNP 32505 70 2 _ _ NNP 32505 70 3 5 5 CD 32505 70 4 . . . 32505 71 1 Dorsum dorsum NN 32505 71 2 brown brown JJ 32505 71 3 with with IN 32505 71 4 irregular irregular JJ 32505 71 5 dorsolateral dorsolateral JJ 32505 71 6 stripes stripe NNS 32505 71 7 and and CC 32505 71 8 interrupted interrupt VBN 32505 71 9 paravertebral paravertebral JJ 32505 71 10 stripes stripe NNS 32505 71 11 ; ; : 32505 71 12 two two CD 32505 71 13 transverse transverse JJ 32505 71 14 bars bar NNS 32505 71 15 on on IN 32505 71 16 shanks shank NNS 32505 71 17 ; ; : 32505 71 18 interorbital interorbital NNP 32505 71 19 bar bar NN 32505 71 20 present present NN 32505 71 21 _ _ NNP 32505 71 22 H. H. NNP 32505 72 1 staufferi staufferi NNP 32505 72 2 staufferi staufferi NNP 32505 72 3 _ _ NNP 32505 72 4 Dorsum Dorsum NNP 32505 72 5 gray gray NN 32505 72 6 with with IN 32505 72 7 complete complete JJ 32505 72 8 dorsolateral dorsolateral JJ 32505 72 9 and and CC 32505 72 10 paravertebral paravertebral JJ 32505 72 11 stripes stripe NNS 32505 72 12 ; ; , 32505 72 13 longitudinal longitudinal JJ 32505 72 14 stripe stripe NN 32505 72 15 on on IN 32505 72 16 shank shank JJ 32505 72 17 ; ; : 32505 72 18 interorbital interorbital NNP 32505 72 19 bar bar NN 32505 72 20 absent absent NN 32505 72 21 _ _ NNP 32505 72 22 H. H. NNP 32505 73 1 staufferi staufferi NNP 32505 73 2 altae altae NNP 32505 73 3 _ _ NNP 32505 73 4 Key Key NNP 32505 73 5 to to IN 32505 73 6 Known Known NNP 32505 73 7 Tadpoles Tadpoles NNP 32505 73 8 1 1 CD 32505 73 9 . . . 32505 74 1 Entire entire JJ 32505 74 2 lower low JJR 32505 74 3 beak beak JJ 32505 74 4 black black NN 32505 74 5 ; ; : 32505 74 6 beaks beak NNS 32505 74 7 moderate moderate JJ 32505 74 8 - - HYPH 32505 74 9 sized sized JJ 32505 74 10 , , , 32505 74 11 serrate serrate NN 32505 74 12 ; ; : 32505 74 13 dorsal dorsal NN 32505 74 14 fin fin NNP 32505 74 15 high high JJ 32505 74 16 , , , 32505 74 17 extending extend VBG 32505 74 18 to to IN 32505 74 19 middle middle NN 32505 74 20 of of IN 32505 74 21 back back JJ 32505 74 22 2 2 CD 32505 74 23 No no DT 32505 74 24 more more JJR 32505 74 25 than than IN 32505 74 26 half half NN 32505 74 27 of of IN 32505 74 28 lower low JJR 32505 74 29 beak beak JJ 32505 74 30 black black NN 32505 74 31 ; ; : 32505 74 32 beaks beak NNS 32505 74 33 small small JJ 32505 74 34 , , , 32505 74 35 finely finely RB 32505 74 36 serrate serrate VB 32505 74 37 ; ; : 32505 74 38 dorsal dorsal NN 32505 74 39 fin fin NNP 32505 74 40 lower low JJR 32505 74 41 , , , 32505 74 42 barely barely RB 32505 74 43 extending extend VBG 32505 74 44 onto onto IN 32505 74 45 body body NN 32505 74 46 3 3 CD 32505 74 47 2 2 CD 32505 74 48 . . . 32505 75 1 Papillae Papillae NNP 32505 75 2 present present VBP 32505 75 3 only only RB 32505 75 4 laterally laterally RB 32505 75 5 _ _ NNP 32505 75 6 H. H. NNP 32505 76 1 boulengeri boulengeri NNP 32505 76 2 _ _ NNP 32505 76 3 Papillae Papillae NNP 32505 76 4 present present VBP 32505 76 5 laterally laterally RB 32505 76 6 and and CC 32505 76 7 ventrally ventrally RB 32505 76 8 _ _ NNP 32505 76 9 H. H. NNP 32505 77 1 foliamorta foliamorta NNP 32505 77 2 _ _ NNP 32505 77 3 3 3 CD 32505 77 4 . . . 32505 78 1 Distinct distinct JJ 32505 78 2 brown brown JJ 32505 78 3 stripe stripe NN 32505 78 4 from from IN 32505 78 5 nostril nostril NNP 32505 78 6 to to IN 32505 78 7 eye eye NN 32505 78 8 ; ; : 32505 78 9 two two CD 32505 78 10 stripes stripe NNS 32505 78 11 below below IN 32505 78 12 eye eye NN 32505 78 13 , , , 32505 78 14 _ _ NNP 32505 78 15 H. H. NNP 32505 79 1 elaeochroa elaeochroa NNP 32505 79 2 _ _ NNP 32505 79 3 Faint Faint NNP 32505 79 4 stripe stripe NN 32505 79 5 from from IN 32505 79 6 nostril nostril NNP 32505 79 7 to to IN 32505 79 8 eye eye NN 32505 79 9 ; ; : 32505 79 10 no no DT 32505 79 11 stripe stripe NN 32505 79 12 below below IN 32505 79 13 eye eye NN 32505 79 14 _ _ NNP 32505 79 15 H. H. NNP 32505 80 1 staufferi staufferi NNP 32505 80 2 _ _ NNP 32505 80 3 ACCOUNTS ACCOUNTS NNP 32505 80 4 OF of IN 32505 80 5 SPECIES SPECIES NNP 32505 80 6 AND and CC 32505 80 7 SUBSPECIES SUBSPECIES NNP 32505 80 8 _ _ NNP 32505 80 9 Hyla Hyla NNP 32505 80 10 boulengeri boulengeri NN 32505 80 11 _ _ NNP 32505 80 12 ( ( -LRB- 32505 80 13 Cope Cope NNP 32505 80 14 ) ) -RRB- 32505 80 15 _ _ NNP 32505 80 16 Scytopis Scytopis NNP 32505 80 17 boulengeri boulengeri NN 32505 80 18 _ _ NNP 32505 80 19 Cope Cope NNP 32505 80 20 , , , 32505 80 21 Bull Bull NNP 32505 80 22 . . . 32505 81 1 U.S. U.S. NNP 32505 81 2 Natl Natl NNP 32505 81 3 . . . 32505 82 1 Mus Mus NNP 32505 82 2 . . NNP 32505 82 3 , , , 32505 82 4 32:12 32:12 CD 32505 82 5 , , , 32505 82 6 December December NNP 32505 82 7 1 1 CD 32505 82 8 , , , 32505 82 9 1887 1887 CD 32505 82 10 [ [ -LRB- 32505 82 11 Holotype.--USNM Holotype.--USNM NNP 32505 82 12 13974 13974 CD 32505 82 13 , , , 32505 82 14 from from IN 32505 82 15 " " `` 32505 82 16 Nicaragua Nicaragua NNP 32505 82 17 " " '' 32505 82 18 ; ; : 32505 82 19 J. J. NNP 32505 83 1 A. A. NNP 32505 83 2 McNiel McNiel NNP 32505 83 3 , , , 32505 83 4 collector collector NN 32505 83 5 ] ] -RRB- 32505 83 6 . . . 32505 84 1 _ _ NNP 32505 84 2 Hyla Hyla NNP 32505 84 3 boulengeri boulengeri NN 32505 84 4 : : : 32505 84 5 _ _ NNP 32505 84 6 Günther Günther NNP 32505 84 7 , , , 32505 84 8 Biologia Biologia NNP 32505 84 9 Centrali Centrali NNP 32505 84 10 - - HYPH 32505 84 11 Americana Americana NNP 32505 84 12 , , , 32505 84 13 Reptilia Reptilia NNP 32505 84 14 and and CC 32505 84 15 Batrachia Batrachia NNP 32505 84 16 , , , 32505 84 17 p. p. NN 32505 84 18 267 267 CD 32505 84 19 , , , 32505 84 20 June June NNP 32505 84 21 1901 1901 CD 32505 84 22 . . . 32505 85 1 Noble Noble NNP 32505 85 2 , , , 32505 85 3 Bull Bull NNP 32505 85 4 . . . 32505 86 1 Amer amer NN 32505 86 2 . . . 32505 87 1 Mus mus NN 32505 87 2 . . . 32505 88 1 Nat Nat NNP 32505 88 2 . . . 32505 89 1 Hist Hist NNP 32505 89 2 , , , 32505 89 3 38:339 38:339 CD 32505 89 4 , , , 32505 89 5 June June NNP 32505 89 6 1918 1918 CD 32505 89 7 . . . 32505 90 1 Taylor Taylor NNP 32505 90 2 , , , 32505 90 3 Univ Univ NNP 32505 90 4 . . . 32505 91 1 Kansas Kansas NNP 32505 91 2 Sci Sci NNP 32505 91 3 . . . 32505 92 1 Bull Bull NNP 32505 92 2 . . NNP 32505 92 3 , , , 32505 92 4 35:856 35:856 CD 32505 92 5 , , , 32505 92 6 July July NNP 32505 92 7 1 1 CD 32505 92 8 , , , 32505 92 9 1952 1952 CD 32505 92 10 . . . 32505 93 1 _ _ NNP 32505 93 2 Diagnosis._--Size diagnosis._--size VB 32505 93 3 large large JJ 32505 93 4 ( ( -LRB- 32505 93 5 Male male NN 32505 93 6 to to IN 32505 93 7 49 49 CD 32505 93 8 mm mm NNS 32505 93 9 . . NNP 32505 93 10 , , , 32505 93 11 Female Female NNP 32505 93 12 to to IN 32505 93 13 53 53 CD 32505 93 14 mm mm NNS 32505 93 15 . . . 32505 94 1 ) ) -RRB- 32505 94 2 ; ; : 32505 94 3 skull skull VB 32505 94 4 as as RB 32505 94 5 long long RB 32505 94 6 as as IN 32505 94 7 wide wide JJ 32505 94 8 ; ; : 32505 94 9 frontoparietal frontoparietal JJ 32505 94 10 fontanelle fontanelle NN 32505 94 11 present present NN 32505 94 12 ; ; : 32505 94 13 snout snout NN 32505 94 14 subacuminate subacuminate NN 32505 94 15 ; ; : 32505 94 16 canthus canthus NNP 32505 94 17 not not RB 32505 94 18 pronounced pronounce VBN 32505 94 19 ; ; : 32505 94 20 choanae choanae RB 32505 94 21 large large JJ 32505 94 22 ; ; : 32505 94 23 tongue tongue NN 32505 94 24 cordiform cordiform NN 32505 94 25 , , , 32505 94 26 slightly slightly RB 32505 94 27 longer long JJR 32505 94 28 than than IN 32505 94 29 broad broad JJ 32505 94 30 ; ; : 32505 94 31 interorbital interorbital NNP 32505 94 32 triangle triangle NNP 32505 94 33 tubercular tubercular NNP 32505 94 34 ; ; : 32505 94 35 skin skin NN 32505 94 36 on on IN 32505 94 37 dorsum dorsum NN 32505 94 38 tuberculate tuberculate NN 32505 94 39 ; ; : 32505 94 40 tarsal tarsal NN 32505 94 41 fold fold VB 32505 94 42 reduced reduced JJ 32505 94 43 or or CC 32505 94 44 absent absent JJ 32505 94 45 ; ; : 32505 94 46 thighs thigh NNS 32505 94 47 , , , 32505 94 48 shanks shank NNS 32505 94 49 , , , 32505 94 50 and and CC 32505 94 51 tarsi tarsi NNP 32505 94 52 boldly boldly RB 32505 94 53 barred bar VBN 32505 94 54 with with IN 32505 94 55 dark dark JJ 32505 94 56 brown brown JJ 32505 94 57 and and CC 32505 94 58 pale pale JJ 32505 94 59 yellow yellow NN 32505 94 60 - - HYPH 32505 94 61 green green JJ 32505 94 62 in in IN 32505 94 63 life life NN 32505 94 64 . . . 32505 95 1 _ _ NNP 32505 95 2 Description._--Head Description._--Head NNP 32505 95 3 flattened flatten VBD 32505 95 4 , , , 32505 95 5 longer longer RB 32505 95 6 than than IN 32505 95 7 wide wide JJ 32505 95 8 ; ; : 32505 95 9 snout snout NN 32505 95 10 projecting projecting NN 32505 95 11 beyond beyond IN 32505 95 12 lower low JJR 32505 95 13 lip lip NN 32505 95 14 ; ; : 32505 95 15 loreal loreal NN 32505 95 16 region region NN 32505 95 17 oblique oblique NNP 32505 95 18 ; ; : 32505 95 19 canthus canthus NNP 32505 95 20 not not RB 32505 95 21 pronounced pronounced JJ 32505 95 22 ; ; : 32505 95 23 length length NN 32505 95 24 of of IN 32505 95 25 eye eye NN 32505 95 26 less less JJR 32505 95 27 than than IN 32505 95 28 interorbital interorbital JJ 32505 95 29 distance distance NN 32505 95 30 ; ; , 32505 95 31 tympanum tympanum VB 32505 95 32 large large JJ 32505 95 33 , , , 32505 95 34 about about RB 32505 95 35 70 70 CD 32505 95 36 per per NN 32505 95 37 cent cent NN 32505 95 38 of of IN 32505 95 39 diameter diameter NN 32505 95 40 of of IN 32505 95 41 eye eye NN 32505 95 42 ; ; : 32505 95 43 interorbital interorbital NNP 32505 95 44 triangle triangle NNP 32505 95 45 distinct distinct JJ 32505 95 46 ; ; : 32505 95 47 arms arm NNS 32505 95 48 short short JJ 32505 95 49 ; ; : 32505 95 50 fingers finger NNS 32505 95 51 lacking lacking JJ 32505 95 52 web web NN 32505 95 53 ; ; : 32505 95 54 palmar palmar NNP 32505 95 55 tubercle tubercle NN 32505 95 56 tripartite tripartite NN 32505 95 57 ; ; , 32505 95 58 subarticular subarticular JJ 32505 95 59 tubercles tubercle NNS 32505 95 60 distinct distinct JJ 32505 95 61 ; ; : 32505 95 62 long long JJ 32505 95 63 tubercle tubercle NN 32505 95 64 on on IN 32505 95 65 base base NN 32505 95 66 of of IN 32505 95 67 first first JJ 32505 95 68 finger finger NN 32505 95 69 ; ; : 32505 95 70 discs discs NNP 32505 95 71 truncate truncate NNP 32505 95 72 ; ; : 32505 95 73 legs leg NNS 32505 95 74 long long RB 32505 95 75 ; ; : 32505 95 76 tarsal tarsal JJ 32505 95 77 fold fold VB 32505 95 78 reduced reduced JJ 32505 95 79 or or CC 32505 95 80 absent absent JJ 32505 95 81 ; ; : 32505 95 82 inner inner JJ 32505 95 83 metatarsal metatarsal NN 32505 95 84 tubercle tubercle NN 32505 95 85 rounded round VBD 32505 95 86 , , , 32505 95 87 larger large JJR 32505 95 88 than than IN 32505 95 89 outer outer JJ 32505 95 90 , , , 32505 95 91 both both DT 32505 95 92 elevated elevated JJ 32505 95 93 ; ; : 32505 95 94 subarticular subarticular JJ 32505 95 95 tubercles tubercle NNS 32505 95 96 distinct distinct JJ 32505 95 97 ; ; : 32505 95 98 one one CD 32505 95 99 phalanx phalanx NN 32505 95 100 free free JJ 32505 95 101 of of IN 32505 95 102 web web NN 32505 95 103 on on IN 32505 95 104 second second JJ 32505 95 105 , , , 32505 95 106 third third JJ 32505 95 107 , , , 32505 95 108 and and CC 32505 95 109 fifth fifth JJ 32505 95 110 toes toe NNS 32505 95 111 , , , 32505 95 112 three three CD 32505 95 113 free free JJ 32505 95 114 on on IN 32505 95 115 fourth fourth JJ 32505 95 116 toe toe NN 32505 95 117 ( ( -LRB- 32505 95 118 Fig fig NN 32505 95 119 . . . 32505 96 1 1A 1a CD 32505 96 2 and and CC 32505 96 3 B B NNP 32505 96 4 ) ) -RRB- 32505 96 5 ; ; : 32505 96 6 skin skin NN 32505 96 7 tuberculate tuberculate NN 32505 96 8 on on IN 32505 96 9 dorsum dorsum NN 32505 96 10 , , , 32505 96 11 less less RBR 32505 96 12 so so RB 32505 96 13 on on IN 32505 96 14 flanks flank NNS 32505 96 15 ; ; : 32505 96 16 skin skin NN 32505 96 17 of of IN 32505 96 18 belly belly NN 32505 96 19 granular granular JJ 32505 96 20 , , , 32505 96 21 that that IN 32505 96 22 on on IN 32505 96 23 chest chest NN 32505 96 24 and and CC 32505 96 25 throat throat NN 32505 96 26 weakly weakly RB 32505 96 27 granular granular NN 32505 96 28 ; ; : 32505 96 29 tongue tongue NN 32505 96 30 cordiform cordiform NN 32505 96 31 , , , 32505 96 32 longer long JJR 32505 96 33 than than IN 32505 96 34 wide wide JJ 32505 96 35 , , , 32505 96 36 free free JJ 32505 96 37 and and CC 32505 96 38 notched notch VBN 32505 96 39 behind behind RB 32505 96 40 ; ; : 32505 96 41 vocal vocal NN 32505 96 42 slits slit NNS 32505 96 43 large large JJ 32505 96 44 , , , 32505 96 45 lateral lateral JJ 32505 96 46 to to IN 32505 96 47 tongue tongue NN 32505 96 48 . . . 32505 97 1 [ [ -LRB- 32505 97 2 Illustration illustration NN 32505 97 3 : : : 32505 97 4 Fig Fig NNP 32505 97 5 . . . 32505 98 1 1 1 LS 32505 98 2 . . . 32505 99 1 A a DT 32505 99 2 and and CC 32505 99 3 B.--Hand b.--hand NN 32505 99 4 and and CC 32505 99 5 foot foot NN 32505 99 6 of of IN 32505 99 7 _ _ NNP 32505 99 8 Hyla Hyla NNP 32505 99 9 boulengeri boulengeri NN 32505 99 10 _ _ NNP 32505 99 11 ( ( -LRB- 32505 99 12 KU KU NNP 32505 99 13 102173 102173 CD 32505 99 14 ) ) -RRB- 32505 99 15 , , , 32505 99 16 × × NN 32505 99 17 3 3 CD 32505 99 18 . . . 32505 100 1 C C NNP 32505 100 2 and and CC 32505 100 3 D.--Hand D.--Hand NNP 32505 100 4 and and CC 32505 100 5 foot foot NN 32505 100 6 of of IN 32505 100 7 _ _ NNP 32505 100 8 Hyla Hyla NNP 32505 100 9 s. s. NN 32505 100 10 staufferi_× staufferi_× NN 32505 100 11 ( ( -LRB- 32505 100 12 KU KU NNP 32505 100 13 57790 57790 CD 32505 100 14 ) ) -RRB- 32505 100 15 , , , 32505 100 16 × × NNP 32505 100 17 6 6 CD 32505 100 18 ] ] -RRB- 32505 100 19 In in IN 32505 100 20 life life NN 32505 100 21 , , , 32505 100 22 dorsum dorsum NN 32505 100 23 tan tan NN 32505 100 24 or or CC 32505 100 25 brown brown JJ 32505 100 26 with with IN 32505 100 27 dark dark JJ 32505 100 28 spots spot NNS 32505 100 29 on on IN 32505 100 30 snout snout NN 32505 100 31 , , , 32505 100 32 head head NN 32505 100 33 , , , 32505 100 34 and and CC 32505 100 35 scapular scapular JJ 32505 100 36 region region NN 32505 100 37 ; ; : 32505 100 38 interorbital interorbital NNP 32505 100 39 triangle triangle NN 32505 100 40 and and CC 32505 100 41 blotch blotch NN 32505 100 42 posteriorly posteriorly RB 32505 100 43 on on IN 32505 100 44 dorsum dorsum NN 32505 100 45 dark dark JJ 32505 100 46 brown brown NN 32505 100 47 ; ; : 32505 100 48 flanks flank VBZ 32505 100 49 pale pale JJ 32505 100 50 green green JJ 32505 100 51 ; ; : 32505 100 52 groin groin NN 32505 100 53 pale pale JJ 32505 100 54 green green JJ 32505 100 55 or or CC 32505 100 56 orange orange NN 32505 100 57 , , , 32505 100 58 mottled mottle VBN 32505 100 59 with with IN 32505 100 60 dark dark JJ 32505 100 61 brown brown NNP 32505 100 62 ; ; : 32505 100 63 thighs thigh NNS 32505 100 64 tan tan NNP 32505 100 65 or or CC 32505 100 66 brown brown NNP 32505 100 67 above above IN 32505 100 68 with with IN 32505 100 69 dark dark JJ 32505 100 70 transverse transverse JJ 32505 100 71 bars bar NNS 32505 100 72 on on IN 32505 100 73 anterior anterior NNP 32505 100 74 and and CC 32505 100 75 posterior posterior NN 32505 100 76 surfaces surface NNS 32505 100 77 ; ; , 32505 100 78 spaces space NNS 32505 100 79 between between IN 32505 100 80 bars bar NNS 32505 100 81 green green JJ 32505 100 82 or or CC 32505 100 83 orange orange JJ 32505 100 84 ; ; , 32505 100 85 inner inner JJ 32505 100 86 surfaces surface NNS 32505 100 87 of of IN 32505 100 88 shanks shank NNS 32505 100 89 and and CC 32505 100 90 outer outer JJ 32505 100 91 surfaces surface NNS 32505 100 92 of of IN 32505 100 93 tarsi tarsi NNP 32505 100 94 brown brown NNP 32505 100 95 and and CC 32505 100 96 orange orange NNP 32505 100 97 ; ; : 32505 100 98 foot foot NNP 32505 100 99 brown brown NNP 32505 100 100 above above RB 32505 100 101 ; ; : 32505 100 102 forelimbs forelimbs NNP 32505 100 103 brown brown JJ 32505 100 104 and and CC 32505 100 105 pale pale JJ 32505 100 106 green green NN 32505 100 107 above above RB 32505 100 108 , , , 32505 100 109 weakly weakly RB 32505 100 110 barred bar VBN 32505 100 111 ; ; : 32505 100 112 belly belly NN 32505 100 113 creamy creamy JJ 32505 100 114 white white JJ 32505 100 115 with with IN 32505 100 116 scattered scatter VBN 32505 100 117 brown brown JJ 32505 100 118 spots spot NNS 32505 100 119 ; ; : 32505 100 120 vocal vocal JJ 32505 100 121 sac sac NN 32505 100 122 creamy creamy JJ 32505 100 123 white white JJ 32505 100 124 flecked fleck VBN 32505 100 125 with with IN 32505 100 126 brown brown JJ 32505 100 127 ; ; : 32505 100 128 lower low JJR 32505 100 129 jaw jaw NN 32505 100 130 brown brown JJ 32505 100 131 with with IN 32505 100 132 white white JJ 32505 100 133 spots spot NNS 32505 100 134 on on IN 32505 100 135 lips lip NNS 32505 100 136 ( ( -LRB- 32505 100 137 Pl Pl NNP 32505 100 138 . . . 32505 101 1 1A 1a LS 32505 101 2 ) ) -RRB- 32505 101 3 . . . 32505 102 1 In in IN 32505 102 2 preservative preservative JJ 32505 102 3 , , , 32505 102 4 head head NN 32505 102 5 and and CC 32505 102 6 dorsum dorsum NN 32505 102 7 dark dark JJ 32505 102 8 brown brown NN 32505 102 9 with with IN 32505 102 10 triangular triangular JJ 32505 102 11 spot spot NN 32505 102 12 between between IN 32505 102 13 eyes eye NNS 32505 102 14 ; ; : 32505 102 15 dark dark JJ 32505 102 16 spots spot NNS 32505 102 17 on on IN 32505 102 18 head head NN 32505 102 19 and and CC 32505 102 20 scapular scapular JJ 32505 102 21 region region NN 32505 102 22 and and CC 32505 102 23 dark dark JJ 32505 102 24 brown brown JJ 32505 102 25 blotch blotch NN 32505 102 26 posteriorly posteriorly RB 32505 102 27 on on IN 32505 102 28 dorsum dorsum NN 32505 102 29 ; ; , 32505 102 30 flanks flank VBZ 32505 102 31 creamy creamy VBP 32505 102 32 white white JJ 32505 102 33 with with IN 32505 102 34 brown brown JJ 32505 102 35 spots spot NNS 32505 102 36 ; ; : 32505 102 37 groin groin NNP 32505 102 38 creamy creamy JJ 32505 102 39 white white NNP 32505 102 40 mottled mottle VBN 32505 102 41 with with IN 32505 102 42 dark dark JJ 32505 102 43 brown brown NNP 32505 102 44 ; ; : 32505 102 45 thighs thigh NNS 32505 102 46 brown brown NNP 32505 102 47 above above IN 32505 102 48 with with IN 32505 102 49 dark dark JJ 32505 102 50 brown brown JJ 32505 102 51 transverse transverse JJ 32505 102 52 bars bar NNS 32505 102 53 on on IN 32505 102 54 anterior anterior NNP 32505 102 55 and and CC 32505 102 56 posterior posterior NN 32505 102 57 surfaces surface NNS 32505 102 58 ; ; , 32505 102 59 inner inner JJ 32505 102 60 surfaces surface NNS 32505 102 61 of of IN 32505 102 62 shanks shank NNS 32505 102 63 and and CC 32505 102 64 outer outer JJ 32505 102 65 surfaces surface NNS 32505 102 66 of of IN 32505 102 67 tarsi tarsi NNP 32505 102 68 barred bar VBN 32505 102 69 with with IN 32505 102 70 pale pale NNP 32505 102 71 brown brown NNP 32505 102 72 ; ; : 32505 102 73 dorsal dorsal NN 32505 102 74 surface surface NN 32505 102 75 of of IN 32505 102 76 foot foot NN 32505 102 77 mottled mottle VBN 32505 102 78 brown brown JJ 32505 102 79 and and CC 32505 102 80 creamy creamy VB 32505 102 81 white white JJ 32505 102 82 ; ; , 32505 102 83 ventral ventral JJ 32505 102 84 surface surface NN 32505 102 85 of of IN 32505 102 86 foot foot NN 32505 102 87 and and CC 32505 102 88 toes toe VBZ 32505 102 89 pale pale JJ 32505 102 90 brown brown NNP 32505 102 91 ; ; : 32505 102 92 forelimbs forelimb NNS 32505 102 93 faintly faintly RB 32505 102 94 barred bar VBN 32505 102 95 with with IN 32505 102 96 pale pale NNP 32505 102 97 brown brown NNP 32505 102 98 ; ; : 32505 102 99 belly belly NN 32505 102 100 white white JJ 32505 102 101 with with IN 32505 102 102 a a DT 32505 102 103 few few JJ 32505 102 104 pale pale JJ 32505 102 105 brown brown JJ 32505 102 106 spots spot NNS 32505 102 107 ; ; : 32505 102 108 vocal vocal JJ 32505 102 109 sac sac NN 32505 102 110 flecked fleck VBN 32505 102 111 with with IN 32505 102 112 pale pale JJ 32505 102 113 brown brown NNP 32505 102 114 ; ; : 32505 102 115 lower low JJR 32505 102 116 jaw jaw NN 32505 102 117 marked mark VBN 32505 102 118 with with IN 32505 102 119 small small JJ 32505 102 120 white white JJ 32505 102 121 spots spot NNS 32505 102 122 on on IN 32505 102 123 lips lip NNS 32505 102 124 . . . 32505 103 1 _ _ NNP 32505 103 2 Variation._--Geographic variation._--geographic JJ 32505 103 3 variation variation NN 32505 103 4 is be VBZ 32505 103 5 evident evident JJ 32505 103 6 in in IN 32505 103 7 the the DT 32505 103 8 snout snout NN 32505 103 9 - - HYPH 32505 103 10 vent vent NN 32505 103 11 length length NN 32505 103 12 , , , 32505 103 13 tibia tibia NNP 32505 103 14 length length NN 32505 103 15 , , , 32505 103 16 and and CC 32505 103 17 foot foot NN 32505 103 18 length length NN 32505 103 19 , , , 32505 103 20 all all RB 32505 103 21 in in IN 32505 103 22 relation relation NN 32505 103 23 to to TO 32505 103 24 snout snout VB 32505 103 25 - - HYPH 32505 103 26 vent vent NN 32505 103 27 length length NN 32505 103 28 , , , 32505 103 29 and and CC 32505 103 30 the the DT 32505 103 31 relative relative JJ 32505 103 32 size size NN 32505 103 33 of of IN 32505 103 34 the the DT 32505 103 35 tympanum tympanum NN 32505 103 36 to to IN 32505 103 37 the the DT 32505 103 38 eye eye NN 32505 103 39 ( ( -LRB- 32505 103 40 Table table NN 32505 103 41 1 1 CD 32505 103 42 ) ) -RRB- 32505 103 43 . . . 32505 104 1 The the DT 32505 104 2 largest large JJS 32505 104 3 specimens specimen NNS 32505 104 4 are be VBP 32505 104 5 from from IN 32505 104 6 the the DT 32505 104 7 humid humid JJ 32505 104 8 Pacific Pacific NNP 32505 104 9 lowlands lowland NNS 32505 104 10 of of IN 32505 104 11 Costa Costa NNP 32505 104 12 Rica Rica NNP 32505 104 13 ; ; : 32505 104 14 individuals individual NNS 32505 104 15 from from IN 32505 104 16 the the DT 32505 104 17 Caribbean caribbean JJ 32505 104 18 lowlands lowland NNS 32505 104 19 of of IN 32505 104 20 Costa Costa NNP 32505 104 21 Rica Rica NNP 32505 104 22 , , , 32505 104 23 Canal Canal NNP 32505 104 24 Zone Zone NNP 32505 104 25 , , , 32505 104 26 and and CC 32505 104 27 South South NNP 32505 104 28 America America NNP 32505 104 29 are be VBP 32505 104 30 smaller small JJR 32505 104 31 . . . 32505 105 1 A a DT 32505 105 2 general general JJ 32505 105 3 trend trend NN 32505 105 4 for for IN 32505 105 5 increase increase NN 32505 105 6 in in IN 32505 105 7 size size NN 32505 105 8 extends extend NNS 32505 105 9 from from IN 32505 105 10 South South NNP 32505 105 11 America America NNP 32505 105 12 to to IN 32505 105 13 the the DT 32505 105 14 Pacific Pacific NNP 32505 105 15 lowlands lowland NNS 32505 105 16 of of IN 32505 105 17 Costa Costa NNP 32505 105 18 Rica Rica NNP 32505 105 19 . . . 32505 106 1 Most Most JJS 32505 106 2 variation variation NN 32505 106 3 in in IN 32505 106 4 color color NN 32505 106 5 does do VBZ 32505 106 6 not not RB 32505 106 7 seem seem VB 32505 106 8 to to TO 32505 106 9 be be VB 32505 106 10 correlated correlate VBN 32505 106 11 with with IN 32505 106 12 geography geography NN 32505 106 13 ; ; : 32505 106 14 color color NN 32505 106 15 variation variation NN 32505 106 16 is be VBZ 32505 106 17 nearly nearly RB 32505 106 18 as as RB 32505 106 19 great great JJ 32505 106 20 within within IN 32505 106 21 a a DT 32505 106 22 given give VBN 32505 106 23 population population NN 32505 106 24 as as IN 32505 106 25 between between IN 32505 106 26 separated separated JJ 32505 106 27 populations population NNS 32505 106 28 . . . 32505 107 1 However however RB 32505 107 2 , , , 32505 107 3 most most JJS 32505 107 4 specimens specimen NNS 32505 107 5 from from IN 32505 107 6 Rincón Rincón NNP 32505 107 7 de de NNP 32505 107 8 Osa Osa NNP 32505 107 9 , , , 32505 107 10 Puntarenas Puntarenas NNP 32505 107 11 Province Province NNP 32505 107 12 , , , 32505 107 13 Costa Costa NNP 32505 107 14 Rica Rica NNP 32505 107 15 , , , 32505 107 16 are be VBP 32505 107 17 dusky dusky NNP 32505 107 18 brown brown NNP 32505 107 19 , , , 32505 107 20 but but CC 32505 107 21 a a DT 32505 107 22 few few JJ 32505 107 23 are be VBP 32505 107 24 darker dark JJR 32505 107 25 . . . 32505 108 1 In in IN 32505 108 2 comparison comparison NN 32505 108 3 with with IN 32505 108 4 specimens specimen NNS 32505 108 5 from from IN 32505 108 6 the the DT 32505 108 7 Caribbean caribbean JJ 32505 108 8 lowlands lowland NNS 32505 108 9 of of IN 32505 108 10 Central Central NNP 32505 108 11 America America NNP 32505 108 12 , , , 32505 108 13 specimens specimen VBZ 32505 108 14 from from IN 32505 108 15 the the DT 32505 108 16 Pacific Pacific NNP 32505 108 17 slopes slope NNS 32505 108 18 in in IN 32505 108 19 Costa Costa NNP 32505 108 20 Rica Rica NNP 32505 108 21 have have VBP 32505 108 22 a a DT 32505 108 23 darker dark JJR 32505 108 24 interorbital interorbital JJ 32505 108 25 triangle triangle NN 32505 108 26 . . . 32505 109 1 In in IN 32505 109 2 some some DT 32505 109 3 specimens specimen NNS 32505 109 4 from from IN 32505 109 5 the the DT 32505 109 6 latter latter JJ 32505 109 7 area area NN 32505 109 8 rugosities rugositie NNS 32505 109 9 are be VBP 32505 109 10 present present JJ 32505 109 11 on on IN 32505 109 12 the the DT 32505 109 13 borders border NNS 32505 109 14 of of IN 32505 109 15 the the DT 32505 109 16 interorbital interorbital JJ 32505 109 17 triangle triangle NN 32505 109 18 , , , 32505 109 19 on on IN 32505 109 20 the the DT 32505 109 21 snout snout NN 32505 109 22 , , , 32505 109 23 on on IN 32505 109 24 the the DT 32505 109 25 upper upper JJ 32505 109 26 eyelid eyelid NN 32505 109 27 , , , 32505 109 28 and and CC 32505 109 29 on on IN 32505 109 30 the the DT 32505 109 31 heel heel NN 32505 109 32 . . . 32505 110 1 Specimens specimen NNS 32505 110 2 from from IN 32505 110 3 the the DT 32505 110 4 Caribbean caribbean JJ 32505 110 5 lowlands lowland NNS 32505 110 6 are be VBP 32505 110 7 less less RBR 32505 110 8 tuberculate tuberculate JJ 32505 110 9 , , , 32505 110 10 and and CC 32505 110 11 most most JJS 32505 110 12 individuals individual NNS 32505 110 13 from from IN 32505 110 14 there there RB 32505 110 15 lack lack NN 32505 110 16 rugosities rugositie NNS 32505 110 17 on on IN 32505 110 18 the the DT 32505 110 19 tarsus tarsus NN 32505 110 20 . . . 32505 111 1 Living live VBG 32505 111 2 individuals individual NNS 32505 111 3 from from IN 32505 111 4 Puerto Puerto NNP 32505 111 5 Viejo Viejo NNP 32505 111 6 , , , 32505 111 7 Heredia Heredia NNP 32505 111 8 Province Province NNP 32505 111 9 , , , 32505 111 10 Costa Costa NNP 32505 111 11 Rica Rica NNP 32505 111 12 , , , 32505 111 13 and and CC 32505 111 14 from from IN 32505 111 15 the the DT 32505 111 16 Canal Canal NNP 32505 111 17 Zone Zone NNP 32505 111 18 , , , 32505 111 19 Panamá Panamá NNP 32505 111 20 , , , 32505 111 21 are be VBP 32505 111 22 brown brown JJ 32505 111 23 above above RB 32505 111 24 with with IN 32505 111 25 a a DT 32505 111 26 metallic metallic JJ 32505 111 27 green green JJ 32505 111 28 tint tint NN 32505 111 29 . . . 32505 112 1 Rugosities rugositie NNS 32505 112 2 are be VBP 32505 112 3 present present JJ 32505 112 4 on on IN 32505 112 5 the the DT 32505 112 6 posterior posterior NN 32505 112 7 edges edge NNS 32505 112 8 of of IN 32505 112 9 the the DT 32505 112 10 forelimbs forelimb NNS 32505 112 11 in in IN 32505 112 12 some some DT 32505 112 13 specimens specimen NNS 32505 112 14 from from IN 32505 112 15 throughout throughout IN 32505 112 16 the the DT 32505 112 17 range range NN 32505 112 18 . . . 32505 113 1 In in IN 32505 113 2 most most JJS 32505 113 3 respects respect NNS 32505 113 4 , , , 32505 113 5 specimens specimen VBZ 32505 113 6 from from IN 32505 113 7 the the DT 32505 113 8 Canal Canal NNP 32505 113 9 Zone Zone NNP 32505 113 10 resemble resemble VB 32505 113 11 those those DT 32505 113 12 from from IN 32505 113 13 the the DT 32505 113 14 Caribbean caribbean JJ 32505 113 15 lowlands lowland NNS 32505 113 16 of of IN 32505 113 17 Costa Costa NNP 32505 113 18 Rica Rica NNP 32505 113 19 more more JJR 32505 113 20 than than IN 32505 113 21 they -PRON- PRP 32505 113 22 resemble resemble VBP 32505 113 23 those those DT 32505 113 24 from from IN 32505 113 25 the the DT 32505 113 26 Pacific Pacific NNP 32505 113 27 lowlands lowland NNS 32505 113 28 of of IN 32505 113 29 Costa Costa NNP 32505 113 30 Rica Rica NNP 32505 113 31 , , , 32505 113 32 but but CC 32505 113 33 some some DT 32505 113 34 individuals individual NNS 32505 113 35 from from IN 32505 113 36 the the DT 32505 113 37 Canal Canal NNP 32505 113 38 Zone Zone NNP 32505 113 39 are be VBP 32505 113 40 less less RBR 32505 113 41 metallic metallic JJ 32505 113 42 above above RB 32505 113 43 and and CC 32505 113 44 have have VBP 32505 113 45 small small JJ 32505 113 46 white white JJ 32505 113 47 spots spot NNS 32505 113 48 dorsally dorsally RB 32505 113 49 on on IN 32505 113 50 the the DT 32505 113 51 body body NN 32505 113 52 , , , 32505 113 53 head head NN 32505 113 54 , , , 32505 113 55 and and CC 32505 113 56 limbs limb NNS 32505 113 57 . . . 32505 114 1 A a DT 32505 114 2 moderate moderate JJ 32505 114 3 amount amount NN 32505 114 4 of of IN 32505 114 5 color color NN 32505 114 6 change change NN 32505 114 7 from from IN 32505 114 8 night night NN 32505 114 9 to to IN 32505 114 10 day day NN 32505 114 11 has have VBZ 32505 114 12 been be VBN 32505 114 13 noted note VBN 32505 114 14 . . . 32505 115 1 At at IN 32505 115 2 night night NN 32505 115 3 , , , 32505 115 4 a a DT 32505 115 5 male male NN 32505 115 6 from from IN 32505 115 7 Puerto Puerto NNP 32505 115 8 Viejo Viejo NNP 32505 115 9 , , , 32505 115 10 Heredia Heredia NNP 32505 115 11 Province Province NNP 32505 115 12 , , , 32505 115 13 Costa Costa NNP 32505 115 14 Rica Rica NNP 32505 115 15 , , , 32505 115 16 was be VBD 32505 115 17 grayish grayish NNP 32505 115 18 tan tan NNP 32505 115 19 above above IN 32505 115 20 with with IN 32505 115 21 slightly slightly RB 32505 115 22 darker dark JJR 32505 115 23 markings marking NNS 32505 115 24 ; ; : 32505 115 25 the the DT 32505 115 26 flanks flank NNS 32505 115 27 were be VBD 32505 115 28 pale pale JJ 32505 115 29 yellowish yellowish JJ 32505 115 30 green green NN 32505 115 31 . . . 32505 116 1 By by IN 32505 116 2 day day NN 32505 116 3 , , , 32505 116 4 the the DT 32505 116 5 dorsum dorsum NN 32505 116 6 was be VBD 32505 116 7 brown brown JJ 32505 116 8 with with IN 32505 116 9 darker dark JJR 32505 116 10 markings marking NNS 32505 116 11 , , , 32505 116 12 and and CC 32505 116 13 the the DT 32505 116 14 throat throat NN 32505 116 15 was be VBD 32505 116 16 pale pale JJ 32505 116 17 gray gray JJ 32505 116 18 with with IN 32505 116 19 white white JJ 32505 116 20 flecks flecks NNPS 32505 116 21 ; ; : 32505 116 22 the the DT 32505 116 23 rest rest NN 32505 116 24 of of IN 32505 116 25 the the DT 32505 116 26 venter venter NN 32505 116 27 was be VBD 32505 116 28 creamy creamy JJ 32505 116 29 white white JJ 32505 116 30 . . . 32505 117 1 The the DT 32505 117 2 groin groin NN 32505 117 3 was be VBD 32505 117 4 pale pale JJ 32505 117 5 green green JJ 32505 117 6 with with IN 32505 117 7 black black JJ 32505 117 8 mottling mottling NN 32505 117 9 ; ; : 32505 117 10 the the DT 32505 117 11 anterior anterior NN 32505 117 12 and and CC 32505 117 13 posterior posterior NN 32505 117 14 surfaces surface NNS 32505 117 15 of of IN 32505 117 16 the the DT 32505 117 17 thighs thigh NNS 32505 117 18 and and CC 32505 117 19 inner inner JJ 32505 117 20 edges edge NNS 32505 117 21 of of IN 32505 117 22 the the DT 32505 117 23 tarsi tarsi NN 32505 117 24 were be VBD 32505 117 25 greenish greenish JJ 32505 117 26 yellow yellow JJ 32505 117 27 with with IN 32505 117 28 black black JJ 32505 117 29 bars bar NNS 32505 117 30 . . . 32505 118 1 TABLE TABLE NNP 32505 118 2 1.--Geographic 1.--geographic CD 32505 118 3 Variation Variation NNP 32505 118 4 in in IN 32505 118 5 Size Size NNP 32505 118 6 and and CC 32505 118 7 Proportions Proportions NNP 32505 118 8 in in IN 32505 118 9 Males Males NNPS 32505 118 10 of of IN 32505 118 11 _ _ NNP 32505 118 12 Hyla Hyla NNP 32505 118 13 boulengeri boulengeri NN 32505 118 14 _ _ NNP 32505 118 15 . . . 32505 119 1 ( ( -LRB- 32505 119 2 Means mean NNS 32505 119 3 in in IN 32505 119 4 parentheses parenthesis NNS 32505 119 5 below below IN 32505 119 6 observed observed JJ 32505 119 7 ranges range NNS 32505 119 8 . . . 32505 119 9 ) ) -RRB- 32505 120 1 = = NFP 32505 120 2 = = NFP 32505 120 3 = = SYM 32505 120 4 = = SYM 32505 120 5 = = SYM 32505 120 6 = = SYM 32505 120 7 = = SYM 32505 120 8 = = SYM 32505 120 9 = = SYM 32505 120 10 = = SYM 32505 120 11 = = SYM 32505 120 12 = = SYM 32505 120 13 = = SYM 32505 120 14 = = SYM 32505 120 15 = = SYM 32505 120 16 = = SYM 32505 120 17 = = SYM 32505 120 18 = = SYM 32505 120 19 = = SYM 32505 120 20 = = SYM 32505 120 21 = = SYM 32505 120 22 = = SYM 32505 120 23 = = SYM 32505 120 24 = = SYM 32505 120 25 = = SYM 32505 120 26 = = SYM 32505 120 27 = = SYM 32505 120 28 = = SYM 32505 120 29 = = SYM 32505 120 30 = = SYM 32505 120 31 = = SYM 32505 120 32 = = SYM 32505 120 33 = = SYM 32505 120 34 = = SYM 32505 120 35 = = SYM 32505 120 36 = = SYM 32505 120 37 = = SYM 32505 120 38 = = SYM 32505 120 39 = = SYM 32505 120 40 = = SYM 32505 120 41 = = SYM 32505 120 42 = = SYM 32505 120 43 = = SYM 32505 120 44 = = SYM 32505 120 45 = = SYM 32505 120 46 = = SYM 32505 120 47 = = SYM 32505 120 48 = = SYM 32505 120 49 = = SYM 32505 120 50 = = SYM 32505 120 51 = = SYM 32505 120 52 = = SYM 32505 120 53 = = SYM 32505 120 54 = = SYM 32505 120 55 = = SYM 32505 120 56 = = SYM 32505 120 57 = = SYM 32505 120 58 = = SYM 32505 120 59 = = SYM 32505 120 60 = = SYM 32505 120 61 = = SYM 32505 120 62 = = SYM 32505 120 63 = = SYM 32505 120 64 = = SYM 32505 120 65 = = SYM 32505 120 66 = = SYM 32505 120 67 = = SYM 32505 120 68 = = SYM 32505 120 69 = = SYM 32505 120 70 = = SYM 32505 120 71 = = SYM 32505 120 72 = = SYM 32505 120 73 = = SYM 32505 120 74 | | NNP 32505 120 75 | | NNP 32505 120 76 Snout snout NN 32505 120 77 - - HYPH 32505 120 78 vent| vent| NNP 32505 120 79 Tibia Tibia NNP 32505 120 80 | | NNP 32505 120 81 | | NNP 32505 120 82 | | CD 32505 120 83 | | CD 32505 120 84 length length NN 32505 120 85 | | NNP 32505 120 86 length/ length/ NN 32505 120 87 |Tympanum/|Foot |tympanum/|foot NN 32505 120 88 length/ length/ NN 32505 120 89 Locality Locality NNP 32505 120 90 | | NNP 32505 120 91 N N NNP 32505 120 92 | | NNP 32505 120 93 ( ( -LRB- 32505 120 94 mm mm NNP 32505 120 95 . . . 32505 120 96 ) ) -RRB- 32505 121 1 | | CD 32505 121 2 snout snout VBP 32505 121 3 - - HYPH 32505 121 4 vent| vent| CD 32505 121 5 eye eye NN 32505 121 6 | | HYPH 32505 121 7 snout snout NN 32505 121 8 - - HYPH 32505 121 9 vent vent NN 32505 121 10 --------------------+----+-----------+-----------+---------+------------- --------------------+----+-----------+-----------+---------+------------- NNP 32505 121 11 Costa Costa NNP 32505 121 12 Rica Rica NNP 32505 121 13 : : : 32505 121 14 Tilarán Tilarán NNP 32505 121 15 | | NNP 32505 121 16 23 23 CD 32505 121 17 | | CD 32505 121 18 37.4 37.4 CD 32505 121 19 - - HYPH 32505 121 20 48.7 48.7 CD 32505 121 21 | | CD 32505 121 22 0.52 0.52 CD 32505 121 23 - - HYPH 32505 121 24 0.58 0.58 CD 32505 121 25 |0.62 |0.62 CD 32505 121 26 - - HYPH 32505 121 27 0.76| 0.76| CD 32505 121 28 0.39 0.39 CD 32505 121 29 - - HYPH 32505 121 30 0.45 0.45 CD 32505 121 31 | | HYPH 32505 121 32 | | CD 32505 121 33 ( ( -LRB- 32505 121 34 43.8 43.8 CD 32505 121 35 ) ) -RRB- 32505 121 36 | | NNP 32505 121 37 ( ( -LRB- 32505 121 38 0.55 0.55 CD 32505 121 39 ) ) -RRB- 32505 121 40 | | NNP 32505 121 41 ( ( -LRB- 32505 121 42 0.71 0.71 CD 32505 121 43 ) ) -RRB- 32505 121 44 | | NNP 32505 121 45 ( ( -LRB- 32505 121 46 0.42 0.42 CD 32505 121 47 ) ) -RRB- 32505 121 48 | | CD 32505 121 49 | | CD 32505 121 50 | | CD 32505 121 51 | | CD 32505 121 52 | | NNP 32505 121 53 Costa Costa NNP 32505 121 54 Rica Rica NNP 32505 121 55 : : : 32505 121 56 Rincón Rincón NNP 32505 121 57 | | NNP 32505 121 58 10 10 CD 32505 121 59 | | CD 32505 121 60 41.4 41.4 CD 32505 121 61 - - HYPH 32505 121 62 46.1 46.1 CD 32505 121 63 | | CD 32505 121 64 0.54 0.54 CD 32505 121 65 - - HYPH 32505 121 66 0.60 0.60 CD 32505 121 67 |0.68 |0.68 CD 32505 121 68 - - HYPH 32505 121 69 0.80| 0.80| CD 32505 121 70 0.40 0.40 CD 32505 121 71 - - HYPH 32505 121 72 0.45 0.45 CD 32505 121 73 de de NNP 32505 121 74 Osa Osa NNP 32505 121 75 | | NNP 32505 121 76 | | NNP 32505 121 77 ( ( -LRB- 32505 121 78 44.0 44.0 CD 32505 121 79 ) ) -RRB- 32505 121 80 | | NNP 32505 121 81 ( ( -LRB- 32505 121 82 0.57 0.57 CD 32505 121 83 ) ) -RRB- 32505 121 84 | | NNP 32505 121 85 ( ( -LRB- 32505 121 86 0.74 0.74 CD 32505 121 87 ) ) -RRB- 32505 121 88 | | NNP 32505 121 89 ( ( -LRB- 32505 121 90 0.43 0.43 CD 32505 121 91 ) ) -RRB- 32505 121 92 | | CD 32505 121 93 | | CD 32505 121 94 | | CD 32505 121 95 | | CD 32505 121 96 | | NNP 32505 121 97 Costa Costa NNP 32505 121 98 Rica Rica NNP 32505 121 99 : : : 32505 121 100 Alajuela| alajuela| $ 32505 121 101 13 13 CD 32505 121 102 | | CD 32505 121 103 35.6 35.6 CD 32505 121 104 - - HYPH 32505 121 105 43.1 43.1 CD 32505 121 106 | | CD 32505 121 107 0.55 0.55 CD 32505 121 108 - - HYPH 32505 121 109 0.60 0.60 CD 32505 121 110 |0.63 |0.63 CD 32505 121 111 - - HYPH 32505 121 112 0.78| 0.78| CD 32505 121 113 0.41 0.41 CD 32505 121 114 - - HYPH 32505 121 115 0.46 0.46 CD 32505 121 116 Province Province NNP 32505 121 117 | | NNP 32505 121 118 | | NNP 32505 121 119 ( ( -LRB- 32505 121 120 39.8 39.8 CD 32505 121 121 ) ) -RRB- 32505 121 122 | | NNP 32505 121 123 ( ( -LRB- 32505 121 124 0.57 0.57 CD 32505 121 125 ) ) -RRB- 32505 121 126 | | NNP 32505 121 127 ( ( -LRB- 32505 121 128 0.69 0.69 CD 32505 121 129 ) ) -RRB- 32505 121 130 | | NNP 32505 121 131 ( ( -LRB- 32505 121 132 0.44 0.44 CD 32505 121 133 ) ) -RRB- 32505 121 134 | | NNP 32505 121 135 | | CD 32505 121 136 | | CD 32505 121 137 | | CD 32505 121 138 | | NNP 32505 121 139 Costa Costa NNP 32505 121 140 Rica Rica NNP 32505 121 141 : : : 32505 121 142 Puerto Puerto NNP 32505 121 143 | | CD 32505 121 144 25 25 CD 32505 121 145 | | CD 32505 121 146 37.5 37.5 CD 32505 121 147 - - HYPH 32505 121 148 42.9 42.9 CD 32505 121 149 | | CD 32505 121 150 0.51 0.51 CD 32505 121 151 - - HYPH 32505 121 152 0.62 0.62 CD 32505 121 153 |0.63 |0.63 NNS 32505 121 154 - - HYPH 32505 121 155 0.79| 0.79| NNS 32505 121 156 0.38 0.38 CD 32505 121 157 - - SYM 32505 121 158 0.46 0.46 CD 32505 121 159 Viejo Viejo NNP 32505 121 160 | | NNP 32505 121 161 | | NNP 32505 121 162 ( ( -LRB- 32505 121 163 41.6 41.6 CD 32505 121 164 ) ) -RRB- 32505 121 165 | | NNP 32505 121 166 ( ( -LRB- 32505 121 167 0.55 0.55 CD 32505 121 168 ) ) -RRB- 32505 121 169 | | NNP 32505 121 170 ( ( -LRB- 32505 121 171 0.71 0.71 CD 32505 121 172 ) ) -RRB- 32505 121 173 | | NNP 32505 121 174 ( ( -LRB- 32505 121 175 0.43 0.43 CD 32505 121 176 ) ) -RRB- 32505 121 177 | | CD 32505 121 178 | | CD 32505 121 179 | | CD 32505 121 180 | | CD 32505 121 181 | | NNP 32505 121 182 Costa Costa NNP 32505 121 183 Rica Rica NNP 32505 121 184 : : : 32505 121 185 Suretka Suretka NNP 32505 121 186 | | NNP 32505 121 187 9 9 CD 32505 121 188 | | CD 32505 121 189 38.7 38.7 CD 32505 121 190 - - HYPH 32505 121 191 42.0 42.0 CD 32505 121 192 | | CD 32505 121 193 0.56 0.56 CD 32505 121 194 - - HYPH 32505 121 195 0.58 0.58 CD 32505 121 196 |0.53 |0.53 CD 32505 121 197 - - HYPH 32505 121 198 0.72| 0.72| CD 32505 121 199 0.35 0.35 CD 32505 121 200 - - HYPH 32505 121 201 0.45 0.45 CD 32505 121 202 | | HYPH 32505 121 203 | | CD 32505 121 204 ( ( -LRB- 32505 121 205 41.0 41.0 CD 32505 121 206 ) ) -RRB- 32505 121 207 | | NNP 32505 121 208 ( ( -LRB- 32505 121 209 0.56 0.56 CD 32505 121 210 ) ) -RRB- 32505 121 211 | | NNP 32505 121 212 ( ( -LRB- 32505 121 213 0.62 0.62 CD 32505 121 214 ) ) -RRB- 32505 121 215 | | NNP 32505 121 216 ( ( -LRB- 32505 121 217 0.42 0.42 CD 32505 121 218 ) ) -RRB- 32505 121 219 | | CD 32505 121 220 | | CD 32505 121 221 | | CD 32505 121 222 | | CD 32505 121 223 | | NNP 32505 121 224 Panamá Panamá NNPS 32505 121 225 : : : 32505 121 226 Canal Canal NNP 32505 121 227 Zone Zone NNP 32505 121 228 | | NNP 32505 121 229 16 16 CD 32505 121 230 | | CD 32505 121 231 36.7 36.7 CD 32505 121 232 - - HYPH 32505 121 233 42.9 42.9 CD 32505 121 234 | | CD 32505 121 235 0.52 0.52 CD 32505 121 236 - - HYPH 32505 121 237 0.58 0.58 CD 32505 121 238 |0.70 |0.70 CD 32505 121 239 - - HYPH 32505 121 240 0.78| 0.78| CD 32505 121 241 0.40 0.40 CD 32505 121 242 - - HYPH 32505 121 243 0.44 0.44 CD 32505 121 244 | | CD 32505 121 245 | | CD 32505 121 246 ( ( -LRB- 32505 121 247 39.0 39.0 CD 32505 121 248 ) ) -RRB- 32505 121 249 | | NNP 32505 121 250 ( ( -LRB- 32505 121 251 0.54 0.54 CD 32505 121 252 ) ) -RRB- 32505 121 253 | | NNP 32505 121 254 ( ( -LRB- 32505 121 255 0.74 0.74 CD 32505 121 256 ) ) -RRB- 32505 121 257 | | NNP 32505 121 258 ( ( -LRB- 32505 121 259 0.42 0.42 CD 32505 121 260 ) ) -RRB- 32505 121 261 | | CD 32505 121 262 | | CD 32505 121 263 | | CD 32505 121 264 | | CD 32505 121 265 | | NNP 32505 121 266 Venezuela Venezuela NNP 32505 121 267 : : : 32505 121 268 Santomé Santomé NNP 32505 121 269 | | NNP 32505 121 270 4 4 CD 32505 121 271 | | CD 32505 121 272 35.5 35.5 CD 32505 121 273 - - HYPH 32505 121 274 40.9 40.9 CD 32505 121 275 | | CD 32505 121 276 0.45 0.45 CD 32505 121 277 - - HYPH 32505 121 278 0.48 0.48 CD 32505 121 279 |0.63 |0.63 CD 32505 121 280 - - HYPH 32505 121 281 0.67| 0.67| CD 32505 121 282 0.36 0.36 CD 32505 121 283 - - HYPH 32505 121 284 0.40 0.40 CD 32505 121 285 | | HYPH 32505 121 286 | | CD 32505 121 287 ( ( -LRB- 32505 121 288 38.5 38.5 CD 32505 121 289 ) ) -RRB- 32505 121 290 | | NNP 32505 121 291 ( ( -LRB- 32505 121 292 0.46 0.46 CD 32505 121 293 ) ) -RRB- 32505 121 294 | | NNP 32505 121 295 ( ( -LRB- 32505 121 296 0.65 0.65 CD 32505 121 297 ) ) -RRB- 32505 121 298 | | NNP 32505 121 299 ( ( -LRB- 32505 121 300 0.38 0.38 CD 32505 121 301 ) ) -RRB- 32505 121 302 TABLE TABLE NNP 32505 121 303 2.--Comparison 2.--comparison CD 32505 121 304 of of IN 32505 121 305 Mating Mating NNP 32505 121 306 Calls Calls NNPS 32505 121 307 in in IN 32505 121 308 the the DT 32505 121 309 _ _ NNP 32505 121 310 Hyla Hyla NNP 32505 121 311 rubra rubra NN 32505 121 312 _ _ NNP 32505 121 313 Group Group NNP 32505 121 314 . . . 32505 122 1 ( ( -LRB- 32505 122 2 Means mean NNS 32505 122 3 in in IN 32505 122 4 parentheses parenthesis NNS 32505 122 5 below below IN 32505 122 6 observed observed JJ 32505 122 7 ranges range NNS 32505 122 8 . . . 32505 122 9 ) ) -RRB- 32505 123 1 = = NFP 32505 123 2 = = NFP 32505 123 3 = = SYM 32505 123 4 = = SYM 32505 123 5 = = SYM 32505 123 6 = = SYM 32505 123 7 = = SYM 32505 123 8 = = SYM 32505 123 9 = = SYM 32505 123 10 = = SYM 32505 123 11 = = SYM 32505 123 12 = = SYM 32505 123 13 = = SYM 32505 123 14 = = SYM 32505 123 15 = = SYM 32505 123 16 = = SYM 32505 123 17 = = SYM 32505 123 18 = = SYM 32505 123 19 = = SYM 32505 123 20 = = SYM 32505 123 21 = = SYM 32505 123 22 = = SYM 32505 123 23 = = SYM 32505 123 24 = = SYM 32505 123 25 = = SYM 32505 123 26 = = SYM 32505 123 27 = = SYM 32505 123 28 = = SYM 32505 123 29 = = SYM 32505 123 30 = = SYM 32505 123 31 = = SYM 32505 123 32 = = SYM 32505 123 33 = = SYM 32505 123 34 = = SYM 32505 123 35 = = SYM 32505 123 36 = = SYM 32505 123 37 = = SYM 32505 123 38 = = SYM 32505 123 39 = = SYM 32505 123 40 = = SYM 32505 123 41 = = SYM 32505 123 42 = = SYM 32505 123 43 = = SYM 32505 123 44 = = SYM 32505 123 45 = = SYM 32505 123 46 = = SYM 32505 123 47 = = SYM 32505 123 48 = = SYM 32505 123 49 = = SYM 32505 123 50 = = SYM 32505 123 51 = = SYM 32505 123 52 = = SYM 32505 123 53 = = SYM 32505 123 54 = = SYM 32505 123 55 = = SYM 32505 123 56 = = SYM 32505 123 57 = = SYM 32505 123 58 = = SYM 32505 123 59 = = SYM 32505 123 60 = = SYM 32505 123 61 = = SYM 32505 123 62 = = SYM 32505 123 63 = = SYM 32505 123 64 = = SYM 32505 123 65 = = SYM 32505 123 66 = = SYM 32505 123 67 = = SYM 32505 123 68 = = SYM 32505 123 69 = = SYM 32505 123 70 = = SYM 32505 123 71 = = SYM 32505 123 72 = = SYM 32505 123 73 = = SYM 32505 123 74 = = SYM 32505 123 75 = = SYM 32505 123 76 = = SYM 32505 123 77 | | CD 32505 123 78 |Notes| |notes| JJ 32505 123 79 | | CD 32505 123 80 | | CD 32505 123 81 | | CD 32505 123 82 Major major JJ 32505 123 83 frequencies frequency NNS 32505 123 84 | | CD 32505 123 85 | | NNS 32505 123 86 per per IN 32505 123 87 |Duration| |duration| NN 32505 123 88 Pulses|Fundamental| Pulses|Fundamental| '' 32505 123 89 ( ( -LRB- 32505 123 90 cps cps NNP 32505 123 91 ) ) -RRB- 32505 123 92 | | NNP 32505 123 93 |call |call CD 32505 123 94 | | NNS 32505 123 95 of of IN 32505 123 96 note| note| CD 32505 123 97 per per IN 32505 123 98 | | CD 32505 123 99 frequency frequency NN 32505 123 100 |--------------------- |--------------------- NNP 32505 123 101 Species Species NNP 32505 123 102 | | NNP 32505 123 103 N N NNP 32505 123 104 |group| |group| NNP 32505 123 105 ( ( -LRB- 32505 123 106 sec sec NNP 32505 123 107 . . . 32505 123 108 ) ) -RRB- 32505 124 1 | | NNP 32505 124 2 second| second| NFP 32505 124 3 ( ( -LRB- 32505 124 4 cps cps NN 32505 124 5 ) ) -RRB- 32505 124 6 | | NNP 32505 124 7 Lower low JJR 32505 124 8 | | NNP 32505 124 9 Upper Upper NNP 32505 124 10 ---------------+---+----+---------+-------+-----------+---------+----------- ---------------+---+----+---------+-------+-----------+---------+----------- NNP 32505 124 11 H. H. NNP 32505 124 12 boulengeri boulengeri NN 32505 124 13 | | CD 32505 124 14 8 8 CD 32505 124 15 | | CD 32505 124 16 1 1 CD 32505 124 17 | | CD 32505 124 18 0.24- 0.24- CD 32505 124 19 | | CD 32505 124 20 80 80 CD 32505 124 21 - - HYPH 32505 124 22 120| 120| CD 32505 124 23 70 70 CD 32505 124 24 - - SYM 32505 124 25 74 74 CD 32505 124 26 |1400 |1400 NNP 32505 124 27 - - HYPH 32505 124 28 1820|2520 1820|2520 CD 32505 124 29 - - HYPH 32505 124 30 3182 3182 CD 32505 124 31 | | NNP 32505 124 32 | | CD 32505 124 33 | | CD 32505 124 34 0.47 0.47 CD 32505 124 35 | | NNS 32505 124 36 ( ( -LRB- 32505 124 37 101)| 101)| CD 32505 124 38 ( ( -LRB- 32505 124 39 71 71 CD 32505 124 40 ) ) -RRB- 32505 124 41 | | NNP 32505 124 42 ( ( -LRB- 32505 124 43 1611 1611 CD 32505 124 44 ) ) -RRB- 32505 124 45 | | NNP 32505 124 46 ( ( -LRB- 32505 124 47 2840 2840 CD 32505 124 48 ) ) -RRB- 32505 124 49 | | NNP 32505 124 50 | | CD 32505 124 51 | | NNS 32505 124 52 ( ( -LRB- 32505 124 53 0.35 0.35 CD 32505 124 54 ) ) -RRB- 32505 124 55 | | NNP 32505 124 56 | | CD 32505 124 57 | | CD 32505 124 58 | | CD 32505 124 59 | | CD 32505 124 60 | | CD 32505 124 61 | | CD 32505 124 62 | | CD 32505 124 63 | | CD 32505 124 64 | | CD 32505 124 65 | | NNP 32505 124 66 H. H. NNP 32505 124 67 foliamorta foliamorta NNP 32505 124 68 | | NNP 32505 124 69 7 7 CD 32505 124 70 | | CD 32505 124 71 1 1 CD 32505 124 72 | | CD 32505 124 73 0.23- 0.23- CD 32505 124 74 | | CD 32505 124 75 50 50 CD 32505 124 76 - - HYPH 32505 124 77 60 60 CD 32505 124 78 | | CD 32505 124 79 52 52 CD 32505 124 80 - - HYPH 32505 124 81 61 61 CD 32505 124 82 | | CD 32505 124 83 912 912 CD 32505 124 84 - - HYPH 32505 124 85 1026|2736 1026|2736 CD 32505 124 86 - - HYPH 32505 124 87 3477 3477 CD 32505 124 88 | | NNP 32505 124 89 | | CD 32505 124 90 | | CD 32505 124 91 1.86 1.86 CD 32505 124 92 | | NNS 32505 124 93 ( ( -LRB- 32505 124 94 51 51 CD 32505 124 95 ) ) -RRB- 32505 124 96 | | NNP 32505 124 97 ( ( -LRB- 32505 124 98 56 56 CD 32505 124 99 ) ) -RRB- 32505 124 100 | | NNP 32505 124 101 ( ( -LRB- 32505 124 102 918 918 CD 32505 124 103 ) ) -RRB- 32505 124 104 | | NNP 32505 124 105 ( ( -LRB- 32505 124 106 3055 3055 CD 32505 124 107 ) ) -RRB- 32505 124 108 | | CD 32505 124 109 | | CD 32505 124 110 | | CD 32505 124 111 ( ( -LRB- 32505 124 112 0.69 0.69 CD 32505 124 113 ) ) -RRB- 32505 124 114 | | NNP 32505 124 115 | | CD 32505 124 116 | | CD 32505 124 117 | | CD 32505 124 118 | | CD 32505 124 119 | | CD 32505 124 120 | | CD 32505 124 121 | | CD 32505 124 122 | | CD 32505 124 123 | | CD 32505 124 124 | | NNP 32505 124 125 H. H. NNP 32505 124 126 elaeochroa elaeochroa NNP 32505 124 127 |15 |15 CD 32505 124 128 | | CD 32505 124 129 2 2 CD 32505 124 130 - - SYM 32505 124 131 95| 95| CD 32505 124 132 0.12- 0.12- CD 32505 124 133 | | CD 32505 124 134 40 40 CD 32505 124 135 - - HYPH 32505 124 136 50 50 CD 32505 124 137 | | CD 32505 124 138 48 48 CD 32505 124 139 - - HYPH 32505 124 140 65 65 CD 32505 124 141 |1254 |1254 NNP 32505 124 142 - - HYPH 32505 124 143 1586|2562 1586|2562 CD 32505 124 144 - - HYPH 32505 124 145 3477 3477 CD 32505 124 146 | | NNP 32505 124 147 | | CD 32505 124 148 ( ( -LRB- 32505 124 149 19)| 19)| CD 32505 124 150 0.24 0.24 CD 32505 124 151 | | NNS 32505 124 152 ( ( -LRB- 32505 124 153 42 42 CD 32505 124 154 ) ) -RRB- 32505 124 155 | | NNP 32505 124 156 ( ( -LRB- 32505 124 157 57 57 CD 32505 124 158 ) ) -RRB- 32505 124 159 | | NNP 32505 124 160 ( ( -LRB- 32505 124 161 1499 1499 CD 32505 124 162 ) ) -RRB- 32505 124 163 | | NNP 32505 124 164 ( ( -LRB- 32505 124 165 2911 2911 CD 32505 124 166 ) ) -RRB- 32505 124 167 | | NNP 32505 124 168 | | CD 32505 124 169 | | NNS 32505 124 170 ( ( -LRB- 32505 124 171 0.17 0.17 CD 32505 124 172 ) ) -RRB- 32505 124 173 | | NNP 32505 124 174 | | CD 32505 124 175 | | CD 32505 124 176 | | CD 32505 124 177 | | CD 32505 124 178 | | CD 32505 124 179 | | CD 32505 124 180 | | CD 32505 124 181 | | CD 32505 124 182 | | CD 32505 124 183 | | NNP 32505 124 184 H. H. NNP 32505 124 185 s. s. NNP 32505 124 186 staufferi|18 staufferi|18 NNP 32505 124 187 | | NNP 32505 124 188 2 2 CD 32505 124 189 - - HYPH 32505 124 190 77| 77| CD 32505 124 191 0.13- 0.13- CD 32505 124 192 |100 |100 CD 32505 124 193 - - HYPH 32505 124 194 130| 130| CD 32505 124 195 96 96 CD 32505 124 196 - - SYM 32505 124 197 130 130 CD 32505 124 198 |1582 |1582 NNP 32505 124 199 - - HYPH 32505 124 200 1872|1962 1872|1962 CD 32505 124 201 - - HYPH 32505 124 202 3744 3744 CD 32505 124 203 | | CD 32505 124 204 | | CD 32505 124 205 ( ( -LRB- 32505 124 206 23)| 23)| CD 32505 124 207 0.23 0.23 CD 32505 124 208 | | NNS 32505 124 209 ( ( -LRB- 32505 124 210 120 120 CD 32505 124 211 ) ) -RRB- 32505 124 212 | | NNP 32505 124 213 ( ( -LRB- 32505 124 214 106 106 CD 32505 124 215 ) ) -RRB- 32505 124 216 | | NNP 32505 124 217 ( ( -LRB- 32505 124 218 1743 1743 CD 32505 124 219 ) ) -RRB- 32505 124 220 | | NNP 32505 124 221 ( ( -LRB- 32505 124 222 3056 3056 CD 32505 124 223 ) ) -RRB- 32505 124 224 | | NNP 32505 124 225 | | CD 32505 124 226 | | CD 32505 124 227 ( ( -LRB- 32505 124 228 0.18 0.18 CD 32505 124 229 ) ) -RRB- 32505 124 230 | | CD 32505 124 231 | | CD 32505 124 232 | | CD 32505 124 233 | | CD 32505 124 234 | | CD 32505 124 235 | | CD 32505 124 236 | | CD 32505 124 237 | | CD 32505 124 238 | | CD 32505 124 239 | | CD 32505 124 240 | | NNP 32505 124 241 H. H. NNP 32505 124 242 s. s. NNP 32505 124 243 altae altae NNP 32505 124 244 | | NNP 32505 124 245 7 7 CD 32505 124 246 | | CD 32505 124 247 2 2 CD 32505 124 248 - - HYPH 32505 124 249 22| 22| CD 32505 124 250 0.14- 0.14- CD 32505 124 251 |110 |110 JJ 32505 124 252 - - HYPH 32505 124 253 130| 130| CD 32505 124 254 104 104 CD 32505 124 255 - - HYPH 32505 124 256 117 117 CD 32505 124 257 |1853 |1853 NNP 32505 124 258 - - HYPH 32505 124 259 2106|3379 2106|3379 CD 32505 124 260 - - HYPH 32505 124 261 4056 4056 CD 32505 124 262 | | CD 32505 124 263 | | CD 32505 124 264 ( ( -LRB- 32505 124 265 11)| 11)| CD 32505 124 266 0.18 0.18 CD 32505 124 267 | | NNS 32505 124 268 ( ( -LRB- 32505 124 269 120 120 CD 32505 124 270 ) ) -RRB- 32505 124 271 | | NNP 32505 124 272 ( ( -LRB- 32505 124 273 112 112 CD 32505 124 274 ) ) -RRB- 32505 124 275 | | NNP 32505 124 276 ( ( -LRB- 32505 124 277 2008 2008 CD 32505 124 278 ) ) -RRB- 32505 124 279 | | NNP 32505 124 280 ( ( -LRB- 32505 124 281 3775 3775 CD 32505 124 282 ) ) -RRB- 32505 124 283 | | CD 32505 124 284 | | CD 32505 124 285 | | CD 32505 124 286 ( ( -LRB- 32505 124 287 0.15 0.15 CD 32505 124 288 ) ) -RRB- 32505 124 289 | | CD 32505 124 290 | | CD 32505 124 291 | | CD 32505 124 292 | | CD 32505 124 293 _ _ NNP 32505 124 294 Cranial Cranial NNP 32505 124 295 Osteology._--The Osteology._--The NNP 32505 124 296 skull skull NN 32505 124 297 of of IN 32505 124 298 _ _ NNP 32505 124 299 Hyla Hyla NNP 32505 124 300 boulengeri boulengeri NN 32505 124 301 _ _ NNP 32505 124 302 is be VBZ 32505 124 303 as as RB 32505 124 304 long long RB 32505 124 305 as as IN 32505 124 306 it -PRON- PRP 32505 124 307 is be VBZ 32505 124 308 wide wide JJ 32505 124 309 , , , 32505 124 310 and and CC 32505 124 311 is be VBZ 32505 124 312 flat flat JJ 32505 124 313 ; ; : 32505 124 314 the the DT 32505 124 315 premaxillary premaxillary NN 32505 124 316 is be VBZ 32505 124 317 small small JJ 32505 124 318 and and CC 32505 124 319 bears bear VBZ 32505 124 320 13 13 CD 32505 124 321 to to TO 32505 124 322 17 17 CD 32505 124 323 teeth tooth NNS 32505 124 324 ( ( -LRB- 32505 124 325 mean mean VB 32505 124 326 for for IN 32505 124 327 6 6 CD 32505 124 328 specimens specimen NNS 32505 124 329 , , , 32505 124 330 14.9 14.9 CD 32505 124 331 ) ) -RRB- 32505 124 332 . . . 32505 125 1 The the DT 32505 125 2 alary alary JJ 32505 125 3 processes process NNS 32505 125 4 of of IN 32505 125 5 the the DT 32505 125 6 premaxillaries premaxillarie NNS 32505 125 7 are be VBP 32505 125 8 widely widely RB 32505 125 9 separated separate VBN 32505 125 10 , , , 32505 125 11 concave concave NNP 32505 125 12 posteriorly posteriorly RB 32505 125 13 , , , 32505 125 14 and and CC 32505 125 15 vertical vertical JJ 32505 125 16 . . . 32505 126 1 Ventrally ventrally RB 32505 126 2 , , , 32505 126 3 the the DT 32505 126 4 premaxillary premaxillary NN 32505 126 5 is be VBZ 32505 126 6 connected connect VBN 32505 126 7 to to IN 32505 126 8 the the DT 32505 126 9 prevomer prevomer NN 32505 126 10 by by IN 32505 126 11 bony bony NNP 32505 126 12 tissues tissue NNS 32505 126 13 . . . 32505 127 1 The the DT 32505 127 2 maxillary maxillary NN 32505 127 3 is be VBZ 32505 127 4 slender slend JJR 32505 127 5 and and CC 32505 127 6 bears bear VBZ 32505 127 7 70 70 CD 32505 127 8 to to TO 32505 127 9 91 91 CD 32505 127 10 teeth tooth NNS 32505 127 11 ( ( -LRB- 32505 127 12 mean mean VB 32505 127 13 for for IN 32505 127 14 6 6 CD 32505 127 15 specimens specimen NNS 32505 127 16 78.1 78.1 CD 32505 127 17 ) ) -RRB- 32505 127 18 . . . 32505 128 1 The the DT 32505 128 2 pars par NNS 32505 128 3 facialis faciali NNS 32505 128 4 of of IN 32505 128 5 the the DT 32505 128 6 maxillary maxillary NN 32505 128 7 is be VBZ 32505 128 8 laterally laterally RB 32505 128 9 convex convex NNP 32505 128 10 and and CC 32505 128 11 about about RB 32505 128 12 four four CD 32505 128 13 times time NNS 32505 128 14 as as RB 32505 128 15 high high RB 32505 128 16 as as IN 32505 128 17 the the DT 32505 128 18 pars par NNS 32505 128 19 dentalis dentalis NNP 32505 128 20 . . . 32505 129 1 The the DT 32505 129 2 nasal nasal NN 32505 129 3 is be VBZ 32505 129 4 large large JJ 32505 129 5 ( ( -LRB- 32505 129 6 its -PRON- PRP$ 32505 129 7 length length NN 32505 129 8 about about RB 32505 129 9 40 40 CD 32505 129 10 per per NN 32505 129 11 cent cent NN 32505 129 12 of of IN 32505 129 13 total total JJ 32505 129 14 length length NN 32505 129 15 of of IN 32505 129 16 skull skull NN 32505 129 17 ) ) -RRB- 32505 129 18 , , , 32505 129 19 and and CC 32505 129 20 pointed point VBD 32505 129 21 anteriorly anteriorly RB 32505 129 22 and and CC 32505 129 23 posteriorly posteriorly RB 32505 129 24 in in IN 32505 129 25 dorsal dorsal NN 32505 129 26 view view NN 32505 129 27 . . . 32505 130 1 The the DT 32505 130 2 nasals nasal NNS 32505 130 3 are be VBP 32505 130 4 separated separate VBN 32505 130 5 anteromedially anteromedially RB 32505 130 6 by by IN 32505 130 7 the the DT 32505 130 8 cartilaginous cartilaginous JJ 32505 130 9 septum septum NN 32505 130 10 nasi nasi NNP 32505 130 11 . . . 32505 131 1 One one CD 32505 131 2 or or CC 32505 131 3 two two CD 32505 131 4 protuberances protuberance NNS 32505 131 5 are be VBP 32505 131 6 present present JJ 32505 131 7 on on IN 32505 131 8 the the DT 32505 131 9 midlateral midlateral JJ 32505 131 10 concavity concavity NN 32505 131 11 of of IN 32505 131 12 the the DT 32505 131 13 nasal nasal NN 32505 131 14 . . . 32505 132 1 Posteriorly posteriorly RB 32505 132 2 , , , 32505 132 3 the the DT 32505 132 4 nasal nasal NN 32505 132 5 overlaps overlap VBZ 32505 132 6 the the DT 32505 132 7 sphenethmoid sphenethmoid NN 32505 132 8 and and CC 32505 132 9 articulates articulate VBZ 32505 132 10 with with IN 32505 132 11 the the DT 32505 132 12 palatine palatine NN 32505 132 13 . . . 32505 133 1 Dorsally dorsally RB 32505 133 2 the the DT 32505 133 3 sphenethmoid sphenethmoid NN 32505 133 4 is be VBZ 32505 133 5 large large JJ 32505 133 6 , , , 32505 133 7 pentagonal pentagonal JJ 32505 133 8 , , , 32505 133 9 and and CC 32505 133 10 completely completely RB 32505 133 11 ossified ossified JJ 32505 133 12 . . . 32505 134 1 The the DT 32505 134 2 frontoparietal frontoparietal NN 32505 134 3 is be VBZ 32505 134 4 elongate elongate NN 32505 134 5 , , , 32505 134 6 smooth smooth JJ 32505 134 7 , , , 32505 134 8 and and CC 32505 134 9 bears bear VBZ 32505 134 10 a a DT 32505 134 11 small small JJ 32505 134 12 supraorbital supraorbital JJ 32505 134 13 process process NN 32505 134 14 on on IN 32505 134 15 the the DT 32505 134 16 anterior anterior JJ 32505 134 17 edge edge NN 32505 134 18 of of IN 32505 134 19 the the DT 32505 134 20 orbit orbit NN 32505 134 21 . . . 32505 135 1 A a DT 32505 135 2 keyhole keyhole NN 32505 135 3 - - HYPH 32505 135 4 shaped shaped JJ 32505 135 5 frontoparietal frontoparietal JJ 32505 135 6 fontanelle fontanelle NN 32505 135 7 is be VBZ 32505 135 8 present present JJ 32505 135 9 ; ; : 32505 135 10 the the DT 32505 135 11 fontanelle fontanelle NN 32505 135 12 is be VBZ 32505 135 13 narrow narrow JJ 32505 135 14 anteriorly anteriorly RB 32505 135 15 and and CC 32505 135 16 wide wide RB 32505 135 17 posteriorly posteriorly RB 32505 135 18 . . . 32505 136 1 The the DT 32505 136 2 bony bony NN 32505 136 3 part part NN 32505 136 4 of of IN 32505 136 5 the the DT 32505 136 6 proötic proötic NN 32505 136 7 is be VBZ 32505 136 8 separated separate VBN 32505 136 9 dorsally dorsally RB 32505 136 10 from from IN 32505 136 11 the the DT 32505 136 12 squamosal squamosal NN 32505 136 13 by by IN 32505 136 14 the the DT 32505 136 15 cartilaginous cartilaginous JJ 32505 136 16 crista crista NNP 32505 136 17 parotica parotica NNP 32505 136 18 . . . 32505 137 1 The the DT 32505 137 2 squamosal squamosal NN 32505 137 3 is be VBZ 32505 137 4 small small JJ 32505 137 5 , , , 32505 137 6 its -PRON- PRP$ 32505 137 7 anterior anterior JJ 32505 137 8 arm arm NN 32505 137 9 slender slender NN 32505 137 10 and and CC 32505 137 11 pointed point VBD 32505 137 12 . . . 32505 138 1 The the DT 32505 138 2 posterior posterior JJ 32505 138 3 arm arm NN 32505 138 4 of of IN 32505 138 5 the the DT 32505 138 6 squamosal squamosal NN 32505 138 7 is be VBZ 32505 138 8 pointed point VBN 32505 138 9 terminally terminally RB 32505 138 10 and and CC 32505 138 11 articulates articulate NNS 32505 138 12 with with IN 32505 138 13 the the DT 32505 138 14 proötic proötic NN 32505 138 15 medially medially RB 32505 138 16 . . . 32505 139 1 The the DT 32505 139 2 prevomer prevomer NN 32505 139 3 is be VBZ 32505 139 4 large large JJ 32505 139 5 and and CC 32505 139 6 elongate elongate VB 32505 139 7 . . . 32505 140 1 Anteriorly anteriorly RB 32505 140 2 the the DT 32505 140 3 prevomer prevomer NN 32505 140 4 is be VBZ 32505 140 5 connected connect VBN 32505 140 6 to to IN 32505 140 7 the the DT 32505 140 8 maxillary maxillary JJ 32505 140 9 - - HYPH 32505 140 10 premaxillary premaxillary NN 32505 140 11 articulation articulation NN 32505 140 12 ; ; , 32505 140 13 posteriorly posteriorly RB 32505 140 14 , , , 32505 140 15 the the DT 32505 140 16 prevomer prevomer NN 32505 140 17 is be VBZ 32505 140 18 separated separate VBN 32505 140 19 from from IN 32505 140 20 the the DT 32505 140 21 sphenethmoid sphenethmoid NN 32505 140 22 by by IN 32505 140 23 cartilage cartilage NN 32505 140 24 . . . 32505 141 1 Each each DT 32505 141 2 prevomer prevomer NN 32505 141 3 bears bear VBZ 32505 141 4 six six CD 32505 141 5 to to TO 32505 141 6 nine nine CD 32505 141 7 teeth tooth NNS 32505 141 8 . . . 32505 142 1 The the DT 32505 142 2 palatine palatine NN 32505 142 3 is be VBZ 32505 142 4 present present JJ 32505 142 5 and and CC 32505 142 6 edentate edentate NN 32505 142 7 . . . 32505 143 1 The the DT 32505 143 2 anterior anterior JJ 32505 143 3 end end NN 32505 143 4 of of IN 32505 143 5 the the DT 32505 143 6 parasphenoid parasphenoid NN 32505 143 7 is be VBZ 32505 143 8 broad broad JJ 32505 143 9 ( ( -LRB- 32505 143 10 less less RBR 32505 143 11 pointed pointed JJ 32505 143 12 than than IN 32505 143 13 in in IN 32505 143 14 _ _ NNP 32505 143 15 Hyla Hyla NNP 32505 143 16 foliamorta foliamorta NN 32505 143 17 _ _ NNP 32505 143 18 ) ) -RRB- 32505 143 19 . . . 32505 144 1 The the DT 32505 144 2 pterygoid pterygoid NN 32505 144 3 is be VBZ 32505 144 4 slender slend JJR 32505 144 5 and and CC 32505 144 6 well well RB 32505 144 7 developed developed JJ 32505 144 8 . . . 32505 145 1 _ _ NNP 32505 145 2 Natural Natural NNP 32505 145 3 History._--_Hyla History._--_Hyla NNP 32505 145 4 boulengeri boulengeri NN 32505 145 5 _ _ NNP 32505 145 6 inhabits inhabit VBZ 32505 145 7 humid humid NN 32505 145 8 lowland lowland NN 32505 145 9 tropical tropical JJ 32505 145 10 forests forest NNS 32505 145 11 and and CC 32505 145 12 breeds breed NNS 32505 145 13 in in IN 32505 145 14 temporary temporary JJ 32505 145 15 ponds pond NNS 32505 145 16 . . . 32505 146 1 Clasping clasp VBG 32505 146 2 pairs pair NNS 32505 146 3 and and CC 32505 146 4 gravid gravid JJ 32505 146 5 females female NNS 32505 146 6 were be VBD 32505 146 7 observed observe VBN 32505 146 8 at at IN 32505 146 9 Puerto Puerto NNP 32505 146 10 Viejo Viejo NNP 32505 146 11 , , , 32505 146 12 Heredia Heredia NNP 32505 146 13 Province Province NNP 32505 146 14 , , , 32505 146 15 Costa Costa NNP 32505 146 16 Rica Rica NNP 32505 146 17 , , , 32505 146 18 on on IN 32505 146 19 June June NNP 32505 146 20 21 21 CD 32505 146 21 , , , 32505 146 22 1966 1966 CD 32505 146 23 . . . 32505 147 1 Males male NNS 32505 147 2 were be VBD 32505 147 3 calling call VBG 32505 147 4 from from IN 32505 147 5 depressions depression NNS 32505 147 6 in in IN 32505 147 7 decaying decay VBG 32505 147 8 logs log NNS 32505 147 9 and and CC 32505 147 10 stumps stump VBZ 32505 147 11 , , , 32505 147 12 in in IN 32505 147 13 forked forked JJ 32505 147 14 stems stem NNS 32505 147 15 , , , 32505 147 16 and and CC 32505 147 17 from from IN 32505 147 18 leaves leave NNS 32505 147 19 of of IN 32505 147 20 broad broad JJ 32505 147 21 - - HYPH 32505 147 22 leafed leaf VBN 32505 147 23 plants plant NNS 32505 147 24 near near IN 32505 147 25 the the DT 32505 147 26 pond pond NN 32505 147 27 . . . 32505 148 1 Males male NNS 32505 148 2 were be VBD 32505 148 3 observed observe VBN 32505 148 4 in in IN 32505 148 5 late late JJ 32505 148 6 July July NNP 32505 148 7 and and CC 32505 148 8 early early JJ 32505 148 9 August August NNP 32505 148 10 calling call VBG 32505 148 11 from from IN 32505 148 12 _ _ NNP 32505 148 13 Calathea Calathea NNP 32505 148 14 _ _ NNP 32505 148 15 and and CC 32505 148 16 _ _ NNP 32505 148 17 Heliconia Heliconia NNP 32505 148 18 _ _ NNP 32505 148 19 leaves leave VBZ 32505 148 20 at at IN 32505 148 21 the the DT 32505 148 22 edge edge NN 32505 148 23 of of IN 32505 148 24 a a DT 32505 148 25 pond pond NN 32505 148 26 in in IN 32505 148 27 the the DT 32505 148 28 wet wet JJ 32505 148 29 forest forest NN 32505 148 30 of of IN 32505 148 31 the the DT 32505 148 32 Osa Osa NNP 32505 148 33 Peninsula Peninsula NNP 32505 148 34 . . . 32505 149 1 William William NNP 32505 149 2 E. E. NNP 32505 149 3 Duellman Duellman NNP 32505 149 4 informed inform VBD 32505 149 5 me -PRON- PRP 32505 149 6 that that IN 32505 149 7 he -PRON- PRP 32505 149 8 collected collect VBD 32505 149 9 calling call VBG 32505 149 10 males male NNS 32505 149 11 in in IN 32505 149 12 January January NNP 32505 149 13 at at IN 32505 149 14 El El NNP 32505 149 15 Real Real NNP 32505 149 16 , , , 32505 149 17 Darién Darién NNP 32505 149 18 , , , 32505 149 19 and and CC 32505 149 20 in in IN 32505 149 21 March March NNP 32505 149 22 at at IN 32505 149 23 Almirante Almirante NNP 32505 149 24 , , , 32505 149 25 Bocas Bocas NNP 32505 149 26 del del NNP 32505 149 27 Toro Toro NNP 32505 149 28 , , , 32505 149 29 Panamá Panamá NNP 32505 149 30 . . . 32505 150 1 Taylor Taylor NNP 32505 150 2 ( ( -LRB- 32505 150 3 1952 1952 CD 32505 150 4 ) ) -RRB- 32505 150 5 found find VBD 32505 150 6 calling call VBG 32505 150 7 males male NNS 32505 150 8 in in IN 32505 150 9 June June NNP 32505 150 10 at at IN 32505 150 11 Turrialba Turrialba NNP 32505 150 12 , , , 32505 150 13 Cartago Cartago NNP 32505 150 14 Province Province NNP 32505 150 15 , , , 32505 150 16 Costa Costa NNP 32505 150 17 Rica Rica NNP 32505 150 18 , , , 32505 150 19 and and CC 32505 150 20 Dunn Dunn NNP 32505 150 21 ( ( -LRB- 32505 150 22 1931a 1931a CD 32505 150 23 ) ) -RRB- 32505 150 24 observed observe VBD 32505 150 25 males male NNS 32505 150 26 calling call VBG 32505 150 27 in in IN 32505 150 28 July July NNP 32505 150 29 , , , 32505 150 30 November November NNP 32505 150 31 , , , 32505 150 32 and and CC 32505 150 33 December December NNP 32505 150 34 in in IN 32505 150 35 Panamá Panamá NNP 32505 150 36 . . . 32505 151 1 Gravid gravid JJ 32505 151 2 females female NNS 32505 151 3 have have VBP 32505 151 4 been be VBN 32505 151 5 found find VBN 32505 151 6 from from IN 32505 151 7 April April NNP 32505 151 8 to to IN 32505 151 9 August August NNP 32505 151 10 . . . 32505 152 1 Breeding breeding NN 32505 152 2 activities activity NNS 32505 152 3 of of IN 32505 152 4 _ _ NNP 32505 152 5 Hyla Hyla NNP 32505 152 6 boulengeri boulengeri NN 32505 152 7 _ _ NNP 32505 152 8 always always RB 32505 152 9 seem seem VBP 32505 152 10 to to TO 32505 152 11 be be VB 32505 152 12 associated associate VBN 32505 152 13 with with IN 32505 152 14 temporary temporary JJ 32505 152 15 ponds pond NNS 32505 152 16 ; ; : 32505 152 17 in in IN 32505 152 18 Central Central NNP 32505 152 19 America America NNP 32505 152 20 breeding breed VBG 32505 152 21 apparently apparently RB 32505 152 22 takes take VBZ 32505 152 23 place place NN 32505 152 24 throughout throughout IN 32505 152 25 most most JJS 32505 152 26 of of IN 32505 152 27 the the DT 32505 152 28 year year NN 32505 152 29 . . . 32505 153 1 The the DT 32505 153 2 mating mating NN 32505 153 3 call call NN 32505 153 4 of of IN 32505 153 5 _ _ NNP 32505 153 6 Hyla Hyla NNP 32505 153 7 boulengeri boulengeri NN 32505 153 8 _ _ NNP 32505 153 9 consists consist VBZ 32505 153 10 of of IN 32505 153 11 one one CD 32505 153 12 short short JJ 32505 153 13 , , , 32505 153 14 moderately moderately RB 32505 153 15 low low RB 32505 153 16 - - HYPH 32505 153 17 pitched pitched JJ 32505 153 18 note note NN 32505 153 19 . . . 32505 154 1 Each each DT 32505 154 2 note note NN 32505 154 3 has have VBZ 32505 154 4 a a DT 32505 154 5 duration duration NN 32505 154 6 of of IN 32505 154 7 0.24 0.24 CD 32505 154 8 to to TO 32505 154 9 0.47 0.47 CD 32505 154 10 second second JJ 32505 154 11 and and CC 32505 154 12 is be VBZ 32505 154 13 repeated repeat VBN 32505 154 14 at at IN 32505 154 15 intervals interval NNS 32505 154 16 of of IN 32505 154 17 one one CD 32505 154 18 second second JJ 32505 154 19 to to IN 32505 154 20 several several JJ 32505 154 21 minutes minute NNS 32505 154 22 . . . 32505 155 1 The the DT 32505 155 2 notes note NNS 32505 155 3 have have VBP 32505 155 4 80 80 CD 32505 155 5 to to TO 32505 155 6 120 120 CD 32505 155 7 pulses pulse NNS 32505 155 8 per per IN 32505 155 9 second second NN 32505 155 10 , , , 32505 155 11 a a DT 32505 155 12 fundamental fundamental JJ 32505 155 13 frequency frequency NN 32505 155 14 of of IN 32505 155 15 about about RB 32505 155 16 70 70 CD 32505 155 17 cycles cycle NNS 32505 155 18 per per IN 32505 155 19 second second NN 32505 155 20 , , , 32505 155 21 and and CC 32505 155 22 a a DT 32505 155 23 dominant dominant JJ 32505 155 24 frequency frequency NN 32505 155 25 of of IN 32505 155 26 2,840 2,840 CD 32505 155 27 cycles cycle NNS 32505 155 28 per per IN 32505 155 29 second second JJ 32505 155 30 ( ( -LRB- 32505 155 31 Table table NN 32505 155 32 2 2 CD 32505 155 33 , , , 32505 155 34 Pl Pl NNP 32505 155 35 . . . 32505 156 1 3A 3A NNP 32505 156 2 ) ) -RRB- 32505 156 3 . . . 32505 157 1 The the DT 32505 157 2 eggs egg NNS 32505 157 3 are be VBP 32505 157 4 deposited deposit VBN 32505 157 5 in in IN 32505 157 6 a a DT 32505 157 7 mass mass NN 32505 157 8 in in IN 32505 157 9 the the DT 32505 157 10 water water NN 32505 157 11 . . . 32505 158 1 No no DT 32505 158 2 information information NN 32505 158 3 is be VBZ 32505 158 4 available available JJ 32505 158 5 concerning concern VBG 32505 158 6 early early JJ 32505 158 7 development development NN 32505 158 8 . . . 32505 159 1 Tadpoles tadpole NNS 32505 159 2 in in IN 32505 159 3 advanced advanced JJ 32505 159 4 stages stage NNS 32505 159 5 of of IN 32505 159 6 development development NN 32505 159 7 were be VBD 32505 159 8 found find VBN 32505 159 9 in in IN 32505 159 10 a a DT 32505 159 11 temporary temporary JJ 32505 159 12 pond pond NN 32505 159 13 at at IN 32505 159 14 Rincón Rincón NNP 32505 159 15 de de NNP 32505 159 16 Osa Osa NNP 32505 159 17 , , , 32505 159 18 Puntarenas Puntarenas NNP 32505 159 19 Province Province NNP 32505 159 20 , , , 32505 159 21 Costa Costa NNP 32505 159 22 Rica Rica NNP 32505 159 23 . . . 32505 160 1 The the DT 32505 160 2 pond pond NN 32505 160 3 was be VBD 32505 160 4 about about RB 32505 160 5 10 10 CD 32505 160 6 cm cm NNS 32505 160 7 . . . 32505 161 1 deep deep RB 32505 161 2 , , , 32505 161 3 had have VBD 32505 161 4 a a DT 32505 161 5 muddy muddy JJ 32505 161 6 bottom bottom NN 32505 161 7 and and CC 32505 161 8 lacked lack VBD 32505 161 9 vegetation vegetation NN 32505 161 10 . . . 32505 162 1 Three three CD 32505 162 2 recently recently RB 32505 162 3 metamorphosed metamorphose VBN 32505 162 4 young young JJ 32505 162 5 were be VBD 32505 162 6 found find VBN 32505 162 7 in in IN 32505 162 8 mid mid NNP 32505 162 9 - - NNP 32505 162 10 August August NNP 32505 162 11 , , , 32505 162 12 1966 1966 CD 32505 162 13 , , , 32505 162 14 on on IN 32505 162 15 grass grass NN 32505 162 16 at at IN 32505 162 17 the the DT 32505 162 18 edge edge NN 32505 162 19 of of IN 32505 162 20 another another DT 32505 162 21 temporary temporary JJ 32505 162 22 pond pond NN 32505 162 23 in in IN 32505 162 24 the the DT 32505 162 25 forest forest NN 32505 162 26 . . . 32505 163 1 _ _ NNP 32505 163 2 Tadpoles_--Twelve tadpoles_--twelve CD 32505 163 3 tadpoles tadpole NNS 32505 163 4 are be VBP 32505 163 5 available available JJ 32505 163 6 . . . 32505 164 1 These these DT 32505 164 2 were be VBD 32505 164 3 collected collect VBN 32505 164 4 at at IN 32505 164 5 Rincón Rincón NNP 32505 164 6 de de NNP 32505 164 7 Osa Osa NNP 32505 164 8 , , , 32505 164 9 Puntarenas Puntarenas NNP 32505 164 10 Province Province NNP 32505 164 11 , , , 32505 164 12 Costa Costa NNP 32505 164 13 Rica Rica NNP 32505 164 14 . . . 32505 165 1 The the DT 32505 165 2 maximum maximum JJ 32505 165 3 size size NN 32505 165 4 represented represent VBD 32505 165 5 is be VBZ 32505 165 6 34.0 34.0 CD 32505 165 7 mm mm NNP 32505 165 8 . . NNP 32505 165 9 , , , 32505 165 10 total total JJ 32505 165 11 length length NN 32505 165 12 ( ( -LRB- 32505 165 13 stage stage VB 32505 165 14 42 42 CD 32505 165 15 of of IN 32505 165 16 development development NN 32505 165 17 ) ) -RRB- 32505 165 18 . . . 32505 166 1 A a DT 32505 166 2 typical typical JJ 32505 166 3 tadpole tadpole NN 32505 166 4 in in IN 32505 166 5 stage stage NN 32505 166 6 36 36 CD 32505 166 7 of of IN 32505 166 8 development development NN 32505 166 9 ( ( -LRB- 32505 166 10 KU KU NNP 32505 166 11 104295 104295 CD 32505 166 12 ) ) -RRB- 32505 166 13 has have VBZ 32505 166 14 a a DT 32505 166 15 body body NN 32505 166 16 length length NN 32505 166 17 of of IN 32505 166 18 12.0 12.0 CD 32505 166 19 mm mm NNP 32505 166 20 . . NNP 32505 166 21 , , , 32505 166 22 tail tail NN 32505 166 23 length length NN 32505 166 24 of of IN 32505 166 25 20.0 20.0 CD 32505 166 26 mm mm NNP 32505 166 27 . . NNP 32505 166 28 , , , 32505 166 29 and and CC 32505 166 30 total total JJ 32505 166 31 length length NN 32505 166 32 of of IN 32505 166 33 32.0 32.0 CD 32505 166 34 mm mm NNS 32505 166 35 . . . 32505 167 1 Other other JJ 32505 167 2 characters character NNS 32505 167 3 are be VBP 32505 167 4 as as IN 32505 167 5 follows follow VBZ 32505 167 6 : : : 32505 167 7 depth depth NN 32505 167 8 of of IN 32505 167 9 tail tail NN 32505 167 10 equal equal JJ 32505 167 11 to to IN 32505 167 12 length length NN 32505 167 13 of of IN 32505 167 14 body body NN 32505 167 15 ; ; : 32505 167 16 body body NN 32505 167 17 deeper deeply RBR 32505 167 18 than than IN 32505 167 19 wide wide JJ 32505 167 20 ; ; : 32505 167 21 distance distance NN 32505 167 22 between between IN 32505 167 23 eye eye NN 32505 167 24 and and CC 32505 167 25 nostril nostril NNP 32505 167 26 equal equal JJ 32505 167 27 to to IN 32505 167 28 that that DT 32505 167 29 between between IN 32505 167 30 nostril nostril NNP 32505 167 31 and and CC 32505 167 32 tip tip NN 32505 167 33 of of IN 32505 167 34 snout snout NNP 32505 167 35 ; ; : 32505 167 36 mouth mouth NNP 32505 167 37 anteroventral anteroventral NNP 32505 167 38 , , , 32505 167 39 upper upper JJ 32505 167 40 and and CC 32505 167 41 lower low JJR 32505 167 42 lips lip NNS 32505 167 43 bare bare VBP 32505 167 44 ; ; : 32505 167 45 papillae papillae NNP 32505 167 46 present present VBP 32505 167 47 laterally laterally RB 32505 167 48 ; ; : 32505 167 49 tooth tooth NN 32505 167 50 rows row VBZ 32505 167 51 2/3 2/3 CD 32505 167 52 ; ; : 32505 167 53 upper upper JJ 32505 167 54 rows row NNS 32505 167 55 about about IN 32505 167 56 equal equal JJ 32505 167 57 in in IN 32505 167 58 length length NN 32505 167 59 ; ; : 32505 167 60 first first RB 32505 167 61 upper upper JJ 32505 167 62 row row NN 32505 167 63 slightly slightly RB 32505 167 64 , , , 32505 167 65 and and CC 32505 167 66 second second JJ 32505 167 67 upper upper JJ 32505 167 68 row row NN 32505 167 69 widely widely RB 32505 167 70 , , , 32505 167 71 interrupted interrupt VBN 32505 167 72 medially medially RB 32505 167 73 ; ; : 32505 167 74 lower low JJR 32505 167 75 rows row NNS 32505 167 76 about about IN 32505 167 77 equal equal JJ 32505 167 78 in in IN 32505 167 79 length length NN 32505 167 80 , , , 32505 167 81 shorter short JJR 32505 167 82 than than IN 32505 167 83 upper upper JJ 32505 167 84 rows row NNS 32505 167 85 ; ; : 32505 167 86 third third RB 32505 167 87 lower low JJR 32505 167 88 row row NN 32505 167 89 containing contain VBG 32505 167 90 5 5 CD 32505 167 91 - - SYM 32505 167 92 10 10 CD 32505 167 93 large large JJ 32505 167 94 teeth tooth NNS 32505 167 95 ; ; : 32505 167 96 beak beak NNP 32505 167 97 strong strong JJ 32505 167 98 , , , 32505 167 99 serrate serrate NN 32505 167 100 ; ; : 32505 167 101 spiracle spiracle NNP 32505 167 102 nearer nearer NNP 32505 167 103 anus anus NN 32505 167 104 than than IN 32505 167 105 eye eye NN 32505 167 106 ; ; : 32505 167 107 anal anal JJ 32505 167 108 aperture aperture NN 32505 167 109 not not RB 32505 167 110 extending extend VBG 32505 167 111 to to IN 32505 167 112 border border NN 32505 167 113 of of IN 32505 167 114 ventral ventral JJ 32505 167 115 fin fin NN 32505 167 116 ; ; , 32505 167 117 caudal caudal NN 32505 167 118 musculature musculature NN 32505 167 119 slender slender NN 32505 167 120 posteriorly posteriorly RB 32505 167 121 , , , 32505 167 122 extending extend VBG 32505 167 123 to to IN 32505 167 124 tip tip NN 32505 167 125 of of IN 32505 167 126 pointed pointed JJ 32505 167 127 tail tail NN 32505 167 128 ; ; : 32505 167 129 dorsal dorsal NN 32505 167 130 fin fin NN 32505 167 131 extending extend VBG 32505 167 132 to to IN 32505 167 133 middle middle NN 32505 167 134 of of IN 32505 167 135 body body NN 32505 167 136 , , , 32505 167 137 slightly slightly RB 32505 167 138 deeper deep JJR 32505 167 139 than than IN 32505 167 140 ventral ventral JJ 32505 167 141 fin fin NN 32505 167 142 ; ; : 32505 167 143 posterior posterior RB 32505 167 144 three three CD 32505 167 145 fourths fourth NNS 32505 167 146 of of IN 32505 167 147 tail tail NN 32505 167 148 spotted spot VBN 32505 167 149 ; ; : 32505 167 150 rest rest NN 32505 167 151 of of IN 32505 167 152 tail tail NN 32505 167 153 and and CC 32505 167 154 body body NN 32505 167 155 gray gray JJ 32505 167 156 - - HYPH 32505 167 157 brown brown JJ 32505 167 158 or or CC 32505 167 159 transparent transparent JJ 32505 167 160 ; ; : 32505 167 161 hindlimbs hindlimb NNS 32505 167 162 flecked fleck VBN 32505 167 163 or or CC 32505 167 164 spotted spot VBN 32505 167 165 with with IN 32505 167 166 brown brown JJ 32505 167 167 ( ( -LRB- 32505 167 168 Table table NN 32505 167 169 3 3 CD 32505 167 170 , , , 32505 167 171 Fig Fig NNP 32505 167 172 . . . 32505 168 1 2A 2A NNP 32505 168 2 and and CC 32505 168 3 3A 3A NNP 32505 168 4 ) ) -RRB- 32505 168 5 . . . 32505 169 1 TABLE TABLE NNS 32505 169 2 3.--Sizes 3.--sizes CD 32505 169 3 of of IN 32505 169 4 Tadpoles Tadpoles NNP 32505 169 5 of of IN 32505 169 6 _ _ NNP 32505 169 7 Hyla Hyla NNP 32505 169 8 boulengeri boulengeri NN 32505 169 9 _ _ NNP 32505 169 10 in in IN 32505 169 11 Relation Relation NNP 32505 169 12 to to IN 32505 169 13 Developmental Developmental NNP 32505 169 14 Stages Stages NNPS 32505 169 15 . . . 32505 170 1 ( ( -LRB- 32505 170 2 Means mean NNS 32505 170 3 in in IN 32505 170 4 parentheses parenthesis NNS 32505 170 5 below below IN 32505 170 6 observed observed JJ 32505 170 7 ranges range NNS 32505 170 8 ; ; : 32505 170 9 measurements measurement NNS 32505 170 10 in in IN 32505 170 11 mm mm NNP 32505 170 12 . . . 32505 170 13 ) ) -RRB- 32505 171 1 = = NFP 32505 171 2 = = NFP 32505 171 3 = = SYM 32505 171 4 = = SYM 32505 171 5 = = SYM 32505 171 6 = = SYM 32505 171 7 = = SYM 32505 171 8 = = SYM 32505 171 9 = = SYM 32505 171 10 = = SYM 32505 171 11 = = SYM 32505 171 12 = = SYM 32505 171 13 = = SYM 32505 171 14 = = SYM 32505 171 15 = = SYM 32505 171 16 = = SYM 32505 171 17 = = SYM 32505 171 18 = = SYM 32505 171 19 = = SYM 32505 171 20 = = SYM 32505 171 21 = = SYM 32505 171 22 = = SYM 32505 171 23 = = SYM 32505 171 24 = = SYM 32505 171 25 = = SYM 32505 171 26 = = SYM 32505 171 27 = = SYM 32505 171 28 = = SYM 32505 171 29 = = SYM 32505 171 30 = = SYM 32505 171 31 = = SYM 32505 171 32 = = SYM 32505 171 33 = = SYM 32505 171 34 = = SYM 32505 171 35 = = SYM 32505 171 36 = = SYM 32505 171 37 = = SYM 32505 171 38 = = SYM 32505 171 39 = = SYM 32505 171 40 = = SYM 32505 171 41 = = SYM 32505 171 42 = = SYM 32505 171 43 = = SYM 32505 171 44 = = SYM 32505 171 45 = = SYM 32505 171 46 = = SYM 32505 171 47 = = SYM 32505 171 48 = = SYM 32505 171 49 = = SYM 32505 171 50 = = SYM 32505 171 51 = = SYM 32505 171 52 = = SYM 32505 171 53 = = SYM 32505 171 54 = = NFP 32505 171 55 = = NFP 32505 171 56 Stage stage NN 32505 171 57 | | NNP 32505 171 58 N N NNP 32505 171 59 | | NNP 32505 171 60 Body Body NNP 32505 171 61 length length NN 32505 171 62 | | NNP 32505 171 63 Tail Tail NNP 32505 171 64 length length NN 32505 171 65 | | NNP 32505 171 66 Total total JJ 32505 171 67 length length NN 32505 171 68 --------+---+-------------+-------------+-------------- --------+---+-------------+-------------+-------------- NN 32505 171 69 30 30 CD 32505 171 70 | | CD 32505 171 71 1 1 CD 32505 171 72 | | CD 32505 171 73 11.0 11.0 CD 32505 171 74 | | CD 32505 171 75 22.2 22.2 CD 32505 171 76 | | CD 32505 171 77 33.2 33.2 CD 32505 171 78 | | CD 32505 171 79 | | CD 32505 171 80 | | CD 32505 171 81 | | CD 32505 171 82 35 35 CD 32505 171 83 | | CD 32505 171 84 1 1 CD 32505 171 85 | | CD 32505 171 86 11.0 11.0 CD 32505 171 87 | | CD 32505 171 88 12.0 12.0 CD 32505 171 89 | | CD 32505 171 90 23.0 23.0 CD 32505 171 91 | | CD 32505 171 92 | | CD 32505 171 93 | | CD 32505 171 94 | | CD 32505 171 95 36 36 CD 32505 171 96 | | CD 32505 171 97 3 3 CD 32505 171 98 | | CD 32505 171 99 9.5 9.5 CD 32505 171 100 - - HYPH 32505 171 101 12.0 12.0 CD 32505 171 102 | | CD 32505 171 103 20.0 20.0 CD 32505 171 104 - - HYPH 32505 171 105 21.5 21.5 CD 32505 171 106 | | CD 32505 171 107 31.0 31.0 CD 32505 171 108 - - HYPH 32505 171 109 32.0 32.0 CD 32505 171 110 | | CD 32505 171 111 | | CD 32505 171 112 ( ( -LRB- 32505 171 113 11.2 11.2 CD 32505 171 114 ) ) -RRB- 32505 171 115 | | NNP 32505 171 116 ( ( -LRB- 32505 171 117 20.5 20.5 CD 32505 171 118 ) ) -RRB- 32505 171 119 | | NNP 32505 171 120 ( ( -LRB- 32505 171 121 31.7 31.7 CD 32505 171 122 ) ) -RRB- 32505 171 123 | | NNP 32505 171 124 | | CD 32505 171 125 | | CD 32505 171 126 | | CD 32505 171 127 38 38 CD 32505 171 128 | | CD 32505 171 129 2 2 CD 32505 171 130 | | CD 32505 171 131 11.5 11.5 CD 32505 171 132 | | CD 32505 171 133 22.0 22.0 CD 32505 171 134 | | CD 32505 171 135 33.5 33.5 CD 32505 171 136 | | CD 32505 171 137 | | CD 32505 171 138 | | CD 32505 171 139 | | CD 32505 171 140 42 42 CD 32505 171 141 | | CD 32505 171 142 2 2 CD 32505 171 143 | | CD 32505 171 144 10.5 10.5 CD 32505 171 145 - - HYPH 32505 171 146 13.0 13.0 CD 32505 171 147 | | CD 32505 171 148 21.0 21.0 CD 32505 171 149 - - HYPH 32505 171 150 22.0 22.0 CD 32505 171 151 | | CD 32505 171 152 32.5 32.5 CD 32505 171 153 - - HYPH 32505 171 154 34.0 34.0 CD 32505 171 155 | | CD 32505 171 156 | | CD 32505 171 157 ( ( -LRB- 32505 171 158 11.8 11.8 CD 32505 171 159 ) ) -RRB- 32505 171 160 | | NNP 32505 171 161 ( ( -LRB- 32505 171 162 21.5 21.5 CD 32505 171 163 ) ) -RRB- 32505 171 164 | | NNP 32505 171 165 ( ( -LRB- 32505 171 166 33.3 33.3 CD 32505 171 167 ) ) -RRB- 32505 171 168 | | NNP 32505 171 169 | | CD 32505 171 170 | | CD 32505 171 171 | | CD 32505 171 172 44 44 CD 32505 171 173 | | CD 32505 171 174 2 2 CD 32505 171 175 | | CD 32505 171 176 14.0 14.0 CD 32505 171 177 - - HYPH 32505 171 178 15.0 15.0 CD 32505 171 179 | | CD 32505 171 180 8.0 8.0 CD 32505 171 181 - - HYPH 32505 171 182 15.0 15.0 CD 32505 171 183 | | CD 32505 171 184 22.0 22.0 CD 32505 171 185 - - HYPH 32505 171 186 30.0 30.0 CD 32505 171 187 | | CD 32505 171 188 | | CD 32505 171 189 ( ( -LRB- 32505 171 190 14.5 14.5 CD 32505 171 191 ) ) -RRB- 32505 171 192 | | NNP 32505 171 193 ( ( -LRB- 32505 171 194 12.5 12.5 CD 32505 171 195 ) ) -RRB- 32505 171 196 | | NNP 32505 171 197 ( ( -LRB- 32505 171 198 26.0 26.0 CD 32505 171 199 ) ) -RRB- 32505 171 200 | | NNP 32505 171 201 | | CD 32505 171 202 | | CD 32505 171 203 | | CD 32505 171 204 46 46 CD 32505 171 205 | | CD 32505 171 206 1 1 CD 32505 171 207 | | CD 32505 171 208 15.0 15.0 CD 32505 171 209 | | CD 32505 171 210 15.0 15.0 CD 32505 171 211 | | NNS 32505 171 212 A a DT 32505 171 213 recently recently RB 32505 171 214 metamorphosed metamorphose VBN 32505 171 215 young young NNP 32505 171 216 has have VBZ 32505 171 217 a a DT 32505 171 218 snout snout NN 32505 171 219 - - HYPH 32505 171 220 vent vent NN 32505 171 221 length length NN 32505 171 222 of of IN 32505 171 223 15 15 CD 32505 171 224 mm mm NN 32505 171 225 . . . 32505 172 1 ; ; : 32505 172 2 the the DT 32505 172 3 head head NN 32505 172 4 is be VBZ 32505 172 5 as as RB 32505 172 6 long long RB 32505 172 7 as as IN 32505 172 8 wide wide JJ 32505 172 9 , , , 32505 172 10 the the DT 32505 172 11 eyes eye NNS 32505 172 12 are be VBP 32505 172 13 prominent prominent JJ 32505 172 14 ; ; : 32505 172 15 the the DT 32505 172 16 limbs limb NNS 32505 172 17 are be VBP 32505 172 18 weakly weakly RB 32505 172 19 barred bar VBN 32505 172 20 ; ; : 32505 172 21 the the DT 32505 172 22 skin skin NN 32505 172 23 is be VBZ 32505 172 24 rugose rugose VB 32505 172 25 above above RB 32505 172 26 and and CC 32505 172 27 granular granular JJ 32505 172 28 below below RB 32505 172 29 . . . 32505 173 1 The the DT 32505 173 2 venter venter NN 32505 173 3 is be VBZ 32505 173 4 immaculate immaculate JJ 32505 173 5 ; ; : 32505 173 6 the the DT 32505 173 7 dorsum dorsum NN 32505 173 8 and and CC 32505 173 9 limbs limb NNS 32505 173 10 are be VBP 32505 173 11 gray gray JJ 32505 173 12 - - HYPH 32505 173 13 brown brown JJ 32505 173 14 in in IN 32505 173 15 preservative preservative JJ 32505 173 16 ( ( -LRB- 32505 173 17 pale pale JJ 32505 173 18 green green NN 32505 173 19 in in IN 32505 173 20 life life NN 32505 173 21 ) ) -RRB- 32505 173 22 . . . 32505 174 1 The the DT 32505 174 2 interorbital interorbital JJ 32505 174 3 space space NN 32505 174 4 , , , 32505 174 5 supratympanic supratympanic NNP 32505 174 6 fold fold NNP 32505 174 7 , , , 32505 174 8 and and CC 32505 174 9 scapular scapular JJ 32505 174 10 region region NN 32505 174 11 are be VBP 32505 174 12 darker dark JJR 32505 174 13 than than IN 32505 174 14 the the DT 32505 174 15 rest rest NN 32505 174 16 of of IN 32505 174 17 the the DT 32505 174 18 body body NN 32505 174 19 ; ; : 32505 174 20 the the DT 32505 174 21 fingers finger NNS 32505 174 22 lack lack NN 32505 174 23 webbing webbing NN 32505 174 24 ; ; : 32505 174 25 the the DT 32505 174 26 webbing webbing NN 32505 174 27 on on IN 32505 174 28 the the DT 32505 174 29 foot foot NN 32505 174 30 is be VBZ 32505 174 31 the the DT 32505 174 32 same same JJ 32505 174 33 as as IN 32505 174 34 in in IN 32505 174 35 adults adult NNS 32505 174 36 ; ; : 32505 174 37 small small JJ 32505 174 38 metatarsal metatarsal NN 32505 174 39 tubercles tubercle NNS 32505 174 40 are be VBP 32505 174 41 present present JJ 32505 174 42 , , , 32505 174 43 but but CC 32505 174 44 the the DT 32505 174 45 tarsal tarsal NN 32505 174 46 fold fold NN 32505 174 47 is be VBZ 32505 174 48 absent absent JJ 32505 174 49 . . . 32505 175 1 [ [ -LRB- 32505 175 2 Illustration illustration NN 32505 175 3 : : : 32505 175 4 Fig Fig NNP 32505 175 5 . . . 32505 176 1 2 2 LS 32505 176 2 . . . 32505 177 1 Tadpoles tadpole NNS 32505 177 2 of of IN 32505 177 3 ( ( -LRB- 32505 177 4 A a NN 32505 177 5 ) ) -RRB- 32505 177 6 _ _ NNP 32505 177 7 Hyla Hyla NNP 32505 177 8 boulengeri boulengeri NN 32505 177 9 _ _ NNP 32505 177 10 ( ( -LRB- 32505 177 11 KU KU NNP 32505 177 12 104295 104295 CD 32505 177 13 ) ) -RRB- 32505 177 14 and and CC 32505 177 15 ( ( -LRB- 32505 177 16 B b NN 32505 177 17 ) ) -RRB- 32505 177 18 _ _ NNP 32505 177 19 Hyla Hyla NNP 32505 177 20 elaeochroa elaeochroa NN 32505 177 21 _ _ NNP 32505 177 22 ( ( -LRB- 32505 177 23 KU KU NNP 32505 177 24 104134 104134 CD 32505 177 25 ) ) -RRB- 32505 177 26 , , , 32505 177 27 × × NN 32505 177 28 3 3 CD 32505 177 29 . . . 32505 177 30 ] ] -RRB- 32505 178 1 [ [ -LRB- 32505 178 2 Illustration illustration NN 32505 178 3 : : : 32505 178 4 Fig Fig NNP 32505 178 5 . . . 32505 179 1 3 3 LS 32505 179 2 . . . 32505 180 1 Mouthparts mouthpart NNS 32505 180 2 of of IN 32505 180 3 tadpoles tadpole NNS 32505 180 4 of of IN 32505 180 5 ( ( -LRB- 32505 180 6 A a NN 32505 180 7 ) ) -RRB- 32505 180 8 _ _ NNP 32505 180 9 Hyla Hyla NNP 32505 180 10 boulengeri boulengeri NN 32505 180 11 _ _ NNP 32505 180 12 ( ( -LRB- 32505 180 13 KU KU NNP 32505 180 14 104295 104295 CD 32505 180 15 ) ) -RRB- 32505 180 16 and and CC 32505 180 17 ( ( -LRB- 32505 180 18 B b NN 32505 180 19 ) ) -RRB- 32505 180 20 _ _ NNP 32505 180 21 Hyla Hyla NNP 32505 180 22 elaeochroa elaeochroa NN 32505 180 23 _ _ NNP 32505 180 24 ( ( -LRB- 32505 180 25 KU KU NNP 32505 180 26 104134 104134 CD 32505 180 27 ) ) -RRB- 32505 180 28 , , , 32505 180 29 × × CD 32505 180 30 25 25 CD 32505 180 31 . . . 32505 180 32 ] ] -RRB- 32505 181 1 _ _ NNP 32505 181 2 Remarks._--Cope Remarks._--Cope NNP 32505 181 3 ( ( -LRB- 32505 181 4 1887:12 1887:12 CD 32505 181 5 ) ) -RRB- 32505 181 6 described describe VBD 32505 181 7 _ _ NNP 32505 181 8 Scytopis Scytopis NNP 32505 181 9 boulengeri boulengeri NN 32505 181 10 _ _ NNP 32505 181 11 from from IN 32505 181 12 Nicaragua Nicaragua NNP 32505 181 13 . . . 32505 182 1 Günther Günther NNP 32505 182 2 ( ( -LRB- 32505 182 3 1901:267 1901:267 CD 32505 182 4 ) ) -RRB- 32505 182 5 placed place VBD 32505 182 6 _ _ NNP 32505 182 7 boulengeri boulengeri NN 32505 182 8 _ _ NNP 32505 182 9 in in IN 32505 182 10 the the DT 32505 182 11 genus genus NN 32505 182 12 _ _ NNP 32505 182 13 Hyla Hyla NNP 32505 182 14 _ _ NNP 32505 182 15 , , , 32505 182 16 and and CC 32505 182 17 stated state VBD 32505 182 18 that that IN 32505 182 19 Cope Cope NNP 32505 182 20 possibly possibly RB 32505 182 21 placed place VBD 32505 182 22 _ _ NNP 32505 182 23 boulengeri boulengeri NN 32505 182 24 _ _ NNP 32505 182 25 in in IN 32505 182 26 the the DT 32505 182 27 genus genus NN 32505 182 28 _ _ NNP 32505 182 29 Scytopis Scytopis NNP 32505 182 30 _ _ NNP 32505 182 31 on on IN 32505 182 32 the the DT 32505 182 33 supposition supposition NN 32505 182 34 that that IN 32505 182 35 it -PRON- PRP 32505 182 36 had have VBD 32505 182 37 an an DT 32505 182 38 accumulation accumulation NN 32505 182 39 of of IN 32505 182 40 " " `` 32505 182 41 sebaceous sebaceous JJ 32505 182 42 glands gland NNS 32505 182 43 " " '' 32505 182 44 above above IN 32505 182 45 the the DT 32505 182 46 tympanum tympanum NN 32505 182 47 . . . 32505 183 1 Noble Noble NNP 32505 183 2 ( ( -LRB- 32505 183 3 1918:339 1918:339 CD 32505 183 4 ) ) -RRB- 32505 183 5 redescribed redescribe VBD 32505 183 6 _ _ NNP 32505 183 7 Hyla Hyla NNP 32505 183 8 boulengeri boulengeri NN 32505 183 9 _ _ NNP 32505 183 10 on on IN 32505 183 11 the the DT 32505 183 12 basis basis NN 32505 183 13 of of IN 32505 183 14 three three CD 32505 183 15 specimens specimen NNS 32505 183 16 from from IN 32505 183 17 Zelaya Zelaya NNP 32505 183 18 Province Province NNP 32505 183 19 , , , 32505 183 20 Nicaragua Nicaragua NNP 32505 183 21 , , , 32505 183 22 and and CC 32505 183 23 noted note VBD 32505 183 24 that that IN 32505 183 25 the the DT 32505 183 26 glands gland NNS 32505 183 27 were be VBD 32505 183 28 not not RB 32505 183 29 prominent prominent JJ 32505 183 30 in in IN 32505 183 31 any any DT 32505 183 32 of of IN 32505 183 33 the the DT 32505 183 34 specimens specimen NNS 32505 183 35 . . . 32505 184 1 Duellman duellman NN 32505 184 2 ( ( -LRB- 32505 184 3 1956:8 1956:8 CD 32505 184 4 ) ) -RRB- 32505 184 5 showed show VBD 32505 184 6 that that IN 32505 184 7 _ _ NNP 32505 184 8 Scytopis Scytopis NNP 32505 184 9 hebes hebe VBZ 32505 184 10 _ _ NNP 32505 184 11 ( ( -LRB- 32505 184 12 generotype generotype NN 32505 184 13 of of IN 32505 184 14 _ _ NNP 32505 184 15 Scytopis Scytopis NNP 32505 184 16 _ _ NNP 32505 184 17 by by IN 32505 184 18 monotypy monotypy NNP 32505 184 19 ) ) -RRB- 32505 184 20 is be VBZ 32505 184 21 a a DT 32505 184 22 Phrynohyas Phrynohyas NNP 32505 184 23 , , , 32505 184 24 and and CC 32505 184 25 thus thus RB 32505 184 26 placed place VBN 32505 184 27 _ _ NNP 32505 184 28 Scytopis Scytopis NNP 32505 184 29 _ _ NNP 32505 184 30 Cope Cope NNP 32505 184 31 , , , 32505 184 32 1862 1862 CD 32505 184 33 , , , 32505 184 34 in in IN 32505 184 35 the the DT 32505 184 36 synonymy synonymy NN 32505 184 37 of of IN 32505 184 38 _ _ NNP 32505 184 39 Phrynohyas Phrynohyas NNP 32505 184 40 _ _ NNP 32505 184 41 Fitzinger Fitzinger NNP 32505 184 42 , , , 32505 184 43 1843 1843 CD 32505 184 44 . . . 32505 185 1 Dunn Dunn NNP 32505 185 2 and and CC 32505 185 3 Emlen Emlen NNP 32505 185 4 ( ( -LRB- 32505 185 5 1932:25 1932:25 CD 32505 185 6 ) ) -RRB- 32505 185 7 placed place VBD 32505 185 8 _ _ NNP 32505 185 9 Hyla Hyla NNP 32505 185 10 lancasteri lancasteri NN 32505 185 11 _ _ NNP 32505 185 12 Barbour Barbour NNP 32505 185 13 in in IN 32505 185 14 the the DT 32505 185 15 synonymy synonymy NN 32505 185 16 of of IN 32505 185 17 _ _ NNP 32505 185 18 Hyla Hyla NNP 32505 185 19 boulengeri boulengeri NN 32505 185 20 _ _ NNP 32505 185 21 ; ; : 32505 185 22 the the DT 32505 185 23 former former JJ 32505 185 24 was be VBD 32505 185 25 known know VBN 32505 185 26 solely solely RB 32505 185 27 from from IN 32505 185 28 one one CD 32505 185 29 juvenile juvenile NN 32505 185 30 . . . 32505 186 1 They -PRON- PRP 32505 186 2 made make VBD 32505 186 3 no no DT 32505 186 4 qualifying qualifying NN 32505 186 5 statements statement NNS 32505 186 6 , , , 32505 186 7 but but CC 32505 186 8 probably probably RB 32505 186 9 they -PRON- PRP 32505 186 10 were be VBD 32505 186 11 impressed impress VBN 32505 186 12 by by IN 32505 186 13 the the DT 32505 186 14 strongly strongly RB 32505 186 15 barred barred JJ 32505 186 16 thighs thigh NNS 32505 186 17 , , , 32505 186 18 a a DT 32505 186 19 coloration coloration NN 32505 186 20 known know VBN 32505 186 21 among among IN 32505 186 22 Central central JJ 32505 186 23 American american JJ 32505 186 24 hylids hylid NNS 32505 186 25 at at IN 32505 186 26 that that DT 32505 186 27 time time NN 32505 186 28 only only RB 32505 186 29 in in IN 32505 186 30 _ _ NNP 32505 186 31 Hyla Hyla NNP 32505 186 32 boulengeri boulengeri NN 32505 186 33 _ _ NNP 32505 186 34 ( ( -LRB- 32505 186 35 Duellman Duellman NNP 32505 186 36 , , , 32505 186 37 1966a:271 1966a:271 CD 32505 186 38 ) ) -RRB- 32505 186 39 . . . 32505 187 1 Taylor Taylor NNP 32505 187 2 ( ( -LRB- 32505 187 3 1952:856 1952:856 CD 32505 187 4 ) ) -RRB- 32505 187 5 followed follow VBD 32505 187 6 Dunn Dunn NNP 32505 187 7 and and CC 32505 187 8 Emlen Emlen NNP 32505 187 9 with with IN 32505 187 10 reservation reservation NN 32505 187 11 and and CC 32505 187 12 noted note VBD 32505 187 13 some some DT 32505 187 14 differences difference NNS 32505 187 15 . . . 32505 188 1 Duellman Duellman NNP 32505 188 2 ( ( -LRB- 32505 188 3 1966a:271 1966a:271 NNP 32505 188 4 ) ) -RRB- 32505 188 5 showed show VBD 32505 188 6 that that IN 32505 188 7 the the DT 32505 188 8 holotype holotype NN 32505 188 9 of of IN 32505 188 10 _ _ NNP 32505 188 11 lancasteri lancasteri NN 32505 188 12 _ _ NNP 32505 188 13 was be VBD 32505 188 14 a a DT 32505 188 15 juvenile juvenile NN 32505 188 16 of of IN 32505 188 17 a a DT 32505 188 18 species species NN 32505 188 19 subsequently subsequently RB 32505 188 20 named name VBN 32505 188 21 as as IN 32505 188 22 _ _ NNP 32505 188 23 Hyla Hyla NNP 32505 188 24 moraviaensis moraviaensis NN 32505 188 25 _ _ NNP 32505 188 26 by by IN 32505 188 27 Taylor Taylor NNP 32505 188 28 ( ( -LRB- 32505 188 29 1952:865 1952:865 CD 32505 188 30 ) ) -RRB- 32505 188 31 . . . 32505 189 1 In in IN 32505 189 2 Central Central NNP 32505 189 3 America America NNP 32505 189 4 , , , 32505 189 5 _ _ NNP 32505 189 6 Hyla Hyla NNP 32505 189 7 boulengeri boulengeri NN 32505 189 8 _ _ NNP 32505 189 9 can can MD 32505 189 10 be be VB 32505 189 11 confused confuse VBN 32505 189 12 only only RB 32505 189 13 with with IN 32505 189 14 _ _ NNP 32505 189 15 Hyla Hyla NNP 32505 189 16 foliamorta foliamorta NN 32505 189 17 ; ; : 32505 189 18 _ _ NNP 32505 189 19 the the DT 32505 189 20 latter latter NN 32505 189 21 is be VBZ 32505 189 22 restricted restrict VBN 32505 189 23 to to IN 32505 189 24 central central JJ 32505 189 25 and and CC 32505 189 26 eastern eastern JJ 32505 189 27 Panamá Panamá NNP 32505 189 28 and and CC 32505 189 29 northern northern JJ 32505 189 30 Colombia Colombia NNP 32505 189 31 . . . 32505 190 1 The the DT 32505 190 2 snout snout NN 32505 190 3 of of IN 32505 190 4 _ _ NNP 32505 190 5 foliamorta foliamorta NNP 32505 190 6 _ _ NNP 32505 190 7 is be VBZ 32505 190 8 more more RBR 32505 190 9 pointed pointed JJ 32505 190 10 and and CC 32505 190 11 protruding protrude VBG 32505 190 12 , , , 32505 190 13 and and CC 32505 190 14 the the DT 32505 190 15 vocal vocal JJ 32505 190 16 sac sac NN 32505 190 17 is be VBZ 32505 190 18 darker dark JJR 32505 190 19 than than IN 32505 190 20 in in IN 32505 190 21 _ _ NNP 32505 190 22 boulengeri boulengeri NNP 32505 190 23 _ _ NNP 32505 190 24 ; ; : 32505 190 25 the the DT 32505 190 26 groin groin NN 32505 190 27 of of IN 32505 190 28 _ _ NNP 32505 190 29 foliamorta foliamorta NNP 32505 190 30 _ _ NNP 32505 190 31 usually usually RB 32505 190 32 is be VBZ 32505 190 33 creamy creamy JJ 32505 190 34 white white JJ 32505 190 35 , , , 32505 190 36 whereas whereas IN 32505 190 37 _ _ NNP 32505 190 38 boulengeri boulengeri NN 32505 190 39 _ _ NNP 32505 190 40 usually usually RB 32505 190 41 has have VBZ 32505 190 42 a a DT 32505 190 43 dark dark JJ 32505 190 44 spot spot NN 32505 190 45 . . . 32505 191 1 The the DT 32505 191 2 skulls skull NNS 32505 191 3 differ differ VBP 32505 191 4 in in IN 32505 191 5 that that DT 32505 191 6 _ _ NNP 32505 191 7 boulengeri boulengeri NN 32505 191 8 _ _ NNP 32505 191 9 has have VBZ 32505 191 10 a a DT 32505 191 11 frontoparietal frontoparietal JJ 32505 191 12 fontanelle fontanelle NN 32505 191 13 , , , 32505 191 14 the the DT 32505 191 15 prevomer prevomer NN 32505 191 16 is be VBZ 32505 191 17 larger large JJR 32505 191 18 and and CC 32505 191 19 elongate elongate VB 32505 191 20 , , , 32505 191 21 anteriorly anteriorly RB 32505 191 22 connected connect VBN 32505 191 23 to to IN 32505 191 24 the the DT 32505 191 25 premaxillary premaxillary NN 32505 191 26 , , , 32505 191 27 and and CC 32505 191 28 posteriorly posteriorly RB 32505 191 29 separated separate VBN 32505 191 30 from from IN 32505 191 31 the the DT 32505 191 32 sphenethmoid sphenethmoid NN 32505 191 33 by by IN 32505 191 34 cartilage cartilage NN 32505 191 35 ; ; : 32505 191 36 _ _ NNP 32505 191 37 foliamorta foliamorta NNP 32505 191 38 _ _ NNP 32505 191 39 lacks lack VBZ 32505 191 40 a a DT 32505 191 41 fontanelle fontanelle NN 32505 191 42 , , , 32505 191 43 the the DT 32505 191 44 prevomer prevomer NN 32505 191 45 is be VBZ 32505 191 46 smaller small JJR 32505 191 47 , , , 32505 191 48 anteriorly anteriorly RB 32505 191 49 separated separate VBN 32505 191 50 from from IN 32505 191 51 the the DT 32505 191 52 premaxillary premaxillary NN 32505 191 53 by by IN 32505 191 54 cartilage cartilage NN 32505 191 55 , , , 32505 191 56 but but CC 32505 191 57 connected connect VBN 32505 191 58 by by IN 32505 191 59 a a DT 32505 191 60 bony bony NN 32505 191 61 suture suture NN 32505 191 62 to to IN 32505 191 63 the the DT 32505 191 64 sphenethmoid sphenethmoid NN 32505 191 65 . . . 32505 192 1 The the DT 32505 192 2 mating mating NN 32505 192 3 call call NN 32505 192 4 of of IN 32505 192 5 _ _ NNP 32505 192 6 boulengeri boulengeri NN 32505 192 7 _ _ NNP 32505 192 8 differs differ VBZ 32505 192 9 by by IN 32505 192 10 having have VBG 32505 192 11 shorter short JJR 32505 192 12 notes note NNS 32505 192 13 , , , 32505 192 14 twice twice RB 32505 192 15 as as RB 32505 192 16 many many JJ 32505 192 17 pulses pulse NNS 32505 192 18 per per IN 32505 192 19 second second NN 32505 192 20 , , , 32505 192 21 a a DT 32505 192 22 higher high JJR 32505 192 23 fundamental fundamental JJ 32505 192 24 frequency frequency NN 32505 192 25 , , , 32505 192 26 and and CC 32505 192 27 more more RBR 32505 192 28 closely closely RB 32505 192 29 approximated approximated JJ 32505 192 30 major major JJ 32505 192 31 frequencies frequency NNS 32505 192 32 than than IN 32505 192 33 does do VBZ 32505 192 34 that that DT 32505 192 35 of of IN 32505 192 36 _ _ NNP 32505 192 37 foliamorta foliamorta NNP 32505 192 38 _ _ NNP 32505 192 39 . . . 32505 193 1 _ _ NNP 32505 193 2 Hyla Hyla NNP 32505 193 3 boulengeri boulengeri NN 32505 193 4 _ _ NNP 32505 193 5 need nee MD 32505 193 6 not not RB 32505 193 7 be be VB 32505 193 8 compared compare VBN 32505 193 9 in in IN 32505 193 10 detail detail NN 32505 193 11 with with IN 32505 193 12 the the DT 32505 193 13 other other JJ 32505 193 14 Central central JJ 32505 193 15 American american JJ 32505 193 16 members member NNS 32505 193 17 of of IN 32505 193 18 the the DT 32505 193 19 _ _ NNP 32505 193 20 Hyla Hyla NNP 32505 193 21 rubra rubra NN 32505 193 22 _ _ NNP 32505 193 23 group group NN 32505 193 24 , , , 32505 193 25 because because IN 32505 193 26 all all DT 32505 193 27 of of IN 32505 193 28 them -PRON- PRP 32505 193 29 are be VBP 32505 193 30 smaller small JJR 32505 193 31 and and CC 32505 193 32 have have VBP 32505 193 33 shorter short JJR 32505 193 34 snouts snout NNS 32505 193 35 , , , 32505 193 36 smoother smoother NN 32505 193 37 skin skin NN 32505 193 38 , , , 32505 193 39 and and CC 32505 193 40 dissimilar dissimilar JJ 32505 193 41 color color NN 32505 193 42 patterns pattern NNS 32505 193 43 . . . 32505 194 1 _ _ NNP 32505 194 2 Distribution._--In distribution._--in NN 32505 194 3 Central Central NNP 32505 194 4 America America NNP 32505 194 5 _ _ NNP 32505 194 6 Hyla Hyla NNP 32505 194 7 boulengeri boulengeri NN 32505 194 8 _ _ NNP 32505 194 9 inhabits inhabit VBZ 32505 194 10 the the DT 32505 194 11 forested forested JJ 32505 194 12 lowlands lowland NNS 32505 194 13 in in IN 32505 194 14 locally locally RB 32505 194 15 humid humid JJ 32505 194 16 areas area NNS 32505 194 17 in in IN 32505 194 18 Guanacaste Guanacaste NNP 32505 194 19 Province Province NNP 32505 194 20 , , , 32505 194 21 Costa Costa NNP 32505 194 22 Rica Rica NNP 32505 194 23 , , , 32505 194 24 and and CC 32505 194 25 in in IN 32505 194 26 the the DT 32505 194 27 humid humid NNP 32505 194 28 Golfo Golfo NNP 32505 194 29 Dulce Dulce NNP 32505 194 30 region region NN 32505 194 31 of of IN 32505 194 32 Costa Costa NNP 32505 194 33 Rica Rica NNP 32505 194 34 ; ; : 32505 194 35 it -PRON- PRP 32505 194 36 occurs occur VBZ 32505 194 37 on on IN 32505 194 38 the the DT 32505 194 39 Carribbean carribbean JJ 32505 194 40 lowlands lowland NNS 32505 194 41 from from IN 32505 194 42 central central JJ 32505 194 43 Nicaragua Nicaragua NNP 32505 194 44 to to IN 32505 194 45 South South NNP 32505 194 46 America America NNP 32505 194 47 , , , 32505 194 48 where where WRB 32505 194 49 it -PRON- PRP 32505 194 50 ranges range VBZ 32505 194 51 to to IN 32505 194 52 Guyana Guyana NNP 32505 194 53 and and CC 32505 194 54 Ecuador Ecuador NNP 32505 194 55 . . . 32505 195 1 The the DT 32505 195 2 highest high JJS 32505 195 3 elevations elevation NNS 32505 195 4 where where WRB 32505 195 5 _ _ NNP 32505 195 6 H. H. NNP 32505 196 1 boulengeri boulengeri NNP 32505 196 2 _ _ NNP 32505 196 3 has have VBZ 32505 196 4 been be VBN 32505 196 5 found find VBN 32505 196 6 are be VBP 32505 196 7 620 620 CD 32505 196 8 meters meter NNS 32505 196 9 at at IN 32505 196 10 Turrialba Turrialba NNP 32505 196 11 , , , 32505 196 12 Cartago Cartago NNP 32505 196 13 Province Province NNP 32505 196 14 , , , 32505 196 15 and and CC 32505 196 16 700 700 CD 32505 196 17 meters meter NNS 32505 196 18 at at IN 32505 196 19 Tilarán Tilarán NNP 32505 196 20 , , , 32505 196 21 Guanacaste Guanacaste NNP 32505 196 22 Province Province NNP 32505 196 23 , , , 32505 196 24 Costa Costa NNP 32505 196 25 Rica Rica NNP 32505 196 26 ( ( -LRB- 32505 196 27 Fig Fig NNP 32505 196 28 . . . 32505 197 1 4 4 LS 32505 197 2 ) ) -RRB- 32505 197 3 . . . 32505 198 1 _ _ NNP 32505 198 2 Specimens Specimens NNP 32505 198 3 Examined._--Costa Examined._--Costa NNP 32505 198 4 Rica Rica NNP 32505 198 5 : : : 32505 198 6 _ _ NNP 32505 198 7 Alajuela Alajuela NNP 32505 198 8 _ _ NNP 32505 198 9 : : : 32505 198 10 9 9 CD 32505 198 11 km km NN 32505 198 12 N n CD 32505 198 13 Ciudad Ciudad NNP 32505 198 14 Quesada Quesada NNP 32505 198 15 , , , 32505 198 16 near near IN 32505 198 17 La La NNP 32505 198 18 Florencia Florencia NNP 32505 198 19 , , , 32505 198 20 USC USC NNP 32505 198 21 8059 8059 CD 32505 198 22 ( ( -LRB- 32505 198 23 4 4 CD 32505 198 24 ) ) -RRB- 32505 198 25 ; ; : 32505 198 26 18 18 CD 32505 198 27 km km NN 32505 198 28 N N NNP 32505 198 29 Florencia Florencia NNP 32505 198 30 , , , 32505 198 31 USC USC NNP 32505 198 32 2624 2624 CD 32505 198 33 ; ; : 32505 198 34 Laguna Laguna NNP 32505 198 35 Monte Monte NNP 32505 198 36 Alegre Alegre NNP 32505 198 37 , , , 32505 198 38 KU KU NNP 32505 198 39 64334 64334 CD 32505 198 40 ; ; : 32505 198 41 Las Las NNP 32505 198 42 Playuelas Playuelas NNP 32505 198 43 , , , 32505 198 44 11 11 CD 32505 198 45 km km NN 32505 198 46 S S NNP 32505 198 47 Los Los NNP 32505 198 48 Chiles Chiles NNP 32505 198 49 , , , 32505 198 50 USC USC NNP 32505 198 51 7216 7216 CD 32505 198 52 , , , 32505 198 53 7217 7217 CD 32505 198 54 ( ( -LRB- 32505 198 55 2 2 CD 32505 198 56 ) ) -RRB- 32505 198 57 , , , 32505 198 58 7219 7219 CD 32505 198 59 ; ; : 32505 198 60 3 3 LS 32505 198 61 km km NN 32505 198 62 NE NE NNP 32505 198 63 Muelle Muelle NNP 32505 198 64 del del NNP 32505 198 65 Arenal Arenal NNP 32505 198 66 , , , 32505 198 67 USC USC NNP 32505 198 68 2644 2644 CD 32505 198 69 ( ( -LRB- 32505 198 70 5 5 CD 32505 198 71 ) ) -RRB- 32505 198 72 . . . 32505 199 1 _ _ NNP 32505 199 2 Cartago Cartago NNP 32505 199 3 _ _ NNP 32505 199 4 : : : 32505 199 5 Turrialba Turrialba NNP 32505 199 6 , , , 32505 199 7 KU KU NNP 32505 199 8 24741 24741 CD 32505 199 9 . . . 32505 200 1 _ _ NNP 32505 200 2 Guanacaste Guanacaste NNP 32505 200 3 _ _ NNP 32505 200 4 : : : 32505 200 5 7 7 CD 32505 200 6 km km NN 32505 200 7 N N NNP 32505 200 8 Liberia Liberia NNP 32505 200 9 , , , 32505 200 10 USC USC NNP 32505 200 11 8096 8096 CD 32505 200 12 ( ( -LRB- 32505 200 13 2 2 CD 32505 200 14 ) ) -RRB- 32505 200 15 , , , 32505 200 16 8138 8138 CD 32505 200 17 ( ( -LRB- 32505 200 18 6 6 CD 32505 200 19 ) ) -RRB- 32505 200 20 ; ; : 32505 200 21 13.6 13.6 CD 32505 200 22 km km NN 32505 200 23 N N NNP 32505 200 24 Liberia Liberia NNP 32505 200 25 , , , 32505 200 26 USC USC NNP 32505 200 27 8151 8151 CD 32505 200 28 , , , 32505 200 29 8171 8171 CD 32505 200 30 ( ( -LRB- 32505 200 31 2 2 CD 32505 200 32 ) ) -RRB- 32505 200 33 ; ; : 32505 200 34 20.5 20.5 LS 32505 200 35 km km NN 32505 200 36 S S NNP 32505 200 37 Liberia Liberia NNP 32505 200 38 , , , 32505 200 39 USC USC NNP 32505 200 40 8205 8205 CD 32505 200 41 ; ; : 32505 200 42 Taboga Taboga NNP 32505 200 43 , , , 32505 200 44 20 20 CD 32505 200 45 km km NN 32505 200 46 SE SE NNP 32505 200 47 Las Las NNP 32505 200 48 Cañas Cañas NNP 32505 200 49 , , , 32505 200 50 KU KU NNP 32505 200 51 102170 102170 CD 32505 200 52 , , , 32505 200 53 USC USC NNP 32505 200 54 7166 7166 CD 32505 200 55 ; ; : 32505 200 56 4 4 LS 32505 200 57 km km NN 32505 200 58 NE NE NNP 32505 200 59 Tilarán Tilarán NNP 32505 200 60 , , , 32505 200 61 USC USC NNP 32505 200 62 8023 8023 CD 32505 200 63 ; ; : 32505 200 64 6 6 CD 32505 200 65 km km NN 32505 200 66 NE NE NNP 32505 200 67 Tilarán Tilarán NNP 32505 200 68 , , , 32505 200 69 USC USC NNP 32505 200 70 523 523 CD 32505 200 71 ( ( -LRB- 32505 200 72 3 3 CD 32505 200 73 ) ) -RRB- 32505 200 74 , , , 32505 200 75 6262 6262 CD 32505 200 76 , , , 32505 200 77 7019 7019 CD 32505 200 78 . . . 32505 201 1 _ _ NNP 32505 201 2 Heredia Heredia NNP 32505 201 3 _ _ NNP 32505 201 4 : : : 32505 201 5 Puerto Puerto NNP 32505 201 6 Viejo Viejo NNP 32505 201 7 , , , 32505 201 8 KU KU NNP 32505 201 9 64323 64323 CD 32505 201 10 - - SYM 32505 201 11 7 7 CD 32505 201 12 ( ( -LRB- 32505 201 13 skeletons skeleton NNS 32505 201 14 ) ) -RRB- 32505 201 15 , , , 32505 201 16 104351 104351 CD 32505 201 17 - - SYM 32505 201 18 3 3 CD 32505 201 19 ( ( -LRB- 32505 201 20 skeletons skeleton NNS 32505 201 21 ) ) -RRB- 32505 201 22 , , , 32505 201 23 64330 64330 CD 32505 201 24 - - SYM 32505 201 25 3 3 CD 32505 201 26 , , , 32505 201 27 103592 103592 CD 32505 201 28 - - SYM 32505 201 29 620 620 CD 32505 201 30 ; ; : 32505 201 31 1 1 CD 32505 201 32 km km NN 32505 201 33 NE NE NNP 32505 201 34 Puerto Puerto NNP 32505 201 35 Viejo Viejo NNP 32505 201 36 , , , 32505 201 37 UMMZ UMMZ NNP 32505 201 38 126042 126042 CD 32505 201 39 ; ; : 32505 201 40 1 1 LS 32505 201 41 km km NN 32505 201 42 S S NNP 32505 201 43 Puerto Puerto NNP 32505 201 44 Viejo Viejo NNP 32505 201 45 , , , 32505 201 46 KU KU NNP 32505 201 47 84983 84983 CD 32505 201 48 - - SYM 32505 201 49 4 4 CD 32505 201 50 ( ( -LRB- 32505 201 51 skeletons skeleton NNS 32505 201 52 ) ) -RRB- 32505 201 53 , , , 32505 201 54 86317 86317 CD 32505 201 55 - - SYM 32505 201 56 22 22 CD 32505 201 57 , , , 32505 201 58 87774 87774 CD 32505 201 59 ( ( -LRB- 32505 201 60 skeleton skeleton NNP 32505 201 61 ) ) -RRB- 32505 201 62 ; ; : 32505 201 63 4.2 4.2 CD 32505 201 64 km km NN 32505 201 65 W W NNP 32505 201 66 Puerto Puerto NNP 32505 201 67 Viejo Viejo NNP 32505 201 68 , , , 32505 201 69 KU KU NNP 32505 201 70 64329 64329 CD 32505 201 71 , , , 32505 201 72 64328 64328 CD 32505 201 73 ( ( -LRB- 32505 201 74 skeleton skeleton NNP 32505 201 75 ) ) -RRB- 32505 201 76 . . . 32505 202 1 _ _ NNP 32505 202 2 Limón Limón NNP 32505 202 3 _ _ NNP 32505 202 4 : : : 32505 202 5 Mountain Mountain NNP 32505 202 6 Cow Cow NNP 32505 202 7 Creek Creek NNP 32505 202 8 , , , 32505 202 9 near near IN 32505 202 10 Banano Banano NNP 32505 202 11 , , , 32505 202 12 KU KU NNP 32505 202 13 37031 37031 CD 32505 202 14 , , , 32505 202 15 41067 41067 CD 32505 202 16 ( ( -LRB- 32505 202 17 skeleton skeleton NNP 32505 202 18 ) ) -RRB- 32505 202 19 ; ; : 32505 202 20 3 3 LS 32505 202 21 km km NN 32505 202 22 S S NNP 32505 202 23 Río Río NNP 32505 202 24 Tortuguero Tortuguero NNP 32505 202 25 , , , 32505 202 26 AMNH AMNH NNP 32505 202 27 69057 69057 CD 32505 202 28 ; ; : 32505 202 29 Suretka Suretka NNP 32505 202 30 , , , 32505 202 31 KU KU NNP 32505 202 32 36482 36482 CD 32505 202 33 - - SYM 32505 202 34 8 8 CD 32505 202 35 , , , 32505 202 36 36699 36699 CD 32505 202 37 . . . 32505 203 1 _ _ NNP 32505 203 2 Puntarenas Puntarenas NNP 32505 203 3 _ _ NNP 32505 203 4 : : : 32505 203 5 4.8 4.8 CD 32505 203 6 km km NN 32505 203 7 S S NNP 32505 203 8 Bahía Bahía NNP 32505 203 9 Rincón Rincón NNP 32505 203 10 on on IN 32505 203 11 NW NW NNP 32505 203 12 side side NN 32505 203 13 Río Río NNP 32505 203 14 Rincón Rincón NNP 32505 203 15 , , , 32505 203 16 USC USC NNP 32505 203 17 705 705 CD 32505 203 18 ; ; : 32505 203 19 Parrita Parrita NNP 32505 203 20 , , , 32505 203 21 USC USC NNP 32505 203 22 6163 6163 CD 32505 203 23 ; ; : 32505 203 24 4.5 4.5 CD 32505 203 25 km km NN 32505 203 26 W W NNP 32505 203 27 Rincón Rincón NNP 32505 203 28 de de NNP 32505 203 29 Osa Osa NNP 32505 203 30 , , , 32505 203 31 KU KU NNP 32505 203 32 102177 102177 CD 32505 203 33 - - SYM 32505 203 34 9 9 CD 32505 203 35 , , , 32505 203 36 104295 104295 CD 32505 203 37 - - SYM 32505 203 38 6 6 CD 32505 203 39 ( ( -LRB- 32505 203 40 tadpoles tadpole NNS 32505 203 41 ) ) -RRB- 32505 203 42 ; ; : 32505 203 43 6 6 CD 32505 203 44 km km NN 32505 203 45 SW SW NNP 32505 203 46 Rincón Rincón NNP 32505 203 47 de de NNP 32505 203 48 Osa Osa NNP 32505 203 49 , , , 32505 203 50 KU KU NNP 32505 203 51 102171 102171 CD 32505 203 52 - - SYM 32505 203 53 6 6 CD 32505 203 54 ; ; : 32505 203 55 4.4 4.4 CD 32505 203 56 km km NN 32505 203 57 NW NW NNP 32505 203 58 Villa Villa NNP 32505 203 59 Neilly Neilly NNP 32505 203 60 , , , 32505 203 61 USC USC NNP 32505 203 62 8003 8003 CD 32505 203 63 ; ; : 32505 203 64 10.5 10.5 CD 32505 203 65 km km NN 32505 203 66 WNW WNW NNP 32505 203 67 Villa Villa NNP 32505 203 68 Neilly Neilly NNP 32505 203 69 , , , 32505 203 70 KU KU NNP 32505 203 71 64321 64321 CD 32505 203 72 . . . 32505 204 1 _ _ NNP 32505 204 2 San San NNP 32505 204 3 José José NNP 32505 204 4 _ _ NNP 32505 204 5 : : : 32505 204 6 21 21 CD 32505 204 7 km km NN 32505 204 8 WSW WSW NNP 32505 204 9 San San NNP 32505 204 10 Isidro Isidro NNP 32505 204 11 el el NNP 32505 204 12 General General NNP 32505 204 13 , , , 32505 204 14 KU KU NNP 32505 204 15 34104 34104 CD 32505 204 16 - - SYM 32505 204 17 6 6 CD 32505 204 18 . . . 32505 205 1 [ [ -LRB- 32505 205 2 Illustration illustration NN 32505 205 3 : : : 32505 205 4 Fig Fig NNP 32505 205 5 . . . 32505 206 1 4 4 LS 32505 206 2 . . . 32505 207 1 Map Map NNP 32505 207 2 showing show VBG 32505 207 3 locality locality NN 32505 207 4 records record NNS 32505 207 5 for for IN 32505 207 6 _ _ NNP 32505 207 7 Hyla Hyla NNP 32505 207 8 boulengeri boulengeri NN 32505 207 9 _ _ NNP 32505 207 10 ( ( -LRB- 32505 207 11 circles circle NNS 32505 207 12 ) ) -RRB- 32505 207 13 and and CC 32505 207 14 _ _ NNP 32505 207 15 H. H. NNP 32505 208 1 foliamorta foliamorta NNP 32505 208 2 _ _ NNP 32505 208 3 ( ( -LRB- 32505 208 4 dots dot NNS 32505 208 5 ) ) -RRB- 32505 208 6 . . . 32505 208 7 ] ] -RRB- 32505 209 1 Panamá panamá NN 32505 209 2 : : : 32505 209 3 _ _ NNP 32505 209 4 Bocas Bocas NNP 32505 209 5 del del NNP 32505 209 6 Toro Toro NNP 32505 209 7 _ _ NNP 32505 209 8 : : : 32505 209 9 3.2 3.2 CD 32505 209 10 km km NN 32505 209 11 W W NNP 32505 209 12 Almirante Almirante NNP 32505 209 13 , , , 32505 209 14 KU KU NNP 32505 209 15 95978 95978 CD 32505 209 16 . . . 32505 210 1 _ _ NNP 32505 210 2 Canal Canal NNP 32505 210 3 Zone Zone NNP 32505 210 4 _ _ NNP 32505 210 5 : : : 32505 210 6 Barro Barro NNP 32505 210 7 Colorado Colorado NNP 32505 210 8 Island Island NNP 32505 210 9 , , , 32505 210 10 FMNH FMNH NNP 32505 210 11 13379 13379 CD 32505 210 12 ; ; : 32505 210 13 near near IN 32505 210 14 Clayton Clayton NNP 32505 210 15 Reservation Reservation NNP 32505 210 16 , , , 32505 210 17 UIMNH UIMNH NNP 32505 210 18 42000 42000 CD 32505 210 19 ; ; : 32505 210 20 2.6 2.6 CD 32505 210 21 km km NN 32505 210 22 SW SW NNP 32505 210 23 Fort Fort NNP 32505 210 24 Kobbe Kobbe NNP 32505 210 25 , , , 32505 210 26 KU KU NNP 32505 210 27 95977 95977 CD 32505 210 28 ; ; : 32505 210 29 Miraflores Miraflores NNP 32505 210 30 Locks Locks NNPS 32505 210 31 , , , 32505 210 32 AMNH AMNH NNP 32505 210 33 69764 69764 CD 32505 210 34 - - SYM 32505 210 35 5 5 CD 32505 210 36 ; ; : 32505 210 37 Summit Summit NNP 32505 210 38 , , , 32505 210 39 AMNH AMNH NNP 32505 210 40 73445 73445 CD 32505 210 41 , , , 32505 210 42 KU KU NNP 32505 210 43 97777 97777 CD 32505 210 44 , , , 32505 210 45 101540 101540 CD 32505 210 46 - - SYM 32505 210 47 9 9 CD 32505 210 48 , , , 32505 210 49 104350 104350 CD 32505 210 50 ( ( -LRB- 32505 210 51 skeleton skeleton NN 32505 210 52 ) ) -RRB- 32505 210 53 . . . 32505 211 1 _ _ NNP 32505 211 2 Colón Colón NNP 32505 211 3 _ _ NNP 32505 211 4 : : : 32505 211 5 Río Río NNP 32505 211 6 Gatuncillo Gatuncillo NNP 32505 211 7 , , , 32505 211 8 near near IN 32505 211 9 Nuevo Nuevo NNP 32505 211 10 San San NNP 32505 211 11 Juan Juan NNP 32505 211 12 , , , 32505 211 13 KU KU NNP 32505 211 14 95976 95976 CD 32505 211 15 . . . 32505 212 1 _ _ NNP 32505 212 2 Darién Darién NNP 32505 212 3 _ _ NNP 32505 212 4 : : : 32505 212 5 El El NNP 32505 212 6 Real Real NNP 32505 212 7 , , , 32505 212 8 KU KU NNP 32505 212 9 80451 80451 CD 32505 212 10 - - SYM 32505 212 11 3 3 CD 32505 212 12 . . . 32505 213 1 _ _ NNP 32505 213 2 Hyla Hyla NNP 32505 213 3 foliamorta foliamorta NN 32505 213 4 _ _ NNP 32505 213 5 Fouquette Fouquette NNP 32505 213 6 _ _ NNP 32505 213 7 Hyla Hyla NNP 32505 213 8 foliamorta foliamorta NN 32505 213 9 _ _ NNP 32505 213 10 Fouquette Fouquette NNP 32505 213 11 , , , 32505 213 12 Herpetologica Herpetologica NNP 32505 213 13 , , , 32505 213 14 14:125 14:125 CD 32505 213 15 , , , 32505 213 16 April April NNP 32505 213 17 25 25 CD 32505 213 18 , , , 32505 213 19 1958 1958 CD 32505 213 20 [ [ -LRB- 32505 213 21 Holotype.--TNHC Holotype.--TNHC NNP 32505 213 22 23109 23109 CD 32505 213 23 , , , 32505 213 24 11 11 CD 32505 213 25 km km NN 32505 213 26 . . . 32505 214 1 NW NW NNP 32505 214 2 Miraflores Miraflores NNP 32505 214 3 Locks Locks NNP 32505 214 4 , , , 32505 214 5 Canal Canal NNP 32505 214 6 Zone Zone NNP 32505 214 7 , , , 32505 214 8 Panamá Panamá NNP 32505 214 9 ; ; : 32505 214 10 M. M. NNP 32505 214 11 J. J. NNP 32505 214 12 Fouquette Fouquette NNP 32505 214 13 , , , 32505 214 14 Jr. Jr. NNP 32505 214 15 collector collector NN 32505 214 16 ] ] -RRB- 32505 214 17 . . . 32505 215 1 _ _ NNP 32505 215 2 Diagnosis._--Size Diagnosis._--Size NNP 32505 215 3 medium medium NN 32505 215 4 ( ( -LRB- 32505 215 5 Male male NN 32505 215 6 to to IN 32505 215 7 43 43 CD 32505 215 8 mm mm NNS 32505 215 9 . . NNP 32505 215 10 , , , 32505 215 11 Female Female NNP 32505 215 12 to to IN 32505 215 13 41 41 CD 32505 215 14 mm mm NNS 32505 215 15 . . . 32505 216 1 ) ) -RRB- 32505 216 2 ; ; : 32505 216 3 skull skull NNP 32505 216 4 longer long RBR 32505 216 5 than than IN 32505 216 6 wide wide JJ 32505 216 7 ; ; : 32505 216 8 frontoparietal frontoparietal JJ 32505 216 9 fontanelle fontanelle NN 32505 216 10 absent absent NN 32505 216 11 ; ; : 32505 216 12 snout snout NN 32505 216 13 acuminate acuminate NN 32505 216 14 , , , 32505 216 15 projecting projecting NN 32505 216 16 ; ; : 32505 216 17 interorbital interorbital NNP 32505 216 18 triangle triangle NNP 32505 216 19 bordered border VBN 32505 216 20 by by IN 32505 216 21 white white JJ 32505 216 22 lines line NNS 32505 216 23 ; ; : 32505 216 24 scapular scapular JJ 32505 216 25 region region NN 32505 216 26 having have VBG 32505 216 27 two two CD 32505 216 28 or or CC 32505 216 29 more more JJR 32505 216 30 elongate elongate NN 32505 216 31 spots spot NNS 32505 216 32 ; ; : 32505 216 33 dorsum dorsum NN 32505 216 34 smooth smooth JJ 32505 216 35 ; ; : 32505 216 36 vocal vocal JJ 32505 216 37 sac sac NN 32505 216 38 dark dark JJ 32505 216 39 gray gray NNP 32505 216 40 ; ; : 32505 216 41 groin groin NNP 32505 216 42 creamy creamy NNP 32505 216 43 white white NNP 32505 216 44 ; ; : 32505 216 45 traces trace NNS 32505 216 46 of of IN 32505 216 47 web web NN 32505 216 48 between between IN 32505 216 49 fingers finger NNS 32505 216 50 . . . 32505 217 1 _ _ NNP 32505 217 2 Description._--Head Description._--Head NNP 32505 217 3 flattened flatten VBD 32505 217 4 , , , 32505 217 5 longer longer RB 32505 217 6 than than IN 32505 217 7 wide wide JJ 32505 217 8 ; ; : 32505 217 9 snout snout NN 32505 217 10 flat flat JJ 32505 217 11 , , , 32505 217 12 pointed pointed JJ 32505 217 13 , , , 32505 217 14 protruding protrude VBG 32505 217 15 beyond beyond IN 32505 217 16 lower low JJR 32505 217 17 lip lip NN 32505 217 18 ; ; : 32505 217 19 loreal loreal NN 32505 217 20 region region NN 32505 217 21 slightly slightly RB 32505 217 22 concave concave NNP 32505 217 23 ; ; : 32505 217 24 canthus canthus NNP 32505 217 25 moderately moderately RB 32505 217 26 prominent prominent JJ 32505 217 27 ; ; : 32505 217 28 eyes eye NNS 32505 217 29 smaller small JJR 32505 217 30 than than IN 32505 217 31 interorbital interorbital NNP 32505 217 32 space space NN 32505 217 33 ; ; : 32505 217 34 tympanum tympanum NN 32505 217 35 distinct distinct JJ 32505 217 36 , , , 32505 217 37 55 55 CD 32505 217 38 to to TO 32505 217 39 75 75 CD 32505 217 40 per per NN 32505 217 41 cent cent NN 32505 217 42 of of IN 32505 217 43 diameter diameter NN 32505 217 44 of of IN 32505 217 45 eye eye NN 32505 217 46 , , , 32505 217 47 smaller small JJR 32505 217 48 than than IN 32505 217 49 internarial internarial JJ 32505 217 50 space space NN 32505 217 51 ; ; : 32505 217 52 arms arm NNS 32505 217 53 short short JJ 32505 217 54 ; ; : 32505 217 55 fingers finger NNS 32505 217 56 having have VBG 32505 217 57 rudimentary rudimentary NNP 32505 217 58 webs webs NN 32505 217 59 ; ; : 32505 217 60 median median JJ 32505 217 61 palmar palmar NNP 32505 217 62 tubercle tubercle NN 32505 217 63 tripartite tripartite NNP 32505 217 64 ; ; , 32505 217 65 inner inner JJ 32505 217 66 palmar palmar NNP 32505 217 67 tubercle tubercle NN 32505 217 68 on on IN 32505 217 69 base base NN 32505 217 70 of of IN 32505 217 71 first first NNP 32505 217 72 finger finger NNP 32505 217 73 flat flat JJ 32505 217 74 ; ; : 32505 217 75 subarticular subarticular JJ 32505 217 76 tubercles tubercle NNS 32505 217 77 distinct distinct JJ 32505 217 78 ; ; : 32505 217 79 discs disc NNS 32505 217 80 of of IN 32505 217 81 fingers finger NNS 32505 217 82 smaller small JJR 32505 217 83 than than IN 32505 217 84 diameter diameter NN 32505 217 85 of of IN 32505 217 86 tympanum tympanum NN 32505 217 87 ; ; : 32505 217 88 legs leg NNS 32505 217 89 long long RB 32505 217 90 ; ; : 32505 217 91 tarsal tarsal JJ 32505 217 92 fold fold VB 32505 217 93 lacking lacking JJ 32505 217 94 ; ; : 32505 217 95 inner inner JJ 32505 217 96 metatarsal metatarsal NN 32505 217 97 tubercle tubercle NN 32505 217 98 larger large JJR 32505 217 99 than than IN 32505 217 100 outer outer JJ 32505 217 101 ; ; : 32505 217 102 one one CD 32505 217 103 phalanx phalanx NN 32505 217 104 free free JJ 32505 217 105 on on IN 32505 217 106 second second JJ 32505 217 107 , , , 32505 217 108 third third JJ 32505 217 109 , , , 32505 217 110 and and CC 32505 217 111 fifth fifth JJ 32505 217 112 toes toe NNS 32505 217 113 , , , 32505 217 114 two two CD 32505 217 115 and and CC 32505 217 116 one one CD 32505 217 117 half half NN 32505 217 118 phalanges phalanx NNS 32505 217 119 free free JJ 32505 217 120 on on IN 32505 217 121 fourth fourth JJ 32505 217 122 toe toe NN 32505 217 123 ; ; , 32505 217 124 narrow narrow JJ 32505 217 125 fringe fringe NN 32505 217 126 continuing continue VBG 32505 217 127 from from IN 32505 217 128 web web NN 32505 217 129 to to IN 32505 217 130 discs disc NNS 32505 217 131 of of IN 32505 217 132 toes toe NNS 32505 217 133 ; ; : 32505 217 134 discs disc NNS 32505 217 135 of of IN 32505 217 136 toes toe NNS 32505 217 137 about about IN 32505 217 138 the the DT 32505 217 139 size size NN 32505 217 140 of of IN 32505 217 141 those those DT 32505 217 142 on on IN 32505 217 143 fingers finger NNS 32505 217 144 ; ; : 32505 217 145 skin skin NN 32505 217 146 smooth smooth JJ 32505 217 147 on on IN 32505 217 148 dorsum dorsum NN 32505 217 149 and and CC 32505 217 150 flanks flank NNS 32505 217 151 , , , 32505 217 152 that that IN 32505 217 153 on on IN 32505 217 154 belly belly NN 32505 217 155 and and CC 32505 217 156 posterior posterior NN 32505 217 157 part part NN 32505 217 158 of of IN 32505 217 159 thighs thigh NNS 32505 217 160 granular granular JJ 32505 217 161 ; ; : 32505 217 162 tongue tongue NN 32505 217 163 oval oval NN 32505 217 164 , , , 32505 217 165 longer longer RB 32505 217 166 than than IN 32505 217 167 wide wide JJ 32505 217 168 ; ; : 32505 217 169 vocal vocal JJ 32505 217 170 slits slit NNS 32505 217 171 oblique oblique NNP 32505 217 172 , , , 32505 217 173 about about RB 32505 217 174 one one CD 32505 217 175 half half JJ 32505 217 176 length length NN 32505 217 177 of of IN 32505 217 178 tongue tongue NN 32505 217 179 . . . 32505 218 1 In in IN 32505 218 2 life life NN 32505 218 3 , , , 32505 218 4 dorsum dorsum NN 32505 218 5 pale pale JJ 32505 218 6 tan tan NN 32505 218 7 to to IN 32505 218 8 pale pale JJ 32505 218 9 reddish reddish JJ 32505 218 10 brown brown NN 32505 218 11 with with IN 32505 218 12 irregular irregular JJ 32505 218 13 reddish reddish JJ 32505 218 14 brown brown JJ 32505 218 15 markings marking NNS 32505 218 16 ; ; : 32505 218 17 small small JJ 32505 218 18 dark dark JJ 32505 218 19 spots spot NNS 32505 218 20 on on IN 32505 218 21 head head NN 32505 218 22 ; ; : 32505 218 23 distinct distinct JJ 32505 218 24 dark dark JJ 32505 218 25 brown brown JJ 32505 218 26 triangular triangular NNP 32505 218 27 mark mark NN 32505 218 28 between between IN 32505 218 29 eyes eye NNS 32505 218 30 , , , 32505 218 31 bordered border VBN 32505 218 32 by by IN 32505 218 33 thin thin JJ 32505 218 34 white white JJ 32505 218 35 lines line NNS 32505 218 36 ; ; : 32505 218 37 apex apex NNP 32505 218 38 of of IN 32505 218 39 triangle triangle NNP 32505 218 40 always always RB 32505 218 41 directed direct VBD 32505 218 42 backward backward RB 32505 218 43 ; ; : 32505 218 44 supratympanic supratympanic NNP 32505 218 45 fold fold VBP 32505 218 46 with with IN 32505 218 47 black black JJ 32505 218 48 edge edge NN 32505 218 49 ; ; , 32505 218 50 scapular scapular JJ 32505 218 51 region region NN 32505 218 52 having have VBG 32505 218 53 two two CD 32505 218 54 to to TO 32505 218 55 five five CD 32505 218 56 small small JJ 32505 218 57 , , , 32505 218 58 elongate elongate VB 32505 218 59 black black JJ 32505 218 60 spots spot NNS 32505 218 61 ; ; : 32505 218 62 belly belly NN 32505 218 63 creamy creamy VB 32505 218 64 tan tan NNP 32505 218 65 with with IN 32505 218 66 small small JJ 32505 218 67 brown brown JJ 32505 218 68 spots spot NNS 32505 218 69 ; ; : 32505 218 70 vocal vocal JJ 32505 218 71 sac sac NN 32505 218 72 uniformly uniformly RB 32505 218 73 dark dark JJ 32505 218 74 brown brown NNP 32505 218 75 with with IN 32505 218 76 scattered scatter VBN 32505 218 77 creamy creamy NNP 32505 218 78 tan tan NNP 32505 218 79 flecks fleck NNS 32505 218 80 ; ; : 32505 218 81 upper upper JJ 32505 218 82 jaw jaw NN 32505 218 83 dark dark JJ 32505 218 84 brown brown NNP 32505 218 85 ; ; : 32505 218 86 limbs limb NNS 32505 218 87 creamy creamy VBP 32505 218 88 white white RB 32505 218 89 below below RB 32505 218 90 with with IN 32505 218 91 scattered scatter VBN 32505 218 92 brown brown JJ 32505 218 93 spots spot NNS 32505 218 94 ; ; : 32505 218 95 groin groin NNP 32505 218 96 marked mark VBN 32505 218 97 with with IN 32505 218 98 small small JJ 32505 218 99 brown brown JJ 32505 218 100 spots spot NNS 32505 218 101 in in IN 32505 218 102 some some DT 32505 218 103 specimens specimen NNS 32505 218 104 ; ; : 32505 218 105 anterior anterior NNP 32505 218 106 and and CC 32505 218 107 posterior posterior NN 32505 218 108 surfaces surface NNS 32505 218 109 of of IN 32505 218 110 thighs thigh NNS 32505 218 111 yellow yellow JJ 32505 218 112 - - HYPH 32505 218 113 orange orange NN 32505 218 114 with with IN 32505 218 115 three three CD 32505 218 116 distinct distinct JJ 32505 218 117 black black JJ 32505 218 118 blotches blotch NNS 32505 218 119 ; ; : 32505 218 120 two two CD 32505 218 121 dark dark JJ 32505 218 122 bands band NNS 32505 218 123 on on IN 32505 218 124 upper upper JJ 32505 218 125 surface surface NN 32505 218 126 of of IN 32505 218 127 shanks shank NNS 32505 218 128 ; ; : 32505 218 129 webbing web VBG 32505 218 130 of of IN 32505 218 131 feet foot NNS 32505 218 132 yellowish yellowish JJ 32505 218 133 tan tan NN 32505 218 134 with with IN 32505 218 135 brown brown JJ 32505 218 136 mottlings mottling NNS 32505 218 137 ( ( -LRB- 32505 218 138 Pl Pl NNP 32505 218 139 . . . 32505 219 1 1B 1B NNP 32505 219 2 ) ) -RRB- 32505 219 3 . . . 32505 220 1 In in IN 32505 220 2 preservative preservative JJ 32505 220 3 , , , 32505 220 4 dorsum dorsum NN 32505 220 5 brown brown JJ 32505 220 6 or or CC 32505 220 7 gray gray JJ 32505 220 8 with with IN 32505 220 9 darker dark JJR 32505 220 10 markings marking NNS 32505 220 11 ; ; , 32505 220 12 interorbital interorbital NNP 32505 220 13 triangle triangle NNP 32505 220 14 distinct distinct NNP 32505 220 15 , , , 32505 220 16 bordered border VBN 32505 220 17 by by IN 32505 220 18 white white JJ 32505 220 19 lines line NNS 32505 220 20 ; ; : 32505 220 21 supratympanic supratympanic NNP 32505 220 22 fold fold VBP 32505 220 23 with with IN 32505 220 24 black black JJ 32505 220 25 edge edge NN 32505 220 26 ; ; : 32505 220 27 two two CD 32505 220 28 or or CC 32505 220 29 more more JJR 32505 220 30 small small JJ 32505 220 31 elongate elongate NN 32505 220 32 black black JJ 32505 220 33 spots spot NNS 32505 220 34 in in IN 32505 220 35 scapular scapular JJ 32505 220 36 region region NN 32505 220 37 ; ; : 32505 220 38 belly belly NN 32505 220 39 white white JJ 32505 220 40 with with IN 32505 220 41 numerous numerous JJ 32505 220 42 brown brown JJ 32505 220 43 flecks fleck NNS 32505 220 44 ; ; : 32505 220 45 edge edge NN 32505 220 46 of of IN 32505 220 47 upper upper JJ 32505 220 48 lip lip NNP 32505 220 49 dark dark JJ 32505 220 50 brown brown NNP 32505 220 51 ; ; : 32505 220 52 vocal vocal JJ 32505 220 53 sac sac NN 32505 220 54 dark dark JJ 32505 220 55 gray gray NN 32505 220 56 ; ; : 32505 220 57 undersides underside NNS 32505 220 58 of of IN 32505 220 59 limbs limb NNS 32505 220 60 creamy creamy VBP 32505 220 61 white white NNP 32505 220 62 ; ; : 32505 220 63 groin groin NN 32505 220 64 creamy creamy JJ 32505 220 65 white white JJ 32505 220 66 with with IN 32505 220 67 or or CC 32505 220 68 without without IN 32505 220 69 brown brown JJ 32505 220 70 spots spot NNS 32505 220 71 ; ; : 32505 220 72 anterior anterior NNP 32505 220 73 and and CC 32505 220 74 posterior posterior NN 32505 220 75 surfaces surface NNS 32505 220 76 of of IN 32505 220 77 thighs thigh NNS 32505 220 78 having have VBG 32505 220 79 three three CD 32505 220 80 black black JJ 32505 220 81 blotches blotch NNS 32505 220 82 separated separate VBN 32505 220 83 by by IN 32505 220 84 creamy creamy JJ 32505 220 85 white white JJ 32505 220 86 spaces space NNS 32505 220 87 ; ; : 32505 220 88 shanks shank NNS 32505 220 89 having have VBG 32505 220 90 two two CD 32505 220 91 brown brown JJ 32505 220 92 bands band NNS 32505 220 93 ; ; : 32505 220 94 webbing web VBG 32505 220 95 of of IN 32505 220 96 feet foot NNS 32505 220 97 mottled mottle VBN 32505 220 98 with with IN 32505 220 99 brown brown NNP 32505 220 100 . . . 32505 221 1 _ _ NNP 32505 221 2 Variation._--Twenty Variation._--Twenty NNP 32505 221 3 - - HYPH 32505 221 4 eight eight CD 32505 221 5 breeding breeding NN 32505 221 6 males male NNS 32505 221 7 from from IN 32505 221 8 the the DT 32505 221 9 area area NN 32505 221 10 between between IN 32505 221 11 Chepo Chepo NNP 32505 221 12 and and CC 32505 221 13 Tocumen Tocumen NNP 32505 221 14 , , , 32505 221 15 Panamá Panamá NNP 32505 221 16 , , , 32505 221 17 have have VBP 32505 221 18 snout snout NN 32505 221 19 - - HYPH 32505 221 20 vent vent NN 32505 221 21 lengths length NNS 32505 221 22 of of IN 32505 221 23 39.0 39.0 CD 32505 221 24 mm mm NNS 32505 221 25 . . . 32505 222 1 to to IN 32505 222 2 46.0 46.0 CD 32505 222 3 mm mm NNS 32505 222 4 . . . 32505 223 1 ( ( -LRB- 32505 223 2 mean mean NNP 32505 223 3 42.5 42.5 CD 32505 223 4 mm mm NNP 32505 223 5 . . . 32505 223 6 ) ) -RRB- 32505 223 7 . . . 32505 224 1 In in IN 32505 224 2 these these DT 32505 224 3 specimens specimen NNS 32505 224 4 , , , 32505 224 5 the the DT 32505 224 6 ratio ratio NN 32505 224 7 of of IN 32505 224 8 the the DT 32505 224 9 tibia tibia NNP 32505 224 10 length length NN 32505 224 11 to to IN 32505 224 12 the the DT 32505 224 13 snout snout NN 32505 224 14 - - HYPH 32505 224 15 vent vent NN 32505 224 16 length length NN 32505 224 17 is be VBZ 32505 224 18 0.54 0.54 CD 32505 224 19 to to TO 32505 224 20 0.61 0.61 CD 32505 224 21 ( ( -LRB- 32505 224 22 mean mean JJ 32505 224 23 , , , 32505 224 24 0.57 0.57 CD 32505 224 25 ) ) -RRB- 32505 224 26 ; ; : 32505 224 27 the the DT 32505 224 28 ratio ratio NN 32505 224 29 of of IN 32505 224 30 the the DT 32505 224 31 diameter diameter NN 32505 224 32 of of IN 32505 224 33 the the DT 32505 224 34 tympanum tympanum NN 32505 224 35 to to IN 32505 224 36 that that DT 32505 224 37 of of IN 32505 224 38 the the DT 32505 224 39 eye eye NN 32505 224 40 is be VBZ 32505 224 41 0.55 0.55 CD 32505 224 42 to to TO 32505 224 43 0.75 0.75 CD 32505 224 44 ( ( -LRB- 32505 224 45 mean mean NN 32505 224 46 , , , 32505 224 47 0.67 0.67 CD 32505 224 48 ) ) -RRB- 32505 224 49 . . . 32505 225 1 One one CD 32505 225 2 female female NN 32505 225 3 has have VBZ 32505 225 4 a a DT 32505 225 5 snout snout NN 32505 225 6 - - HYPH 32505 225 7 vent vent NN 32505 225 8 length length NN 32505 225 9 of of IN 32505 225 10 41.0 41.0 CD 32505 225 11 mm mm NNP 32505 225 12 . . NNP 32505 225 13 , , , 32505 225 14 tibia tibia NNP 32505 225 15 / / SYM 32505 225 16 snout snout NN 32505 225 17 - - HYPH 32505 225 18 vent vent NN 32505 225 19 length length NN 32505 225 20 ratio ratio NN 32505 225 21 of of IN 32505 225 22 0.57 0.57 CD 32505 225 23 , , , 32505 225 24 and and CC 32505 225 25 tympanum tympanum NN 32505 225 26 / / SYM 32505 225 27 eye eye NN 32505 225 28 ratio ratio NN 32505 225 29 of of IN 32505 225 30 0.76 0.76 CD 32505 225 31 . . . 32505 226 1 Two two CD 32505 226 2 to to TO 32505 226 3 five five CD 32505 226 4 ( ( -LRB- 32505 226 5 usually usually RB 32505 226 6 three three CD 32505 226 7 ) ) -RRB- 32505 226 8 elongate elongate VB 32505 226 9 black black JJ 32505 226 10 spots spot NNS 32505 226 11 are be VBP 32505 226 12 present present JJ 32505 226 13 in in IN 32505 226 14 the the DT 32505 226 15 scapular scapular JJ 32505 226 16 region region NN 32505 226 17 in in IN 32505 226 18 different different JJ 32505 226 19 individuals individual NNS 32505 226 20 . . . 32505 227 1 The the DT 32505 227 2 flanks flank NNS 32505 227 3 in in IN 32505 227 4 some some DT 32505 227 5 are be VBP 32505 227 6 spotted spot VBN 32505 227 7 with with IN 32505 227 8 brown brown NNP 32505 227 9 ; ; : 32505 227 10 in in IN 32505 227 11 others other NNS 32505 227 12 they -PRON- PRP 32505 227 13 are be VBP 32505 227 14 creamy creamy JJ 32505 227 15 white white JJ 32505 227 16 . . . 32505 228 1 A a DT 32505 228 2 small small JJ 32505 228 3 black black JJ 32505 228 4 spot spot NN 32505 228 5 is be VBZ 32505 228 6 present present JJ 32505 228 7 in in IN 32505 228 8 the the DT 32505 228 9 groin groin NN 32505 228 10 of of IN 32505 228 11 some some DT 32505 228 12 specimens specimen NNS 32505 228 13 . . . 32505 229 1 Usually usually RB 32505 229 2 two two CD 32505 229 3 to to TO 32505 229 4 four four CD 32505 229 5 blotches blotch NNS 32505 229 6 are be VBP 32505 229 7 present present JJ 32505 229 8 on on IN 32505 229 9 the the DT 32505 229 10 anterior anterior NN 32505 229 11 and and CC 32505 229 12 posterior posterior NN 32505 229 13 surfaces surface NNS 32505 229 14 of of IN 32505 229 15 the the DT 32505 229 16 thighs thigh NNS 32505 229 17 ; ; : 32505 229 18 in in IN 32505 229 19 some some DT 32505 229 20 specimens specimen NNS 32505 229 21 the the DT 32505 229 22 blotches blotch NNS 32505 229 23 are be VBP 32505 229 24 reduced reduce VBN 32505 229 25 to to IN 32505 229 26 small small JJ 32505 229 27 spots spot NNS 32505 229 28 . . . 32505 230 1 One one CD 32505 230 2 or or CC 32505 230 3 two two CD 32505 230 4 brown brown JJ 32505 230 5 spots spot NNS 32505 230 6 are be VBP 32505 230 7 present present JJ 32505 230 8 proximally proximally RB 32505 230 9 on on IN 32505 230 10 the the DT 32505 230 11 shanks shank NNS 32505 230 12 in in IN 32505 230 13 most most JJS 32505 230 14 specimens specimen NNS 32505 230 15 . . . 32505 231 1 In in IN 32505 231 2 some some DT 32505 231 3 individuals individual NNS 32505 231 4 tuberculations tuberculation NNS 32505 231 5 are be VBP 32505 231 6 scattered scatter VBN 32505 231 7 on on IN 32505 231 8 the the DT 32505 231 9 head head NN 32505 231 10 and and CC 32505 231 11 in in IN 32505 231 12 the the DT 32505 231 13 tympanic tympanic JJ 32505 231 14 and and CC 32505 231 15 scapular scapular JJ 32505 231 16 regions region NNS 32505 231 17 , , , 32505 231 18 but but CC 32505 231 19 the the DT 32505 231 20 dorsum dorsum NN 32505 231 21 is be VBZ 32505 231 22 smooth smooth JJ 32505 231 23 in in IN 32505 231 24 most most JJS 32505 231 25 specimens specimen NNS 32505 231 26 ; ; : 32505 231 27 the the DT 32505 231 28 belly belly NN 32505 231 29 is be VBZ 32505 231 30 creamy creamy JJ 32505 231 31 white white JJ 32505 231 32 flecked fleck VBN 32505 231 33 with with IN 32505 231 34 brown brown NNP 32505 231 35 . . . 32505 232 1 _ _ NNP 32505 232 2 Cranial Cranial NNP 32505 232 3 Osteology._--The Osteology._--The NNP 32505 232 4 skull skull NN 32505 232 5 of of IN 32505 232 6 _ _ NNP 32505 232 7 Hyla Hyla NNP 32505 232 8 foliamorta foliamorta NN 32505 232 9 _ _ NNP 32505 232 10 is be VBZ 32505 232 11 flat flat JJ 32505 232 12 and and CC 32505 232 13 longer long JJR 32505 232 14 than than IN 32505 232 15 it -PRON- PRP 32505 232 16 is be VBZ 32505 232 17 wide wide JJ 32505 232 18 . . . 32505 233 1 The the DT 32505 233 2 premaxillary premaxillary NN 32505 233 3 is be VBZ 32505 233 4 small small JJ 32505 233 5 and and CC 32505 233 6 bears bear VBZ 32505 233 7 13 13 CD 32505 233 8 to to TO 32505 233 9 16 16 CD 32505 233 10 teeth tooth NNS 32505 233 11 ( ( -LRB- 32505 233 12 mean mean VB 32505 233 13 for for IN 32505 233 14 2 2 CD 32505 233 15 specimens specimen NNS 32505 233 16 , , , 32505 233 17 14.8 14.8 CD 32505 233 18 ) ) -RRB- 32505 233 19 . . . 32505 234 1 The the DT 32505 234 2 alary alary JJ 32505 234 3 process process NN 32505 234 4 of of IN 32505 234 5 the the DT 32505 234 6 premaxillary premaxillary NN 32505 234 7 is be VBZ 32505 234 8 vertical vertical JJ 32505 234 9 and and CC 32505 234 10 concave concave NNP 32505 234 11 posteriorly posteriorly RB 32505 234 12 . . . 32505 235 1 Ventrally ventrally RB 32505 235 2 , , , 32505 235 3 the the DT 32505 235 4 premaxillary premaxillary NN 32505 235 5 is be VBZ 32505 235 6 completely completely RB 32505 235 7 separated separate VBN 32505 235 8 from from IN 32505 235 9 the the DT 32505 235 10 prevomer prevomer NN 32505 235 11 by by IN 32505 235 12 cartilage cartilage NN 32505 235 13 . . . 32505 236 1 The the DT 32505 236 2 maxillary maxillary NN 32505 236 3 is be VBZ 32505 236 4 slender slend JJR 32505 236 5 ; ; : 32505 236 6 each each DT 32505 236 7 bears bear VBZ 32505 236 8 77 77 CD 32505 236 9 to to TO 32505 236 10 84 84 CD 32505 236 11 teeth tooth NNS 32505 236 12 ( ( -LRB- 32505 236 13 mean mean VB 32505 236 14 for for IN 32505 236 15 2 2 CD 32505 236 16 specimens specimen NNS 32505 236 17 , , , 32505 236 18 81 81 CD 32505 236 19 ) ) -RRB- 32505 236 20 . . . 32505 237 1 The the DT 32505 237 2 pars par NNS 32505 237 3 facialis faciali NNS 32505 237 4 of of IN 32505 237 5 the the DT 32505 237 6 maxillary maxillary NN 32505 237 7 is be VBZ 32505 237 8 laterally laterally RB 32505 237 9 convex convex NNP 32505 237 10 and and CC 32505 237 11 less less JJR 32505 237 12 than than IN 32505 237 13 three three CD 32505 237 14 times time NNS 32505 237 15 the the DT 32505 237 16 height height NN 32505 237 17 of of IN 32505 237 18 the the DT 32505 237 19 pars par NNS 32505 237 20 dentalis dentalis NNP 32505 237 21 . . . 32505 238 1 The the DT 32505 238 2 nasal nasal NN 32505 238 3 is be VBZ 32505 238 4 large large JJ 32505 238 5 and and CC 32505 238 6 pointed point VBN 32505 238 7 anteriorly anteriorly RB 32505 238 8 and and CC 32505 238 9 posteriorly posteriorly RB 32505 238 10 in in IN 32505 238 11 dorsal dorsal NN 32505 238 12 view view NN 32505 238 13 . . . 32505 239 1 The the DT 32505 239 2 length length NN 32505 239 3 of of IN 32505 239 4 the the DT 32505 239 5 nasal nasal NN 32505 239 6 comprises comprise VBZ 32505 239 7 about about IN 32505 239 8 40 40 CD 32505 239 9 per per NN 32505 239 10 cent cent NN 32505 239 11 of of IN 32505 239 12 the the DT 32505 239 13 total total JJ 32505 239 14 length length NN 32505 239 15 of of IN 32505 239 16 the the DT 32505 239 17 skull skull NN 32505 239 18 . . . 32505 240 1 The the DT 32505 240 2 nasals nasal NNS 32505 240 3 are be VBP 32505 240 4 separated separate VBN 32505 240 5 anteromedially anteromedially RB 32505 240 6 by by IN 32505 240 7 the the DT 32505 240 8 cartilaginous cartilaginous JJ 32505 240 9 septum septum NN 32505 240 10 nasi nasi NNP 32505 240 11 . . . 32505 241 1 One one CD 32505 241 2 protuberance protuberance NN 32505 241 3 is be VBZ 32505 241 4 present present JJ 32505 241 5 on on IN 32505 241 6 the the DT 32505 241 7 midlateral midlateral JJ 32505 241 8 concavity concavity NN 32505 241 9 of of IN 32505 241 10 the the DT 32505 241 11 nasal nasal NN 32505 241 12 . . . 32505 242 1 Posteriorly posteriorly RB 32505 242 2 , , , 32505 242 3 the the DT 32505 242 4 nasal nasal NN 32505 242 5 overlaps overlap VBZ 32505 242 6 the the DT 32505 242 7 sphenethmoid sphenethmoid NN 32505 242 8 ; ; : 32505 242 9 posterolaterally posterolaterally RB 32505 242 10 the the DT 32505 242 11 nasal nasal NN 32505 242 12 articulates articulate VBZ 32505 242 13 with with IN 32505 242 14 the the DT 32505 242 15 palatine palatine NN 32505 242 16 . . . 32505 243 1 The the DT 32505 243 2 sphenethmoid sphenethmoid NN 32505 243 3 is be VBZ 32505 243 4 completely completely RB 32505 243 5 ossified ossified JJ 32505 243 6 and and CC 32505 243 7 pentagonal pentagonal JJ 32505 243 8 in in IN 32505 243 9 dorsal dorsal NN 32505 243 10 view view NN 32505 243 11 . . . 32505 244 1 The the DT 32505 244 2 frontoparietal frontoparietal NN 32505 244 3 is be VBZ 32505 244 4 elongate elongate NN 32505 244 5 , , , 32505 244 6 without without IN 32505 244 7 a a DT 32505 244 8 pronounced pronounce VBN 32505 244 9 anterior anterior JJ 32505 244 10 supraorbital supraorbital NN 32505 244 11 process process NN 32505 244 12 . . . 32505 245 1 The the DT 32505 245 2 frontoparientals frontopariental NNS 32505 245 3 are be VBP 32505 245 4 sutured suture VBN 32505 245 5 medially medially RB 32505 245 6 throughout throughout IN 32505 245 7 their -PRON- PRP$ 32505 245 8 lengths length NNS 32505 245 9 ; ; : 32505 245 10 the the DT 32505 245 11 frontoparietal frontoparietal JJ 32505 245 12 fontanelle fontanelle NN 32505 245 13 is be VBZ 32505 245 14 absent absent JJ 32505 245 15 . . . 32505 246 1 The the DT 32505 246 2 bony bony NN 32505 246 3 part part NN 32505 246 4 of of IN 32505 246 5 the the DT 32505 246 6 proötic proötic NN 32505 246 7 is be VBZ 32505 246 8 narrowly narrowly RB 32505 246 9 separated separate VBN 32505 246 10 dorsolaterally dorsolaterally RB 32505 246 11 from from IN 32505 246 12 the the DT 32505 246 13 squamosal squamosal NN 32505 246 14 by by IN 32505 246 15 the the DT 32505 246 16 cartilaginous cartilaginous JJ 32505 246 17 crista crista NNP 32505 246 18 parotica parotica NNP 32505 246 19 . . . 32505 247 1 The the DT 32505 247 2 squamosal squamosal NN 32505 247 3 is be VBZ 32505 247 4 large large JJ 32505 247 5 ; ; : 32505 247 6 the the DT 32505 247 7 anterior anterior JJ 32505 247 8 arm arm NN 32505 247 9 is be VBZ 32505 247 10 pointed point VBN 32505 247 11 . . . 32505 248 1 The the DT 32505 248 2 posterior posterior JJ 32505 248 3 arm arm NN 32505 248 4 of of IN 32505 248 5 the the DT 32505 248 6 squamosal squamosal NN 32505 248 7 is be VBZ 32505 248 8 broad broad JJ 32505 248 9 , , , 32505 248 10 rounded round VBN 32505 248 11 terminally terminally RB 32505 248 12 , , , 32505 248 13 and and CC 32505 248 14 articulates articulate VBZ 32505 248 15 with with IN 32505 248 16 the the DT 32505 248 17 proötic proötic NN 32505 248 18 medially medially RB 32505 248 19 . . . 32505 249 1 The the DT 32505 249 2 prevomer prevomer NN 32505 249 3 is be VBZ 32505 249 4 short short JJ 32505 249 5 and and CC 32505 249 6 separated separate VBN 32505 249 7 anteriorly anteriorly RB 32505 249 8 from from IN 32505 249 9 the the DT 32505 249 10 premaxillary premaxillary NN 32505 249 11 and and CC 32505 249 12 maxillary maxillary NNP 32505 249 13 by by IN 32505 249 14 cartilage cartilage NN 32505 249 15 . . . 32505 250 1 The the DT 32505 250 2 posterior posterior JJ 32505 250 3 margin margin NN 32505 250 4 of of IN 32505 250 5 the the DT 32505 250 6 prevomer prevomer NN 32505 250 7 has have VBZ 32505 250 8 a a DT 32505 250 9 bony bony NN 32505 250 10 articulation articulation NN 32505 250 11 with with IN 32505 250 12 the the DT 32505 250 13 sphenethmoid sphenethmoid NN 32505 250 14 . . . 32505 251 1 Each each DT 32505 251 2 prevomer prevomer NN 32505 251 3 bears bear VBZ 32505 251 4 five five CD 32505 251 5 to to TO 32505 251 6 seven seven CD 32505 251 7 teeth tooth NNS 32505 251 8 . . . 32505 252 1 The the DT 32505 252 2 palatine palatine NN 32505 252 3 is be VBZ 32505 252 4 small small JJ 32505 252 5 and and CC 32505 252 6 edentate edentate JJ 32505 252 7 . . . 32505 253 1 The the DT 32505 253 2 anterior anterior JJ 32505 253 3 end end NN 32505 253 4 of of IN 32505 253 5 the the DT 32505 253 6 parasphenoid parasphenoid NN 32505 253 7 is be VBZ 32505 253 8 narrow narrow JJ 32505 253 9 ( ( -LRB- 32505 253 10 more more RBR 32505 253 11 pointed pointed JJ 32505 253 12 than than IN 32505 253 13 in in IN 32505 253 14 _ _ NNP 32505 253 15 Hyla Hyla NNP 32505 253 16 boulengeri boulengeri NN 32505 253 17 _ _ NNP 32505 253 18 ) ) -RRB- 32505 253 19 . . . 32505 254 1 The the DT 32505 254 2 pterygoid pterygoid NN 32505 254 3 is be VBZ 32505 254 4 slender slend JJR 32505 254 5 and and CC 32505 254 6 well well RB 32505 254 7 developed developed JJ 32505 254 8 ( ( -LRB- 32505 254 9 Fig fig NN 32505 254 10 . . . 32505 255 1 5A 5a RB 32505 255 2 ) ) -RRB- 32505 255 3 . . . 32505 256 1 [ [ -LRB- 32505 256 2 Illustration illustration NN 32505 256 3 : : : 32505 256 4 Fig Fig NNP 32505 256 5 . . . 32505 257 1 5 5 CD 32505 257 2 . . . 32505 258 1 Dorsal dorsal NN 32505 258 2 views view NNS 32505 258 3 of of IN 32505 258 4 the the DT 32505 258 5 skulls skull NNS 32505 258 6 of of IN 32505 258 7 ( ( -LRB- 32505 258 8 A a NN 32505 258 9 ) ) -RRB- 32505 258 10 _ _ NNP 32505 258 11 Hyla Hyla NNP 32505 258 12 foliamorta foliamorta NN 32505 258 13 _ _ NNP 32505 258 14 ( ( -LRB- 32505 258 15 KU KU NNP 32505 258 16 77687 77687 CD 32505 258 17 ) ) -RRB- 32505 258 18 and and CC 32505 258 19 ( ( -LRB- 32505 258 20 B b NN 32505 258 21 ) ) -RRB- 32505 258 22 _ _ NNP 32505 258 23 H. H. NNP 32505 259 1 elaeochroa elaeochroa NNP 32505 259 2 _ _ NNP 32505 259 3 ( ( -LRB- 32505 259 4 KU KU NNP 32505 259 5 68289 68289 CD 32505 259 6 ) ) -RRB- 32505 259 7 , , , 32505 259 8 × × NN 32505 259 9 3 3 CD 32505 259 10 . . . 32505 259 11 ] ] -RRB- 32505 260 1 _ _ NNP 32505 260 2 Natural Natural NNP 32505 260 3 History._--_Hyla History._--_Hyla NNP 32505 260 4 foliamorta foliamorta NNP 32505 260 5 _ _ NNP 32505 260 6 inhabits inhabit NNS 32505 260 7 lowland lowland VB 32505 260 8 forests forest NNS 32505 260 9 in in IN 32505 260 10 eastern eastern JJ 32505 260 11 Panamá Panamá NNP 32505 260 12 and and CC 32505 260 13 breeds breed NNS 32505 260 14 in in IN 32505 260 15 temporary temporary JJ 32505 260 16 ponds pond NNS 32505 260 17 . . . 32505 261 1 Males male NNS 32505 261 2 have have VBP 32505 261 3 been be VBN 32505 261 4 observed observe VBN 32505 261 5 calling call VBG 32505 261 6 from from IN 32505 261 7 grasses grass NNS 32505 261 8 , , , 32505 261 9 bushes bush NNS 32505 261 10 , , , 32505 261 11 and and CC 32505 261 12 emergent emergent JJ 32505 261 13 vegetation vegetation NN 32505 261 14 near near IN 32505 261 15 temporary temporary JJ 32505 261 16 ponds pond NNS 32505 261 17 and and CC 32505 261 18 ditches ditch NNS 32505 261 19 along along IN 32505 261 20 roads road NNS 32505 261 21 . . . 32505 262 1 William William NNP 32505 262 2 E. E. NNP 32505 262 3 Duellman Duellman NNP 32505 262 4 informed inform VBD 32505 262 5 me -PRON- PRP 32505 262 6 that that IN 32505 262 7 he -PRON- PRP 32505 262 8 found find VBD 32505 262 9 a a DT 32505 262 10 breeding breeding NN 32505 262 11 congregation congregation NN 32505 262 12 of of IN 32505 262 13 this this DT 32505 262 14 species specie NNS 32505 262 15 in in IN 32505 262 16 June June NNP 32505 262 17 near near IN 32505 262 18 Chepo Chepo NNP 32505 262 19 , , , 32505 262 20 Panamá Panamá NNP 32505 262 21 , , , 32505 262 22 where where WRB 32505 262 23 males male NNS 32505 262 24 were be VBD 32505 262 25 calling call VBG 32505 262 26 from from IN 32505 262 27 spiny spiny JJ 32505 262 28 palms palm NNS 32505 262 29 at at IN 32505 262 30 the the DT 32505 262 31 edge edge NN 32505 262 32 of of IN 32505 262 33 a a DT 32505 262 34 woodland woodland JJ 32505 262 35 pond pond NN 32505 262 36 . . . 32505 263 1 Fouquette Fouquette NNP 32505 263 2 ( ( -LRB- 32505 263 3 1958 1958 CD 32505 263 4 ) ) -RRB- 32505 263 5 found find VBD 32505 263 6 calling call VBG 32505 263 7 males male NNS 32505 263 8 in in IN 32505 263 9 May May NNP 32505 263 10 , , , 32505 263 11 August August NNP 32505 263 12 , , , 32505 263 13 and and CC 32505 263 14 September September NNP 32505 263 15 near near IN 32505 263 16 Miraflores Miraflores NNP 32505 263 17 Locks Locks NNP 32505 263 18 , , , 32505 263 19 Canal Canal NNP 32505 263 20 Zone Zone NNP 32505 263 21 . . . 32505 264 1 Calling call VBG 32505 264 2 stations station NNS 32505 264 3 vary vary VBP 32505 264 4 from from IN 32505 264 5 one one CD 32505 264 6 to to IN 32505 264 7 two two CD 32505 264 8 meters meter NNS 32505 264 9 above above IN 32505 264 10 ground ground NN 32505 264 11 . . . 32505 265 1 No no DT 32505 265 2 clasping clasp VBG 32505 265 3 pairs pair NNS 32505 265 4 have have VBP 32505 265 5 been be VBN 32505 265 6 found find VBN 32505 265 7 ; ; : 32505 265 8 only only RB 32505 265 9 one one CD 32505 265 10 female female NN 32505 265 11 is be VBZ 32505 265 12 known know VBN 32505 265 13 ( ( -LRB- 32505 265 14 KU KU NNP 32505 265 15 101589 101589 CD 32505 265 16 , , , 32505 265 17 from from IN 32505 265 18 8 8 CD 32505 265 19 km km NN 32505 265 20 NE NE NNP 32505 265 21 Tocumen Tocumen NNP 32505 265 22 , , , 32505 265 23 Panamá Panamá NNP 32505 265 24 ) ) -RRB- 32505 265 25 ; ; : 32505 265 26 this this DT 32505 265 27 gravid gravid JJ 32505 265 28 individual individual NN 32505 265 29 was be VBD 32505 265 30 collected collect VBN 32505 265 31 in in IN 32505 265 32 early early JJ 32505 265 33 June June NNP 32505 265 34 . . . 32505 266 1 The the DT 32505 266 2 mating mating NN 32505 266 3 call call NN 32505 266 4 of of IN 32505 266 5 _ _ NNP 32505 266 6 Hyla Hyla NNP 32505 266 7 foliamorta foliamorta NN 32505 266 8 _ _ NNP 32505 266 9 consists consist VBZ 32505 266 10 of of IN 32505 266 11 one one CD 32505 266 12 pulsed pulse VBN 32505 266 13 , , , 32505 266 14 low low RB 32505 266 15 - - HYPH 32505 266 16 pitched pitched JJ 32505 266 17 , , , 32505 266 18 moderate moderate JJ 32505 266 19 trill trill NN 32505 266 20 of of IN 32505 266 21 about about RB 32505 266 22 O.5 o.5 JJ 32505 266 23 second second JJ 32505 266 24 duration duration NN 32505 266 25 . . . 32505 267 1 Each each DT 32505 267 2 note note NN 32505 267 3 is be VBZ 32505 267 4 repeated repeat VBN 32505 267 5 at at IN 32505 267 6 intervals interval NNS 32505 267 7 of of IN 32505 267 8 5 5 CD 32505 267 9 seconds second NNS 32505 267 10 to to IN 32505 267 11 a a DT 32505 267 12 few few JJ 32505 267 13 minutes minute NNS 32505 267 14 . . . 32505 268 1 The the DT 32505 268 2 notes note NNS 32505 268 3 have have VBP 32505 268 4 about about RB 32505 268 5 50 50 CD 32505 268 6 pulses pulse NNS 32505 268 7 per per IN 32505 268 8 second second NN 32505 268 9 , , , 32505 268 10 a a DT 32505 268 11 fundamental fundamental JJ 32505 268 12 frequency frequency NN 32505 268 13 of of IN 32505 268 14 56 56 CD 32505 268 15 cycles cycle NNS 32505 268 16 per per IN 32505 268 17 second second NN 32505 268 18 and and CC 32505 268 19 a a DT 32505 268 20 dominant dominant JJ 32505 268 21 frequency frequency NN 32505 268 22 of of IN 32505 268 23 about about RB 32505 268 24 3000 3000 CD 32505 268 25 cycles cycle NNS 32505 268 26 per per IN 32505 268 27 second second JJ 32505 268 28 ( ( -LRB- 32505 268 29 Table table NN 32505 268 30 2 2 CD 32505 268 31 , , , 32505 268 32 Pl Pl NNP 32505 268 33 . . . 32505 269 1 3B 3B NNP 32505 269 2 ) ) -RRB- 32505 269 3 . . . 32505 270 1 Egg egg NN 32505 270 2 deposition deposition NN 32505 270 3 sites site NNS 32505 270 4 are be VBP 32505 270 5 unknown unknown JJ 32505 270 6 . . . 32505 271 1 No no DT 32505 271 2 information information NN 32505 271 3 is be VBZ 32505 271 4 available available JJ 32505 271 5 concerning concern VBG 32505 271 6 early early JJ 32505 271 7 development development NN 32505 271 8 , , , 32505 271 9 and and CC 32505 271 10 little little JJ 32505 271 11 is be VBZ 32505 271 12 known know VBN 32505 271 13 about about IN 32505 271 14 the the DT 32505 271 15 breeding breeding NN 32505 271 16 season season NN 32505 271 17 of of IN 32505 271 18 _ _ NNP 32505 271 19 Hyla Hyla NNP 32505 271 20 foliamorta foliamorta NN 32505 271 21 _ _ NNP 32505 271 22 . . . 32505 272 1 Probably probably RB 32505 272 2 its -PRON- PRP$ 32505 272 3 breeding breeding NN 32505 272 4 activities activity NNS 32505 272 5 are be VBP 32505 272 6 restricted restrict VBN 32505 272 7 to to IN 32505 272 8 the the DT 32505 272 9 rainy rainy JJ 32505 272 10 months month NNS 32505 272 11 . . . 32505 273 1 _ _ NNP 32505 273 2 Tadpoles._--Eight tadpoles._--eight CD 32505 273 3 tadpoles tadpole NNS 32505 273 4 were be VBD 32505 273 5 collected collect VBN 32505 273 6 from from IN 32505 273 7 a a DT 32505 273 8 weedy weedy JJ 32505 273 9 temporary temporary JJ 32505 273 10 pond pond NN 32505 273 11 near near IN 32505 273 12 Chepo Chepo NNP 32505 273 13 , , , 32505 273 14 Panamá Panamá NNP 32505 273 15 , , , 32505 273 16 in in IN 32505 273 17 early early JJ 32505 273 18 June June NNP 32505 273 19 . . . 32505 274 1 A a DT 32505 274 2 typical typical JJ 32505 274 3 tadpole tadpole NN 32505 274 4 in in IN 32505 274 5 stage stage NN 32505 274 6 35 35 CD 32505 274 7 of of IN 32505 274 8 development development NN 32505 274 9 ( ( -LRB- 32505 274 10 KU KU NNP 32505 274 11 104244 104244 CD 32505 274 12 ) ) -RRB- 32505 274 13 has have VBZ 32505 274 14 a a DT 32505 274 15 body body NN 32505 274 16 length length NN 32505 274 17 of of IN 32505 274 18 9.5 9.5 CD 32505 274 19 mm mm NNP 32505 274 20 . . NNP 32505 274 21 , , , 32505 274 22 tail tail NN 32505 274 23 length length NN 32505 274 24 of of IN 32505 274 25 25.0 25.0 CD 32505 274 26 mm mm NNP 32505 274 27 . . NNP 32505 274 28 , , , 32505 274 29 and and CC 32505 274 30 a a DT 32505 274 31 total total JJ 32505 274 32 length length NN 32505 274 33 of of IN 32505 274 34 34.5 34.5 CD 32505 274 35 mm mm NN 32505 274 36 . . . 32505 275 1 Other other JJ 32505 275 2 characters character NNS 32505 275 3 are be VBP 32505 275 4 as as IN 32505 275 5 follows follow VBZ 32505 275 6 : : : 32505 275 7 depth depth NN 32505 275 8 of of IN 32505 275 9 tail tail NN 32505 275 10 equal equal JJ 32505 275 11 to to IN 32505 275 12 length length NN 32505 275 13 of of IN 32505 275 14 body body NN 32505 275 15 ; ; : 32505 275 16 body body NN 32505 275 17 deeper deeply RBR 32505 275 18 than than IN 32505 275 19 wide wide JJ 32505 275 20 ; ; : 32505 275 21 distance distance NN 32505 275 22 between between IN 32505 275 23 eye eye NN 32505 275 24 and and CC 32505 275 25 nostril nostril NNP 32505 275 26 equal equal JJ 32505 275 27 to to TO 32505 275 28 distance distance NN 32505 275 29 between between IN 32505 275 30 eye eye NN 32505 275 31 and and CC 32505 275 32 spiracle spiracle NNP 32505 275 33 ; ; : 32505 275 34 mouth mouth NNP 32505 275 35 anteroventral anteroventral NNP 32505 275 36 ; ; : 32505 275 37 median median JJ 32505 275 38 part part NN 32505 275 39 of of IN 32505 275 40 upper upper NNP 32505 275 41 lip lip NNP 32505 275 42 bare bare NNP 32505 275 43 ; ; : 32505 275 44 rest rest NN 32505 275 45 of of IN 32505 275 46 lip lip NN 32505 275 47 having have VBG 32505 275 48 one one CD 32505 275 49 row row NN 32505 275 50 of of IN 32505 275 51 papillae papillae NNP 32505 275 52 ; ; : 32505 275 53 a a DT 32505 275 54 few few JJ 32505 275 55 other other JJ 32505 275 56 rows row NNS 32505 275 57 of of IN 32505 275 58 small small JJ 32505 275 59 papillae papilla NNS 32505 275 60 at at IN 32505 275 61 corners corner NNS 32505 275 62 of of IN 32505 275 63 mouth mouth NN 32505 275 64 ; ; , 32505 275 65 tooth tooth NN 32505 275 66 rows row VBZ 32505 275 67 2/3 2/3 CD 32505 275 68 ; ; : 32505 275 69 first first RB 32505 275 70 upper upper JJ 32505 275 71 row row NN 32505 275 72 entire entire JJ 32505 275 73 , , , 32505 275 74 second second JJ 32505 275 75 upper upper JJ 32505 275 76 row row NN 32505 275 77 interrupted interrupt VBN 32505 275 78 medially medially RB 32505 275 79 , , , 32505 275 80 shorter short JJR 32505 275 81 than than IN 32505 275 82 first first RB 32505 275 83 ; ; : 32505 275 84 lower low JJR 32505 275 85 rows row NNS 32505 275 86 shorter short JJR 32505 275 87 than than IN 32505 275 88 upper upper JJ 32505 275 89 rows row NNS 32505 275 90 , , , 32505 275 91 third third RB 32505 275 92 shortest short JJS 32505 275 93 ; ; : 32505 275 94 beak beak VB 32505 275 95 moderately moderately RB 32505 275 96 robust robust JJ 32505 275 97 ; ; : 32505 275 98 spiracle spiracle NNP 32505 275 99 nearer nearer NNP 32505 275 100 eye eye NN 32505 275 101 than than IN 32505 275 102 anus anus NN 32505 275 103 ; ; : 32505 275 104 anal anal JJ 32505 275 105 tube tube NN 32505 275 106 short short JJ 32505 275 107 , , , 32505 275 108 aperture aperture JJ 32505 275 109 not not RB 32505 275 110 extending extend VBG 32505 275 111 to to IN 32505 275 112 border border NN 32505 275 113 of of IN 32505 275 114 ventral ventral JJ 32505 275 115 fin fin NN 32505 275 116 ; ; , 32505 275 117 caudal caudal NN 32505 275 118 musculature musculature NN 32505 275 119 slender slender NN 32505 275 120 , , , 32505 275 121 extending extend VBG 32505 275 122 to to IN 32505 275 123 tip tip NN 32505 275 124 of of IN 32505 275 125 pointed pointed JJ 32505 275 126 tail tail NN 32505 275 127 ; ; : 32505 275 128 dorsal dorsal NN 32505 275 129 fin fin NN 32505 275 130 extending extend VBG 32505 275 131 onto onto IN 32505 275 132 body body NN 32505 275 133 ( ( -LRB- 32505 275 134 Table table NN 32505 275 135 4 4 CD 32505 275 136 ) ) -RRB- 32505 275 137 . . . 32505 276 1 TABLE TABLE NNP 32505 276 2 4.--Sizes 4.--sizes CD 32505 276 3 of of IN 32505 276 4 Tadpoles Tadpoles NNP 32505 276 5 of of IN 32505 276 6 _ _ NNP 32505 276 7 Hyla Hyla NNP 32505 276 8 foliamorta foliamorta NN 32505 276 9 _ _ NNP 32505 276 10 in in IN 32505 276 11 Relation Relation NNP 32505 276 12 to to IN 32505 276 13 Developmental Developmental NNP 32505 276 14 Stages Stages NNPS 32505 276 15 . . . 32505 277 1 ( ( -LRB- 32505 277 2 Means mean NNS 32505 277 3 in in IN 32505 277 4 parentheses parenthesis NNS 32505 277 5 below below IN 32505 277 6 observed observed JJ 32505 277 7 ranges range NNS 32505 277 8 ; ; : 32505 277 9 measurements measurement NNS 32505 277 10 in in IN 32505 277 11 mm mm NNP 32505 277 12 . . . 32505 277 13 ) ) -RRB- 32505 278 1 = = NFP 32505 278 2 = = NFP 32505 278 3 = = SYM 32505 278 4 = = SYM 32505 278 5 = = SYM 32505 278 6 = = SYM 32505 278 7 = = SYM 32505 278 8 = = SYM 32505 278 9 = = SYM 32505 278 10 = = SYM 32505 278 11 = = SYM 32505 278 12 = = SYM 32505 278 13 = = SYM 32505 278 14 = = SYM 32505 278 15 = = SYM 32505 278 16 = = SYM 32505 278 17 = = SYM 32505 278 18 = = SYM 32505 278 19 = = SYM 32505 278 20 = = SYM 32505 278 21 = = SYM 32505 278 22 = = SYM 32505 278 23 = = SYM 32505 278 24 = = SYM 32505 278 25 = = SYM 32505 278 26 = = SYM 32505 278 27 = = SYM 32505 278 28 = = SYM 32505 278 29 = = SYM 32505 278 30 = = SYM 32505 278 31 = = SYM 32505 278 32 = = SYM 32505 278 33 = = SYM 32505 278 34 = = SYM 32505 278 35 = = SYM 32505 278 36 = = SYM 32505 278 37 = = SYM 32505 278 38 = = SYM 32505 278 39 = = SYM 32505 278 40 = = SYM 32505 278 41 = = SYM 32505 278 42 = = SYM 32505 278 43 = = SYM 32505 278 44 = = SYM 32505 278 45 = = SYM 32505 278 46 = = SYM 32505 278 47 = = SYM 32505 278 48 = = SYM 32505 278 49 = = SYM 32505 278 50 = = SYM 32505 278 51 = = SYM 32505 278 52 = = SYM 32505 278 53 = = SYM 32505 278 54 = = NFP 32505 278 55 = = NFP 32505 278 56 Stage stage NN 32505 278 57 | | NNP 32505 278 58 N N NNP 32505 278 59 | | NNP 32505 278 60 Body Body NNP 32505 278 61 length length NN 32505 278 62 | | NNP 32505 278 63 Tail Tail NNP 32505 278 64 length length NN 32505 278 65 | | NNP 32505 278 66 Total total JJ 32505 278 67 length length NN 32505 278 68 --------+---+-------------+-------------+-------------- --------+---+-------------+-------------+-------------- NN 32505 278 69 25 25 CD 32505 278 70 | | CD 32505 278 71 2 2 CD 32505 278 72 | | CD 32505 278 73 5.0 5.0 CD 32505 278 74 - - HYPH 32505 278 75 5.2 5.2 CD 32505 278 76 | | CD 32505 278 77 8.0 8.0 CD 32505 278 78 - - HYPH 32505 278 79 8.5 8.5 CD 32505 278 80 | | CD 32505 278 81 13.0 13.0 CD 32505 278 82 - - HYPH 32505 278 83 13.7 13.7 CD 32505 278 84 | | CD 32505 278 85 | | CD 32505 278 86 ( ( -LRB- 32505 278 87 5.1 5.1 CD 32505 278 88 ) ) -RRB- 32505 278 89 | | NNP 32505 278 90 ( ( -LRB- 32505 278 91 8.3 8.3 CD 32505 278 92 ) ) -RRB- 32505 278 93 | | NNP 32505 278 94 ( ( -LRB- 32505 278 95 13.4 13.4 CD 32505 278 96 ) ) -RRB- 32505 278 97 | | CD 32505 278 98 | | CD 32505 278 99 | | CD 32505 278 100 | | CD 32505 278 101 26 26 CD 32505 278 102 | | CD 32505 278 103 3 3 CD 32505 278 104 | | CD 32505 278 105 7.0 7.0 CD 32505 278 106 - - HYPH 32505 278 107 7.5 7.5 CD 32505 278 108 | | CD 32505 278 109 12.0 12.0 CD 32505 278 110 - - HYPH 32505 278 111 12.4 12.4 CD 32505 278 112 | | CD 32505 278 113 17.0 17.0 CD 32505 278 114 - - HYPH 32505 278 115 19.5 19.5 CD 32505 278 116 | | CD 32505 278 117 | | CD 32505 278 118 ( ( -LRB- 32505 278 119 7.2 7.2 CD 32505 278 120 ) ) -RRB- 32505 278 121 | | NNP 32505 278 122 ( ( -LRB- 32505 278 123 12.1 12.1 CD 32505 278 124 ) ) -RRB- 32505 278 125 | | NNP 32505 278 126 ( ( -LRB- 32505 278 127 18.6 18.6 CD 32505 278 128 ) ) -RRB- 32505 278 129 | | NNP 32505 278 130 | | CD 32505 278 131 | | CD 32505 278 132 | | CD 32505 278 133 28 28 CD 32505 278 134 | | CD 32505 278 135 2 2 CD 32505 278 136 | | CD 32505 278 137 6.5 6.5 CD 32505 278 138 - - HYPH 32505 278 139 7.0 7.0 CD 32505 278 140 | | CD 32505 278 141 18.0 18.0 CD 32505 278 142 | | CD 32505 278 143 25.0 25.0 CD 32505 278 144 | | CD 32505 278 145 | | CD 32505 278 146 ( ( -LRB- 32505 278 147 6.8 6.8 CD 32505 278 148 ) ) -RRB- 32505 278 149 | | CD 32505 278 150 | | CD 32505 278 151 | | CD 32505 278 152 | | CD 32505 278 153 | | CD 32505 278 154 | | CD 32505 278 155 35 35 CD 32505 278 156 | | CD 32505 278 157 1 1 CD 32505 278 158 | | CD 32505 278 159 9.5 9.5 CD 32505 278 160 | | CD 32505 278 161 25.0 25.0 CD 32505 278 162 | | CD 32505 278 163 34.5 34.5 CD 32505 278 164 In in IN 32505 278 165 life life NN 32505 278 166 , , , 32505 278 167 yellow yellow NN 32505 278 168 above above RB 32505 278 169 , , , 32505 278 170 white white JJ 32505 278 171 below below RB 32505 278 172 ; ; : 32505 278 173 caudal caudal JJ 32505 278 174 fin fin NNP 32505 278 175 greenish greenish NNP 32505 278 176 yellow yellow NNP 32505 278 177 with with IN 32505 278 178 black black JJ 32505 278 179 or or CC 32505 278 180 gray gray JJ 32505 278 181 reticulations reticulation NNS 32505 278 182 ; ; , 32505 278 183 dark dark JJ 32505 278 184 line line NN 32505 278 185 from from IN 32505 278 186 snout snout NN 32505 278 187 to to IN 32505 278 188 eye eye NN 32505 278 189 ; ; : 32505 278 190 dark dark JJ 32505 278 191 spot spot NN 32505 278 192 behind behind IN 32505 278 193 eye eye NN 32505 278 194 ; ; : 32505 278 195 tail tail NN 32505 278 196 unpigmented unpigmente VBN 32505 278 197 except except IN 32505 278 198 for for IN 32505 278 199 fine fine JJ 32505 278 200 dark dark JJ 32505 278 201 reticulations reticulation NNS 32505 278 202 . . . 32505 279 1 In in IN 32505 279 2 preservative preservative JJ 32505 279 3 body body NN 32505 279 4 creamy creamy NN 32505 279 5 white white JJ 32505 279 6 , , , 32505 279 7 transparent transparent JJ 32505 279 8 below below IN 32505 279 9 with with IN 32505 279 10 dark dark JJ 32505 279 11 pigment pigment NN 32505 279 12 above above IN 32505 279 13 in in IN 32505 279 14 some some DT 32505 279 15 specimens specimen NNS 32505 279 16 . . . 32505 280 1 _ _ NNP 32505 280 2 Remarks._--_Hyla Remarks._--_Hyla NNP 32505 280 3 foliamorta foliamorta NNP 32505 280 4 _ _ NNP 32505 280 5 can can MD 32505 280 6 be be VB 32505 280 7 confused confuse VBN 32505 280 8 only only RB 32505 280 9 with with IN 32505 280 10 _ _ NNP 32505 280 11 Hyla Hyla NNP 32505 280 12 boulengeri boulengeri NN 32505 280 13 _ _ NNP 32505 280 14 . . . 32505 281 1 The the DT 32505 281 2 differences difference NNS 32505 281 3 between between IN 32505 281 4 adults adult NNS 32505 281 5 of of IN 32505 281 6 these these DT 32505 281 7 species specie NNS 32505 281 8 were be VBD 32505 281 9 discussed discuss VBN 32505 281 10 in in IN 32505 281 11 _ _ NNP 32505 281 12 Remarks Remarks NNP 32505 281 13 _ _ NNP 32505 281 14 on on IN 32505 281 15 _ _ NNP 32505 281 16 H. H. NNP 32505 282 1 boulengeri boulengeri NNP 32505 282 2 _ _ NNP 32505 282 3 . . . 32505 283 1 The the DT 32505 283 2 tadpoles tadpole NNS 32505 283 3 of of IN 32505 283 4 _ _ NNP 32505 283 5 foliamorta foliamorta NNP 32505 283 6 _ _ NNP 32505 283 7 have have VBP 32505 283 8 labial labial JJ 32505 283 9 papillae papilla NNS 32505 283 10 on on IN 32505 283 11 the the DT 32505 283 12 lower low JJR 32505 283 13 lip lip NN 32505 283 14 and and CC 32505 283 15 a a DT 32505 283 16 stripe stripe NN 32505 283 17 between between IN 32505 283 18 the the DT 32505 283 19 eye eye NN 32505 283 20 and and CC 32505 283 21 the the DT 32505 283 22 tip tip NN 32505 283 23 of of IN 32505 283 24 the the DT 32505 283 25 snout snout NN 32505 283 26 . . . 32505 284 1 By by IN 32505 284 2 comparison comparison NN 32505 284 3 the the DT 32505 284 4 tadpoles tadpole NNS 32505 284 5 of of IN 32505 284 6 _ _ NNP 32505 284 7 boulengeri boulengeri NN 32505 284 8 _ _ NNP 32505 284 9 have have VBP 32505 284 10 a a DT 32505 284 11 bare bare JJ 32505 284 12 lower low JJR 32505 284 13 lip lip NN 32505 284 14 and and CC 32505 284 15 no no DT 32505 284 16 stripe stripe NN 32505 284 17 between between IN 32505 284 18 the the DT 32505 284 19 eye eye NN 32505 284 20 and and CC 32505 284 21 the the DT 32505 284 22 tip tip NN 32505 284 23 of of IN 32505 284 24 the the DT 32505 284 25 snout snout NN 32505 284 26 . . . 32505 285 1 _ _ NNP 32505 285 2 Distribution._--_Hyla Distribution._--_Hyla NNP 32505 285 3 foliamorta foliamorta NNP 32505 285 4 _ _ NNP 32505 285 5 inhabits inhabit VBZ 32505 285 6 the the DT 32505 285 7 subhumid subhumid NN 32505 285 8 Pacific Pacific NNP 32505 285 9 lowlands lowland NNS 32505 285 10 ( ( -LRB- 32505 285 11 elevations elevation NNS 32505 285 12 of of IN 32505 285 13 less less JJR 32505 285 14 than than IN 32505 285 15 100 100 CD 32505 285 16 meters meter NNS 32505 285 17 ) ) -RRB- 32505 285 18 of of IN 32505 285 19 Central Central NNP 32505 285 20 Panamá Panamá NNP 32505 285 21 and and CC 32505 285 22 Caribbean caribbean JJ 32505 285 23 lowlands lowland NNS 32505 285 24 of of IN 32505 285 25 northern northern JJ 32505 285 26 Colombia Colombia NNP 32505 285 27 ( ( -LRB- 32505 285 28 Fig Fig NNP 32505 285 29 . . . 32505 286 1 4 4 LS 32505 286 2 ) ) -RRB- 32505 286 3 . . . 32505 287 1 _ _ NNP 32505 287 2 Specimens Specimens NNP 32505 287 3 Examined._--Panamá Examined._--Panamá NNPS 32505 287 4 : : : 32505 287 5 _ _ NNP 32505 287 6 Panamá Panamá NNP 32505 287 7 _ _ NNP 32505 287 8 : : : 32505 287 9 3 3 CD 32505 287 10 km km NN 32505 287 11 WSW WSW NNP 32505 287 12 Chepo Chepo NNP 32505 287 13 , , , 32505 287 14 KU KU NNP 32505 287 15 77164 77164 CD 32505 287 16 - - SYM 32505 287 17 9 9 CD 32505 287 18 , , , 32505 287 19 101573 101573 CD 32505 287 20 - - SYM 32505 287 21 5 5 CD 32505 287 22 , , , 32505 287 23 104243 104243 CD 32505 287 24 - - SYM 32505 287 25 4 4 CD 32505 287 26 ( ( -LRB- 32505 287 27 tadpoles tadpole NNS 32505 287 28 ) ) -RRB- 32505 287 29 ; ; : 32505 287 30 6 6 CD 32505 287 31 km km NN 32505 287 32 WSW WSW NNP 32505 287 33 Chepo Chepo NNP 32505 287 34 , , , 32505 287 35 KU KU NNP 32505 287 36 77170 77170 CD 32505 287 37 , , , 32505 287 38 101576 101576 CD 32505 287 39 - - SYM 32505 287 40 8 8 CD 32505 287 41 ; ; : 32505 287 42 1.5 1.5 CD 32505 287 43 km km NN 32505 287 44 SW SW NNP 32505 287 45 Naranjal Naranjal NNP 32505 287 46 , , , 32505 287 47 KU KU NNP 32505 287 48 77171 77171 CD 32505 287 49 , , , 32505 287 50 77687 77687 CD 32505 287 51 ( ( -LRB- 32505 287 52 skeleton skeleton NNP 32505 287 53 ) ) -RRB- 32505 287 54 ; ; : 32505 287 55 2 2 LS 32505 287 56 km km NN 32505 287 57 N N NNP 32505 287 58 Tocumen Tocumen NNP 32505 287 59 , , , 32505 287 60 KU KU NNP 32505 287 61 101579 101579 CD 32505 287 62 - - SYM 32505 287 63 83 83 CD 32505 287 64 , , , 32505 287 65 104349 104349 CD 32505 287 66 ( ( -LRB- 32505 287 67 skeleton skeleton NNP 32505 287 68 ) ) -RRB- 32505 287 69 ; ; : 32505 287 70 8 8 CD 32505 287 71 km km NN 32505 287 72 NE NE NNP 32505 287 73 Tocumen Tocumen NNP 32505 287 74 , , , 32505 287 75 KU KU NNP 32505 287 76 101584 101584 CD 32505 287 77 - - SYM 32505 287 78 92 92 CD 32505 287 79 . . . 32505 288 1 No no DT 32505 288 2 specific specific JJ 32505 288 3 locality locality NN 32505 288 4 : : : 32505 288 5 TNHC TNHC NNP 32505 288 6 24401 24401 CD 32505 288 7 . . . 32505 289 1 _ _ NNP 32505 289 2 Hyla Hyla NNP 32505 289 3 rubra rubra NN 32505 289 4 _ _ NNP 32505 289 5 Laurenti Laurenti NNP 32505 289 6 _ _ NNP 32505 289 7 Hyla Hyla NNP 32505 289 8 rubra rubra NN 32505 289 9 _ _ NNP 32505 289 10 Laurenti Laurenti NNP 32505 289 11 , , , 32505 289 12 Synopsis Synopsis NNP 32505 289 13 Reptilium Reptilium NNP 32505 289 14 Emendatum Emendatum NNP 32505 289 15 , , , 32505 289 16 p. p. NN 32505 289 17 35 35 CD 32505 289 18 , , , 32505 289 19 1768 1768 CD 32505 289 20 . . . 32505 290 1 Daudin Daudin NNP 32505 290 2 , , , 32505 290 3 Hist Hist NNP 32505 290 4 . . . 32505 291 1 Nat Nat NNP 32505 291 2 . . . 32505 292 1 Rainettes Rainettes NNP 32505 292 2 Grenouilles Grenouilles NNP 32505 292 3 Crapauds Crapauds NNP 32505 292 4 , , , 32505 292 5 II:26 II:26 NNP 32505 292 6 , , , 32505 292 7 1802 1802 CD 32505 292 8 . . . 32505 293 1 Daudin Daudin NNP 32505 293 2 , , , 32505 293 3 Hist Hist NNP 32505 293 4 . . . 32505 294 1 Nat Nat NNP 32505 294 2 . . . 32505 295 1 Particuliere Particuliere NNP 32505 295 2 Reptiles Reptiles NNP 32505 295 3 , , , 32505 295 4 8:53 8:53 CD 32505 295 5 , , , 32505 295 6 1803 1803 CD 32505 295 7 . . . 32505 296 1 Günther Günther NNP 32505 296 2 , , , 32505 296 3 Catalogue Catalogue NNP 32505 296 4 Batrachia Batrachia NNP 32505 296 5 Salientia Salientia NNP 32505 296 6 Brit Brit NNP 32505 296 7 . . . 32505 297 1 Mus Mus NNP 32505 297 2 . . NNP 32505 297 3 , , , 32505 297 4 p. p. NN 32505 297 5 110 110 CD 32505 297 6 , , , 32505 297 7 1859 1859 CD 32505 297 8 . . . 32505 298 1 Boulenger Boulenger NNP 32505 298 2 , , , 32505 298 3 Catalogue Catalogue NNP 32505 298 4 Batrachia Batrachia NNP 32505 298 5 Salientia Salientia NNP 32505 298 6 s. s. NNP 32505 298 7 Ecaudata Ecaudata NNP 32505 298 8 , , , 32505 298 9 p. p. NN 32505 298 10 403 403 CD 32505 298 11 , , , 32505 298 12 February February NNP 32505 298 13 1 1 CD 32505 298 14 , , , 32505 298 15 1882 1882 CD 32505 298 16 . . . 32505 299 1 Dunn Dunn NNP 32505 299 2 , , , 32505 299 3 Occas Occas NNP 32505 299 4 . . . 32505 300 1 Papers Papers NNP 32505 300 2 , , , 32505 300 3 Boston Boston NNP 32505 300 4 Soc Soc NNP 32505 300 5 . . . 32505 301 1 Nat Nat NNP 32505 301 2 . . . 32505 302 1 Hist Hist NNP 32505 302 2 . . . 32505 302 3 , , , 32505 302 4 5:413 5:413 CD 32505 302 5 , , , 32505 302 6 October October NNP 32505 302 7 10 10 CD 32505 302 8 , , , 32505 302 9 1931 1931 CD 32505 302 10 . . . 32505 303 1 _ _ NNP 32505 303 2 Hyla Hyla NNP 32505 303 3 elaeochroa elaeochroa NN 32505 303 4 _ _ NNP 32505 303 5 ( ( -LRB- 32505 303 6 part part NN 32505 303 7 ) ) -RRB- 32505 303 8 : : : 32505 303 9 Dunn Dunn NNP 32505 303 10 and and CC 32505 303 11 Emlen Emlen NNP 32505 303 12 , , , 32505 303 13 Proc Proc NNP 32505 303 14 . . . 32505 304 1 Acad Acad NNS 32505 304 2 . . . 32505 305 1 Nat Nat NNP 32505 305 2 . . . 32505 306 1 Sci Sci NNP 32505 306 2 . . . 32505 307 1 Philadelphia Philadelphia NNP 32505 307 2 , , , 32505 307 3 84:25 84:25 CD 32505 307 4 , , , 32505 307 5 March March NNP 32505 307 6 22 22 CD 32505 307 7 , , , 32505 307 8 1932 1932 CD 32505 307 9 . . . 32505 308 1 _ _ NNP 32505 308 2 Diagnosis._--Size Diagnosis._--Size NNP 32505 308 3 medium medium NN 32505 308 4 ; ; : 32505 308 5 skull skull NNP 32505 308 6 longer long RBR 32505 308 7 than than IN 32505 308 8 wide wide JJ 32505 308 9 ; ; : 32505 308 10 frontoparietal frontoparietal JJ 32505 308 11 fontanelle fontanelle NN 32505 308 12 absent absent NN 32505 308 13 in in IN 32505 308 14 adults adult NNS 32505 308 15 ; ; : 32505 308 16 snout snout NNP 32505 308 17 subovoid subovoid NNP 32505 308 18 ; ; : 32505 308 19 choanae choanae NNP 32505 308 20 rounded round VBD 32505 308 21 ; ; : 32505 308 22 dorsal dorsal NN 32505 308 23 stripes stripe NNS 32505 308 24 present present JJ 32505 308 25 ; ; : 32505 308 26 black black JJ 32505 308 27 vermiculations vermiculation NNS 32505 308 28 on on IN 32505 308 29 posterior posterior NN 32505 308 30 surfaces surface NNS 32505 308 31 of of IN 32505 308 32 thighs thigh NNS 32505 308 33 . . . 32505 309 1 _ _ NNP 32505 309 2 Description._--Head Description._--Head NNP 32505 309 3 flat flat RB 32505 309 4 , , , 32505 309 5 longer longer RB 32505 309 6 than than IN 32505 309 7 wide wide JJ 32505 309 8 ; ; : 32505 309 9 snout snout NN 32505 309 10 long long RB 32505 309 11 , , , 32505 309 12 subovoid subovoid JJ 32505 309 13 , , , 32505 309 14 slightly slightly RB 32505 309 15 protruding protrude VBG 32505 309 16 beyond beyond IN 32505 309 17 lower low JJR 32505 309 18 lip lip NN 32505 309 19 ; ; : 32505 309 20 loreal loreal NN 32505 309 21 oblique oblique NNP 32505 309 22 , , , 32505 309 23 concave concave NNP 32505 309 24 ; ; : 32505 309 25 canthus canthus NNP 32505 309 26 rounded round VBD 32505 309 27 , , , 32505 309 28 indistinct indistinct JJ 32505 309 29 ; ; : 32505 309 30 diameter diameter NN 32505 309 31 of of IN 32505 309 32 eye eye NN 32505 309 33 about about IN 32505 309 34 equal equal JJ 32505 309 35 to to IN 32505 309 36 interorbital interorbital JJ 32505 309 37 space space NN 32505 309 38 ; ; : 32505 309 39 tympanum tympanum FW 32505 309 40 large large JJ 32505 309 41 , , , 32505 309 42 about about RB 32505 309 43 three three CD 32505 309 44 fifths fifth NNS 32505 309 45 diameter diameter NN 32505 309 46 of of IN 32505 309 47 eye eye NN 32505 309 48 , , , 32505 309 49 smaller small JJR 32505 309 50 than than IN 32505 309 51 internarial internarial JJ 32505 309 52 distance distance NN 32505 309 53 ; ; : 32505 309 54 supratympanic supratympanic NNP 32505 309 55 fold fold VBP 32505 309 56 indistinct indistinct JJ 32505 309 57 ; ; : 32505 309 58 arms arm NNS 32505 309 59 short short JJ 32505 309 60 ; ; : 32505 309 61 fingers finger NNS 32505 309 62 free free JJ 32505 309 63 of of IN 32505 309 64 webs webs NN 32505 309 65 ; ; : 32505 309 66 subarticular subarticular JJ 32505 309 67 tubercles tubercle NNS 32505 309 68 distinct distinct JJ 32505 309 69 ; ; : 32505 309 70 median median JJ 32505 309 71 palmar palmar NNP 32505 309 72 tubercle tubercle NN 32505 309 73 large large JJ 32505 309 74 , , , 32505 309 75 bifid bifid VBP 32505 309 76 ; ; : 32505 309 77 inner inner JJ 32505 309 78 palmar palmar NN 32505 309 79 tubercle tubercle NN 32505 309 80 on on IN 32505 309 81 base base NN 32505 309 82 of of IN 32505 309 83 first first NNP 32505 309 84 finger finger NNP 32505 309 85 flat flat JJ 32505 309 86 , , , 32505 309 87 elongate elongate NN 32505 309 88 ; ; : 32505 309 89 disc disc NN 32505 309 90 of of IN 32505 309 91 third third JJ 32505 309 92 finger finger NN 32505 309 93 about about IN 32505 309 94 one one CD 32505 309 95 half half JJ 32505 309 96 diameter diameter NN 32505 309 97 of of IN 32505 309 98 tympanum tympanum NN 32505 309 99 ; ; : 32505 309 100 legs leg NNS 32505 309 101 moderately moderately RB 32505 309 102 long long RB 32505 309 103 ; ; : 32505 309 104 tarsal tarsal JJ 32505 309 105 fold fold VB 32505 309 106 absent absent JJ 32505 309 107 ; ; : 32505 309 108 inner inner JJ 32505 309 109 metatarsal metatarsal NN 32505 309 110 tubercle tubercle NN 32505 309 111 distinct distinct NNP 32505 309 112 , , , 32505 309 113 oval oval NN 32505 309 114 ; ; : 32505 309 115 toes toe VBZ 32505 309 116 about about RB 32505 309 117 half half NN 32505 309 118 webbed web VBN 32505 309 119 ; ; : 32505 309 120 web web NN 32505 309 121 on on IN 32505 309 122 fourth fourth JJ 32505 309 123 toe toe NN 32505 309 124 extending extend VBG 32505 309 125 to to IN 32505 309 126 disc disc NN 32505 309 127 ; ; : 32505 309 128 discs disc NNS 32505 309 129 of of IN 32505 309 130 toes toe NNS 32505 309 131 about about IN 32505 309 132 size size NN 32505 309 133 of of IN 32505 309 134 those those DT 32505 309 135 on on IN 32505 309 136 fingers finger NNS 32505 309 137 ; ; : 32505 309 138 skin skin NN 32505 309 139 smooth smooth JJ 32505 309 140 above above RB 32505 309 141 with with IN 32505 309 142 small small JJ 32505 309 143 granules granule NNS 32505 309 144 on on IN 32505 309 145 head head NN 32505 309 146 and and CC 32505 309 147 in in IN 32505 309 148 scapular scapular JJ 32505 309 149 region region NN 32505 309 150 in in IN 32505 309 151 some some DT 32505 309 152 specimens specimen NNS 32505 309 153 ; ; : 32505 309 154 skin skin NN 32505 309 155 on on IN 32505 309 156 flanks flank NNS 32505 309 157 , , , 32505 309 158 throat throat NN 32505 309 159 , , , 32505 309 160 belly belly NN 32505 309 161 , , , 32505 309 162 and and CC 32505 309 163 lower low JJR 32505 309 164 surfaces surface NNS 32505 309 165 of of IN 32505 309 166 thighs thigh NNS 32505 309 167 granular granular JJ 32505 309 168 ; ; : 32505 309 169 tongue tongue NN 32505 309 170 oval oval NN 32505 309 171 , , , 32505 309 172 longer longer RB 32505 309 173 than than IN 32505 309 174 wide wide JJ 32505 309 175 , , , 32505 309 176 not not RB 32505 309 177 free free JJ 32505 309 178 behind behind RB 32505 309 179 ; ; : 32505 309 180 choanae choanae JJ 32505 309 181 small small JJ 32505 309 182 , , , 32505 309 183 oval oval JJ 32505 309 184 ; ; : 32505 309 185 vocal vocal NN 32505 309 186 slits slit NNS 32505 309 187 long long JJ 32505 309 188 , , , 32505 309 189 lateral lateral JJ 32505 309 190 to to IN 32505 309 191 tongue tongue NN 32505 309 192 . . . 32505 310 1 In in IN 32505 310 2 preservative preservative JJ 32505 310 3 , , , 32505 310 4 dorsum dorsum NN 32505 310 5 pale pale NN 32505 310 6 brown brown NNP 32505 310 7 with with IN 32505 310 8 darker dark JJR 32505 310 9 dorsolateral dorsolateral JJ 32505 310 10 stripes stripe NNS 32505 310 11 ; ; , 32505 310 12 narrow narrow JJ 32505 310 13 dark dark JJ 32505 310 14 brown brown JJ 32505 310 15 line line NN 32505 310 16 from from IN 32505 310 17 nostril nostril NNP 32505 310 18 to to IN 32505 310 19 eye eye NN 32505 310 20 ; ; : 32505 310 21 groin groin NN 32505 310 22 , , , 32505 310 23 anterior anterior NNP 32505 310 24 surface surface NN 32505 310 25 of of IN 32505 310 26 thighs thigh NNS 32505 310 27 , , , 32505 310 28 and and CC 32505 310 29 posteroventral posteroventral JJ 32505 310 30 surfaces surface NNS 32505 310 31 of of IN 32505 310 32 shanks shank NNS 32505 310 33 creamy creamy NNP 32505 310 34 tan tan NNP 32505 310 35 with with IN 32505 310 36 dark dark JJ 32505 310 37 brown brown JJ 32505 310 38 vermiculations vermiculation NNS 32505 310 39 ; ; : 32505 310 40 white white JJ 32505 310 41 spots spot NNS 32505 310 42 present present JJ 32505 310 43 on on IN 32505 310 44 thighs thigh NNS 32505 310 45 in in IN 32505 310 46 some some DT 32505 310 47 specimens specimen NNS 32505 310 48 ; ; : 32505 310 49 throat throat NN 32505 310 50 flecked fleck VBN 32505 310 51 with with IN 32505 310 52 brown brown JJ 32505 310 53 ; ; : 32505 310 54 belly belly NN 32505 310 55 creamy creamy JJ 32505 310 56 white white JJ 32505 310 57 or or CC 32505 310 58 gray gray JJ 32505 310 59 . . . 32505 311 1 _ _ NNP 32505 311 2 Remarks._--The Remarks._--The NNP 32505 311 3 taxonomic taxonomic JJ 32505 311 4 history history NN 32505 311 5 of of IN 32505 311 6 _ _ NNP 32505 311 7 Hyla Hyla NNP 32505 311 8 rubra rubra NN 32505 311 9 _ _ NNP 32505 311 10 Laurenti Laurenti NNP 32505 311 11 is be VBZ 32505 311 12 confused confused JJ 32505 311 13 . . . 32505 312 1 Seba Seba NNP 32505 312 2 ( ( -LRB- 32505 312 3 1734:70 1734:70 CD 32505 312 4 ) ) -RRB- 32505 312 5 illustrated illustrate VBD 32505 312 6 and and CC 32505 312 7 diagnosed diagnose VBD 32505 312 8 a a DT 32505 312 9 frog frog NN 32505 312 10 for for IN 32505 312 11 which which WDT 32505 312 12 he -PRON- PRP 32505 312 13 used use VBD 32505 312 14 the the DT 32505 312 15 name name NN 32505 312 16 _ _ NNP 32505 312 17 Ranula Ranula NNP 32505 312 18 , , , 32505 312 19 Americana Americana NNP 32505 312 20 , , , 32505 312 21 Rubra Rubra NNP 32505 312 22 _ _ NNP 32505 312 23 . . . 32505 313 1 Linnaeus Linnaeus NNP 32505 313 2 ( ( -LRB- 32505 313 3 1758:213 1758:213 CD 32505 313 4 ) ) -RRB- 32505 313 5 considered consider VBD 32505 313 6 Seba Seba NNP 32505 313 7 's 's POS 32505 313 8 frog frog NN 32505 313 9 to to TO 32505 313 10 be be VB 32505 313 11 a a DT 32505 313 12 variety variety NN 32505 313 13 of of IN 32505 313 14 _ _ NNP 32505 313 15 Hyla Hyla NNP 32505 313 16 arborea arborea NN 32505 313 17 _ _ NNP 32505 313 18 . . . 32505 314 1 Laurenti laurenti NN 32505 314 2 ( ( -LRB- 32505 314 3 1768:35 1768:35 CD 32505 314 4 ) ) -RRB- 32505 314 5 apparently apparently RB 32505 314 6 examined examine VBD 32505 314 7 the the DT 32505 314 8 same same JJ 32505 314 9 individual individual NN 32505 314 10 that that WDT 32505 314 11 Seba Seba NNP 32505 314 12 called call VBD 32505 314 13 _ _ NNP 32505 314 14 Ranula Ranula NNP 32505 314 15 , , , 32505 314 16 Americana Americana NNP 32505 314 17 , , , 32505 314 18 Rubra Rubra NNP 32505 314 19 _ _ NNP 32505 314 20 . . . 32505 315 1 For for IN 32505 315 2 this this DT 32505 315 3 specimen speciman NNS 32505 315 4 , , , 32505 315 5 Laurenti Laurenti NNP 32505 315 6 used use VBD 32505 315 7 the the DT 32505 315 8 binominal binominal NN 32505 315 9 _ _ NNP 32505 315 10 Hyla Hyla NNP 32505 315 11 rubra rubra NN 32505 315 12 _ _ NNP 32505 315 13 and and CC 32505 315 14 provided provide VBD 32505 315 15 a a DT 32505 315 16 brief brief JJ 32505 315 17 diagnosis diagnosis NN 32505 315 18 . . . 32505 316 1 The the DT 32505 316 2 type type NN 32505 316 3 locality locality NN 32505 316 4 was be VBD 32505 316 5 given give VBN 32505 316 6 as as IN 32505 316 7 America America NNP 32505 316 8 . . . 32505 317 1 Daudin Daudin NNP 32505 317 2 ( ( -LRB- 32505 317 3 1802:26 1802:26 NNP 32505 317 4 ) ) -RRB- 32505 317 5 redescribed redescribe VBD 32505 317 6 the the DT 32505 317 7 same same JJ 32505 317 8 specimen(s specimen(s NNP 32505 317 9 ? ? . 32505 317 10 ) ) -RRB- 32505 318 1 treated treat VBN 32505 318 2 by by IN 32505 318 3 Seba Seba NNP 32505 318 4 and and CC 32505 318 5 Laurenti Laurenti NNP 32505 318 6 and and CC 32505 318 7 provided provide VBD 32505 318 8 a a DT 32505 318 9 fairly fairly RB 32505 318 10 good good JJ 32505 318 11 description description NN 32505 318 12 and and CC 32505 318 13 figures figure NNS 32505 318 14 . . . 32505 319 1 Daudin Daudin NNP 32505 319 2 restricted restrict VBD 32505 319 3 the the DT 32505 319 4 type type NN 32505 319 5 locality locality NN 32505 319 6 to to IN 32505 319 7 Surinam Surinam NNP 32505 319 8 and and CC 32505 319 9 indicated indicate VBD 32505 319 10 that that IN 32505 319 11 Marin Marin NNP 32505 319 12 de de FW 32505 319 13 Baize Baize NNP 32505 319 14 was be VBD 32505 319 15 the the DT 32505 319 16 probable probable JJ 32505 319 17 collector collector NN 32505 319 18 . . . 32505 320 1 Daudin Daudin NNP 32505 320 2 ( ( -LRB- 32505 320 3 1802:26 1802:26 CD 32505 320 4 and and CC 32505 320 5 1803:53 1803:53 CD 32505 320 6 ) ) -RRB- 32505 320 7 neglected neglect VBD 32505 320 8 to to TO 32505 320 9 consider consider VB 32505 320 10 Laurenti Laurenti NNP 32505 320 11 's 's POS 32505 320 12 work work NN 32505 320 13 , , , 32505 320 14 but but CC 32505 320 15 he -PRON- PRP 32505 320 16 applied apply VBD 32505 320 17 the the DT 32505 320 18 same same JJ 32505 320 19 name name NN 32505 320 20 used use VBN 32505 320 21 by by IN 32505 320 22 Laurenti Laurenti NNP 32505 320 23 . . . 32505 321 1 Most Most JJS 32505 321 2 authors author NNS 32505 321 3 have have VBP 32505 321 4 credited credit VBN 32505 321 5 _ _ NNP 32505 321 6 Hyla Hyla NNP 32505 321 7 rubra rubra RB 32505 321 8 _ _ NNP 32505 321 9 to to IN 32505 321 10 Daudin Daudin NNP 32505 321 11 , , , 32505 321 12 but but CC 32505 321 13 Rivero Rivero NNP 32505 321 14 ( ( -LRB- 32505 321 15 1961:120 1961:120 CD 32505 321 16 ) ) -RRB- 32505 321 17 noted note VBD 32505 321 18 that that IN 32505 321 19 _ _ NNP 32505 321 20 Hyla Hyla NNP 32505 321 21 rubra rubra NN 32505 321 22 _ _ NNP 32505 321 23 Laurenti Laurenti NNP 32505 321 24 , , , 32505 321 25 1768 1768 CD 32505 321 26 , , , 32505 321 27 has have VBZ 32505 321 28 priority priority NN 32505 321 29 over over IN 32505 321 30 _ _ NNP 32505 321 31 Hyla Hyla NNP 32505 321 32 rubra rubra NN 32505 321 33 _ _ NNP 32505 321 34 Daudin Daudin NNP 32505 321 35 , , , 32505 321 36 1802 1802 CD 32505 321 37 . . . 32505 322 1 Since since IN 32505 322 2 both both CC 32505 322 3 Laurenti Laurenti NNP 32505 322 4 and and CC 32505 322 5 Daudin Daudin NNP 32505 322 6 worked work VBD 32505 322 7 on on IN 32505 322 8 Seba Seba NNP 32505 322 9 's 's POS 32505 322 10 material material NN 32505 322 11 , , , 32505 322 12 it -PRON- PRP 32505 322 13 is be VBZ 32505 322 14 reasonable reasonable JJ 32505 322 15 to to TO 32505 322 16 assume assume VB 32505 322 17 that that IN 32505 322 18 Daudin Daudin NNP 32505 322 19 redescribed redescribe VBD 32505 322 20 the the DT 32505 322 21 same same JJ 32505 322 22 frog frog NN 32505 322 23 that that WDT 32505 322 24 was be VBD 32505 322 25 named name VBN 32505 322 26 by by IN 32505 322 27 Laurenti Laurenti NNP 32505 322 28 ; ; : 32505 322 29 this this DT 32505 322 30 was be VBD 32505 322 31 not not RB 32505 322 32 an an DT 32505 322 33 uncommon uncommon JJ 32505 322 34 practice practice NN 32505 322 35 in in IN 32505 322 36 the the DT 32505 322 37 early early JJ 32505 322 38 nineteenth nineteenth JJ 32505 322 39 century century NN 32505 322 40 . . . 32505 323 1 Thus thus RB 32505 323 2 I -PRON- PRP 32505 323 3 conclude conclude VBP 32505 323 4 that that IN 32505 323 5 _ _ NNP 32505 323 6 Hyla Hyla NNP 32505 323 7 rubra rubra NN 32505 323 8 _ _ NNP 32505 323 9 Daudin Daudin NNP 32505 323 10 , , , 32505 323 11 1802 1802 CD 32505 323 12 , , , 32505 323 13 is be VBZ 32505 323 14 a a DT 32505 323 15 junior junior JJ 32505 323 16 synonym synonym NN 32505 323 17 of of IN 32505 323 18 _ _ NNP 32505 323 19 Hyla Hyla NNP 32505 323 20 rubra rubra NN 32505 323 21 _ _ NNP 32505 323 22 Laurenti Laurenti NNP 32505 323 23 , , , 32505 323 24 1768 1768 CD 32505 323 25 . . . 32505 324 1 Dunn Dunn NNP 32505 324 2 ( ( -LRB- 32505 324 3 1931a:413 1931a:413 NNP 32505 324 4 ) ) -RRB- 32505 324 5 first first RB 32505 324 6 reported report VBD 32505 324 7 _ _ NNP 32505 324 8 Hyla Hyla NNP 32505 324 9 rubra rubra NN 32505 324 10 _ _ NNP 32505 324 11 from from IN 32505 324 12 Central Central NNP 32505 324 13 America America NNP 32505 324 14 ; ; : 32505 324 15 he -PRON- PRP 32505 324 16 recorded record VBD 32505 324 17 the the DT 32505 324 18 species specie NNS 32505 324 19 from from IN 32505 324 20 the the DT 32505 324 21 Canal Canal NNP 32505 324 22 Zone Zone NNP 32505 324 23 and and CC 32505 324 24 San San NNP 32505 324 25 Pablo Pablo NNP 32505 324 26 , , , 32505 324 27 Panamá Panamá NNP 32505 324 28 . . . 32505 325 1 I -PRON- PRP 32505 325 2 have have VBP 32505 325 3 examined examine VBN 32505 325 4 the the DT 32505 325 5 material material NN 32505 325 6 of of IN 32505 325 7 _ _ NNP 32505 325 8 Hyla Hyla NNP 32505 325 9 rubra rubra NN 32505 325 10 _ _ NNP 32505 325 11 from from IN 32505 325 12 Panamá Panamá NNP 32505 325 13 deposited deposit VBN 32505 325 14 in in IN 32505 325 15 various various JJ 32505 325 16 museums museum NNS 32505 325 17 . . . 32505 326 1 Most Most JJS 32505 326 2 of of IN 32505 326 3 the the DT 32505 326 4 specimens specimen NNS 32505 326 5 are be VBP 32505 326 6 faded fade VBN 32505 326 7 , , , 32505 326 8 discolored discolor VBN 32505 326 9 , , , 32505 326 10 and and CC 32505 326 11 do do VBP 32505 326 12 not not RB 32505 326 13 have have VB 32505 326 14 distinct distinct JJ 32505 326 15 brown brown JJ 32505 326 16 vermiculations vermiculation NNS 32505 326 17 on on IN 32505 326 18 the the DT 32505 326 19 thighs thigh NNS 32505 326 20 . . . 32505 327 1 The the DT 32505 327 2 specimens specimen NNS 32505 327 3 seem seem VBP 32505 327 4 to to TO 32505 327 5 be be VB 32505 327 6 more more JJR 32505 327 7 like like IN 32505 327 8 _ _ NNP 32505 327 9 Hyla Hyla NNP 32505 327 10 rubra rubra RB 32505 327 11 _ _ NNP 32505 327 12 than than IN 32505 327 13 any any DT 32505 327 14 of of IN 32505 327 15 the the DT 32505 327 16 other other JJ 32505 327 17 species specie NNS 32505 327 18 in in IN 32505 327 19 the the DT 32505 327 20 _ _ NNP 32505 327 21 rubra rubra NNP 32505 327 22 _ _ NNP 32505 327 23 group group NN 32505 327 24 . . . 32505 328 1 The the DT 32505 328 2 presence presence NN 32505 328 3 of of IN 32505 328 4 oval oval NNP 32505 328 5 choanae choanae NN 32505 328 6 and and CC 32505 328 7 a a DT 32505 328 8 tympanum tympanum NN 32505 328 9 larger large JJR 32505 328 10 than than IN 32505 328 11 the the DT 32505 328 12 largest large JJS 32505 328 13 finger finger NN 32505 328 14 disc disc NN 32505 328 15 separate separate VBP 32505 328 16 these these DT 32505 328 17 specimens specimen NNS 32505 328 18 from from IN 32505 328 19 _ _ NNP 32505 328 20 Hyla Hyla NNP 32505 328 21 elaeochroa elaeochroa NN 32505 328 22 _ _ NNP 32505 328 23 , , , 32505 328 24 a a DT 32505 328 25 species species NN 32505 328 26 with with IN 32505 328 27 which which WDT 32505 328 28 _ _ NNP 32505 328 29 rubra rubra NNP 32505 328 30 _ _ NNP 32505 328 31 has have VBZ 32505 328 32 been be VBN 32505 328 33 confused confuse VBN 32505 328 34 . . . 32505 329 1 _ _ NNP 32505 329 2 Hyla Hyla NNP 32505 329 3 elaeochroa elaeochroa NN 32505 329 4 _ _ NNP 32505 329 5 does do VBZ 32505 329 6 not not RB 32505 329 7 occur occur VB 32505 329 8 in in IN 32505 329 9 the the DT 32505 329 10 Canal Canal NNP 32505 329 11 Zone Zone NNP 32505 329 12 or or CC 32505 329 13 eastern eastern NNP 32505 329 14 Panamá Panamá NNP 32505 329 15 . . . 32505 330 1 All all DT 32505 330 2 museum museum NN 32505 330 3 specimens specimen NNS 32505 330 4 from from IN 32505 330 5 Nicaragua Nicaragua NNP 32505 330 6 , , , 32505 330 7 Costa Costa NNP 32505 330 8 Rica Rica NNP 32505 330 9 , , , 32505 330 10 and and CC 32505 330 11 western western JJ 32505 330 12 Panamá Panamá NNP 32505 330 13 that that WDT 32505 330 14 have have VBP 32505 330 15 been be VBN 32505 330 16 called call VBN 32505 330 17 _ _ NNP 32505 330 18 Hyla Hyla NNP 32505 330 19 rubra rubra NN 32505 330 20 _ _ NNP 32505 330 21 , , , 32505 330 22 plus plus CC 32505 330 23 those those DT 32505 330 24 mentioned mention VBN 32505 330 25 by by IN 32505 330 26 Dunn Dunn NNP 32505 330 27 and and CC 32505 330 28 Emlen Emlen NNP 32505 330 29 ( ( -LRB- 32505 330 30 1932:25 1932:25 CD 32505 330 31 ) ) -RRB- 32505 330 32 and and CC 32505 330 33 Dunn Dunn NNP 32505 330 34 ( ( -LRB- 32505 330 35 1933:61 1933:61 CD 32505 330 36 ) ) -RRB- 32505 330 37 are be VBP 32505 330 38 _ _ NNP 32505 330 39 Hyla Hyla NNP 32505 330 40 elaeochroa elaeochroa NN 32505 330 41 _ _ NNP 32505 330 42 . . . 32505 331 1 The the DT 32505 331 2 taxonomic taxonomic JJ 32505 331 3 status status NN 32505 331 4 of of IN 32505 331 5 the the DT 32505 331 6 many many JJ 32505 331 7 South south JJ 32505 331 8 American american JJ 32505 331 9 populations population NNS 32505 331 10 referred refer VBD 32505 331 11 to to IN 32505 331 12 _ _ NNP 32505 331 13 Hyla Hyla NNP 32505 331 14 rubra rubra NN 32505 331 15 _ _ NNP 32505 331 16 and and CC 32505 331 17 of of IN 32505 331 18 other other JJ 32505 331 19 populations population NNS 32505 331 20 now now RB 32505 331 21 recognized recognize VBN 32505 331 22 as as IN 32505 331 23 different different JJ 32505 331 24 species specie NNS 32505 331 25 is be VBZ 32505 331 26 not not RB 32505 331 27 clear clear JJ 32505 331 28 at at IN 32505 331 29 the the DT 32505 331 30 present present JJ 32505 331 31 time time NN 32505 331 32 . . . 32505 332 1 Considerable considerable JJ 32505 332 2 variation variation NN 32505 332 3 in in IN 32505 332 4 external external JJ 32505 332 5 characters character NNS 32505 332 6 and and CC 32505 332 7 in in IN 32505 332 8 cranial cranial JJ 32505 332 9 features feature NNS 32505 332 10 has have VBZ 32505 332 11 been be VBN 32505 332 12 observed observe VBN 32505 332 13 in in IN 32505 332 14 South South NNP 32505 332 15 American American NNP 32505 332 16 _ _ NNP 32505 332 17 rubra rubra NNP 32505 332 18 _ _ NNP 32505 332 19 . . . 32505 333 1 A a DT 32505 333 2 review review NN 32505 333 3 of of IN 32505 333 4 the the DT 32505 333 5 taxonomy taxonomy NN 32505 333 6 of of IN 32505 333 7 these these DT 32505 333 8 populations population NNS 32505 333 9 is be VBZ 32505 333 10 beyond beyond IN 32505 333 11 the the DT 32505 333 12 scope scope NN 32505 333 13 of of IN 32505 333 14 this this DT 32505 333 15 paper paper NN 32505 333 16 . . . 32505 334 1 Possibly possibly RB 32505 334 2 the the DT 32505 334 3 Central central JJ 32505 334 4 American american JJ 32505 334 5 specimens specimens NNP 32505 334 6 herein herein NNP 32505 334 7 referred refer VBD 32505 334 8 to to IN 32505 334 9 _ _ NNP 32505 334 10 rubra rubra NNP 32505 334 11 _ _ NNP 32505 334 12 will will MD 32505 334 13 ultimately ultimately RB 32505 334 14 be be VB 32505 334 15 found find VBN 32505 334 16 to to TO 32505 334 17 be be VB 32505 334 18 specifically specifically RB 32505 334 19 distinct distinct JJ 32505 334 20 from from IN 32505 334 21 those those DT 32505 334 22 in in IN 32505 334 23 Surinam Surinam NNP 32505 334 24 . . . 32505 335 1 Since since IN 32505 335 2 I -PRON- PRP 32505 335 3 have have VBP 32505 335 4 no no DT 32505 335 5 osteological osteological JJ 32505 335 6 material material NN 32505 335 7 from from IN 32505 335 8 Central Central NNP 32505 335 9 America America NNP 32505 335 10 , , , 32505 335 11 I -PRON- PRP 32505 335 12 have have VBP 32505 335 13 been be VBN 32505 335 14 unable unable JJ 32505 335 15 to to TO 32505 335 16 describe describe VB 32505 335 17 the the DT 32505 335 18 cranium cranium NN 32505 335 19 in in IN 32505 335 20 this this DT 32505 335 21 account account NN 32505 335 22 . . . 32505 336 1 Furthermore furthermore RB 32505 336 2 , , , 32505 336 3 I -PRON- PRP 32505 336 4 have have VBP 32505 336 5 no no DT 32505 336 6 data data NN 32505 336 7 on on IN 32505 336 8 the the DT 32505 336 9 ecology ecology NN 32505 336 10 and and CC 32505 336 11 life life NN 32505 336 12 history history NN 32505 336 13 of of IN 32505 336 14 _ _ NNP 32505 336 15 rubra rubra NNP 32505 336 16 _ _ NNP 32505 336 17 in in IN 32505 336 18 Central Central NNP 32505 336 19 America America NNP 32505 336 20 . . . 32505 337 1 _ _ NNP 32505 337 2 Distribution._--_Hyla Distribution._--_Hyla NNP 32505 337 3 rubra rubra NNP 32505 337 4 _ _ NNP 32505 337 5 inhabits inhabit NNS 32505 337 6 lowland lowland VBP 32505 337 7 tropical tropical JJ 32505 337 8 forests forest NNS 32505 337 9 from from IN 32505 337 10 central central JJ 32505 337 11 - - HYPH 32505 337 12 eastern eastern JJ 32505 337 13 Panamá Panamá NNP 32505 337 14 to to IN 32505 337 15 northern northern JJ 32505 337 16 South South NNP 32505 337 17 America America NNP 32505 337 18 and and CC 32505 337 19 thence thence NNP 32505 337 20 through through IN 32505 337 21 lowlands lowland NNS 32505 337 22 east east NNP 32505 337 23 of of IN 32505 337 24 the the DT 32505 337 25 Andes Andes NNP 32505 337 26 to to TO 32505 337 27 northern northern JJ 32505 337 28 Argentina Argentina NNP 32505 337 29 ( ( -LRB- 32505 337 30 Fig Fig NNP 32505 337 31 . . . 32505 338 1 6 6 LS 32505 338 2 ) ) -RRB- 32505 338 3 . . . 32505 339 1 _ _ NNP 32505 339 2 Specimens Specimens NNP 32505 339 3 Examined._--Panamá Examined._--Panamá NNPS 32505 339 4 : : : 32505 339 5 _ _ NNP 32505 339 6 Canal Canal NNP 32505 339 7 Zone Zone NNP 32505 339 8 _ _ NNP 32505 339 9 : : : 32505 339 10 Gatun Gatun NNP 32505 339 11 , , , 32505 339 12 UMMZ UMMZ NNP 32505 339 13 52720 52720 CD 32505 339 14 ( ( -LRB- 32505 339 15 2 2 CD 32505 339 16 ) ) -RRB- 32505 339 17 ; ; : 32505 339 18 Madden Madden NNP 32505 339 19 Dam Dam NNP 32505 339 20 , , , 32505 339 21 FMNH FMNH NNP 32505 339 22 67820 67820 CD 32505 339 23 ; ; : 32505 339 24 no no DT 32505 339 25 specific specific JJ 32505 339 26 locality locality NN 32505 339 27 , , , 32505 339 28 UMMZ UMMZ NNP 32505 339 29 56517 56517 CD 32505 339 30 ( ( -LRB- 32505 339 31 3 3 CD 32505 339 32 ) ) -RRB- 32505 339 33 , , , 32505 339 34 USNM USNM NNP 32505 339 35 37863 37863 CD 32505 339 36 . . . 32505 340 1 _ _ NNP 32505 340 2 Colón Colón NNP 32505 340 3 _ _ NNP 32505 340 4 : : : 32505 340 5 Cerro Cerro NNP 32505 340 6 Bruja Bruja NNP 32505 340 7 , , , 32505 340 8 MCZ MCZ NNP 32505 340 9 13248 13248 CD 32505 340 10 . . . 32505 341 1 _ _ NNP 32505 341 2 Darién Darién NNP 32505 341 3 _ _ NNP 32505 341 4 : : : 32505 341 5 El El NNP 32505 341 6 Real Real NNP 32505 341 7 , , , 32505 341 8 USNM USNM NNP 32505 341 9 140569 140569 CD 32505 341 10 - - SYM 32505 341 11 70 70 CD 32505 341 12 , , , 32505 341 13 140573 140573 CD 32505 341 14 . . . 32505 342 1 _ _ NNP 32505 342 2 Panamá Panamá NNP 32505 342 3 _ _ NNP 32505 342 4 : : : 32505 342 5 Juan Juan NNP 32505 342 6 Díaz Díaz NNP 32505 342 7 , , , 32505 342 8 MCZ MCZ NNP 32505 342 9 17973 17973 CD 32505 342 10 ; ; : 32505 342 11 Las Las NNP 32505 342 12 Sabanas Sabanas NNP 32505 342 13 , , , 32505 342 14 MCZ MCZ NNP 32505 342 15 17581 17581 CD 32505 342 16 ; ; : 32505 342 17 Río Río NNP 32505 342 18 Trinidad Trinidad NNP 32505 342 19 , , , 32505 342 20 UMMZ UMMZ NNP 32505 342 21 64003 64003 CD 32505 342 22 ; ; : 32505 342 23 San San NNP 32505 342 24 Pablo Pablo NNP 32505 342 25 , , , 32505 342 26 MCZ MCZ NNP 32505 342 27 1398 1398 CD 32505 342 28 - - SYM 32505 342 29 9 9 CD 32505 342 30 . . . 32505 343 1 _ _ NNP 32505 343 2 Hyla Hyla NNP 32505 343 3 elaeochroa elaeochroa NN 32505 343 4 _ _ NNP 32505 343 5 Cope Cope NNP 32505 343 6 _ _ NNP 32505 343 7 Hyla Hyla NNP 32505 343 8 elaeochroa elaeochroa NN 32505 343 9 _ _ NNP 32505 343 10 Cope Cope NNP 32505 343 11 , , , 32505 343 12 Jour Jour NNP 32505 343 13 . . . 32505 344 1 Acad Acad NNS 32505 344 2 . . . 32505 345 1 Nat Nat NNP 32505 345 2 . . . 32505 346 1 Sci Sci NNP 32505 346 2 . . . 32505 347 1 Philadelphia Philadelphia NNP 32505 347 2 , , , 32505 347 3 8:105 8:105 CD 32505 347 4 , , , 32505 347 5 1876 1876 CD 32505 347 6 [ [ -LRB- 32505 347 7 Holotype.--USNM Holotype.--USNM NNP 32505 347 8 30689 30689 CD 32505 347 9 , , , 32505 347 10 Sipurio Sipurio NNP 32505 347 11 , , , 32505 347 12 Limón Limón NNP 32505 347 13 Province Province NNP 32505 347 14 , , , 32505 347 15 Costa Costa NNP 32505 347 16 Rica Rica NNP 32505 347 17 ; ; : 32505 347 18 William William NNP 32505 347 19 M. M. NNP 32505 347 20 Gabb Gabb NNP 32505 347 21 collector collector NN 32505 347 22 ] ] -RRB- 32505 347 23 . . . 32505 348 1 Günther Günther NNP 32505 348 2 , , , 32505 348 3 Biologia Biologia NNP 32505 348 4 Centrali Centrali NNP 32505 348 5 - - HYPH 32505 348 6 Americana Americana NNP 32505 348 7 , , , 32505 348 8 Reptilia Reptilia NNP 32505 348 9 and and CC 32505 348 10 Batrachia Batrachia NNP 32505 348 11 , , , 32505 348 12 p. p. NN 32505 348 13 265 265 CD 32505 348 14 , , , 32505 348 15 June June NNP 32505 348 16 1901 1901 CD 32505 348 17 . . . 32505 349 1 Taylor Taylor NNP 32505 349 2 , , , 32505 349 3 Univ Univ NNP 32505 349 4 . . . 32505 350 1 Kansas Kansas NNP 32505 350 2 Sci Sci NNP 32505 350 3 . . . 32505 351 1 Bull Bull NNP 32505 351 2 . . NNP 32505 351 3 , , , 32505 351 4 35:859 35:859 CD 32505 351 5 , , , 32505 351 6 July July NNP 32505 351 7 1 1 CD 32505 351 8 , , , 32505 351 9 1952 1952 CD 32505 351 10 . . . 32505 352 1 Duellman Duellman NNP 32505 352 2 , , , 32505 352 3 Univ Univ NNP 32505 352 4 . . . 32505 353 1 Kansas Kansas NNP 32505 353 2 Publ Publ NNP 32505 353 3 . . . 32505 353 4 , , , 32505 353 5 Mus mus NN 32505 353 6 . . . 32505 354 1 Nat Nat NNP 32505 354 2 . . . 32505 355 1 Hist Hist NNP 32505 355 2 . . . 32505 355 3 , , , 32505 355 4 17:270 17:270 CD 32505 355 5 , , , 32505 355 6 June June NNP 32505 355 7 17 17 CD 32505 355 8 , , , 32505 355 9 1966 1966 CD 32505 355 10 . . . 32505 356 1 _ _ NNP 32505 356 2 Hyla Hyla NNP 32505 356 3 quinquevittata quinquevittata NN 32505 356 4 _ _ NNP 32505 356 5 Cope Cope NNP 32505 356 6 , , , 32505 356 7 Proc Proc NNP 32505 356 8 . . . 32505 357 1 Amer amer NN 32505 357 2 . . . 32505 358 1 Philos Philos NNP 32505 358 2 . . . 32505 359 1 Soc Soc NNP 32505 359 2 . . NNP 32505 359 3 , , , 32505 359 4 23:273 23:273 CD 32505 359 5 , , , 32505 359 6 April April NNP 32505 359 7 1887 1887 CD 32505 359 8 [ [ -LRB- 32505 359 9 Holotype.--USNM Holotype.--USNM NNP 32505 359 10 14187 14187 CD 32505 359 11 , , , 32505 359 12 Nicaragua Nicaragua NNP 32505 359 13 ; ; : 32505 359 14 J. J. NNP 32505 359 15 F. F. NNP 32505 359 16 Bransford Bransford NNP 32505 359 17 collector collector NN 32505 359 18 ] ] -RRB- 32505 359 19 . . . 32505 360 1 Günther Günther NNP 32505 360 2 , , , 32505 360 3 Biologia Biologia NNP 32505 360 4 Centrali Centrali NNP 32505 360 5 - - HYPH 32505 360 6 Americana Americana NNP 32505 360 7 , , , 32505 360 8 Reptilia Reptilia NNP 32505 360 9 and and CC 32505 360 10 Batrachia Batrachia NNP 32505 360 11 , , , 32505 360 12 p. p. NN 32505 360 13 268 268 CD 32505 360 14 , , , 32505 360 15 June June NNP 32505 360 16 1901 1901 CD 32505 360 17 . . . 32505 361 1 Noble Noble NNP 32505 361 2 , , , 32505 361 3 Bull Bull NNP 32505 361 4 . . . 32505 362 1 Amer amer NN 32505 362 2 . . . 32505 363 1 Mus mus NN 32505 363 2 . . . 32505 364 1 Nat Nat NNP 32505 364 2 . . . 32505 365 1 Hist Hist NNP 32505 365 2 . . . 32505 365 3 , , , 32505 365 4 38:340 38:340 CD 32505 365 5 , , , 32505 365 6 June June NNP 32505 365 7 1918 1918 CD 32505 365 8 . . . 32505 366 1 _ _ NNP 32505 366 2 Hyla Hyla NNP 32505 366 3 rubra rubra NN 32505 366 4 _ _ NNP 32505 366 5 ( ( -LRB- 32505 366 6 part part NN 32505 366 7 ) ) -RRB- 32505 366 8 : : : 32505 366 9 Dunn Dunn NNP 32505 366 10 and and CC 32505 366 11 Emlen Emlen NNP 32505 366 12 , , , 32505 366 13 Proc Proc NNP 32505 366 14 . . . 32505 367 1 Acad Acad NNS 32505 367 2 . . . 32505 368 1 Nat Nat NNP 32505 368 2 . . . 32505 369 1 Sci Sci NNP 32505 369 2 . . . 32505 370 1 Philadelphia Philadelphia NNP 32505 370 2 , , , 32505 370 3 84:25 84:25 CD 32505 370 4 , , , 32505 370 5 March March NNP 32505 370 6 22 22 CD 32505 370 7 , , , 32505 370 8 1932 1932 CD 32505 370 9 . . . 32505 371 1 _ _ NNP 32505 371 2 Hyla Hyla NNP 32505 371 3 dulcensis dulcensis NN 32505 371 4 _ _ NNP 32505 371 5 Taylor Taylor NNP 32505 371 6 , , , 32505 371 7 Univ Univ NNP 32505 371 8 . . . 32505 372 1 Kansas Kansas NNP 32505 372 2 Sci Sci NNP 32505 372 3 . . . 32505 373 1 Bull Bull NNP 32505 373 2 . . . 32505 373 3 , , , 32505 373 4 39:37 39:37 CD 32505 373 5 , , , 32505 373 6 November November NNP 32505 373 7 18 18 CD 32505 373 8 , , , 32505 373 9 1958 1958 CD 32505 373 10 [ [ -LRB- 32505 373 11 Holotype.--KU Holotype.--KU NNP 32505 373 12 32168 32168 CD 32505 373 13 , , , 32505 373 14 Golfito Golfito NNP 32505 373 15 , , , 32505 373 16 Puntarenas Puntarenas NNP 32505 373 17 Province Province NNP 32505 373 18 , , , 32505 373 19 Costa Costa NNP 32505 373 20 Rica Rica NNP 32505 373 21 ; ; : 32505 373 22 Edward Edward NNP 32505 373 23 H. H. NNP 32505 373 24 Taylor Taylor NNP 32505 373 25 collector collector NN 32505 373 26 ] ] -RRB- 32505 373 27 . . . 32505 374 1 _ _ NNP 32505 374 2 Diagnosis._--Size Diagnosis._--Size NNP 32505 374 3 medium medium NN 32505 374 4 ( ( -LRB- 32505 374 5 Male male NN 32505 374 6 to to IN 32505 374 7 38 38 CD 32505 374 8 mm mm NNS 32505 374 9 . . NNP 32505 374 10 , , , 32505 374 11 Female Female NNP 32505 374 12 to to IN 32505 374 13 40 40 CD 32505 374 14 mm mm NNS 32505 374 15 . . . 32505 375 1 ) ) -RRB- 32505 375 2 ; ; : 32505 375 3 skull skull VB 32505 375 4 wider wider RBR 32505 375 5 than than IN 32505 375 6 long long RB 32505 375 7 ; ; : 32505 375 8 nasals nasal NNS 32505 375 9 truncate truncate VBP 32505 375 10 anteriorly anteriorly RB 32505 375 11 ; ; : 32505 375 12 frontoparietal frontoparietal JJ 32505 375 13 fontanelle fontanelle NNP 32505 375 14 moderate moderate NNP 32505 375 15 in in IN 32505 375 16 size size NN 32505 375 17 ; ; : 32505 375 18 snout snout VB 32505 375 19 slightly slightly RB 32505 375 20 protruding protrude VBG 32505 375 21 ; ; : 32505 375 22 tympanum tympanum NN 32505 375 23 about about IN 32505 375 24 size size NN 32505 375 25 of of IN 32505 375 26 largest large JJS 32505 375 27 discs disc NNS 32505 375 28 on on IN 32505 375 29 fingers finger NNS 32505 375 30 ; ; : 32505 375 31 dorsum dorsum NN 32505 375 32 marked mark VBN 32505 375 33 by by IN 32505 375 34 longitudinal longitudinal JJ 32505 375 35 stripes stripe NNS 32505 375 36 ; ; , 32505 375 37 dark dark JJ 32505 375 38 stripe stripe NN 32505 375 39 between between IN 32505 375 40 eye eye NN 32505 375 41 and and CC 32505 375 42 nostril nostril NNP 32505 375 43 ; ; : 32505 375 44 in in IN 32505 375 45 life life NN 32505 375 46 tan tan NN 32505 375 47 to to IN 32505 375 48 olive olive NN 32505 375 49 - - HYPH 32505 375 50 green green JJ 32505 375 51 with with IN 32505 375 52 or or CC 32505 375 53 without without IN 32505 375 54 dark dark JJ 32505 375 55 mark mark NN 32505 375 56 between between IN 32505 375 57 eyes eye NNS 32505 375 58 ; ; : 32505 375 59 bones bone NNS 32505 375 60 greenish greenish JJ 32505 375 61 blue blue NNP 32505 375 62 . . . 32505 376 1 _ _ NNP 32505 376 2 Description._--Head Description._--Head NNP 32505 376 3 flat flat RB 32505 376 4 , , , 32505 376 5 longer longer RB 32505 376 6 than than IN 32505 376 7 wide wide JJ 32505 376 8 ; ; : 32505 376 9 snout snout NN 32505 376 10 long long RB 32505 376 11 , , , 32505 376 12 rounded round VBD 32505 376 13 , , , 32505 376 14 protruding protrude VBG 32505 376 15 beyond beyond IN 32505 376 16 mouth mouth NN 32505 376 17 ; ; : 32505 376 18 canthus canthus NNP 32505 376 19 indistinct indistinct NNP 32505 376 20 ; ; : 32505 376 21 length length NN 32505 376 22 of of IN 32505 376 23 eye eye NN 32505 376 24 equal equal JJ 32505 376 25 to to IN 32505 376 26 interorbital interorbital JJ 32505 376 27 distance distance NN 32505 376 28 ; ; , 32505 376 29 loreal loreal NN 32505 376 30 region region NN 32505 376 31 not not RB 32505 376 32 pronounced pronounce VBN 32505 376 33 ; ; : 32505 376 34 tympanum tympanum VB 32505 376 35 distinct distinct JJ 32505 376 36 and and CC 32505 376 37 about about RB 32505 376 38 two two CD 32505 376 39 - - HYPH 32505 376 40 fifths fifth NNS 32505 376 41 diameter diameter NN 32505 376 42 of of IN 32505 376 43 eye eye NN 32505 376 44 ; ; : 32505 376 45 interorbital interorbital NNP 32505 376 46 triangle triangle NNP 32505 376 47 present present JJ 32505 376 48 or or CC 32505 376 49 absent absent JJ 32505 376 50 ; ; : 32505 376 51 arms arm NNS 32505 376 52 short short JJ 32505 376 53 ; ; : 32505 376 54 trace trace NN 32505 376 55 of of IN 32505 376 56 web web NN 32505 376 57 between between IN 32505 376 58 fingers finger NNS 32505 376 59 , , , 32505 376 60 extending extend VBG 32505 376 61 as as IN 32505 376 62 fringe fringe NN 32505 376 63 along along IN 32505 376 64 sides side NNS 32505 376 65 of of IN 32505 376 66 fingers finger NNS 32505 376 67 ; ; : 32505 376 68 first first RB 32505 376 69 finger finger NN 32505 376 70 very very RB 32505 376 71 short short JJ 32505 376 72 with with IN 32505 376 73 small small JJ 32505 376 74 disc disc NN 32505 376 75 ; ; : 32505 376 76 other other JJ 32505 376 77 discs disc NNS 32505 376 78 about about IN 32505 376 79 size size NN 32505 376 80 of of IN 32505 376 81 those those DT 32505 376 82 on on IN 32505 376 83 toes toe NNS 32505 376 84 ; ; , 32505 376 85 discs disc NNS 32505 376 86 on on IN 32505 376 87 third third JJ 32505 376 88 finger finger NN 32505 376 89 and and CC 32505 376 90 fourth fourth JJ 32505 376 91 toe toe NN 32505 376 92 as as RB 32505 376 93 large large JJ 32505 376 94 as as IN 32505 376 95 tympanum tympanum NN 32505 376 96 ; ; : 32505 376 97 outer outer JJ 32505 376 98 palmar palmar NNP 32505 376 99 tubercle tubercle NN 32505 376 100 moderate moderate JJ 32505 376 101 in in IN 32505 376 102 size size NN 32505 376 103 , , , 32505 376 104 partly partly RB 32505 376 105 bifid bifid VBP 32505 376 106 ; ; : 32505 376 107 inner inner JJ 32505 376 108 palmar palmar NNP 32505 376 109 tubercle tubercle NN 32505 376 110 large large NNP 32505 376 111 , , , 32505 376 112 elongate elongate VB 32505 376 113 , , , 32505 376 114 flat flat JJ 32505 376 115 ; ; : 32505 376 116 subarticular subarticular JJ 32505 376 117 tubercles tubercle NNS 32505 376 118 distinct distinct JJ 32505 376 119 ; ; : 32505 376 120 legs leg NNS 32505 376 121 moderately moderately RB 32505 376 122 long long RB 32505 376 123 ; ; : 32505 376 124 tarsal tarsal JJ 32505 376 125 fold fold VB 32505 376 126 absent absent JJ 32505 376 127 ; ; : 32505 376 128 inner inner JJ 32505 376 129 metatarsal metatarsal NN 32505 376 130 tubercle tubercle NN 32505 376 131 flat flat JJ 32505 376 132 ; ; : 32505 376 133 outer outer JJ 32505 376 134 metatarsal metatarsal NN 32505 376 135 tubercle tubercle NN 32505 376 136 smaller smaller RBR 32505 376 137 , , , 32505 376 138 indistinct indistinct NNP 32505 376 139 ; ; : 32505 376 140 subarticular subarticular JJ 32505 376 141 tubercles tubercle NNS 32505 376 142 moderate moderate JJ 32505 376 143 in in IN 32505 376 144 size size NN 32505 376 145 ; ; , 32505 376 146 fringe fringe NN 32505 376 147 on on IN 32505 376 148 toes toe NNS 32505 376 149 to to TO 32505 376 150 tip tip VB 32505 376 151 of of IN 32505 376 152 disc disc NN 32505 376 153 of of IN 32505 376 154 second second JJ 32505 376 155 toe toe NN 32505 376 156 ; ; : 32505 376 157 rest rest NN 32505 376 158 of of IN 32505 376 159 toes toe NNS 32505 376 160 about about RB 32505 376 161 two two CD 32505 376 162 - - HYPH 32505 376 163 thirds third NNS 32505 376 164 webbed web VBN 32505 376 165 ; ; : 32505 376 166 foot foot NN 32505 376 167 length length NN 32505 376 168 about about RB 32505 376 169 two two CD 32505 376 170 fifths fifth NNS 32505 376 171 snout snout NN 32505 376 172 - - HYPH 32505 376 173 vent vent NN 32505 376 174 length length NN 32505 376 175 ; ; : 32505 376 176 tibia tibia NNP 32505 376 177 length length NN 32505 376 178 about about RB 32505 376 179 one one CD 32505 376 180 half half JJ 32505 376 181 snout snout NN 32505 376 182 - - HYPH 32505 376 183 vent vent NN 32505 376 184 length length NN 32505 376 185 ; ; : 32505 376 186 skin skin NN 32505 376 187 above above IN 32505 376 188 smooth smooth JJ 32505 376 189 or or CC 32505 376 190 with with IN 32505 376 191 minute minute JJ 32505 376 192 pustules pustule NNS 32505 376 193 ; ; : 32505 376 194 belly belly NN 32505 376 195 finely finely RB 32505 376 196 granular granular JJ 32505 376 197 ; ; , 32505 376 198 ventral ventral JJ 32505 376 199 surfaces surface NNS 32505 376 200 of of IN 32505 376 201 thighs thigh NNS 32505 376 202 and and CC 32505 376 203 areas area NNS 32505 376 204 below below IN 32505 376 205 anus anus NN 32505 376 206 granular granular NN 32505 376 207 ; ; : 32505 376 208 skin skin NN 32505 376 209 on on IN 32505 376 210 ventral ventral JJ 32505 376 211 surfaces surface NNS 32505 376 212 of of IN 32505 376 213 limbs limb NNS 32505 376 214 smooth smooth VBP 32505 376 215 ; ; : 32505 376 216 tongue tongue NN 32505 376 217 relatively relatively RB 32505 376 218 large large JJ 32505 376 219 , , , 32505 376 220 longer long JJR 32505 376 221 than than IN 32505 376 222 wide wide JJ 32505 376 223 , , , 32505 376 224 barely barely RB 32505 376 225 notched notch VBN 32505 376 226 behind behind RB 32505 376 227 ; ; : 32505 376 228 vocal vocal NN 32505 376 229 slits slit NNS 32505 376 230 elongate elongate VBP 32505 376 231 , , , 32505 376 232 lateral lateral JJ 32505 376 233 to to IN 32505 376 234 tongue tongue NN 32505 376 235 ; ; : 32505 376 236 choanae choanae JJ 32505 376 237 medium medium NN 32505 376 238 in in IN 32505 376 239 size size NN 32505 376 240 . . . 32505 377 1 In in IN 32505 377 2 life life NN 32505 377 3 , , , 32505 377 4 dorsum dorsum NN 32505 377 5 yellowish yellowish NNP 32505 377 6 brown brown NNP 32505 377 7 , , , 32505 377 8 olive olive JJ 32505 377 9 green green NN 32505 377 10 , , , 32505 377 11 or or CC 32505 377 12 grayish grayish VB 32505 377 13 brown brown JJ 32505 377 14 with with IN 32505 377 15 dark dark JJ 32505 377 16 brown brown JJ 32505 377 17 spots spot NNS 32505 377 18 on on IN 32505 377 19 snout snout NN 32505 377 20 , , , 32505 377 21 dark dark JJ 32505 377 22 brown brown JJ 32505 377 23 stripe stripe NN 32505 377 24 from from IN 32505 377 25 nostril nostril NNP 32505 377 26 to to IN 32505 377 27 eye eye NN 32505 377 28 , , , 32505 377 29 dark dark JJ 32505 377 30 yellow yellow JJ 32505 377 31 - - HYPH 32505 377 32 brown brown JJ 32505 377 33 interorbital interorbital NN 32505 377 34 triangle triangle NN 32505 377 35 , , , 32505 377 36 and and CC 32505 377 37 dark dark JJ 32505 377 38 supratympanic supratympanic NN 32505 377 39 region region NN 32505 377 40 ; ; : 32505 377 41 generally generally RB 32505 377 42 five five CD 32505 377 43 interrupted interrupted JJ 32505 377 44 longitudinal longitudinal JJ 32505 377 45 dark dark JJ 32505 377 46 brown brown JJ 32505 377 47 stripes stripe NNS 32505 377 48 on on IN 32505 377 49 dorsum dorsum NN 32505 377 50 ( ( -LRB- 32505 377 51 one one CD 32505 377 52 on on IN 32505 377 53 each each DT 32505 377 54 flank flank NN 32505 377 55 , , , 32505 377 56 pair pair NN 32505 377 57 of of IN 32505 377 58 paravertebral paravertebral NN 32505 377 59 and and CC 32505 377 60 one one CD 32505 377 61 vertebral vertebral NN 32505 377 62 ) ) -RRB- 32505 377 63 ; ; : 32505 377 64 flanks flank VBZ 32505 377 65 pale pale JJ 32505 377 66 yellow yellow NNP 32505 377 67 ; ; : 32505 377 68 groin groin NNP 32505 377 69 yellowish yellowish NNP 32505 377 70 brown brown NNP 32505 377 71 ; ; : 32505 377 72 thighs thigh NNS 32505 377 73 marked mark VBN 32505 377 74 with with IN 32505 377 75 one one CD 32505 377 76 or or CC 32505 377 77 two two CD 32505 377 78 transverse transverse JJ 32505 377 79 yellow yellow JJ 32505 377 80 - - HYPH 32505 377 81 brown brown JJ 32505 377 82 blotches blotch NNS 32505 377 83 ; ; : 32505 377 84 shanks shank VBZ 32505 377 85 with with IN 32505 377 86 two two CD 32505 377 87 or or CC 32505 377 88 three three CD 32505 377 89 yellow yellow JJ 32505 377 90 - - HYPH 32505 377 91 brown brown JJ 32505 377 92 blotches blotch NNS 32505 377 93 above above RB 32505 377 94 ; ; : 32505 377 95 spaces space NNS 32505 377 96 between between IN 32505 377 97 blotches blotch NNS 32505 377 98 on on IN 32505 377 99 thighs thigh NNS 32505 377 100 , , , 32505 377 101 shanks shank NNS 32505 377 102 , , , 32505 377 103 tarsi tarsi NNP 32505 377 104 , , , 32505 377 105 and and CC 32505 377 106 feet foot NNS 32505 377 107 yellow yellow JJ 32505 377 108 ; ; : 32505 377 109 brown brown JJ 32505 377 110 spots spot NNS 32505 377 111 on on IN 32505 377 112 tarsi tarsi NNP 32505 377 113 and and CC 32505 377 114 in in IN 32505 377 115 some some DT 32505 377 116 specimens specimen NNS 32505 377 117 on on IN 32505 377 118 feet foot NNS 32505 377 119 ; ; , 32505 377 120 arm arm NN 32505 377 121 pale pale JJ 32505 377 122 yellow yellow NN 32505 377 123 with with IN 32505 377 124 pale pale JJ 32505 377 125 brown brown JJ 32505 377 126 spots spot NNS 32505 377 127 ; ; : 32505 377 128 belly belly NN 32505 377 129 creamy creamy JJ 32505 377 130 white white JJ 32505 377 131 having have VBG 32505 377 132 slight slight JJ 32505 377 133 blue blue JJ 32505 377 134 - - HYPH 32505 377 135 green green JJ 32505 377 136 tint tint NN 32505 377 137 ; ; : 32505 377 138 vocal vocal JJ 32505 377 139 sac sac NN 32505 377 140 and and CC 32505 377 141 chin chin NNP 32505 377 142 yellow yellow NNP 32505 377 143 ; ; : 32505 377 144 axillary axillary JJ 32505 377 145 region region NN 32505 377 146 yellow yellow NN 32505 377 147 , , , 32505 377 148 blue blue JJ 32505 377 149 - - HYPH 32505 377 150 green green JJ 32505 377 151 in in IN 32505 377 152 some some DT 32505 377 153 specimens specimen NNS 32505 377 154 ( ( -LRB- 32505 377 155 Pl Pl NNP 32505 377 156 . . . 32505 378 1 2A 2a LS 32505 378 2 ) ) -RRB- 32505 378 3 . . . 32505 379 1 In in IN 32505 379 2 preservative preservative JJ 32505 379 3 , , , 32505 379 4 head head NN 32505 379 5 and and CC 32505 379 6 dorsum dorsum NN 32505 379 7 yellowish yellowish JJ 32505 379 8 brown brown NN 32505 379 9 ; ; : 32505 379 10 dark dark JJ 32505 379 11 brown brown JJ 32505 379 12 stripe stripe NN 32505 379 13 from from IN 32505 379 14 nostril nostril NNP 32505 379 15 to to IN 32505 379 16 eye eye NN 32505 379 17 ; ; : 32505 379 18 dark dark JJ 32505 379 19 brown brown JJ 32505 379 20 spots spot NNS 32505 379 21 on on IN 32505 379 22 snout snout NN 32505 379 23 ; ; : 32505 379 24 a a DT 32505 379 25 dark dark JJ 32505 379 26 brown brown JJ 32505 379 27 interorbital interorbital NN 32505 379 28 triangle triangle NN 32505 379 29 with with IN 32505 379 30 apex apex NNP 32505 379 31 directed direct VBN 32505 379 32 backward backward RB 32505 379 33 ; ; : 32505 379 34 dark dark JJ 32505 379 35 brown brown JJ 32505 379 36 supratympanic supratympanic NN 32505 379 37 region region NN 32505 379 38 ; ; : 32505 379 39 dorsal dorsal NN 32505 379 40 stripes stripe NNS 32505 379 41 same same JJ 32505 379 42 as as IN 32505 379 43 in in IN 32505 379 44 living live VBG 32505 379 45 individuals individual NNS 32505 379 46 ; ; : 32505 379 47 flanks flank VBZ 32505 379 48 pale pale JJ 32505 379 49 yellow yellow NN 32505 379 50 with with IN 32505 379 51 brown brown JJ 32505 379 52 spots spot NNS 32505 379 53 in in IN 32505 379 54 some some DT 32505 379 55 specimens specimen NNS 32505 379 56 ; ; : 32505 379 57 groin groin NNP 32505 379 58 creamy creamy JJ 32505 379 59 white white NNP 32505 379 60 ; ; : 32505 379 61 thighs thigh NNS 32505 379 62 and and CC 32505 379 63 shanks shank NNS 32505 379 64 having have VBG 32505 379 65 or or CC 32505 379 66 lacking lack VBG 32505 379 67 transverse transverse JJ 32505 379 68 dark dark JJ 32505 379 69 brown brown JJ 32505 379 70 blotches blotch NNS 32505 379 71 ; ; , 32505 379 72 spaces space NNS 32505 379 73 between between IN 32505 379 74 blotches blotch NNS 32505 379 75 creamy creamy VBP 32505 379 76 white white JJ 32505 379 77 or or CC 32505 379 78 yellow yellow JJ 32505 379 79 - - HYPH 32505 379 80 brown brown JJ 32505 379 81 ; ; : 32505 379 82 arms arm NNS 32505 379 83 pale pale JJ 32505 379 84 yellowish yellowish NNP 32505 379 85 brown brown NNP 32505 379 86 ; ; : 32505 379 87 belly belly NN 32505 379 88 and and CC 32505 379 89 vocal vocal NN 32505 379 90 sac sac NN 32505 379 91 creamy creamy JJ 32505 379 92 white white NN 32505 379 93 . . . 32505 380 1 _ _ NNP 32505 380 2 Variation._--Geographic variation._--geographic JJ 32505 380 3 variation variation NN 32505 380 4 in in IN 32505 380 5 size size NN 32505 380 6 and and CC 32505 380 7 some some DT 32505 380 8 proportions proportion NNS 32505 380 9 , , , 32505 380 10 such such JJ 32505 380 11 as as IN 32505 380 12 the the DT 32505 380 13 ratio ratio NN 32505 380 14 of of IN 32505 380 15 tibia tibia NNP 32505 380 16 length length NN 32505 380 17 to to TO 32505 380 18 snout snout VB 32505 380 19 - - HYPH 32505 380 20 vent vent NN 32505 380 21 length length NN 32505 380 22 and and CC 32505 380 23 the the DT 32505 380 24 ratio ratio NN 32505 380 25 of of IN 32505 380 26 the the DT 32505 380 27 diameter diameter NN 32505 380 28 of of IN 32505 380 29 the the DT 32505 380 30 tympanum tympanum NN 32505 380 31 to to IN 32505 380 32 that that DT 32505 380 33 of of IN 32505 380 34 the the DT 32505 380 35 eye eye NN 32505 380 36 , , , 32505 380 37 have have VBP 32505 380 38 been be VBN 32505 380 39 observed observe VBN 32505 380 40 in in IN 32505 380 41 this this DT 32505 380 42 species species NN 32505 380 43 . . . 32505 381 1 The the DT 32505 381 2 largest large JJS 32505 381 3 individuals individual NNS 32505 381 4 are be VBP 32505 381 5 from from IN 32505 381 6 the the DT 32505 381 7 Golfo Golfo NNP 32505 381 8 Dulce Dulce NNP 32505 381 9 region region NN 32505 381 10 ( ( -LRB- 32505 381 11 samples sample NNS 32505 381 12 from from IN 32505 381 13 Piedras Piedras NNP 32505 381 14 Blancas Blancas NNP 32505 381 15 and and CC 32505 381 16 Rincón Rincón NNP 32505 381 17 de de NNP 32505 381 18 Osa Osa NNP 32505 381 19 ) ) -RRB- 32505 381 20 , , , 32505 381 21 Puntarenas Puntarenas NNP 32505 381 22 Province Province NNP 32505 381 23 , , , 32505 381 24 Costa Costa NNP 32505 381 25 Rica Rica NNP 32505 381 26 . . . 32505 382 1 The the DT 32505 382 2 smallest small JJS 32505 382 3 individuals individual NNS 32505 382 4 are be VBP 32505 382 5 from from IN 32505 382 6 El El NNP 32505 382 7 Recreo Recreo NNP 32505 382 8 , , , 32505 382 9 Zelaya Zelaya NNP 32505 382 10 Province Province NNP 32505 382 11 , , , 32505 382 12 Nicaragua Nicaragua NNP 32505 382 13 , , , 32505 382 14 and and CC 32505 382 15 from from IN 32505 382 16 the the DT 32505 382 17 Caribbean caribbean JJ 32505 382 18 lowlands lowland NNS 32505 382 19 of of IN 32505 382 20 Costa Costa NNP 32505 382 21 Rica Rica NNP 32505 382 22 . . . 32505 383 1 The the DT 32505 383 2 diameter diameter NN 32505 383 3 of of IN 32505 383 4 the the DT 32505 383 5 tympanum tympanum NN 32505 383 6 is be VBZ 32505 383 7 proportionately proportionately RB 32505 383 8 larger large JJR 32505 383 9 ( ( -LRB- 32505 383 10 relative relative JJ 32505 383 11 to to IN 32505 383 12 the the DT 32505 383 13 size size NN 32505 383 14 of of IN 32505 383 15 the the DT 32505 383 16 eye eye NN 32505 383 17 ) ) -RRB- 32505 383 18 in in IN 32505 383 19 males male NNS 32505 383 20 from from IN 32505 383 21 Tilarán Tilarán NNP 32505 383 22 , , , 32505 383 23 Guanacaste Guanacaste NNP 32505 383 24 Province Province NNP 32505 383 25 ; ; : 32505 383 26 the the DT 32505 383 27 tympanum tympanum NN 32505 383 28 is be VBZ 32505 383 29 nearly nearly RB 32505 383 30 as as RB 32505 383 31 large large JJ 32505 383 32 in in IN 32505 383 33 males male NNS 32505 383 34 from from IN 32505 383 35 Piedras Piedras NNP 32505 383 36 Blancas Blancas NNP 32505 383 37 , , , 32505 383 38 Puntarenas Puntarenas NNP 32505 383 39 Province Province NNP 32505 383 40 , , , 32505 383 41 and and CC 32505 383 42 Puerto Puerto NNP 32505 383 43 Viejo Viejo NNP 32505 383 44 , , , 32505 383 45 Heredia Heredia NNP 32505 383 46 Province Province NNP 32505 383 47 , , , 32505 383 48 Costa Costa NNP 32505 383 49 Rica Rica NNP 32505 383 50 . . . 32505 384 1 The the DT 32505 384 2 lowest low JJS 32505 384 3 ratios ratio NNS 32505 384 4 occur occur VBP 32505 384 5 in in IN 32505 384 6 individuals individual NNS 32505 384 7 from from IN 32505 384 8 Almirante Almirante NNP 32505 384 9 , , , 32505 384 10 Bocas Bocas NNP 32505 384 11 del del NNP 32505 384 12 Toro Toro NNP 32505 384 13 , , , 32505 384 14 Panamá Panamá NNP 32505 384 15 , , , 32505 384 16 in in IN 32505 384 17 specimens specimen NNS 32505 384 18 from from IN 32505 384 19 the the DT 32505 384 20 Caribbean caribbean JJ 32505 384 21 lowlands lowland NNS 32505 384 22 of of IN 32505 384 23 Costa Costa NNP 32505 384 24 Rica Rica NNP 32505 384 25 ( ( -LRB- 32505 384 26 except except IN 32505 384 27 Puerto Puerto NNP 32505 384 28 Viejo Viejo NNP 32505 384 29 ) ) -RRB- 32505 384 30 , , , 32505 384 31 and and CC 32505 384 32 in in IN 32505 384 33 those those DT 32505 384 34 from from IN 32505 384 35 El El NNP 32505 384 36 Recreo Recreo NNP 32505 384 37 , , , 32505 384 38 Zelaya Zelaya NNP 32505 384 39 Province Province NNP 32505 384 40 , , , 32505 384 41 Nicaragua Nicaragua NNP 32505 384 42 . . . 32505 385 1 In in IN 32505 385 2 general general JJ 32505 385 3 , , , 32505 385 4 the the DT 32505 385 5 tympanum tympanum NN 32505 385 6 is be VBZ 32505 385 7 proportionately proportionately RB 32505 385 8 larger large JJR 32505 385 9 in in IN 32505 385 10 females female NNS 32505 385 11 than than IN 32505 385 12 in in IN 32505 385 13 males male NNS 32505 385 14 ; ; : 32505 385 15 the the DT 32505 385 16 tympanum tympanum NN 32505 385 17 is be VBZ 32505 385 18 largest large JJS 32505 385 19 in in IN 32505 385 20 females female NNS 32505 385 21 from from IN 32505 385 22 the the DT 32505 385 23 Pacific Pacific NNP 32505 385 24 lowlands lowland NNS 32505 385 25 of of IN 32505 385 26 Costa Costa NNP 32505 385 27 Rica Rica NNP 32505 385 28 ( ( -LRB- 32505 385 29 Table Table NNP 32505 385 30 5 5 CD 32505 385 31 ) ) -RRB- 32505 385 32 . . . 32505 386 1 Color color NN 32505 386 2 variation variation NN 32505 386 3 has have VBZ 32505 386 4 been be VBN 32505 386 5 observed observe VBN 32505 386 6 in in IN 32505 386 7 individuals individual NNS 32505 386 8 from from IN 32505 386 9 the the DT 32505 386 10 same same JJ 32505 386 11 population population NN 32505 386 12 , , , 32505 386 13 as as RB 32505 386 14 well well RB 32505 386 15 as as IN 32505 386 16 in in IN 32505 386 17 individuals individual NNS 32505 386 18 from from IN 32505 386 19 different different JJ 32505 386 20 localities locality NNS 32505 386 21 , , , 32505 386 22 between between IN 32505 386 23 males male NNS 32505 386 24 and and CC 32505 386 25 females female NNS 32505 386 26 , , , 32505 386 27 and and CC 32505 386 28 from from IN 32505 386 29 night night NN 32505 386 30 to to IN 32505 386 31 day day NN 32505 386 32 . . . 32505 387 1 In in IN 32505 387 2 life life NN 32505 387 3 , , , 32505 387 4 most most JJS 32505 387 5 individuals individual NNS 32505 387 6 from from IN 32505 387 7 the the DT 32505 387 8 Pacific Pacific NNP 32505 387 9 lowlands lowland NNS 32505 387 10 of of IN 32505 387 11 Costa Costa NNP 32505 387 12 Rica Rica NNP 32505 387 13 are be VBP 32505 387 14 dark dark JJ 32505 387 15 tan tan NNP 32505 387 16 to to IN 32505 387 17 greenish greenish NNP 32505 387 18 gray gray NNP 32505 387 19 above above RB 32505 387 20 with with IN 32505 387 21 dark dark JJ 32505 387 22 brown brown JJ 32505 387 23 longitudinal longitudinal JJ 32505 387 24 stripes stripe NNS 32505 387 25 that that WDT 32505 387 26 are be VBP 32505 387 27 entire entire JJ 32505 387 28 or or CC 32505 387 29 broken break VBN 32505 387 30 , , , 32505 387 31 but but CC 32505 387 32 some some DT 32505 387 33 specimens specimen NNS 32505 387 34 ( ( -LRB- 32505 387 35 mostly mostly RB 32505 387 36 males male NNS 32505 387 37 ) ) -RRB- 32505 387 38 are be VBP 32505 387 39 dusky dusky NNP 32505 387 40 brown brown NNP 32505 387 41 and and CC 32505 387 42 lack lack NN 32505 387 43 longitudinal longitudinal JJ 32505 387 44 stripes stripe NNS 32505 387 45 or or CC 32505 387 46 an an DT 32505 387 47 interorbital interorbital JJ 32505 387 48 triangle triangle NN 32505 387 49 ; ; : 32505 387 50 females female NNS 32505 387 51 usually usually RB 32505 387 52 have have VBP 32505 387 53 the the DT 32505 387 54 dark dark JJ 32505 387 55 interorbital interorbital NN 32505 387 56 triangle triangle NN 32505 387 57 and and CC 32505 387 58 the the DT 32505 387 59 stripes stripe NNS 32505 387 60 on on IN 32505 387 61 the the DT 32505 387 62 dorsum dorsum NN 32505 387 63 . . . 32505 388 1 Individuals individual NNS 32505 388 2 from from IN 32505 388 3 Turrialba Turrialba NNP 32505 388 4 , , , 32505 388 5 Cartago Cartago NNP 32505 388 6 Province Province NNP 32505 388 7 , , , 32505 388 8 Costa Costa NNP 32505 388 9 Rica Rica NNP 32505 388 10 , , , 32505 388 11 are be VBP 32505 388 12 pale pale JJ 32505 388 13 olive olive JJ 32505 388 14 - - HYPH 32505 388 15 tan tan NN 32505 388 16 with with IN 32505 388 17 olive olive JJ 32505 388 18 - - HYPH 32505 388 19 brown brown JJ 32505 388 20 markings marking NNS 32505 388 21 . . . 32505 389 1 Individuals individual NNS 32505 389 2 from from IN 32505 389 3 Puerto Puerto NNP 32505 389 4 Viejo Viejo NNP 32505 389 5 , , , 32505 389 6 Heredia Heredia NNP 32505 389 7 Province Province NNP 32505 389 8 , , , 32505 389 9 Costa Costa NNP 32505 389 10 Rica Rica NNP 32505 389 11 , , , 32505 389 12 are be VBP 32505 389 13 uniformly uniformly RB 32505 389 14 yellowish yellowish JJ 32505 389 15 brown brown JJ 32505 389 16 with with IN 32505 389 17 or or CC 32505 389 18 without without IN 32505 389 19 dark dark JJ 32505 389 20 longitudinal longitudinal JJ 32505 389 21 stripes stripe NNS 32505 389 22 . . . 32505 390 1 Specimens specimen NNS 32505 390 2 from from IN 32505 390 3 El El NNP 32505 390 4 Recreo Recreo NNP 32505 390 5 , , , 32505 390 6 Zelaya Zelaya NNP 32505 390 7 Province Province NNP 32505 390 8 , , , 32505 390 9 Nicaragua Nicaragua NNP 32505 390 10 , , , 32505 390 11 are be VBP 32505 390 12 like like IN 32505 390 13 those those DT 32505 390 14 from from IN 32505 390 15 Puerto Puerto NNP 32505 390 16 Viejo Viejo NNP 32505 390 17 . . . 32505 391 1 Males male NNS 32505 391 2 from from IN 32505 391 3 Almirante Almirante NNP 32505 391 4 , , , 32505 391 5 Bocas Bocas NNP 32505 391 6 del del NNP 32505 391 7 Toro Toro NNP 32505 391 8 , , , 32505 391 9 Panamá Panamá NNP 32505 391 10 , , , 32505 391 11 are be VBP 32505 391 12 pale pale JJ 32505 391 13 brown brown JJ 32505 391 14 with with IN 32505 391 15 dark dark JJ 32505 391 16 brown brown JJ 32505 391 17 longitudinal longitudinal JJ 32505 391 18 stripes stripe NNS 32505 391 19 and and CC 32505 391 20 an an DT 32505 391 21 indistinct indistinct JJ 32505 391 22 interorbital interorbital NN 32505 391 23 triangle triangle NN 32505 391 24 . . . 32505 392 1 Females female NNS 32505 392 2 have have VBP 32505 392 3 a a DT 32505 392 4 distinct distinct JJ 32505 392 5 interorbital interorbital NN 32505 392 6 triangle triangle NN 32505 392 7 and and CC 32505 392 8 dark dark JJ 32505 392 9 brown brown JJ 32505 392 10 blotches blotch NNS 32505 392 11 on on IN 32505 392 12 the the DT 32505 392 13 thighs thigh NNS 32505 392 14 and and CC 32505 392 15 shanks shank NNS 32505 392 16 . . . 32505 393 1 By by IN 32505 393 2 night night NN 32505 393 3 , , , 32505 393 4 the the DT 32505 393 5 dorsum dorsum NN 32505 393 6 usually usually RB 32505 393 7 is be VBZ 32505 393 8 pale pale JJ 32505 393 9 yellow yellow JJ 32505 393 10 , , , 32505 393 11 and and CC 32505 393 12 the the DT 32505 393 13 belly belly NN 32505 393 14 is be VBZ 32505 393 15 creamy creamy JJ 32505 393 16 white white JJ 32505 393 17 . . . 32505 394 1 By by IN 32505 394 2 day day NN 32505 394 3 , , , 32505 394 4 the the DT 32505 394 5 dorsum dorsum NN 32505 394 6 is be VBZ 32505 394 7 dark dark JJ 32505 394 8 tan tan NNP 32505 394 9 ; ; : 32505 394 10 the the DT 32505 394 11 stripes stripe NNS 32505 394 12 and and CC 32505 394 13 spots spot NNS 32505 394 14 are be VBP 32505 394 15 darker dark JJR 32505 394 16 , , , 32505 394 17 and and CC 32505 394 18 the the DT 32505 394 19 belly belly NN 32505 394 20 is be VBZ 32505 394 21 yellowish yellowish JJ 32505 394 22 white white JJ 32505 394 23 . . . 32505 395 1 Taylor Taylor NNP 32505 395 2 ( ( -LRB- 32505 395 3 1952 1952 CD 32505 395 4 ) ) -RRB- 32505 395 5 noticed notice VBD 32505 395 6 that that IN 32505 395 7 considerable considerable JJ 32505 395 8 variation variation NN 32505 395 9 in in IN 32505 395 10 color color NN 32505 395 11 pattern pattern NN 32505 395 12 occurred occur VBD 32505 395 13 from from IN 32505 395 14 night night NN 32505 395 15 to to IN 32505 395 16 day day NN 32505 395 17 in in IN 32505 395 18 individuals individual NNS 32505 395 19 from from IN 32505 395 20 Turrialba Turrialba NNP 32505 395 21 , , , 32505 395 22 Cartago Cartago NNP 32505 395 23 Province Province NNP 32505 395 24 , , , 32505 395 25 Costa Costa NNP 32505 395 26 Rica Rica NNP 32505 395 27 . . . 32505 396 1 At at IN 32505 396 2 night night NN 32505 396 3 some some DT 32505 396 4 individuals individual NNS 32505 396 5 lacked lack VBD 32505 396 6 a a DT 32505 396 7 dorsal dorsal NN 32505 396 8 pattern pattern NN 32505 396 9 , , , 32505 396 10 but but CC 32505 396 11 by by IN 32505 396 12 day day NN 32505 396 13 many many JJ 32505 396 14 of of IN 32505 396 15 these these DT 32505 396 16 individuals individual NNS 32505 396 17 developed develop VBD 32505 396 18 dorsal dorsal NN 32505 396 19 stripes stripe NNS 32505 396 20 . . . 32505 397 1 _ _ NNP 32505 397 2 Cranial Cranial NNP 32505 397 3 Osteology._--The Osteology._--The NNP 32505 397 4 skull skull NN 32505 397 5 of of IN 32505 397 6 _ _ NNP 32505 397 7 Hyla Hyla NNP 32505 397 8 elaeochroa elaeochroa NN 32505 397 9 _ _ NNP 32505 397 10 is be VBZ 32505 397 11 slightly slightly RB 32505 397 12 wider wide JJR 32505 397 13 than than IN 32505 397 14 it -PRON- PRP 32505 397 15 is be VBZ 32505 397 16 long long JJ 32505 397 17 , , , 32505 397 18 and and CC 32505 397 19 flat flat JJ 32505 397 20 . . . 32505 398 1 The the DT 32505 398 2 premaxillary premaxillary NN 32505 398 3 is be VBZ 32505 398 4 small small JJ 32505 398 5 and and CC 32505 398 6 bears bear VBZ 32505 398 7 10 10 CD 32505 398 8 to to TO 32505 398 9 15 15 CD 32505 398 10 teeth tooth NNS 32505 398 11 ( ( -LRB- 32505 398 12 mean mean VB 32505 398 13 for for IN 32505 398 14 9 9 CD 32505 398 15 specimens specimen NNS 32505 398 16 , , , 32505 398 17 12.3 12.3 CD 32505 398 18 ) ) -RRB- 32505 398 19 . . . 32505 399 1 The the DT 32505 399 2 alary alary JJ 32505 399 3 process process NN 32505 399 4 of of IN 32505 399 5 the the DT 32505 399 6 premaxillary premaxillary NN 32505 399 7 is be VBZ 32505 399 8 small small JJ 32505 399 9 , , , 32505 399 10 vertical vertical JJ 32505 399 11 , , , 32505 399 12 and and CC 32505 399 13 slightly slightly RB 32505 399 14 concave concave VBP 32505 399 15 posteriorly posteriorly RB 32505 399 16 . . . 32505 400 1 Ventrally ventrally RB 32505 400 2 , , , 32505 400 3 the the DT 32505 400 4 premaxillary premaxillary NN 32505 400 5 is be VBZ 32505 400 6 partially partially RB 32505 400 7 united united JJ 32505 400 8 to to IN 32505 400 9 the the DT 32505 400 10 prevomer prevomer NN 32505 400 11 by by IN 32505 400 12 ossification ossification NN 32505 400 13 . . . 32505 401 1 The the DT 32505 401 2 maxillary maxillary NN 32505 401 3 is be VBZ 32505 401 4 slender slend JJR 32505 401 5 and and CC 32505 401 6 bears bear VBZ 32505 401 7 70 70 CD 32505 401 8 to to TO 32505 401 9 82 82 CD 32505 401 10 teeth tooth NNS 32505 401 11 ( ( -LRB- 32505 401 12 mean mean VB 32505 401 13 for for IN 32505 401 14 9 9 CD 32505 401 15 specimens specimen NNS 32505 401 16 , , , 32505 401 17 74.3 74.3 CD 32505 401 18 ) ) -RRB- 32505 401 19 . . . 32505 402 1 The the DT 32505 402 2 pars par NNS 32505 402 3 facialis faciali NNS 32505 402 4 of of IN 32505 402 5 the the DT 32505 402 6 maxillary maxillary NN 32505 402 7 is be VBZ 32505 402 8 laterally laterally RB 32505 402 9 convex convex NNP 32505 402 10 and and CC 32505 402 11 is be VBZ 32505 402 12 about about RB 32505 402 13 twice twice PDT 32505 402 14 as as RB 32505 402 15 high high JJ 32505 402 16 as as IN 32505 402 17 the the DT 32505 402 18 pars par NNS 32505 402 19 dentalis dentalis NNP 32505 402 20 . . . 32505 403 1 The the DT 32505 403 2 nasal nasal NN 32505 403 3 is be VBZ 32505 403 4 large large JJ 32505 403 5 , , , 32505 403 6 robust robust JJ 32505 403 7 , , , 32505 403 8 anteriorly anteriorly RB 32505 403 9 truncate truncate JJ 32505 403 10 , , , 32505 403 11 but but CC 32505 403 12 pointed point VBD 32505 403 13 posteriorly posteriorly RB 32505 403 14 in in IN 32505 403 15 dorsal dorsal NN 32505 403 16 view view NN 32505 403 17 . . . 32505 404 1 The the DT 32505 404 2 nasal nasal NN 32505 404 3 comprises comprise VBZ 32505 404 4 about about IN 32505 404 5 45 45 CD 32505 404 6 per per NN 32505 404 7 cent cent NN 32505 404 8 of of IN 32505 404 9 the the DT 32505 404 10 total total JJ 32505 404 11 length length NN 32505 404 12 of of IN 32505 404 13 the the DT 32505 404 14 skull skull NN 32505 404 15 . . . 32505 405 1 There there EX 32505 405 2 is be VBZ 32505 405 3 an an DT 32505 405 4 anterior anterior JJ 32505 405 5 cartilaginous cartilaginous JJ 32505 405 6 septum septum NN 32505 405 7 nasi nasi NNP 32505 405 8 separating separate VBG 32505 405 9 the the DT 32505 405 10 two two CD 32505 405 11 nasals nasal NNS 32505 405 12 ; ; : 32505 405 13 the the DT 32505 405 14 latter latter JJ 32505 405 15 overlap overlap VBP 32505 405 16 the the DT 32505 405 17 sphenethmoid sphenethmoid NN 32505 405 18 posteriorly posteriorly RB 32505 405 19 . . . 32505 406 1 Each each DT 32505 406 2 nasal nasal NN 32505 406 3 bears bear VBZ 32505 406 4 a a DT 32505 406 5 shallow shallow JJ 32505 406 6 concavity concavity NN 32505 406 7 in in IN 32505 406 8 the the DT 32505 406 9 midlateral midlateral JJ 32505 406 10 side side NN 32505 406 11 and and CC 32505 406 12 lacks lack VBZ 32505 406 13 a a DT 32505 406 14 maxillary maxillary JJ 32505 406 15 process process NN 32505 406 16 . . . 32505 407 1 Dorsally dorsally RB 32505 407 2 , , , 32505 407 3 the the DT 32505 407 4 sphenethmoid sphenethmoid NN 32505 407 5 is be VBZ 32505 407 6 wider wide JJR 32505 407 7 than than IN 32505 407 8 long long JJ 32505 407 9 , , , 32505 407 10 roughly roughly RB 32505 407 11 pentagonal pentagonal JJ 32505 407 12 in in IN 32505 407 13 shape shape NN 32505 407 14 ; ; : 32505 407 15 the the DT 32505 407 16 frontoparietal frontoparietal NN 32505 407 17 is be VBZ 32505 407 18 elongate elongate NN 32505 407 19 , , , 32505 407 20 smooth smooth JJ 32505 407 21 , , , 32505 407 22 and and CC 32505 407 23 bears bear VBZ 32505 407 24 a a DT 32505 407 25 small small JJ 32505 407 26 anterior anterior JJ 32505 407 27 supraorbital supraorbital NN 32505 407 28 process process NN 32505 407 29 . . . 32505 408 1 The the DT 32505 408 2 sphenethmoid sphenethmoid JJ 32505 408 3 and and CC 32505 408 4 frontoparietal frontoparietal JJ 32505 408 5 form form NN 32505 408 6 the the DT 32505 408 7 anterior anterior JJ 32505 408 8 margin margin NN 32505 408 9 of of IN 32505 408 10 the the DT 32505 408 11 frontoparietal frontoparietal JJ 32505 408 12 fontanelle fontanelle NNP 32505 408 13 ; ; : 32505 408 14 the the DT 32505 408 15 fontanelle fontanelle NN 32505 408 16 is be VBZ 32505 408 17 narrow narrow JJ 32505 408 18 anteriorly anteriorly RB 32505 408 19 and and CC 32505 408 20 wider wider RBR 32505 408 21 posteriorly posteriorly RB 32505 408 22 ( ( -LRB- 32505 408 23 Fig fig NN 32505 408 24 . . . 32505 409 1 5B 5b JJ 32505 409 2 ) ) -RRB- 32505 409 3 . . . 32505 410 1 The the DT 32505 410 2 entire entire JJ 32505 410 3 distal distal JJ 32505 410 4 surface surface NN 32505 410 5 of of IN 32505 410 6 the the DT 32505 410 7 proötic proötic NN 32505 410 8 is be VBZ 32505 410 9 in in IN 32505 410 10 contact contact NN 32505 410 11 with with IN 32505 410 12 the the DT 32505 410 13 posterior posterior NN 32505 410 14 arm arm NN 32505 410 15 of of IN 32505 410 16 the the DT 32505 410 17 squamosal squamosal NN 32505 410 18 . . . 32505 411 1 A a DT 32505 411 2 narrow narrow JJ 32505 411 3 cartilaginous cartilaginous JJ 32505 411 4 crista crista NN 32505 411 5 parotica parotica NN 32505 411 6 is be VBZ 32505 411 7 visible visible JJ 32505 411 8 dorsally dorsally RB 32505 411 9 in in IN 32505 411 10 some some DT 32505 411 11 specimens specimen NNS 32505 411 12 . . . 32505 412 1 The the DT 32505 412 2 squamosal squamosal NN 32505 412 3 is be VBZ 32505 412 4 broad broad JJ 32505 412 5 posteriorly posteriorly RB 32505 412 6 but but CC 32505 412 7 its -PRON- PRP$ 32505 412 8 anterior anterior JJ 32505 412 9 arm arm NN 32505 412 10 is be VBZ 32505 412 11 slender slend JJR 32505 412 12 and and CC 32505 412 13 not not RB 32505 412 14 in in IN 32505 412 15 contact contact NN 32505 412 16 with with IN 32505 412 17 the the DT 32505 412 18 maxillary maxillary NN 32505 412 19 . . . 32505 413 1 TABLE TABLE NNP 32505 413 2 5.--Geographic 5.--Geographic NNP 32505 413 3 Variation Variation NNP 32505 413 4 in in IN 32505 413 5 Size Size NNP 32505 413 6 and and CC 32505 413 7 Proportions Proportions NNP 32505 413 8 in in IN 32505 413 9 Males Males NNPS 32505 413 10 of of IN 32505 413 11 _ _ NNP 32505 413 12 Hyla Hyla NNP 32505 413 13 elaeochroa elaeochroa NN 32505 413 14 _ _ NNP 32505 413 15 . . . 32505 414 1 ( ( -LRB- 32505 414 2 Means mean NNS 32505 414 3 in in IN 32505 414 4 parentheses parenthesis NNS 32505 414 5 below below IN 32505 414 6 observed observed JJ 32505 414 7 ranges range NNS 32505 414 8 . . . 32505 414 9 ) ) -RRB- 32505 415 1 = = NFP 32505 415 2 = = NFP 32505 415 3 = = SYM 32505 415 4 = = SYM 32505 415 5 = = SYM 32505 415 6 = = SYM 32505 415 7 = = SYM 32505 415 8 = = SYM 32505 415 9 = = SYM 32505 415 10 = = SYM 32505 415 11 = = SYM 32505 415 12 = = SYM 32505 415 13 = = SYM 32505 415 14 = = SYM 32505 415 15 = = SYM 32505 415 16 = = SYM 32505 415 17 = = SYM 32505 415 18 = = SYM 32505 415 19 = = SYM 32505 415 20 = = SYM 32505 415 21 = = SYM 32505 415 22 = = SYM 32505 415 23 = = SYM 32505 415 24 = = SYM 32505 415 25 = = SYM 32505 415 26 = = SYM 32505 415 27 = = SYM 32505 415 28 = = SYM 32505 415 29 = = SYM 32505 415 30 = = SYM 32505 415 31 = = SYM 32505 415 32 = = SYM 32505 415 33 = = SYM 32505 415 34 = = SYM 32505 415 35 = = SYM 32505 415 36 = = SYM 32505 415 37 = = SYM 32505 415 38 = = SYM 32505 415 39 = = SYM 32505 415 40 = = SYM 32505 415 41 = = SYM 32505 415 42 = = SYM 32505 415 43 = = SYM 32505 415 44 = = SYM 32505 415 45 = = SYM 32505 415 46 = = SYM 32505 415 47 = = SYM 32505 415 48 = = SYM 32505 415 49 = = SYM 32505 415 50 = = SYM 32505 415 51 = = SYM 32505 415 52 = = SYM 32505 415 53 = = SYM 32505 415 54 = = SYM 32505 415 55 = = SYM 32505 415 56 = = SYM 32505 415 57 = = SYM 32505 415 58 = = SYM 32505 415 59 = = SYM 32505 415 60 = = SYM 32505 415 61 = = SYM 32505 415 62 = = SYM 32505 415 63 = = SYM 32505 415 64 = = SYM 32505 415 65 = = SYM 32505 415 66 = = SYM 32505 415 67 = = SYM 32505 415 68 = = SYM 32505 415 69 = = SYM 32505 415 70 = = SYM 32505 415 71 = = SYM 32505 415 72 = = SYM 32505 415 73 = = SYM 32505 415 74 = = SYM 32505 415 75 | | CD 32505 415 76 |Snout |snout NN 32505 415 77 - - HYPH 32505 415 78 vent| vent| CD 32505 415 79 Tibia Tibia NNP 32505 415 80 | | NNP 32505 415 81 | | NNP 32505 415 82 | | CD 32505 415 83 | | CD 32505 415 84 length length NN 32505 415 85 | | NNP 32505 415 86 length/ length/ NN 32505 415 87 |Tympanum/|Foot |tympanum/|foot NN 32505 415 88 length/ length/ NN 32505 415 89 Locality Locality NNP 32505 415 90 | | NNP 32505 415 91 N N NNP 32505 415 92 | | NNP 32505 415 93 ( ( -LRB- 32505 415 94 mm mm NNP 32505 415 95 . . . 32505 415 96 ) ) -RRB- 32505 416 1 | | CD 32505 416 2 snout snout NN 32505 416 3 - - HYPH 32505 416 4 vent vent NN 32505 416 5 | | NNP 32505 416 6 eye eye NN 32505 416 7 | | HYPH 32505 416 8 snout snout NN 32505 416 9 - - HYPH 32505 416 10 vent vent NN 32505 416 11 ---------------------+----+----------+------------+---------+------------- ---------------------+----+----------+------------+---------+------------- NN 32505 416 12 Nicaragua Nicaragua NNP 32505 416 13 : : : 32505 416 14 El El NNP 32505 416 15 Recreo Recreo NNP 32505 416 16 | | NNP 32505 416 17 9 9 CD 32505 416 18 | | CD 32505 416 19 28.0 28.0 CD 32505 416 20 - - HYPH 32505 416 21 30.3| 30.3| CD 32505 416 22 0.51 0.51 CD 32505 416 23 - - HYPH 32505 416 24 0.57 0.57 CD 32505 416 25 |0.47 |0.47 CD 32505 416 26 - - HYPH 32505 416 27 0.59| 0.59| CD 32505 416 28 0.39 0.39 CD 32505 416 29 - - HYPH 32505 416 30 0.54 0.54 CD 32505 416 31 | | HYPH 32505 416 32 | | CD 32505 416 33 ( ( -LRB- 32505 416 34 29.3 29.3 CD 32505 416 35 ) ) -RRB- 32505 416 36 | | NNP 32505 416 37 ( ( -LRB- 32505 416 38 0.55 0.55 CD 32505 416 39 ) ) -RRB- 32505 416 40 | | NNP 32505 416 41 ( ( -LRB- 32505 416 42 0.51 0.51 CD 32505 416 43 ) ) -RRB- 32505 416 44 | | NNP 32505 416 45 ( ( -LRB- 32505 416 46 0.41 0.41 CD 32505 416 47 ) ) -RRB- 32505 416 48 | | NNP 32505 416 49 | | CD 32505 416 50 | | CD 32505 416 51 | | CD 32505 416 52 | | NNP 32505 416 53 Costa Costa NNP 32505 416 54 Rica Rica NNP 32505 416 55 : : : 32505 416 56 Tilarán Tilarán NNP 32505 416 57 | | NNP 32505 416 58 21 21 CD 32505 416 59 | | CD 32505 416 60 28.8 28.8 CD 32505 416 61 - - HYPH 32505 416 62 33.6| 33.6| CD 32505 416 63 0.47 0.47 CD 32505 416 64 - - HYPH 32505 416 65 0.57 0.57 CD 32505 416 66 |0.48 |0.48 CD 32505 416 67 - - HYPH 32505 416 68 0.65| 0.65| CD 32505 416 69 0.40 0.40 CD 32505 416 70 - - HYPH 32505 416 71 0.46 0.46 CD 32505 416 72 | | CD 32505 416 73 | | CD 32505 416 74 ( ( -LRB- 32505 416 75 30.6 30.6 CD 32505 416 76 ) ) -RRB- 32505 416 77 | | NNP 32505 416 78 ( ( -LRB- 32505 416 79 0.52 0.52 CD 32505 416 80 ) ) -RRB- 32505 416 81 | | NNP 32505 416 82 ( ( -LRB- 32505 416 83 0.59 0.59 CD 32505 416 84 ) ) -RRB- 32505 416 85 | | NNP 32505 416 86 ( ( -LRB- 32505 416 87 0.41 0.41 CD 32505 416 88 ) ) -RRB- 32505 416 89 | | NNP 32505 416 90 | | CD 32505 416 91 | | CD 32505 416 92 | | CD 32505 416 93 | | NNP 32505 416 94 Costa Costa NNP 32505 416 95 Rica Rica NNP 32505 416 96 : : : 32505 416 97 Puerto Puerto NNP 32505 416 98 | | CD 32505 416 99 22 22 CD 32505 416 100 | | CD 32505 416 101 26.3 26.3 CD 32505 416 102 - - HYPH 32505 416 103 32.4| 32.4| CD 32505 416 104 0.49 0.49 CD 32505 416 105 - - HYPH 32505 416 106 0.54 0.54 CD 32505 416 107 |0.48 |0.48 CD 32505 416 108 - - HYPH 32505 416 109 0.65| 0.65| CD 32505 416 110 0.38 0.38 CD 32505 416 111 - - HYPH 32505 416 112 0.45 0.45 CD 32505 416 113 Viejo Viejo NNP 32505 416 114 | | NNP 32505 416 115 | | NNP 32505 416 116 ( ( -LRB- 32505 416 117 29.7 29.7 CD 32505 416 118 ) ) -RRB- 32505 416 119 | | NNP 32505 416 120 ( ( -LRB- 32505 416 121 0.52 0.52 CD 32505 416 122 ) ) -RRB- 32505 416 123 | | NNP 32505 416 124 ( ( -LRB- 32505 416 125 0.57 0.57 CD 32505 416 126 ) ) -RRB- 32505 416 127 | | NNP 32505 416 128 ( ( -LRB- 32505 416 129 0.42 0.42 CD 32505 416 130 ) ) -RRB- 32505 416 131 | | CD 32505 416 132 | | CD 32505 416 133 | | CD 32505 416 134 | | CD 32505 416 135 | | NNP 32505 416 136 Costa Costa NNP 32505 416 137 Rica Rica NNP 32505 416 138 : : : 32505 416 139 Turrialba| Turrialba| NNP 32505 416 140 95 95 CD 32505 416 141 | | CD 32505 416 142 28.1 28.1 CD 32505 416 143 - - HYPH 32505 416 144 35.0| 35.0| CD 32505 416 145 0.47 0.47 CD 32505 416 146 - - HYPH 32505 416 147 0.56 0.56 CD 32505 416 148 |0.47 |0.47 CD 32505 416 149 - - HYPH 32505 416 150 0.68| 0.68| CD 32505 416 151 0.37 0.37 CD 32505 416 152 - - HYPH 32505 416 153 0.46 0.46 CD 32505 416 154 | | CD 32505 416 155 | | CD 32505 416 156 ( ( -LRB- 32505 416 157 30.6 30.6 CD 32505 416 158 ) ) -RRB- 32505 416 159 | | NNP 32505 416 160 ( ( -LRB- 32505 416 161 0.51 0.51 CD 32505 416 162 ) ) -RRB- 32505 416 163 | | NNP 32505 416 164 ( ( -LRB- 32505 416 165 0.56 0.56 CD 32505 416 166 ) ) -RRB- 32505 416 167 | | NNP 32505 416 168 ( ( -LRB- 32505 416 169 0.41 0.41 CD 32505 416 170 ) ) -RRB- 32505 416 171 | | NNP 32505 416 172 | | CD 32505 416 173 | | CD 32505 416 174 | | CD 32505 416 175 | | NNP 32505 416 176 Costa Costa NNP 32505 416 177 Rica Rica NNP 32505 416 178 : : : 32505 416 179 Bataán Bataán NNP 32505 416 180 , , , 32505 416 181 | | NNP 32505 416 182 26 26 CD 32505 416 183 | | CD 32505 416 184 26.3 26.3 CD 32505 416 185 - - HYPH 32505 416 186 32.7| 32.7| CD 32505 416 187 0.47 0.47 CD 32505 416 188 - - HYPH 32505 416 189 0.54 0.54 CD 32505 416 190 |0.45 |0.45 NNS 32505 416 191 - - : 32505 416 192 0.66| 0.66| CD 32505 416 193 0.36 0.36 CD 32505 416 194 - - HYPH 32505 416 195 0.44 0.44 CD 32505 416 196 Limón Limón NNP 32505 416 197 , , , 32505 416 198 and and CC 32505 416 199 Suretka Suretka NNP 32505 416 200 | | NNP 32505 416 201 | | NNP 32505 416 202 ( ( -LRB- 32505 416 203 30.0 30.0 CD 32505 416 204 ) ) -RRB- 32505 416 205 | | NNP 32505 416 206 ( ( -LRB- 32505 416 207 0.51 0.51 CD 32505 416 208 ) ) -RRB- 32505 416 209 | | NNP 32505 416 210 ( ( -LRB- 32505 416 211 0.50 0.50 CD 32505 416 212 ) ) -RRB- 32505 416 213 | | NNP 32505 416 214 ( ( -LRB- 32505 416 215 0.41 0.41 CD 32505 416 216 ) ) -RRB- 32505 416 217 | | NNP 32505 416 218 | | CD 32505 416 219 | | CD 32505 416 220 | | CD 32505 416 221 | | NNP 32505 416 222 Costa Costa NNP 32505 416 223 Rica Rica NNP 32505 416 224 : : : 32505 416 225 Piedras Piedras NNP 32505 416 226 | | NNP 32505 416 227 21 21 CD 32505 416 228 | | CD 32505 416 229 33.3 33.3 CD 32505 416 230 - - HYPH 32505 416 231 37.7| 37.7| CD 32505 416 232 0.50 0.50 CD 32505 416 233 - - HYPH 32505 416 234 0.54 0.54 CD 32505 416 235 |0.48 |0.48 CD 32505 416 236 - - HYPH 32505 416 237 0.64| 0.64| CD 32505 416 238 0.40 0.40 CD 32505 416 239 - - HYPH 32505 416 240 0.46 0.46 CD 32505 416 241 Blancas Blancas NNP 32505 416 242 | | NNP 32505 416 243 | | NNP 32505 416 244 ( ( -LRB- 32505 416 245 35.2 35.2 CD 32505 416 246 ) ) -RRB- 32505 416 247 | | NNP 32505 416 248 ( ( -LRB- 32505 416 249 0.51 0.51 CD 32505 416 250 ) ) -RRB- 32505 416 251 | | NNP 32505 416 252 ( ( -LRB- 32505 416 253 0.57 0.57 CD 32505 416 254 ) ) -RRB- 32505 416 255 | | NNP 32505 416 256 ( ( -LRB- 32505 416 257 0.43 0.43 CD 32505 416 258 ) ) -RRB- 32505 416 259 | | CD 32505 416 260 | | CD 32505 416 261 | | CD 32505 416 262 | | CD 32505 416 263 | | NNP 32505 416 264 Costa Costa NNP 32505 416 265 Rica Rica NNP 32505 416 266 : : : 32505 416 267 Rincón Rincón NNP 32505 416 268 de| de| NNP 32505 416 269 24 24 CD 32505 416 270 | | CD 32505 416 271 31.4 31.4 CD 32505 416 272 - - HYPH 32505 416 273 35.9| 35.9| CD 32505 416 274 0.50 0.50 CD 32505 416 275 - - HYPH 32505 416 276 0.56 0.56 CD 32505 416 277 |0.45 |0.45 : 32505 416 278 - - HYPH 32505 416 279 0.61| 0.61| CD 32505 416 280 0.40 0.40 CD 32505 416 281 - - HYPH 32505 416 282 0.46 0.46 CD 32505 416 283 Osa Osa NNP 32505 416 284 | | NNP 32505 416 285 | | NNP 32505 416 286 ( ( -LRB- 32505 416 287 34.1 34.1 CD 32505 416 288 ) ) -RRB- 32505 416 289 | | NNP 32505 416 290 ( ( -LRB- 32505 416 291 0.53 0.53 CD 32505 416 292 ) ) -RRB- 32505 416 293 | | NNP 32505 416 294 ( ( -LRB- 32505 416 295 0.54 0.54 CD 32505 416 296 ) ) -RRB- 32505 416 297 | | NNP 32505 416 298 ( ( -LRB- 32505 416 299 0.43 0.43 CD 32505 416 300 ) ) -RRB- 32505 416 301 | | CD 32505 416 302 | | CD 32505 416 303 | | CD 32505 416 304 | | CD 32505 416 305 | | NNP 32505 416 306 Panamá Panamá NNP 32505 416 307 : : : 32505 416 308 Bocas Bocas NNP 32505 416 309 del del NNP 32505 416 310 | | NNP 32505 416 311 6 6 CD 32505 416 312 | | CD 32505 416 313 31.0 31.0 CD 32505 416 314 - - HYPH 32505 416 315 33.5| 33.5| CD 32505 416 316 0.49 0.49 CD 32505 416 317 - - HYPH 32505 416 318 0.54 0.54 CD 32505 416 319 |0.47 |0.47 CD 32505 416 320 - - HYPH 32505 416 321 0.50| 0.50| CD 32505 416 322 0.41 0.41 CD 32505 416 323 - - HYPH 32505 416 324 0.43 0.43 CD 32505 416 325 Toro Toro NNP 32505 416 326 | | NNP 32505 416 327 | | NNP 32505 416 328 ( ( -LRB- 32505 416 329 32.1 32.1 CD 32505 416 330 ) ) -RRB- 32505 416 331 | | NNP 32505 416 332 ( ( -LRB- 32505 416 333 0.51 0.51 CD 32505 416 334 ) ) -RRB- 32505 416 335 | | NNP 32505 416 336 ( ( -LRB- 32505 416 337 0.48 0.48 CD 32505 416 338 ) ) -RRB- 32505 416 339 | | NNP 32505 416 340 ( ( -LRB- 32505 416 341 0.42 0.42 CD 32505 416 342 ) ) -RRB- 32505 416 343 [ [ -LRB- 32505 416 344 Illustration illustration NN 32505 416 345 : : : 32505 416 346 PLATE PLATE NNP 32505 416 347 1 1 CD 32505 416 348 Living live VBG 32505 416 349 _ _ NNP 32505 416 350 Hyla Hyla NNP 32505 416 351 _ _ NNP 32505 416 352 : : : 32505 416 353 ( ( -LRB- 32505 416 354 A a NN 32505 416 355 ) ) -RRB- 32505 416 356 _ _ NNP 32505 416 357 H. H. NNP 32505 417 1 boulengeri boulengeri NNP 32505 417 2 _ _ NNP 32505 417 3 ( ( -LRB- 32505 417 4 KU KU NNP 32505 417 5 86322 86322 CD 32505 417 6 ) ) -RRB- 32505 417 7 and and CC 32505 417 8 ( ( -LRB- 32505 417 9 B b NN 32505 417 10 ) ) -RRB- 32505 417 11 _ _ NNP 32505 417 12 H. H. NNP 32505 418 1 foliamorta foliamorta NNP 32505 418 2 _ _ NNP 32505 418 3 ( ( -LRB- 32505 418 4 KU KU NNP 32505 418 5 101576 101576 CD 32505 418 6 ) ) -RRB- 32505 418 7 , , , 32505 418 8 × × NN 32505 418 9 2 2 CD 32505 418 10 . . . 32505 418 11 ] ] -RRB- 32505 419 1 [ [ -LRB- 32505 419 2 Illustration illustration NN 32505 419 3 : : : 32505 419 4 PLATE PLATE NNP 32505 419 5 2 2 CD 32505 419 6 Living live VBG 32505 419 7 _ _ NNP 32505 419 8 Hyla Hyla NNP 32505 419 9 : : : 32505 419 10 _ _ NNP 32505 419 11 ( ( -LRB- 32505 419 12 A A NNP 32505 419 13 ) ) -RRB- 32505 419 14 _ _ NNP 32505 419 15 H. H. NNP 32505 420 1 elaeochroa elaeochroa NNP 32505 420 2 _ _ NNP 32505 420 3 ( ( -LRB- 32505 420 4 KU KU NNP 32505 420 5 91688 91688 CD 32505 420 6 ) ) -RRB- 32505 420 7 , , , 32505 420 8 ( ( -LRB- 32505 420 9 B b NN 32505 420 10 ) ) -RRB- 32505 420 11 _ _ NNP 32505 420 12 H. H. NNP 32505 421 1 staufferi staufferi NNP 32505 421 2 staufferi staufferi NNP 32505 421 3 _ _ NNP 32505 421 4 ( ( -LRB- 32505 421 5 KU KU NNP 32505 421 6 57791 57791 CD 32505 421 7 ) ) -RRB- 32505 421 8 , , , 32505 421 9 and and CC 32505 421 10 ( ( -LRB- 32505 421 11 C C NNP 32505 421 12 ) ) -RRB- 32505 421 13 _ _ NNP 32505 421 14 H. H. NNP 32505 422 1 staufferi staufferi NNP 32505 422 2 altae altae NNP 32505 422 3 _ _ NNP 32505 422 4 ( ( -LRB- 32505 422 5 KU KU NNP 32505 422 6 101688 101688 CD 32505 422 7 ) ) -RRB- 32505 422 8 , , , 32505 422 9 × × NN 32505 422 10 2 2 CD 32505 422 11 . . . 32505 422 12 ] ] -RRB- 32505 423 1 [ [ -LRB- 32505 423 2 Illustration illustration NN 32505 423 3 : : : 32505 423 4 PLATE PLATE NNP 32505 423 5 3 3 CD 32505 423 6 Audiospectrograms audiospectrogram NNS 32505 423 7 and and CC 32505 423 8 sections section NNS 32505 423 9 of of IN 32505 423 10 mating mate VBG 32505 423 11 calls call NNS 32505 423 12 of of IN 32505 423 13 ( ( -LRB- 32505 423 14 A a NN 32505 423 15 ) ) -RRB- 32505 423 16 _ _ NNP 32505 423 17 Hyla Hyla NNP 32505 423 18 boulengeri boulengeri NN 32505 423 19 _ _ NNP 32505 423 20 ( ( -LRB- 32505 423 21 KU KU NNP 32505 423 22 Tape Tape NNP 32505 423 23 No no UH 32505 423 24 . . . 32505 424 1 511 511 CD 32505 424 2 ) ) -RRB- 32505 424 3 and and CC 32505 424 4 ( ( -LRB- 32505 424 5 B b NN 32505 424 6 ) ) -RRB- 32505 424 7 _ _ NNP 32505 424 8 H. H. NNP 32505 425 1 foliamorta foliamorta NNP 32505 425 2 _ _ NNP 32505 425 3 ( ( -LRB- 32505 425 4 KU KU NNP 32505 425 5 Tape Tape NNP 32505 425 6 No no UH 32505 425 7 . . . 32505 426 1 511 511 CD 32505 426 2 ) ) -RRB- 32505 426 3 and and CC 32505 426 4 ( ( -LRB- 32505 426 5 B b NN 32505 426 6 ) ) -RRB- 32505 426 7 _ _ NNP 32505 426 8 H. H. NNP 32505 427 1 foliamorta foliamorta NNP 32505 427 2 _ _ NNP 32505 427 3 ( ( -LRB- 32505 427 4 KU KU NNP 32505 427 5 Tape Tape NNP 32505 427 6 No no UH 32505 427 7 . . . 32505 428 1 288 288 CD 32505 428 2 ) ) -RRB- 32505 428 3 . . . 32505 428 4 ] ] -RRB- 32505 429 1 [ [ -LRB- 32505 429 2 Illustration illustration NN 32505 429 3 : : : 32505 429 4 PLATE PLATE NNP 32505 429 5 4 4 CD 32505 429 6 Audiospectrograms audiospectrogram NNS 32505 429 7 and and CC 32505 429 8 sections section NNS 32505 429 9 of of IN 32505 429 10 mating mate VBG 32505 429 11 calls call NNS 32505 429 12 of of IN 32505 429 13 ( ( -LRB- 32505 429 14 A a NN 32505 429 15 ) ) -RRB- 32505 429 16 _ _ NNP 32505 429 17 Hyla Hyla NNP 32505 429 18 elaeochroa elaeochroa NN 32505 429 19 _ _ NNP 32505 429 20 ( ( -LRB- 32505 429 21 KU KU NNP 32505 429 22 Tape Tape NNP 32505 429 23 No no UH 32505 429 24 . . . 32505 430 1 97 97 CD 32505 430 2 ) ) -RRB- 32505 430 3 , , , 32505 430 4 ( ( -LRB- 32505 430 5 B b NN 32505 430 6 ) ) -RRB- 32505 430 7 _ _ NNP 32505 430 8 H. H. NNP 32505 431 1 s. s. NNP 32505 431 2 staufferi staufferi NNP 32505 431 3 _ _ NNP 32505 431 4 ( ( -LRB- 32505 431 5 KU KU NNP 32505 431 6 Tape Tape NNP 32505 431 7 No no UH 32505 431 8 . . . 32505 432 1 93 93 CD 32505 432 2 ) ) -RRB- 32505 432 3 , , , 32505 432 4 and and CC 32505 432 5 ( ( -LRB- 32505 432 6 C C NNP 32505 432 7 ) ) -RRB- 32505 432 8 _ _ NNP 32505 432 9 H. H. NNP 32505 433 1 staufferi staufferi NNP 32505 433 2 altae altae NNP 32505 433 3 _ _ NNP 32505 433 4 ( ( -LRB- 32505 433 5 KU KU NNP 32505 433 6 Tape Tape NNP 32505 433 7 No no UH 32505 433 8 . . . 32505 434 1 502 502 CD 32505 434 2 ) ) -RRB- 32505 434 3 . . . 32505 434 4 ] ] -RRB- 32505 435 1 The the DT 32505 435 2 prevomer prevomer NN 32505 435 3 is be VBZ 32505 435 4 short short JJ 32505 435 5 , , , 32505 435 6 and and CC 32505 435 7 broadest broad JJS 32505 435 8 anteriorly anteriorly RB 32505 435 9 . . . 32505 436 1 The the DT 32505 436 2 prevomer prevomer NN 32505 436 3 is be VBZ 32505 436 4 joined join VBN 32505 436 5 to to IN 32505 436 6 the the DT 32505 436 7 premaxillary premaxillary NN 32505 436 8 by by IN 32505 436 9 ossification ossification NN 32505 436 10 . . . 32505 437 1 The the DT 32505 437 2 posterior posterior JJ 32505 437 3 margin margin NN 32505 437 4 of of IN 32505 437 5 the the DT 32505 437 6 prevomer prevomer NN 32505 437 7 bears bear VBZ 32505 437 8 a a DT 32505 437 9 narrow narrow JJ 32505 437 10 cartilaginous cartilaginous JJ 32505 437 11 articulation articulation NN 32505 437 12 with with IN 32505 437 13 the the DT 32505 437 14 sphenethmoid sphenethmoid NN 32505 437 15 . . . 32505 438 1 The the DT 32505 438 2 anterolateral anterolateral JJ 32505 438 3 and and CC 32505 438 4 posterolateral posterolateral JJ 32505 438 5 processes process NNS 32505 438 6 of of IN 32505 438 7 the the DT 32505 438 8 prevomer prevomer NN 32505 438 9 form form NN 32505 438 10 an an DT 32505 438 11 incomplete incomplete JJ 32505 438 12 bony bony NN 32505 438 13 margin margin NN 32505 438 14 to to IN 32505 438 15 the the DT 32505 438 16 small small JJ 32505 438 17 choanae choanae NN 32505 438 18 ; ; : 32505 438 19 each each DT 32505 438 20 prevomer prevomer NN 32505 438 21 bears bear VBZ 32505 438 22 four four CD 32505 438 23 to to TO 32505 438 24 seven seven CD 32505 438 25 teeth tooth NNS 32505 438 26 . . . 32505 439 1 The the DT 32505 439 2 palatine palatine NN 32505 439 3 is be VBZ 32505 439 4 small small JJ 32505 439 5 , , , 32505 439 6 curved curve VBN 32505 439 7 anteriorly anteriorly RB 32505 439 8 and and CC 32505 439 9 edentate edentate NN 32505 439 10 . . . 32505 440 1 The the DT 32505 440 2 anterior anterior JJ 32505 440 3 part part NN 32505 440 4 of of IN 32505 440 5 the the DT 32505 440 6 parasphenoid parasphenoid NN 32505 440 7 is be VBZ 32505 440 8 robust robust JJ 32505 440 9 and and CC 32505 440 10 ends end VBZ 32505 440 11 in in IN 32505 440 12 a a DT 32505 440 13 point point NN 32505 440 14 . . . 32505 441 1 The the DT 32505 441 2 pterygoid pterygoid NN 32505 441 3 is be VBZ 32505 441 4 slender slend JJR 32505 441 5 and and CC 32505 441 6 weakly weakly RB 32505 441 7 developed develop VBD 32505 441 8 . . . 32505 442 1 _ _ NNP 32505 442 2 Natural Natural NNP 32505 442 3 History._--_Hyla History._--_Hyla NNP 32505 442 4 elaeochroa elaeochroa NNP 32505 442 5 _ _ NNP 32505 442 6 inhabits inhabit VBZ 32505 442 7 humid humid NN 32505 442 8 lowland lowland NN 32505 442 9 tropical tropical JJ 32505 442 10 forests forest NNS 32505 442 11 in in IN 32505 442 12 lower low JJR 32505 442 13 Central Central NNP 32505 442 14 America America NNP 32505 442 15 and and CC 32505 442 16 breeds breed NNS 32505 442 17 in in IN 32505 442 18 temporary temporary JJ 32505 442 19 ponds pond NNS 32505 442 20 . . . 32505 443 1 Clasping clasp VBG 32505 443 2 pairs pair NNS 32505 443 3 , , , 32505 443 4 gravid gravid JJ 32505 443 5 females female NNS 32505 443 6 , , , 32505 443 7 and and CC 32505 443 8 calling call VBG 32505 443 9 males male NNS 32505 443 10 have have VBP 32505 443 11 been be VBN 32505 443 12 found find VBN 32505 443 13 mostly mostly RB 32505 443 14 in in IN 32505 443 15 June June NNP 32505 443 16 , , , 32505 443 17 July July NNP 32505 443 18 , , , 32505 443 19 and and CC 32505 443 20 August August NNP 32505 443 21 . . . 32505 444 1 William William NNP 32505 444 2 E. E. NNP 32505 444 3 Duellman Duellman NNP 32505 444 4 informed inform VBD 32505 444 5 me -PRON- PRP 32505 444 6 that that IN 32505 444 7 he -PRON- PRP 32505 444 8 also also RB 32505 444 9 found find VBD 32505 444 10 males male NNS 32505 444 11 calling call VBG 32505 444 12 in in IN 32505 444 13 mid mid NNP 32505 444 14 - - NNP 32505 444 15 February February NNP 32505 444 16 , , , 32505 444 17 late late JJ 32505 444 18 April April NNP 32505 444 19 , , , 32505 444 20 and and CC 32505 444 21 May. May NNP 32505 445 1 Duellman Duellman NNP 32505 445 2 ( ( -LRB- 32505 445 3 1967 1967 CD 32505 445 4 ) ) -RRB- 32505 445 5 reported report VBD 32505 445 6 detailed detailed JJ 32505 445 7 observations observation NNS 32505 445 8 of of IN 32505 445 9 the the DT 32505 445 10 social social JJ 32505 445 11 organization organization NN 32505 445 12 in in IN 32505 445 13 the the DT 32505 445 14 mating mating NN 32505 445 15 call call NN 32505 445 16 of of IN 32505 445 17 _ _ NNP 32505 445 18 Hyla Hyla NNP 32505 445 19 elaeochroa elaeochroa NN 32505 445 20 _ _ NNP 32505 445 21 . . . 32505 446 1 The the DT 32505 446 2 choruses chorus NNS 32505 446 3 in in IN 32505 446 4 this this DT 32505 446 5 species specie NNS 32505 446 6 are be VBP 32505 446 7 initially initially RB 32505 446 8 organized organize VBN 32505 446 9 , , , 32505 446 10 but but CC 32505 446 11 when when WRB 32505 446 12 many many JJ 32505 446 13 individuals individual NNS 32505 446 14 call call VBP 32505 446 15 , , , 32505 446 16 the the DT 32505 446 17 chorus chorus NN 32505 446 18 loses lose VBZ 32505 446 19 organization organization NN 32505 446 20 . . . 32505 447 1 I -PRON- PRP 32505 447 2 observed observe VBD 32505 447 3 this this DT 32505 447 4 species specie NNS 32505 447 5 breeding breed VBG 32505 447 6 in in IN 32505 447 7 a a DT 32505 447 8 temporary temporary JJ 32505 447 9 pond pond NN 32505 447 10 at at IN 32505 447 11 Puerto Puerto NNP 32505 447 12 Viejo Viejo NNP 32505 447 13 , , , 32505 447 14 Heredia Heredia NNP 32505 447 15 Province Province NNP 32505 447 16 , , , 32505 447 17 Cost Cost NNP 32505 447 18 Rica Rica NNP 32505 447 19 , , , 32505 447 20 in in IN 32505 447 21 late late JJ 32505 447 22 June June NNP 32505 447 23 . . . 32505 448 1 Calling call VBG 32505 448 2 males male NNS 32505 448 3 and and CC 32505 448 4 clasping clasp VBG 32505 448 5 pairs pair NNS 32505 448 6 were be VBD 32505 448 7 extremely extremely RB 32505 448 8 abundant abundant JJ 32505 448 9 within within IN 32505 448 10 a a DT 32505 448 11 few few JJ 32505 448 12 hours hour NNS 32505 448 13 after after IN 32505 448 14 a a DT 32505 448 15 heavy heavy JJ 32505 448 16 rain rain NN 32505 448 17 . . . 32505 449 1 Males male NNS 32505 449 2 were be VBD 32505 449 3 mostly mostly RB 32505 449 4 found find VBN 32505 449 5 calling call VBG 32505 449 6 from from IN 32505 449 7 low low JJ 32505 449 8 emergent emergent JJ 32505 449 9 herbs herb NNS 32505 449 10 in in IN 32505 449 11 the the DT 32505 449 12 pond pond NN 32505 449 13 and and CC 32505 449 14 less less RBR 32505 449 15 commonly commonly RB 32505 449 16 from from IN 32505 449 17 bushes bush NNS 32505 449 18 and and CC 32505 449 19 trees tree NNS 32505 449 20 to to IN 32505 449 21 heights height NNS 32505 449 22 of of IN 32505 449 23 six six CD 32505 449 24 meters meter NNS 32505 449 25 above above IN 32505 449 26 the the DT 32505 449 27 water water NN 32505 449 28 . . . 32505 450 1 Calling call VBG 32505 450 2 males male NNS 32505 450 3 were be VBD 32505 450 4 also also RB 32505 450 5 observed observe VBN 32505 450 6 at at IN 32505 450 7 Ricón Ricón NNP 32505 450 8 de de NNP 32505 450 9 Osa Osa NNP 32505 450 10 , , , 32505 450 11 Puntarenas Puntarenas NNP 32505 450 12 Province Province NNP 32505 450 13 , , , 32505 450 14 Costa Costa NNP 32505 450 15 Rica Rica NNP 32505 450 16 , , , 32505 450 17 in in IN 32505 450 18 late late JJ 32505 450 19 July July NNP 32505 450 20 . . . 32505 451 1 These these DT 32505 451 2 breeding breeding NN 32505 451 3 individuals individual NNS 32505 451 4 were be VBD 32505 451 5 found find VBN 32505 451 6 in in IN 32505 451 7 a a DT 32505 451 8 shallow shallow JJ 32505 451 9 pond pond NN 32505 451 10 at at IN 32505 451 11 the the DT 32505 451 12 edge edge NN 32505 451 13 of of IN 32505 451 14 a a DT 32505 451 15 wet wet JJ 32505 451 16 forest forest NN 32505 451 17 . . . 32505 452 1 Calling call VBG 32505 452 2 stations station NNS 32505 452 3 were be VBD 32505 452 4 less less JJR 32505 452 5 than than IN 32505 452 6 two two CD 32505 452 7 meters meter NNS 32505 452 8 in in IN 32505 452 9 height height NN 32505 452 10 . . . 32505 453 1 John John NNP 32505 453 2 D. D. NNP 32505 453 3 Lynch Lynch NNP 32505 453 4 informed inform VBD 32505 453 5 me -PRON- PRP 32505 453 6 that that IN 32505 453 7 after after IN 32505 453 8 a a DT 32505 453 9 heavy heavy JJ 32505 453 10 rain rain NN 32505 453 11 in in IN 32505 453 12 early early JJ 32505 453 13 August August NNP 32505 453 14 , , , 32505 453 15 he -PRON- PRP 32505 453 16 found find VBD 32505 453 17 several several JJ 32505 453 18 hundred hundred CD 32505 453 19 individuals individual NNS 32505 453 20 congregated congregate VBN 32505 453 21 in in IN 32505 453 22 a a DT 32505 453 23 small small JJ 32505 453 24 grassy grassy JJ 32505 453 25 pond pond NN 32505 453 26 less less JJR 32505 453 27 than than IN 32505 453 28 a a DT 32505 453 29 foot foot NN 32505 453 30 deep deep RB 32505 453 31 , , , 32505 453 32 at at IN 32505 453 33 Rincón Rincón NNP 32505 453 34 de de NNP 32505 453 35 Osa Osa NNP 32505 453 36 . . . 32505 454 1 Males male NNS 32505 454 2 were be VBD 32505 454 3 calling call VBG 32505 454 4 from from IN 32505 454 5 sites site NNS 32505 454 6 on on IN 32505 454 7 grass grass NN 32505 454 8 stems stem VBZ 32505 454 9 a a DT 32505 454 10 few few JJ 32505 454 11 centimeters centimeter NNS 32505 454 12 above above IN 32505 454 13 the the DT 32505 454 14 water water NN 32505 454 15 . . . 32505 455 1 The the DT 32505 455 2 mating mating NN 32505 455 3 call call NN 32505 455 4 of of IN 32505 455 5 _ _ NNP 32505 455 6 Hyla Hyla NNP 32505 455 7 elaeochroa elaeochroa NN 32505 455 8 _ _ NNP 32505 455 9 consists consist VBZ 32505 455 10 of of IN 32505 455 11 short short JJ 32505 455 12 notes note NNS 32505 455 13 , , , 32505 455 14 repeated repeat VBN 32505 455 15 at at IN 32505 455 16 intervals interval NNS 32505 455 17 of of IN 32505 455 18 about about RB 32505 455 19 0.40 0.40 CD 32505 455 20 second second JJ 32505 455 21 . . . 32505 456 1 Each each DT 32505 456 2 note note NN 32505 456 3 has have VBZ 32505 456 4 a a DT 32505 456 5 duration duration NN 32505 456 6 of of IN 32505 456 7 0.12 0.12 CD 32505 456 8 to to IN 32505 456 9 0.24 0.24 CD 32505 456 10 second second JJ 32505 456 11 . . . 32505 457 1 The the DT 32505 457 2 fundamental fundamental JJ 32505 457 3 frequency frequency NN 32505 457 4 varies vary VBZ 32505 457 5 from from IN 32505 457 6 48 48 CD 32505 457 7 to to TO 32505 457 8 65 65 CD 32505 457 9 cycles cycle NNS 32505 457 10 per per IN 32505 457 11 second second NN 32505 457 12 , , , 32505 457 13 and and CC 32505 457 14 the the DT 32505 457 15 notes note NNS 32505 457 16 have have VBP 32505 457 17 40 40 CD 32505 457 18 - - SYM 32505 457 19 50 50 CD 32505 457 20 pulses pulse NNS 32505 457 21 per per IN 32505 457 22 second second NN 32505 457 23 ; ; : 32505 457 24 the the DT 32505 457 25 dominant dominant JJ 32505 457 26 frequency frequency NN 32505 457 27 is be VBZ 32505 457 28 at at IN 32505 457 29 about about IN 32505 457 30 2,900 2,900 CD 32505 457 31 cycles cycle NNS 32505 457 32 per per IN 32505 457 33 second second JJ 32505 457 34 ( ( -LRB- 32505 457 35 Table table NN 32505 457 36 2 2 CD 32505 457 37 , , , 32505 457 38 Pl Pl NNP 32505 457 39 . . . 32505 458 1 4A 4A NNP 32505 458 2 ) ) -RRB- 32505 458 3 . . . 32505 459 1 The the DT 32505 459 2 eggs egg NNS 32505 459 3 are be VBP 32505 459 4 deposited deposit VBN 32505 459 5 in in IN 32505 459 6 a a DT 32505 459 7 mass mass NN 32505 459 8 in in IN 32505 459 9 the the DT 32505 459 10 water water NN 32505 459 11 near near IN 32505 459 12 floating float VBG 32505 459 13 vegetation vegetation NN 32505 459 14 . . . 32505 460 1 William William NNP 32505 460 2 E. E. NNP 32505 460 3 Duellman Duellman NNP 32505 460 4 informed inform VBD 32505 460 5 me -PRON- PRP 32505 460 6 that that IN 32505 460 7 he -PRON- PRP 32505 460 8 observed observe VBD 32505 460 9 hatchlings hatchling NNS 32505 460 10 oriented orient VBN 32505 460 11 vertically vertically RB 32505 460 12 with with IN 32505 460 13 the the DT 32505 460 14 tip tip NN 32505 460 15 of of IN 32505 460 16 the the DT 32505 460 17 mouth mouth NN 32505 460 18 at at IN 32505 460 19 the the DT 32505 460 20 surface surface NN 32505 460 21 of of IN 32505 460 22 the the DT 32505 460 23 water water NN 32505 460 24 . . . 32505 461 1 They -PRON- PRP 32505 461 2 gradually gradually RB 32505 461 3 sank sink VBD 32505 461 4 to to IN 32505 461 5 bottom bottom NN 32505 461 6 , , , 32505 461 7 but but CC 32505 461 8 swam swam NNP 32505 461 9 back back RB 32505 461 10 to to IN 32505 461 11 surface surface NN 32505 461 12 again again RB 32505 461 13 . . . 32505 462 1 No no DT 32505 462 2 additional additional JJ 32505 462 3 information information NN 32505 462 4 is be VBZ 32505 462 5 available available JJ 32505 462 6 concerning concern VBG 32505 462 7 early early JJ 32505 462 8 development development NN 32505 462 9 . . . 32505 463 1 Tadpoles tadpole NNS 32505 463 2 have have VBP 32505 463 3 been be VBN 32505 463 4 found find VBN 32505 463 5 in in IN 32505 463 6 shallow shallow JJ 32505 463 7 grassy grassy JJ 32505 463 8 ponds pond NNS 32505 463 9 in in IN 32505 463 10 clearings clearing NNS 32505 463 11 and and CC 32505 463 12 in in IN 32505 463 13 temporary temporary JJ 32505 463 14 woodland woodland JJ 32505 463 15 ponds pond NNS 32505 463 16 . . . 32505 464 1 _ _ NNP 32505 464 2 Tadpoles._--Three tadpoles._--three CD 32505 464 3 hundred hundred CD 32505 464 4 and and CC 32505 464 5 thirty thirty CD 32505 464 6 - - HYPH 32505 464 7 one one CD 32505 464 8 tadpoles tadpole NNS 32505 464 9 in in IN 32505 464 10 various various JJ 32505 464 11 stages stage NNS 32505 464 12 of of IN 32505 464 13 development development NN 32505 464 14 are be VBP 32505 464 15 available available JJ 32505 464 16 . . . 32505 465 1 Thirty thirty CD 32505 465 2 - - HYPH 32505 465 3 five five CD 32505 465 4 tadpoles tadpole NNS 32505 465 5 in in IN 32505 465 6 stage stage NN 32505 465 7 35 35 CD 32505 465 8 have have VBP 32505 465 9 a a DT 32505 465 10 mean mean JJ 32505 465 11 body body NN 32505 465 12 length length NN 32505 465 13 of of IN 32505 465 14 8.1 8.1 CD 32505 465 15 mm mm CD 32505 465 16 . . . 32505 466 1 ( ( -LRB- 32505 466 2 8.0 8.0 CD 32505 466 3 - - SYM 32505 466 4 9.0 9.0 CD 32505 466 5 mm mm CD 32505 466 6 . . . 32505 467 1 ) ) -RRB- 32505 467 2 , , , 32505 467 3 tail tail NN 32505 467 4 length length NN 32505 467 5 of of IN 32505 467 6 17.7 17.7 CD 32505 467 7 mm mm NNS 32505 467 8 . . . 32505 468 1 ( ( -LRB- 32505 468 2 15.0 15.0 CD 32505 468 3 - - SYM 32505 468 4 19.5 19.5 CD 32505 468 5 mm mm CD 32505 468 6 . . . 32505 469 1 ) ) -RRB- 32505 469 2 , , , 32505 469 3 and and CC 32505 469 4 total total JJ 32505 469 5 length length NN 32505 469 6 of of IN 32505 469 7 25.9 25.9 CD 32505 469 8 mm mm NNP 32505 469 9 . . . 32505 470 1 ( ( -LRB- 32505 470 2 23.0 23.0 CD 32505 470 3 - - SYM 32505 470 4 27.5 27.5 CD 32505 470 5 mm mm NNP 32505 470 6 . . . 32505 470 7 ) ) -RRB- 32505 470 8 . . . 32505 471 1 The the DT 32505 471 2 largest large JJS 32505 471 3 tadpole tadpole NN 32505 471 4 examined examine VBN 32505 471 5 is be VBZ 32505 471 6 in in IN 32505 471 7 stage stage NN 32505 471 8 40 40 CD 32505 471 9 and and CC 32505 471 10 has have VBZ 32505 471 11 a a DT 32505 471 12 total total JJ 32505 471 13 length length NN 32505 471 14 of of IN 32505 471 15 34.5 34.5 CD 32505 471 16 mm mm NN 32505 471 17 . . . 32505 472 1 ( ( -LRB- 32505 472 2 Table table NN 32505 472 3 6 6 CD 32505 472 4 ) ) -RRB- 32505 472 5 . . . 32505 473 1 A a DT 32505 473 2 typical typical JJ 32505 473 3 tadpole tadpole NN 32505 473 4 , , , 32505 473 5 stage stage VB 32505 473 6 35 35 CD 32505 473 7 of of IN 32505 473 8 development development NN 32505 473 9 ( ( -LRB- 32505 473 10 KU KU NNP 32505 473 11 104134 104134 CD 32505 473 12 , , , 32505 473 13 from from IN 32505 473 14 Puerto Puerto NNP 32505 473 15 Viejo Viejo NNP 32505 473 16 , , , 32505 473 17 Heredia Heredia NNP 32505 473 18 Province Province NNP 32505 473 19 , , , 32505 473 20 Costa Costa NNP 32505 473 21 Rica Rica NNP 32505 473 22 ) ) -RRB- 32505 473 23 , , , 32505 473 24 has have VBZ 32505 473 25 a a DT 32505 473 26 body body NN 32505 473 27 length length NN 32505 473 28 of of IN 32505 473 29 9.1 9.1 CD 32505 473 30 mm mm CD 32505 473 31 . . . 32505 473 32 , , , 32505 473 33 tail tail NN 32505 473 34 length length NN 32505 473 35 of of IN 32505 473 36 17.7 17.7 CD 32505 473 37 mm mm NNP 32505 473 38 . . NNP 32505 473 39 , , , 32505 473 40 and and CC 32505 473 41 a a DT 32505 473 42 total total JJ 32505 473 43 length length NN 32505 473 44 of of IN 32505 473 45 26.8 26.8 CD 32505 473 46 mm mm NNS 32505 473 47 . . . 32505 474 1 Other other JJ 32505 474 2 characters character NNS 32505 474 3 are be VBP 32505 474 4 as as IN 32505 474 5 follows follow VBZ 32505 474 6 : : : 32505 474 7 body body NN 32505 474 8 depressed depress VBN 32505 474 9 anteriorly anteriorly RB 32505 474 10 ; ; : 32505 474 11 body body NN 32505 474 12 length length NN 32505 474 13 greater great JJR 32505 474 14 than than IN 32505 474 15 depth depth NN 32505 474 16 of of IN 32505 474 17 tail tail NN 32505 474 18 ; ; : 32505 474 19 internarial internarial JJ 32505 474 20 space space NN 32505 474 21 as as RB 32505 474 22 broad broad JJ 32505 474 23 as as IN 32505 474 24 interorbital interorbital NNP 32505 474 25 distance distance NN 32505 474 26 ; ; : 32505 474 27 nostril nostril NNP 32505 474 28 equidistant equidistant NN 32505 474 29 between between IN 32505 474 30 eye eye NN 32505 474 31 and and CC 32505 474 32 tip tip NN 32505 474 33 of of IN 32505 474 34 snout snout NN 32505 474 35 ; ; : 32505 474 36 eyes eye NNS 32505 474 37 moderately moderately RB 32505 474 38 large large JJ 32505 474 39 ; ; : 32505 474 40 mouth mouth NNP 32505 474 41 anteroventral anteroventral NNP 32505 474 42 and and CC 32505 474 43 triangular triangular NNP 32505 474 44 ; ; : 32505 474 45 median median NNP 32505 474 46 fourth fourth RB 32505 474 47 of of IN 32505 474 48 upper upper JJ 32505 474 49 lip lip NNP 32505 474 50 bare bare NNP 32505 474 51 ; ; : 32505 474 52 rest rest NN 32505 474 53 of of IN 32505 474 54 lip lip NN 32505 474 55 bordered border VBN 32505 474 56 by by IN 32505 474 57 one one CD 32505 474 58 row row NN 32505 474 59 of of IN 32505 474 60 papillae papillae NNP 32505 474 61 ; ; , 32505 474 62 clumps clump NNS 32505 474 63 of of IN 32505 474 64 small small JJ 32505 474 65 papillae papilla NNS 32505 474 66 at at IN 32505 474 67 corners corner NNS 32505 474 68 of of IN 32505 474 69 mouth mouth NN 32505 474 70 ; ; , 32505 474 71 tooth tooth NN 32505 474 72 rows row VBZ 32505 474 73 2/3 2/3 CD 32505 474 74 ; ; : 32505 474 75 upper upper JJ 32505 474 76 rows row NNS 32505 474 77 equal equal JJ 32505 474 78 in in IN 32505 474 79 length length NN 32505 474 80 ; ; : 32505 474 81 second second JJ 32505 474 82 row row NN 32505 474 83 interrupted interrupt VBN 32505 474 84 medially medially RB 32505 474 85 ; ; : 32505 474 86 lower low JJR 32505 474 87 rows row NNS 32505 474 88 shorter short JJR 32505 474 89 than than IN 32505 474 90 upper upper JJ 32505 474 91 rows row NNS 32505 474 92 , , , 32505 474 93 diminishing diminish VBG 32505 474 94 in in IN 32505 474 95 length length NN 32505 474 96 ; ; : 32505 474 97 beak beak JJ 32505 474 98 rather rather RB 32505 474 99 weak weak JJ 32505 474 100 with with IN 32505 474 101 small small JJ 32505 474 102 serrations serration NNS 32505 474 103 ; ; : 32505 474 104 spiracle spiracle NNP 32505 474 105 short short JJ 32505 474 106 and and CC 32505 474 107 nearer nearer JJ 32505 474 108 eyes eye NNS 32505 474 109 than than IN 32505 474 110 anus anus NN 32505 474 111 ; ; : 32505 474 112 anal anal JJ 32505 474 113 opening opening NN 32505 474 114 not not RB 32505 474 115 reaching reach VBG 32505 474 116 edge edge NN 32505 474 117 of of IN 32505 474 118 ventral ventral JJ 32505 474 119 fin fin NN 32505 474 120 ; ; , 32505 474 121 caudal caudal NN 32505 474 122 musculature musculature NN 32505 474 123 attenuated attenuate VBD 32505 474 124 distally distally RB 32505 474 125 ( ( -LRB- 32505 474 126 Figs fig NNS 32505 474 127 . . . 32505 475 1 2B 2B NNP 32505 475 2 and and CC 32505 475 3 3B 3B NNP 32505 475 4 ) ) -RRB- 32505 475 5 . . . 32505 476 1 TABLE TABLE NNP 32505 476 2 6.--Sizes 6.--sizes CD 32505 476 3 of of IN 32505 476 4 Tadpoles Tadpoles NNP 32505 476 5 of of IN 32505 476 6 _ _ NNP 32505 476 7 Hyla Hyla NNP 32505 476 8 elaeochroa elaeochroa NN 32505 476 9 _ _ NNP 32505 476 10 in in IN 32505 476 11 Relation Relation NNP 32505 476 12 to to IN 32505 476 13 Developmental Developmental NNP 32505 476 14 Stages Stages NNPS 32505 476 15 . . . 32505 477 1 ( ( -LRB- 32505 477 2 Means mean NNS 32505 477 3 in in IN 32505 477 4 parentheses parenthesis NNS 32505 477 5 below below IN 32505 477 6 observed observed JJ 32505 477 7 ranges range NNS 32505 477 8 ; ; : 32505 477 9 measurements measurement NNS 32505 477 10 in in IN 32505 477 11 mm mm NNP 32505 477 12 . . . 32505 477 13 ) ) -RRB- 32505 478 1 ------+---+-------------+-------------+-------------- ------+---+-------------+-------------+-------------- NFP 32505 478 2 Stage stage NN 32505 478 3 | | NNP 32505 478 4 N N NNP 32505 478 5 | | NNP 32505 478 6 Body Body NNP 32505 478 7 length length NN 32505 478 8 | | NNP 32505 478 9 Tail Tail NNP 32505 478 10 length length NN 32505 478 11 | | NNP 32505 478 12 Total total JJ 32505 478 13 length length NN 32505 478 14 ------+---+-------------+-------------+-------------- ------+---+-------------+-------------+-------------- NNP 32505 478 15 24 24 CD 32505 478 16 | | CD 32505 478 17 2 2 CD 32505 478 18 | | CD 32505 478 19 4.0 4.0 CD 32505 478 20 - - CD 32505 478 21 4.0 4.0 CD 32505 478 22 | | CD 32505 478 23 8.5 8.5 CD 32505 478 24 - - HYPH 32505 478 25 9.0 9.0 CD 32505 478 26 | | CD 32505 478 27 12.5 12.5 CD 32505 478 28 - - HYPH 32505 478 29 13.0 13.0 CD 32505 478 30 | | CD 32505 478 31 | | CD 32505 478 32 ( ( -LRB- 32505 478 33 4.0 4.0 CD 32505 478 34 ) ) -RRB- 32505 478 35 | | NNP 32505 478 36 ( ( -LRB- 32505 478 37 8.8 8.8 CD 32505 478 38 ) ) -RRB- 32505 478 39 | | NNP 32505 478 40 ( ( -LRB- 32505 478 41 12.8 12.8 CD 32505 478 42 ) ) -RRB- 32505 478 43 | | CD 32505 478 44 | | CD 32505 478 45 | | CD 32505 478 46 | | CD 32505 478 47 25 25 CD 32505 478 48 |64 |64 CD 32505 478 49 | | CD 32505 478 50 5.0 5.0 CD 32505 478 51 - - HYPH 32505 478 52 6.5 6.5 CD 32505 478 53 | | CD 32505 478 54 8.5 8.5 CD 32505 478 55 - - HYPH 32505 478 56 15.0 15.0 CD 32505 478 57 | | CD 32505 478 58 13.5 13.5 CD 32505 478 59 - - HYPH 32505 478 60 21.5 21.5 CD 32505 478 61 | | CD 32505 478 62 | | CD 32505 478 63 ( ( -LRB- 32505 478 64 5.7 5.7 CD 32505 478 65 ) ) -RRB- 32505 478 66 | | NNP 32505 478 67 ( ( -LRB- 32505 478 68 11.8 11.8 CD 32505 478 69 ) ) -RRB- 32505 478 70 | | NNP 32505 478 71 ( ( -LRB- 32505 478 72 17.6 17.6 CD 32505 478 73 ) ) -RRB- 32505 478 74 | | CD 32505 478 75 | | CD 32505 478 76 | | CD 32505 478 77 | | CD 32505 478 78 27 27 CD 32505 478 79 |30 |30 CD 32505 478 80 | | CD 32505 478 81 7.0 7.0 CD 32505 478 82 - - HYPH 32505 478 83 7.5 7.5 CD 32505 478 84 | | CD 32505 478 85 13.0 13.0 CD 32505 478 86 - - HYPH 32505 478 87 16.0 16.0 CD 32505 478 88 | | CD 32505 478 89 20.0 20.0 CD 32505 478 90 - - HYPH 32505 478 91 23.0 23.0 CD 32505 478 92 | | CD 32505 478 93 | | CD 32505 478 94 ( ( -LRB- 32505 478 95 7.1 7.1 CD 32505 478 96 ) ) -RRB- 32505 478 97 | | NNP 32505 478 98 ( ( -LRB- 32505 478 99 14.2 14.2 CD 32505 478 100 ) ) -RRB- 32505 478 101 | | NNP 32505 478 102 ( ( -LRB- 32505 478 103 21.3 21.3 CD 32505 478 104 ) ) -RRB- 32505 478 105 | | NNP 32505 478 106 | | CD 32505 478 107 | | CD 32505 478 108 | | CD 32505 478 109 30 30 CD 32505 478 110 |15 |15 CD 32505 478 111 | | CD 32505 478 112 7.0 7.0 CD 32505 478 113 - - HYPH 32505 478 114 8.0 8.0 CD 32505 478 115 | | CD 32505 478 116 13.0 13.0 CD 32505 478 117 - - HYPH 32505 478 118 16.5 16.5 CD 32505 478 119 | | CD 32505 478 120 20.0 20.0 CD 32505 478 121 - - HYPH 32505 478 122 24.0 24.0 CD 32505 478 123 | | CD 32505 478 124 | | CD 32505 478 125 ( ( -LRB- 32505 478 126 7.3 7.3 CD 32505 478 127 ) ) -RRB- 32505 478 128 | | NNP 32505 478 129 ( ( -LRB- 32505 478 130 15.0 15.0 CD 32505 478 131 ) ) -RRB- 32505 478 132 | | NNP 32505 478 133 ( ( -LRB- 32505 478 134 22.4 22.4 CD 32505 478 135 ) ) -RRB- 32505 478 136 | | NNP 32505 478 137 | | NNP 32505 478 138 | | CD 32505 478 139 | | CD 32505 478 140 32 32 CD 32505 478 141 |30 |30 CD 32505 478 142 | | CD 32505 478 143 7.5 7.5 CD 32505 478 144 - - HYPH 32505 478 145 8.5 8.5 CD 32505 478 146 | | CD 32505 478 147 15.0 15.0 CD 32505 478 148 - - HYPH 32505 478 149 17.0 17.0 CD 32505 478 150 | | CD 32505 478 151 22.5 22.5 CD 32505 478 152 - - HYPH 32505 478 153 25.0 25.0 CD 32505 478 154 | | CD 32505 478 155 | | CD 32505 478 156 ( ( -LRB- 32505 478 157 7.8 7.8 CD 32505 478 158 ) ) -RRB- 32505 478 159 | | NNP 32505 478 160 ( ( -LRB- 32505 478 161 16.1 16.1 CD 32505 478 162 ) ) -RRB- 32505 478 163 | | NNP 32505 478 164 ( ( -LRB- 32505 478 165 23.8 23.8 CD 32505 478 166 ) ) -RRB- 32505 478 167 | | NNP 32505 478 168 | | CD 32505 478 169 | | CD 32505 478 170 | | CD 32505 478 171 35 35 CD 32505 478 172 |35 |35 CD 32505 478 173 | | CD 32505 478 174 8.0 8.0 CD 32505 478 175 - - HYPH 32505 478 176 9.0 9.0 CD 32505 478 177 | | CD 32505 478 178 15.0 15.0 CD 32505 478 179 - - HYPH 32505 478 180 19.5 19.5 CD 32505 478 181 | | CD 32505 478 182 23.0 23.0 CD 32505 478 183 - - HYPH 32505 478 184 27.5 27.5 CD 32505 478 185 | | CD 32505 478 186 | | CD 32505 478 187 ( ( -LRB- 32505 478 188 8.1 8.1 CD 32505 478 189 ) ) -RRB- 32505 478 190 | | NNP 32505 478 191 ( ( -LRB- 32505 478 192 17.7 17.7 CD 32505 478 193 ) ) -RRB- 32505 478 194 | | NNP 32505 478 195 ( ( -LRB- 32505 478 196 25.9 25.9 CD 32505 478 197 ) ) -RRB- 32505 478 198 | | NNP 32505 478 199 | | NNP 32505 478 200 | | CD 32505 478 201 | | CD 32505 478 202 37 37 CD 32505 478 203 |22 |22 CD 32505 478 204 | | CD 32505 478 205 8.5 8.5 CD 32505 478 206 - - HYPH 32505 478 207 9.5 9.5 CD 32505 478 208 | | CD 32505 478 209 16.0 16.0 CD 32505 478 210 - - HYPH 32505 478 211 22.0 22.0 CD 32505 478 212 | | CD 32505 478 213 25.0 25.0 CD 32505 478 214 - - HYPH 32505 478 215 31.0 31.0 CD 32505 478 216 | | CD 32505 478 217 | | CD 32505 478 218 ( ( -LRB- 32505 478 219 9.0 9.0 CD 32505 478 220 ) ) -RRB- 32505 478 221 | | NNP 32505 478 222 ( ( -LRB- 32505 478 223 18.8 18.8 CD 32505 478 224 ) ) -RRB- 32505 478 225 | | NNP 32505 478 226 ( ( -LRB- 32505 478 227 27.8 27.8 CD 32505 478 228 ) ) -RRB- 32505 478 229 | | NNP 32505 478 230 | | NNP 32505 478 231 | | CD 32505 478 232 | | CD 32505 478 233 39 39 CD 32505 478 234 |14 |14 CD 32505 478 235 | | CD 32505 478 236 9.5 9.5 CD 32505 478 237 - - HYPH 32505 478 238 10.5 10.5 CD 32505 478 239 | | CD 32505 478 240 19.0 19.0 CD 32505 478 241 - - HYPH 32505 478 242 24.9 24.9 CD 32505 478 243 | | CD 32505 478 244 28.5 28.5 CD 32505 478 245 - - HYPH 32505 478 246 33.5 33.5 CD 32505 478 247 | | CD 32505 478 248 | | NNS 32505 478 249 ( ( -LRB- 32505 478 250 9.9 9.9 CD 32505 478 251 ) ) -RRB- 32505 478 252 | | CD 32505 478 253 ( ( -LRB- 32505 478 254 21.1 21.1 CD 32505 478 255 ) ) -RRB- 32505 478 256 | | NNP 32505 478 257 ( ( -LRB- 32505 478 258 31.0 31.0 CD 32505 478 259 ) ) -RRB- 32505 478 260 | | NNP 32505 478 261 | | NNP 32505 478 262 | | CD 32505 478 263 | | CD 32505 478 264 40 40 CD 32505 478 265 |27 |27 CD 32505 478 266 | | CD 32505 478 267 7.0 7.0 CD 32505 478 268 - - HYPH 32505 478 269 11.5 11.5 CD 32505 478 270 | | CD 32505 478 271 15.0 15.0 CD 32505 478 272 - - HYPH 32505 478 273 23.0 23.0 CD 32505 478 274 | | CD 32505 478 275 23.0 23.0 CD 32505 478 276 - - HYPH 32505 478 277 34.5 34.5 CD 32505 478 278 | | CD 32505 478 279 | | NNS 32505 478 280 ( ( -LRB- 32505 478 281 9.1 9.1 CD 32505 478 282 ) ) -RRB- 32505 478 283 | | CD 32505 478 284 ( ( -LRB- 32505 478 285 22.0 22.0 CD 32505 478 286 ) ) -RRB- 32505 478 287 | | NNP 32505 478 288 ( ( -LRB- 32505 478 289 31.2 31.2 CD 32505 478 290 ) ) -RRB- 32505 478 291 | | NNP 32505 478 292 | | NNP 32505 478 293 | | CD 32505 478 294 | | CD 32505 478 295 43 43 CD 32505 478 296 |10 |10 CD 32505 478 297 | | CD 32505 478 298 8.0 8.0 CD 32505 478 299 - - HYPH 32505 478 300 12.0 12.0 CD 32505 478 301 | | CD 32505 478 302 11.0 11.0 CD 32505 478 303 - - HYPH 32505 478 304 17.0 17.0 CD 32505 478 305 | | CD 32505 478 306 20.0 20.0 CD 32505 478 307 - - HYPH 32505 478 308 26.0 26.0 CD 32505 478 309 | | CD 32505 478 310 | | CD 32505 478 311 ( ( -LRB- 32505 478 312 10.2 10.2 CD 32505 478 313 ) ) -RRB- 32505 478 314 | | NNP 32505 478 315 ( ( -LRB- 32505 478 316 13.5 13.5 CD 32505 478 317 ) ) -RRB- 32505 478 318 | | NNP 32505 478 319 ( ( -LRB- 32505 478 320 23.7 23.7 CD 32505 478 321 ) ) -RRB- 32505 478 322 | | NNP 32505 478 323 | | CD 32505 478 324 | | CD 32505 478 325 | | CD 32505 478 326 45 45 CD 32505 478 327 |16 |16 CD 32505 478 328 | | CD 32505 478 329 10.0 10.0 CD 32505 478 330 - - HYPH 32505 478 331 12.0 12.0 CD 32505 478 332 | | CD 32505 478 333 1.0 1.0 CD 32505 478 334 - - HYPH 32505 478 335 7.0 7.0 CD 32505 478 336 | | CD 32505 478 337 12.0 12.0 CD 32505 478 338 - - HYPH 32505 478 339 17.0 17.0 CD 32505 478 340 | | CD 32505 478 341 | | CD 32505 478 342 ( ( -LRB- 32505 478 343 11.2 11.2 CD 32505 478 344 ) ) -RRB- 32505 478 345 | | NNP 32505 478 346 ( ( -LRB- 32505 478 347 3.4 3.4 CD 32505 478 348 ) ) -RRB- 32505 478 349 | | NNP 32505 478 350 ( ( -LRB- 32505 478 351 14.6 14.6 CD 32505 478 352 ) ) -RRB- 32505 478 353 | | NNP 32505 478 354 | | NNP 32505 478 355 | | CD 32505 478 356 | | CD 32505 478 357 46 46 CD 32505 478 358 |45 |45 CD 32505 478 359 | | CD 32505 478 360 11.0 11.0 CD 32505 478 361 - - HYPH 32505 478 362 13.0 13.0 CD 32505 478 363 | | CD 32505 478 364 | | CD 32505 478 365 | | CD 32505 478 366 | | CD 32505 478 367 ( ( -LRB- 32505 478 368 11.8 11.8 CD 32505 478 369 ) ) -RRB- 32505 478 370 | | CD 32505 478 371 | | NNS 32505 478 372 In in IN 32505 478 373 life life NN 32505 478 374 , , , 32505 478 375 dorsum dorsum NN 32505 478 376 yellowish yellowish JJ 32505 478 377 tan tan NNP 32505 478 378 with with IN 32505 478 379 gray gray JJ 32505 478 380 - - HYPH 32505 478 381 brown brown JJ 32505 478 382 mottling mottling NN 32505 478 383 ; ; : 32505 478 384 belly belly NN 32505 478 385 and and CC 32505 478 386 ventrolateral ventrolateral JJ 32505 478 387 surfaces surface NNS 32505 478 388 silvery silvery JJ 32505 478 389 - - HYPH 32505 478 390 gold gold JJ 32505 478 391 or or CC 32505 478 392 white white JJ 32505 478 393 ; ; : 32505 478 394 black black JJ 32505 478 395 stripe stripe NN 32505 478 396 from from IN 32505 478 397 tip tip NN 32505 478 398 of of IN 32505 478 399 snout snout NN 32505 478 400 to to IN 32505 478 401 eye eye NN 32505 478 402 ; ; : 32505 478 403 two two CD 32505 478 404 black black JJ 32505 478 405 blotches blotch NNS 32505 478 406 below below IN 32505 478 407 eye eye NN 32505 478 408 , , , 32505 478 409 another another DT 32505 478 410 blotch blotch NN 32505 478 411 extending extend VBG 32505 478 412 from from IN 32505 478 413 eye eye NN 32505 478 414 to to IN 32505 478 415 base base NN 32505 478 416 of of IN 32505 478 417 caudal caudal JJ 32505 478 418 musculature musculature NN 32505 478 419 ; ; : 32505 478 420 caudal caudal JJ 32505 478 421 musculature musculature NN 32505 478 422 and and CC 32505 478 423 fins fin VBZ 32505 478 424 gray gray JJ 32505 478 425 - - HYPH 32505 478 426 brown brown JJ 32505 478 427 . . . 32505 479 1 In in IN 32505 479 2 preservative preservative JJ 32505 479 3 , , , 32505 479 4 yellowish yellowish NNP 32505 479 5 tan tan NNP 32505 479 6 and and CC 32505 479 7 silvery silvery JJ 32505 479 8 - - HYPH 32505 479 9 gold gold JJ 32505 479 10 colors color NNS 32505 479 11 lost lose VBD 32505 479 12 ; ; : 32505 479 13 black black JJ 32505 479 14 reticulations reticulation NNS 32505 479 15 present present JJ 32505 479 16 on on IN 32505 479 17 tail tail NN 32505 479 18 . . . 32505 480 1 _ _ NNP 32505 480 2 Remarks._--Cope Remarks._--Cope NNP 32505 480 3 ( ( -LRB- 32505 480 4 1876:105 1876:105 CD 32505 480 5 ) ) -RRB- 32505 480 6 described describe VBN 32505 480 7 _ _ NNP 32505 480 8 Hyla Hyla NNP 32505 480 9 elaeochroa elaeochroa NN 32505 480 10 _ _ NNP 32505 480 11 from from IN 32505 480 12 Sipurio Sipurio NNP 32505 480 13 , , , 32505 480 14 Limón Limón NNP 32505 480 15 Province Province NNP 32505 480 16 , , , 32505 480 17 Costa Costa NNP 32505 480 18 Rica Rica NNP 32505 480 19 . . . 32505 481 1 He -PRON- PRP 32505 481 2 based base VBD 32505 481 3 his -PRON- PRP$ 32505 481 4 description description NN 32505 481 5 on on IN 32505 481 6 a a DT 32505 481 7 small small JJ 32505 481 8 specimen speciman NNS 32505 481 9 , , , 32505 481 10 26.0 26.0 CD 32505 481 11 mm mm NNP 32505 481 12 . . . 32505 482 1 in in IN 32505 482 2 snout snout NN 32505 482 3 - - HYPH 32505 482 4 vent vent NN 32505 482 5 length length NN 32505 482 6 , , , 32505 482 7 having have VBG 32505 482 8 a a DT 32505 482 9 dorsum dorsum NN 32505 482 10 uniformly uniformly RB 32505 482 11 colored color VBN 32505 482 12 and and CC 32505 482 13 lacking lack VBG 32505 482 14 an an DT 32505 482 15 interorbital interorbital JJ 32505 482 16 triangle triangle NN 32505 482 17 and and CC 32505 482 18 blotches blotch NNS 32505 482 19 on on IN 32505 482 20 the the DT 32505 482 21 thighs thigh NNS 32505 482 22 . . . 32505 483 1 Cope Cope NNP 32505 483 2 ( ( -LRB- 32505 483 3 1887 1887 CD 32505 483 4 ) ) -RRB- 32505 483 5 described describe VBD 32505 483 6 pigmented pigment VBN 32505 483 7 specimens specimen NNS 32505 483 8 from from IN 32505 483 9 Nicaragua Nicaragua NNP 32505 483 10 as as IN 32505 483 11 _ _ NNP 32505 483 12 Hyla Hyla NNP 32505 483 13 quinquevittata quinquevittata NN 32505 483 14 _ _ NNP 32505 483 15 , , , 32505 483 16 which which WDT 32505 483 17 he -PRON- PRP 32505 483 18 diagnosed diagnose VBD 32505 483 19 as as IN 32505 483 20 having have VBG 32505 483 21 dark dark JJ 32505 483 22 brown brown JJ 32505 483 23 bars bar NNS 32505 483 24 on on IN 32505 483 25 the the DT 32505 483 26 hind hind NN 32505 483 27 limbs limb NNS 32505 483 28 and and CC 32505 483 29 five five CD 32505 483 30 dark dark JJ 32505 483 31 brown brown JJ 32505 483 32 longitudinal longitudinal JJ 32505 483 33 stripes stripe NNS 32505 483 34 on on IN 32505 483 35 the the DT 32505 483 36 dorsum dorsum NN 32505 483 37 , , , 32505 483 38 the the DT 32505 483 39 median median JJ 32505 483 40 one one CD 32505 483 41 of of IN 32505 483 42 which which WDT 32505 483 43 was be VBD 32505 483 44 expanded expand VBN 32505 483 45 anteriorly anteriorly RB 32505 483 46 so so IN 32505 483 47 as as IN 32505 483 48 to to TO 32505 483 49 form form VB 32505 483 50 a a DT 32505 483 51 large large JJ 32505 483 52 triangular triangular JJ 32505 483 53 spot spot NN 32505 483 54 between between IN 32505 483 55 the the DT 32505 483 56 eyes eye NNS 32505 483 57 . . . 32505 484 1 He -PRON- PRP 32505 484 2 thought think VBD 32505 484 3 this this DT 32505 484 4 species species NN 32505 484 5 was be VBD 32505 484 6 related relate VBN 32505 484 7 to to IN 32505 484 8 _ _ NNP 32505 484 9 Hyla Hyla NNP 32505 484 10 eximia eximia NN 32505 484 11 _ _ NNP 32505 484 12 Baird Baird NNP 32505 484 13 and and CC 32505 484 14 noted note VBD 32505 484 15 that that IN 32505 484 16 " " `` 32505 484 17 the the DT 32505 484 18 hinder hinder NN 32505 484 19 legs leg NNS 32505 484 20 are be VBP 32505 484 21 much much RB 32505 484 22 larger large JJR 32505 484 23 ; ; : 32505 484 24 the the DT 32505 484 25 muzzle muzzle NN 32505 484 26 is be VBZ 32505 484 27 more more JJR 32505 484 28 acuminate acuminate NN 32505 484 29 and and CC 32505 484 30 the the DT 32505 484 31 color color NN 32505 484 32 bands band NNS 32505 484 33 are be VBP 32505 484 34 much much RB 32505 484 35 wider wide JJR 32505 484 36 " " '' 32505 484 37 than than IN 32505 484 38 in in IN 32505 484 39 _ _ NNP 32505 484 40 eximia eximia NN 32505 484 41 _ _ NNP 32505 484 42 . . . 32505 485 1 Cope Cope NNP 32505 485 2 did do VBD 32505 485 3 not not RB 32505 485 4 compare compare VB 32505 485 5 _ _ NNP 32505 485 6 quinquevittata quinquevittata NN 32505 485 7 _ _ NNP 32505 485 8 with with IN 32505 485 9 _ _ NNP 32505 485 10 elaeochroa elaeochroa NNP 32505 485 11 _ _ NNP 32505 485 12 , , , 32505 485 13 which which WDT 32505 485 14 he -PRON- PRP 32505 485 15 had have VBD 32505 485 16 described describe VBN 32505 485 17 ten ten CD 32505 485 18 years year NNS 32505 485 19 before before RB 32505 485 20 . . . 32505 486 1 Günther Günther NNP 32505 486 2 ( ( -LRB- 32505 486 3 1901:268 1901:268 CD 32505 486 4 ) ) -RRB- 32505 486 5 , , , 32505 486 6 Noble Noble NNP 32505 486 7 ( ( -LRB- 32505 486 8 1918:340 1918:340 CD 32505 486 9 ) ) -RRB- 32505 486 10 , , , 32505 486 11 and and CC 32505 486 12 Nieden Nieden NNP 32505 486 13 ( ( -LRB- 32505 486 14 1923:251 1923:251 CD 32505 486 15 ) ) -RRB- 32505 486 16 regarded regard VBD 32505 486 17 both both CC 32505 486 18 _ _ NNP 32505 486 19 elaeochroa elaeochroa NNP 32505 486 20 _ _ NNP 32505 486 21 and and CC 32505 486 22 _ _ NNP 32505 486 23 quinquevittata quinquevittata NN 32505 486 24 _ _ NNP 32505 486 25 as as IN 32505 486 26 valid valid JJ 32505 486 27 species specie NNS 32505 486 28 . . . 32505 487 1 Dunn Dunn NNP 32505 487 2 and and CC 32505 487 3 Emlen Emlen NNP 32505 487 4 ( ( -LRB- 32505 487 5 1932:25 1932:25 CD 32505 487 6 ) ) -RRB- 32505 487 7 regarded regard VBD 32505 487 8 both both DT 32505 487 9 as as IN 32505 487 10 synonyms synonyms NN 32505 487 11 of of IN 32505 487 12 _ _ NNP 32505 487 13 Hyla Hyla NNP 32505 487 14 rubra rubra NN 32505 487 15 _ _ NNP 32505 487 16 , , , 32505 487 17 but but CC 32505 487 18 they -PRON- PRP 32505 487 19 made make VBD 32505 487 20 no no DT 32505 487 21 qualifying qualifying NN 32505 487 22 statements statement NNS 32505 487 23 . . . 32505 488 1 Taylor Taylor NNP 32505 488 2 ( ( -LRB- 32505 488 3 1952:859 1952:859 CD 32505 488 4 ) ) -RRB- 32505 488 5 placed place VBN 32505 488 6 _ _ NNP 32505 488 7 quinquevittata quinquevittata NN 32505 488 8 _ _ NNP 32505 488 9 as as IN 32505 488 10 a a DT 32505 488 11 synonym synonym NN 32505 488 12 of of IN 32505 488 13 _ _ NNP 32505 488 14 elaeochroa elaeochroa NNP 32505 488 15 _ _ NNP 32505 488 16 and and CC 32505 488 17 indicated indicate VBD 32505 488 18 that that IN 32505 488 19 _ _ NNP 32505 488 20 rubra rubra NNP 32505 488 21 _ _ NNP 32505 488 22 was be VBD 32505 488 23 another another DT 32505 488 24 species species NN 32505 488 25 . . . 32505 489 1 Taylor Taylor NNP 32505 489 2 ( ( -LRB- 32505 489 3 1958:37 1958:37 CD 32505 489 4 ) ) -RRB- 32505 489 5 described describe VBD 32505 489 6 _ _ NNP 32505 489 7 Hyla Hyla NNP 32505 489 8 dulcensis dulcensis NN 32505 489 9 _ _ NNP 32505 489 10 from from IN 32505 489 11 the the DT 32505 489 12 humid humid JJ 32505 489 13 tropical tropical JJ 32505 489 14 forests forest NNS 32505 489 15 of of IN 32505 489 16 Golfo Golfo NNP 32505 489 17 Dulce Dulce NNP 32505 489 18 , , , 32505 489 19 Puntarenas Puntarenas NNP 32505 489 20 Province Province NNP 32505 489 21 , , , 32505 489 22 Costa Costa NNP 32505 489 23 Rica Rica NNP 32505 489 24 . . . 32505 490 1 He -PRON- PRP 32505 490 2 thought think VBD 32505 490 3 this this DT 32505 490 4 species species NN 32505 490 5 was be VBD 32505 490 6 " " `` 32505 490 7 related related JJ 32505 490 8 to to IN 32505 490 9 _ _ NNP 32505 490 10 H. H. NNP 32505 491 1 elaeochroa elaeochroa NNP 32505 491 2 _ _ NNP 32505 491 3 but but CC 32505 491 4 differs differ VBZ 32505 491 5 in in IN 32505 491 6 its -PRON- PRP$ 32505 491 7 somewhat somewhat RB 32505 491 8 larger large JJR 32505 491 9 size size NN 32505 491 10 , , , 32505 491 11 smaller small JJR 32505 491 12 finger finger NN 32505 491 13 and and CC 32505 491 14 toe toe NN 32505 491 15 discs disc NNS 32505 491 16 , , , 32505 491 17 the the DT 32505 491 18 obsolete obsolete JJ 32505 491 19 canthus canthus NNP 32505 491 20 rostralis rostralis NNP 32505 491 21 ; ; : 32505 491 22 the the DT 32505 491 23 loreal loreal NN 32505 491 24 region region NN 32505 491 25 concave concave NNP 32505 491 26 and and CC 32505 491 27 the the DT 32505 491 28 choanae choanae NN 32505 491 29 larger large JJR 32505 491 30 . . . 32505 491 31 " " '' 32505 492 1 Duellman Duellman NNP 32505 492 2 ( ( -LRB- 32505 492 3 1966a:270 1966a:270 CD 32505 492 4 ) ) -RRB- 32505 492 5 compared compare VBN 32505 492 6 adults adult NNS 32505 492 7 , , , 32505 492 8 tadpoles tadpole NNS 32505 492 9 , , , 32505 492 10 and and CC 32505 492 11 mating mating NN 32505 492 12 calls call NNS 32505 492 13 of of IN 32505 492 14 _ _ NNP 32505 492 15 dulcensis dulcensis NN 32505 492 16 _ _ NNP 32505 492 17 and and CC 32505 492 18 _ _ NNP 32505 492 19 elaeochroa elaeochroa NNP 32505 492 20 _ _ NNP 32505 492 21 and and CC 32505 492 22 concluded conclude VBD 32505 492 23 that that IN 32505 492 24 a a DT 32505 492 25 single single JJ 32505 492 26 species specie NNS 32505 492 27 was be VBD 32505 492 28 involved involve VBN 32505 492 29 . . . 32505 493 1 _ _ NNP 32505 493 2 Hyla Hyla NNP 32505 493 3 elaeochroa elaeochroa NN 32505 493 4 _ _ NNP 32505 493 5 can can MD 32505 493 6 be be VB 32505 493 7 easily easily RB 32505 493 8 confused confuse VBN 32505 493 9 with with IN 32505 493 10 the the DT 32505 493 11 closely closely RB 32505 493 12 related relate VBN 32505 493 13 _ _ NNP 32505 493 14 Hyla Hyla NNP 32505 493 15 staufferi staufferi NN 32505 493 16 _ _ NNP 32505 493 17 . . . 32505 494 1 Although although IN 32505 494 2 the the DT 32505 494 3 durations duration NNS 32505 494 4 of of IN 32505 494 5 the the DT 32505 494 6 calls call NNS 32505 494 7 are be VBP 32505 494 8 similar similar JJ 32505 494 9 , , , 32505 494 10 the the DT 32505 494 11 call call NN 32505 494 12 of of IN 32505 494 13 _ _ NNP 32505 494 14 elaeochroa elaeochroa NNP 32505 494 15 _ _ NNP 32505 494 16 has have VBZ 32505 494 17 only only RB 32505 494 18 about about RB 32505 494 19 one one CD 32505 494 20 third third NN 32505 494 21 the the DT 32505 494 22 number number NN 32505 494 23 of of IN 32505 494 24 pulses pulse NNS 32505 494 25 per per IN 32505 494 26 second second NN 32505 494 27 , , , 32505 494 28 a a DT 32505 494 29 much much RB 32505 494 30 lower low JJR 32505 494 31 fundamental fundamental JJ 32505 494 32 frequency frequency NN 32505 494 33 , , , 32505 494 34 and and CC 32505 494 35 a a DT 32505 494 36 lower low JJR 32505 494 37 dominant dominant JJ 32505 494 38 frequency frequency NN 32505 494 39 than than IN 32505 494 40 that that DT 32505 494 41 of of IN 32505 494 42 _ _ NNP 32505 494 43 staufferi staufferi NN 32505 494 44 _ _ NNP 32505 494 45 . . . 32505 495 1 _ _ NNP 32505 495 2 Hyla Hyla NNP 32505 495 3 elaeochroa elaeochroa NN 32505 495 4 _ _ NNP 32505 495 5 is be VBZ 32505 495 6 larger large JJR 32505 495 7 and and CC 32505 495 8 has have VBZ 32505 495 9 a a DT 32505 495 10 less less RBR 32505 495 11 pointed pointed JJ 32505 495 12 snout snout NN 32505 495 13 than than IN 32505 495 14 does do VBZ 32505 495 15 _ _ NNP 32505 495 16 staufferi staufferi NN 32505 495 17 _ _ NNP 32505 495 18 . . . 32505 496 1 Although although IN 32505 496 2 the the DT 32505 496 3 skulls skull NNS 32505 496 4 of of IN 32505 496 5 the the DT 32505 496 6 two two CD 32505 496 7 species specie NNS 32505 496 8 are be VBP 32505 496 9 similar similar JJ 32505 496 10 , , , 32505 496 11 that that DT 32505 496 12 of of IN 32505 496 13 _ _ NNP 32505 496 14 elaeochroa elaeochroa NNP 32505 496 15 _ _ NNP 32505 496 16 differs differ VBZ 32505 496 17 in in IN 32505 496 18 having have VBG 32505 496 19 broad broad JJ 32505 496 20 palatines palatine NNS 32505 496 21 and and CC 32505 496 22 comparatively comparatively RB 32505 496 23 larger large JJR 32505 496 24 nasals nasal NNS 32505 496 25 that that WDT 32505 496 26 are be VBP 32505 496 27 truncate truncate JJ 32505 496 28 anteriorly anteriorly RB 32505 496 29 . . . 32505 497 1 In in IN 32505 497 2 _ _ NNP 32505 497 3 staufferi staufferi NN 32505 497 4 _ _ NNP 32505 497 5 the the DT 32505 497 6 nasal nasal NN 32505 497 7 is be VBZ 32505 497 8 rounded round VBN 32505 497 9 anteriorly anteriorly RB 32505 497 10 and and CC 32505 497 11 the the DT 32505 497 12 palatine palatine NN 32505 497 13 is be VBZ 32505 497 14 absent absent JJ 32505 497 15 . . . 32505 498 1 _ _ NNP 32505 498 2 Distribution._--_Hyla Distribution._--_Hyla NNP 32505 498 3 elaeochroa elaeochroa NNP 32505 498 4 _ _ NNP 32505 498 5 occurs occur VBZ 32505 498 6 on on IN 32505 498 7 the the DT 32505 498 8 Caribbean caribbean JJ 32505 498 9 lowlands lowland NNS 32505 498 10 from from IN 32505 498 11 western western JJ 32505 498 12 Panamá Panamá NNP 32505 498 13 through through IN 32505 498 14 Costa Costa NNP 32505 498 15 Rica Rica NNP 32505 498 16 to to TO 32505 498 17 eastern eastern JJ 32505 498 18 Nicaragua Nicaragua NNP 32505 498 19 , , , 32505 498 20 and and CC 32505 498 21 on on IN 32505 498 22 the the DT 32505 498 23 Pacific Pacific NNP 32505 498 24 lowlands lowland NNS 32505 498 25 of of IN 32505 498 26 southeastern southeastern JJ 32505 498 27 Costa Costa NNP 32505 498 28 Rica Rica NNP 32505 498 29 and and CC 32505 498 30 extreme extreme JJ 32505 498 31 western western JJ 32505 498 32 Panamá Panamá NNP 32505 498 33 . . . 32505 499 1 Most Most JJS 32505 499 2 localities locality NNS 32505 499 3 where where WRB 32505 499 4 it -PRON- PRP 32505 499 5 has have VBZ 32505 499 6 been be VBN 32505 499 7 collected collect VBN 32505 499 8 are be VBP 32505 499 9 below below IN 32505 499 10 800 800 CD 32505 499 11 meters meter NNS 32505 499 12 , , , 32505 499 13 but but CC 32505 499 14 the the DT 32505 499 15 species specie NNS 32505 499 16 has have VBZ 32505 499 17 been be VBN 32505 499 18 found find VBN 32505 499 19 at at IN 32505 499 20 two two CD 32505 499 21 localities locality NNS 32505 499 22 above above IN 32505 499 23 1000 1000 CD 32505 499 24 meters meter NNS 32505 499 25 ( ( -LRB- 32505 499 26 El El NNP 32505 499 27 Silencio Silencio NNP 32505 499 28 and and CC 32505 499 29 Pacuare Pacuare NNP 32505 499 30 , , , 32505 499 31 Cartago Cartago NNP 32505 499 32 Province Province NNP 32505 499 33 ) ) -RRB- 32505 499 34 on on IN 32505 499 35 the the DT 32505 499 36 Caribbean caribbean JJ 32505 499 37 slopes slope NNS 32505 499 38 of of IN 32505 499 39 the the DT 32505 499 40 Cordillera Cordillera NNP 32505 499 41 de de IN 32505 499 42 Talamanca Talamanca NNP 32505 499 43 , , , 32505 499 44 Costa Costa NNP 32505 499 45 Rica Rica NNP 32505 499 46 ( ( -LRB- 32505 499 47 Fig Fig NNP 32505 499 48 . . . 32505 500 1 6 6 LS 32505 500 2 ) ) -RRB- 32505 500 3 . . . 32505 501 1 _ _ NNP 32505 501 2 Specimens Specimens NNP 32505 501 3 Examined._--Nicaragua Examined._--Nicaragua NNP 32505 501 4 : : : 32505 501 5 _ _ NNP 32505 501 6 Zelaya Zelaya NNP 32505 501 7 _ _ NNP 32505 501 8 : : : 32505 501 9 El El NNP 32505 501 10 Recreo Recreo NNP 32505 501 11 , , , 32505 501 12 UMMZ UMMZ NNP 32505 501 13 79721 79721 CD 32505 501 14 ( ( -LRB- 32505 501 15 9 9 CD 32505 501 16 ) ) -RRB- 32505 501 17 . . . 32505 502 1 Costa Costa NNP 32505 502 2 Rica Rica NNP 32505 502 3 : : : 32505 502 4 _ _ NNP 32505 502 5 Alajuela Alajuela NNP 32505 502 6 _ _ NNP 32505 502 7 : : : 32505 502 8 Laguna Laguna NNP 32505 502 9 Monte Monte NNP 32505 502 10 Alegre Alegre NNP 32505 502 11 , , , 32505 502 12 KU KU NNP 32505 502 13 64499 64499 CD 32505 502 14 . . . 32505 503 1 _ _ NNP 32505 503 2 Cartago Cartago NNP 32505 503 3 _ _ NNP 32505 503 4 : : : 32505 503 5 2 2 CD 32505 503 6 km km NN 32505 503 7 E E NNP 32505 503 8 Chitaría Chitaría NNP 32505 503 9 , , , 32505 503 10 KU KU NNP 32505 503 11 107058 107058 CD 32505 503 12 ; ; : 32505 503 13 El El NNP 32505 503 14 Silencio Silencio NNP 32505 503 15 , , , 32505 503 16 14.4 14.4 CD 32505 503 17 km km NN 32505 503 18 NE NE NNP 32505 503 19 Turrialba Turrialba NNP 32505 503 20 , , , 32505 503 21 KU KU NNP 32505 503 22 107059 107059 CD 32505 503 23 - - SYM 32505 503 24 60 60 CD 32505 503 25 ; ; : 32505 503 26 4.6 4.6 CD 32505 503 27 km km NN 32505 503 28 ENE ENE NNP 32505 503 29 Pacuare Pacuare NNP 32505 503 30 , , , 32505 503 31 KU KU NNP 32505 503 32 64451 64451 CD 32505 503 33 - - SYM 32505 503 34 75 75 CD 32505 503 35 , , , 32505 503 36 64628 64628 CD 32505 503 37 - - SYM 32505 503 38 37 37 CD 32505 503 39 ; ; : 32505 503 40 4 4 LS 32505 503 41 km km NN 32505 503 42 S S NNP 32505 503 43 Pavones Pavones NNP 32505 503 44 , , , 32505 503 45 KU KU NNP 32505 503 46 64500 64500 CD 32505 503 47 ; ; : 32505 503 48 Turrialba Turrialba NNP 32505 503 49 ( ( -LRB- 32505 503 50 Instituto Instituto NNP 32505 503 51 Interamericano Interamericano NNP 32505 503 52 de de NNP 32505 503 53 Ciéncias Ciéncias NNP 32505 503 54 Agrícolas Agrícolas NNP 32505 503 55 ) ) -RRB- 32505 503 56 , , , 32505 503 57 KU KU NNP 32505 503 58 30305 30305 CD 32505 503 59 - - SYM 32505 503 60 26 26 CD 32505 503 61 , , , 32505 503 62 24616 24616 CD 32505 503 63 - - SYM 32505 503 64 57 57 CD 32505 503 65 , , , 32505 503 66 30337 30337 CD 32505 503 67 - - SYM 32505 503 68 54 54 CD 32505 503 69 , , , 32505 503 70 31776 31776 CD 32505 503 71 - - SYM 32505 503 72 91 91 CD 32505 503 73 , , , 32505 503 74 31803 31803 CD 32505 503 75 , , , 32505 503 76 31807 31807 CD 32505 503 77 - - SYM 32505 503 78 15 15 CD 32505 503 79 , , , 32505 503 80 64413 64413 CD 32505 503 81 - - SYM 32505 503 82 50 50 CD 32505 503 83 , , , 32505 503 84 68283 68283 CD 32505 503 85 - - SYM 32505 503 86 87 87 CD 32505 503 87 ( ( -LRB- 32505 503 88 skeletons skeleton NNS 32505 503 89 ) ) -RRB- 32505 503 90 , , , 32505 503 91 68390 68390 CD 32505 503 92 - - SYM 32505 503 93 1 1 NNP 32505 503 94 ( ( -LRB- 32505 503 95 young young JJ 32505 503 96 ) ) -RRB- 32505 503 97 , , , 32505 503 98 35042 35042 CD 32505 503 99 ( ( -LRB- 32505 503 100 eggs egg NNS 32505 503 101 ) ) -RRB- 32505 503 102 , , , 32505 503 103 25207 25207 CD 32505 503 104 - - SYM 32505 503 105 8 8 CD 32505 503 106 ( ( -LRB- 32505 503 107 skeletons skeleton NNS 32505 503 108 ) ) -RRB- 32505 503 109 , , , 32505 503 110 25221 25221 CD 32505 503 111 ( ( -LRB- 32505 503 112 skeleton skeleton NNP 32505 503 113 ) ) -RRB- 32505 503 114 , , , 32505 503 115 41073 41073 CD 32505 503 116 - - SYM 32505 503 117 83 83 CD 32505 503 118 ( ( -LRB- 32505 503 119 skeletons skeleton NNS 32505 503 120 ) ) -RRB- 32505 503 121 . . . 32505 504 1 _ _ NNP 32505 504 2 Guanacaste Guanacaste NNP 32505 504 3 _ _ NNP 32505 504 4 : : : 32505 504 5 2 2 CD 32505 504 6 km km NN 32505 504 7 E E NNP 32505 504 8 Tilarán Tilarán NNP 32505 504 9 , , , 32505 504 10 KU KU NNP 32505 504 11 86356 86356 CD 32505 504 12 - - SYM 32505 504 13 77 77 CD 32505 504 14 , , , 32505 504 15 87667 87667 CD 32505 504 16 - - SYM 32505 504 17 8 8 CD 32505 504 18 ( ( -LRB- 32505 504 19 young young JJ 32505 504 20 ) ) -RRB- 32505 504 21 . . . 32505 505 1 _ _ NNP 32505 505 2 Heredia Heredia NNP 32505 505 3 _ _ NNP 32505 505 4 : : : 32505 505 5 Puerto Puerto NNP 32505 505 6 Viejo Viejo NNP 32505 505 7 , , , 32505 505 8 KU KU NNP 32505 505 9 36696 36696 CD 32505 505 10 , , , 32505 505 11 46466 46466 CD 32505 505 12 , , , 32505 505 13 64501 64501 CD 32505 505 14 - - SYM 32505 505 15 17 17 CD 32505 505 16 , , , 32505 505 17 68288 68288 CD 32505 505 18 - - SYM 32505 505 19 91 91 CD 32505 505 20 , , , 32505 505 21 68387 68387 CD 32505 505 22 , , , 32505 505 23 68388 68388 CD 32505 505 24 - - SYM 32505 505 25 9 9 CD 32505 505 26 ( ( -LRB- 32505 505 27 young young JJ 32505 505 28 ) ) -RRB- 32505 505 29 , , , 32505 505 30 91803 91803 CD 32505 505 31 ( ( -LRB- 32505 505 32 young young JJ 32505 505 33 ) ) -RRB- 32505 505 34 , , , 32505 505 35 91688 91688 CD 32505 505 36 - - SYM 32505 505 37 9 9 CD 32505 505 38 , , , 32505 505 39 104134 104134 CD 32505 505 40 ( ( -LRB- 32505 505 41 tadpoles tadpole NNS 32505 505 42 ) ) -RRB- 32505 505 43 , , , 32505 505 44 104135 104135 CD 32505 505 45 ( ( -LRB- 32505 505 46 young young JJ 32505 505 47 ) ) -RRB- 32505 505 48 , , , 32505 505 49 104354 104354 CD 32505 505 50 - - SYM 32505 505 51 6 6 CD 32505 505 52 ( ( -LRB- 32505 505 53 skeletons skeleton NNS 32505 505 54 ) ) -RRB- 32505 505 55 ; ; : 32505 505 56 1.5 1.5 CD 32505 505 57 km km NN 32505 505 58 N N NNP 32505 505 59 Puerto Puerto NNP 32505 505 60 Viejo Viejo NNP 32505 505 61 , , , 32505 505 62 KU KU NNP 32505 505 63 64518 64518 CD 32505 505 64 - - HYPH 32505 505 65 23 23 CD 32505 505 66 , , , 32505 505 67 68386 68386 CD 32505 505 68 ( ( -LRB- 32505 505 69 tadpoles tadpole NNS 32505 505 70 ) ) -RRB- 32505 505 71 ; ; : 32505 505 72 1 1 LS 32505 505 73 km km NN 32505 505 74 S S NNP 32505 505 75 Puerto Puerto NNP 32505 505 76 Viejo Viejo NNP 32505 505 77 , , , 32505 505 78 KU KU NNP 32505 505 79 84985 84985 CD 32505 505 80 - - SYM 32505 505 81 6 6 CD 32505 505 82 ( ( -LRB- 32505 505 83 skeletons skeleton NNS 32505 505 84 ) ) -RRB- 32505 505 85 , , , 32505 505 86 87669 87669 CD 32505 505 87 ( ( -LRB- 32505 505 88 young young JJ 32505 505 89 ) ) -RRB- 32505 505 90 , , , 32505 505 91 87772 87772 CD 32505 505 92 - - SYM 32505 505 93 3 3 CD 32505 505 94 ( ( -LRB- 32505 505 95 skeletons skeleton NNS 32505 505 96 ) ) -RRB- 32505 505 97 . . . 32505 506 1 _ _ NNP 32505 506 2 Limón Limón NNP 32505 506 3 _ _ NNP 32505 506 4 : : : 32505 506 5 Bataán Bataán NNP 32505 506 6 , , , 32505 506 7 KU KU NNP 32505 506 8 30327 30327 CD 32505 506 9 - - SYM 32505 506 10 36 36 CD 32505 506 11 ; ; : 32505 506 12 La La NNP 32505 506 13 Lola Lola NNP 32505 506 14 , , , 32505 506 15 KU KU NNP 32505 506 16 64478 64478 CD 32505 506 17 - - SYM 32505 506 18 98 98 CD 32505 506 19 , , , 32505 506 20 68281 68281 CD 32505 506 21 - - HYPH 32505 506 22 2 2 CD 32505 506 23 ( ( -LRB- 32505 506 24 skeletons skeleton NNS 32505 506 25 ) ) -RRB- 32505 506 26 ; ; : 32505 506 27 Los Los NNP 32505 506 28 Diamantes Diamantes NNP 32505 506 29 , , , 32505 506 30 KU KU NNP 32505 506 31 31800 31800 CD 32505 506 32 - - HYPH 32505 506 33 02 02 CD 32505 506 34 , , , 32505 506 35 64476 64476 CD 32505 506 36 - - SYM 32505 506 37 7 7 CD 32505 506 38 ; ; : 32505 506 39 Peralta Peralta NNP 32505 506 40 , , , 32505 506 41 KU KU NNP 32505 506 42 31816 31816 CD 32505 506 43 - - SYM 32505 506 44 21 21 CD 32505 506 45 ; ; : 32505 506 46 Puerto Puerto NNP 32505 506 47 Limón Limón NNP 32505 506 48 , , , 32505 506 49 KU KU NNP 32505 506 50 31792 31792 CD 32505 506 51 - - SYM 32505 506 52 99 99 CD 32505 506 53 ; ; : 32505 506 54 Suretka Suretka NNP 32505 506 55 , , , 32505 506 56 KU KU NNP 32505 506 57 36467 36467 CD 32505 506 58 - - SYM 32505 506 59 79 79 CD 32505 506 60 , , , 32505 506 61 36697 36697 CD 32505 506 62 , , , 32505 506 63 41084 41084 CD 32505 506 64 . . . 32505 507 1 _ _ NNP 32505 507 2 Puntarenas Puntarenas NNP 32505 507 3 _ _ NNP 32505 507 4 : : : 32505 507 5 5 5 CD 32505 507 6 km km NN 32505 507 7 NW NW NNP 32505 507 8 Buenos Buenos NNP 32505 507 9 Aires Aires NNP 32505 507 10 , , , 32505 507 11 KU KU NNP 32505 507 12 107057 107057 CD 32505 507 13 ; ; : 32505 507 14 10 10 CD 32505 507 15 km km NN 32505 507 16 E E NNP 32505 507 17 Esparta Esparta NNP 32505 507 18 , , , 32505 507 19 KU KU NNP 32505 507 20 87666 87666 CD 32505 507 21 ( ( -LRB- 32505 507 22 tadpoles tadpole NNS 32505 507 23 ) ) -RRB- 32505 507 24 ; ; : 32505 507 25 Golfito Golfito NNP 32505 507 26 , , , 32505 507 27 KU KU NNP 32505 507 28 32166 32166 CD 32505 507 29 - - SYM 32505 507 30 8 8 CD 32505 507 31 ; ; : 32505 507 32 8 8 CD 32505 507 33 km km NN 32505 507 34 E E NNP 32505 507 35 Palmer Palmer NNP 32505 507 36 Norte Norte NNP 32505 507 37 KU KU NNP 32505 507 38 93939 93939 CD 32505 507 39 ; ; : 32505 507 40 10.7 10.7 CD 32505 507 41 km km NN 32505 507 42 SE SE NNP 32505 507 43 Palmar Palmar NNP 32505 507 44 Sur Sur NNP 32505 507 45 , , , 32505 507 46 KU KU NNP 32505 507 47 93938 93938 CD 32505 507 48 ( ( -LRB- 32505 507 49 skeleton skeleton NNP 32505 507 50 ) ) -RRB- 32505 507 51 , , , 32505 507 52 93940 93940 CD 32505 507 53 - - SYM 32505 507 54 51 51 CD 32505 507 55 , , , 32505 507 56 93952 93952 CD 32505 507 57 ( ( -LRB- 32505 507 58 eggs egg NNS 32505 507 59 ) ) -RRB- 32505 507 60 , , , 32505 507 61 93953 93953 CD 32505 507 62 - - SYM 32505 507 63 6 6 CD 32505 507 64 ( ( -LRB- 32505 507 65 tadpoles tadpole NNS 32505 507 66 ) ) -RRB- 32505 507 67 ; ; : 32505 507 68 Piedras Piedras NNP 32505 507 69 Blancas Blancas NNP 32505 507 70 , , , 32505 507 71 KU KU NNP 32505 507 72 103646 103646 CD 32505 507 73 - - SYM 32505 507 74 59 59 CD 32505 507 75 ; ; : 32505 507 76 4.5 4.5 CD 32505 507 77 km km NN 32505 507 78 W W NNP 32505 507 79 Rincón Rincón NNP 32505 507 80 de de NNP 32505 507 81 Osa Osa NNP 32505 507 82 , , , 32505 507 83 KU KU NNP 32505 507 84 102208 102208 CD 32505 507 85 - - SYM 32505 507 86 41 41 CD 32505 507 87 , , , 32505 507 88 104298 104298 CD 32505 507 89 ( ( -LRB- 32505 507 90 tadpoles tadpole NNS 32505 507 91 ) ) -RRB- 32505 507 92 . . . 32505 508 1 [ [ -LRB- 32505 508 2 Illustration illustration NN 32505 508 3 : : : 32505 508 4 Fig Fig NNP 32505 508 5 . . . 32505 509 1 6 6 CD 32505 509 2 . . . 32505 510 1 Map Map NNP 32505 510 2 showing show VBG 32505 510 3 locality locality NN 32505 510 4 records record NNS 32505 510 5 for for IN 32505 510 6 _ _ NNP 32505 510 7 Hyla Hyla NNP 32505 510 8 elaeochroa elaeochroa NN 32505 510 9 _ _ NNP 32505 510 10 ( ( -LRB- 32505 510 11 circles circle NNS 32505 510 12 ) ) -RRB- 32505 510 13 and and CC 32505 510 14 _ _ NNP 32505 510 15 H. H. NNP 32505 511 1 rubra rubra NNP 32505 511 2 _ _ NNP 32505 511 3 ( ( -LRB- 32505 511 4 dots dot NNS 32505 511 5 ) ) -RRB- 32505 511 6 . . . 32505 511 7 ] ] -RRB- 32505 512 1 Panamá panamá NN 32505 512 2 : : : 32505 512 3 _ _ NNP 32505 512 4 Bocas Bocas NNP 32505 512 5 del del NNP 32505 512 6 Toro Toro NNP 32505 512 7 _ _ NNP 32505 512 8 : : : 32505 512 9 Almirante Almirante NNP 32505 512 10 , , , 32505 512 11 KU KU NNP 32505 512 12 80079 80079 CD 32505 512 13 ; ; : 32505 512 14 Isla Isla NNP 32505 512 15 Bastimentos Bastimentos NNP 32505 512 16 , , , 32505 512 17 KU KU NNP 32505 512 18 96008 96008 CD 32505 512 19 - - SYM 32505 512 20 11 11 CD 32505 512 21 ; ; : 32505 512 22 Río Río NNP 32505 512 23 Cricamola Cricamola NNP 32505 512 24 , , , 32505 512 25 3.7 3.7 CD 32505 512 26 km km NN 32505 512 27 from from IN 32505 512 28 coast coast NN 32505 512 29 , , , 32505 512 30 KU KU NNP 32505 512 31 96012 96012 CD 32505 512 32 . . . 32505 513 1 _ _ NNP 32505 513 2 Chiriquí Chiriquí NNP 32505 513 3 _ _ NNP 32505 513 4 : : : 32505 513 5 Río Río NNP 32505 513 6 Gariché Gariché NNP 32505 513 7 , , , 32505 513 8 8.3 8.3 CD 32505 513 9 km km NN 32505 513 10 ESE ESE NNP 32505 513 11 Paso Paso NNP 32505 513 12 de de NNP 32505 513 13 Canoas Canoas NNP 32505 513 14 , , , 32505 513 15 KU KU NNP 32505 513 16 101571 101571 CD 32505 513 17 - - SYM 32505 513 18 2 2 CD 32505 513 19 . . . 32505 514 1 _ _ NNP 32505 514 2 Hyla Hyla NNP 32505 514 3 staufferi staufferi NN 32505 514 4 _ _ NNP 32505 514 5 Cope Cope NNP 32505 514 6 _ _ NNP 32505 514 7 Hyla Hyla NNP 32505 514 8 staufferi staufferi NN 32505 514 9 _ _ NNP 32505 514 10 Cope Cope NNP 32505 514 11 , , , 32505 514 12 Proc Proc NNP 32505 514 13 . . . 32505 515 1 Acad Acad NNS 32505 515 2 . . . 32505 516 1 Nat Nat NNP 32505 516 2 . . . 32505 517 1 Sci Sci NNP 32505 517 2 . . . 32505 518 1 Philadelphia Philadelphia NNP 32505 518 2 , , , 32505 518 3 17:165 17:165 CD 32505 518 4 , , , 32505 518 5 October October NNP 32505 518 6 1 1 CD 32505 518 7 , , , 32505 518 8 1865 1865 CD 32505 518 9 [ [ -LRB- 32505 518 10 Holotype.--USNM Holotype.--USNM NNP 32505 518 11 15317 15317 CD 32505 518 12 , , , 32505 518 13 Orizaba Orizaba NNP 32505 518 14 , , , 32505 518 15 Veracruz Veracruz NNP 32505 518 16 , , , 32505 518 17 México México NNP 32505 518 18 ; ; : 32505 518 19 Francis Francis NNP 32505 518 20 Sumichrast Sumichrast NNP 32505 518 21 collector collector NN 32505 518 22 ] ] -RRB- 32505 518 23 . . . 32505 519 1 _ _ NNP 32505 519 2 Diagnosis._--Small Diagnosis._--Small NNP 32505 519 3 frogs frog NNS 32505 519 4 ( ( -LRB- 32505 519 5 Male male NN 32505 519 6 to to TO 32505 519 7 29 29 CD 32505 519 8 mm mm NNS 32505 519 9 . . NNP 32505 519 10 , , , 32505 519 11 Female female JJ 32505 519 12 to to IN 32505 519 13 31.6 31.6 CD 32505 519 14 mm mm NNS 32505 519 15 . . . 32505 520 1 ) ) -RRB- 32505 520 2 ; ; : 32505 520 3 skull skull NNP 32505 520 4 longer long RBR 32505 520 5 than than IN 32505 520 6 wide wide JJ 32505 520 7 ; ; : 32505 520 8 palatine palatine JJ 32505 520 9 absent absent NN 32505 520 10 ; ; : 32505 520 11 large large JJ 32505 520 12 cartilaginous cartilaginous JJ 32505 520 13 crista crista NNP 32505 520 14 parotica parotica NN 32505 520 15 present present NN 32505 520 16 ; ; : 32505 520 17 snout snout NN 32505 520 18 flat flat JJ 32505 520 19 , , , 32505 520 20 elongate elongate NN 32505 520 21 and and CC 32505 520 22 protruding protrude VBG 32505 520 23 ; ; : 32505 520 24 dark dark JJ 32505 520 25 interorbital interorbital NNP 32505 520 26 bar bar NN 32505 520 27 and and CC 32505 520 28 dorsal dorsal NN 32505 520 29 stripes stripe NNS 32505 520 30 usually usually RB 32505 520 31 present present JJ 32505 520 32 . . . 32505 521 1 _ _ NNP 32505 521 2 Description._--Head Description._--Head NNP 32505 521 3 flat flat RB 32505 521 4 , , , 32505 521 5 especially especially RB 32505 521 6 in in IN 32505 521 7 females female NNS 32505 521 8 , , , 32505 521 9 longer long JJR 32505 521 10 than than IN 32505 521 11 wide wide JJ 32505 521 12 ; ; : 32505 521 13 snout snout NN 32505 521 14 long long RB 32505 521 15 , , , 32505 521 16 protruding protrude VBG 32505 521 17 beyond beyond IN 32505 521 18 mouth mouth NN 32505 521 19 ; ; , 32505 521 20 loreal loreal NN 32505 521 21 region region NN 32505 521 22 concave concave NNP 32505 521 23 ; ; : 32505 521 24 canthus canthus NNP 32505 521 25 ill ill RB 32505 521 26 - - HYPH 32505 521 27 defined define VBN 32505 521 28 ; ; : 32505 521 29 length length NN 32505 521 30 of of IN 32505 521 31 eye eye NN 32505 521 32 greater great JJR 32505 521 33 than than IN 32505 521 34 internarial internarial JJ 32505 521 35 distance distance NN 32505 521 36 or or CC 32505 521 37 width width NN 32505 521 38 of of IN 32505 521 39 eyelid eyelid NN 32505 521 40 ; ; : 32505 521 41 length length NN 32505 521 42 of of IN 32505 521 43 eye eye NN 32505 521 44 less less JJR 32505 521 45 than than IN 32505 521 46 interorbital interorbital JJ 32505 521 47 space space NN 32505 521 48 ; ; : 32505 521 49 tympanum tympanum NN 32505 521 50 distinct distinct JJ 32505 521 51 ; ; : 32505 521 52 interorbital interorbital JJ 32505 521 53 spot spot NN 32505 521 54 irregular irregular JJ 32505 521 55 ; ; : 32505 521 56 supratympanic supratympanic JJ 32505 521 57 fold fold VB 32505 521 58 faint faint JJ 32505 521 59 ; ; : 32505 521 60 arms arm NNS 32505 521 61 short short JJ 32505 521 62 ; ; : 32505 521 63 fingers finger NNS 32505 521 64 free free JJ 32505 521 65 of of IN 32505 521 66 webs webs NN 32505 521 67 ; ; : 32505 521 68 discs disc NNS 32505 521 69 on on IN 32505 521 70 third third JJ 32505 521 71 and and CC 32505 521 72 fourth fourth JJ 32505 521 73 fingers finger NNS 32505 521 74 equal equal JJ 32505 521 75 to to IN 32505 521 76 diameter diameter NN 32505 521 77 of of IN 32505 521 78 tympanum tympanum NN 32505 521 79 ; ; : 32505 521 80 inner inner JJ 32505 521 81 metatarsal metatarsal NN 32505 521 82 tubercle tubercle NN 32505 521 83 on on IN 32505 521 84 base base NN 32505 521 85 of of IN 32505 521 86 first first NNP 32505 521 87 finger finger NN 32505 521 88 distinct distinct JJ 32505 521 89 ; ; : 32505 521 90 first first RB 32505 521 91 finger finger NN 32505 521 92 shorter short JJR 32505 521 93 than than IN 32505 521 94 second second JJ 32505 521 95 ; ; : 32505 521 96 palmar palmar NNP 32505 521 97 tubercle tubercle NN 32505 521 98 distinct distinct JJ 32505 521 99 ( ( -LRB- 32505 521 100 Fig fig NN 32505 521 101 . . . 32505 522 1 1C 1C NNP 32505 522 2 ) ) -RRB- 32505 522 3 ; ; : 32505 522 4 legs leg NNS 32505 522 5 short short JJ 32505 522 6 ( ( -LRB- 32505 522 7 usually usually RB 32505 522 8 less less JJR 32505 522 9 than than IN 32505 522 10 50 50 CD 32505 522 11 per per IN 32505 522 12 cent cent NN 32505 522 13 of of IN 32505 522 14 snout snout NN 32505 522 15 - - HYPH 32505 522 16 vent vent NN 32505 522 17 length length NN 32505 522 18 ) ) -RRB- 32505 522 19 ; ; : 32505 522 20 tarsal tarsal JJ 32505 522 21 fold fold VB 32505 522 22 absent absent JJ 32505 522 23 ; ; : 32505 522 24 metatarsal metatarsal NN 32505 522 25 tubercles tubercle VBZ 32505 522 26 small small JJ 32505 522 27 , , , 32505 522 28 outer outer JJ 32505 522 29 tubercle tubercle NN 32505 522 30 smaller small JJR 32505 522 31 than than IN 32505 522 32 inner inner JJ 32505 522 33 ; ; : 32505 522 34 subarticular subarticular JJ 32505 522 35 tubercles tubercle NNS 32505 522 36 small small JJ 32505 522 37 , , , 32505 522 38 simple simple JJ 32505 522 39 , , , 32505 522 40 distinct distinct JJ 32505 522 41 ; ; : 32505 522 42 toes toe VBZ 32505 522 43 less less JJR 32505 522 44 than than IN 32505 522 45 half half NN 32505 522 46 webbed web VBN 32505 522 47 ( ( -LRB- 32505 522 48 Fig fig NN 32505 522 49 . . . 32505 523 1 1D 1D NNP 32505 523 2 ) ) -RRB- 32505 523 3 ; ; : 32505 523 4 skin skin NN 32505 523 5 smooth smooth JJ 32505 523 6 above above RB 32505 523 7 with with IN 32505 523 8 a a DT 32505 523 9 few few JJ 32505 523 10 small small JJ 32505 523 11 pustules pustule NNS 32505 523 12 on on IN 32505 523 13 head head NN 32505 523 14 , , , 32505 523 15 scapular scapular JJ 32505 523 16 region region NN 32505 523 17 , , , 32505 523 18 flanks flank NNS 32505 523 19 , , , 32505 523 20 and and CC 32505 523 21 supratympanic supratympanic JJ 32505 523 22 region region NN 32505 523 23 ; ; : 32505 523 24 arms arm NNS 32505 523 25 and and CC 32505 523 26 legs leg NNS 32505 523 27 smooth smooth JJ 32505 523 28 ; ; : 32505 523 29 skin skin NN 32505 523 30 of of IN 32505 523 31 belly belly NN 32505 523 32 coarsely coarsely RB 32505 523 33 granular granular JJ 32505 523 34 ; ; : 32505 523 35 posteroventral posteroventral JJ 32505 523 36 surfaces surface NNS 32505 523 37 of of IN 32505 523 38 thighs thigh NNS 32505 523 39 finely finely RB 32505 523 40 granular granular JJ 32505 523 41 ; ; : 32505 523 42 tongue tongue NN 32505 523 43 small small JJ 32505 523 44 , , , 32505 523 45 rounded rounded JJ 32505 523 46 , , , 32505 523 47 longer long JJR 32505 523 48 than than IN 32505 523 49 wide wide JJ 32505 523 50 , , , 32505 523 51 slightly slightly RB 32505 523 52 free free JJ 32505 523 53 and and CC 32505 523 54 notched notch VBN 32505 523 55 posteriorly posteriorly RB 32505 523 56 ; ; : 32505 523 57 vocal vocal NN 32505 523 58 slits slit NNS 32505 523 59 small small JJ 32505 523 60 , , , 32505 523 61 lateral lateral JJ 32505 523 62 to to IN 32505 523 63 tongue tongue NN 32505 523 64 ; ; : 32505 523 65 choanae choanae NNP 32505 523 66 moderate moderate JJ 32505 523 67 in in IN 32505 523 68 size size NN 32505 523 69 . . . 32505 524 1 _ _ NNP 32505 524 2 Variation._--The variation._--the CD 32505 524 3 largest large JJS 32505 524 4 males male NNS 32505 524 5 of of IN 32505 524 6 _ _ NNP 32505 524 7 Hyla Hyla NNP 32505 524 8 staufferi staufferi NN 32505 524 9 _ _ NNP 32505 524 10 are be VBP 32505 524 11 from from IN 32505 524 12 Jalapa Jalapa NNP 32505 524 13 , , , 32505 524 14 Guatemala Guatemala NNP 32505 524 15 , , , 32505 524 16 and and CC 32505 524 17 from from IN 32505 524 18 San San NNP 32505 524 19 Salvador Salvador NNP 32505 524 20 , , , 32505 524 21 El El NNP 32505 524 22 Salvador Salvador NNP 32505 524 23 . . . 32505 525 1 In in IN 32505 525 2 these these DT 32505 525 3 samples sample NNS 32505 525 4 the the DT 32505 525 5 average average JJ 32505 525 6 snout snout NN 32505 525 7 - - HYPH 32505 525 8 vent vent NN 32505 525 9 length length NN 32505 525 10 is be VBZ 32505 525 11 27 27 CD 32505 525 12 mm mm CD 32505 525 13 . . . 32505 526 1 In in IN 32505 526 2 Panamanian panamanian JJ 32505 526 3 specimens specimen VBZ 32505 526 4 the the DT 32505 526 5 average average JJ 32505 526 6 snout snout NN 32505 526 7 - - HYPH 32505 526 8 vent vent NN 32505 526 9 length length NN 32505 526 10 is be VBZ 32505 526 11 23.6 23.6 CD 32505 526 12 mm mm NNS 32505 526 13 . . . 32505 527 1 Slight slight JJ 32505 527 2 variation variation NN 32505 527 3 in in IN 32505 527 4 the the DT 32505 527 5 ratio ratio NN 32505 527 6 of of IN 32505 527 7 tibia tibia NNP 32505 527 8 length length NN 32505 527 9 to to TO 32505 527 10 snout snout VB 32505 527 11 - - HYPH 32505 527 12 vent vent NN 32505 527 13 length length NN 32505 527 14 exists exist VBZ 32505 527 15 throughout throughout IN 32505 527 16 the the DT 32505 527 17 range range NN 32505 527 18 ; ; : 32505 527 19 more more JJR 32505 527 20 variation variation NN 32505 527 21 exists exist VBZ 32505 527 22 in in IN 32505 527 23 the the DT 32505 527 24 ratio ratio NN 32505 527 25 of of IN 32505 527 26 the the DT 32505 527 27 diameter diameter NN 32505 527 28 of of IN 32505 527 29 the the DT 32505 527 30 tympanum tympanum NN 32505 527 31 to to IN 32505 527 32 that that DT 32505 527 33 of of IN 32505 527 34 the the DT 32505 527 35 eye eye NN 32505 527 36 ; ; : 32505 527 37 the the DT 32505 527 38 tympanum tympanum NN 32505 527 39 is be VBZ 32505 527 40 proportionately proportionately RB 32505 527 41 larger large JJR 32505 527 42 in in IN 32505 527 43 northern northern JJ 32505 527 44 populations population NNS 32505 527 45 ( ( -LRB- 32505 527 46 Table table NN 32505 527 47 7 7 CD 32505 527 48 ) ) -RRB- 32505 527 49 . . . 32505 528 1 The the DT 32505 528 2 primary primary JJ 32505 528 3 differences difference NNS 32505 528 4 between between IN 32505 528 5 Panamanian Panamanian NNP 32505 528 6 and and CC 32505 528 7 more more JJR 32505 528 8 northern northern JJ 32505 528 9 populations population NNS 32505 528 10 are be VBP 32505 528 11 in in IN 32505 528 12 size size NN 32505 528 13 , , , 32505 528 14 color color NN 32505 528 15 pattern pattern NN 32505 528 16 on on IN 32505 528 17 the the DT 32505 528 18 dorsum dorsum NN 32505 528 19 and and CC 32505 528 20 shanks shank NNS 32505 528 21 , , , 32505 528 22 amount amount NN 32505 528 23 of of IN 32505 528 24 webbing web VBG 32505 528 25 between between IN 32505 528 26 the the DT 32505 528 27 toes toe NNS 32505 528 28 , , , 32505 528 29 and and CC 32505 528 30 duration duration NN 32505 528 31 of of IN 32505 528 32 notes note NNS 32505 528 33 in in IN 32505 528 34 the the DT 32505 528 35 mating mating NN 32505 528 36 call call NN 32505 528 37 ( ( -LRB- 32505 528 38 Table table NN 32505 528 39 2 2 CD 32505 528 40 , , , 32505 528 41 Pl Pl NNP 32505 528 42 . . . 32505 529 1 4 4 LS 32505 529 2 ) ) -RRB- 32505 529 3 . . . 32505 530 1 The the DT 32505 530 2 color color NN 32505 530 3 in in IN 32505 530 4 Panamanian Panamanian NNP 32505 530 5 _ _ NNP 32505 530 6 staufferi staufferi NN 32505 530 7 _ _ NNP 32505 530 8 is be VBZ 32505 530 9 gray gray JJ 32505 530 10 or or CC 32505 530 11 gray gray JJ 32505 530 12 - - HYPH 32505 530 13 brown brown JJ 32505 530 14 with with IN 32505 530 15 a a DT 32505 530 16 pair pair NN 32505 530 17 of of IN 32505 530 18 distinct distinct JJ 32505 530 19 , , , 32505 530 20 complete complete JJ 32505 530 21 , , , 32505 530 22 dark dark JJ 32505 530 23 brown brown JJ 32505 530 24 dorsolateral dorsolateral JJ 32505 530 25 stripes stripe NNS 32505 530 26 , , , 32505 530 27 a a DT 32505 530 28 pair pair NN 32505 530 29 of of IN 32505 530 30 entire entire JJ 32505 530 31 paravertebral paravertebral JJ 32505 530 32 stripes stripe NNS 32505 530 33 , , , 32505 530 34 and and CC 32505 530 35 in in IN 32505 530 36 some some DT 32505 530 37 specimens specimen NNS 32505 530 38 a a DT 32505 530 39 vertebral vertebral JJ 32505 530 40 stripe stripe NN 32505 530 41 . . . 32505 531 1 About about RB 32505 531 2 five five CD 32505 531 3 per per IN 32505 531 4 cent cent NN 32505 531 5 of of IN 32505 531 6 the the DT 32505 531 7 individuals individual NNS 32505 531 8 have have VBP 32505 531 9 interrupted interrupt VBN 32505 531 10 stripes stripe NNS 32505 531 11 on on IN 32505 531 12 the the DT 32505 531 13 dorsum dorsum NN 32505 531 14 , , , 32505 531 15 whereas whereas IN 32505 531 16 in in IN 32505 531 17 the the DT 32505 531 18 more more RBR 32505 531 19 northern northern JJ 32505 531 20 populations population NNS 32505 531 21 complete complete JJ 32505 531 22 paravertebral paravertebral JJ 32505 531 23 stripes stripe NNS 32505 531 24 are be VBP 32505 531 25 present present JJ 32505 531 26 in in IN 32505 531 27 less less JJR 32505 531 28 ten ten CD 32505 531 29 per per IN 32505 531 30 cent cent NN 32505 531 31 of of IN 32505 531 32 the the DT 32505 531 33 specimens specimen NNS 32505 531 34 ; ; : 32505 531 35 when when WRB 32505 531 36 complete complete JJ 32505 531 37 stripes stripe NNS 32505 531 38 are be VBP 32505 531 39 present present JJ 32505 531 40 , , , 32505 531 41 they -PRON- PRP 32505 531 42 are be VBP 32505 531 43 irregular irregular JJ 32505 531 44 . . . 32505 532 1 The the DT 32505 532 2 dorsal dorsal NN 32505 532 3 ground ground NN 32505 532 4 color color NN 32505 532 5 in in IN 32505 532 6 non non JJ 32505 532 7 - - JJ 32505 532 8 Panamanian panamanian JJ 32505 532 9 specimens specimen NNS 32505 532 10 is be VBZ 32505 532 11 brown brown JJ 32505 532 12 , , , 32505 532 13 olive olive JJ 32505 532 14 - - HYPH 32505 532 15 brown brown JJ 32505 532 16 , , , 32505 532 17 or or CC 32505 532 18 dark dark JJ 32505 532 19 brown brown NN 32505 532 20 . . . 32505 533 1 Transverse Transverse NNP 32505 533 2 bars bar NNS 32505 533 3 are be VBP 32505 533 4 present present JJ 32505 533 5 on on IN 32505 533 6 the the DT 32505 533 7 shanks shank NNS 32505 533 8 in in IN 32505 533 9 _ _ NNP 32505 533 10 Hyla Hyla NNP 32505 533 11 staufferi staufferi NN 32505 533 12 _ _ NNP 32505 533 13 from from IN 32505 533 14 Costa Costa NNP 32505 533 15 Rica Rica NNP 32505 533 16 northward northward RB 32505 533 17 to to IN 32505 533 18 México México NNP 32505 533 19 , , , 32505 533 20 whereas whereas IN 32505 533 21 in in IN 32505 533 22 Panamá Panamá NNP 32505 533 23 all all PDT 32505 533 24 the the DT 32505 533 25 individuals individual NNS 32505 533 26 have have VBP 32505 533 27 a a DT 32505 533 28 longitudinal longitudinal JJ 32505 533 29 stripe stripe NN 32505 533 30 on on IN 32505 533 31 the the DT 32505 533 32 shank shank JJ 32505 533 33 ( ( -LRB- 32505 533 34 Table table NN 32505 533 35 7 7 CD 32505 533 36 , , , 32505 533 37 Pl Pl NNP 32505 533 38 . . . 32505 534 1 2 2 LS 32505 534 2 ) ) -RRB- 32505 534 3 . . . 32505 535 1 The the DT 32505 535 2 interorbital interorbital JJ 32505 535 3 spot spot NN 32505 535 4 or or CC 32505 535 5 bar bar NN 32505 535 6 is be VBZ 32505 535 7 more more RBR 32505 535 8 noticeable noticeable JJ 32505 535 9 in in IN 32505 535 10 northern northern JJ 32505 535 11 populations population NNS 32505 535 12 than than IN 32505 535 13 in in IN 32505 535 14 specimens specimen NNS 32505 535 15 from from IN 32505 535 16 Panamá Panamá NNP 32505 535 17 . . . 32505 536 1 Frogs frog NNS 32505 536 2 from from IN 32505 536 3 Costa Costa NNP 32505 536 4 Rica Rica NNP 32505 536 5 and and CC 32505 536 6 northward northward RB 32505 536 7 have have VBP 32505 536 8 the the DT 32505 536 9 toes toe NNS 32505 536 10 about about IN 32505 536 11 three three CD 32505 536 12 fourths fourth NNS 32505 536 13 webbed web VBN 32505 536 14 , , , 32505 536 15 whereas whereas IN 32505 536 16 in in IN 32505 536 17 Panamá Panamá NNP 32505 536 18 the the DT 32505 536 19 toes toe NNS 32505 536 20 are be VBP 32505 536 21 about about RB 32505 536 22 two two CD 32505 536 23 fifths fifth NNS 32505 536 24 webbed web VBN 32505 536 25 . . . 32505 537 1 The the DT 32505 537 2 mating mating NN 32505 537 3 calls call NNS 32505 537 4 of of IN 32505 537 5 the the DT 32505 537 6 northern northern JJ 32505 537 7 and and CC 32505 537 8 Panamanian panamanian JJ 32505 537 9 populations population NNS 32505 537 10 are be VBP 32505 537 11 similar similar JJ 32505 537 12 , , , 32505 537 13 but but CC 32505 537 14 the the DT 32505 537 15 notes note NNS 32505 537 16 have have VBP 32505 537 17 a a DT 32505 537 18 longer long JJR 32505 537 19 duration duration NN 32505 537 20 in in IN 32505 537 21 the the DT 32505 537 22 northern northern JJ 32505 537 23 populations population NNS 32505 537 24 and and CC 32505 537 25 a a DT 32505 537 26 higher high JJR 32505 537 27 dominant dominant JJ 32505 537 28 frequency frequency NN 32505 537 29 in in IN 32505 537 30 Panamanian panamanian JJ 32505 537 31 populations population NNS 32505 537 32 . . . 32505 538 1 _ _ NNP 32505 538 2 Hyla Hyla NNP 32505 538 3 staufferi staufferi NN 32505 538 4 _ _ NNP 32505 538 5 is be VBZ 32505 538 6 the the DT 32505 538 7 most most RBS 32505 538 8 variable variable JJ 32505 538 9 member member NN 32505 538 10 of of IN 32505 538 11 the the DT 32505 538 12 _ _ NNP 32505 538 13 Hyla Hyla NNP 32505 538 14 rubra rubra NN 32505 538 15 _ _ NNP 32505 538 16 group group NN 32505 538 17 in in IN 32505 538 18 Central Central NNP 32505 538 19 America America NNP 32505 538 20 . . . 32505 539 1 The the DT 32505 539 2 Panamanian panamanian JJ 32505 539 3 populations population NNS 32505 539 4 are be VBP 32505 539 5 geographically geographically RB 32505 539 6 separated separate VBN 32505 539 7 from from IN 32505 539 8 the the DT 32505 539 9 Costa costa JJ 32505 539 10 Rican rican JJ 32505 539 11 and and CC 32505 539 12 more more JJR 32505 539 13 northern northern JJ 32505 539 14 populations population NNS 32505 539 15 by by IN 32505 539 16 an an DT 32505 539 17 area area NN 32505 539 18 of of IN 32505 539 19 tropical tropical JJ 32505 539 20 rainforest rainforest NN 32505 539 21 in in IN 32505 539 22 the the DT 32505 539 23 Golfo Golfo NNP 32505 539 24 Dulce Dulce NNP 32505 539 25 region region NN 32505 539 26 in in IN 32505 539 27 southeastern southeastern JJ 32505 539 28 Costa Costa NNP 32505 539 29 Rica Rica NNP 32505 539 30 and and CC 32505 539 31 adjacent adjacent NNP 32505 539 32 Panamá Panamá NNP 32505 539 33 . . . 32505 540 1 _ _ NNP 32505 540 2 Hyla Hyla NNP 32505 540 3 staufferi staufferi NN 32505 540 4 _ _ NNP 32505 540 5 does do VBZ 32505 540 6 not not RB 32505 540 7 occur occur VB 32505 540 8 on on IN 32505 540 9 the the DT 32505 540 10 Caribbean caribbean JJ 32505 540 11 versant versant NN 32505 540 12 of of IN 32505 540 13 Costa Costa NNP 32505 540 14 Rica Rica NNP 32505 540 15 and and CC 32505 540 16 Panamá Panamá NNP 32505 540 17 . . . 32505 541 1 The the DT 32505 541 2 Golfo Golfo NNP 32505 541 3 Dulce Dulce NNP 32505 541 4 region region NN 32505 541 5 and and CC 32505 541 6 the the DT 32505 541 7 Caribbean caribbean JJ 32505 541 8 versant versant NN 32505 541 9 are be VBP 32505 541 10 humid humid JJ 32505 541 11 and and CC 32505 541 12 inhabited inhabit VBN 32505 541 13 by by IN 32505 541 14 _ _ NNP 32505 541 15 Hyla Hyla NNP 32505 541 16 elaeochroa elaeochroa NN 32505 541 17 _ _ NNP 32505 541 18 . . . 32505 542 1 _ _ NNP 32505 542 2 Hyla Hyla NNP 32505 542 3 staufferi staufferi NN 32505 542 4 _ _ NNP 32505 542 5 is be VBZ 32505 542 6 an an DT 32505 542 7 inhabitant inhabitant NN 32505 542 8 of of IN 32505 542 9 subhumid subhumid NN 32505 542 10 and and CC 32505 542 11 xeric xeric JJ 32505 542 12 areas area NNS 32505 542 13 . . . 32505 543 1 On on IN 32505 543 2 the the DT 32505 543 3 basis basis NN 32505 543 4 of of IN 32505 543 5 the the DT 32505 543 6 discontinuous discontinuous JJ 32505 543 7 variation variation NN 32505 543 8 in in IN 32505 543 9 several several JJ 32505 543 10 characters character NNS 32505 543 11 which which WDT 32505 543 12 correlate correlate VBP 32505 543 13 with with IN 32505 543 14 the the DT 32505 543 15 disjunct disjunct JJ 32505 543 16 distribution distribution NN 32505 543 17 of of IN 32505 543 18 the the DT 32505 543 19 two two CD 32505 543 20 populations population NNS 32505 543 21 , , , 32505 543 22 two two CD 32505 543 23 subspecies subspecie NNS 32505 543 24 of of IN 32505 543 25 _ _ NNP 32505 543 26 Hyla Hyla NNP 32505 543 27 staufferi staufferi NN 32505 543 28 _ _ NNP 32505 543 29 are be VBP 32505 543 30 recognized recognize VBN 32505 543 31 . . . 32505 544 1 The the DT 32505 544 2 accounts account NNS 32505 544 3 that that WDT 32505 544 4 follow follow VBP 32505 544 5 apply apply VBP 32505 544 6 equally equally RB 32505 544 7 to to IN 32505 544 8 each each DT 32505 544 9 . . . 32505 545 1 _ _ NNP 32505 545 2 Cranial Cranial NNP 32505 545 3 Osteology._--The Osteology._--The NNP 32505 545 4 skull skull NN 32505 545 5 of of IN 32505 545 6 _ _ NNP 32505 545 7 Hyla Hyla NNP 32505 545 8 staufferi staufferi NN 32505 545 9 _ _ NNP 32505 545 10 is be VBZ 32505 545 11 flat flat JJ 32505 545 12 and and CC 32505 545 13 longer long JJR 32505 545 14 than than IN 32505 545 15 wide wide JJ 32505 545 16 . . . 32505 546 1 The the DT 32505 546 2 premaxillary premaxillary NN 32505 546 3 is be VBZ 32505 546 4 small small JJ 32505 546 5 and and CC 32505 546 6 bears bear VBZ 32505 546 7 9 9 CD 32505 546 8 to to TO 32505 546 9 13 13 CD 32505 546 10 teeth tooth NNS 32505 546 11 ( ( -LRB- 32505 546 12 mean mean VB 32505 546 13 for for IN 32505 546 14 5 5 CD 32505 546 15 specimens specimen NNS 32505 546 16 , , , 32505 546 17 11.3 11.3 CD 32505 546 18 ) ) -RRB- 32505 546 19 . . . 32505 547 1 The the DT 32505 547 2 alary alary JJ 32505 547 3 process process NN 32505 547 4 of of IN 32505 547 5 the the DT 32505 547 6 premaxillary premaxillary NN 32505 547 7 is be VBZ 32505 547 8 small small JJ 32505 547 9 , , , 32505 547 10 concave concave NNP 32505 547 11 posteriorly posteriorly JJ 32505 547 12 and and CC 32505 547 13 vertical vertical JJ 32505 547 14 . . . 32505 548 1 Ventrally ventrally RB 32505 548 2 , , , 32505 548 3 the the DT 32505 548 4 premaxillary premaxillary NN 32505 548 5 is be VBZ 32505 548 6 united united JJ 32505 548 7 to to IN 32505 548 8 the the DT 32505 548 9 prevomers prevomer NNS 32505 548 10 by by IN 32505 548 11 partially partially RB 32505 548 12 ossified ossify VBN 32505 548 13 cartilage cartilage NN 32505 548 14 . . . 32505 549 1 The the DT 32505 549 2 maxillary maxillary NN 32505 549 3 is be VBZ 32505 549 4 slender slend JJR 32505 549 5 and and CC 32505 549 6 usually usually RB 32505 549 7 bears bear VBZ 32505 549 8 49 49 CD 32505 549 9 to to TO 32505 549 10 70 70 CD 32505 549 11 teeth tooth NNS 32505 549 12 ( ( -LRB- 32505 549 13 mean mean VB 32505 549 14 for for IN 32505 549 15 5 5 CD 32505 549 16 specimens specimen NNS 32505 549 17 , , , 32505 549 18 60.7 60.7 CD 32505 549 19 ) ) -RRB- 32505 549 20 . . . 32505 550 1 The the DT 32505 550 2 pars par NNS 32505 550 3 facialis faciali NNS 32505 550 4 of of IN 32505 550 5 the the DT 32505 550 6 maxillary maxillary NN 32505 550 7 is be VBZ 32505 550 8 convex convex NNP 32505 550 9 and and CC 32505 550 10 less less JJR 32505 550 11 than than IN 32505 550 12 twice twice PDT 32505 550 13 the the DT 32505 550 14 height height NN 32505 550 15 of of IN 32505 550 16 the the DT 32505 550 17 pars par NNS 32505 550 18 dentalis dentalis NNP 32505 550 19 . . . 32505 551 1 The the DT 32505 551 2 nasal nasal NN 32505 551 3 is be VBZ 32505 551 4 large large JJ 32505 551 5 , , , 32505 551 6 rounded round VBN 32505 551 7 anteriorly anteriorly RB 32505 551 8 , , , 32505 551 9 and and CC 32505 551 10 pointed point VBD 32505 551 11 posteriorly posteriorly RB 32505 551 12 in in IN 32505 551 13 dorsal dorsal NN 32505 551 14 view view NN 32505 551 15 . . . 32505 552 1 The the DT 32505 552 2 nasal nasal NN 32505 552 3 comprises comprise VBZ 32505 552 4 about about IN 32505 552 5 40 40 CD 32505 552 6 per per NN 32505 552 7 cent cent NN 32505 552 8 of of IN 32505 552 9 the the DT 32505 552 10 total total JJ 32505 552 11 length length NN 32505 552 12 of of IN 32505 552 13 the the DT 32505 552 14 skull skull NN 32505 552 15 . . . 32505 553 1 Anteromedially anteromedially RB 32505 553 2 the the DT 32505 553 3 two two CD 32505 553 4 nasals nasal NNS 32505 553 5 converge converge VBP 32505 553 6 ; ; : 32505 553 7 posteriorly posteriorly RB 32505 553 8 they -PRON- PRP 32505 553 9 overlap overlap VBP 32505 553 10 the the DT 32505 553 11 sphenethmoid sphenethmoid NN 32505 553 12 . . . 32505 554 1 The the DT 32505 554 2 nasals nasal NNS 32505 554 3 lack lack VBP 32505 554 4 a a DT 32505 554 5 concavity concavity NN 32505 554 6 in in IN 32505 554 7 the the DT 32505 554 8 midlateral midlateral JJ 32505 554 9 surface surface NN 32505 554 10 . . . 32505 555 1 Dorsally dorsally RB 32505 555 2 , , , 32505 555 3 the the DT 32505 555 4 sphenethmoid sphenethmoid NN 32505 555 5 is be VBZ 32505 555 6 wider wide JJR 32505 555 7 than than IN 32505 555 8 long long JJ 32505 555 9 , , , 32505 555 10 roughly roughly RB 32505 555 11 pentagonal pentagonal JJ 32505 555 12 in in IN 32505 555 13 shape shape NN 32505 555 14 ; ; : 32505 555 15 the the DT 32505 555 16 frontoparietal frontoparietal NN 32505 555 17 is be VBZ 32505 555 18 elongate elongate NN 32505 555 19 , , , 32505 555 20 narrow narrow JJ 32505 555 21 , , , 32505 555 22 and and CC 32505 555 23 smooth smooth JJ 32505 555 24 , , , 32505 555 25 with with IN 32505 555 26 a a DT 32505 555 27 small small JJ 32505 555 28 supraorbital supraorbital JJ 32505 555 29 process process NN 32505 555 30 anteriorly anteriorly RB 32505 555 31 . . . 32505 556 1 The the DT 32505 556 2 frontoparietal frontoparietal JJ 32505 556 3 fontanelle fontanelle NN 32505 556 4 is be VBZ 32505 556 5 narrow narrow JJ 32505 556 6 anteriorly anteriorly RB 32505 556 7 and and CC 32505 556 8 wide wide RB 32505 556 9 posteriorly posteriorly RB 32505 556 10 . . . 32505 557 1 TABLE TABLE NNP 32505 557 2 7.--Geographic 7.--Geographic NNP 32505 557 3 Variation Variation NNP 32505 557 4 in in IN 32505 557 5 Size Size NNP 32505 557 6 and and CC 32505 557 7 Color Color NNP 32505 557 8 in in IN 32505 557 9 Males Males NNPS 32505 557 10 of of IN 32505 557 11 _ _ NNP 32505 557 12 Hyla Hyla NNP 32505 557 13 staufferi staufferi NN 32505 557 14 _ _ NNP 32505 557 15 . . . 32505 558 1 ( ( -LRB- 32505 558 2 Means mean NNS 32505 558 3 in in IN 32505 558 4 parentheses parenthesis NNS 32505 558 5 below below IN 32505 558 6 observed observed JJ 32505 558 7 ranges range NNS 32505 558 8 . . . 32505 558 9 ) ) -RRB- 32505 559 1 = = NFP 32505 559 2 = = NFP 32505 559 3 = = SYM 32505 559 4 = = SYM 32505 559 5 = = SYM 32505 559 6 = = SYM 32505 559 7 = = SYM 32505 559 8 = = SYM 32505 559 9 = = SYM 32505 559 10 = = SYM 32505 559 11 = = SYM 32505 559 12 = = SYM 32505 559 13 = = SYM 32505 559 14 = = SYM 32505 559 15 = = SYM 32505 559 16 = = SYM 32505 559 17 = = SYM 32505 559 18 = = SYM 32505 559 19 = = SYM 32505 559 20 = = SYM 32505 559 21 = = SYM 32505 559 22 = = SYM 32505 559 23 = = SYM 32505 559 24 = = SYM 32505 559 25 = = SYM 32505 559 26 = = SYM 32505 559 27 = = SYM 32505 559 28 = = SYM 32505 559 29 = = SYM 32505 559 30 = = SYM 32505 559 31 = = SYM 32505 559 32 = = SYM 32505 559 33 = = SYM 32505 559 34 = = SYM 32505 559 35 = = SYM 32505 559 36 = = SYM 32505 559 37 = = SYM 32505 559 38 = = SYM 32505 559 39 = = SYM 32505 559 40 = = SYM 32505 559 41 = = SYM 32505 559 42 = = SYM 32505 559 43 = = SYM 32505 559 44 = = SYM 32505 559 45 = = SYM 32505 559 46 = = SYM 32505 559 47 = = SYM 32505 559 48 = = SYM 32505 559 49 = = SYM 32505 559 50 = = SYM 32505 559 51 = = SYM 32505 559 52 = = SYM 32505 559 53 = = SYM 32505 559 54 = = SYM 32505 559 55 = = SYM 32505 559 56 = = SYM 32505 559 57 = = SYM 32505 559 58 = = SYM 32505 559 59 = = SYM 32505 559 60 = = SYM 32505 559 61 = = SYM 32505 559 62 = = SYM 32505 559 63 = = SYM 32505 559 64 = = SYM 32505 559 65 = = SYM 32505 559 66 | | CD 32505 559 67 | | CD 32505 559 68 |Complete |complete CD 32505 559 69 dorsal| dorsal| CD 32505 559 70 | | CD 32505 559 71 |Snout |snout NN 32505 559 72 - - HYPH 32505 559 73 vent vent NN 32505 559 74 | | NNP 32505 559 75 stripes stripe NNS 32505 559 76 |Barred |barre VBN 32505 559 77 shanks shank VBZ 32505 559 78 Locality Locality NNP 32505 559 79 | | NNP 32505 559 80 N N NNP 32505 559 81 |length |length CD 32505 559 82 ( ( -LRB- 32505 559 83 mm mm NNP 32505 559 84 . . . 32505 560 1 ) ) -RRB- 32505 560 2 | | NNP 32505 560 3 ( ( -LRB- 32505 560 4 per per IN 32505 560 5 cent cent NNP 32505 560 6 ) ) -RRB- 32505 560 7 | | NNP 32505 560 8 ( ( -LRB- 32505 560 9 per per IN 32505 560 10 cent cent NNP 32505 560 11 ) ) -RRB- 32505 560 12 ---------------+-----+------------+---------------+-------------- ---------------+-----+------------+---------------+-------------- NFP 32505 560 13 Veracruz Veracruz NNP 32505 560 14 | | NNP 32505 560 15 47 47 CD 32505 560 16 | | CD 32505 560 17 23.0 23.0 CD 32505 560 18 - - HYPH 32505 560 19 27.3 27.3 CD 32505 560 20 | | CD 32505 560 21 0.0 0.0 CD 32505 560 22 | | CD 32505 560 23 100 100 CD 32505 560 24 | | CD 32505 560 25 | | CD 32505 560 26 ( ( -LRB- 32505 560 27 25.4 25.4 CD 32505 560 28 ) ) -RRB- 32505 560 29 | | CD 32505 560 30 | | CD 32505 560 31 | | CD 32505 560 32 | | CD 32505 560 33 | | CD 32505 560 34 | | NNP 32505 560 35 Campeche Campeche NNP 32505 560 36 | | NNP 32505 560 37 20 20 CD 32505 560 38 | | CD 32505 560 39 24.6 24.6 CD 32505 560 40 - - HYPH 32505 560 41 27.5 27.5 CD 32505 560 42 | | CD 32505 560 43 0.0 0.0 CD 32505 560 44 | | CD 32505 560 45 100 100 CD 32505 560 46 | | CD 32505 560 47 | | CD 32505 560 48 ( ( -LRB- 32505 560 49 25.5 25.5 CD 32505 560 50 ) ) -RRB- 32505 560 51 | | NNP 32505 560 52 | | CD 32505 560 53 | | CD 32505 560 54 | | CD 32505 560 55 | | CD 32505 560 56 | | NNP 32505 560 57 Oaxaca Oaxaca NNP 32505 560 58 | | NNP 32505 560 59 75 75 CD 32505 560 60 | | CD 32505 560 61 24.0 24.0 CD 32505 560 62 - - HYPH 32505 560 63 28.7 28.7 CD 32505 560 64 | | CD 32505 560 65 9.3 9.3 CD 32505 560 66 | | CD 32505 560 67 100 100 CD 32505 560 68 | | CD 32505 560 69 | | NNS 32505 560 70 ( ( -LRB- 32505 560 71 26.4 26.4 CD 32505 560 72 ) ) -RRB- 32505 560 73 | | NNP 32505 560 74 | | CD 32505 560 75 | | CD 32505 560 76 | | CD 32505 560 77 | | CD 32505 560 78 | | NNP 32505 560 79 Chiapas Chiapas NNP 32505 560 80 | | CD 32505 560 81 20 20 CD 32505 560 82 | | CD 32505 560 83 23.2 23.2 CD 32505 560 84 - - HYPH 32505 560 85 27.8 27.8 CD 32505 560 86 | | CD 32505 560 87 10.0 10.0 CD 32505 560 88 | | CD 32505 560 89 100 100 CD 32505 560 90 | | CD 32505 560 91 | | CD 32505 560 92 ( ( -LRB- 32505 560 93 25.5 25.5 CD 32505 560 94 ) ) -RRB- 32505 560 95 | | NNP 32505 560 96 | | CD 32505 560 97 | | CD 32505 560 98 | | CD 32505 560 99 | | CD 32505 560 100 | | NNP 32505 560 101 Guatemala Guatemala NNP 32505 560 102 | | NNP 32505 560 103 22 22 CD 32505 560 104 | | CD 32505 560 105 25.0 25.0 CD 32505 560 106 - - HYPH 32505 560 107 29.0 29.0 CD 32505 560 108 | | CD 32505 560 109 10.9 10.9 CD 32505 560 110 | | CD 32505 560 111 100 100 CD 32505 560 112 | | CD 32505 560 113 | | CD 32505 560 114 ( ( -LRB- 32505 560 115 26.9 26.9 CD 32505 560 116 ) ) -RRB- 32505 560 117 | | NNP 32505 560 118 | | CD 32505 560 119 | | CD 32505 560 120 | | CD 32505 560 121 | | CD 32505 560 122 | | CD 32505 560 123 El El NNP 32505 560 124 Salvador Salvador NNP 32505 560 125 | | NNP 32505 560 126 21 21 CD 32505 560 127 | | CD 32505 560 128 24.7 24.7 CD 32505 560 129 - - HYPH 32505 560 130 28.6 28.6 CD 32505 560 131 | | CD 32505 560 132 0.0 0.0 CD 32505 560 133 | | CD 32505 560 134 100 100 CD 32505 560 135 | | CD 32505 560 136 | | CD 32505 560 137 ( ( -LRB- 32505 560 138 27.0 27.0 CD 32505 560 139 ) ) -RRB- 32505 560 140 | | CD 32505 560 141 | | CD 32505 560 142 | | CD 32505 560 143 | | CD 32505 560 144 | | CD 32505 560 145 | | NNP 32505 560 146 Honduras Honduras NNP 32505 560 147 | | NNP 32505 560 148 34 34 CD 32505 560 149 | | CD 32505 560 150 20.6 20.6 CD 32505 560 151 - - HYPH 32505 560 152 27.0 27.0 CD 32505 560 153 | | CD 32505 560 154 3.3 3.3 CD 32505 560 155 | | CD 32505 560 156 100 100 CD 32505 560 157 | | CD 32505 560 158 | | CD 32505 560 159 ( ( -LRB- 32505 560 160 24.9 24.9 CD 32505 560 161 ) ) -RRB- 32505 560 162 | | CD 32505 560 163 | | CD 32505 560 164 | | CD 32505 560 165 | | CD 32505 560 166 | | CD 32505 560 167 | | CD 32505 560 168 Nicaragua Nicaragua NNP 32505 560 169 | | NNP 32505 560 170 67 67 CD 32505 560 171 | | CD 32505 560 172 21.5 21.5 CD 32505 560 173 - - HYPH 32505 560 174 26.8 26.8 CD 32505 560 175 | | CD 32505 560 176 3.0 3.0 CD 32505 560 177 | | CD 32505 560 178 92.7 92.7 CD 32505 560 179 | | CD 32505 560 180 | | CD 32505 560 181 ( ( -LRB- 32505 560 182 24.9 24.9 CD 32505 560 183 ) ) -RRB- 32505 560 184 | | CD 32505 560 185 | | CD 32505 560 186 | | CD 32505 560 187 | | CD 32505 560 188 | | CD 32505 560 189 | | NNP 32505 560 190 Costa Costa NNP 32505 560 191 Rica Rica NNP 32505 560 192 | | NNP 32505 560 193 54 54 CD 32505 560 194 | | CD 32505 560 195 20.7 20.7 CD 32505 560 196 - - HYPH 32505 560 197 26.6 26.6 CD 32505 560 198 | | CD 32505 560 199 5.5 5.5 CD 32505 560 200 | | CD 32505 560 201 98.1 98.1 CD 32505 560 202 | | CD 32505 560 203 | | CD 32505 560 204 ( ( -LRB- 32505 560 205 24.2 24.2 CD 32505 560 206 ) ) -RRB- 32505 560 207 | | NNP 32505 560 208 | | CD 32505 560 209 | | CD 32505 560 210 | | CD 32505 560 211 | | CD 32505 560 212 | | NNP 32505 560 213 Total total JJ 32505 560 214 | | NNP 32505 560 215 360 360 CD 32505 560 216 | | CD 32505 560 217 20.7 20.7 CD 32505 560 218 - - HYPH 32505 560 219 29.0 29.0 CD 32505 560 220 | | CD 32505 560 221 5.4 5.4 CD 32505 560 222 | | CD 32505 560 223 98.3 98.3 CD 32505 560 224 Non non JJ 32505 560 225 - - JJ 32505 560 226 Panamanian panamanian JJ 32505 560 227 | | NNP 32505 560 228 | | CD 32505 560 229 ( ( -LRB- 32505 560 230 25.9 25.9 CD 32505 560 231 ) ) -RRB- 32505 560 232 | | NNP 32505 560 233 | | CD 32505 560 234 | | CD 32505 560 235 | | CD 32505 560 236 | | CD 32505 560 237 | | NNP 32505 560 238 Panamá Panamá NNP 32505 560 239 | | NNP 32505 560 240 72 72 CD 32505 560 241 | | CD 32505 560 242 21.7 21.7 CD 32505 560 243 - - HYPH 32505 560 244 26.0 26.0 CD 32505 560 245 | | CD 32505 560 246 94.5 94.5 CD 32505 560 247 | | CD 32505 560 248 0.0 0.0 CD 32505 560 249 | | CD 32505 560 250 | | CD 32505 560 251 ( ( -LRB- 32505 560 252 23.6 23.6 CD 32505 560 253 ) ) -RRB- 32505 560 254 | | CD 32505 560 255 | | VBD 32505 560 256 Only only RB 32505 560 257 a a DT 32505 560 258 narrow narrow JJ 32505 560 259 connection connection NN 32505 560 260 exists exist VBZ 32505 560 261 between between IN 32505 560 262 the the DT 32505 560 263 posterior posterior NN 32505 560 264 , , , 32505 560 265 pointed pointed JJ 32505 560 266 arm arm NN 32505 560 267 of of IN 32505 560 268 the the DT 32505 560 269 squamosal squamosal NN 32505 560 270 and and CC 32505 560 271 the the DT 32505 560 272 lateral lateral JJ 32505 560 273 edge edge NN 32505 560 274 of of IN 32505 560 275 the the DT 32505 560 276 proötic proötic NN 32505 560 277 . . . 32505 561 1 The the DT 32505 561 2 crista crista NN 32505 561 3 parotica parotica NN 32505 561 4 is be VBZ 32505 561 5 visible visible JJ 32505 561 6 dorsally dorsally RB 32505 561 7 along along IN 32505 561 8 the the DT 32505 561 9 lateral lateral JJ 32505 561 10 edge edge NN 32505 561 11 of of IN 32505 561 12 the the DT 32505 561 13 bony bony NNP 32505 561 14 proötic proötic NN 32505 561 15 . . . 32505 562 1 The the DT 32505 562 2 squamosal squamosal NN 32505 562 3 is be VBZ 32505 562 4 narrow narrow JJ 32505 562 5 anteriorly anteriorly RB 32505 562 6 and and CC 32505 562 7 posteriorly posteriorly RB 32505 562 8 . . . 32505 563 1 The the DT 32505 563 2 prevomers prevomer NNS 32505 563 3 are be VBP 32505 563 4 short short JJ 32505 563 5 and and CC 32505 563 6 separated separate VBN 32505 563 7 anteriorly anteriorly RB 32505 563 8 by by IN 32505 563 9 partly partly RB 32505 563 10 ossified ossify VBN 32505 563 11 cartilage cartilage NN 32505 563 12 of of IN 32505 563 13 the the DT 32505 563 14 overlying overlie VBG 32505 563 15 solum solum NNP 32505 563 16 nasi nasi NNP 32505 563 17 . . . 32505 564 1 The the DT 32505 564 2 prevomer prevomer NN 32505 564 3 is be VBZ 32505 564 4 joined join VBN 32505 564 5 to to IN 32505 564 6 the the DT 32505 564 7 premaxillary premaxillary NN 32505 564 8 by by IN 32505 564 9 cartilage cartilage NN 32505 564 10 . . . 32505 565 1 The the DT 32505 565 2 posterior posterior JJ 32505 565 3 margin margin NN 32505 565 4 of of IN 32505 565 5 the the DT 32505 565 6 prevomer prevomer NN 32505 565 7 articulates articulate NNS 32505 565 8 directly directly RB 32505 565 9 with with IN 32505 565 10 the the DT 32505 565 11 sphenethmoid sphenethmoid NN 32505 565 12 . . . 32505 566 1 The the DT 32505 566 2 anterolateral anterolateral JJ 32505 566 3 and and CC 32505 566 4 posterolateral posterolateral JJ 32505 566 5 processes process NNS 32505 566 6 of of IN 32505 566 7 the the DT 32505 566 8 prevomers prevomer NNS 32505 566 9 form form VBP 32505 566 10 the the DT 32505 566 11 incomplete incomplete JJ 32505 566 12 bony bony NNP 32505 566 13 internal internal JJ 32505 566 14 margin margin NN 32505 566 15 of of IN 32505 566 16 the the DT 32505 566 17 choanae choanae NN 32505 566 18 . . . 32505 567 1 Each each DT 32505 567 2 prevomer prevomer NN 32505 567 3 bears bear VBZ 32505 567 4 three three CD 32505 567 5 to to TO 32505 567 6 six six CD 32505 567 7 teeth tooth NNS 32505 567 8 . . . 32505 568 1 The the DT 32505 568 2 palatine palatine NN 32505 568 3 is be VBZ 32505 568 4 absent absent JJ 32505 568 5 . . . 32505 569 1 The the DT 32505 569 2 anterior anterior JJ 32505 569 3 part part NN 32505 569 4 of of IN 32505 569 5 the the DT 32505 569 6 parasphenoid parasphenoid NN 32505 569 7 is be VBZ 32505 569 8 narrow narrow JJ 32505 569 9 and and CC 32505 569 10 ends end VBZ 32505 569 11 in in IN 32505 569 12 a a DT 32505 569 13 point point NN 32505 569 14 . . . 32505 570 1 The the DT 32505 570 2 pterygoid pterygoid NN 32505 570 3 is be VBZ 32505 570 4 slender slend JJR 32505 570 5 and and CC 32505 570 6 weakly weakly RB 32505 570 7 developed develop VBD 32505 570 8 . . . 32505 571 1 _ _ NNP 32505 571 2 Natural Natural NNP 32505 571 3 History._--Throughout history._--throughout CD 32505 571 4 its -PRON- PRP$ 32505 571 5 range range NN 32505 571 6 _ _ NNP 32505 571 7 Hyla Hyla NNP 32505 571 8 staufferi staufferi NN 32505 571 9 _ _ NNP 32505 571 10 occurs occur VBZ 32505 571 11 in in IN 32505 571 12 subhumid subhumid NN 32505 571 13 forests forest NNS 32505 571 14 and and CC 32505 571 15 savannas savanna NNS 32505 571 16 ; ; : 32505 571 17 consequently consequently RB 32505 571 18 , , , 32505 571 19 the the DT 32505 571 20 breeding breeding NN 32505 571 21 activities activity NNS 32505 571 22 are be VBP 32505 571 23 limited limit VBN 32505 571 24 by by IN 32505 571 25 the the DT 32505 571 26 seasonal seasonal JJ 32505 571 27 occurrence occurrence NN 32505 571 28 of of IN 32505 571 29 rainfall rainfall NN 32505 571 30 , , , 32505 571 31 which which WDT 32505 571 32 accumulates accumulate VBZ 32505 571 33 in in IN 32505 571 34 temporary temporary JJ 32505 571 35 ponds pond NNS 32505 571 36 where where WRB 32505 571 37 this this DT 32505 571 38 species species NN 32505 571 39 breeds breed NNS 32505 571 40 . . . 32505 572 1 Clasping clasp VBG 32505 572 2 pairs pair NNS 32505 572 3 and and CC 32505 572 4 gravid gravid JJ 32505 572 5 females female NNS 32505 572 6 have have VBP 32505 572 7 been be VBN 32505 572 8 found find VBN 32505 572 9 mostly mostly RB 32505 572 10 from from IN 32505 572 11 June June NNP 32505 572 12 to to IN 32505 572 13 August August NNP 32505 572 14 throughout throughout IN 32505 572 15 its -PRON- PRP$ 32505 572 16 range range NN 32505 572 17 . . . 32505 573 1 This this DT 32505 573 2 species species NN 32505 573 3 was be VBD 32505 573 4 observed observe VBN 32505 573 5 calling call VBG 32505 573 6 at at IN 32505 573 7 Finca Finca NNP 32505 573 8 Taboga Taboga NNP 32505 573 9 , , , 32505 573 10 Guanacaste Guanacaste NNP 32505 573 11 Province Province NNP 32505 573 12 , , , 32505 573 13 Costa Costa NNP 32505 573 14 Rica Rica NNP 32505 573 15 , , , 32505 573 16 in in IN 32505 573 17 mid mid NNP 32505 573 18 - - NNP 32505 573 19 July july NN 32505 573 20 . . . 32505 574 1 The the DT 32505 574 2 males male NNS 32505 574 3 were be VBD 32505 574 4 calling call VBG 32505 574 5 from from IN 32505 574 6 temporary temporary JJ 32505 574 7 grassy grassy JJ 32505 574 8 and and CC 32505 574 9 weedy weedy JJ 32505 574 10 ponds pond NNS 32505 574 11 in in IN 32505 574 12 which which WDT 32505 574 13 _ _ NNP 32505 574 14 Hyla Hyla NNP 32505 574 15 microcephala microcephala NN 32505 574 16 _ _ NNP 32505 574 17 also also RB 32505 574 18 was be VBD 32505 574 19 calling call VBG 32505 574 20 , , , 32505 574 21 but but CC 32505 574 22 the the DT 32505 574 23 two two CD 32505 574 24 species specie NNS 32505 574 25 had have VBD 32505 574 26 different different JJ 32505 574 27 calling calling NN 32505 574 28 sites site NNS 32505 574 29 . . . 32505 575 1 _ _ NNP 32505 575 2 Hyla Hyla NNP 32505 575 3 staufferi staufferi NN 32505 575 4 _ _ NNP 32505 575 5 called call VBD 32505 575 6 at at IN 32505 575 7 stations station NNS 32505 575 8 at at IN 32505 575 9 heights height NNS 32505 575 10 of of IN 32505 575 11 five five CD 32505 575 12 to to TO 32505 575 13 80 80 CD 32505 575 14 cm cm NNS 32505 575 15 . . . 32505 576 1 near near IN 32505 576 2 the the DT 32505 576 3 edge edge NN 32505 576 4 of of IN 32505 576 5 the the DT 32505 576 6 pond pond NN 32505 576 7 , , , 32505 576 8 whereas whereas IN 32505 576 9 _ _ NNP 32505 576 10 Hyla Hyla NNP 32505 576 11 microcephala microcephala NN 32505 576 12 _ _ NNP 32505 576 13 called call VBD 32505 576 14 from from IN 32505 576 15 emergent emergent JJ 32505 576 16 vegetation vegetation NN 32505 576 17 in in IN 32505 576 18 the the DT 32505 576 19 middle middle NN 32505 576 20 of of IN 32505 576 21 the the DT 32505 576 22 pond pond NN 32505 576 23 . . . 32505 577 1 Charles Charles NNP 32505 577 2 W. W. NNP 32505 577 3 Myers Myers NNP 32505 577 4 informed inform VBD 32505 577 5 me -PRON- PRP 32505 577 6 that that IN 32505 577 7 at at IN 32505 577 8 Penonomé Penonomé NNP 32505 577 9 , , , 32505 577 10 Coclé Coclé NNP 32505 577 11 , , , 32505 577 12 Panamá Panamá NNP 32505 577 13 , , , 32505 577 14 he -PRON- PRP 32505 577 15 found find VBD 32505 577 16 _ _ NNP 32505 577 17 staufferi staufferi NN 32505 577 18 _ _ NNP 32505 577 19 calling call VBG 32505 577 20 from from IN 32505 577 21 grass grass NN 32505 577 22 in in IN 32505 577 23 puddles puddle NNS 32505 577 24 where where WRB 32505 577 25 _ _ NNP 32505 577 26 microcephala microcephala NN 32505 577 27 _ _ NNP 32505 577 28 was be VBD 32505 577 29 absent absent JJ 32505 577 30 , , , 32505 577 31 and and CC 32505 577 32 at at IN 32505 577 33 El El NNP 32505 577 34 Caño Caño NNP 32505 577 35 , , , 32505 577 36 Coclé Coclé NNP 32505 577 37 , , , 32505 577 38 Panamá Panamá NNP 32505 577 39 , , , 32505 577 40 _ _ NNP 32505 577 41 staufferi staufferi NN 32505 577 42 _ _ NNP 32505 577 43 was be VBD 32505 577 44 calling call VBG 32505 577 45 from from IN 32505 577 46 higher high JJR 32505 577 47 sites site NNS 32505 577 48 ( ( -LRB- 32505 577 49 " " `` 32505 577 50 several several JJ 32505 577 51 inches inch NNS 32505 577 52 to to IN 32505 577 53 a a DT 32505 577 54 few few JJ 32505 577 55 feet foot NNS 32505 577 56 above above IN 32505 577 57 water water NN 32505 577 58 " " '' 32505 577 59 ) ) -RRB- 32505 577 60 than than IN 32505 577 61 _ _ NNP 32505 577 62 microcephala microcephala NNP 32505 577 63 _ _ NNP 32505 577 64 . . . 32505 578 1 Stuart Stuart NNP 32505 578 2 ( ( -LRB- 32505 578 3 1948:34 1948:34 CD 32505 578 4 ) ) -RRB- 32505 578 5 reported report VBD 32505 578 6 breeding breeding NN 32505 578 7 individuals individual NNS 32505 578 8 from from IN 32505 578 9 La La NNP 32505 578 10 Libertad Libertad NNP 32505 578 11 , , , 32505 578 12 Guatemala Guatemala NNP 32505 578 13 , , , 32505 578 14 after after IN 32505 578 15 rainfall rainfall NN 32505 578 16 in in IN 32505 578 17 late late JJ 32505 578 18 May May NNP 32505 578 19 , , , 32505 578 20 and and CC 32505 578 21 Schmidt Schmidt NNP 32505 578 22 and and CC 32505 578 23 Stuart Stuart NNP 32505 578 24 ( ( -LRB- 32505 578 25 1941:239 1941:239 CD 32505 578 26 ) ) -RRB- 32505 578 27 reported report VBD 32505 578 28 _ _ NNP 32505 578 29 staufferi staufferi NN 32505 578 30 _ _ NNP 32505 578 31 breeding breeding NN 32505 578 32 in in IN 32505 578 33 July July NNP 32505 578 34 in in IN 32505 578 35 the the DT 32505 578 36 Salamá Salamá NNP 32505 578 37 basin basin NN 32505 578 38 , , , 32505 578 39 Alta Alta NNP 32505 578 40 Verapaz Verapaz NNP 32505 578 41 , , , 32505 578 42 Guatemala Guatemala NNP 32505 578 43 . . . 32505 579 1 Stuart Stuart NNP 32505 579 2 ( ( -LRB- 32505 579 3 1935:38 1935:38 CD 32505 579 4 ) ) -RRB- 32505 579 5 and and CC 32505 579 6 Duellman Duellman NNP 32505 579 7 ( ( -LRB- 32505 579 8 1960:63 1960:63 CD 32505 579 9 and and CC 32505 579 10 1963:226 1963:226 CD 32505 579 11 ) ) -RRB- 32505 579 12 agreed agree VBD 32505 579 13 that that IN 32505 579 14 this this DT 32505 579 15 species species NN 32505 579 16 breeds breed VBZ 32505 579 17 early early RB 32505 579 18 in in IN 32505 579 19 the the DT 32505 579 20 rainy rainy JJ 32505 579 21 season season NN 32505 579 22 . . . 32505 580 1 However however RB 32505 580 2 , , , 32505 580 3 Rand Rand NNP 32505 580 4 ( ( -LRB- 32505 580 5 1957:519 1957:519 CD 32505 580 6 ) ) -RRB- 32505 580 7 stated state VBD 32505 580 8 that that IN 32505 580 9 in in IN 32505 580 10 El El NNP 32505 580 11 Salvador Salvador NNP 32505 580 12 " " '' 32505 580 13 these these DT 32505 580 14 frogs frog NNS 32505 580 15 did do VBD 32505 580 16 not not RB 32505 580 17 begin begin VB 32505 580 18 to to TO 32505 580 19 call call VB 32505 580 20 until until IN 32505 580 21 almost almost RB 32505 580 22 a a DT 32505 580 23 month month NN 32505 580 24 and and CC 32505 580 25 a a DT 32505 580 26 half half NN 32505 580 27 after after IN 32505 580 28 the the DT 32505 580 29 beginning beginning NN 32505 580 30 of of IN 32505 580 31 the the DT 32505 580 32 rains rain NNS 32505 580 33 . . . 32505 580 34 " " '' 32505 581 1 Blair Blair NNP 32505 581 2 ( ( -LRB- 32505 581 3 1960:133 1960:133 CD 32505 581 4 ) ) -RRB- 32505 581 5 reported report VBD 32505 581 6 that that IN 32505 581 7 males male NNS 32505 581 8 call call VBP 32505 581 9 in in IN 32505 581 10 June June NNP 32505 581 11 and and CC 32505 581 12 July July NNP 32505 581 13 in in IN 32505 581 14 Chiapas Chiapas NNP 32505 581 15 , , , 32505 581 16 Oaxaca Oaxaca NNP 32505 581 17 , , , 32505 581 18 Veracruz Veracruz NNP 32505 581 19 , , , 32505 581 20 and and CC 32505 581 21 Tamaulipas Tamaulipas NNP 32505 581 22 , , , 32505 581 23 México México NNP 32505 581 24 . . . 32505 582 1 The the DT 32505 582 2 mating mating NN 32505 582 3 call call NN 32505 582 4 of of IN 32505 582 5 this this DT 32505 582 6 species species NN 32505 582 7 is be VBZ 32505 582 8 a a DT 32505 582 9 series series NN 32505 582 10 of of IN 32505 582 11 closely closely RB 32505 582 12 spaced space VBN 32505 582 13 notes note NNS 32505 582 14 having have VBG 32505 582 15 a a DT 32505 582 16 fundamental fundamental JJ 32505 582 17 frequency frequency NN 32505 582 18 of of IN 32505 582 19 about about RB 32505 582 20 100 100 CD 32505 582 21 cycles cycle NNS 32505 582 22 per per IN 32505 582 23 second second NN 32505 582 24 . . . 32505 583 1 Each each DT 32505 583 2 note note NN 32505 583 3 has have VBZ 32505 583 4 a a DT 32505 583 5 duration duration NN 32505 583 6 of of IN 32505 583 7 0.13 0.13 CD 32505 583 8 to to IN 32505 583 9 0.23 0.23 CD 32505 583 10 second second RB 32505 583 11 , , , 32505 583 12 repeated repeat VBN 32505 583 13 at at IN 32505 583 14 intervals interval NNS 32505 583 15 that that WDT 32505 583 16 are be VBP 32505 583 17 longer long JJR 32505 583 18 than than IN 32505 583 19 the the DT 32505 583 20 duration duration NN 32505 583 21 of of IN 32505 583 22 the the DT 32505 583 23 call call NN 32505 583 24 . . . 32505 584 1 The the DT 32505 584 2 notes note NNS 32505 584 3 are be VBP 32505 584 4 moderately moderately RB 32505 584 5 low low RB 32505 584 6 - - HYPH 32505 584 7 pitched pitched JJ 32505 584 8 and and CC 32505 584 9 have have VBP 32505 584 10 a a DT 32505 584 11 dominant dominant JJ 32505 584 12 frequency frequency NN 32505 584 13 of of IN 32505 584 14 more more JJR 32505 584 15 than than IN 32505 584 16 3,000 3,000 CD 32505 584 17 cycles cycle NNS 32505 584 18 per per IN 32505 584 19 second second NN 32505 584 20 and and CC 32505 584 21 about about RB 32505 584 22 120 120 CD 32505 584 23 pulses pulse NNS 32505 584 24 per per IN 32505 584 25 second second JJ 32505 584 26 ( ( -LRB- 32505 584 27 Table table NN 32505 584 28 2 2 CD 32505 584 29 ) ) -RRB- 32505 584 30 . . . 32505 585 1 _ _ NNP 32505 585 2 Tadpoles._--Measurements tadpoles._--measurement NNS 32505 585 3 of of IN 32505 585 4 the the DT 32505 585 5 33 33 CD 32505 585 6 tadpoles tadpole NNS 32505 585 7 that that WDT 32505 585 8 are be VBP 32505 585 9 available available JJ 32505 585 10 are be VBP 32505 585 11 given give VBN 32505 585 12 in in IN 32505 585 13 Table table NN 32505 585 14 8 8 CD 32505 585 15 . . . 32505 586 1 The the DT 32505 586 2 largest large JJS 32505 586 3 tadpole tadpole NN 32505 586 4 examined examine VBN 32505 586 5 is be VBZ 32505 586 6 in in IN 32505 586 7 stage stage NN 32505 586 8 38 38 CD 32505 586 9 and and CC 32505 586 10 has have VBZ 32505 586 11 a a DT 32505 586 12 total total JJ 32505 586 13 length length NN 32505 586 14 of of IN 32505 586 15 29.5 29.5 CD 32505 586 16 mm mm CD 32505 586 17 . . . 32505 587 1 A a DT 32505 587 2 typical typical JJ 32505 587 3 tadpole tadpole NN 32505 587 4 in in IN 32505 587 5 stage stage NN 32505 587 6 38 38 CD 32505 587 7 of of IN 32505 587 8 development development NN 32505 587 9 ( ( -LRB- 32505 587 10 KU KU NNP 32505 587 11 104162 104162 CD 32505 587 12 , , , 32505 587 13 5 5 CD 32505 587 14 km km NN 32505 587 15 ESE ESE NNP 32505 587 16 Córdoba Córdoba NNP 32505 587 17 , , , 32505 587 18 Veracruz Veracruz NNP 32505 587 19 , , , 32505 587 20 México México NNP 32505 587 21 ) ) -RRB- 32505 587 22 has have VBZ 32505 587 23 a a DT 32505 587 24 body body NN 32505 587 25 length length NN 32505 587 26 of of IN 32505 587 27 10 10 CD 32505 587 28 mm mm NNS 32505 587 29 . . NNP 32505 587 30 , , , 32505 587 31 tail tail NN 32505 587 32 length length NN 32505 587 33 of of IN 32505 587 34 19.5 19.5 CD 32505 587 35 mm mm NNP 32505 587 36 . . NNP 32505 587 37 , , , 32505 587 38 and and CC 32505 587 39 a a DT 32505 587 40 total total JJ 32505 587 41 length length NN 32505 587 42 of of IN 32505 587 43 29.5 29.5 CD 32505 587 44 mm mm CD 32505 587 45 . . . 32505 588 1 Other other JJ 32505 588 2 characters character NNS 32505 588 3 are be VBP 32505 588 4 as as IN 32505 588 5 follows follow VBZ 32505 588 6 : : : 32505 588 7 body body NN 32505 588 8 as as RB 32505 588 9 deep deep JJ 32505 588 10 as as IN 32505 588 11 wide wide JJ 32505 588 12 , , , 32505 588 13 depressed depress VBN 32505 588 14 anteriorly anteriorly RB 32505 588 15 ; ; : 32505 588 16 body body NN 32505 588 17 as as RB 32505 588 18 long long RB 32505 588 19 as as IN 32505 588 20 depth depth NN 32505 588 21 of of IN 32505 588 22 tail tail NN 32505 588 23 ; ; : 32505 588 24 interorbital interorbital NN 32505 588 25 space space NN 32505 588 26 greater great JJR 32505 588 27 than than IN 32505 588 28 distance distance NN 32505 588 29 between between IN 32505 588 30 eye eye NN 32505 588 31 and and CC 32505 588 32 snout snout NN 32505 588 33 but but CC 32505 588 34 equal equal JJ 32505 588 35 to to IN 32505 588 36 internarial internarial JJ 32505 588 37 space space NN 32505 588 38 ; ; : 32505 588 39 nostril nostril NNP 32505 588 40 equidistant equidistant NN 32505 588 41 between between IN 32505 588 42 eye eye NN 32505 588 43 and and CC 32505 588 44 tip tip NN 32505 588 45 of of IN 32505 588 46 snout snout NN 32505 588 47 ; ; : 32505 588 48 distance distance NN 32505 588 49 between between IN 32505 588 50 spiracle spiracle NNP 32505 588 51 and and CC 32505 588 52 eye eye NN 32505 588 53 less less JJR 32505 588 54 than than IN 32505 588 55 distance distance NN 32505 588 56 between between IN 32505 588 57 eye eye NN 32505 588 58 and and CC 32505 588 59 snout snout NN 32505 588 60 ; ; : 32505 588 61 eyes eye NNS 32505 588 62 large large JJ 32505 588 63 , , , 32505 588 64 situated situate VBN 32505 588 65 dorsolaterally dorsolaterally RB 32505 588 66 ; ; : 32505 588 67 mouth mouth NNP 32505 588 68 anteroventral anteroventral NNP 32505 588 69 , , , 32505 588 70 approximately approximately RB 32505 588 71 triangular triangular JJ 32505 588 72 in in IN 32505 588 73 outline outline NN 32505 588 74 ; ; : 32505 588 75 one one CD 32505 588 76 row row NN 32505 588 77 of of IN 32505 588 78 papillae papillae NNP 32505 588 79 covering cover VBG 32505 588 80 lower low JJR 32505 588 81 lip lip NN 32505 588 82 and and CC 32505 588 83 all all DT 32505 588 84 except except IN 32505 588 85 median median JJ 32505 588 86 fourth fourth RB 32505 588 87 of of IN 32505 588 88 upper upper JJ 32505 588 89 lip lip NN 32505 588 90 ; ; : 32505 588 91 scattered scatter VBN 32505 588 92 papillae papillae NNP 32505 588 93 at at IN 32505 588 94 corners corner NNS 32505 588 95 of of IN 32505 588 96 mouth mouth NN 32505 588 97 ; ; , 32505 588 98 tooth tooth NN 32505 588 99 rows row VBZ 32505 588 100 2/3 2/3 CD 32505 588 101 ; ; : 32505 588 102 first first RB 32505 588 103 upper upper JJ 32505 588 104 row row NN 32505 588 105 entire entire JJ 32505 588 106 , , , 32505 588 107 second second JJ 32505 588 108 row row NN 32505 588 109 interrupted interrupt VBN 32505 588 110 medially medially RB 32505 588 111 , , , 32505 588 112 shorter short JJR 32505 588 113 than than IN 32505 588 114 first first RB 32505 588 115 ; ; : 32505 588 116 lower low JJR 32505 588 117 rows row NNS 32505 588 118 shorter short JJR 32505 588 119 than than IN 32505 588 120 upper upper JJ 32505 588 121 rows row NNS 32505 588 122 ; ; , 32505 588 123 beak beak JJ 32505 588 124 weak weak JJ 32505 588 125 ; ; : 32505 588 126 spiracle spiracle NNP 32505 588 127 short short JJ 32505 588 128 and and CC 32505 588 129 nearer nearer JJ 32505 588 130 eyes eye NNS 32505 588 131 than than IN 32505 588 132 anus anus NN 32505 588 133 ; ; : 32505 588 134 anal anal JJ 32505 588 135 opening opening NN 32505 588 136 not not RB 32505 588 137 reaching reach VBG 32505 588 138 edge edge NN 32505 588 139 of of IN 32505 588 140 ventral ventral JJ 32505 588 141 fin fin NN 32505 588 142 ; ; : 32505 588 143 dorsal dorsal NN 32505 588 144 fin fin NNP 32505 588 145 barely barely RB 32505 588 146 extending extend VBG 32505 588 147 onto onto IN 32505 588 148 body body NN 32505 588 149 ; ; : 32505 588 150 caudal caudal JJ 32505 588 151 musculature musculature NN 32505 588 152 pointed point VBD 32505 588 153 distally distally RB 32505 588 154 . . . 32505 589 1 TABLE table NN 32505 589 2 8.--Sizes 8.--sizes CD 32505 589 3 of of IN 32505 589 4 Tadpoles Tadpoles NNP 32505 589 5 of of IN 32505 589 6 _ _ NNP 32505 589 7 Hyla Hyla NNP 32505 589 8 s. s. NNP 32505 589 9 staufferi staufferi NNP 32505 589 10 _ _ NNP 32505 589 11 in in IN 32505 589 12 Relation Relation NNP 32505 589 13 to to IN 32505 589 14 Developmental Developmental NNP 32505 589 15 Stages Stages NNPS 32505 589 16 . . . 32505 590 1 ( ( -LRB- 32505 590 2 Means mean NNS 32505 590 3 in in IN 32505 590 4 parentheses parenthesis NNS 32505 590 5 below below IN 32505 590 6 observed observed JJ 32505 590 7 ranges range NNS 32505 590 8 ; ; : 32505 590 9 measurements measurement NNS 32505 590 10 in in IN 32505 590 11 mm mm NNP 32505 590 12 . . . 32505 590 13 ) ) -RRB- 32505 591 1 = = NFP 32505 591 2 = = NFP 32505 591 3 = = SYM 32505 591 4 = = SYM 32505 591 5 = = SYM 32505 591 6 = = SYM 32505 591 7 = = SYM 32505 591 8 = = SYM 32505 591 9 = = SYM 32505 591 10 = = SYM 32505 591 11 = = SYM 32505 591 12 = = SYM 32505 591 13 = = SYM 32505 591 14 = = SYM 32505 591 15 = = SYM 32505 591 16 = = SYM 32505 591 17 = = SYM 32505 591 18 = = SYM 32505 591 19 = = SYM 32505 591 20 = = SYM 32505 591 21 = = SYM 32505 591 22 = = SYM 32505 591 23 = = SYM 32505 591 24 = = SYM 32505 591 25 = = SYM 32505 591 26 = = SYM 32505 591 27 = = SYM 32505 591 28 = = SYM 32505 591 29 = = SYM 32505 591 30 = = SYM 32505 591 31 = = SYM 32505 591 32 = = SYM 32505 591 33 = = SYM 32505 591 34 = = SYM 32505 591 35 = = SYM 32505 591 36 = = SYM 32505 591 37 = = SYM 32505 591 38 = = SYM 32505 591 39 = = SYM 32505 591 40 = = SYM 32505 591 41 = = SYM 32505 591 42 = = SYM 32505 591 43 = = SYM 32505 591 44 = = SYM 32505 591 45 = = SYM 32505 591 46 = = SYM 32505 591 47 = = SYM 32505 591 48 = = SYM 32505 591 49 = = SYM 32505 591 50 = = SYM 32505 591 51 = = SYM 32505 591 52 = = SYM 32505 591 53 = = NFP 32505 591 54 = = NFP 32505 591 55 Stage stage NN 32505 591 56 | | NNP 32505 591 57 N N NNP 32505 591 58 | | NNP 32505 591 59 Body Body NNP 32505 591 60 length| length| . 32505 591 61 Tail Tail NNP 32505 591 62 length length NN 32505 591 63 | | NNP 32505 591 64 Total total JJ 32505 591 65 length length NN 32505 591 66 --------+---+------------+-------------+-------------- --------+---+------------+-------------+-------------- NN 32505 591 67 25 25 CD 32505 591 68 | | CD 32505 591 69 3 3 CD 32505 591 70 | | CD 32505 591 71 6.0 6.0 CD 32505 591 72 - - HYPH 32505 591 73 7.0 7.0 CD 32505 591 74 | | CD 32505 591 75 12.0 12.0 CD 32505 591 76 - - HYPH 32505 591 77 13.0 13.0 CD 32505 591 78 | | CD 32505 591 79 18.0 18.0 CD 32505 591 80 - - HYPH 32505 591 81 20.0 20.0 CD 32505 591 82 | | CD 32505 591 83 | | CD 32505 591 84 ( ( -LRB- 32505 591 85 6.7 6.7 CD 32505 591 86 ) ) -RRB- 32505 591 87 | | NNP 32505 591 88 ( ( -LRB- 32505 591 89 12.5 12.5 CD 32505 591 90 ) ) -RRB- 32505 591 91 | | NNP 32505 591 92 ( ( -LRB- 32505 591 93 19.2 19.2 CD 32505 591 94 ) ) -RRB- 32505 591 95 | | NNP 32505 591 96 | | CD 32505 591 97 | | CD 32505 591 98 | | CD 32505 591 99 26 26 CD 32505 591 100 | | CD 32505 591 101 2 2 CD 32505 591 102 | | CD 32505 591 103 7.0 7.0 CD 32505 591 104 - - HYPH 32505 591 105 7.5 7.5 CD 32505 591 106 | | CD 32505 591 107 14.0 14.0 CD 32505 591 108 - - HYPH 32505 591 109 15.0 15.0 CD 32505 591 110 | | CD 32505 591 111 21.5 21.5 CD 32505 591 112 - - HYPH 32505 591 113 22.0 22.0 CD 32505 591 114 | | CD 32505 591 115 | | NNS 32505 591 116 ( ( -LRB- 32505 591 117 7.3 7.3 CD 32505 591 118 ) ) -RRB- 32505 591 119 | | NNP 32505 591 120 ( ( -LRB- 32505 591 121 14.5 14.5 CD 32505 591 122 ) ) -RRB- 32505 591 123 | | NNP 32505 591 124 ( ( -LRB- 32505 591 125 21.8 21.8 CD 32505 591 126 ) ) -RRB- 32505 591 127 | | NNP 32505 591 128 | | CD 32505 591 129 | | CD 32505 591 130 | | CD 32505 591 131 27 27 CD 32505 591 132 | | CD 32505 591 133 9 9 CD 32505 591 134 | | CD 32505 591 135 7.0 7.0 CD 32505 591 136 - - HYPH 32505 591 137 8.0 8.0 CD 32505 591 138 | | CD 32505 591 139 13.0 13.0 CD 32505 591 140 - - HYPH 32505 591 141 17.0 17.0 CD 32505 591 142 | | CD 32505 591 143 21.0 21.0 CD 32505 591 144 - - HYPH 32505 591 145 25.0 25.0 CD 32505 591 146 | | CD 32505 591 147 | | CD 32505 591 148 ( ( -LRB- 32505 591 149 7.6 7.6 CD 32505 591 150 ) ) -RRB- 32505 591 151 | | NNP 32505 591 152 ( ( -LRB- 32505 591 153 14.5 14.5 CD 32505 591 154 ) ) -RRB- 32505 591 155 | | NNP 32505 591 156 ( ( -LRB- 32505 591 157 22.0 22.0 CD 32505 591 158 ) ) -RRB- 32505 591 159 | | CD 32505 591 160 | | CD 32505 591 161 | | CD 32505 591 162 | | CD 32505 591 163 32 32 CD 32505 591 164 | | CD 32505 591 165 1 1 CD 32505 591 166 | | CD 32505 591 167 8.5 8.5 CD 32505 591 168 | | CD 32505 591 169 15.5 15.5 CD 32505 591 170 | | CD 32505 591 171 24.0 24.0 CD 32505 591 172 | | CD 32505 591 173 | | CD 32505 591 174 | | CD 32505 591 175 | | CD 32505 591 176 36 36 CD 32505 591 177 | | CD 32505 591 178 2 2 CD 32505 591 179 | | CD 32505 591 180 8.0 8.0 CD 32505 591 181 - - HYPH 32505 591 182 10.0 10.0 CD 32505 591 183 | | CD 32505 591 184 16.5 16.5 CD 32505 591 185 - - HYPH 32505 591 186 17.0 17.0 CD 32505 591 187 | | CD 32505 591 188 25.0 25.0 CD 32505 591 189 - - HYPH 32505 591 190 26.5 26.5 CD 32505 591 191 | | CD 32505 591 192 | | NNS 32505 591 193 ( ( -LRB- 32505 591 194 9.0 9.0 CD 32505 591 195 ) ) -RRB- 32505 591 196 | | NNP 32505 591 197 ( ( -LRB- 32505 591 198 16.8 16.8 CD 32505 591 199 ) ) -RRB- 32505 591 200 | | NNP 32505 591 201 ( ( -LRB- 32505 591 202 25.8 25.8 CD 32505 591 203 ) ) -RRB- 32505 591 204 | | CD 32505 591 205 | | CD 32505 591 206 | | CD 32505 591 207 | | CD 32505 591 208 38 38 CD 32505 591 209 | | CD 32505 591 210 6 6 CD 32505 591 211 | | CD 32505 591 212 9.0 9.0 CD 32505 591 213 - - HYPH 32505 591 214 10.0 10.0 CD 32505 591 215 | | CD 32505 591 216 19.0 19.0 CD 32505 591 217 - - HYPH 32505 591 218 20.5 20.5 CD 32505 591 219 | | CD 32505 591 220 28.0 28.0 CD 32505 591 221 - - HYPH 32505 591 222 29.5 29.5 CD 32505 591 223 | | CD 32505 591 224 | | CD 32505 591 225 ( ( -LRB- 32505 591 226 9.6 9.6 CD 32505 591 227 ) ) -RRB- 32505 591 228 | | CD 32505 591 229 ( ( -LRB- 32505 591 230 19.5 19.5 CD 32505 591 231 ) ) -RRB- 32505 591 232 | | NNP 32505 591 233 ( ( -LRB- 32505 591 234 29.1 29.1 CD 32505 591 235 ) ) -RRB- 32505 591 236 | | CD 32505 591 237 | | CD 32505 591 238 | | CD 32505 591 239 | | CD 32505 591 240 41 41 CD 32505 591 241 | | CD 32505 591 242 1 1 CD 32505 591 243 | | CD 32505 591 244 10.0 10.0 CD 32505 591 245 | | CD 32505 591 246 14.0 14.0 CD 32505 591 247 | | CD 32505 591 248 24.0 24.0 CD 32505 591 249 | | CD 32505 591 250 | | CD 32505 591 251 | | CD 32505 591 252 | | CD 32505 591 253 42 42 CD 32505 591 254 | | CD 32505 591 255 6 6 CD 32505 591 256 | | CD 32505 591 257 11.0 11.0 CD 32505 591 258 - - HYPH 32505 591 259 14.0 14.0 CD 32505 591 260 | | CD 32505 591 261 10.0 10.0 CD 32505 591 262 - - HYPH 32505 591 263 13.0 13.0 CD 32505 591 264 | | CD 32505 591 265 20.0 20.0 CD 32505 591 266 - - HYPH 32505 591 267 29.0 29.0 CD 32505 591 268 | | CD 32505 591 269 | | CD 32505 591 270 ( ( -LRB- 32505 591 271 11.8 11.8 CD 32505 591 272 ) ) -RRB- 32505 591 273 | | NNP 32505 591 274 ( ( -LRB- 32505 591 275 11.9 11.9 CD 32505 591 276 ) ) -RRB- 32505 591 277 | | NNP 32505 591 278 ( ( -LRB- 32505 591 279 24.8 24.8 CD 32505 591 280 ) ) -RRB- 32505 591 281 | | CD 32505 591 282 | | CD 32505 591 283 | | CD 32505 591 284 | | CD 32505 591 285 45 45 CD 32505 591 286 | | CD 32505 591 287 1 1 CD 32505 591 288 | | CD 32505 591 289 12.5 12.5 CD 32505 591 290 | | CD 32505 591 291 0.5 0.5 CD 32505 591 292 | | CD 32505 591 293 13.0 13.0 CD 32505 591 294 | | CD 32505 591 295 | | CD 32505 591 296 | | CD 32505 591 297 | | CD 32505 591 298 46 46 CD 32505 591 299 | | CD 32505 591 300 1 1 CD 32505 591 301 | | CD 32505 591 302 13.0 13.0 CD 32505 591 303 | | CD 32505 591 304 -- -- : 32505 591 305 | | NNS 32505 591 306 -- -- : 32505 591 307 In in IN 32505 591 308 life life NN 32505 591 309 , , , 32505 591 310 body body NN 32505 591 311 pale pale JJ 32505 591 312 olive olive JJ 32505 591 313 - - HYPH 32505 591 314 tan tan NN 32505 591 315 , , , 32505 591 316 belly belly NN 32505 591 317 silvery silvery RB 32505 591 318 white white JJ 32505 591 319 with with IN 32505 591 320 pinkish pinkish JJ 32505 591 321 - - HYPH 32505 591 322 orange orange NNP 32505 591 323 reticulations reticulation NNS 32505 591 324 in in IN 32505 591 325 some some DT 32505 591 326 specimens specimen NNS 32505 591 327 ; ; : 32505 591 328 tail tail NN 32505 591 329 creamy creamy JJ 32505 591 330 white white JJ 32505 591 331 with with IN 32505 591 332 silvery silvery JJ 32505 591 333 flecks fleck NNS 32505 591 334 and and CC 32505 591 335 black black JJ 32505 591 336 or or CC 32505 591 337 brown brown JJ 32505 591 338 reticulations reticulation NNS 32505 591 339 . . . 32505 592 1 In in IN 32505 592 2 preservative preservative JJ 32505 592 3 , , , 32505 592 4 tan tan NNP 32505 592 5 and and CC 32505 592 6 pinkish pinkish NNP 32505 592 7 - - HYPH 32505 592 8 orange orange NNP 32505 592 9 coloration coloration NN 32505 592 10 lost lose VBD 32505 592 11 ; ; : 32505 592 12 body body NN 32505 592 13 transparent transparent NN 32505 592 14 , , , 32505 592 15 reticulations reticulation NNS 32505 592 16 on on IN 32505 592 17 tail tail NN 32505 592 18 present present JJ 32505 592 19 . . . 32505 593 1 _ _ NNP 32505 593 2 Remarks._--_Hyla Remarks._--_Hyla NNP 32505 593 3 staufferi staufferi NN 32505 593 4 _ _ NNP 32505 593 5 was be VBD 32505 593 6 described describe VBN 32505 593 7 by by IN 32505 593 8 Cope Cope NNP 32505 593 9 ( ( -LRB- 32505 593 10 1865:195 1865:195 CD 32505 593 11 ) ) -RRB- 32505 593 12 on on IN 32505 593 13 the the DT 32505 593 14 basis basis NN 32505 593 15 of of IN 32505 593 16 specimens specimen NNS 32505 593 17 from from IN 32505 593 18 Orizaba Orizaba NNP 32505 593 19 , , , 32505 593 20 Veracruz Veracruz NNP 32505 593 21 , , , 32505 593 22 México México NNP 32505 593 23 . . . 32505 594 1 He -PRON- PRP 32505 594 2 described describe VBD 32505 594 3 the the DT 32505 594 4 color color NN 32505 594 5 pattern pattern NN 32505 594 6 as as IN 32505 594 7 " " `` 32505 594 8 color color NN 32505 594 9 above above IN 32505 594 10 dark dark JJ 32505 594 11 olive olive NN 32505 594 12 , , , 32505 594 13 with with IN 32505 594 14 a a DT 32505 594 15 short short JJ 32505 594 16 black black JJ 32505 594 17 bar bar NN 32505 594 18 over over IN 32505 594 19 each each DT 32505 594 20 scapula scapula NN 32505 594 21 , , , 32505 594 22 and and CC 32505 594 23 one one CD 32505 594 24 from from IN 32505 594 25 eye eye NN 32505 594 26 to to IN 32505 594 27 eye eye NN 32505 594 28 , , , 32505 594 29 with with IN 32505 594 30 a a DT 32505 594 31 trace trace NN 32505 594 32 along along IN 32505 594 33 the the DT 32505 594 34 coccyx coccyx NN 32505 594 35 . . . 32505 594 36 " " '' 32505 595 1 Cope Cope NNP 32505 595 2 ( ( -LRB- 32505 595 3 1887:14 1887:14 CD 32505 595 4 ) ) -RRB- 32505 595 5 placed place VBN 32505 595 6 _ _ NNP 32505 595 7 staufferi staufferi NN 32505 595 8 _ _ NNP 32505 595 9 as as IN 32505 595 10 a a DT 32505 595 11 subspecies subspecie NNS 32505 595 12 of of IN 32505 595 13 _ _ NNP 32505 595 14 Hyla Hyla NNP 32505 595 15 eximia eximia NN 32505 595 16 _ _ NNP 32505 595 17 , , , 32505 595 18 but but CC 32505 595 19 he -PRON- PRP 32505 595 20 did do VBD 32505 595 21 not not RB 32505 595 22 justify justify VB 32505 595 23 his -PRON- PRP$ 32505 595 24 action action NN 32505 595 25 . . . 32505 596 1 Günther Günther NNP 32505 596 2 ( ( -LRB- 32505 596 3 1901:262 1901:262 CD 32505 596 4 ) ) -RRB- 32505 596 5 also also RB 32505 596 6 considered consider VBN 32505 596 7 _ _ NNP 32505 596 8 staufferi staufferi NN 32505 596 9 _ _ NNP 32505 596 10 to to TO 32505 596 11 be be VB 32505 596 12 conspecific conspecific NN 32505 596 13 with with IN 32505 596 14 _ _ NNP 32505 596 15 eximia eximia NN 32505 596 16 _ _ NNP 32505 596 17 without without IN 32505 596 18 making make VBG 32505 596 19 any any DT 32505 596 20 qualifying qualify VBG 32505 596 21 statement statement NN 32505 596 22 . . . 32505 597 1 Dunn Dunn NNP 32505 597 2 and and CC 32505 597 3 Emlen Emlen NNP 32505 597 4 ( ( -LRB- 32505 597 5 1932:24 1932:24 CD 32505 597 6 ) ) -RRB- 32505 597 7 named name VBN 32505 597 8 _ _ NNP 32505 597 9 Hyla Hyla NNP 32505 597 10 culex culex NN 32505 597 11 _ _ NNP 32505 597 12 from from IN 32505 597 13 Tela Tela NNP 32505 597 14 , , , 32505 597 15 Honduras Honduras NNP 32505 597 16 , , , 32505 597 17 on on IN 32505 597 18 the the DT 32505 597 19 basis basis NN 32505 597 20 of of IN 32505 597 21 a a DT 32505 597 22 male male NN 32505 597 23 ( ( -LRB- 32505 597 24 MCZ MCZ NNP 32505 597 25 16098 16098 CD 32505 597 26 ) ) -RRB- 32505 597 27 having have VBG 32505 597 28 a a DT 32505 597 29 snout snout NN 32505 597 30 - - HYPH 32505 597 31 vent vent NN 32505 597 32 length length NN 32505 597 33 of of IN 32505 597 34 25.1 25.1 CD 32505 597 35 mm mm NNP 32505 597 36 . . NNP 32505 597 37 , , , 32505 597 38 and and CC 32505 597 39 a a DT 32505 597 40 female female JJ 32505 597 41 ( ( -LRB- 32505 597 42 USNM USNM NNP 32505 597 43 20267 20267 CD 32505 597 44 ) ) -RRB- 32505 597 45 from from IN 32505 597 46 Patuca Patuca NNP 32505 597 47 , , , 32505 597 48 Honduras Honduras NNP 32505 597 49 . . . 32505 598 1 They -PRON- PRP 32505 598 2 diagnosed diagnose VBD 32505 598 3 the the DT 32505 598 4 species specie NNS 32505 598 5 as as IN 32505 598 6 having have VBG 32505 598 7 " " `` 32505 598 8 discs disc NNS 32505 598 9 larger large JJR 32505 598 10 than than IN 32505 598 11 tympanum tympanum NN 32505 598 12 ... ... : 32505 598 13 black black JJ 32505 598 14 interorbital interorbital NN 32505 598 15 triangle triangle NN 32505 598 16 , , , 32505 598 17 traces trace NNS 32505 598 18 of of IN 32505 598 19 black black JJ 32505 598 20 dorsal dorsal NN 32505 598 21 marking marking NN 32505 598 22 ; ; : 32505 598 23 three three CD 32505 598 24 black black JJ 32505 598 25 bars bar NNS 32505 598 26 on on IN 32505 598 27 anterior anterior NNP 32505 598 28 and and CC 32505 598 29 posterior posterior NN 32505 598 30 face face NN 32505 598 31 of of IN 32505 598 32 thighs thigh NNS 32505 598 33 , , , 32505 598 34 two two CD 32505 598 35 black black JJ 32505 598 36 bars bar NNS 32505 598 37 on on IN 32505 598 38 tibia tibia NNP 32505 598 39 , , , 32505 598 40 on on IN 32505 598 41 tarsus tarsus NNP 32505 598 42 and and CC 32505 598 43 on on IN 32505 598 44 forearm forearm NN 32505 598 45 . . . 32505 598 46 " " '' 32505 599 1 The the DT 32505 599 2 holotype holotype NN 32505 599 3 now now RB 32505 599 4 is be VBZ 32505 599 5 faded fade VBN 32505 599 6 but but CC 32505 599 7 has have VBZ 32505 599 8 some some DT 32505 599 9 of of IN 32505 599 10 the the DT 32505 599 11 pattern pattern NN 32505 599 12 described describe VBN 32505 599 13 . . . 32505 600 1 Dunn Dunn NNP 32505 600 2 and and CC 32505 600 3 Emlen Emlen NNP 32505 600 4 did do VBD 32505 600 5 not not RB 32505 600 6 compare compare VB 32505 600 7 _ _ NNP 32505 600 8 culex culex NN 32505 600 9 _ _ NNP 32505 600 10 with with IN 32505 600 11 _ _ NNP 32505 600 12 staufferi staufferi NN 32505 600 13 _ _ NNP 32505 600 14 but but CC 32505 600 15 did do VBD 32505 600 16 compare compare VB 32505 600 17 it -PRON- PRP 32505 600 18 with with IN 32505 600 19 _ _ NNP 32505 600 20 boulengeri boulengeri NN 32505 600 21 _ _ NNP 32505 600 22 and and CC 32505 600 23 _ _ NNP 32505 600 24 rubra rubra NNP 32505 600 25 _ _ NNP 32505 600 26 . . . 32505 601 1 Dunn Dunn NNP 32505 601 2 ( ( -LRB- 32505 601 3 1933:61 1933:61 CD 32505 601 4 ) ) -RRB- 32505 601 5 named name VBN 32505 601 6 _ _ NNP 32505 601 7 Hyla Hyla NNP 32505 601 8 altae altae NN 32505 601 9 _ _ NNP 32505 601 10 from from IN 32505 601 11 Summit Summit NNP 32505 601 12 , , , 32505 601 13 Canal Canal NNP 32505 601 14 Zone Zone NNP 32505 601 15 . . . 32505 602 1 His -PRON- PRP$ 32505 602 2 description description NN 32505 602 3 was be VBD 32505 602 4 based base VBN 32505 602 5 on on IN 32505 602 6 a a DT 32505 602 7 male male NN 32505 602 8 ( ( -LRB- 32505 602 9 MCZ MCZ NNP 32505 602 10 17972 17972 CD 32505 602 11 ) ) -RRB- 32505 602 12 having have VBG 32505 602 13 a a DT 32505 602 14 snout snout NN 32505 602 15 - - HYPH 32505 602 16 vent vent NN 32505 602 17 length length NN 32505 602 18 of of IN 32505 602 19 25.1 25.1 CD 32505 602 20 mm mm NNP 32505 602 21 . . NNP 32505 602 22 , , , 32505 602 23 the the DT 32505 602 24 color color NN 32505 602 25 pattern pattern NN 32505 602 26 was be VBD 32505 602 27 described describe VBN 32505 602 28 as as IN 32505 602 29 " " `` 32505 602 30 gray gray JJ 32505 602 31 with with IN 32505 602 32 four four CD 32505 602 33 darker dark JJR 32505 602 34 dorsal dorsal NN 32505 602 35 stripes stripe NNS 32505 602 36 ... ... : 32505 602 37 a a DT 32505 602 38 faint faint JJ 32505 602 39 trace trace NN 32505 602 40 of of IN 32505 602 41 mid mid JJ 32505 602 42 - - JJ 32505 602 43 dorsal dorsal NN 32505 602 44 striping striping NN 32505 602 45 .... .... . 32505 602 46 " " '' 32505 602 47 Dunn Dunn NNP 32505 602 48 defined define VBD 32505 602 49 the the DT 32505 602 50 _ _ NNP 32505 602 51 Hyla Hyla NNP 32505 602 52 rubra rubra NN 32505 602 53 _ _ NNP 32505 602 54 group group NN 32505 602 55 and and CC 32505 602 56 recognized recognize VBD 32505 602 57 _ _ NNP 32505 602 58 boulengeri boulengeri NN 32505 602 59 _ _ NNP 32505 602 60 , , , 32505 602 61 _ _ NNP 32505 602 62 altae altae NN 32505 602 63 _ _ NNP 32505 602 64 , , , 32505 602 65 _ _ NNP 32505 602 66 culex culex NN 32505 602 67 _ _ NNP 32505 602 68 , , , 32505 602 69 and and CC 32505 602 70 _ _ NNP 32505 602 71 rubra rubra NNP 32505 602 72 _ _ NNP 32505 602 73 as as IN 32505 602 74 members member NNS 32505 602 75 . . . 32505 603 1 _ _ NNP 32505 603 2 Hyla Hyla NNP 32505 603 3 elaeochroa elaeochroa NN 32505 603 4 _ _ NNP 32505 603 5 and and CC 32505 603 6 _ _ NNP 32505 603 7 staufferi staufferi NN 32505 603 8 _ _ NNP 32505 603 9 were be VBD 32505 603 10 omitted omit VBN 32505 603 11 from from IN 32505 603 12 his -PRON- PRP$ 32505 603 13 key key NN 32505 603 14 to to IN 32505 603 15 the the DT 32505 603 16 group group NN 32505 603 17 in in IN 32505 603 18 Central Central NNP 32505 603 19 America America NNP 32505 603 20 . . . 32505 604 1 Kellogg Kellogg NNP 32505 604 2 ( ( -LRB- 32505 604 3 1932:174 1932:174 CD 32505 604 4 ) ) -RRB- 32505 604 5 compared compare VBN 32505 604 6 _ _ NNP 32505 604 7 staufferi staufferi NN 32505 604 8 _ _ NNP 32505 604 9 with with IN 32505 604 10 _ _ NNP 32505 604 11 eximia eximia NN 32505 604 12 _ _ NNP 32505 604 13 and and CC 32505 604 14 concluded conclude VBD 32505 604 15 that that IN 32505 604 16 the the DT 32505 604 17 two two CD 32505 604 18 were be VBD 32505 604 19 probably probably RB 32505 604 20 distinct distinct JJ 32505 604 21 species specie NNS 32505 604 22 . . . 32505 605 1 Stuart Stuart NNP 32505 605 2 ( ( -LRB- 32505 605 3 1935:38 1935:38 CD 32505 605 4 ) ) -RRB- 32505 605 5 considered consider VBN 32505 605 6 _ _ NNP 32505 605 7 altae altae NN 32505 605 8 _ _ NNP 32505 605 9 to to TO 32505 605 10 be be VB 32505 605 11 a a DT 32505 605 12 synonym synonym NN 32505 605 13 of of IN 32505 605 14 _ _ NNP 32505 605 15 culex culex NN 32505 605 16 _ _ NNP 32505 605 17 . . . 32505 606 1 Gaige Gaige NNP 32505 606 2 ( ( -LRB- 32505 606 3 1936:293 1936:293 CD 32505 606 4 ) ) -RRB- 32505 606 5 considered consider VBN 32505 606 6 _ _ NNP 32505 606 7 altae altae NN 32505 606 8 _ _ NNP 32505 606 9 and and CC 32505 606 10 _ _ NNP 32505 606 11 culex culex NN 32505 606 12 _ _ NNP 32505 606 13 to to TO 32505 606 14 be be VB 32505 606 15 conspecific conspecific NN 32505 606 16 but but CC 32505 606 17 regarded regard VBN 32505 606 18 _ _ NNP 32505 606 19 staufferi staufferi NN 32505 606 20 _ _ NNP 32505 606 21 as as IN 32505 606 22 a a DT 32505 606 23 different different JJ 32505 606 24 species species NN 32505 606 25 . . . 32505 607 1 She -PRON- PRP 32505 607 2 also also RB 32505 607 3 suggested suggest VBD 32505 607 4 that that IN 32505 607 5 _ _ NNP 32505 607 6 staufferi staufferi NN 32505 607 7 _ _ NNP 32505 607 8 was be VBD 32505 607 9 not not RB 32505 607 10 related relate VBN 32505 607 11 to to IN 32505 607 12 _ _ NNP 32505 607 13 eximia eximia NN 32505 607 14 _ _ NNP 32505 607 15 but but CC 32505 607 16 belonged belong VBD 32505 607 17 to to IN 32505 607 18 the the DT 32505 607 19 _ _ NNP 32505 607 20 rubra rubra NNP 32505 607 21 _ _ NNP 32505 607 22 group group NN 32505 607 23 . . . 32505 608 1 Taylor Taylor NNP 32505 608 2 ( ( -LRB- 32505 608 3 1952:865 1952:865 CD 32505 608 4 ) ) -RRB- 32505 608 5 and and CC 32505 608 6 Duellman Duellman NNP 32505 608 7 ( ( -LRB- 32505 608 8 1966a:274 1966a:274 NNP 32505 608 9 ) ) -RRB- 32505 608 10 considered consider VBN 32505 608 11 _ _ NNP 32505 608 12 altae altae NN 32505 608 13 _ _ NNP 32505 608 14 and and CC 32505 608 15 _ _ NNP 32505 608 16 culex culex NN 32505 608 17 _ _ NNP 32505 608 18 to to TO 32505 608 19 be be VB 32505 608 20 synonyms synonyms JJ 32505 608 21 of of IN 32505 608 22 _ _ NNP 32505 608 23 staufferi staufferi NN 32505 608 24 _ _ NNP 32505 608 25 . . . 32505 609 1 The the DT 32505 609 2 only only JJ 32505 609 3 other other JJ 32505 609 4 worker worker NN 32505 609 5 besides besides IN 32505 609 6 Cope Cope NNP 32505 609 7 and and CC 32505 609 8 Günther Günther NNP 32505 609 9 to to TO 32505 609 10 consider consider VB 32505 609 11 _ _ NNP 32505 609 12 Hyla Hyla NNP 32505 609 13 staufferi staufferi NN 32505 609 14 _ _ NNP 32505 609 15 as as IN 32505 609 16 a a DT 32505 609 17 member member NN 32505 609 18 of of IN 32505 609 19 the the DT 32505 609 20 _ _ NNP 32505 609 21 eximia eximia NN 32505 609 22 _ _ NNP 32505 609 23 group group NN 32505 609 24 was be VBD 32505 609 25 Blair Blair NNP 32505 609 26 ( ( -LRB- 32505 609 27 1960:129 1960:129 CD 32505 609 28 ) ) -RRB- 32505 609 29 , , , 32505 609 30 who who WP 32505 609 31 suggested suggest VBD 32505 609 32 the the DT 32505 609 33 relationship relationship NN 32505 609 34 on on IN 32505 609 35 the the DT 32505 609 36 basis basis NN 32505 609 37 of of IN 32505 609 38 similarities similarity NNS 32505 609 39 in in IN 32505 609 40 the the DT 32505 609 41 structure structure NN 32505 609 42 of of IN 32505 609 43 the the DT 32505 609 44 calls call NNS 32505 609 45 of of IN 32505 609 46 _ _ NNP 32505 609 47 eximia eximia NN 32505 609 48 _ _ NNP 32505 609 49 and and CC 32505 609 50 _ _ NNP 32505 609 51 staufferi staufferi NN 32505 609 52 _ _ NNP 32505 609 53 . . . 32505 610 1 Taylor Taylor NNP 32505 610 2 ( ( -LRB- 32505 610 3 1938:421 1938:421 CD 32505 610 4 ) ) -RRB- 32505 610 5 and and CC 32505 610 6 Smith Smith NNP 32505 610 7 and and CC 32505 610 8 Taylor Taylor NNP 32505 610 9 ( ( -LRB- 32505 610 10 1948:78 1948:78 CD 32505 610 11 ) ) -RRB- 32505 610 12 excluded exclude VBD 32505 610 13 _ _ NNP 32505 610 14 staufferi staufferi NN 32505 610 15 _ _ NNP 32505 610 16 from from IN 32505 610 17 the the DT 32505 610 18 _ _ NNP 32505 610 19 eximia eximia NN 32505 610 20 _ _ NNP 32505 610 21 group group NN 32505 610 22 on on IN 32505 610 23 the the DT 32505 610 24 basis basis NN 32505 610 25 of of IN 32505 610 26 morphological morphological JJ 32505 610 27 characteristics characteristic NNS 32505 610 28 . . . 32505 611 1 I -PRON- PRP 32505 611 2 consider consider VBP 32505 611 3 _ _ NNP 32505 611 4 culex culex NN 32505 611 5 _ _ NNP 32505 611 6 to to TO 32505 611 7 be be VB 32505 611 8 inseparable inseparable JJ 32505 611 9 from from IN 32505 611 10 _ _ NNP 32505 611 11 staufferi staufferi NN 32505 611 12 _ _ NNP 32505 611 13 , , , 32505 611 14 whereas whereas IN 32505 611 15 _ _ NNP 32505 611 16 altae altae NN 32505 611 17 _ _ NNP 32505 611 18 is be VBZ 32505 611 19 recognizable recognizable JJ 32505 611 20 as as IN 32505 611 21 a a DT 32505 611 22 Panamanian panamanian JJ 32505 611 23 subspecies subspecie NNS 32505 611 24 of of IN 32505 611 25 _ _ NNP 32505 611 26 staufferi staufferi NN 32505 611 27 _ _ NNP 32505 611 28 . . . 32505 612 1 _ _ NNP 32505 612 2 Hyla Hyla NNP 32505 612 3 staufferi staufferi NN 32505 612 4 staufferi staufferi NN 32505 612 5 _ _ NNP 32505 612 6 Cope Cope NNP 32505 612 7 , , , 32505 612 8 New New NNP 32505 612 9 Combination Combination NNP 32505 612 10 _ _ NNP 32505 612 11 Hyla Hyla NNP 32505 612 12 staufferi staufferi NN 32505 612 13 _ _ NNP 32505 612 14 Cope Cope NNP 32505 612 15 , , , 32505 612 16 Proc Proc NNP 32505 612 17 . . . 32505 613 1 Acad Acad NNS 32505 613 2 . . . 32505 614 1 Nat Nat NNP 32505 614 2 . . . 32505 615 1 Sci Sci NNP 32505 615 2 . . . 32505 616 1 Philadelphia Philadelphia NNP 32505 616 2 , , , 32505 616 3 17:195 17:195 CD 32505 616 4 , , , 32505 616 5 October October NNP 32505 616 6 1865 1865 CD 32505 616 7 [ [ -LRB- 32505 616 8 Holotype.--USNM Holotype.--USNM NNP 32505 616 9 15317 15317 CD 32505 616 10 , , , 32505 616 11 Orizaba Orizaba NNP 32505 616 12 , , , 32505 616 13 Veracruz Veracruz NNP 32505 616 14 , , , 32505 616 15 México México NNP 32505 616 16 ; ; : 32505 616 17 Francis Francis NNP 32505 616 18 Sumichrast Sumichrast NNP 32505 616 19 collector collector NN 32505 616 20 ] ] -RRB- 32505 616 21 , , , 32505 616 22 Brocchi Brocchi NNP 32505 616 23 , , , 32505 616 24 Mission Mission NNP 32505 616 25 Scientifique Scientifique NNP 32505 616 26 au au NNP 32505 616 27 Mexique Mexique NNP 32505 616 28 et et NNP 32505 616 29 dans dan VBZ 32505 616 30 L'Amerique L'Amerique NNP 32505 616 31 Centrale Centrale NNP 32505 616 32 , , , 32505 616 33 1881 1881 CD 32505 616 34 , , , 32505 616 35 p. p. NN 32505 616 36 36 36 CD 32505 616 37 . . . 32505 617 1 Boulenger Boulenger NNP 32505 617 2 , , , 32505 617 3 Catalogue Catalogue NNP 32505 617 4 , , , 32505 617 5 of of IN 32505 617 6 the the DT 32505 617 7 Bratrachia Bratrachia NNP 32505 617 8 Salientia Salientia NNP 32505 617 9 s. s. NNP 32505 617 10 Ecaudata Ecaudata NNP 32505 617 11 , , , 32505 617 12 p. p. NN 32505 617 13 400 400 CD 32505 617 14 , , , 32505 617 15 February February NNP 32505 617 16 1 1 CD 32505 617 17 , , , 32505 617 18 1882 1882 CD 32505 617 19 . . . 32505 618 1 Kellogg Kellogg NNP 32505 618 2 , , , 32505 618 3 Bull Bull NNP 32505 618 4 . . . 32505 619 1 U.S. U.S. NNP 32505 619 2 Natl Natl NNP 32505 619 3 . . . 32505 620 1 Mus Mus NNP 32505 620 2 . . NNP 32505 620 3 , , , 32505 620 4 160:173 160:173 CD 32505 620 5 , , , 32505 620 6 March March NNP 32505 620 7 31 31 CD 32505 620 8 , , , 32505 620 9 1932 1932 CD 32505 620 10 . . . 32505 621 1 Smith Smith NNP 32505 621 2 and and CC 32505 621 3 Taylor Taylor NNP 32505 621 4 , , , 32505 621 5 Bull Bull NNP 32505 621 6 . . . 32505 622 1 U.S. U.S. NNP 32505 622 2 Natl Natl NNP 32505 622 3 . . . 32505 623 1 Mus Mus NNP 32505 623 2 . . NNP 32505 623 3 , , , 32505 623 4 194:88 194:88 CD 32505 623 5 , , , 32505 623 6 1948 1948 CD 32505 623 7 . . . 32505 624 1 Taylor Taylor NNP 32505 624 2 , , , 32505 624 3 Univ Univ NNP 32505 624 4 . . . 32505 625 1 Kansas Kansas NNP 32505 625 2 Sci Sci NNP 32505 625 3 . . . 32505 626 1 Bull Bull NNP 32505 626 2 . . NNP 32505 626 3 , , , 32505 626 4 35:862 35:862 CD 32505 626 5 , , , 32505 626 6 July July NNP 32505 626 7 1 1 CD 32505 626 8 , , , 32505 626 9 1952 1952 CD 32505 626 10 . . . 32505 627 1 Rand Rand NNP 32505 627 2 , , , 32505 627 3 Fieldiana Fieldiana NNP 32505 627 4 Zool Zool NNP 32505 627 5 . . . 32505 628 1 Chicago Chicago NNP 32505 628 2 Nat Nat NNP 32505 628 3 . . . 32505 629 1 Hist hist NN 32505 629 2 . . . 32505 630 1 Mus Mus NNP 32505 630 2 . . NNP 32505 630 3 , , , 32505 630 4 34:518 34:518 CD 32505 630 5 , , , 32505 630 6 April April NNP 32505 630 7 18 18 CD 32505 630 8 , , , 32505 630 9 1957 1957 CD 32505 630 10 . . . 32505 631 1 Duellman Duellman NNP 32505 631 2 , , , 32505 631 3 Univ Univ NNP 32505 631 4 . . . 32505 632 1 Kansas Kansas NNP 32505 632 2 Publ Publ NNP 32505 632 3 , , , 32505 632 4 Mus Mus NNP 32505 632 5 . . . 32505 633 1 Nat Nat NNP 32505 633 2 . . . 32505 634 1 Hist Hist NNP 32505 634 2 . . NNP 32505 634 3 , , , 32505 634 4 17:274 17:274 CD 32505 634 5 , , , 32505 634 6 June June NNP 32505 634 7 17 17 CD 32505 634 8 , , , 32505 634 9 1966 1966 CD 32505 634 10 . . . 32505 635 1 _ _ NNP 32505 635 2 Hyla Hyla NNP 32505 635 3 eximia eximia NN 32505 635 4 staufferi staufferi NN 32505 635 5 _ _ NNP 32505 635 6 Cope Cope NNP 32505 635 7 , , , 32505 635 8 Bull Bull NNP 32505 635 9 . . . 32505 636 1 U.S. U.S. NNP 32505 636 2 Natl Natl NNP 32505 636 3 . . . 32505 637 1 Mus Mus NNP 32505 637 2 . . . 32505 637 3 , , , 32505 637 4 32:14 32:14 CD 32505 637 5 , , , 32505 637 6 January January NNP 32505 637 7 16 16 CD 32505 637 8 , , , 32505 637 9 1887 1887 CD 32505 637 10 . . . 32505 638 1 _ _ NNP 32505 638 2 Hyla Hyla NNP 32505 638 3 eximia eximia NN 32505 638 4 _ _ NNP 32505 638 5 ( ( -LRB- 32505 638 6 part part NN 32505 638 7 ) ) -RRB- 32505 638 8 : : : 32505 638 9 Günther Günther NNP 32505 638 10 , , , 32505 638 11 Biologia Biologia NNP 32505 638 12 Centrali Centrali NNP 32505 638 13 - - HYPH 32505 638 14 Americana Americana NNP 32505 638 15 , , , 32505 638 16 Reptilia Reptilia NNP 32505 638 17 and and CC 32505 638 18 Batrachia Batrachia NNP 32505 638 19 , , , 32505 638 20 p. p. NN 32505 638 21 261 261 CD 32505 638 22 , , , 32505 638 23 June June NNP 32505 638 24 1901 1901 CD 32505 638 25 . . . 32505 639 1 Nieden Nieden NNP 32505 639 2 , , , 32505 639 3 Das Das NNP 32505 639 4 Tierreich Tierreich NNP 32505 639 5 , , , 32505 639 6 Anura Anura NNP 32505 639 7 I I NNP 32505 639 8 , , , 32505 639 9 p. p. NN 32505 639 10 245 245 CD 32505 639 11 , , , 32505 639 12 June June NNP 32505 639 13 1923 1923 CD 32505 639 14 . . . 32505 640 1 _ _ NNP 32505 640 2 Hyla Hyla NNP 32505 640 3 culex culex NN 32505 640 4 _ _ NNP 32505 640 5 Dunn Dunn NNP 32505 640 6 and and CC 32505 640 7 Emlen Emlen NNP 32505 640 8 , , , 32505 640 9 Proc Proc NNP 32505 640 10 . . . 32505 641 1 Acad Acad NNS 32505 641 2 . . . 32505 642 1 Nat Nat NNP 32505 642 2 . . . 32505 643 1 Sci Sci NNP 32505 643 2 . . . 32505 644 1 Philadelphia Philadelphia NNP 32505 644 2 , , , 32505 644 3 84:24 84:24 CD 32505 644 4 , , , 32505 644 5 March March NNP 32505 644 6 22 22 CD 32505 644 7 , , , 32505 644 8 1932 1932 CD 32505 644 9 [ [ -LRB- 32505 644 10 Holotype.--MCZ Holotype.--MCZ NNP 32505 644 11 16098 16098 CD 32505 644 12 , , , 32505 644 13 Tela Tela NNP 32505 644 14 Honduras Honduras NNP 32505 644 15 ; ; : 32505 644 16 Raymond Raymond NNP 32505 644 17 A. A. NNP 32505 644 18 Stadelman Stadelman NNP 32505 644 19 collector collector NN 32505 644 20 ] ] -RRB- 32505 644 21 . . . 32505 645 1 Stuart Stuart NNP 32505 645 2 , , , 32505 645 3 Misc Misc NNP 32505 645 4 . . . 32505 646 1 Publ Publ NNS 32505 646 2 . . . 32505 646 3 , , , 32505 646 4 Univ Univ NNP 32505 646 5 . . . 32505 647 1 Michigan Michigan NNP 32505 647 2 Mus Mus NNP 32505 647 3 . . . 32505 648 1 Zool Zool NNP 32505 648 2 . . . 32505 648 3 , , , 32505 648 4 29:38 29:38 CD 32505 648 5 , , , 32505 648 6 October October NNP 32505 648 7 1935 1935 CD 32505 648 8 . . . 32505 649 1 Gaige Gaige NNP 32505 649 2 , , , 32505 649 3 Carnegie Carnegie NNP 32505 649 4 Inst Inst NNP 32505 649 5 . . . 32505 650 1 Washington Washington NNP 32505 650 2 Publ Publ NNP 32505 650 3 . . . 32505 650 4 , , , 32505 650 5 457:293 457:293 CD 32505 650 6 , , , 32505 650 7 1936 1936 CD 32505 650 8 . . . 32505 651 1 _ _ NNP 32505 651 2 Diagnosis._--Small Diagnosis._--Small NNP 32505 651 3 frogs frog NNS 32505 651 4 ( ( -LRB- 32505 651 5 Male male NN 32505 651 6 to to TO 32505 651 7 29 29 CD 32505 651 8 mm mm NNS 32505 651 9 . . NNP 32505 651 10 , , , 32505 651 11 Female female JJ 32505 651 12 to to IN 32505 651 13 31.6 31.6 CD 32505 651 14 mm mm NNS 32505 651 15 . . . 32505 652 1 ) ) -RRB- 32505 652 2 ; ; : 32505 652 3 dorsolateral dorsolateral JJ 32505 652 4 stripes stripes NNPS 32505 652 5 irregular irregular NNP 32505 652 6 ; ; , 32505 652 7 paravertebral paravertebral JJ 32505 652 8 stripes stripe NNS 32505 652 9 usually usually RB 32505 652 10 broken break VBN 32505 652 11 ; ; : 32505 652 12 two two CD 32505 652 13 or or CC 32505 652 14 three three CD 32505 652 15 transverse transverse JJ 32505 652 16 bars bar NNS 32505 652 17 on on IN 32505 652 18 shanks shank NNS 32505 652 19 ; ; : 32505 652 20 thighs thigh NNS 32505 652 21 spotted spot VBD 32505 652 22 or or CC 32505 652 23 not not RB 32505 652 24 ; ; : 32505 652 25 arms arm NNS 32505 652 26 usually usually RB 32505 652 27 barred bar VBN 32505 652 28 ; ; : 32505 652 29 interorbital interorbital NNP 32505 652 30 bar bar NN 32505 652 31 usually usually RB 32505 652 32 present present JJ 32505 652 33 ; ; : 32505 652 34 toes toe NNS 32505 652 35 about about RB 32505 652 36 three three CD 32505 652 37 fourths fourth NNS 32505 652 38 webbed web VBN 32505 652 39 ; ; : 32505 652 40 color color NN 32505 652 41 brown brown NNP 32505 652 42 , , , 32505 652 43 tan tan NNP 32505 652 44 , , , 32505 652 45 or or CC 32505 652 46 olive olive NN 32505 652 47 - - HYPH 32505 652 48 green green JJ 32505 652 49 . . . 32505 653 1 _ _ NNP 32505 653 2 Variation._--Three variation._--three CD 32505 653 3 hundred hundred CD 32505 653 4 and and CC 32505 653 5 sixty sixty CD 32505 653 6 males male NNS 32505 653 7 chosen choose VBN 32505 653 8 at at IN 32505 653 9 random random JJ 32505 653 10 from from IN 32505 653 11 throughout throughout IN 32505 653 12 the the DT 32505 653 13 range range NN 32505 653 14 have have VBP 32505 653 15 snout snout VBN 32505 653 16 - - HYPH 32505 653 17 vent vent NN 32505 653 18 lengths length NNS 32505 653 19 of of IN 32505 653 20 20.7 20.7 CD 32505 653 21 to to TO 32505 653 22 29 29 CD 32505 653 23 mm mm NNS 32505 653 24 . . . 32505 654 1 ( ( -LRB- 32505 654 2 25.9 25.9 CD 32505 654 3 mm mm NNP 32505 654 4 . . . 32505 654 5 ) ) -RRB- 32505 654 6 . . . 32505 655 1 The the DT 32505 655 2 smallest small JJS 32505 655 3 individuals individual NNS 32505 655 4 are be VBP 32505 655 5 from from IN 32505 655 6 Costa Costa NNP 32505 655 7 Rica Rica NNP 32505 655 8 and and CC 32505 655 9 Nicaragua Nicaragua NNP 32505 655 10 ( ( -LRB- 32505 655 11 means mean VBZ 32505 655 12 24.2 24.2 CD 32505 655 13 and and CC 32505 655 14 24.4 24.4 CD 32505 655 15 mm mm NNP 32505 655 16 . . NNP 32505 655 17 , , , 32505 655 18 respectively respectively RB 32505 655 19 ) ) -RRB- 32505 655 20 . . . 32505 656 1 The the DT 32505 656 2 largest large JJS 32505 656 3 individuals individual NNS 32505 656 4 are be VBP 32505 656 5 from from IN 32505 656 6 Guatemala Guatemala NNP 32505 656 7 and and CC 32505 656 8 El El NNP 32505 656 9 Salvador Salvador NNP 32505 656 10 ( ( -LRB- 32505 656 11 mean mean NN 32505 656 12 of of IN 32505 656 13 each each DT 32505 656 14 27.0 27.0 CD 32505 656 15 mm mm NNP 32505 656 16 . . . 32505 656 17 ) ) -RRB- 32505 656 18 . . . 32505 657 1 The the DT 32505 657 2 ratio ratio NN 32505 657 3 of of IN 32505 657 4 the the DT 32505 657 5 diameter diameter NN 32505 657 6 of of IN 32505 657 7 the the DT 32505 657 8 tympanum tympanum NN 32505 657 9 to to IN 32505 657 10 that that DT 32505 657 11 of of IN 32505 657 12 the the DT 32505 657 13 eye eye NN 32505 657 14 is be VBZ 32505 657 15 more more JJR 32505 657 16 than than IN 32505 657 17 60 60 CD 32505 657 18 per per NN 32505 657 19 cent cent NN 32505 657 20 in in IN 32505 657 21 most most JJS 32505 657 22 samples sample NNS 32505 657 23 , , , 32505 657 24 but but CC 32505 657 25 in in IN 32505 657 26 those those DT 32505 657 27 from from IN 32505 657 28 Costa Costa NNP 32505 657 29 Rica Rica NNP 32505 657 30 and and CC 32505 657 31 British British NNP 32505 657 32 Honduras Honduras NNP 32505 657 33 it -PRON- PRP 32505 657 34 is be VBZ 32505 657 35 smaller small JJR 32505 657 36 . . . 32505 658 1 The the DT 32505 658 2 color color NN 32505 658 3 pattern pattern NN 32505 658 4 is be VBZ 32505 658 5 highly highly RB 32505 658 6 variable variable JJ 32505 658 7 . . . 32505 659 1 Some some DT 32505 659 2 specimens specimen NNS 32505 659 3 are be VBP 32505 659 4 dark dark JJ 32505 659 5 brown brown JJ 32505 659 6 or or CC 32505 659 7 pale pale JJ 32505 659 8 brown brown JJ 32505 659 9 in in IN 32505 659 10 color color NN 32505 659 11 . . . 32505 660 1 Incomplete incomplete JJ 32505 660 2 dorsal dorsal NN 32505 660 3 stripes stripe NNS 32505 660 4 are be VBP 32505 660 5 present present JJ 32505 660 6 in in IN 32505 660 7 94.6 94.6 CD 32505 660 8 per per IN 32505 660 9 cent cent NN 32505 660 10 of of IN 32505 660 11 the the DT 32505 660 12 specimens specimen NNS 32505 660 13 , , , 32505 660 14 and and CC 32505 660 15 transverse transverse JJ 32505 660 16 bars bar NNS 32505 660 17 are be VBP 32505 660 18 present present JJ 32505 660 19 on on IN 32505 660 20 the the DT 32505 660 21 shanks shank NNS 32505 660 22 in in IN 32505 660 23 98.3 98.3 CD 32505 660 24 per per IN 32505 660 25 cent cent NN 32505 660 26 of of IN 32505 660 27 the the DT 32505 660 28 specimens specimen NNS 32505 660 29 . . . 32505 661 1 The the DT 32505 661 2 interorbital interorbital JJ 32505 661 3 spot spot NN 32505 661 4 varies vary VBZ 32505 661 5 from from IN 32505 661 6 transverse transverse NNP 32505 661 7 to to IN 32505 661 8 longitudinal longitudinal VB 32505 661 9 in in IN 32505 661 10 position position NN 32505 661 11 , , , 32505 661 12 and and CC 32505 661 13 an an DT 32505 661 14 irregular irregular JJ 32505 661 15 white white JJ 32505 661 16 line line NN 32505 661 17 extends extend VBZ 32505 661 18 from from IN 32505 661 19 the the DT 32505 661 20 upper upper JJ 32505 661 21 jaw jaw NN 32505 661 22 to to IN 32505 661 23 the the DT 32505 661 24 arm arm NN 32505 661 25 in in IN 32505 661 26 some some DT 32505 661 27 specimens specimen NNS 32505 661 28 ( ( -LRB- 32505 661 29 Table table NN 32505 661 30 7 7 CD 32505 661 31 ) ) -RRB- 32505 661 32 . . . 32505 662 1 _ _ NNP 32505 662 2 Distribution._--_Hyla Distribution._--_Hyla NNP 32505 662 3 staufferi staufferi NNP 32505 662 4 staufferi staufferi NNP 32505 662 5 _ _ NNP 32505 662 6 inhabits inhabit VBZ 32505 662 7 savanna savanna NN 32505 662 8 and and CC 32505 662 9 subhumid subhumid NN 32505 662 10 and and CC 32505 662 11 xeric xeric JJ 32505 662 12 forests forest NNS 32505 662 13 in in IN 32505 662 14 the the DT 32505 662 15 lowlands lowland NNS 32505 662 16 and and CC 32505 662 17 moderate moderate JJ 32505 662 18 elevations elevation NNS 32505 662 19 from from IN 32505 662 20 southern southern JJ 32505 662 21 Tamaulipas Tamaulipas NNP 32505 662 22 southward southward RB 32505 662 23 to to IN 32505 662 24 Nicaragua Nicaragua NNP 32505 662 25 on on IN 32505 662 26 the the DT 32505 662 27 Caribbean caribbean JJ 32505 662 28 versant versant NN 32505 662 29 and and CC 32505 662 30 from from IN 32505 662 31 Guerrero Guerrero NNP 32505 662 32 , , , 32505 662 33 México México NNP 32505 662 34 to to IN 32505 662 35 northwestern northwestern JJ 32505 662 36 Costa Costa NNP 32505 662 37 Rica Rica NNP 32505 662 38 on on IN 32505 662 39 the the DT 32505 662 40 Pacific Pacific NNP 32505 662 41 lowlands lowland NNS 32505 662 42 ( ( -LRB- 32505 662 43 Fig fig NN 32505 662 44 . . . 32505 663 1 7 7 LS 32505 663 2 ) ) -RRB- 32505 663 3 . . . 32505 664 1 Duellman duellman NN 32505 664 2 ( ( -LRB- 32505 664 3 1963:226 1963:226 CD 32505 664 4 ) ) -RRB- 32505 664 5 commented comment VBD 32505 664 6 that that IN 32505 664 7 a a DT 32505 664 8 specimen speciman NNS 32505 664 9 from from IN 32505 664 10 Chinajá Chinajá NNP 32505 664 11 , , , 32505 664 12 Guatemala Guatemala NNP 32505 664 13 , , , 32505 664 14 possibly possibly RB 32505 664 15 was be VBD 32505 664 16 transported transport VBN 32505 664 17 there there RB 32505 664 18 in in IN 32505 664 19 the the DT 32505 664 20 cargo cargo NN 32505 664 21 from from IN 32505 664 22 Toocog Toocog NNP 32505 664 23 , , , 32505 664 24 because because IN 32505 664 25 with with IN 32505 664 26 this this DT 32505 664 27 one one CD 32505 664 28 exception exception NN 32505 664 29 the the DT 32505 664 30 species species NN 32505 664 31 is be VBZ 32505 664 32 unknown unknown JJ 32505 664 33 in in IN 32505 664 34 tropical tropical JJ 32505 664 35 rainforest rainforest NN 32505 664 36 in in IN 32505 664 37 Guatemala Guatemala NNP 32505 664 38 . . . 32505 665 1 [ [ -LRB- 32505 665 2 Illustration illustration NN 32505 665 3 : : : 32505 665 4 Fig Fig NNP 32505 665 5 . . . 32505 666 1 7 7 LS 32505 666 2 . . . 32505 667 1 Map Map NNP 32505 667 2 showing show VBG 32505 667 3 locality locality NN 32505 667 4 records record NNS 32505 667 5 for for IN 32505 667 6 _ _ NNP 32505 667 7 Hyla Hyla NNP 32505 667 8 staufferi staufferi NN 32505 667 9 staufferi staufferi NN 32505 667 10 _ _ NNP 32505 667 11 ( ( -LRB- 32505 667 12 circles circles NNP 32505 667 13 ) ) -RRB- 32505 667 14 and and CC 32505 667 15 _ _ NNP 32505 667 16 H. H. NNP 32505 668 1 staufferi staufferi NNP 32505 668 2 altae altae NNP 32505 668 3 _ _ NNP 32505 668 4 ( ( -LRB- 32505 668 5 dots dot NNS 32505 668 6 ) ) -RRB- 32505 668 7 . . . 32505 668 8 ] ] -RRB- 32505 669 1 _ _ NNP 32505 669 2 Specimens Specimens NNP 32505 669 3 Examined._--México Examined._--México NNP 32505 669 4 : : : 32505 669 5 _ _ NNP 32505 669 6 Campeche Campeche NNP 32505 669 7 _ _ NNP 32505 669 8 : : : 32505 669 9 5 5 CD 32505 669 10 km km NN 32505 669 11 S S NNP 32505 669 12 Champotón Champotón NNP 32505 669 13 KU KU NNP 32505 669 14 71296 71296 CD 32505 669 15 - - SYM 32505 669 16 7 7 CD 32505 669 17 ; ; : 32505 669 18 7 7 CD 32505 669 19 km km NN 32505 669 20 W W NNP 32505 669 21 Escárcega Escárcega NNP 32505 669 22 , , , 32505 669 23 KU KU NNP 32505 669 24 71298 71298 CD 32505 669 25 - - SYM 32505 669 26 308 308 CD 32505 669 27 ; ; : 32505 669 28 13 13 CD 32505 669 29 km km NN 32505 669 30 W w NN 32505 669 31 , , , 32505 669 32 1 1 CD 32505 669 33 km km NN 32505 669 34 N N NNP 32505 669 35 Escárcega Escárcega NNP 32505 669 36 , , , 32505 669 37 KU KU NNP 32505 669 38 71309 71309 CD 32505 669 39 - - SYM 32505 669 40 10 10 CD 32505 669 41 , , , 32505 669 42 75090 75090 CD 32505 669 43 - - SYM 32505 669 44 4 4 CD 32505 669 45 . . . 32505 670 1 _ _ NNP 32505 670 2 Chiapas Chiapas NNP 32505 670 3 _ _ NNP 32505 670 4 : : : 32505 670 5 32 32 CD 32505 670 6 km km NN 32505 670 7 S S NNP 32505 670 8 Arriaga Arriaga NNP 32505 670 9 , , , 32505 670 10 KU KU NNP 32505 670 11 57789 57789 CD 32505 670 12 - - SYM 32505 670 13 92 92 CD 32505 670 14 ; ; : 32505 670 15 4 4 CD 32505 670 16 km km NN 32505 670 17 N N NNP 32505 670 18 Ixtapa Ixtapa NNP 32505 670 19 , , , 32505 670 20 KU KU NNP 32505 670 21 5776 5776 CD 32505 670 22 - - SYM 32505 670 23 81 81 CD 32505 670 24 ; ; : 32505 670 25 3.6 3.6 CD 32505 670 26 km km NN 32505 670 27 SW SW NNP 32505 670 28 Las Las NNP 32505 670 29 Cruces Cruces NNPS 32505 670 30 , , , 32505 670 31 KU KU NNP 32505 670 32 37740 37740 CD 32505 670 33 ; ; : 32505 670 34 17 17 CD 32505 670 35 km km NN 32505 670 36 S S NNP 32505 670 37 Las Las NNP 32505 670 38 Cruces Cruces NNPS 32505 670 39 , , , 32505 670 40 KU KU NNP 32505 670 41 57793 57793 CD 32505 670 42 - - SYM 32505 670 43 4 4 CD 32505 670 44 ; ; : 32505 670 45 24 24 CD 32505 670 46 km km NN 32505 670 47 S S NNP 32505 670 48 Las Las NNP 32505 670 49 Cruces Cruces NNPS 32505 670 50 , , , 32505 670 51 KU KU NNP 32505 670 52 104160 104160 CD 32505 670 53 ( ( -LRB- 32505 670 54 tadpoles tadpole NNS 32505 670 55 ) ) -RRB- 32505 670 56 ; ; : 32505 670 57 11 11 CD 32505 670 58 km km NN 32505 670 59 S S NNP 32505 670 60 Tapachula Tapachula NNP 32505 670 61 , , , 32505 670 62 KU KU NNP 32505 670 63 57782 57782 CD 32505 670 64 - - SYM 32505 670 65 8 8 CD 32505 670 66 , , , 32505 670 67 60000 60000 CD 32505 670 68 ( ( -LRB- 32505 670 69 young young JJ 32505 670 70 ) ) -RRB- 32505 670 71 . . . 32505 671 1 _ _ NNP 32505 671 2 Guerrero Guerrero NNP 32505 671 3 _ _ NNP 32505 671 4 : : : 32505 671 5 El El NNP 32505 671 6 Limoncito Limoncito NNP 32505 671 7 , , , 32505 671 8 near near IN 32505 671 9 La La NNP 32505 671 10 Venta Venta NNP 32505 671 11 , , , 32505 671 12 KU KU NNP 32505 671 13 31392 31392 CD 32505 671 14 - - SYM 32505 671 15 401 401 CD 32505 671 16 ; ; : 32505 671 17 Mexcala Mexcala NNP 32505 671 18 , , , 32505 671 19 near near IN 32505 671 20 Balsa Balsa NNP 32505 671 21 River River NNP 32505 671 22 , , , 32505 671 23 KU KU NNP 32505 671 24 31391 31391 CD 32505 671 25 ; ; : 32505 671 26 Organos Organos NNP 32505 671 27 , , , 32505 671 28 S S NNP 32505 671 29 El El NNP 32505 671 30 Trienta Trienta NNP 32505 671 31 , , , 32505 671 32 KU KU NNP 32505 671 33 31390 31390 CD 32505 671 34 . . . 32505 672 1 _ _ NNP 32505 672 2 Oaxaca Oaxaca NNP 32505 672 3 _ _ NNP 32505 672 4 : : : 32505 672 5 26 26 CD 32505 672 6 km km NN 32505 672 7 N N NNP 32505 672 8 Matías Matías NNP 32505 672 9 Romero Romero NNP 32505 672 10 , , , 32505 672 11 KU KU NNP 32505 672 12 33878 33878 CD 32505 672 13 - - SYM 32505 672 14 82 82 CD 32505 672 15 ; ; : 32505 672 16 2.5 2.5 CD 32505 672 17 km km NN 32505 672 18 S S NNP 32505 672 19 Pochutla Pochutla NNP 32505 672 20 , , , 32505 672 21 KU KU NNP 32505 672 22 59924 59924 CD 32505 672 23 - - SYM 32505 672 24 7 7 CD 32505 672 25 ( ( -LRB- 32505 672 26 skeletons skeleton NNS 32505 672 27 ) ) -RRB- 32505 672 28 ; ; : 32505 672 29 5 5 LS 32505 672 30 km km NN 32505 672 31 S S NNP 32505 672 32 Pochutla Pochutla NNP 32505 672 33 , , , 32505 672 34 KU KU NNP 32505 672 35 57795 57795 CD 32505 672 36 - - SYM 32505 672 37 801 801 CD 32505 672 38 ; ; : 32505 672 39 3.2 3.2 CD 32505 672 40 km km NN 32505 672 41 E E NNP 32505 672 42 Tapanatepec Tapanatepec NNP 32505 672 43 , , , 32505 672 44 KU KU NNP 32505 672 45 37877 37877 CD 32505 672 46 - - SYM 32505 672 47 902 902 CD 32505 672 48 ; ; : 32505 672 49 17.6 17.6 CD 32505 672 50 km km NN 32505 672 51 WNW WNW NNP 32505 672 52 Tapanatepec Tapanatepec NNP 32505 672 53 , , , 32505 672 54 KU KU NNP 32505 672 55 65033 65033 CD 32505 672 56 - - SYM 32505 672 57 4 4 CD 32505 672 58 ; ; : 32505 672 59 Temascal temascal JJ 32505 672 60 , , , 32505 672 61 USC USC NNP 32505 672 62 8243 8243 CD 32505 672 63 ( ( -LRB- 32505 672 64 8) 8) CD 32505 672 65 ; ; : 32505 672 66 3.2 3.2 CD 32505 672 67 km km NN 32505 672 68 S S NNP 32505 672 69 Tolocita Tolocita NNP 32505 672 70 , , , 32505 672 71 KU KU NNP 32505 672 72 39657 39657 CD 32505 672 73 - - SYM 32505 672 74 8 8 CD 32505 672 75 ; ; : 32505 672 76 0.5 0.5 CD 32505 672 77 km km NN 32505 672 78 Tuxtepec Tuxtepec NNP 32505 672 79 , , , 32505 672 80 KU KU NNP 32505 672 81 87073 87073 CD 32505 672 82 - - SYM 32505 672 83 81 81 CD 32505 672 84 , , , 32505 672 85 87610 87610 CD 32505 672 86 ( ( -LRB- 32505 672 87 tadpoles tadpole NNS 32505 672 88 ) ) -RRB- 32505 672 89 ; ; : 32505 672 90 17 17 CD 32505 672 91 km km NN 32505 672 92 S S NNP 32505 672 93 Tuxtepec Tuxtepec NNP 32505 672 94 , , , 32505 672 95 KU KU NNP 32505 672 96 65035 65035 CD 32505 672 97 - - SYM 32505 672 98 7 7 CD 32505 672 99 ; ; : 32505 672 100 1 1 LS 32505 672 101 km km NN 32505 672 102 W W NNP 32505 672 103 Zanatepec Zanatepec NNP 32505 672 104 , , , 32505 672 105 KU KU NNP 32505 672 106 104161 104161 CD 32505 672 107 ( ( -LRB- 32505 672 108 tadpoles tadpole NNS 32505 672 109 ) ) -RRB- 32505 672 110 . . . 32505 673 1 _ _ NNP 32505 673 2 Quintana Quintana NNP 32505 673 3 Roo Roo NNP 32505 673 4 _ _ NNP 32505 673 5 : : : 32505 673 6 Isla Isla NNP 32505 673 7 Cozumel Cozumel NNP 32505 673 8 , , , 32505 673 9 3.5 3.5 CD 32505 673 10 km km NN 32505 673 11 N N NNP 32505 673 12 San San NNP 32505 673 13 Miguel Miguel NNP 32505 673 14 , , , 32505 673 15 KU KU NNP 32505 673 16 71710 71710 CD 32505 673 17 - - SYM 32505 673 18 11 11 CD 32505 673 19 ( ( -LRB- 32505 673 20 young young JJ 32505 673 21 ) ) -RRB- 32505 673 22 . . . 32505 674 1 _ _ NNP 32505 674 2 San San NNP 32505 674 3 Luis Luis NNP 32505 674 4 Potosí Potosí NNP 32505 674 5 _ _ NNP 32505 674 6 : : : 32505 674 7 Valles Valles NNP 32505 674 8 , , , 32505 674 9 KU KU NNP 32505 674 10 31490 31490 CD 32505 674 11 . . . 32505 675 1 _ _ NNP 32505 675 2 Tabasco Tabasco NNP 32505 675 3 _ _ NNP 32505 675 4 : : : 32505 675 5 Teapa Teapa NNP 32505 675 6 , , , 32505 675 7 UMMZ UMMZ NNP 32505 675 8 118887 118887 CD 32505 675 9 ( ( -LRB- 32505 675 10 3 3 CD 32505 675 11 ) ) -RRB- 32505 675 12 , , , 32505 675 13 119203 119203 CD 32505 675 14 ( ( -LRB- 32505 675 15 13 13 CD 32505 675 16 ) ) -RRB- 32505 675 17 ; ; : 32505 675 18 9.6 9.6 CD 32505 675 19 km km NN 32505 675 20 N N NNP 32505 675 21 Teapa Teapa NNP 32505 675 22 , , , 32505 675 23 UMMZ UMMZ NNP 32505 675 24 119202 119202 CD 32505 675 25 ; ; : 32505 675 26 24 24 CD 32505 675 27 km km NN 32505 675 28 N N NNP 32505 675 29 Teapa Teapa NNP 32505 675 30 , , , 32505 675 31 UMMZ UMMZ NNP 32505 675 32 119961 119961 CD 32505 675 33 ( ( -LRB- 32505 675 34 5 5 CD 32505 675 35 ) ) -RRB- 32505 675 36 ; ; : 32505 675 37 29 29 CD 32505 675 38 km km NN 32505 675 39 N N NNP 32505 675 40 Teapa Teapa NNP 32505 675 41 , , , 32505 675 42 UMMZ UMMZ NNP 32505 675 43 119960 119960 CD 32505 675 44 ; ; : 32505 675 45 3.5 3.5 CD 32505 675 46 km km NN 32505 675 47 S S NNP 32505 675 48 Villahermosa Villahermosa NNP 32505 675 49 , , , 32505 675 50 UMMZ UMMZ NNP 32505 675 51 119201 119201 CD 32505 675 52 ( ( -LRB- 32505 675 53 2 2 CD 32505 675 54 ) ) -RRB- 32505 675 55 ; ; : 32505 675 56 17.6 17.6 CD 32505 675 57 km km NN 32505 675 58 S S NNP 32505 675 59 Villahermosa Villahermosa NNP 32505 675 60 , , , 32505 675 61 UMMZ UMMZ NNP 32505 675 62 119200 119200 CD 32505 675 63 ( ( -LRB- 32505 675 64 8) 8) CD 32505 675 65 . . . 32505 676 1 _ _ NNP 32505 676 2 Tamaulipas Tamaulipas NNP 32505 676 3 _ _ NNP 32505 676 4 : : : 32505 676 5 1 1 CD 32505 676 6 km km NN 32505 676 7 E E NNP 32505 676 8 Chamal Chamal NNP 32505 676 9 , , , 32505 676 10 UMMZ UMMZ NNP 32505 676 11 110706 110706 CD 32505 676 12 ; ; : 32505 676 13 Gómez Gómez NNP 32505 676 14 Farías Farías NNP 32505 676 15 , , , 32505 676 16 UMMZ UMMZ NNP 32505 676 17 110701 110701 CD 32505 676 18 ( ( -LRB- 32505 676 19 3 3 CD 32505 676 20 ) ) -RRB- 32505 676 21 ; ; : 32505 676 22 5 5 CD 32505 676 23 km km NN 32505 676 24 SE SE NNP 32505 676 25 Gómez Gómez NNP 32505 676 26 Farías Farías NNP 32505 676 27 , , , 32505 676 28 UMMZ UMMZ NNP 32505 676 29 110705 110705 CD 32505 676 30 ; ; : 32505 676 31 8 8 CD 32505 676 32 km km NN 32505 676 33 NE NE NNP 32505 676 34 Gómez Gómez NNP 32505 676 35 Farías Farías NNP 32505 676 36 , , , 32505 676 37 UMMZ UMMZ NNP 32505 676 38 11282 11282 CD 32505 676 39 ( ( -LRB- 32505 676 40 2 2 CD 32505 676 41 ) ) -RRB- 32505 676 42 , , , 32505 676 43 11283 11283 CD 32505 676 44 ( ( -LRB- 32505 676 45 3 3 CD 32505 676 46 ) ) -RRB- 32505 676 47 ; ; : 32505 676 48 Kilometer kilometer NN 32505 676 49 615 615 CD 32505 676 50 between between IN 32505 676 51 Río Río NNP 32505 676 52 Limón Limón NNP 32505 676 53 and and CC 32505 676 54 Llera Llera NNP 32505 676 55 , , , 32505 676 56 UMMZ UMMZ NNP 32505 676 57 80455 80455 CD 32505 676 58 ( ( -LRB- 32505 676 59 2 2 CD 32505 676 60 ) ) -RRB- 32505 676 61 ; ; : 32505 676 62 5 5 LS 32505 676 63 km km NN 32505 676 64 W W NNP 32505 676 65 San San NNP 32505 676 66 Geraldo Geraldo NNP 32505 676 67 , , , 32505 676 68 UMMZ UMMZ NNP 32505 676 69 110702 110702 CD 32505 676 70 ( ( -LRB- 32505 676 71 4 4 CD 32505 676 72 ) ) -RRB- 32505 676 73 , , , 32505 676 74 110703 110703 CD 32505 676 75 ( ( -LRB- 32505 676 76 3 3 CD 32505 676 77 ) ) -RRB- 32505 676 78 ; ; : 32505 676 79 8 8 CD 32505 676 80 km km NN 32505 676 81 W W NNP 32505 676 82 San San NNP 32505 676 83 Geraldo Geraldo NNP 32505 676 84 , , , 32505 676 85 near near IN 32505 676 86 Río Río NNP 32505 676 87 Frío Frío NNP 32505 676 88 , , , 32505 676 89 UMMZ UMMZ NNP 32505 676 90 110704 110704 CD 32505 676 91 ( ( -LRB- 32505 676 92 5 5 CD 32505 676 93 ) ) -RRB- 32505 676 94 . . . 32505 677 1 _ _ NNP 32505 677 2 Veracruz Veracruz NNP 32505 677 3 _ _ NNP 32505 677 4 : : : 32505 677 5 3 3 CD 32505 677 6 km km NN 32505 677 7 SW SW NNP 32505 677 8 Boca Boca NNP 32505 677 9 del del NNP 32505 677 10 Río Río NNP 32505 677 11 , , , 32505 677 12 KU KU NNP 32505 677 13 10494 10494 CD 32505 677 14 - - SYM 32505 677 15 8 8 CD 32505 677 16 ; ; : 32505 677 17 5 5 CD 32505 677 18 km km NN 32505 677 19 SW SW NNP 32505 677 20 Boca Boca NNP 32505 677 21 del del NNP 32505 677 22 Río Río NNP 32505 677 23 , , , 32505 677 24 KU KU NNP 32505 677 25 23701 23701 CD 32505 677 26 ; ; : 32505 677 27 5 5 CD 32505 677 28 km km NN 32505 677 29 ESE ESE NNP 32505 677 30 Córdoba Córdoba NNP 32505 677 31 , , , 32505 677 32 KU KU NNP 32505 677 33 104162 104162 CD 32505 677 34 ( ( -LRB- 32505 677 35 tadpoles tadpole NNS 32505 677 36 ) ) -RRB- 32505 677 37 ; ; : 32505 677 38 Cuautlapán Cuautlapán NNP 32505 677 39 , , , 32505 677 40 KU KU NNP 32505 677 41 57098 57098 CD 32505 677 42 - - SYM 32505 677 43 102 102 CD 32505 677 44 , , , 32505 677 45 26787 26787 CD 32505 677 46 ; ; : 32505 677 47 Hacienda Hacienda NNP 32505 677 48 Tamiahua Tamiahua NNP 32505 677 49 , , , 32505 677 50 Cabo Cabo NNP 32505 677 51 Rojo Rojo NNP 32505 677 52 , , , 32505 677 53 KU KU NNP 32505 677 54 62871 62871 CD 32505 677 55 ; ; : 32505 677 56 2 2 CD 32505 677 57 km km NN 32505 677 58 ENE ENE NNP 32505 677 59 Mata Mata NNP 32505 677 60 Oscura Oscura NNP 32505 677 61 , , , 32505 677 62 KU KU NNP 32505 677 63 105627 105627 CD 32505 677 64 ; ; : 32505 677 65 5 5 CD 32505 677 66 km km NN 32505 677 67 SE SE NNP 32505 677 68 Paso Paso NNP 32505 677 69 del del NNP 32505 677 70 Toro Toro NNP 32505 677 71 , , , 32505 677 72 KU KU NNP 32505 677 73 40144 40144 CD 32505 677 74 ; ; : 32505 677 75 Portrero Portrero NNP 32505 677 76 Viejo Viejo NNP 32505 677 77 , , , 32505 677 78 KU KU NNP 32505 677 79 23911 23911 CD 32505 677 80 - - SYM 32505 677 81 2 2 CD 32505 677 82 , , , 32505 677 83 26786 26786 CD 32505 677 84 , , , 32505 677 85 27413 27413 CD 32505 677 86 , , , 32505 677 87 57094 57094 CD 32505 677 88 - - SYM 32505 677 89 7 7 CD 32505 677 90 . . . 32505 678 1 Guatemala Guatemala NNP 32505 678 2 : : : 32505 678 3 _ _ NNP 32505 678 4 Alta Alta NNP 32505 678 5 Verapaz Verapaz NNP 32505 678 6 _ _ NNP 32505 678 7 : : : 32505 678 8 Chinajá Chinajá NNP 32505 678 9 , , , 32505 678 10 KU KU NNP 32505 678 11 57769 57769 CD 32505 678 12 ; ; : 32505 678 13 Finca Finca NNP 32505 678 14 La La NNP 32505 678 15 Cubilquitz Cubilquitz NNP 32505 678 16 , , , 32505 678 17 UMMZ UMMZ NNP 32505 678 18 90871 90871 CD 32505 678 19 , , , 32505 678 20 90872 90872 CD 32505 678 21 ( ( -LRB- 32505 678 22 5 5 CD 32505 678 23 ) ) -RRB- 32505 678 24 , , , 32505 678 25 91379 91379 CD 32505 678 26 ( ( -LRB- 32505 678 27 2 2 CD 32505 678 28 ) ) -RRB- 32505 678 29 . . . 32505 679 1 _ _ NNP 32505 679 2 Baja Baja NNP 32505 679 3 Verapaz Verapaz NNP 32505 679 4 _ _ NNP 32505 679 5 : : : 32505 679 6 1 1 LS 32505 679 7 km km NN 32505 679 8 S S NNP 32505 679 9 San San NNP 32505 679 10 Jerónimo Jerónimo NNP 32505 679 11 , , , 32505 679 12 UMMZ UMMZ NNP 32505 679 13 84077 84077 CD 32505 679 14 ( ( -LRB- 32505 679 15 7 7 CD 32505 679 16 ) ) -RRB- 32505 679 17 , , , 32505 679 18 84078 84078 CD 32505 679 19 ( ( -LRB- 32505 679 20 14 14 CD 32505 679 21 ) ) -RRB- 32505 679 22 . . . 32505 680 1 _ _ NNP 32505 680 2 Chiquimula Chiquimula NNP 32505 680 3 _ _ NNP 32505 680 4 : : : 32505 680 5 1.6 1.6 CD 32505 680 6 km km NN 32505 680 7 SE SE NNP 32505 680 8 Chiquimula Chiquimula NNP 32505 680 9 , , , 32505 680 10 UMMZ UMMZ NNP 32505 680 11 98114 98114 CD 32505 680 12 ( ( -LRB- 32505 680 13 2 2 CD 32505 680 14 ) ) -RRB- 32505 680 15 ; ; : 32505 680 16 Esquipulas Esquipulas NNP 32505 680 17 , , , 32505 680 18 UMMZ UMMZ NNP 32505 680 19 106784 106784 CD 32505 680 20 ( ( -LRB- 32505 680 21 4 4 CD 32505 680 22 ) ) -RRB- 32505 680 23 , , , 32505 680 24 106785 106785 CD 32505 680 25 ( ( -LRB- 32505 680 26 14 14 CD 32505 680 27 ) ) -RRB- 32505 680 28 . . . 32505 681 1 _ _ NNP 32505 681 2 El El NNP 32505 681 3 Petén Petén NNP 32505 681 4 _ _ NNP 32505 681 5 : : : 32505 681 6 No no DT 32505 681 7 specific specific JJ 32505 681 8 locality locality NN 32505 681 9 , , , 32505 681 10 USNM USNM NNP 32505 681 11 25143 25143 CD 32505 681 12 , , , 32505 681 13 24825 24825 CD 32505 681 14 - - SYM 32505 681 15 6 6 CD 32505 681 16 ; ; : 32505 681 17 La La NNP 32505 681 18 Libertad Libertad NNP 32505 681 19 , , , 32505 681 20 FMNH FMNH NNP 32505 681 21 27096 27096 CD 32505 681 22 - - SYM 32505 681 23 7 7 CD 32505 681 24 , , , 32505 681 25 KU KU NNP 32505 681 26 57770 57770 CD 32505 681 27 , , , 32505 681 28 UMMZ UMMZ NNP 32505 681 29 75339 75339 CD 32505 681 30 ( ( -LRB- 32505 681 31 15 15 CD 32505 681 32 ) ) -RRB- 32505 681 33 , , , 32505 681 34 75340 75340 CD 32505 681 35 ( ( -LRB- 32505 681 36 15 15 CD 32505 681 37 ) ) -RRB- 32505 681 38 , , , 32505 681 39 UMMZ UMMZ NNP 32505 681 40 94341 94341 CD 32505 681 41 - - SYM 32505 681 42 2 2 CD 32505 681 43 . . . 32505 682 1 _ _ NNP 32505 682 2 Esquintla Esquintla NNP 32505 682 3 _ _ NNP 32505 682 4 : : : 32505 682 5 20 20 CD 32505 682 6 km km NN 32505 682 7 N N NNP 32505 682 8 San San NNP 32505 682 9 José José NNP 32505 682 10 , , , 32505 682 11 AMNH AMNH NNP 32505 682 12 74369 74369 CD 32505 682 13 - - SYM 32505 682 14 76 76 CD 32505 682 15 . . . 32505 683 1 _ _ NNP 32505 683 2 Guatamala Guatamala NNP 32505 683 3 _ _ NNP 32505 683 4 : : : 32505 683 5 16 16 CD 32505 683 6 km km NN 32505 683 7 NE NE NNP 32505 683 8 Guatamala Guatamala NNP 32505 683 9 , , , 32505 683 10 KU KU NNP 32505 683 11 43539 43539 CD 32505 683 12 . . . 32505 684 1 Izábal izábal JJ 32505 684 2 : : : 32505 684 3 Puerto Puerto NNP 32505 684 4 Barrios Barrios NNPS 32505 684 5 , , , 32505 684 6 TCWC TCWC NNP 32505 684 7 16671 16671 CD 32505 684 8 - - SYM 32505 684 9 73 73 CD 32505 684 10 , , , 32505 684 11 16646 16646 CD 32505 684 12 - - SYM 32505 684 13 56 56 CD 32505 684 14 ; ; : 32505 684 15 2.5 2.5 CD 32505 684 16 km km NN 32505 684 17 NE NE NNP 32505 684 18 Río Río NNP 32505 684 19 Blanco Blanco NNP 32505 684 20 , , , 32505 684 21 KU KU NNP 32505 684 22 57774 57774 CD 32505 684 23 - - SYM 32505 684 24 5 5 CD 32505 684 25 . . . 32505 685 1 _ _ NNP 32505 685 2 Jalapa Jalapa NNP 32505 685 3 _ _ NNP 32505 685 4 : : : 32505 685 5 Jalapa Jalapa NNP 32505 685 6 , , , 32505 685 7 UMMZ UMMZ NNP 32505 685 8 106788 106788 CD 32505 685 9 ( ( -LRB- 32505 685 10 44 44 CD 32505 685 11 ) ) -RRB- 32505 685 12 . . . 32505 686 1 _ _ NNP 32505 686 2 Jutiapa Jutiapa NNP 32505 686 3 _ _ NNP 32505 686 4 : : : 32505 686 5 Finca Finca NNP 32505 686 6 La La NNP 32505 686 7 Trinidad Trinidad NNP 32505 686 8 , , , 32505 686 9 UMMZ UMMZ NNP 32505 686 10 107730 107730 CD 32505 686 11 ( ( -LRB- 32505 686 12 12 12 CD 32505 686 13 ) ) -RRB- 32505 686 14 , , , 32505 686 15 107731 107731 CD 32505 686 16 ( ( -LRB- 32505 686 17 16 16 CD 32505 686 18 ) ) -RRB- 32505 686 19 ; ; : 32505 686 20 Jutiapa Jutiapa NNP 32505 686 21 , , , 32505 686 22 UMMZ UMMZ NNP 32505 686 23 106786 106786 CD 32505 686 24 ( ( -LRB- 32505 686 25 2 2 CD 32505 686 26 ) ) -RRB- 32505 686 27 . . . 32505 687 1 _ _ NNP 32505 687 2 Zacapa Zacapa NNP 32505 687 3 _ _ NNP 32505 687 4 : : : 32505 687 5 14 14 CD 32505 687 6 km km NN 32505 687 7 ENE ENE NNP 32505 687 8 Mayuelas Mayuelas NNP 32505 687 9 , , , 32505 687 10 KU KU NNP 32505 687 11 57773 57773 CD 32505 687 12 ; ; : 32505 687 13 7 7 CD 32505 687 14 km km NN 32505 687 15 ENE ENE NNP 32505 687 16 Río Río NNP 32505 687 17 Hondo Hondo NNP 32505 687 18 , , , 32505 687 19 KU KU NNP 32505 687 20 57771 57771 CD 32505 687 21 - - SYM 32505 687 22 2 2 CD 32505 687 23 , , , 32505 687 24 59999 59999 CD 32505 687 25 ( ( -LRB- 32505 687 26 young young JJ 32505 687 27 ) ) -RRB- 32505 687 28 . . . 32505 688 1 British british JJ 32505 688 2 Honduras Honduras NNP 32505 688 3 : : : 32505 688 4 BELIZE BELIZE NNP 32505 688 5 : : : 32505 688 6 Belize Belize NNP 32505 688 7 , , , 32505 688 8 FMNH FMNH NNP 32505 688 9 4406 4406 CD 32505 688 10 . . . 32505 689 1 _ _ NNP 32505 689 2 El El NNP 32505 689 3 Cayo Cayo NNP 32505 689 4 _ _ NNP 32505 689 5 : : : 32505 689 6 San San NNP 32505 689 7 Agustín Agustín NNP 32505 689 8 , , , 32505 689 9 UMMZ UMMZ NNP 32505 689 10 80741 80741 CD 32505 689 11 ( ( -LRB- 32505 689 12 8) 8) CD 32505 689 13 . . . 32505 690 1 _ _ NNP 32505 690 2 Stann Stann NNP 32505 690 3 Creek Creek NNP 32505 690 4 _ _ NNP 32505 690 5 : : : 32505 690 6 10 10 CD 32505 690 7 km km NN 32505 690 8 S S NNP 32505 690 9 Stann Stann NNP 32505 690 10 Creek Creek NNP 32505 690 11 on on IN 32505 690 12 Hummingbird Hummingbird NNP 32505 690 13 Highway Highway NNP 32505 690 14 , , , 32505 690 15 UMMZ UMMZ NNP 32505 690 16 125720 125720 CD 32505 690 17 - - SYM 32505 690 18 1 1 CD 32505 690 19 . . . 32505 691 1 El El NNP 32505 691 2 Salvador Salvador NNP 32505 691 3 : : : 32505 691 4 _ _ NNP 32505 691 5 Cuscatlán Cuscatlán NNP 32505 691 6 _ _ NNP 32505 691 7 : : : 32505 691 8 7 7 CD 32505 691 9 km km NN 32505 691 10 WNW WNW NNP 32505 691 11 Cojutepeque Cojutepeque NNP 32505 691 12 , , , 32505 691 13 TNHC TNHC NNP 32505 691 14 32004 32004 CD 32505 691 15 - - SYM 32505 691 16 10 10 CD 32505 691 17 . . . 32505 692 1 _ _ NNP 32505 692 2 La La NNP 32505 692 3 Libertad Libertad NNP 32505 692 4 _ _ NNP 32505 692 5 : : : 32505 692 6 16 16 CD 32505 692 7 km km NN 32505 692 8 NW NW NNP 32505 692 9 Santa Santa NNP 32505 692 10 Tecla Tecla NNP 32505 692 11 , , , 32505 692 12 KU KU NNP 32505 692 13 43540 43540 CD 32505 692 14 - - SYM 32505 692 15 1 1 CD 32505 692 16 . . . 32505 693 1 _ _ NNP 32505 693 2 La La NNP 32505 693 3 Union Union NNP 32505 693 4 _ _ NNP 32505 693 5 : : : 32505 693 6 2.5 2.5 CD 32505 693 7 km km NN 32505 693 8 Santa Santa NNP 32505 693 9 Rosa Rosa NNP 32505 693 10 , , , 32505 693 11 TCWC TCWC NNP 32505 693 12 16669 16669 CD 32505 693 13 - - SYM 32505 693 14 70 70 CD 32505 693 15 . . . 32505 694 1 _ _ NNP 32505 694 2 Morazán Morazán NNP 32505 694 3 _ _ NNP 32505 694 4 : : : 32505 694 5 Dividendero Dividendero NNP 32505 694 6 , , , 32505 694 7 USNM USNM NNP 32505 694 8 73288 73288 CD 32505 694 9 - - SYM 32505 694 10 92 92 CD 32505 694 11 . . . 32505 695 1 _ _ NNP 32505 695 2 San San NNP 32505 695 3 Salvador Salvador NNP 32505 695 4 _ _ NNP 32505 695 5 : : : 32505 695 6 San San NNP 32505 695 7 Salvador Salvador NNP 32505 695 8 , , , 32505 695 9 FMNH FMNH NNP 32505 695 10 65101 65101 CD 32505 695 11 - - SYM 32505 695 12 06 06 CD 32505 695 13 , , , 32505 695 14 KU KU NNP 32505 695 15 61932 61932 CD 32505 695 16 - - SYM 32505 695 17 44 44 CD 32505 695 18 , , , 32505 695 19 61989 61989 CD 32505 695 20 - - SYM 32505 695 21 92 92 CD 32505 695 22 , , , 32505 695 23 62152 62152 CD 32505 695 24 ( ( -LRB- 32505 695 25 eggs egg NNS 32505 695 26 ) ) -RRB- 32505 695 27 , , , 32505 695 28 USNM USNM NNP 32505 695 29 117588 117588 CD 32505 695 30 , , , 32505 695 31 118391 118391 CD 32505 695 32 ( ( -LRB- 32505 695 33 3 3 CD 32505 695 34 ) ) -RRB- 32505 695 35 , , , 32505 695 36 118394 118394 CD 32505 695 37 ; ; : 32505 695 38 1.6 1.6 CD 32505 695 39 km km NN 32505 695 40 NW NW NNP 32505 695 41 San San NNP 32505 695 42 Salvador Salvador NNP 32505 695 43 , , , 32505 695 44 KU KU NNP 32505 695 45 43162 43162 CD 32505 695 46 - - SYM 32505 695 47 3 3 CD 32505 695 48 . . . 32505 696 1 Honduras hondura NNS 32505 696 2 : : : 32505 696 3 _ _ NNP 32505 696 4 Atlantidad Atlantidad NNP 32505 696 5 _ _ NNP 32505 696 6 : : : 32505 696 7 Ceiba Ceiba NNP 32505 696 8 , , , 32505 696 9 USNM USNM NNP 32505 696 10 117592 117592 CD 32505 696 11 . . . 32505 697 1 _ _ NNP 32505 697 2 Choluteca Choluteca NNP 32505 697 3 _ _ NNP 32505 697 4 : : : 32505 697 5 Choluteca Choluteca NNP 32505 697 6 , , , 32505 697 7 KU KU NNP 32505 697 8 85361 85361 CD 32505 697 9 - - SYM 32505 697 10 6 6 CD 32505 697 11 ; ; : 32505 697 12 2 2 CD 32505 697 13 km km NN 32505 697 14 E E NNP 32505 697 15 Choluteca Choluteca NNP 32505 697 16 , , , 32505 697 17 UMMZ UMMZ NNP 32505 697 18 118395 118395 CD 32505 697 19 ( ( -LRB- 32505 697 20 7 7 CD 32505 697 21 ) ) -RRB- 32505 697 22 ; ; : 32505 697 23 3.2 3.2 CD 32505 697 24 km km NN 32505 697 25 NE NE NNP 32505 697 26 Choluteca Choluteca NNP 32505 697 27 , , , 32505 697 28 KU KU NNP 32505 697 29 100500 100500 CD 32505 697 30 - - SYM 32505 697 31 01 01 CD 32505 697 32 ; ; : 32505 697 33 6.2 6.2 CD 32505 697 34 km km NN 32505 697 35 E E NNP 32505 697 36 Choluteca Choluteca NNP 32505 697 37 , , , 32505 697 38 KU KU NNP 32505 697 39 65046 65046 CD 32505 697 40 - - SYM 32505 697 41 56 56 CD 32505 697 42 ; ; : 32505 697 43 10 10 CD 32505 697 44 km km NN 32505 697 45 E E NNP 32505 697 46 Choluteca Choluteca NNP 32505 697 47 , , , 32505 697 48 KU KU NNP 32505 697 49 65045 65045 CD 32505 697 50 : : : 32505 697 51 5 5 CD 32505 697 52 km km NN 32505 697 53 S S NNP 32505 697 54 Choluteca Choluteca NNP 32505 697 55 , , , 32505 697 56 USC USC NNP 32505 697 57 2700 2700 CD 32505 697 58 ( ( -LRB- 32505 697 59 4 4 CD 32505 697 60 ) ) -RRB- 32505 697 61 . . . 32505 698 1 _ _ NNP 32505 698 2 Colón Colón NNP 32505 698 3 _ _ NNP 32505 698 4 : : : 32505 698 5 Isla Isla NNP 32505 698 6 Guanaja Guanaja NNP 32505 698 7 ( ( -LRB- 32505 698 8 Islas Islas NNP 32505 698 9 de de NNP 32505 698 10 la la NNP 32505 698 11 Bahía Bahía NNP 32505 698 12 ) ) -RRB- 32505 698 13 , , , 32505 698 14 TCWC TCWC NNP 32505 698 15 21551 21551 CD 32505 698 16 , , , 32505 698 17 TNHC TNHC NNP 32505 698 18 32011 32011 CD 32505 698 19 . . . 32505 699 1 _ _ NNP 32505 699 2 Cortés Cortés NNP 32505 699 3 _ _ NNP 32505 699 4 : : : 32505 699 5 Agua Agua NNP 32505 699 6 Azul Azul NNP 32505 699 7 , , , 32505 699 8 TCWC TCWC NNP 32505 699 9 19178 19178 CD 32505 699 10 - - SYM 32505 699 11 9 9 CD 32505 699 12 ; ; : 32505 699 13 East east JJ 32505 699 14 side side NN 32505 699 15 Lago Lago NNP 32505 699 16 Yojoa Yojoa NNP 32505 699 17 , , , 32505 699 18 KU KU NNP 32505 699 19 65038 65038 CD 32505 699 20 - - SYM 32505 699 21 44 44 CD 32505 699 22 . . . 32505 700 1 _ _ NNP 32505 700 2 El El NNP 32505 700 3 Paraiso Paraiso NNP 32505 700 4 _ _ NNP 32505 700 5 : : : 32505 700 6 Valle Valle NNP 32505 700 7 de de NNP 32505 700 8 Jamastrán Jamastrán NNP 32505 700 9 , , , 32505 700 10 AMNH AMNH NNP 32505 700 11 54800 54800 CD 32505 700 12 - - SYM 32505 700 13 04 04 CD 32505 700 14 . . . 32505 701 1 _ _ NNP 32505 701 2 Francisco Francisco NNP 32505 701 3 Morazán Morazán NNP 32505 701 4 _ _ NNP 32505 701 5 : : : 32505 701 6 Escuela Escuela NNP 32505 701 7 Agrícola Agrícola NNP 32505 701 8 Panamericana Panamericana NNP 32505 701 9 , , , 32505 701 10 AMNH AMNH NNP 32505 701 11 54963 54963 CD 32505 701 12 - - SYM 32505 701 13 73 73 CD 32505 701 14 ; ; : 32505 701 15 14.5 14.5 CD 32505 701 16 km km NN 32505 701 17 NW NW NNP 32505 701 18 Comayaguela Comayaguela NNP 32505 701 19 , , , 32505 701 20 KU KU NNP 32505 701 21 100499 100499 CD 32505 701 22 ; ; : 32505 701 23 El El NNP 32505 701 24 Zamorano Zamorano NNP 32505 701 25 , , , 32505 701 26 KU KU NNP 32505 701 27 103224 103224 CD 32505 701 28 ; ; : 32505 701 29 29 29 CD 32505 701 30 km km NN 32505 701 31 N N NNP 32505 701 32 Tegucigalpa Tegucigalpa NNP 32505 701 33 , , , 32505 701 34 TNHC TNHC NNP 32505 701 35 32003 32003 CD 32505 701 36 , , , 32505 701 37 32012 32012 CD 32505 701 38 . . . 32505 702 1 Nicaragua Nicaragua NNP 32505 702 2 : : : 32505 702 3 _ _ NNP 32505 702 4 Chinandega Chinandega NNP 32505 702 5 _ _ NNP 32505 702 6 : : : 32505 702 7 Finca Finca NNP 32505 702 8 San San NNP 32505 702 9 Isidro Isidro NNP 32505 702 10 , , , 32505 702 11 10 10 CD 32505 702 12 km km NN 32505 702 13 S S NNP 32505 702 14 Chinandega Chinandega NNP 32505 702 15 , , , 32505 702 16 KU KU NNP 32505 702 17 85311 85311 CD 32505 702 18 - - SYM 32505 702 19 33 33 CD 32505 702 20 . . . 32505 703 1 _ _ NNP 32505 703 2 Managua Managua NNP 32505 703 3 _ _ NNP 32505 703 4 : : : 32505 703 5 13 13 CD 32505 703 6 km km NN 32505 703 7 E E NNP 32505 703 8 Managua Managua NNP 32505 703 9 , , , 32505 703 10 KU KU NNP 32505 703 11 85339 85339 CD 32505 703 12 ; ; : 32505 703 13 2 2 LS 32505 703 14 km km NN 32505 703 15 S S NNP 32505 703 16 Tipitapa Tipitapa NNP 32505 703 17 , , , 32505 703 18 KU KU NNP 32505 703 19 85334 85334 CD 32505 703 20 - - SYM 32505 703 21 8 8 CD 32505 703 22 . . . 32505 704 1 _ _ NNP 32505 704 2 Rivas Rivas NNP 32505 704 3 _ _ NNP 32505 704 4 : : : 32505 704 5 9.5 9.5 CD 32505 704 6 km km NN 32505 704 7 SE SE NNP 32505 704 8 Rivas Rivas NNP 32505 704 9 , , , 32505 704 10 KU KU NNP 32505 704 11 85355 85355 CD 32505 704 12 - - SYM 32505 704 13 6 6 CD 32505 704 14 ; ; : 32505 704 15 18 18 CD 32505 704 16 km km NN 32505 704 17 SE SE NNP 32505 704 18 Rivas Rivas NNP 32505 704 19 , , , 32505 704 20 KU KU NNP 32505 704 21 85354 85354 CD 32505 704 22 ; ; : 32505 704 23 7.7 7.7 CD 32505 704 24 km km NN 32505 704 25 NE NE NNP 32505 704 26 San San NNP 32505 704 27 Juan Juan NNP 32505 704 28 del del NNP 32505 704 29 Sur Sur NNP 32505 704 30 , , , 32505 704 31 KU KU NNP 32505 704 32 85346 85346 CD 32505 704 33 - - SYM 32505 704 34 53 53 CD 32505 704 35 ; ; : 32505 704 36 16.5 16.5 CD 32505 704 37 km km NN 32505 704 38 NE NE NNP 32505 704 39 San San NNP 32505 704 40 Juan Juan NNP 32505 704 41 del del NNP 32505 704 42 Sur Sur NNP 32505 704 43 , , , 32505 704 44 KU KU NNP 32505 704 45 85340 85340 CD 32505 704 46 - - SYM 32505 704 47 5 5 CD 32505 704 48 ; ; : 32505 704 49 5 5 CD 32505 704 50 km km NN 32505 704 51 SE SE NNP 32505 704 52 San San NNP 32505 704 53 Pablo Pablo NNP 32505 704 54 , , , 32505 704 55 KU KU NNP 32505 704 56 43151 43151 CD 32505 704 57 - - SYM 32505 704 58 61 61 CD 32505 704 59 . . . 32505 705 1 _ _ NNP 32505 705 2 Zelaya Zelaya NNP 32505 705 3 _ _ NNP 32505 705 4 : : : 32505 705 5 Isla Isla NNP 32505 705 6 Grande Grande NNP 32505 705 7 del del NNP 32505 705 8 Maiz Maiz NNP 32505 705 9 , , , 32505 705 10 KU KU NNP 32505 705 11 85357 85357 CD 32505 705 12 - - SYM 32505 705 13 60 60 CD 32505 705 14 . . . 32505 706 1 Costa Costa NNP 32505 706 2 Rica Rica NNP 32505 706 3 : : : 32505 706 4 _ _ NNP 32505 706 5 Alajuela Alajuela NNP 32505 706 6 _ _ NNP 32505 706 7 : : : 32505 706 8 _ _ NNP 32505 706 9 Los Los NNP 32505 706 10 Chiles Chiles NNP 32505 706 11 _ _ NNP 32505 706 12 , , , 32505 706 13 USC USC NNP 32505 706 14 7215 7215 CD 32505 706 15 ( ( -LRB- 32505 706 16 2 2 CD 32505 706 17 ) ) -RRB- 32505 706 18 , , , 32505 706 19 7217 7217 CD 32505 706 20 . . . 32505 707 1 _ _ NNP 32505 707 2 Guanacaste Guanacaste NNP 32505 707 3 _ _ NNP 32505 707 4 : : : 32505 707 5 4 4 CD 32505 707 6 km km NN 32505 707 7 W W NNP 32505 707 8 Bagaces Bagaces NNPS 32505 707 9 , , , 32505 707 10 USC USC NNP 32505 707 11 7019 7019 CD 32505 707 12 ( ( -LRB- 32505 707 13 5 5 CD 32505 707 14 ) ) -RRB- 32505 707 15 ; ; : 32505 707 16 Finca Finca NNP 32505 707 17 Taboga Taboga NNP 32505 707 18 , , , 32505 707 19 KU KU NNP 32505 707 20 102265 102265 CD 32505 707 21 - - SYM 32505 707 22 5 5 CD 32505 707 23 ; ; : 32505 707 24 12 12 CD 32505 707 25 km km NN 32505 707 26 S S NNP 32505 707 27 La La NNP 32505 707 28 Cruz Cruz NNP 32505 707 29 , , , 32505 707 30 USC USC NNP 32505 707 31 8091 8091 CD 32505 707 32 ; ; : 32505 707 33 Las Las NNP 32505 707 34 Cañas Cañas NNP 32505 707 35 , , , 32505 707 36 KU KU NNP 32505 707 37 41113 41113 CD 32505 707 38 ( ( -LRB- 32505 707 39 skeleton skeleton NNP 32505 707 40 ) ) -RRB- 32505 707 41 ; ; : 32505 707 42 27 27 CD 32505 707 43 km km NN 32505 707 44 N N NNP 32505 707 45 Las Las NNP 32505 707 46 Cañas Cañas NNP 32505 707 47 , , , 32505 707 48 USC USC NNP 32505 707 49 8171 8171 CD 32505 707 50 ( ( -LRB- 32505 707 51 5 5 CD 32505 707 52 ) ) -RRB- 32505 707 53 ; ; : 32505 707 54 Guardia Guardia NNP 32505 707 55 , , , 32505 707 56 Río Río NNP 32505 707 57 Tempisque Tempisque NNP 32505 707 58 , , , 32505 707 59 USC USC NNP 32505 707 60 8214 8214 CD 32505 707 61 ; ; : 32505 707 62 10 10 CD 32505 707 63 km km NN 32505 707 64 N N NNP 32505 707 65 Guardia Guardia NNP 32505 707 66 , , , 32505 707 67 KU KU NNP 32505 707 68 102266 102266 CD 32505 707 69 - - SYM 32505 707 70 7 7 CD 32505 707 71 ; ; : 32505 707 72 1.6 1.6 CD 32505 707 73 km km NN 32505 707 74 N N NNP 32505 707 75 Guayabo Guayabo NNP 32505 707 76 de de NNP 32505 707 77 Bagaces Bagaces NNP 32505 707 78 , , , 32505 707 79 USC USC NNP 32505 707 80 7023 7023 CD 32505 707 81 ( ( -LRB- 32505 707 82 3 3 CD 32505 707 83 ) ) -RRB- 32505 707 84 ; ; : 32505 707 85 Liberia Liberia NNP 32505 707 86 , , , 32505 707 87 KU KU NNP 32505 707 88 36510 36510 CD 32505 707 89 - - SYM 32505 707 90 22 22 CD 32505 707 91 ; ; : 32505 707 92 4 4 CD 32505 707 93 km km NN 32505 707 94 W W NNP 32505 707 95 Liberia Liberia NNP 32505 707 96 , , , 32505 707 97 KU KU NNP 32505 707 98 36449 36449 CD 32505 707 99 - - SYM 32505 707 100 64 64 CD 32505 707 101 , , , 32505 707 102 USC USC NNP 32505 707 103 102 102 CD 32505 707 104 ( ( -LRB- 32505 707 105 10 10 CD 32505 707 106 ) ) -RRB- 32505 707 107 , , , 32505 707 108 103 103 CD 32505 707 109 ( ( -LRB- 32505 707 110 9 9 CD 32505 707 111 ) ) -RRB- 32505 707 112 , , , 32505 707 113 104 104 CD 32505 707 114 ( ( -LRB- 32505 707 115 7 7 CD 32505 707 116 ) ) -RRB- 32505 707 117 , , , 32505 707 118 105 105 CD 32505 707 119 ; ; : 32505 707 120 6 6 CD 32505 707 121 km km NN 32505 707 122 N N NNP 32505 707 123 Liberia Liberia NNP 32505 707 124 . . . 32505 708 1 USC USC NNP 32505 708 2 8096 8096 CD 32505 708 3 ; ; : 32505 708 4 8 8 CD 32505 708 5 km km NN 32505 708 6 NNW NNW NNP 32505 708 7 Liberia Liberia NNP 32505 708 8 , , , 32505 708 9 KU KU NNP 32505 708 10 65032 65032 CD 32505 708 11 ; ; : 32505 708 12 14.5 14.5 CD 32505 708 13 km km NN 32505 708 14 N N NNP 32505 708 15 Liberia Liberia NNP 32505 708 16 , , , 32505 708 17 USC USC NNP 32505 708 18 8079 8079 CD 32505 708 19 , , , 32505 708 20 8138 8138 CD 32505 708 21 ( ( -LRB- 32505 708 22 2 2 CD 32505 708 23 ) ) -RRB- 32505 708 24 ; ; : 32505 708 25 14.5 14.5 CD 32505 708 26 km km NN 32505 708 27 S S NNP 32505 708 28 Liberia Liberia NNP 32505 708 29 , , , 32505 708 30 USC USC NNP 32505 708 31 8238 8238 CD 32505 708 32 ( ( -LRB- 32505 708 33 5 5 CD 32505 708 34 ) ) -RRB- 32505 708 35 ; ; : 32505 708 36 6 6 CD 32505 708 37 km km NN 32505 708 38 N N NNP 32505 708 39 Nicoya Nicoya NNP 32505 708 40 , , , 32505 708 41 USC USC NNP 32505 708 42 8229 8229 CD 32505 708 43 ( ( -LRB- 32505 708 44 11 11 CD 32505 708 45 ) ) -RRB- 32505 708 46 ; ; : 32505 708 47 4 4 LS 32505 708 48 km km NN 32505 708 49 S S NNP 32505 708 50 Nicoya Nicoya NNP 32505 708 51 , , , 32505 708 52 USC USC NNP 32505 708 53 8230 8230 CD 32505 708 54 , , , 32505 708 55 8231 8231 CD 32505 708 56 ; ; : 32505 708 57 Peñas Peñas NNP 32505 708 58 Blancas Blancas NNP 32505 708 59 , , , 32505 708 60 KU KU NNP 32505 708 61 102263 102263 CD 32505 708 62 ; ; : 32505 708 63 8.6 8.6 CD 32505 708 64 km km NN 32505 708 65 ESE ESE NNP 32505 708 66 Playa Playa NNP 32505 708 67 del del NNP 32505 708 68 Coco Coco NNP 32505 708 69 , , , 32505 708 70 USC USC NNP 32505 708 71 8137 8137 CD 32505 708 72 ( ( -LRB- 32505 708 73 14 14 CD 32505 708 74 ) ) -RRB- 32505 708 75 ; ; : 32505 708 76 21 21 CD 32505 708 77 km km NN 32505 708 78 E E NNP 32505 708 79 Playa Playa NNP 32505 708 80 del del NNP 32505 708 81 Coco Coco NNP 32505 708 82 , , , 32505 708 83 USC USC NNP 32505 708 84 8138 8138 CD 32505 708 85 ( ( -LRB- 32505 708 86 2 2 CD 32505 708 87 ) ) -RRB- 32505 708 88 ; ; : 32505 708 89 Santa Santa NNP 32505 708 90 Cruz Cruz NNP 32505 708 91 , , , 32505 708 92 USC USC NNP 32505 708 93 8232 8232 CD 32505 708 94 ( ( -LRB- 32505 708 95 2 2 CD 32505 708 96 ) ) -RRB- 32505 708 97 ; ; : 32505 708 98 3 3 LS 32505 708 99 km km NN 32505 708 100 E E NNP 32505 708 101 Santa Santa NNP 32505 708 102 Rosa Rosa NNP 32505 708 103 , , , 32505 708 104 TCWC TCWC NNP 32505 708 105 16663 16663 CD 32505 708 106 - - SYM 32505 708 107 68 68 CD 32505 708 108 ; ; : 32505 708 109 Tenorio Tenorio NNP 32505 708 110 , , , 32505 708 111 KU KU NNP 32505 708 112 32159 32159 CD 32505 708 113 ; ; : 32505 708 114 Tilarán Tilarán NNP 32505 708 115 , , , 32505 708 116 KU KU NNP 32505 708 117 36509 36509 CD 32505 708 118 . . . 32505 709 1 _ _ NNP 32505 709 2 Puntarenas Puntarenas NNP 32505 709 3 _ _ NNP 32505 709 4 : : : 32505 709 5 10 10 CD 32505 709 6 km km NN 32505 709 7 WNW WNW NNP 32505 709 8 Esparta Esparta NNP 32505 709 9 , , , 32505 709 10 KU KU NNP 32505 709 11 65022 65022 CD 32505 709 12 - - SYM 32505 709 13 9 9 CD 32505 709 14 , , , 32505 709 15 68614 68614 CD 32505 709 16 ( ( -LRB- 32505 709 17 skeleton skeleton NNP 32505 709 18 ) ) -RRB- 32505 709 19 ; ; : 32505 709 20 4.5 4.5 CD 32505 709 21 km km NN 32505 709 22 WNW WNW NNP 32505 709 23 Esparta Esparta NNP 32505 709 24 , , , 32505 709 25 KU KU NNP 32505 709 26 65030 65030 CD 32505 709 27 ; ; : 32505 709 28 12 12 CD 32505 709 29 km km NN 32505 709 30 WNW WNW NNP 32505 709 31 Esparta Esparta NNP 32505 709 32 , , , 32505 709 33 KU KU NNP 32505 709 34 65031 65031 CD 32505 709 35 ; ; : 32505 709 36 6 6 CD 32505 709 37 km km NN 32505 709 38 E E NNP 32505 709 39 Esparta Esparta NNP 32505 709 40 , , , 32505 709 41 KU KU NNP 32505 709 42 86477 86477 CD 32505 709 43 ; ; : 32505 709 44 Hotel Hotel NNP 32505 709 45 Maribella Maribella NNP 32505 709 46 , , , 32505 709 47 KU KU NNP 32505 709 48 32157 32157 CD 32505 709 49 - - SYM 32505 709 50 8 8 CD 32505 709 51 ; ; : 32505 709 52 3 3 LS 32505 709 53 km km NN 32505 709 54 W W NNP 32505 709 55 Puntarenas Puntarenas NNP 32505 709 56 , , , 32505 709 57 TCWC TCWC NNP 32505 709 58 16657 16657 CD 32505 709 59 - - SYM 32505 709 60 62 62 CD 32505 709 61 . . . 32505 710 1 _ _ NNP 32505 710 2 Hyla Hyla NNP 32505 710 3 staufferi staufferi NN 32505 710 4 altae altae NN 32505 710 5 _ _ NNP 32505 710 6 Dunn Dunn NNP 32505 710 7 , , , 32505 710 8 New New NNP 32505 710 9 Combination Combination NNP 32505 710 10 _ _ NNP 32505 710 11 Hyla Hyla NNP 32505 710 12 altae altae NN 32505 710 13 _ _ NNP 32505 710 14 Dunn Dunn NNP 32505 710 15 , , , 32505 710 16 Occas Occas NNP 32505 710 17 . . . 32505 711 1 Papers Papers NNP 32505 711 2 Boston Boston NNP 32505 711 3 Soc Soc NNP 32505 711 4 . . . 32505 712 1 Nat Nat NNP 32505 712 2 . . . 32505 713 1 Hist Hist NNP 32505 713 2 . . . 32505 713 3 , , , 32505 713 4 8:61 8:61 CD 32505 713 5 , , , 32505 713 6 June June NNP 32505 713 7 7 7 CD 32505 713 8 , , , 32505 713 9 1933 1933 CD 32505 713 10 [ [ -LRB- 32505 713 11 Holotype.--MCZ Holotype.--MCZ NNP 32505 713 12 17972 17972 CD 32505 713 13 , , , 32505 713 14 Summit Summit NNP 32505 713 15 , , , 32505 713 16 Canal Canal NNP 32505 713 17 Zone Zone NNP 32505 713 18 , , , 32505 713 19 Panamá Panamá NNP 32505 713 20 ; ; : 32505 713 21 Emmett Emmett NNP 32505 713 22 R. R. NNP 32505 713 23 Dunn Dunn NNP 32505 713 24 collector collector NN 32505 713 25 ] ] -RRB- 32505 713 26 . . . 32505 714 1 _ _ NNP 32505 714 2 Hyla Hyla NNP 32505 714 3 culex culex NN 32505 714 4 _ _ NNP 32505 714 5 : : : 32505 714 6 Stuart Stuart NNP 32505 714 7 , , , 32505 714 8 Misc Misc NNP 32505 714 9 . . . 32505 715 1 Publ Publ NNS 32505 715 2 . . . 32505 716 1 Univ Univ NNP 32505 716 2 . . . 32505 717 1 Michigan Michigan NNP 32505 717 2 Mus Mus NNP 32505 717 3 . . . 32505 718 1 Zool Zool NNP 32505 718 2 . . . 32505 718 3 , , , 32505 718 4 29:38 29:38 CD 32505 718 5 , , , 32505 718 6 October October NNP 32505 718 7 1 1 CD 32505 718 8 , , , 32505 718 9 1935 1935 CD 32505 718 10 . . . 32505 719 1 Gaige Gaige NNP 32505 719 2 , , , 32505 719 3 Carnegie Carnegie NNP 32505 719 4 Inst Inst NNP 32505 719 5 . . . 32505 720 1 Washington Washington NNP 32505 720 2 Publ Publ NNP 32505 720 3 . . . 32505 720 4 , , , 32505 720 5 457:293 457:293 CD 32505 720 6 , , , 32505 720 7 1936 1936 CD 32505 720 8 . . . 32505 721 1 _ _ NNP 32505 721 2 Hyla Hyla NNP 32505 721 3 staufferi staufferi NN 32505 721 4 _ _ NNP 32505 721 5 : : : 32505 721 6 Taylor Taylor NNP 32505 721 7 , , , 32505 721 8 Univ Univ NNP 32505 721 9 . . . 32505 722 1 Kansas Kansas NNP 32505 722 2 Sci Sci NNP 32505 722 3 . . . 32505 723 1 Bull Bull NNP 32505 723 2 . . NNP 32505 723 3 , , , 32505 723 4 35:862 35:862 CD 32505 723 5 , , , 32505 723 6 July July NNP 32505 723 7 1 1 CD 32505 723 8 , , , 32505 723 9 1952 1952 CD 32505 723 10 . . . 32505 724 1 Duellman Duellman NNP 32505 724 2 , , , 32505 724 3 Univ Univ NNP 32505 724 4 . . . 32505 725 1 Kansas Kansas NNP 32505 725 2 Publ Publ NNP 32505 725 3 . . . 32505 725 4 , , , 32505 725 5 Mus mus NN 32505 725 6 . . . 32505 726 1 Nat Nat NNP 32505 726 2 . . . 32505 727 1 Hist Hist NNP 32505 727 2 . . NNP 32505 727 3 , , , 32505 727 4 17:274 17:274 CD 32505 727 5 , , , 32505 727 6 June June NNP 32505 727 7 17 17 CD 32505 727 8 , , , 32505 727 9 1966 1966 CD 32505 727 10 . . . 32505 728 1 _ _ NNP 32505 728 2 Diagnosis._--Small Diagnosis._--Small NNP 32505 728 3 frogs frog NNS 32505 728 4 ( ( -LRB- 32505 728 5 Male male NN 32505 728 6 to to IN 32505 728 7 26 26 CD 32505 728 8 mm mm NN 32505 728 9 . . NNP 32505 728 10 , , , 32505 728 11 Female female JJ 32505 728 12 to to IN 32505 728 13 27 27 CD 32505 728 14 mm mm NNS 32505 728 15 . . . 32505 729 1 ) ) -RRB- 32505 729 2 ; ; : 32505 729 3 dorsolateral dorsolateral JJ 32505 729 4 and and CC 32505 729 5 paravertebral paravertebral JJ 32505 729 6 stripes stripe NNS 32505 729 7 complete complete JJ 32505 729 8 ; ; : 32505 729 9 longitudinal longitudinal JJ 32505 729 10 dark dark JJ 32505 729 11 gray gray JJ 32505 729 12 stripe stripe NN 32505 729 13 on on IN 32505 729 14 shank shank JJ 32505 729 15 ; ; : 32505 729 16 thighs thigh NNS 32505 729 17 unmarked unmarked JJ 32505 729 18 ; ; : 32505 729 19 interorbital interorbital NNP 32505 729 20 bar bar NN 32505 729 21 usually usually RB 32505 729 22 absent absent JJ 32505 729 23 ; ; : 32505 729 24 toes toe VBZ 32505 729 25 about about RB 32505 729 26 three three CD 32505 729 27 fifths fifth NNS 32505 729 28 webbed web VBN 32505 729 29 ; ; : 32505 729 30 gray gray JJ 32505 729 31 to to IN 32505 729 32 brownish brownish NNP 32505 729 33 gray gray NNP 32505 729 34 above above RB 32505 729 35 . . . 32505 730 1 _ _ NNP 32505 730 2 Variation._--_Hyla Variation._--_Hyla NNP 32505 730 3 staufferi staufferi NNP 32505 730 4 altae altae NNP 32505 730 5 _ _ NNP 32505 730 6 is be VBZ 32505 730 7 less less RBR 32505 730 8 variable variable JJ 32505 730 9 in in IN 32505 730 10 size size NN 32505 730 11 , , , 32505 730 12 proportions proportion NNS 32505 730 13 , , , 32505 730 14 and and CC 32505 730 15 color color NN 32505 730 16 pattern pattern NN 32505 730 17 than than IN 32505 730 18 is be VBZ 32505 730 19 _ _ NNP 32505 730 20 H. H. NNP 32505 731 1 s. s. NNP 32505 731 2 staufferi staufferi NNP 32505 731 3 _ _ NNP 32505 731 4 . . . 32505 732 1 The the DT 32505 732 2 size size NN 32505 732 3 varies vary VBZ 32505 732 4 from from IN 32505 732 5 21.7 21.7 CD 32505 732 6 to to IN 32505 732 7 26 26 CD 32505 732 8 mm mm NNS 32505 732 9 . . . 32505 733 1 ( ( -LRB- 32505 733 2 23.6 23.6 CD 32505 733 3 ) ) -RRB- 32505 733 4 in in IN 32505 733 5 72 72 CD 32505 733 6 males male NNS 32505 733 7 . . . 32505 734 1 The the DT 32505 734 2 ratio ratio NN 32505 734 3 of of IN 32505 734 4 tibia tibia NNP 32505 734 5 to to TO 32505 734 6 snout snout VB 32505 734 7 - - HYPH 32505 734 8 vent vent NN 32505 734 9 length length NN 32505 734 10 is be VBZ 32505 734 11 0.42 0.42 CD 32505 734 12 to to IN 32505 734 13 0.50 0.50 CD 32505 734 14 ( ( -LRB- 32505 734 15 0.45 0.45 CD 32505 734 16 ) ) -RRB- 32505 734 17 , , , 32505 734 18 slightly slightly RB 32505 734 19 less less JJR 32505 734 20 than than IN 32505 734 21 in in IN 32505 734 22 the the DT 32505 734 23 northern northern JJ 32505 734 24 subspecies subspecie NNS 32505 734 25 . . . 32505 735 1 In in IN 32505 735 2 color color NN 32505 735 3 pattern pattern NN 32505 735 4 94.5 94.5 CD 32505 735 5 per per IN 32505 735 6 cent cent NN 32505 735 7 of of IN 32505 735 8 the the DT 32505 735 9 individuals individual NNS 32505 735 10 have have VBP 32505 735 11 complete complete JJ 32505 735 12 dorsal dorsal NN 32505 735 13 stripes stripe NNS 32505 735 14 , , , 32505 735 15 and and CC 32505 735 16 all all DT 32505 735 17 have have VBP 32505 735 18 a a DT 32505 735 19 longitudinal longitudinal JJ 32505 735 20 stripe stripe NN 32505 735 21 on on IN 32505 735 22 the the DT 32505 735 23 shank shank JJ 32505 735 24 ( ( -LRB- 32505 735 25 Table table NN 32505 735 26 7 7 CD 32505 735 27 ) ) -RRB- 32505 735 28 . . . 32505 736 1 _ _ NNP 32505 736 2 Distribution._--This distribution._--this DT 32505 736 3 subspecies subspecie NNS 32505 736 4 is be VBZ 32505 736 5 restricted restrict VBN 32505 736 6 to to TO 32505 736 7 subhumid subhumid NN 32505 736 8 forests forest NNS 32505 736 9 and and CC 32505 736 10 savannas savanna NNS 32505 736 11 on on IN 32505 736 12 the the DT 32505 736 13 Pacific Pacific NNP 32505 736 14 lowlands lowland NNS 32505 736 15 of of IN 32505 736 16 Panamá Panamá NNP 32505 736 17 . . . 32505 737 1 _ _ NNP 32505 737 2 Hyla Hyla NNP 32505 737 3 s. s. NN 32505 737 4 altae altae NNP 32505 737 5 _ _ NNP 32505 737 6 is be VBZ 32505 737 7 presently presently RB 32505 737 8 known know VBN 32505 737 9 to to TO 32505 737 10 occur occur VB 32505 737 11 from from IN 32505 737 12 Chepo Chepo NNP 32505 737 13 in in IN 32505 737 14 east east JJ 32505 737 15 - - HYPH 32505 737 16 central central JJ 32505 737 17 Panamá Panamá NNP 32505 737 18 through through IN 32505 737 19 the the DT 32505 737 20 Azuero Azuero NNP 32505 737 21 Peninsula Peninsula NNP 32505 737 22 to to IN 32505 737 23 Concepción Concepción NNP 32505 737 24 , , , 32505 737 25 Chiriquí Chiriquí NNP 32505 737 26 , , , 32505 737 27 in in IN 32505 737 28 western western JJ 32505 737 29 Panamá Panamá NNP 32505 737 30 ( ( -LRB- 32505 737 31 Fig Fig NNP 32505 737 32 . . . 32505 738 1 7 7 LS 32505 738 2 ) ) -RRB- 32505 738 3 . . . 32505 739 1 _ _ NNP 32505 739 2 Specimens Specimens NNP 32505 739 3 Examined._--Panamá Examined._--Panamá NNPS 32505 739 4 : : : 32505 739 5 _ _ NNP 32505 739 6 Canal Canal NNP 32505 739 7 Zone Zone NNP 32505 739 8 _ _ NNP 32505 739 9 : : : 32505 739 10 No no DT 32505 739 11 specific specific JJ 32505 739 12 locality locality NN 32505 739 13 , , , 32505 739 14 TNHC TNHC NNP 32505 739 15 24406 24406 CD 32505 739 16 ; ; : 32505 739 17 2.8 2.8 CD 32505 739 18 km km NN 32505 739 19 SW SW NNP 32505 739 20 Fort Fort NNP 32505 739 21 Kobbe Kobbe NNP 32505 739 22 , , , 32505 739 23 KU KU NNP 32505 739 24 101679 101679 CD 32505 739 25 . . . 32505 740 1 _ _ NNP 32505 740 2 Chiriquí Chiriquí NNP 32505 740 3 _ _ NNP 32505 740 4 : : : 32505 740 5 14.4 14.4 CD 32505 740 6 km km NN 32505 740 7 E e NN 32505 740 8 Concepción Concepción NNP 32505 740 9 , , , 32505 740 10 AMNH AMNH NNP 32505 740 11 69799 69799 CD 32505 740 12 - - SYM 32505 740 13 801 801 CD 32505 740 14 ; ; : 32505 740 15 6.6 6.6 CD 32505 740 16 km km NN 32505 740 17 N N NNP 32505 740 18 David David NNP 32505 740 19 , , , 32505 740 20 TNHC TNHC NNP 32505 740 21 32013 32013 CD 32505 740 22 - - SYM 32505 740 23 4 4 CD 32505 740 24 ; ; : 32505 740 25 2 2 CD 32505 740 26 km km NN 32505 740 27 S S NNP 32505 740 28 David David NNP 32505 740 29 , , , 32505 740 30 AMNH AMNH NNP 32505 740 31 68802 68802 CD 32505 740 32 . . . 32505 741 1 _ _ NNP 32505 741 2 Coclé Coclé NNP 32505 741 3 _ _ NNP 32505 741 4 : : : 32505 741 5 1 1 CD 32505 741 6 km km NN 32505 741 7 NE NE NNP 32505 741 8 El El NNP 32505 741 9 Caño Caño NNP 32505 741 10 , , , 32505 741 11 KU KU NNP 32505 741 12 101662 101662 CD 32505 741 13 - - SYM 32505 741 14 75 75 CD 32505 741 15 ; ; : 32505 741 16 El El NNP 32505 741 17 Valle Valle NNP 32505 741 18 de de NNP 32505 741 19 Antón Antón NNP 32505 741 20 , , , 32505 741 21 AMNH AMNH NNP 32505 741 22 59601 59601 CD 32505 741 23 - - SYM 32505 741 24 5 5 CD 32505 741 25 , , , 32505 741 26 KU KU NNP 32505 741 27 77333 77333 CD 32505 741 28 - - SYM 32505 741 29 47 47 CD 32505 741 30 ; ; : 32505 741 31 7 7 CD 32505 741 32 km km NN 32505 741 33 SSW SSW NNP 32505 741 34 Penonomé Penonomé NNP 32505 741 35 , , , 32505 741 36 KU KU NNP 32505 741 37 101654 101654 CD 32505 741 38 - - SYM 32505 741 39 61 61 CD 32505 741 40 . . . 32505 742 1 _ _ NNP 32505 742 2 Los Los NNP 32505 742 3 Santos Santos NNP 32505 742 4 _ _ NNP 32505 742 5 : : : 32505 742 6 Tonosí Tonosí NNP 32505 742 7 , , , 32505 742 8 KU KU NNP 32505 742 9 101246 101246 CD 32505 742 10 ( ( -LRB- 32505 742 11 tadpoles tadpole NNS 32505 742 12 ) ) -RRB- 32505 742 13 , , , 32505 742 14 101697 101697 CD 32505 742 15 - - SYM 32505 742 16 701 701 CD 32505 742 17 . . . 32505 743 1 _ _ NNP 32505 743 2 Panamá Panamá NNP 32505 743 3 _ _ NNP 32505 743 4 : : : 32505 743 5 2 2 CD 32505 743 6 km km NN 32505 743 7 WSW WSW NNP 32505 743 8 Chepo Chepo NNP 32505 743 9 , , , 32505 743 10 KU KU NNP 32505 743 11 101680 101680 CD 32505 743 12 - - SYM 32505 743 13 8 8 CD 32505 743 14 ; ; : 32505 743 15 6 6 CD 32505 743 16 km km NN 32505 743 17 WSW WSW NNP 32505 743 18 Chepo Chepo NNP 32505 743 19 , , , 32505 743 20 KU KU NNP 32505 743 21 77324 77324 CD 32505 743 22 - - SYM 32505 743 23 27 27 CD 32505 743 24 ; ; : 32505 743 25 El El NNP 32505 743 26 Cangrejo Cangrejo NNP 32505 743 27 ( ( -LRB- 32505 743 28 Panamá Panamá NNP 32505 743 29 ) ) -RRB- 32505 743 30 , , , 32505 743 31 KU KU NNP 32505 743 32 101676 101676 CD 32505 743 33 - - SYM 32505 743 34 8 8 CD 32505 743 35 ; ; : 32505 743 36 Nueva Nueva NNP 32505 743 37 Gorgona Gorgona NNP 32505 743 38 , , , 32505 743 39 AMNH AMNH NNP 32505 743 40 69991 69991 CD 32505 743 41 , , , 32505 743 42 69798 69798 CD 32505 743 43 ; ; : 32505 743 44 1.5 1.5 CD 32505 743 45 km km NN 32505 743 46 W W NNP 32505 743 47 Pacora Pacora NNP 32505 743 48 , , , 32505 743 49 KU KU NNP 32505 743 50 77328 77328 CD 32505 743 51 - - SYM 32505 743 52 32 32 CD 32505 743 53 ; ; : 32505 743 54 2 2 CD 32505 743 55 km km NN 32505 743 56 N N NNP 32505 743 57 Tocumen Tocumen NNP 32505 743 58 , , , 32505 743 59 KU KU NNP 32505 743 60 101689 101689 CD 32505 743 61 - - SYM 32505 743 62 95 95 CD 32505 743 63 ; ; : 32505 743 64 8 8 CD 32505 743 65 km km NN 32505 743 66 NE NE NNP 32505 743 67 Tocumen Tocumen NNP 32505 743 68 , , , 32505 743 69 KU KU NNP 32505 743 70 101696 101696 CD 32505 743 71 . . . 32505 744 1 EVOLUTIONARY evolutionary NN 32505 744 2 HISTORY HISTORY NNP 32505 744 3 My -PRON- PRP$ 32505 744 4 assumptions assumption NNS 32505 744 5 regarding regard VBG 32505 744 6 the the DT 32505 744 7 evolutionary evolutionary JJ 32505 744 8 history history NN 32505 744 9 of of IN 32505 744 10 the the DT 32505 744 11 _ _ NNP 32505 744 12 Hyla Hyla NNP 32505 744 13 rubra rubra NN 32505 744 14 _ _ NNP 32505 744 15 group group NN 32505 744 16 in in IN 32505 744 17 Central Central NNP 32505 744 18 America America NNP 32505 744 19 were be VBD 32505 744 20 derived derive VBN 32505 744 21 partly partly RB 32505 744 22 from from IN 32505 744 23 interpretations interpretation NNS 32505 744 24 of of IN 32505 744 25 the the DT 32505 744 26 evolutionary evolutionary JJ 32505 744 27 history history NN 32505 744 28 of of IN 32505 744 29 other other JJ 32505 744 30 animal animal NN 32505 744 31 groups group NNS 32505 744 32 ( ( -LRB- 32505 744 33 Simpson Simpson NNP 32505 744 34 , , , 32505 744 35 1943 1943 CD 32505 744 36 , , , 32505 744 37 1965 1965 CD 32505 744 38 ; ; : 32505 744 39 Dunn Dunn NNP 32505 744 40 , , , 32505 744 41 1931b 1931b CD 32505 744 42 ; ; : 32505 744 43 Stuart Stuart NNP 32505 744 44 , , , 32505 744 45 1950 1950 CD 32505 744 46 ; ; : 32505 744 47 Duellman Duellman NNP 32505 744 48 , , , 32505 744 49 1958 1958 CD 32505 744 50 , , , 32505 744 51 1960 1960 CD 32505 744 52 , , , 32505 744 53 1963 1963 CD 32505 744 54 , , , 32505 744 55 1965 1965 CD 32505 744 56 ; ; : 32505 744 57 and and CC 32505 744 58 Duellman Duellman NNP 32505 744 59 and and CC 32505 744 60 Trueb Trueb NNP 32505 744 61 , , , 32505 744 62 1966 1966 CD 32505 744 63 ) ) -RRB- 32505 744 64 . . . 32505 745 1 The the DT 32505 745 2 origin origin NN 32505 745 3 and and CC 32505 745 4 early early JJ 32505 745 5 evolution evolution NN 32505 745 6 of of IN 32505 745 7 the the DT 32505 745 8 group group NN 32505 745 9 probably probably RB 32505 745 10 occurred occur VBD 32505 745 11 prior prior RB 32505 745 12 to to IN 32505 745 13 the the DT 32505 745 14 Mid Mid NNP 32505 745 15 - - HYPH 32505 745 16 Pliocene Pliocene NNP 32505 745 17 in in IN 32505 745 18 the the DT 32505 745 19 lowlands lowland NNS 32505 745 20 of of IN 32505 745 21 South South NNP 32505 745 22 America America NNP 32505 745 23 , , , 32505 745 24 because because IN 32505 745 25 the the DT 32505 745 26 greatest great JJS 32505 745 27 diversity diversity NN 32505 745 28 of of IN 32505 745 29 the the DT 32505 745 30 group group NN 32505 745 31 is be VBZ 32505 745 32 in in IN 32505 745 33 Brazil Brazil NNP 32505 745 34 . . . 32505 746 1 Differentiation differentiation NN 32505 746 2 into into IN 32505 746 3 two two CD 32505 746 4 or or CC 32505 746 5 more more JJR 32505 746 6 subgroups subgroup NNS 32505 746 7 took take VBD 32505 746 8 place place NN 32505 746 9 in in IN 32505 746 10 South South NNP 32505 746 11 America America NNP 32505 746 12 prior prior RB 32505 746 13 to to IN 32505 746 14 the the DT 32505 746 15 late late JJ 32505 746 16 Pliocene Pliocene NNP 32505 746 17 . . . 32505 747 1 At at IN 32505 747 2 the the DT 32505 747 3 end end NN 32505 747 4 of of IN 32505 747 5 the the DT 32505 747 6 Pliocene Pliocene NNP 32505 747 7 , , , 32505 747 8 shortly shortly RB 32505 747 9 after after IN 32505 747 10 the the DT 32505 747 11 closure closure NN 32505 747 12 of of IN 32505 747 13 the the DT 32505 747 14 Colombian Colombian NNP 32505 747 15 Portal Portal NNP 32505 747 16 , , , 32505 747 17 many many JJ 32505 747 18 South south JJ 32505 747 19 American american JJ 32505 747 20 animals animal NNS 32505 747 21 migrated migrate VBN 32505 747 22 into into IN 32505 747 23 Central Central NNP 32505 747 24 America America NNP 32505 747 25 ( ( -LRB- 32505 747 26 Simpson Simpson NNP 32505 747 27 , , , 32505 747 28 1943 1943 CD 32505 747 29 , , , 32505 747 30 Maldonado Maldonado NNP 32505 747 31 - - HYPH 32505 747 32 Koerdell Koerdell NNP 32505 747 33 , , , 32505 747 34 1964 1964 CD 32505 747 35 , , , 32505 747 36 and and CC 32505 747 37 Savage Savage NNP 32505 747 38 , , , 32505 747 39 1966 1966 CD 32505 747 40 ) ) -RRB- 32505 747 41 . . . 32505 748 1 It -PRON- PRP 32505 748 2 is be VBZ 32505 748 3 likely likely JJ 32505 748 4 that that IN 32505 748 5 the the DT 32505 748 6 _ _ NNP 32505 748 7 Hyla Hyla NNP 32505 748 8 rubra rubra NN 32505 748 9 _ _ NNP 32505 748 10 group group NN 32505 748 11 entered enter VBD 32505 748 12 Central Central NNP 32505 748 13 America America NNP 32505 748 14 at at IN 32505 748 15 that that DT 32505 748 16 time time NN 32505 748 17 ; ; : 32505 748 18 apparently apparently RB 32505 748 19 two two CD 32505 748 20 stocks stock NNS 32505 748 21 ( ( -LRB- 32505 748 22 _ _ NNP 32505 748 23 rubra rubra NNP 32505 748 24 - - HYPH 32505 748 25 elaeochroa elaeochroa NNP 32505 748 26 - - HYPH 32505 748 27 staufferi staufferi NNP 32505 748 28 _ _ NNP 32505 748 29 stock stock NN 32505 748 30 and and CC 32505 748 31 _ _ NNP 32505 748 32 boulengeri boulengeri NN 32505 748 33 - - HYPH 32505 748 34 foliamorta foliamorta NNP 32505 748 35 _ _ NNP 32505 748 36 stock stock NN 32505 748 37 ) ) -RRB- 32505 748 38 migrated migrate VBN 32505 748 39 into into IN 32505 748 40 Central Central NNP 32505 748 41 America America NNP 32505 748 42 . . . 32505 749 1 _ _ NNP 32505 749 2 Hyla Hyla NNP 32505 749 3 elaeochroa elaeochroa NN 32505 749 4 _ _ NNP 32505 749 5 is be VBZ 32505 749 6 closely closely RB 32505 749 7 related relate VBN 32505 749 8 to to IN 32505 749 9 _ _ NNP 32505 749 10 rubra rubra NNP 32505 749 11 _ _ NNP 32505 749 12 and and CC 32505 749 13 probably probably RB 32505 749 14 differentiated differentiate VBD 32505 749 15 from from IN 32505 749 16 _ _ NNP 32505 749 17 rubra rubra NNP 32505 749 18 _ _ NNP 32505 749 19 through through IN 32505 749 20 spatial spatial JJ 32505 749 21 isolation isolation NN 32505 749 22 . . . 32505 750 1 Thus thus RB 32505 750 2 , , , 32505 750 3 we -PRON- PRP 32505 750 4 have have VBP 32505 750 5 _ _ NNP 32505 750 6 elaeochroa elaeochroa NNP 32505 750 7 _ _ NNP 32505 750 8 in in IN 32505 750 9 Central Central NNP 32505 750 10 America America NNP 32505 750 11 and and CC 32505 750 12 _ _ NNP 32505 750 13 rubra rubra NNP 32505 750 14 _ _ NNP 32505 750 15 in in IN 32505 750 16 South South NNP 32505 750 17 America America NNP 32505 750 18 ; ; : 32505 750 19 most most RBS 32505 750 20 likely likely RB 32505 750 21 only only RB 32505 750 22 in in IN 32505 750 23 relatively relatively RB 32505 750 24 recent recent JJ 32505 750 25 times time NNS 32505 750 26 has have VBZ 32505 750 27 _ _ NNP 32505 750 28 rubra rubra NNP 32505 750 29 _ _ NNP 32505 750 30 migrated migrate VBN 32505 750 31 into into IN 32505 750 32 eastern eastern JJ 32505 750 33 Panamá Panamá NNP 32505 750 34 from from IN 32505 750 35 northern northern JJ 32505 750 36 South South NNP 32505 750 37 America America NNP 32505 750 38 . . . 32505 751 1 The the DT 32505 751 2 differentiation differentiation NN 32505 751 3 and and CC 32505 751 4 dispersal dispersal NN 32505 751 5 of of IN 32505 751 6 _ _ NNP 32505 751 7 elaeochroa elaeochroa NNP 32505 751 8 _ _ NNP 32505 751 9 and and CC 32505 751 10 _ _ NNP 32505 751 11 staufferi staufferi NN 32505 751 12 _ _ NNP 32505 751 13 took take VBD 32505 751 14 place place NN 32505 751 15 in in IN 32505 751 16 Central Central NNP 32505 751 17 America America NNP 32505 751 18 after after IN 32505 751 19 the the DT 32505 751 20 Pliocene Pliocene NNP 32505 751 21 . . . 32505 752 1 Probably probably RB 32505 752 2 the the DT 32505 752 3 events event NNS 32505 752 4 of of IN 32505 752 5 the the DT 32505 752 6 Pleistocene Pleistocene NNP 32505 752 7 resulted result VBD 32505 752 8 in in IN 32505 752 9 the the DT 32505 752 10 isolation isolation NN 32505 752 11 of of IN 32505 752 12 populations population NNS 32505 752 13 . . . 32505 753 1 One one CD 32505 753 2 of of IN 32505 753 3 these these DT 32505 753 4 ( ( -LRB- 32505 753 5 _ _ NNP 32505 753 6 Hyla Hyla NNP 32505 753 7 staufferi staufferi NN 32505 753 8 _ _ NNP 32505 753 9 stock stock NN 32505 753 10 ) ) -RRB- 32505 753 11 was be VBD 32505 753 12 restricted restrict VBN 32505 753 13 in in IN 32505 753 14 the the DT 32505 753 15 subhumid subhumid JJ 32505 753 16 Pacific Pacific NNP 32505 753 17 lowlands lowland NNS 32505 753 18 , , , 32505 753 19 whereas whereas IN 32505 753 20 the the DT 32505 753 21 _ _ NNP 32505 753 22 Hyla Hyla NNP 32505 753 23 elaeochroa elaeochroa NN 32505 753 24 _ _ NNP 32505 753 25 stock stock NN 32505 753 26 occupied occupy VBD 32505 753 27 the the DT 32505 753 28 tropical tropical JJ 32505 753 29 wet wet JJ 32505 753 30 forests forest NNS 32505 753 31 of of IN 32505 753 32 the the DT 32505 753 33 Caribbean caribbean JJ 32505 753 34 lowlands lowland NNS 32505 753 35 . . . 32505 754 1 _ _ NNP 32505 754 2 Hyla Hyla NNP 32505 754 3 elaeochroa elaeochroa NN 32505 754 4 _ _ NNP 32505 754 5 apparently apparently RB 32505 754 6 more more RBR 32505 754 7 closely closely RB 32505 754 8 resembled resemble VBN 32505 754 9 the the DT 32505 754 10 parental parental JJ 32505 754 11 stock stock NN 32505 754 12 by by IN 32505 754 13 being be VBG 32505 754 14 restricted restrict VBN 32505 754 15 to to IN 32505 754 16 the the DT 32505 754 17 tropical tropical JJ 32505 754 18 rain rain NN 32505 754 19 forests forest NNS 32505 754 20 , , , 32505 754 21 whereas whereas IN 32505 754 22 _ _ NNP 32505 754 23 staufferi staufferi NN 32505 754 24 _ _ NNP 32505 754 25 adapted adapt VBD 32505 754 26 to to IN 32505 754 27 subhumid subhumid NN 32505 754 28 environments environment NNS 32505 754 29 and and CC 32505 754 30 thereby thereby RB 32505 754 31 was be VBD 32505 754 32 able able JJ 32505 754 33 to to TO 32505 754 34 disperse disperse VB 32505 754 35 throughout throughout IN 32505 754 36 most most JJS 32505 754 37 of of IN 32505 754 38 the the DT 32505 754 39 subhumid subhumid NN 32505 754 40 regions region NNS 32505 754 41 of of IN 32505 754 42 Central Central NNP 32505 754 43 America America NNP 32505 754 44 . . . 32505 755 1 After after IN 32505 755 2 geographical geographical JJ 32505 755 3 separation separation NN 32505 755 4 took take VBD 32505 755 5 place place NN 32505 755 6 the the DT 32505 755 7 initial initial JJ 32505 755 8 genetic genetic JJ 32505 755 9 divergence divergence NN 32505 755 10 between between IN 32505 755 11 the the DT 32505 755 12 two two CD 32505 755 13 populations population NNS 32505 755 14 was be VBD 32505 755 15 maintained maintain VBN 32505 755 16 by by IN 32505 755 17 means mean NNS 32505 755 18 of of IN 32505 755 19 ecological ecological JJ 32505 755 20 and and CC 32505 755 21 ethological ethological JJ 32505 755 22 isolating isolating NN 32505 755 23 mechanisms mechanism NNS 32505 755 24 . . . 32505 756 1 Under under IN 32505 756 2 these these DT 32505 756 3 circumstances circumstance NNS 32505 756 4 it -PRON- PRP 32505 756 5 can can MD 32505 756 6 be be VB 32505 756 7 supposed suppose VBN 32505 756 8 that that IN 32505 756 9 the the DT 32505 756 10 different different JJ 32505 756 11 ecological ecological JJ 32505 756 12 preferences preference NNS 32505 756 13 of of IN 32505 756 14 _ _ NNP 32505 756 15 elaeochroa elaeochroa NNP 32505 756 16 _ _ NNP 32505 756 17 and and CC 32505 756 18 _ _ NNP 32505 756 19 staufferi staufferi NN 32505 756 20 _ _ NNP 32505 756 21 depend depend VBP 32505 756 22 on on IN 32505 756 23 the the DT 32505 756 24 climatic climatic JJ 32505 756 25 changes change NNS 32505 756 26 that that WDT 32505 756 27 took take VBD 32505 756 28 place place NN 32505 756 29 during during IN 32505 756 30 the the DT 32505 756 31 Pleistocene Pleistocene NNP 32505 756 32 . . . 32505 757 1 On on IN 32505 757 2 this this DT 32505 757 3 basis basis NN 32505 757 4 it -PRON- PRP 32505 757 5 may may MD 32505 757 6 be be VB 32505 757 7 proposed propose VBN 32505 757 8 that that IN 32505 757 9 when when WRB 32505 757 10 the the DT 32505 757 11 original original JJ 32505 757 12 prototype prototype NN 32505 757 13 broke break VBD 32505 757 14 up up RP 32505 757 15 into into IN 32505 757 16 the the DT 32505 757 17 two two CD 32505 757 18 incipient incipient JJ 32505 757 19 species specie NNS 32505 757 20 , , , 32505 757 21 the the DT 32505 757 22 _ _ NNP 32505 757 23 staufferi staufferi NN 32505 757 24 _ _ NNP 32505 757 25 stock stock NN 32505 757 26 became become VBD 32505 757 27 physiologically physiologically RB 32505 757 28 and and CC 32505 757 29 behaviorally behaviorally RB 32505 757 30 adapted adapt VBN 32505 757 31 to to IN 32505 757 32 subhumid subhumid NN 32505 757 33 conditions condition NNS 32505 757 34 and and CC 32505 757 35 dispersed disperse VBN 32505 757 36 into into IN 32505 757 37 dry dry JJ 32505 757 38 areas area NNS 32505 757 39 of of IN 32505 757 40 the the DT 32505 757 41 lowlands lowland NNS 32505 757 42 of of IN 32505 757 43 Middle Middle NNP 32505 757 44 America America NNP 32505 757 45 . . . 32505 758 1 The the DT 32505 758 2 tropical tropical JJ 32505 758 3 evergreen evergreen JJ 32505 758 4 forests forest NNS 32505 758 5 on on IN 32505 758 6 the the DT 32505 758 7 Caribbean caribbean JJ 32505 758 8 side side NN 32505 758 9 of of IN 32505 758 10 lower low JJR 32505 758 11 Central Central NNP 32505 758 12 America America NNP 32505 758 13 and and CC 32505 758 14 the the DT 32505 758 15 uplift uplift NN 32505 758 16 of of IN 32505 758 17 the the DT 32505 758 18 Talamanca Talamanca NNP 32505 758 19 range range NN 32505 758 20 in in IN 32505 758 21 the the DT 32505 758 22 Pliocene Pliocene NNP 32505 758 23 were be VBD 32505 758 24 barriers barrier NNS 32505 758 25 to to IN 32505 758 26 the the DT 32505 758 27 dispersal dispersal NN 32505 758 28 of of IN 32505 758 29 _ _ NNP 32505 758 30 staufferi staufferi NN 32505 758 31 _ _ NNP 32505 758 32 . . . 32505 759 1 Consequently consequently RB 32505 759 2 , , , 32505 759 3 this this DT 32505 759 4 frog frog NN 32505 759 5 dispersed disperse VBN 32505 759 6 along along IN 32505 759 7 the the DT 32505 759 8 Pacific Pacific NNP 32505 759 9 lowlands lowland NNS 32505 759 10 . . . 32505 760 1 At at IN 32505 760 2 the the DT 32505 760 3 present present JJ 32505 760 4 time time NN 32505 760 5 _ _ NNP 32505 760 6 staufferi staufferi NN 32505 760 7 _ _ NNP 32505 760 8 occupies occupy VBZ 32505 760 9 the the DT 32505 760 10 length length NN 32505 760 11 of of IN 32505 760 12 the the DT 32505 760 13 Pacific Pacific NNP 32505 760 14 lowlands lowland NNS 32505 760 15 in in IN 32505 760 16 Central Central NNP 32505 760 17 America America NNP 32505 760 18 , , , 32505 760 19 except except IN 32505 760 20 in in IN 32505 760 21 the the DT 32505 760 22 rainforest rainforest NN 32505 760 23 of of IN 32505 760 24 the the DT 32505 760 25 Golfo Golfo NNP 32505 760 26 Duce Duce NNP 32505 760 27 region region NN 32505 760 28 , , , 32505 760 29 which which WDT 32505 760 30 apparently apparently RB 32505 760 31 is be VBZ 32505 760 32 a a DT 32505 760 33 relict relict JJ 32505 760 34 stand stand NN 32505 760 35 and and CC 32505 760 36 now now RB 32505 760 37 separates separate VBZ 32505 760 38 the the DT 32505 760 39 ranges range NNS 32505 760 40 of of IN 32505 760 41 two two CD 32505 760 42 subspecies subspecie NNS 32505 760 43 of of IN 32505 760 44 _ _ NNP 32505 760 45 Hyla Hyla NNP 32505 760 46 staufferi staufferi NN 32505 760 47 _ _ NNP 32505 760 48 . . . 32505 761 1 This this DT 32505 761 2 species specie NNS 32505 761 3 crossed cross VBD 32505 761 4 the the DT 32505 761 5 central central JJ 32505 761 6 Nicaraguan nicaraguan JJ 32505 761 7 lowlands lowland NNS 32505 761 8 and and CC 32505 761 9 reached reach VBD 32505 761 10 the the DT 32505 761 11 Caribbean caribbean JJ 32505 761 12 lowlands lowland NNS 32505 761 13 of of IN 32505 761 14 Nicaragua Nicaragua NNP 32505 761 15 and and CC 32505 761 16 nuclear nuclear JJ 32505 761 17 Central Central NNP 32505 761 18 America America NNP 32505 761 19 . . . 32505 762 1 The the DT 32505 762 2 species specie NNS 32505 762 3 migrated migrate VBN 32505 762 4 through through IN 32505 762 5 the the DT 32505 762 6 subhumid subhumid NNP 32505 762 7 corridor corridor NNP 32505 762 8 in in IN 32505 762 9 northern northern JJ 32505 762 10 Honduras Honduras NNP 32505 762 11 and and CC 32505 762 12 eastern eastern JJ 32505 762 13 Guatemala Guatemala NNP 32505 762 14 ( ( -LRB- 32505 762 15 Comayagua Comayagua NNP 32505 762 16 Valley Valley NNP 32505 762 17 in in IN 32505 762 18 Honduras Honduras NNP 32505 762 19 and and CC 32505 762 20 the the DT 32505 762 21 Motagua Motagua NNP 32505 762 22 Valley Valley NNP 32505 762 23 of of IN 32505 762 24 Guatemala Guatemala NNP 32505 762 25 ) ) -RRB- 32505 762 26 to to IN 32505 762 27 the the DT 32505 762 28 Isthmus Isthmus NNP 32505 762 29 of of IN 32505 762 30 Tehuantepec Tehuantepec NNP 32505 762 31 . . . 32505 763 1 Duellman Duellman NNP 32505 763 2 ( ( -LRB- 32505 763 3 1960 1960 CD 32505 763 4 ) ) -RRB- 32505 763 5 hypothesized hypothesize VBD 32505 763 6 " " '' 32505 763 7 that that IN 32505 763 8 during during IN 32505 763 9 the the DT 32505 763 10 times time NNS 32505 763 11 of of IN 32505 763 12 glacial glacial JJ 32505 763 13 advances advance NNS 32505 763 14 ( ( -LRB- 32505 763 15 Pleistocene Pleistocene NNP 32505 763 16 ) ) -RRB- 32505 763 17 the the DT 32505 763 18 lowlands lowland NNS 32505 763 19 of of IN 32505 763 20 the the DT 32505 763 21 Isthmus Isthmus NNP 32505 763 22 probably probably RB 32505 763 23 were be VBD 32505 763 24 more more RBR 32505 763 25 extensive extensive JJ 32505 763 26 and and CC 32505 763 27 had have VBD 32505 763 28 more more JJR 32505 763 29 semiarid semiarid NN 32505 763 30 tropical tropical JJ 32505 763 31 environments environment NNS 32505 763 32 than than IN 32505 763 33 at at IN 32505 763 34 the the DT 32505 763 35 present present NN 32505 763 36 " " '' 32505 763 37 and and CC 32505 763 38 that that IN 32505 763 39 when when WRB 32505 763 40 semiarid semiarid NN 32505 763 41 environments environment NNS 32505 763 42 were be VBD 32505 763 43 continuous continuous JJ 32505 763 44 from from IN 32505 763 45 the the DT 32505 763 46 Pacific Pacific NNP 32505 763 47 slope slope NN 32505 763 48 across across IN 32505 763 49 the the DT 32505 763 50 isthmus isthmus NN 32505 763 51 to to IN 32505 763 52 the the DT 32505 763 53 Gulf Gulf NNP 32505 763 54 lowlands lowland NNS 32505 763 55 _ _ NNP 32505 763 56 staufferi staufferi NN 32505 763 57 _ _ NNP 32505 763 58 and and CC 32505 763 59 other other JJ 32505 763 60 amphibians amphibian NNS 32505 763 61 migrated migrate VBN 32505 763 62 northward northward RB 32505 763 63 to to IN 32505 763 64 southeastern southeastern JJ 32505 763 65 Tamaulipas Tamaulipas NNP 32505 763 66 , , , 32505 763 67 México México NNP 32505 763 68 . . . 32505 764 1 _ _ NNP 32505 764 2 Hyla Hyla NNP 32505 764 3 elaeochroa elaeochroa NN 32505 764 4 _ _ NNP 32505 764 5 dispersed disperse VBD 32505 764 6 along along IN 32505 764 7 Caribbean caribbean JJ 32505 764 8 lowland lowland NN 32505 764 9 routes route NNS 32505 764 10 . . . 32505 765 1 This this DT 32505 765 2 species specie NNS 32505 765 3 not not RB 32505 765 4 only only RB 32505 765 5 occurs occur VBZ 32505 765 6 in in IN 32505 765 7 the the DT 32505 765 8 wet wet JJ 32505 765 9 forests forest NNS 32505 765 10 of of IN 32505 765 11 the the DT 32505 765 12 Golfo Golfo NNP 32505 765 13 Dulce Dulce NNP 32505 765 14 region region NN 32505 765 15 but but CC 32505 765 16 also also RB 32505 765 17 in in IN 32505 765 18 Guanacaste Guanacaste NNP 32505 765 19 . . . 32505 766 1 It -PRON- PRP 32505 766 2 is be VBZ 32505 766 3 possible possible JJ 32505 766 4 that that IN 32505 766 5 _ _ NNP 32505 766 6 elaeochroa elaeochroa NNP 32505 766 7 _ _ NNP 32505 766 8 entered enter VBD 32505 766 9 Guanacaste Guanacaste NNP 32505 766 10 and and CC 32505 766 11 moved move VBD 32505 766 12 to to IN 32505 766 13 the the DT 32505 766 14 Golfo Golfo NNP 32505 766 15 Dulce Dulce NNP 32505 766 16 region region NN 32505 766 17 when when WRB 32505 766 18 the the DT 32505 766 19 intervening intervening NN 32505 766 20 area area NN 32505 766 21 was be VBD 32505 766 22 less less JJR 32505 766 23 xeric xeric JJ 32505 766 24 than than IN 32505 766 25 now now RB 32505 766 26 ( ( -LRB- 32505 766 27 Duellman duellman NN 32505 766 28 , , , 32505 766 29 1966b 1966b CD 32505 766 30 ) ) -RRB- 32505 766 31 . . . 32505 767 1 _ _ NNP 32505 767 2 Hyla Hyla NNP 32505 767 3 elaeochroa elaeochroa NN 32505 767 4 _ _ NNP 32505 767 5 extended extend VBD 32505 767 6 its -PRON- PRP$ 32505 767 7 range range NN 32505 767 8 to to IN 32505 767 9 eastern eastern JJ 32505 767 10 Nicaragua Nicaragua NNP 32505 767 11 , , , 32505 767 12 but but CC 32505 767 13 even even RB 32505 767 14 though though IN 32505 767 15 northeastern northeastern JJ 32505 767 16 Nicaragua Nicaragua NNP 32505 767 17 has have VBZ 32505 767 18 over over IN 32505 767 19 2,000 2,000 CD 32505 767 20 mm mm CD 32505 767 21 . . . 32505 768 1 of of IN 32505 768 2 precipitation precipitation NN 32505 768 3 annually annually RB 32505 768 4 ( ( -LRB- 32505 768 5 Vivo Vivo NNP 32505 768 6 Escoto Escoto NNP 32505 768 7 , , , 32505 768 8 1964 1964 CD 32505 768 9 ) ) -RRB- 32505 768 10 , , , 32505 768 11 this this DT 32505 768 12 species species NN 32505 768 13 has have VBZ 32505 768 14 not not RB 32505 768 15 spread spread VBN 32505 768 16 into into IN 32505 768 17 Honduras Honduras NNP 32505 768 18 and and CC 32505 768 19 Guatemala Guatemala NNP 32505 768 20 . . . 32505 769 1 _ _ NNP 32505 769 2 Hyla Hyla NNP 32505 769 3 boulengeri boulengeri NN 32505 769 4 _ _ NNP 32505 769 5 is be VBZ 32505 769 6 widespread widespread JJ 32505 769 7 in in IN 32505 769 8 Amazonian amazonian JJ 32505 769 9 and and CC 32505 769 10 northern northern JJ 32505 769 11 South South NNP 32505 769 12 America America NNP 32505 769 13 , , , 32505 769 14 whereas whereas IN 32505 769 15 _ _ NNP 32505 769 16 foliamorta foliamorta NNP 32505 769 17 _ _ NNP 32505 769 18 occurs occur VBZ 32505 769 19 only only RB 32505 769 20 in in IN 32505 769 21 eastern eastern JJ 32505 769 22 Panamá Panamá NNP 32505 769 23 and and CC 32505 769 24 in in IN 32505 769 25 north north NN 32505 769 26 - - HYPH 32505 769 27 central central JJ 32505 769 28 Colombia Colombia NNP 32505 769 29 . . . 32505 770 1 The the DT 32505 770 2 ancestral ancestral JJ 32505 770 3 _ _ NNP 32505 770 4 boulengeri boulengeri NN 32505 770 5 - - HYPH 32505 770 6 foliamorta foliamorta NNP 32505 770 7 _ _ NNP 32505 770 8 stock stock NN 32505 770 9 probably probably RB 32505 770 10 invaded invade VBD 32505 770 11 Central Central NNP 32505 770 12 America America NNP 32505 770 13 in in IN 32505 770 14 the the DT 32505 770 15 late late JJ 32505 770 16 Pliocene Pliocene NNP 32505 770 17 and and CC 32505 770 18 dispersed disperse VBD 32505 770 19 through through IN 32505 770 20 humid humid JJ 32505 770 21 forested forested JJ 32505 770 22 environments environment NNS 32505 770 23 to to IN 32505 770 24 Nicaragua Nicaragua NNP 32505 770 25 . . . 32505 771 1 Apparently apparently RB 32505 771 2 a a DT 32505 771 3 peripheral peripheral JJ 32505 771 4 population population NN 32505 771 5 established establish VBD 32505 771 6 itself -PRON- PRP 32505 771 7 in in IN 32505 771 8 the the DT 32505 771 9 dry dry JJ 32505 771 10 Pacific Pacific NNP 32505 771 11 lowlands lowland NNS 32505 771 12 of of IN 32505 771 13 Panamá Panamá NNP 32505 771 14 . . . 32505 772 1 This this DT 32505 772 2 population population NN 32505 772 3 differentiated differentiate VBD 32505 772 4 from from IN 32505 772 5 _ _ NNP 32505 772 6 boulengeri boulengeri NN 32505 772 7 _ _ NNP 32505 772 8 of of IN 32505 772 9 the the DT 32505 772 10 humid humid JJ 32505 772 11 Caribbean caribbean JJ 32505 772 12 lowlands lowland NNS 32505 772 13 and and CC 32505 772 14 evolved evolve VBD 32505 772 15 into into IN 32505 772 16 _ _ NNP 32505 772 17 foliamorta foliamorta NNP 32505 772 18 _ _ NNP 32505 772 19 , , , 32505 772 20 which which WDT 32505 772 21 subsequently subsequently RB 32505 772 22 expanded expand VBD 32505 772 23 its -PRON- PRP$ 32505 772 24 range range NN 32505 772 25 into into IN 32505 772 26 Colombia Colombia NNP 32505 772 27 . . . 32505 773 1 LITERATURE LITERATURE NNP 32505 773 2 CITED cite VBD 32505 773 3 BLAIR BLAIR NNP 32505 773 4 , , , 32505 773 5 W. W. NNP 32505 773 6 F. F. NNP 32505 773 7 1960 1960 CD 32505 773 8 . . . 32505 774 1 Mating mate VBG 32505 774 2 call call NN 32505 774 3 as as IN 32505 774 4 evidence evidence NN 32505 774 5 of of IN 32505 774 6 relations relation NNS 32505 774 7 in in IN 32505 774 8 the the DT 32505 774 9 _ _ NNP 32505 774 10 Hyla Hyla NNP 32505 774 11 eximia eximia NN 32505 774 12 _ _ NNP 32505 774 13 group group NN 32505 774 14 . . . 32505 775 1 Southwestern Southwestern NNP 32505 775 2 Nat Nat NNP 32505 775 3 . . . 32505 775 4 , , , 32505 775 5 5(3):129 5(3):129 CD 32505 775 6 - - SYM 32505 775 7 135 135 CD 32505 775 8 . . . 32505 776 1 November November NNP 32505 776 2 1 1 CD 32505 776 3 . . . 32505 777 1 BOULENGER BOULENGER NNP 32505 777 2 , , , 32505 777 3 G. G. NNP 32505 777 4 A. A. NNP 32505 778 1 1882 1882 CD 32505 778 2 . . . 32505 779 1 Catalogue catalogue NN 32505 779 2 of of IN 32505 779 3 the the DT 32505 779 4 Batrachia Batrachia NNP 32505 779 5 Salientia Salientia NNP 32505 779 6 s. s. NNP 32505 779 7 Ecaudata Ecaudata NNP 32505 779 8 , , , 32505 779 9 2nd 2nd NN 32505 779 10 . . . 32505 780 1 ed ed NNP 32505 780 2 . . NNP 32505 780 3 , , , 32505 780 4 pp pp NNP 32505 780 5 . . . 32505 781 1 1 1 CD 32505 781 2 - - SYM 32505 781 3 503 503 CD 32505 781 4 , , , 32505 781 5 pls pls UH 32505 781 6 . . . 32505 782 1 1 1 CD 32505 782 2 - - SYM 32505 782 3 30 30 CD 32505 782 4 . . . 32505 783 1 February February NNP 32505 783 2 . . . 32505 784 1 BROCCHI BROCCHI NNP 32505 784 2 , , , 32505 784 3 P. P. NNP 32505 784 4 1881 1881 CD 32505 784 5 - - SYM 32505 784 6 1883 1883 CD 32505 784 7 . . . 32505 785 1 Etude Etude NNP 32505 785 2 des des NNP 32505 785 3 batraciens batracien NNS 32505 785 4 de de IN 32505 785 5 l'Amerique l'Amerique NNP 32505 785 6 Centrale Centrale NNP 32505 785 7 . . . 32505 786 1 Mission Mission NNP 32505 786 2 Scientifique Scientifique NNP 32505 786 3 au au NNP 32505 786 4 Mexique Mexique NNP 32505 786 5 et et NNP 32505 786 6 dans dan VBZ 32505 786 7 l'Amerique l'Amerique NNP 32505 786 8 Centrale Centrale NNP 32505 786 9 , , , 32505 786 10 Liv Liv NNP 32505 786 11 . . . 32505 787 1 1 1 CD 32505 787 2 , , , 32505 787 3 pp pp NNP 32505 787 4 . . . 32505 788 1 1 1 CD 32505 788 2 - - SYM 32505 788 3 122 122 CD 32505 788 4 , , , 32505 788 5 pls pls UH 32505 788 6 . . . 32505 789 1 1 1 CD 32505 789 2 - - SYM 32505 789 3 21 21 CD 32505 789 4 . . . 32505 790 1 COCHRAN COCHRAN NNP 32505 790 2 , , , 32505 790 3 D. D. NNP 32505 790 4 M. M. NNP 32505 790 5 1955 1955 CD 32505 790 6 . . . 32505 791 1 Frogs frog NNS 32505 791 2 of of IN 32505 791 3 southeastern southeastern JJ 32505 791 4 Brazil Brazil NNP 32505 791 5 . . . 32505 792 1 Bull Bull NNP 32505 792 2 . . . 32505 793 1 U.S. U.S. NNP 32505 793 2 Natl Natl NNP 32505 793 3 . . . 32505 794 1 Mus Mus NNP 32505 794 2 . . NNP 32505 794 3 , , , 32505 794 4 206:1 206:1 CD 32505 794 5 - - SYM 32505 794 6 423 423 CD 32505 794 7 . . . 32505 795 1 COPE COPE NNP 32505 795 2 , , , 32505 795 3 E. E. NNP 32505 795 4 D. D. NNP 32505 795 5 1865 1865 CD 32505 795 6 . . . 32505 796 1 Third third JJ 32505 796 2 contribution contribution NN 32505 796 3 to to IN 32505 796 4 the the DT 32505 796 5 Herpetology Herpetology NNP 32505 796 6 of of IN 32505 796 7 Tropical Tropical NNP 32505 796 8 America America NNP 32505 796 9 . . . 32505 797 1 Proc Proc NNP 32505 797 2 . . . 32505 798 1 Acad Acad NNS 32505 798 2 . . . 32505 799 1 Nat Nat NNP 32505 799 2 . . . 32505 800 1 Sci Sci NNP 32505 800 2 . . . 32505 801 1 Philadelphia Philadelphia NNP 32505 801 2 , , , 32505 801 3 17:195 17:195 CD 32505 801 4 . . . 32505 802 1 1876 1876 CD 32505 802 2 . . . 32505 803 1 On on IN 32505 803 2 the the DT 32505 803 3 batrachia batrachia NNS 32505 803 4 and and CC 32505 803 5 reptilia reptilia NNS 32505 803 6 of of IN 32505 803 7 Costa Costa NNP 32505 803 8 Rica Rica NNP 32505 803 9 . . . 32505 804 1 Jour Jour NNP 32505 804 2 . . . 32505 805 1 Acad Acad NNS 32505 805 2 . . . 32505 806 1 Nat Nat NNP 32505 806 2 . . . 32505 807 1 Sci Sci NNP 32505 807 2 . . . 32505 808 1 Philadelphia Philadelphia NNP 32505 808 2 , , , 32505 808 3 new new JJ 32505 808 4 series series NN 32505 808 5 , , , 32505 808 6 8:93 8:93 CD 32505 808 7 - - SYM 32505 808 8 154 154 CD 32505 808 9 , , , 32505 808 10 pls pls UH 32505 808 11 . . . 32505 809 1 23 23 CD 32505 809 2 - - SYM 32505 809 3 28 28 CD 32505 809 4 . . . 32505 810 1 1887 1887 CD 32505 810 2 . . . 32505 811 1 Thirteenth thirteenth JJ 32505 811 2 contribution contribution NN 32505 811 3 to to IN 32505 811 4 the the DT 32505 811 5 Herpetology Herpetology NNP 32505 811 6 of of IN 32505 811 7 Tropical Tropical NNP 32505 811 8 America America NNP 32505 811 9 . . . 32505 812 1 Proc Proc NNP 32505 812 2 . . . 32505 813 1 Amer amer NN 32505 813 2 . . . 32505 814 1 Philos Philos NNP 32505 814 2 . . . 32505 815 1 Society society NN 32505 815 2 , , , 32505 815 3 23:273 23:273 CD 32505 815 4 . . . 32505 816 1 April April NNP 32505 816 2 . . . 32505 817 1 DAUDIN DAUDIN NNP 32505 817 2 , , , 32505 817 3 F. F. NNP 32505 817 4 M. M. NNP 32505 817 5 1802 1802 CD 32505 817 6 . . . 32505 818 1 Histoire Histoire NNP 32505 818 2 naturelle naturelle NNP 32505 818 3 des des FW 32505 818 4 rainettes rainettes NNP 32505 818 5 , , , 32505 818 6 des des NNP 32505 818 7 grenouilles grenouilles NNP 32505 818 8 et et FW 32505 818 9 des des FW 32505 818 10 crapauds crapaud VBZ 32505 818 11 . . . 32505 819 1 Paris Paris NNP 32505 819 2 , , , 32505 819 3 pp pp NNP 32505 819 4 . . . 32505 820 1 1 1 CD 32505 820 2 - - SYM 32505 820 3 71 71 CD 32505 820 4 , , , 32505 820 5 pls pls UH 32505 820 6 . . . 32505 821 1 1 1 CD 32505 821 2 - - SYM 32505 821 3 33 33 CD 32505 821 4 . . . 32505 822 1 1803 1803 CD 32505 822 2 . . . 32505 823 1 Histoire Histoire NNP 32505 823 2 naturelle naturelle NNP 32505 823 3 générale générale FW 32505 823 4 et et FW 32505 823 5 particulière particulière NN 32505 823 6 des des NNP 32505 823 7 reptiles reptile VBZ 32505 823 8 . . . 32505 824 1 Paris Paris NNP 32505 824 2 , , , 32505 824 3 8:1 8:1 CD 32505 824 4 - - SYM 32505 824 5 439 439 CD 32505 824 6 . . . 32505 825 1 DUELLMAN DUELLMAN NNP 32505 825 2 , , , 32505 825 3 W. W. NNP 32505 825 4 E. E. NNP 32505 825 5 1956 1956 CD 32505 825 6 . . . 32505 826 1 The the DT 32505 826 2 frogs frog NNS 32505 826 3 of of IN 32505 826 4 the the DT 32505 826 5 hylid hylid JJ 32505 826 6 Genus Genus NNP 32505 826 7 _ _ NNP 32505 826 8 Phrynohyas Phrynohyas NNP 32505 826 9 Fitzinger Fitzinger NNP 32505 826 10 _ _ NNP 32505 826 11 , , , 32505 826 12 1843 1843 CD 32505 826 13 . . . 32505 827 1 Misc Misc NNP 32505 827 2 . . . 32505 828 1 Publ Publ NNS 32505 828 2 . . . 32505 829 1 Mus mus NN 32505 829 2 . . . 32505 830 1 Zool Zool NNP 32505 830 2 . . . 32505 830 3 , , , 32505 830 4 Univ Univ NNP 32505 830 5 . . . 32505 831 1 Mich. Michigan NNP 32505 831 2 , , , 32505 831 3 96:1 96:1 CD 32505 831 4 - - SYM 32505 831 5 47 47 CD 32505 831 6 , , , 32505 831 7 pls pls UH 32505 831 8 . . . 32505 832 1 1 1 CD 32505 832 2 - - SYM 32505 832 3 6 6 CD 32505 832 4 . . . 32505 833 1 February February NNP 32505 833 2 21 21 CD 32505 833 3 . . . 32505 834 1 1958 1958 CD 32505 834 2 . . . 32505 835 1 A a DT 32505 835 2 monographic monographic JJ 32505 835 3 study study NN 32505 835 4 of of IN 32505 835 5 the the DT 32505 835 6 colubrid colubrid NN 32505 835 7 snakes snake NNS 32505 835 8 Genus Genus NNP 32505 835 9 _ _ NNP 32505 835 10 Leptodeira Leptodeira NNP 32505 835 11 _ _ NNP 32505 835 12 . . . 32505 836 1 Bull Bull NNP 32505 836 2 . . . 32505 837 1 Amer amer NN 32505 837 2 . . . 32505 838 1 Mus mus NN 32505 838 2 . . . 32505 839 1 Nat Nat NNP 32505 839 2 . . . 32505 840 1 Hist Hist NNP 32505 840 2 . . . 32505 840 3 , , , 32505 840 4 114:1 114:1 CD 32505 840 5 - - SYM 32505 840 6 152 152 CD 32505 840 7 , , , 32505 840 8 pls pls UH 32505 840 9 . . . 32505 841 1 1 1 CD 32505 841 2 - - SYM 32505 841 3 31 31 CD 32505 841 4 . . . 32505 842 1 February February NNP 32505 842 2 24 24 CD 32505 842 3 . . . 32505 843 1 1960 1960 CD 32505 843 2 . . . 32505 844 1 A a DT 32505 844 2 distributional distributional JJ 32505 844 3 study study NN 32505 844 4 of of IN 32505 844 5 the the DT 32505 844 6 amphibians amphibian NNS 32505 844 7 of of IN 32505 844 8 the the DT 32505 844 9 Isthmus Isthmus NNP 32505 844 10 of of IN 32505 844 11 Tehuantepec Tehuantepec NNP 32505 844 12 , , , 32505 844 13 México México NNP 32505 844 14 . . . 32505 845 1 Univ Univ NNP 32505 845 2 . . . 32505 846 1 Kansas Kansas NNP 32505 846 2 Publ Publ NNP 32505 846 3 . . . 32505 846 4 , , , 32505 846 5 Mus mus NN 32505 846 6 . . . 32505 847 1 Nat Nat NNP 32505 847 2 . . . 32505 848 1 Hist Hist NNP 32505 848 2 , , , 32505 848 3 13:19 13:19 CD 32505 848 4 - - SYM 32505 848 5 72 72 CD 32505 848 6 , , , 32505 848 7 pls pls UH 32505 848 8 . . . 32505 849 1 1 1 CD 32505 849 2 - - SYM 32505 849 3 8 8 CD 32505 849 4 . . . 32505 850 1 August August NNP 32505 850 2 16 16 CD 32505 850 3 . . . 32505 851 1 1963 1963 CD 32505 851 2 . . . 32505 852 1 Amphibians amphibian NNS 32505 852 2 and and CC 32505 852 3 reptiles reptile NNS 32505 852 4 of of IN 32505 852 5 the the DT 32505 852 6 rainforests rainforest NNS 32505 852 7 of of IN 32505 852 8 southern southern JJ 32505 852 9 El El NNP 32505 852 10 Petén Petén NNP 32505 852 11 , , , 32505 852 12 Guatemala Guatemala NNP 32505 852 13 . . . 32505 853 1 Univ Univ NNP 32505 853 2 . . . 32505 854 1 Kansas Kansas NNP 32505 854 2 Publ Publ NNP 32505 854 3 . . . 32505 854 4 , , , 32505 854 5 Mus mus NN 32505 854 6 . . . 32505 855 1 Nat Nat NNP 32505 855 2 . . . 32505 856 1 Hist Hist NNP 32505 856 2 . . . 32505 856 3 , , , 32505 856 4 15:205 15:205 CD 32505 856 5 - - SYM 32505 856 6 249 249 CD 32505 856 7 , , , 32505 856 8 pls pls UH 32505 856 9 . . . 32505 857 1 7 7 CD 32505 857 2 - - SYM 32505 857 3 10 10 CD 32505 857 4 . . . 32505 858 1 October October NNP 32505 858 2 4 4 CD 32505 858 3 . . . 32505 859 1 1965 1965 CD 32505 859 2 . . . 32505 860 1 A a DT 32505 860 2 biogeographic biogeographic JJ 32505 860 3 account account NN 32505 860 4 of of IN 32505 860 5 the the DT 32505 860 6 herpetofauna herpetofauna NN 32505 860 7 of of IN 32505 860 8 Michoacán Michoacán NNP 32505 860 9 , , , 32505 860 10 México México NNP 32505 860 11 . . . 32505 861 1 Univ Univ NNP 32505 861 2 . . . 32505 862 1 Kansas Kansas NNP 32505 862 2 Publ Publ NNP 32505 862 3 . . . 32505 862 4 , , , 32505 862 5 Mus mus NN 32505 862 6 . . . 32505 863 1 Nat Nat NNP 32505 863 2 . . . 32505 864 1 Hist Hist NNP 32505 864 2 . . NNP 32505 864 3 , , , 32505 864 4 15:627 15:627 CD 32505 864 5 - - SYM 32505 864 6 709 709 CD 32505 864 7 , , , 32505 864 8 pls pls UH 32505 864 9 . . . 32505 865 1 29 29 CD 32505 865 2 - - SYM 32505 865 3 36 36 CD 32505 865 4 . . . 32505 866 1 December December NNP 32505 866 2 30 30 CD 32505 866 3 . . . 32505 867 1 1966a 1966a CD 32505 867 2 . . . 32505 868 1 Taxonomic taxonomic JJ 32505 868 2 notes note NNS 32505 868 3 on on IN 32505 868 4 some some DT 32505 868 5 Mexican mexican JJ 32505 868 6 and and CC 32505 868 7 Central central JJ 32505 868 8 American american JJ 32505 868 9 hylid hylid NNP 32505 868 10 frogs frog NNS 32505 868 11 . . . 32505 869 1 Univ Univ NNP 32505 869 2 . . . 32505 870 1 Kansas Kansas NNP 32505 870 2 Publ Publ NNP 32505 870 3 . . . 32505 870 4 , , , 32505 870 5 Mus mus NN 32505 870 6 . . . 32505 871 1 Nat Nat NNP 32505 871 2 . . . 32505 872 1 Hist Hist NNP 32505 872 2 . . . 32505 872 3 , , , 32505 872 4 17:263 17:263 CD 32505 872 5 - - SYM 32505 872 6 279 279 CD 32505 872 7 . . . 32505 873 1 June June NNP 32505 873 2 17 17 CD 32505 873 3 . . . 32505 874 1 1966b 1966b CD 32505 874 2 . . . 32505 875 1 The the DT 32505 875 2 Central central JJ 32505 875 3 American american JJ 32505 875 4 herpetofauna herpetofauna NN 32505 875 5 : : : 32505 875 6 An an DT 32505 875 7 ecological ecological JJ 32505 875 8 perspective perspective NN 32505 875 9 . . . 32505 876 1 Copeia Copeia NNP 32505 876 2 , , , 32505 876 3 4:700 4:700 CD 32505 876 4 - - SYM 32505 876 5 719 719 CD 32505 876 6 . . . 32505 877 1 December December NNP 32505 877 2 23 23 CD 32505 877 3 . . . 32505 878 1 1967 1967 CD 32505 878 2 . . . 32505 879 1 Social social JJ 32505 879 2 organization organization NN 32505 879 3 in in IN 32505 879 4 the the DT 32505 879 5 mating mating NN 32505 879 6 calls call NNS 32505 879 7 of of IN 32505 879 8 some some DT 32505 879 9 Neotropical Neotropical NNP 32505 879 10 Anurans Anurans NNPS 32505 879 11 . . . 32505 880 1 Amer amer NN 32505 880 2 . . . 32505 881 1 Midl Midl NNP 32505 881 2 . . . 32505 882 1 Nat Nat NNP 32505 882 2 . . NNP 32505 882 3 , , , 32505 882 4 77(1):156 77(1):156 CD 32505 882 5 - - SYM 32505 882 6 163 163 CD 32505 882 7 . . . 32505 883 1 January January NNP 32505 883 2 . . . 32505 884 1 DUELLMAN DUELLMAN NNP 32505 884 2 , , , 32505 884 3 W. W. NNP 32505 884 4 E. E. NNP 32505 884 5 , , , 32505 884 6 and and CC 32505 884 7 L. L. NNP 32505 884 8 TRUEB TRUEB NNP 32505 884 9 1966 1966 CD 32505 884 10 . . . 32505 885 1 Neotropical neotropical JJ 32505 885 2 hylid hylid NN 32505 885 3 frogs frog NNS 32505 885 4 , , , 32505 885 5 Genus Genus NNP 32505 885 6 Smilisca Smilisca NNP 32505 885 7 . . . 32505 886 1 Univ Univ NNP 32505 886 2 . . . 32505 887 1 Kansas Kansas NNP 32505 887 2 Publ Publ NNP 32505 887 3 . . . 32505 887 4 , , , 32505 887 5 Mus mus NN 32505 887 6 . . . 32505 888 1 Nat Nat NNP 32505 888 2 . . . 32505 889 1 Hist Hist NNP 32505 889 2 . . NNP 32505 889 3 , , , 32505 889 4 17:281 17:281 CD 32505 889 5 - - SYM 32505 889 6 375 375 CD 32505 889 7 , , , 32505 889 8 pls pls UH 32505 889 9 . . . 32505 890 1 1 1 CD 32505 890 2 - - SYM 32505 890 3 12 12 CD 32505 890 4 . . . 32505 891 1 July July NNP 32505 891 2 14 14 CD 32505 891 3 . . . 32505 892 1 DUNN DUNN NNP 32505 892 2 , , , 32505 892 3 E. E. NNP 32505 892 4 R. R. NNP 32505 892 5 1931a 1931a CD 32505 892 6 . . . 32505 893 1 The the DT 32505 893 2 amphibians amphibian NNS 32505 893 3 of of IN 32505 893 4 Barro Barro NNP 32505 893 5 Colorado Colorado NNP 32505 893 6 Island Island NNP 32505 893 7 . . . 32505 894 1 Occas occas JJ 32505 894 2 . . . 32505 895 1 Papers Papers NNP 32505 895 2 Boston Boston NNP 32505 895 3 Soc Soc NNP 32505 895 4 . . . 32505 896 1 Nat Nat NNP 32505 896 2 . . . 32505 897 1 Hist Hist NNP 32505 897 2 . . . 32505 897 3 , , , 32505 897 4 5:403 5:403 CD 32505 897 5 - - SYM 32505 897 6 421 421 CD 32505 897 7 . . . 32505 898 1 October October NNP 32505 898 2 10 10 CD 32505 898 3 . . . 32505 899 1 1931b 1931b LS 32505 899 2 . . . 32505 900 1 The the DT 32505 900 2 herpetological herpetological JJ 32505 900 3 fauna fauna NN 32505 900 4 of of IN 32505 900 5 the the DT 32505 900 6 Americas Americas NNPS 32505 900 7 . . . 32505 901 1 Copeia Copeia NNP 32505 901 2 , , , 32505 901 3 3:106 3:106 CD 32505 901 4 - - SYM 32505 901 5 119 119 CD 32505 901 6 . . . 32505 902 1 October October NNP 32505 902 2 30 30 CD 32505 902 3 . . . 32505 903 1 1933 1933 CD 32505 903 2 . . . 32505 904 1 A a DT 32505 904 2 new new JJ 32505 904 3 Hyla Hyla NNP 32505 904 4 from from IN 32505 904 5 Panamá Panamá NNP 32505 904 6 Canal Canal NNP 32505 904 7 Zone Zone NNP 32505 904 8 . . . 32505 905 1 Occas occas JJ 32505 905 2 . . . 32505 906 1 Papers Papers NNP 32505 906 2 Boston Boston NNP 32505 906 3 Soc Soc NNP 32505 906 4 . . . 32505 907 1 Nat Nat NNP 32505 907 2 . . . 32505 908 1 Hist Hist NNP 32505 908 2 . . . 32505 908 3 , , , 32505 908 4 8:61 8:61 CD 32505 908 5 - - SYM 32505 908 6 64 64 CD 32505 908 7 . . . 32505 909 1 June June NNP 32505 909 2 7 7 CD 32505 909 3 . . . 32505 910 1 DUNN DUNN NNP 32505 910 2 , , , 32505 910 3 E. E. NNP 32505 910 4 R. R. NNP 32505 910 5 , , , 32505 910 6 and and CC 32505 910 7 J. J. NNP 32505 910 8 EMLEN EMLEN NNP 32505 910 9 1932 1932 CD 32505 910 10 . . . 32505 911 1 Reptiles reptile NNS 32505 911 2 and and CC 32505 911 3 amphibians amphibian NNS 32505 911 4 from from IN 32505 911 5 Honduras Honduras NNP 32505 911 6 . . . 32505 912 1 Proc Proc NNP 32505 912 2 . . . 32505 913 1 Acad Acad NNS 32505 913 2 . . . 32505 914 1 Nat Nat NNP 32505 914 2 . . . 32505 915 1 Sci Sci NNP 32505 915 2 . . . 32505 916 1 Philadelphia Philadelphia NNP 32505 916 2 , , , 32505 916 3 84:24 84:24 CD 32505 916 4 - - SYM 32505 916 5 25 25 CD 32505 916 6 . . . 32505 917 1 March March NNP 32505 917 2 22 22 CD 32505 917 3 . . . 32505 918 1 FOUQUETTE FOUQUETTE NNP 32505 918 2 , , , 32505 918 3 M. M. NNP 32505 918 4 J. J. NNP 32505 919 1 1958 1958 CD 32505 919 2 . . . 32505 920 1 A a DT 32505 920 2 new new JJ 32505 920 3 tree tree NN 32505 920 4 frog frog NN 32505 920 5 , , , 32505 920 6 Genus Genus NNP 32505 920 7 _ _ NNP 32505 920 8 Hyla Hyla NNP 32505 920 9 _ _ NNP 32505 920 10 , , , 32505 920 11 from from IN 32505 920 12 the the DT 32505 920 13 Canal Canal NNP 32505 920 14 Zone Zone NNP 32505 920 15 . . . 32505 921 1 Herpetologica Herpetologica NNP 32505 921 2 , , , 32505 921 3 14:125 14:125 CD 32505 921 4 - - SYM 32505 921 5 128 128 CD 32505 921 6 . . . 32505 922 1 April April NNP 32505 922 2 25 25 CD 32505 922 3 . . . 32505 923 1 GAIGE GAIGE NNP 32505 923 2 , , , 32505 923 3 H. H. NNP 32505 923 4 T. T. NNP 32505 923 5 1936 1936 CD 32505 923 6 . . . 32505 924 1 Some some DT 32505 924 2 reptiles reptile NNS 32505 924 3 and and CC 32505 924 4 amphibians amphibian NNS 32505 924 5 from from IN 32505 924 6 Yucatan Yucatan NNP 32505 924 7 and and CC 32505 924 8 Campeche Campeche NNP 32505 924 9 , , , 32505 924 10 México México NNP 32505 924 11 . . . 32505 925 1 Carnegie Carnegie NNP 32505 925 2 Inst Inst NNP 32505 925 3 . . . 32505 926 1 Washington Washington NNP 32505 926 2 Publ Publ NNP 32505 926 3 . . . 32505 926 4 , , , 32505 926 5 457:289 457:289 CD 32505 926 6 - - SYM 32505 926 7 304 304 CD 32505 926 8 . . . 32505 927 1 GOSNER GOSNER NNP 32505 927 2 , , , 32505 927 3 K. K. NNP 32505 927 4 L. L. NNP 32505 927 5 1960 1960 CD 32505 927 6 . . . 32505 928 1 A a DT 32505 928 2 simplified simplify VBN 32505 928 3 table table NN 32505 928 4 for for IN 32505 928 5 staging stage VBG 32505 928 6 anuran anuran NNP 32505 928 7 embryos embryo NNS 32505 928 8 and and CC 32505 928 9 larvae larvae VBN 32505 928 10 with with IN 32505 928 11 notes note NNS 32505 928 12 on on IN 32505 928 13 identification identification NN 32505 928 14 . . . 32505 929 1 Herpetologica Herpetologica NNP 32505 929 2 , , , 32505 929 3 16:183 16:183 CD 32505 929 4 - - SYM 32505 929 5 190 190 CD 32505 929 6 . . . 32505 930 1 June June NNP 32505 930 2 17 17 CD 32505 930 3 . . . 32505 931 1 GÜNTHER GÜNTHER NNP 32505 931 2 , , , 32505 931 3 A. a. NN 32505 931 4 C. C. NNP 32505 931 5 L. L. NNP 32505 931 6 1859 1859 CD 32505 931 7 . . . 32505 932 1 Catalogue catalogue NN 32505 932 2 of of IN 32505 932 3 the the DT 32505 932 4 Batrachia Batrachia NNP 32505 932 5 Salientia Salientia NNP 32505 932 6 in in IN 32505 932 7 the the DT 32505 932 8 British British NNP 32505 932 9 Museum Museum NNP 32505 932 10 . . . 32505 933 1 pp pp NNP 32505 933 2 . . . 32505 934 1 1 1 CD 32505 934 2 - - SYM 32505 934 3 160 160 CD 32505 934 4 . . . 32505 935 1 1885 1885 CD 32505 935 2 - - SYM 32505 935 3 1902 1902 CD 32505 935 4 . . . 32505 936 1 Biologia Biologia NNP 32505 936 2 Centrali Centrali NNP 32505 936 3 Americana Americana NNP 32505 936 4 . . . 32505 937 1 Reptilia Reptilia NNP 32505 937 2 and and CC 32505 937 3 Batrachia Batrachia NNP 32505 937 4 . . . 32505 938 1 London London NNP 32505 938 2 , , , 32505 938 3 xx xx NN 32505 938 4 + + DT 32505 938 5 326 326 CD 32505 938 6 pp pp NNP 32505 938 7 . . . 32505 938 8 , , , 32505 938 9 pls pls UH 32505 938 10 . . . 32505 939 1 1 1 CD 32505 939 2 - - SYM 32505 939 3 76 76 CD 32505 939 4 . . . 32505 940 1 KELLOGG KELLOGG NNP 32505 940 2 , , , 32505 940 3 R. R. NNP 32505 940 4 1932 1932 CD 32505 940 5 . . . 32505 941 1 Mexican mexican JJ 32505 941 2 tailless tailless NN 32505 941 3 amphibians amphibian NNS 32505 941 4 in in IN 32505 941 5 the the DT 32505 941 6 United United NNP 32505 941 7 States States NNP 32505 941 8 National National NNP 32505 941 9 Museum Museum NNP 32505 941 10 , , , 32505 941 11 Bull Bull NNP 32505 941 12 . . . 32505 942 1 U.S. U.S. NNP 32505 942 2 Natl Natl NNP 32505 942 3 . . . 32505 943 1 Mus Mus NNP 32505 943 2 . . NNP 32505 943 3 , , , 32505 943 4 160:1 160:1 CD 32505 943 5 - - SYM 32505 943 6 224 224 CD 32505 943 7 . . . 32505 944 1 LAURENTI LAURENTI NNP 32505 944 2 , , , 32505 944 3 J. J. NNP 32505 944 4 N. N. NNP 32505 944 5 1768 1768 CD 32505 944 6 . . . 32505 945 1 Specimen Specimen NNP 32505 945 2 medicum medicum NNP 32505 945 3 exhibens exhibens NNP 32505 945 4 synopsin synopsin NNP 32505 945 5 reptilium reptilium NN 32505 945 6 emendatum emendatum NNP 32505 945 7 cum cum NNP 32505 945 8 experimentis experimentis NNP 32505 945 9 circa circa NNP 32505 945 10 venema venema NNP 32505 945 11 et et NNP 32505 945 12 antidota antidota NNP 32505 945 13 reptilium reptilium NNP 32505 945 14 austriacorum austriacorum NNP 32505 945 15 . . . 32505 946 1 Vienna Vienna NNP 32505 946 2 , , , 32505 946 3 pp pp NNP 32505 946 4 . . . 32505 947 1 1 1 LS 32505 947 2 - - SYM 32505 947 3 216 216 CD 32505 947 4 . . . 32505 948 1 LINNAEUS LINNAEUS NNP 32505 948 2 , , , 32505 948 3 C. C. NNP 32505 948 4 1758 1758 CD 32505 948 5 . . . 32505 949 1 Systema systema JJ 32505 949 2 naturae naturae NNS 32505 949 3 . . . 32505 950 1 Holmiae Holmiae NNP 32505 950 2 , , , 32505 950 3 ed ed NNP 32505 950 4 . . . 32505 951 1 10 10 CD 32505 951 2 reformata reformata NN 32505 951 3 , , , 32505 951 4 I:1 I:1 NNP 32505 951 5 - - HYPH 32505 951 6 532 532 CD 32505 951 7 . . . 32505 952 1 MALDONADO MALDONADO NNP 32505 952 2 - - HYPH 32505 952 3 KOERDELL KOERDELL NNP 32505 952 4 , , , 32505 952 5 M. M. NNP 32505 952 6 1964 1964 CD 32505 952 7 . . . 32505 953 1 Geohistory geohistory NN 32505 953 2 and and CC 32505 953 3 paleography paleography NN 32505 953 4 of of IN 32505 953 5 Middle Middle NNP 32505 953 6 America America NNP 32505 953 7 . . . 32505 954 1 In in IN 32505 954 2 R. R. NNP 32505 954 3 Wauchope Wauchope NNP 32505 954 4 and and CC 32505 954 5 R. R. NNP 32505 954 6 C. C. NNP 32505 954 7 West West NNP 32505 954 8 ( ( -LRB- 32505 954 9 Eds Eds NNP 32505 954 10 . . . 32505 954 11 ) ) -RRB- 32505 954 12 . . . 32505 955 1 Handbook Handbook NNP 32505 955 2 of of IN 32505 955 3 Middle middle JJ 32505 955 4 American American NNP 32505 955 5 Indians Indians NNPS 32505 955 6 , , , 32505 955 7 Vol Vol NNP 32505 955 8 . . . 32505 956 1 I -PRON- PRP 32505 956 2 , , , 32505 956 3 Univ Univ NNP 32505 956 4 . . . 32505 957 1 Texas Texas NNP 32505 957 2 Press Press NNP 32505 957 3 , , , 32505 957 4 Austin Austin NNP 32505 957 5 , , , 32505 957 6 570 570 CD 32505 957 7 pp pp NN 32505 957 8 . . . 32505 958 1 NIEDEN NIEDEN NNP 32505 958 2 , , , 32505 958 3 F. F. NNP 32505 958 4 1923 1923 CD 32505 958 5 . . . 32505 959 1 Amphibia Amphibia NNP 32505 959 2 : : : 32505 959 3 Anura Anura NNP 32505 959 4 I. I. NNP 32505 959 5 Das Das NNP 32505 959 6 Tierreich Tierreich NNP 32505 959 7 . . . 32505 960 1 Berlin Berlin NNP 32505 960 2 , , , 32505 960 3 584 584 CD 32505 960 4 pp pp NN 32505 960 5 . . . 32505 961 1 NOBLE NOBLE NNP 32505 961 2 , , , 32505 961 3 G. G. NNP 32505 961 4 K. K. NNP 32505 961 5 1918 1918 CD 32505 961 6 . . . 32505 962 1 The the DT 32505 962 2 amphibians amphibian NNS 32505 962 3 collected collect VBN 32505 962 4 by by IN 32505 962 5 the the DT 32505 962 6 American American NNP 32505 962 7 Museum Museum NNP 32505 962 8 expedition expedition NN 32505 962 9 to to IN 32505 962 10 Nicaragua Nicaragua NNP 32505 962 11 in in IN 32505 962 12 1916 1916 CD 32505 962 13 . . . 32505 963 1 Bull Bull NNP 32505 963 2 . . . 32505 964 1 Amer amer NN 32505 964 2 . . . 32505 965 1 Mus mus NN 32505 965 2 . . . 32505 966 1 Nat Nat NNP 32505 966 2 . . . 32505 967 1 Hist Hist NNP 32505 967 2 . . NNP 32505 967 3 , , , 32505 967 4 38:311 38:311 CD 32505 967 5 - - SYM 32505 967 6 347 347 CD 32505 967 7 . . . 32505 968 1 RAND RAND NNP 32505 968 2 , , , 32505 968 3 A. A. NNP 32505 968 4 S. S. NNP 32505 968 5 1957 1957 CD 32505 968 6 . . . 32505 969 1 Notes note NNS 32505 969 2 on on IN 32505 969 3 amphibians amphibian NNS 32505 969 4 and and CC 32505 969 5 reptiles reptile NNS 32505 969 6 from from IN 32505 969 7 El El NNP 32505 969 8 Salvador Salvador NNP 32505 969 9 . . . 32505 970 1 Fieldiana Fieldiana NNP 32505 970 2 , , , 32505 970 3 Zool Zool NNP 32505 970 4 . . NNP 32505 970 5 , , , 32505 970 6 Chicago Chicago NNP 32505 970 7 Nat Nat NNP 32505 970 8 . . . 32505 971 1 Hist hist NN 32505 971 2 . . . 32505 972 1 Mus Mus NNP 32505 972 2 . . NNP 32505 972 3 , , , 32505 972 4 43:505 43:505 NNP 32505 972 5 - - SYM 32505 972 6 534 534 CD 32505 972 7 . . . 32505 973 1 April April NNP 32505 973 2 18 18 CD 32505 973 3 . . . 32505 974 1 RIVERO RIVERO NNP 32505 974 2 , , , 32505 974 3 J. J. NNP 32505 975 1 A. A. NNP 32505 976 1 1961 1961 CD 32505 976 2 . . . 32505 977 1 Salientia Salientia NNP 32505 977 2 of of IN 32505 977 3 Venezuela Venezuela NNP 32505 977 4 . . . 32505 978 1 Bull Bull NNP 32505 978 2 . . . 32505 979 1 Mus mus NN 32505 979 2 . . . 32505 980 1 Comp Comp NNP 32505 980 2 . . . 32505 981 1 Zool Zool NNP 32505 981 2 . . NNP 32505 981 3 , , , 32505 981 4 126:1 126:1 CD 32505 981 5 - - SYM 32505 981 6 207 207 CD 32505 981 7 . . . 32505 982 1 November November NNP 32505 982 2 . . . 32505 983 1 SAVAGE SAVAGE NNP 32505 983 2 , , , 32505 983 3 J. J. NNP 32505 983 4 M. M. NNP 32505 983 5 1966 1966 CD 32505 983 6 . . . 32505 984 1 The the DT 32505 984 2 origins origin NNS 32505 984 3 and and CC 32505 984 4 history history NN 32505 984 5 of of IN 32505 984 6 the the DT 32505 984 7 Central central JJ 32505 984 8 American american JJ 32505 984 9 herpetofauna herpetofauna NN 32505 984 10 . . . 32505 985 1 Copeia Copeia NNP 32505 985 2 , , , 32505 985 3 4:719 4:719 CD 32505 985 4 - - SYM 32505 985 5 756 756 CD 32505 985 6 . . . 32505 986 1 December December NNP 32505 986 2 23 23 CD 32505 986 3 . . . 32505 987 1 SCHMIDT schmidt NN 32505 987 2 , , , 32505 987 3 K. K. NNP 32505 987 4 P. P. NNP 32505 987 5 , , , 32505 987 6 and and CC 32505 987 7 L. L. NNP 32505 987 8 C. C. NNP 32505 987 9 STUART STUART NNP 32505 987 10 1941 1941 CD 32505 987 11 . . . 32505 988 1 The the DT 32505 988 2 herpetological herpetological JJ 32505 988 3 fauna fauna NN 32505 988 4 of of IN 32505 988 5 the the DT 32505 988 6 Salamá Salamá NNP 32505 988 7 Basin Basin NNP 32505 988 8 , , , 32505 988 9 Baja Baja NNP 32505 988 10 Verapaz Verapaz NNP 32505 988 11 , , , 32505 988 12 Guatemala Guatemala NNP 32505 988 13 . . . 32505 989 1 Field field NN 32505 989 2 Mus Mus NNP 32505 989 3 . . . 32505 990 1 Nat Nat NNP 32505 990 2 . . . 32505 991 1 Hist Hist NNP 32505 991 2 . . NNP 32505 991 3 , , , 32505 991 4 Zool Zool NNP 32505 991 5 . . . 32505 992 1 Ser Ser NNP 32505 992 2 . . NNP 32505 992 3 , , , 32505 992 4 24:233 24:233 CD 32505 992 5 - - HYPH 32505 992 6 247 247 CD 32505 992 7 . . . 32505 993 1 SEBA SEBA NNP 32505 993 2 , , , 32505 993 3 A. a. NN 32505 994 1 1734 1734 CD 32505 994 2 . . . 32505 995 1 Locupletissimi Locupletissimi NNP 32505 995 2 rerum rerum NNP 32505 995 3 naturalium naturalium NN 32505 995 4 thesauri thesauri NNP 32505 995 5 accurata accurata NNP 32505 995 6 descriptio descriptio NNP 32505 995 7 , , , 32505 995 8 et et NNP 32505 995 9 iconibus iconibus NNP 32505 995 10 artificissimis artificissimis NNP 32505 995 11 expressio expressio NNP 32505 995 12 , , , 32505 995 13 per per IN 32505 995 14 universam universam NNP 32505 995 15 physis physis NNP 32505 995 16 historiam historiam NNP 32505 995 17 . . . 32505 996 1 Amsterdam Amsterdam NNP 32505 996 2 , , , 32505 996 3 I -PRON- PRP 32505 996 4 xxxiv xxxiv NN 32505 996 5 + + SYM 32505 996 6 178 178 CD 32505 996 7 pp pp NNP 32505 996 8 . . . 32505 996 9 , , , 32505 996 10 pls pls UH 32505 996 11 . . . 32505 997 1 1 1 CD 32505 997 2 - - SYM 32505 997 3 111 111 CD 32505 997 4 . . . 32505 998 1 SIMPSON SIMPSON NNP 32505 998 2 , , , 32505 998 3 G. G. NNP 32505 998 4 G. G. NNP 32505 998 5 1943 1943 CD 32505 998 6 . . . 32505 999 1 Turtles turtle NNS 32505 999 2 and and CC 32505 999 3 the the DT 32505 999 4 origin origin NN 32505 999 5 of of IN 32505 999 6 the the DT 32505 999 7 fauna fauna NNS 32505 999 8 of of IN 32505 999 9 Latin Latin NNP 32505 999 10 America America NNP 32505 999 11 . . . 32505 1000 1 Amer amer NN 32505 1000 2 . . . 32505 1001 1 Jour Jour NNP 32505 1001 2 . . . 32505 1002 1 Sci Sci NNP 32505 1002 2 . . NNP 32505 1002 3 , , , 32505 1002 4 24:413 24:413 CD 32505 1002 5 - - SYM 32505 1002 6 429 429 CD 32505 1002 7 . . . 32505 1003 1 1965 1965 CD 32505 1003 2 . . . 32505 1004 1 The the DT 32505 1004 2 geography geography NN 32505 1004 3 of of IN 32505 1004 4 evolution evolution NN 32505 1004 5 . . . 32505 1005 1 Chilton Chilton NNP 32505 1005 2 Books Books NNPS 32505 1005 3 Publishers Publishers NNPS 32505 1005 4 . . . 32505 1006 1 Philadelphia Philadelphia NNP 32505 1006 2 , , , 32505 1006 3 pp pp NNP 32505 1006 4 . . . 32505 1007 1 1 1 CD 32505 1007 2 - - SYM 32505 1007 3 349 349 CD 32505 1007 4 . . . 32505 1008 1 July July NNP 32505 1008 2 . . . 32505 1009 1 SMITH SMITH NNP 32505 1009 2 , , , 32505 1009 3 H. H. NNP 32505 1009 4 M. M. NNP 32505 1009 5 and and CC 32505 1009 6 E. E. NNP 32505 1009 7 H. H. NNP 32505 1009 8 TAYLOR TAYLOR NNP 32505 1009 9 1948 1948 CD 32505 1009 10 . . . 32505 1010 1 An an DT 32505 1010 2 annotated annotate VBN 32505 1010 3 checklist checklist NN 32505 1010 4 and and CC 32505 1010 5 key key NN 32505 1010 6 to to IN 32505 1010 7 the the DT 32505 1010 8 amphibians amphibian NNS 32505 1010 9 of of IN 32505 1010 10 México México NNP 32505 1010 11 . . . 32505 1011 1 Bull Bull NNP 32505 1011 2 . . . 32505 1012 1 U.S. U.S. NNP 32505 1012 2 Natl Natl NNP 32505 1012 3 . . . 32505 1013 1 Mus Mus NNP 32505 1013 2 . . NNP 32505 1013 3 , , , 32505 1013 4 194:1 194:1 CD 32505 1013 5 - - SYM 32505 1013 6 118 118 CD 32505 1013 7 . . . 32505 1014 1 STARRETT STARRETT NNP 32505 1014 2 , , , 32505 1014 3 P. P. NNP 32505 1014 4 1960 1960 CD 32505 1014 5 . . . 32505 1015 1 Description description NN 32505 1015 2 of of IN 32505 1015 3 tadpoles tadpole NNS 32505 1015 4 of of IN 32505 1015 5 Middle middle JJ 32505 1015 6 American american JJ 32505 1015 7 frogs frog NNS 32505 1015 8 . . . 32505 1016 1 Misc Misc NNP 32505 1016 2 . . . 32505 1017 1 Publ Publ NNS 32505 1017 2 . . . 32505 1018 1 Mus mus NN 32505 1018 2 . . . 32505 1019 1 Zool Zool NNP 32505 1019 2 . . . 32505 1019 3 , , , 32505 1019 4 Univ Univ NNP 32505 1019 5 . . . 32505 1020 1 Michigan Michigan NNP 32505 1020 2 , , , 32505 1020 3 110:1 110:1 CD 32505 1020 4 - - SYM 32505 1020 5 37 37 CD 32505 1020 6 , , , 32505 1020 7 pl pl NNP 32505 1020 8 . . . 32505 1021 1 1 1 LS 32505 1021 2 . . . 32505 1022 1 STUART STUART NNP 32505 1022 2 , , , 32505 1022 3 L. L. NNP 32505 1022 4 C. C. NNP 32505 1022 5 1935 1935 CD 32505 1022 6 . . . 32505 1023 1 A a DT 32505 1023 2 contribution contribution NN 32505 1023 3 to to IN 32505 1023 4 a a DT 32505 1023 5 knowledge knowledge NN 32505 1023 6 of of IN 32505 1023 7 the the DT 32505 1023 8 herpetology herpetology NN 32505 1023 9 of of IN 32505 1023 10 a a DT 32505 1023 11 portion portion NN 32505 1023 12 of of IN 32505 1023 13 the the DT 32505 1023 14 savanna savanna NN 32505 1023 15 region region NN 32505 1023 16 of of IN 32505 1023 17 Central Central NNP 32505 1023 18 Petén Petén NNP 32505 1023 19 , , , 32505 1023 20 Guatemala Guatemala NNP 32505 1023 21 . . . 32505 1024 1 Misc Misc NNP 32505 1024 2 . . . 32505 1025 1 Publ Publ NNS 32505 1025 2 . . . 32505 1026 1 Mus mus NN 32505 1026 2 . . . 32505 1027 1 Zool Zool NNP 32505 1027 2 . . . 32505 1027 3 , , , 32505 1027 4 Univ Univ NNP 32505 1027 5 . . . 32505 1028 1 Michigan Michigan NNP 32505 1028 2 , , , 32505 1028 3 29:1 29:1 CD 32505 1028 4 - - SYM 32505 1028 5 56 56 CD 32505 1028 6 , , , 32505 1028 7 pls pls UH 32505 1028 8 . . . 32505 1029 1 1 1 CD 32505 1029 2 - - SYM 32505 1029 3 4 4 CD 32505 1029 4 . . . 32505 1030 1 October October NNP 32505 1030 2 1 1 CD 32505 1030 3 . . . 32505 1031 1 1948 1948 CD 32505 1031 2 . . . 32505 1032 1 The the DT 32505 1032 2 amphibians amphibian NNS 32505 1032 3 and and CC 32505 1032 4 reptiles reptile NNS 32505 1032 5 of of IN 32505 1032 6 Alta Alta NNP 32505 1032 7 Verapaz Verapaz NNP 32505 1032 8 , , , 32505 1032 9 Guatemala Guatemala NNP 32505 1032 10 . . . 32505 1033 1 Misc Misc NNP 32505 1033 2 . . . 32505 1034 1 Publ Publ NNS 32505 1034 2 . . . 32505 1035 1 Mus mus NN 32505 1035 2 . . . 32505 1036 1 Zool Zool NNP 32505 1036 2 . . . 32505 1036 3 , , , 32505 1036 4 Univ Univ NNP 32505 1036 5 . . . 32505 1037 1 Michigan Michigan NNP 32505 1037 2 , , , 32505 1037 3 69:1 69:1 CD 32505 1037 4 - - HYPH 32505 1037 5 109 109 CD 32505 1037 6 . . . 32505 1038 1 June June NNP 32505 1038 2 12 12 CD 32505 1038 3 . . . 32505 1039 1 1950 1950 CD 32505 1039 2 . . . 32505 1040 1 A a DT 32505 1040 2 geographic geographic JJ 32505 1040 3 study study NN 32505 1040 4 of of IN 32505 1040 5 the the DT 32505 1040 6 herpetofauna herpetofauna NN 32505 1040 7 of of IN 32505 1040 8 Alta Alta NNP 32505 1040 9 Verapaz Verapaz NNP 32505 1040 10 , , , 32505 1040 11 Guatemala Guatemala NNP 32505 1040 12 . . . 32505 1041 1 Contr Contr NNP 32505 1041 2 . . . 32505 1042 1 Lab lab NN 32505 1042 2 . . . 32505 1043 1 Vert Vert NNP 32505 1043 2 . . . 32505 1044 1 Biol Biol NNP 32505 1044 2 . . . 32505 1044 3 , , , 32505 1044 4 Univ Univ NNP 32505 1044 5 . . . 32505 1045 1 Michigan Michigan NNP 32505 1045 2 , , , 32505 1045 3 45:1 45:1 CD 32505 1045 4 - - SYM 32505 1045 5 77 77 CD 32505 1045 6 , , , 32505 1045 7 pls pls UH 32505 1045 8 . . . 32505 1046 1 1 1 CD 32505 1046 2 - - SYM 32505 1046 3 9 9 CD 32505 1046 4 . . . 32505 1047 1 May. May NNP 32505 1048 1 TAYLOR TAYLOR NNP 32505 1048 2 , , , 32505 1048 3 E. E. NNP 32505 1048 4 H. H. NNP 32505 1048 5 1938 1938 CD 32505 1048 6 . . . 32505 1049 1 Frogs frog NNS 32505 1049 2 of of IN 32505 1049 3 the the DT 32505 1049 4 _ _ NNP 32505 1049 5 Hyla Hyla NNP 32505 1049 6 eximia eximia NN 32505 1049 7 _ _ NNP 32505 1049 8 group group NN 32505 1049 9 , , , 32505 1049 10 with with IN 32505 1049 11 description description NN 32505 1049 12 of of IN 32505 1049 13 two two CD 32505 1049 14 new new JJ 32505 1049 15 species specie NNS 32505 1049 16 . . . 32505 1050 1 Univ Univ NNP 32505 1050 2 . . . 32505 1051 1 Kansas Kansas NNP 32505 1051 2 Sci Sci NNP 32505 1051 3 . . . 32505 1052 1 Bull Bull NNP 32505 1052 2 . . NNP 32505 1052 3 , , , 32505 1052 4 25:421 25:421 CD 32505 1052 5 - - SYM 32505 1052 6 445 445 CD 32505 1052 7 . . . 32505 1053 1 June June NNP 32505 1053 2 1 1 CD 32505 1053 3 . . . 32505 1054 1 1952 1952 CD 32505 1054 2 . . . 32505 1055 1 The the DT 32505 1055 2 frogs frog NNS 32505 1055 3 and and CC 32505 1055 4 toads toad NNS 32505 1055 5 of of IN 32505 1055 6 Costa Costa NNP 32505 1055 7 Rica Rica NNP 32505 1055 8 . . . 32505 1056 1 Univ Univ NNP 32505 1056 2 . . . 32505 1057 1 Kansas Kansas NNP 32505 1057 2 Sci Sci NNP 32505 1057 3 . . . 32505 1058 1 Bull Bull NNP 32505 1058 2 . . . 32505 1058 3 , , , 32505 1058 4 35:577 35:577 CD 32505 1058 5 - - SYM 32505 1058 6 942 942 CD 32505 1058 7 . . . 32505 1059 1 July July NNP 32505 1059 2 1 1 CD 32505 1059 3 . . . 32505 1060 1 1958 1958 CD 32505 1060 2 . . . 32505 1061 1 Additions addition NNS 32505 1061 2 to to IN 32505 1061 3 the the DT 32505 1061 4 known know VBN 32505 1061 5 herpetological herpetological JJ 32505 1061 6 fauna fauna NNS 32505 1061 7 of of IN 32505 1061 8 Costa Costa NNP 32505 1061 9 Rica Rica NNP 32505 1061 10 , , , 32505 1061 11 with with IN 32505 1061 12 comments comment NNS 32505 1061 13 on on IN 32505 1061 14 other other JJ 32505 1061 15 species specie NNS 32505 1061 16 . . . 32505 1062 1 No no UH 32505 1062 2 . . . 32505 1063 1 III iii CD 32505 1063 2 . . . 32505 1064 1 Univ Univ NNP 32505 1064 2 . . . 32505 1065 1 Kansas Kansas NNP 32505 1065 2 Sci Sci NNP 32505 1065 3 . . . 32505 1066 1 Bull Bull NNP 32505 1066 2 . . NNP 32505 1066 3 , , , 32505 1066 4 34:3 34:3 CD 32505 1066 5 - - SYM 32505 1066 6 40 40 CD 32505 1066 7 . . . 32505 1067 1 November November NNP 32505 1067 2 18 18 CD 32505 1067 3 . . . 32505 1068 1 VIVO VIVO NNP 32505 1068 2 ESCOTO ESCOTO NNP 32505 1068 3 , , , 32505 1068 4 J. J. NNP 32505 1069 1 A. A. NNP 32505 1070 1 1964 1964 CD 32505 1070 2 . . . 32505 1071 1 Weather weather NN 32505 1071 2 and and CC 32505 1071 3 climate climate NN 32505 1071 4 of of IN 32505 1071 5 México México NNP 32505 1071 6 and and CC 32505 1071 7 Central Central NNP 32505 1071 8 America America NNP 32505 1071 9 . . . 32505 1072 1 _ _ NNP 32505 1072 2 In in IN 32505 1072 3 _ _ NNP 32505 1072 4 R. R. NNP 32505 1072 5 Wauchope Wauchope NNP 32505 1072 6 and and CC 32505 1072 7 R. R. NNP 32505 1072 8 C. C. NNP 32505 1072 9 West West NNP 32505 1072 10 ( ( -LRB- 32505 1072 11 Eds Eds NNP 32505 1072 12 . . . 32505 1073 1 ) ) -RRB- 32505 1073 2 , , , 32505 1073 3 Handbook Handbook NNP 32505 1073 4 of of IN 32505 1073 5 Middle Middle NNP 32505 1073 6 American American NNP 32505 1073 7 Indians Indians NNPS 32505 1073 8 . . . 32505 1074 1 Vol Vol NNP 32505 1074 2 . . . 32505 1075 1 1 1 LS 32505 1075 2 , , , 32505 1075 3 Univ Univ NNP 32505 1075 4 . . . 32505 1076 1 Texas Texas NNP 32505 1076 2 Press Press NNP 32505 1076 3 , , , 32505 1076 4 Austin Austin NNP 32505 1076 5 , , , 32505 1076 6 570 570 CD 32505 1076 7 pp pp NN 32505 1076 8 . . . 32505 1077 1 _ _ NNP 32505 1077 2 Transmitted Transmitted NNP 32505 1077 3 February February NNP 32505 1077 4 7 7 CD 32505 1077 5 , , , 32505 1077 6 1969 1969 CD 32505 1077 7 . . . 32505 1077 8 _ _ NNP