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TYPICAL FORMS
AND
SPECIAL ENDS IN CREATION
ET
REV. JAMES M'COSH, LL.D.,
PKOFE880K OF LOGIC AND METAPHYSICS IN THE QUEEN'S UNIVEB8ITY IN IRELAND;
AUTHOB OF "TOE METHOD OF TIIE DIVINE GOVEBNMENT,
PHYSICAL AND MOEAL," ETC. ;
AND
GEORGE DICKIE, A.M., M.D.,
raOFESSOK OF NATURAL HISTORY IN TOE QUEEN'S UNIVERSITY IN IRELAND ; AND
AUTHOB OP A NUMliEB OF PAPERS ON ZOOLOGY AND ItOTANY.
TYIIOZ KAI TEAOS.
NEW YOEK:
ROBERT CARTER & BROTHERS,
No. 530 BEOADWAY.
1887.
STEREOTYPED BT
THOMAS B. SMITH,
82 & fc Beekman St., N. Y.
PRINTED B»
£. O. JENKINS,
ADVERTISEMENT
THE AMERICAN PUBLISHER.
The principles now fully explained and illustrated in this work
were first brought before the public in an article on Typical
Forms by Dr. McCosh in the " North British Review" for August
1851. Mr. Hugh Miller wrote a lengthened notice of that
article, describing it as :
" An article at once the most suggestive and ingenious which we have
almost ever perused. The typology of Scripture has formed the subject of
many a volume and many a discourse. It is one of the most obvious and
rudimental truths of the theologian, that he who spoke in parable and
allegory when he walked the earth in the flesh, spoke in his previous rev-
elation ere he had yet put on the nature of man, by type and symbol; and
that there is thus a palpable unity of style maintained between God in
the Old and God in the New Testament. Nay, some of the profounder
theologians went further than this ; and works such as the " Analogy" of
Butler may be regarded in one point of view as critical Essays, written to
establish a yet further identity between the style of Deity in Revelation
and in Nature. "All things are double one against another," said the
wise son of Sirach ; and the celebrated " Treatise" of the most philosophic
of English bishops maybe deemed simply an expansion of the idea. Butler
set himself to seek in the natural world the " double" of the revelations of
the spiritual one, and to argue from the existence and fitness of the natural
type the authenticity and genuineness of the spiritual anti-type. Such, in
short, seems to be the principle of his "Analogy." It has, however, been
reserved for our own times, and hitherto at least for a class of men not
much disposed to conciliate the assertors of the popular theology, whether
at home or abroad — in Protestant or in Popish countries — to find in Nature
analogies which, though they themselves have failed to apply them, seem
to reach further than even those of Butler ; and which, we can have little
doubt, will at no distant date form the staple facts of a department of
theology still very meagerly represented in our literature, and intermediate
in its place and character between the Natural Theology of the Philoso-
ADVERTISEMENT.
phers and the Dogmatic Theology of the Divines. The article in the " North
British" on Typical Forms is a vigorous contribution to this middle depart-
ment of theology, which, like a central area left unbuilt in a street after
the completion of the erections on bpth sides, seems so necessary to the
union of the contiguous fabrics, and to the design of the whole ; and all
that its perusal leaves us to regret is, that its accomplished author, in
whom the reader will, we believe, recognize a most original thinker — a
man already well known in the ethical field, both in our own country and
America — should not have expanded it into a volume. But in the special
field which he has chosen he need not greatly fear a competitor. The sub-
ject is one, too, on which thought ripens slowly ; for, like the agricultural
produce of a new colony, it has all to be raised from the seed ; and the
deeply interesting, but, comparatively brief article of the reviewer, will, we
can not doubt, be yet expanded into a separate treatise, which will prove
none the less fresh, and all the more solid, from the circumstance that it
should have appeared as an article first."
Since the time when the article referred to was written, Dr.
McCosh, in conjunction with Dr. Dickie, has been prosecuting
the subject, and the two have laid a number of their scientific
observations before various learned societies, such as the Bo-
tanical Society of Edinburg, the Natural History Society of
Belfast, and the British Association for the Promotion of Science
at its meetings in 1852 and 1854. Summaries of these have
appeared in the Transactions of the Botanical Society of Edin-
burgh in the Annals of Natural History, in the proceedings of
the British Association, and in the E [rabiirgh New Philosophi-
cal Journal. They were referred to by his Grace the Duke
of Argyle, the President of the British Association, in his open-
ing address in September last in the following language:
" In physiology, what is the meaning of that great law of adherence to
type and pattern, standing behind, as it were, and in reserve, of that other
law by which organic structures are specially adapted to special modes of
life ? What is the relation between these two laws ; and can any light be
cast upon it derived from the history of extinct forms, or from the condi-
tions to which we find that existing forms are subject ? In vegetable phy-
siology do the same or similar laws prevail, or can we trace others, such as
these on the relations between structure, form, and color, of which clear
indications have already been established in communications lately made
»o this Association by Dr. McCosh and Dr. Dickie of Belfast."
CONTENTS,
BOOK FIRST.
PRINCIPLES OF GENERAL ORDER AND SPECIAL ADAPTATION.
CHAPTER I.
NATURE OF THE ORDER PREVAILING IN THE MATERIAL WORLD.
PAGH
Sect. I. — Principles which seem to run through the Structure of
the Cosmos, ...... 1
Sect. II. — Analysis of the Order in Nature, ... 10
CHAPTER II.
nature of the special adaptations in the material world.
Sect. I. — Need of Special Adjustments in order to tho Beneficent
Operation of the Forces of Nature, ... 30
Sect. II. — The Adjustments are designed, and not Casual. — Na-
ture of Chance, ..... 39
Sect. III. — The Obviousness and Completeness of the Special
Adaptations, ...... 56
BOOK SECOND.
CO-ORDINATED SERIES OF FACTS, GIVING INDICATIONS OF COM-
BINED ORDER AND ADAPTATION THROUGHOUT THE
VARIOUS KINGDOMS OF NATURE.
CHAPTER I.
toe minute structure of plants and animals.
Sect. I. — Order in the Structure of the Cell, . . . 69
Sect. II. — Special Modifications of the Coll, . . • ^ 1
VI CONTENTS.
CHAPTER II.
THE FORMS OF PLANTS.
(Ml
Sect. I. — Traces of Order in the Organs of Plants, . . 81
Sect. II. — Traces of Special Adaptation in the Organs of the Plant, 130
CHAPTER III.
THE COLOURS OF PLANTS.
Sect. I.— The Relation of Form and Colour in the Flower, . 146
Sect. II. — Adaptation of the Colours of Plants to the Natural
Tastes of Man, ..... 152
CHAPTER IV.
THE VERTEBRATE SKELETON.
Sect. I. — The Homologies and Homotypes of the Vertebrate
Skeleton, ...... 175
Sect. II. — Special Adaptations in the Structure of the Skeleton, . 192
CHAPTER V.
TEETH.
Sect. I. — Order in the Number, Form, and Structure of Teeth, . 213
Sect. II. — Special Adaptations in the Number, Form, and Struc-
ture of Teeth, ..... 215
CHAPTER VI.
MOLLUSCA.
Sect. I. — Typical Forms of Mollusca, .... 223
Sect. II. — Modifications of the Archetype Mollusc, . . 227
CHAPTER VII.
ARTICTJLATA.
Sect. I. — Homotypal Rings and Appendages, . . . 233
Sect. IT. — Special Modifications of Rings and Appendages, . 237
CONTENTS. VU
CHAPTER VIII.
PAQB
Sect. I. — Typical Forms of Radiata, .... 261
Sect. II. — Adaptation of Radiate Types to Mode of Life, . . 211
CHAPTER IX.
Nervous, vascular, and muscular systems, . . . 280
CHAPTER X.
Community of plan, with special modifications, in the
development of organized beings, . . . 299
CHAPTER XI.
GEOLOGY.
Sect. I. — Traces of Plan in Fossil Remains, . . . 309
Sect. II. — Adaptations of Fossil Organisms to their Functions. —
Preparations for Man, . . . .333
CHAPTER XII.
INORGANIC OBJECTS ON THE EARTH'S SURFACE.
Sect. I. — Crystalline Forms and Chemical Proportions, . . 354
Sect. II. — Adaptations of Inorganic Objects to Animals and Plants
— Physical Geography, . ' . . . 369
CHAPTER XIII.
THE HEAVENS.
Sect. I. — Order in the Movements of the Heavenly Bodies, . 388
Sect. II. — Special Adjustments needed in order to the Harmony
of Cosmical Bodies, 401
Vlil CONTENTS.
BOOK THIRD.
THE INTERPRETATION OF THE FACTS.
CHAPTER I.
PACK
The argument from combined order and adaptation, . 420
CHAPTER II.
CORRESPONDENCE between the laws op the material world and the
FACULTIES OF THE HUMAN MIND.
Sect. I. — The Fantasy, or Imaging Power of the Mind, . . 440
Sect. II. — The Faculties which discover Relations (Correlative), . 449
Sect. III. — The Association of Ideas, .... 473
Sect. IV. — The ^Esthetic Sentiments, .... 481
Supplementary Section. — Brief Historical and Critical Review
of the Theories of the Continental Philosophers as to the Re-
lation between the Laws of the Internal and External Worlds, . 492
CHAPTER III.
TYPICAL SYSTEMS OF NATURE AND REVELATION.
Sect. I.— Old Testament Types, ..... 504
Sect. II. — Typical Numbers of Scripture, . . .518
Sect. III. — Typical System of the New Testament, . . 525
APPENDIX.
Selected List of Plants, illustrating Associations of Colour, and the
Relati ns of Form and Colour, .... 533
Tndex, ........ 537
BOOK FIRST,
CHAPTER I.
NATURE OF THE ORDER PREVAILING IN THE MATERIAL
WORLD.
SECT. I. PRINCIPLES WHICH SEEM TO RUN THROUGH
THE STRUCTURE OF THE COSMOS.
In taking an enlarged view of the constitution of the
material universe, so far as it falls under our notice, it
may be discovered that attention, at once extensive and
minute, is paid to two great principles or methods of
procedure. The one is the Principle of Order, or a
General Plan, Pattern, or Type, to which every given
object is made to conform with more or less precision.
The other is the Principle of Special Adaptation, or
Particular End, by which each object, while constructed
after a general model, is, at the same time, accommo-
dated to the situation which it has to occupy, and # pur-
pose which it is intended to serve. These two principles
are exhibited in not a few inorganic objects, and they
meet in the structure of every plant and every animal.
These two principles are characteristic of intelli-
gence ; they must proceed from intelligence, and they are
addressed to intelligence. They may both be discovered,
though necessarily to a limited extent, in human work-
1
1 PRINCIPLES RUNNING THROUGH
manship. When circumstances admit, man delights to
construct the instruments or utensils which are designed
to serve a common purpose after a common plan, even
when this is by no means essential to the immediate
purpose to be served. Each particular piece of dress or
article of furniture in a country is commonly fashioned
after some general model, so that we are able to guess
its use as soon as we cast our eyes upon it. That there
is so much of this figure no way fitted to accomplish a
special end, is evident from the circumstance that articles
serving the same purpose take — in different ages and
nations, and according to the fashion of the place or time
— somewhat different forms, all of which are equally
convenient. The farmer builds up his grain in stacks,
which have all a like contour, and the merchant packs
his goods in vessels of equal size and similar shape, or
disposes of them in bales of equal weight. It is only
when his possessions are so arranged that man can be
said to have the command of them. Were his property
not so disposed, were his grain gathered into heaps of all
sizes and shapes, were his merchandise scattered in every
corner of the apartment, the possessor would become
bewildered in proportion to the profusion and variety of
his wealth. When things are formed or arranged on
some plan tacitly agreed on, man can recognize every
object at a distance by its physiognomy, and determine
its nature and its end without seeing it in use or ope-
ration.
There are still more frequent and obvious examples
in the works of man of the principle of special adap-
tation. While there is a general regard, so far as it can
be done without immediate inconvenience to the prin-
ciple of order, there is a far more constant attention to
the other principle. In some cases, indeed, little respect
THE MATERIAL WOULD. 6
can be had to the general model ; the sole end aimed at
is the fitting of the instrument to the purpose which it
is meant to serve. In nations low in the scale of civili-
zation, and among persons who have to engage in a hard
struggle to procure the necessaries of life, the general
order is apt to be neglected in the exclusive regard
which must be had to immediate utility. In such cir-
cumstances, individuals care little how an article he con-
structed, provided it serves its practical purpose. But
as man's industrial treasures increase, and the number
of separate works intended to accomplish similar ends
are multiplied, he finds it becoming to institute some
systematic arrangement among them, or devise some
pattern after which to fashion them.
When hard necessity does not forbid, man feels a
pleasure in constructing his works upon a general plan.
Human intelligence delights to employ itself in forming
such models. They seem to have a beauty to the eye,
or rather to the mind, which contemplates them. If it
is a basket that is to be woven, there will commonly be a
regularity in the succession of the plaits, and an aiming
after some ideal form in the shape of the whole. If it
is a water-jug that is to be fashioned, there will be a
general attention paid to symmetry ; not unfrequently
there will be graceful and waving lines in the figure
which strikes the eye. The dwelling which the indi-
vidual erects for his own special accommodation, will
commonly be found to have a door, or some other prom-
inent object, in the center, with a balancing of pillars,
windows, or something else that fixes the attention, on
the one side and the other. As man advances in the
scale of civilization, and comes to have superfluous wealth
and leisure, he pays an increasing attention to symmetry
and ornament. In the urns which he makes to receive.
4 PRINCIPLES RUNNING THROUGH
the ashes of the dead, in. the temples erected by hiui in
honor of the God whom he worships, there is a scrupu-
lous regard had to proportion and outline. As wealth
accumulates and taste is cultivated, the law of order and
ornament comes to he valued for its own sake, and is
followed in the construction of every house, and of every
article of furniture in that house, in the setting of every
jewel, and in the location of every ornament.
In most articles of human workmanship we may dis-
cover a greater or less attention to hoth of the principles
to which we have referred. The farmer's stacks are all
formed after a general mould, but we may observe a
departure from it on either side to suit the quantity or
quality of the grain. The merchant's shop seems to be
regulated by forms or weights, but there is special form
or average weight for every separate article. In some
objects we see a greater regard to general plan, and in
others to special purposes, and this according as persons
wish to give a greater prominence at the time to orna-
ment or to utility.
Now, if this world proceeds from intelligence, and if
it is intended to be contemplated by intelligence, it is
surely not unreasonable to suppose that there may be
traces in it of the same two modes of procedure. In this
treatise we hope to be able to show that there are abun-
dant illustrations of both, by an induction reaching over
all the kingdoms of nature, and extending even into the
kingdoms of grace. Both will be found in the theology
of nature to point to the1 same conclusion ; each furnishes
its appropriate proof of the existence and wisdom of a
Being who hath constructed every thing on a plan, and
made it, at the same time, to serve a purpose. The one,
as well as the other, will be found in the dispensations
of God, in the kingdom of his Son, and point to a most
THE MATERIAL WORLD. 5
interesting analogy between nature and revelation. It
will be expedient to treat of them as so far different,
which they really are, but it will be necessary, at the
same time, to show, what is equally true, that the two
principles are made to correspond the one to the other,
that they meet in a higher unity, and that, after all, they
are but two aspects — in many respects different indeed —
of one Great Truth.*
In certain sections of this treatise it is proposed to
unfold some of the more striking examples of General
Plan. In respect of this order of facts, natural theology
can now take a step in advance, in consequence of what
has been done of late years in the discovery of homologies
by the sciences of comparative anatomy and morpho-
logical botany. But the recent discoveries in regard to
the homology of parts can never set aside the old doctrine
of the teleology of parts, which affirms that every organ
is adapted to a special end. Every organic object is con-
structed after a type, (rtnoc,) and is, at the same time,
* In order to remove misapprehension, it may be necessary hero to estimate how much
truth there is in a statement of Professor Owen, who has done so much to illustrate the
subject of general order. " By whatever means or instruments man aids or supersedes
his natural locomotive organs, such instruments are adapted expressly and immediately
to the cud proposed. He does not fetter himself by the trammels of any common type
of locomotive instrument, and increase his pains by having to adjust the parts and com-
pensate their proportions so as best to perform the end required without deviating from
the pattern previously laid down for all. There is no community of plan or structure
between the boat and the balloon, between Stephenson's engine and Brunei's tunnelling
machinery ; a very remote analogy, if any, can be traced between the instruments de-
vised by man to travel in the air and on the sea, through the earth or along its surface."
(Owen on the Nature of Limbs, p. 9.) There is truth in the remark here made, but it
seems to us to be overstated, and without the necessary corrections. Man does, in many
cases, construct the works which arc to serve a common end upon a common plan.
There is a model structure for the boat, for the steam-engine, for our houses, and our
temples, in which elegance is more or less attended to. But still it is to be admitted
that the harmonies, the correspondences, the compensations, are far more numerous and
beautiful, both in kind and degree, in the works of God than in the works of man. It
is certain that the union of the two principles is not so frequently attended to in human
•s in Divine workmanship. Man is often obliged to sacrifice the one to the other, the
symmetry to the convenience, or the utility to the ornament. It is only in the works
of Deity that we find the two at all tinies in harmonious operation.
6 PKINCIPLES RUNNING THROUGH
made to accomplish a final cause, (lilog.) Throughout
the next Book we purpose to exhibit the traces of General
Order in one series of sections, and the traces of Special
Adaptation in another series of sections, the two being
made to run alongside of each other. While both
will be illustrated, it will be seen, by our adopting this
method, that the two are not contradictory, but coin-
cident ; that they do not cross, but run parallel to each
other. The general conformity to a pattern will be seen
to be all the more curious when contemplated in con-
nection with certain singular deviations ; while the special
modifications will appear all the more wonderful when
exhibited as a departure, and evidently an intentional
departure, to effect a particular end, from a model usually
attended to, nay, to some extent attended to, it may be,
in the very structure which is thus modified. The de-
signed irregularities will thus, by a legitimate reaction,
show that the regularities are also designed ; the excep-
tions in this case emphatically prove the rule. The
nature of the eccentricities demonstrate that, after all,
there is a center round which the revolution is per-
formed ; the deviations point to a disturbing influence
also under the influence of law — in much the same way
as the deviations of an old planet were shown by living
astronomers to point to a previously undiscovered plane-
tary body. The nature, the value, and the relation of the
two principles, will thus come out to view more strikingly
by comparison and contrast when they are 2^1aced in
juxtaposition.
The arguments and illustrations adduced by British
writers for the last age or two in behalf of the Divine
existence, have been taken almost exclusively from the
indications in nature of special adaptation of parts.
Hence, when traces were discovered within the last age
THE MATEKIAL WORLD. 7
of a general pattern, which had no reference to the com-
fort of the animal or the functions of the particular plant,
the discoveiy was represented by some as overturnino;
the whole doctrine of final cause ; not a few viewed the
new doctrine with suspicion or alarm, as seemingly
adverse to religion, while the great body of scientific
men did not know what to make of its religious import.
The question is thus started, Have not the writers on the
theology of nature been of late most unnecessarily nar-
rowing and restricting the argument ? We have found
it most interesting to notice that the philosophers of
ancient Greece and Rome, and not a few of the earlier
writers on the subject in our own country, gave it a
much wider range, and reckoned that they had found
evidence of the existence of God whenever they detected
traces of order and ornament. Let us inquire what
instruction we can gather on this subject from some of
those great luminaries of the ancient world, which, like
stars, send their light down to us through the wide space
which intervenes, and serve, like them, to enlarge and
rectify our ideas of magnitude, and to keep us from
being unduly impressed with the greatness of the near
and the present.
Plato, in the Fourth Book of the Laws, makes Clinias
of Crete, in proving the existence of God from his works,
appeal at once to the order and beauty of the universe,
and does not regard it as at all necessary to dwell on
minute instances of adaptation. He refers to the earth,
the sun, and all the stars, and to the beautiful arrange-
ment of the seasons, divided into months and years, as
evidencing that there is a Divine Being.'"' In the review
of the argument in the Twelfth Book, he repeats, that
the orderly movements of the stars, and other objects,
* B. x. c. 9, where be also brings in the argument from universal consent.
8 PRINCIPLES RUNNING THROUGH
prove that all things were arranged and adorned, not
by mutter or necessity, but according to a Divine fore-
thought and will.* According to the sublime philosophy
of Plato, all things are formed according to unalterable
laws or types, which remain unchanged amidst the flux
of individual objects, and that because they proceed from
eternal ideas, which had been in or before the Divine
mind from all eternity.
A similar style of argument is adopted in Cicero's
Treatise on the Nature of the Gods, the most systematic
work on natural theology which has been handed down
to us from ancient times. The evidence adduced by
Balbus the Stoic, the representative of theism in the
dialogue by which the argument is conducted, is derived
from four sources : first, from the presages of futurity by
gifted men and oracles ; secondly, from the number of
things fit and useful ; thirdly, from prodigies ; fourthly,
and highest of all, from the equable motions of the
heavenly bodies, and from the beauty and order of the
sun, moon, and stars, of which the very sight is sufficient
to convince us that they are not fortuitous.-}" Through-
out his defence, he dwells on the consenting and conspir-
ing motions of the heavenly bodies, on their progressions
and other movements, all constant and according to law ;
he points to the planets, which are regular in their very
wanderings ; and shews how, in all this, there is an order
and a certain likeness to art.J When one observes, he
says, their defined and equable motions, and all things
proceeding in an appointed order, and by a regulated and
unchangeable constancy, he is led to understand not only
that there is an inhabitant in this celestial and divine
dwelling, but a ruler or regulator, and, if we may so
* B. xiii. c. 13.
t Cic. De Nat. Deor., Lib. II. c v. t Lib. 11. c. vil. ; xx. ; xxxll.
THE MATERIAL WORLD. 9
Speak, architect of so great a work and gift.* He speaks
of the harmony arising from dissimilar motions ; and
after quoting largely from the hymn of Aratus, he says,
such order and ornament could not have proceeded from
bodies running together hither and thither, and by
accident.f
Plutarch derives men's general agreement as to the
existence of God, from their observation of the constant
order and motion of the stars.J
In modern times, we have the same line of argument
seized by the profound mind of Newton. Keferring to
the UNIFORMITY IN THE BODIES OF ANIMALS, he Says, " It
must necessarily be confessed that it has been effected
by intelligence and counsel."§ Dr. Samuel Clarke quotes
this language, and asks — " In all the greater species of
animals, where was the necessity for the conformity we
observe in the Number and Likeness of all their prin-
cipal members ?"||
It is very evident that, down to a comparatively late
date, writers on natural theism did not confine their
proof to a mere adaptation of parts, but that along with
this they introduced other considerations, and in parti-
cular, the prevalence of general order. It will not be
difficult to defend the legitimacy of the conviction which
the order and beauty of the universe have produced in un-
sophisticated minds in all ages. In this, as in many other
instances, the philosopher will find it to be his delightful
office, not to set aside the spontaneous beliefs of mankind,
but rather to vindicate and illustrate them by the new
discoveries which advancing science is ever opening.
* Cic. De Nat. Deor., Lib. ii. c. xxsv. t Lib. iL c. xliv.
% Plut. De Plac. i. 6. § Optics.
| Demonstration of Being and Attributes of God.
1*
10 ANALYSIS OF THE
SECT. II. ANALYSIS OF THE ORDER IN" NATURE LAWS
OP NATURE.
The most careless observer is led to notice, that there
is a beautiful regularity running through nature as a
whole, and through every individual part of it. This
was discovered in very early ages of the world's history,
by persons who had no very precise ideas as to its nature,
or the means by which it was produced. The Greeks,
from the time of Pythagoras, embodied their impressions
in the word by which they denoted the visible world,
which they called Cosmos, to denote at once its order
and its beauty, while the Latins styled the world Mundus,
to express their sense of its surpassing loveliness. Ever
since the time when the philosophic spirit was first
awakened, reflecting minds have been speculating as to
the sources of this order, and caught, at a very early age,
glimpses of the truth. The philosophers of the Ionian
School, which arose between 600 B.C. and 500 B.C., re-
ferred it to the power and the varied transformations of
certain elements, which they did their best to classify, as
air, water, earth, and fire, representing the dry, the moist,
the solid, the ethereal. In the speculations of this school,
we have vague anticipations of modern chemistry, and in
particular, of the doctrine of polar forces, in the balanced
strife and friendships of Empedocles, and of that of de-
finite proportions, in the " homoiomera" or equal parts of
Anaxagoras. A rival school arose at a little later date,
among the Greeks in Italy, and ascribed the order of
nature, in a more profound spirit, to the power of Num-
bers. We have no authentic or connected account of
the system of the Pythagoreans, but it is evident, from
the scattered notices which have been handed clown to
us, that they represented numbers, the significance of
ORDER IN NATURE. H
which is so clearly seen in music, as in some mysterious
sense the principia of the universe. Aristotle tells us,
that they considered existing things to be a copy of num-
bers,* and we have extracts preserved from the writings
of some of the disciples of the school, describing numbers
as being in the Divine Mind prior to the existence of
things, as being used as a model (naQiideiy[<<*) in the
formation of objects, and as that by which all things
were brought together and linked in order. Among the
disciples of the same school, and others who arose at a
subsequent date, there was supposed to be a deep mean-
ing in forms ; and the properties of certain figures, such
as the triangle, the square, the parallelogram, the circle,
the ellipse, were investigated with great care, giving us
the science of geometry as the result. A very special in-
terest gathered round certain numbers, such as seven and
ten, and certain figures, such as the circle and triangle,
which came in consequence to be regarded as perfect, or
as sacred. From a still earlier date, and as a manifestation
of the same intellectual propensity, peculiar feelings be-
came associated with certain recurring times and perio-
dical seasons, such as the revolutions of the moon, the
signs of the zodiac, and other cycles, which seemed to
have a deep significancy in the economy of nature. De-
mocritus, who lived 400 B.C., and the Epicureans, who
flourished at a later date, sought for the origin of this
order in the formation of all things out of atoms possessed
of definite forms. The sublime genius of Plato ascribed
it to certain patterns after which all things were fash-
ioned, which patterns he traced back to the eternal ideas
of the Divine Mind. Aristotle, while correcting some of
the extravagances Of his great master, clung resolutely to
the doctrine, that forms were as necessary as matter to
* Mifiqaiv c'lvai ra Svta rdv apiO/taiVi — Metaph. of Aria.
12 ANALYSIS OF THE
the construction of the universe. The Platonists of the
Alexandrian School literally revelled among numbers
and forms, till they lost themselves among their intrica-
cies and windings. The Platonizing Jew who wrote the
Book of Wisdom, caught for a moment a very clear
glimpse of the full truth, when he speaks of God
" having arranged all things in measure, number, and
weight."®
Early science, like youth, is ardent, is eager, and not
having as yet determined either its strength or its weak-
ness, it would attempt every work, and works far beyond
its capacity. Like the giants of the early world, it is
ambitious, and would heap Ossa on Pelion, and mount
to heaven, not by gradual and numerous steps, but by
one b£# bold and presumptuous effort. In following this
method of speculation, the sage — as he meditates on
the banks of the Euphrates or Nile, along which an
early civilisation had sprang up, or in the cities of
Miletus, Elea, or Athens, in which the human spirit
was sharpened by discussion and the love of enterprise
— makes many a shrewd guess ; he anticipates not a
few truths which later discovery confirms ; he awakens a
spirit of inquiry which craves for a more accurate mode
of procedure ; and if he does not settle, he at least starts
questions which must sooner or later be settled. But
his attempt, though characterized by enlargement of
vision and power of vaticination, is, in respect of scien-
tific strictness and certainty of result, a failure, and the
favourite dogma of one school is ever disputed by the
disciples of another school. It turns out that the work
which one man or one school has attempted, needs, in
order to its completion, the combined industry of many
investigators continued through long successive ages.
* lidvra fierpto Kal apifyico Kai arafyoj Jiardjaj.
ORDER IN NATURE. 13
For just as when society makes progress there is a neces-
sity for the division of manual labour, (as Adam Smith
has shewn in the opening chapter of the Wealth of Na-
tions,) so, in order to the advance of science, there is need
of a division of intellectual labour. Most important of
all, there arises, in the midst of the jealousies of rival
schools and the noise of fruitless disputations, a demand
for a surer, even though it should be a slower, method of
investigation, — a method which will give results, be they
many or be they few, which are not of the nature of in-
genious speculations, to be set aside by other ingenious
speculations, but ascertained truths, fixed for ever, and
which all inquirers who come after may use, to help them
to add to the accumulating stores of knowledge. It is
late in the history of the world before such a plan comes
to be systematically unfolded ; and it is to the glory of
our country, a glory not exceeded even by that of the
land which produced Plato and Aristotle, that the first
exposition of it was by Lord Bacon. Since his days,
scientific inquirers, according to their tastes, talents, and
position, have betaken them each to his own field of in-
vestigation, with the view of thoroughly exploring it ;
and as the grand result, we have a settled body of truth,
to which additions will be made from age to age.
But as the deeply-underlying and prompting cause of
all this intellectual activity, there is still the same crav-
ing desire to find out the means by which unity and
order are given to the great Cosmos. In these days we
speak of all tilings being governed by laws ; we lay it
down as a maxim, that the end of all science is the dis-
covery of law. The language may be more correct than
that employed by the ancients, but it is far from being
definite or incapable of misinterpretation. For the ques-
tion occurs, What is meant by laws in this application
14 ANALYSIS OF THE
of the term ? Every one sees that, as thus used, it does
not mean the same thing as when we speak of the laws
of a country, of the moral law, or of the law of Grod; It
is a term with which we cannot dispense, but it is far
from being unambiguous ; it is often used in an unlawful
sense, and at times it is turned to the worst of purposes,
as when it is supposed, that in referring an event to a
law of nature, we have placed it beyond the dominion of
God. When we speak of things being arranged in a
law, or falling out according to a law, we signify, if we
know what we mean, that all phenomena take place in
a regular manner, that is, according to a rule.'* It is
the special office of each science to discover what the
nature of the law is in its own department. This is the
grand aim, so far as it has a grand aim, of all modern
physical investigation, — to determine the rule to which
the particular classes of objects under contemplation
accommodate themselves. But in very proportion as
the sciences have become subdivided and narrowed to
particular facts, is there a desire waxing stronger, among
minds of larger view, to have the light which they have
scattered collected into a focus. As the special sciences
advance, the old question, which has been from the be-
ginning, will anew and anew be started, — What is the
general meaning of the laws which reign throughout the
visible world ? A correct and adequate answer to this
wide question can be given only by a wide induction,
and a combination of the results gained by a vast number
of separate sciences, each conducted on its own principles.
We live in the expectation of the approach of a time
when science — the division of labour having fulfilled its
ends — shall seek to combine its individual truths, and to
* Seo a more minute analysis of the laws of nature in the Method of the Divine Govern-
ment, Physical and Moral, B. ii. o. t
ORDER IN NATURE. 15
realize the dream of its youth, and, as it were, carry us
to a mountain top, whence we may obtain not only a
scattered view of the separate parts, but a connected view
of the whole, and of the relative bearing and direction of
every part. It appears to us that we are approaching
the time when an answer may be given to the old ques-
tion, and that this must be something like the following :
— All things in this world are subordinated to law, and
this law is just the order established in nature by Him
who made nature, and is an order in respect of such
qualities as number, time, colour, and form. We use
the vague languages of such qualities, because science has
not arrived at such a stage as to enable it to determine
what these qualities are with anything like perfect cer-
tainty and precision.*
Every law of nature which can be said to be correctly
ascertained is certainly of this description. We shall
furnish abundant illustrations in the next Book of this
treatise ; in this section we are merely to collect a few
striking examples of the attention paid to each of the
qualities named, and thus prepare the way for entering
upon the separate sciences, when more systematic proof
will be offered.
First, There is an Order in Nature in Respect
of Number.— This important truth, long believed in
before it could plead any scientific evidence in its favour,
was established and brought into prominence when
Kepler unfolded the three laws which have formed, his-
torically, the foundation of modern astronomy. It was
* A more scientific classification would probably give us active property instead of
colour, and including colour. There is a curious combination of active properties con-
stituting individual objects, and enabling us to classify them, which will be referred to
In B. iii. c. 1 & 2, but which cannot be fully cleared up till wc know more of the Ifttto5
forces of nature.
16 ORDER IN RESPECT
the confident expectation that there would be found
some such principle of order which led that ingenious
and persevering sage to make calculation upon calcula-
tion, and devise one hypothesis after another, till, after
nineteen unsuccessful attempts, his fine genius and his
industry . were rewarded by the discovery of the true
laws of the planetary movements. These laws are, —
that the planets move in orbits, which are elliptical in
shape ; that if you draw a line from the planet to the
sun, the areas described by that line in its motion round
the sun are proportional to the times employed in the
motion ; and that the squares of the periodic times are as
the cubes of the distances. . The first of these is a law of
forms, the other two are laws of numbers. The dis-
coveries of Kepler prepared the way for the still more
important ones of Sir Isaac Newton. When the immor-
tal work of this greatest of inductive philosophers was
published, it was seen that the laws of Kepler were
not original but derivative ; but the original law now
unfolded belonged to the same class ; for the law of
gravitation, the best established and the most univer-
sally operative law yet determined, is a law of numbers.
Turning to chemistry, we find that ever since it emerged
•as a science there has been a constantly renewed attempt
to reduce its laws to a numerical expression. The only
laws which can be reckoned as certainly determined
in this science possess this character. The great law
which lies at the basis of all the compositions and
decompositions of substances, is that of definite propor-
tions for equivalents, as expounded by Dalton. In the
same science Gay Lussac discovered an arithmetical law,
regulating the combination of gaseous substances, which
unite in very simple proportions, according to volumes.
Lest it should be thought that we are making a fanciful
OF NUMBER. 17
reduction of the operations of nature, \vc are happy to be
able to bring to our aid the name of Sir John Herschel.
" Chemistry/' says he, " is, in a most pre-eminent degree,
a science of quantity, and to enumerate the discoveries
which have risen from it from the mere determination
of weights and measures, would be nearly to give a
synopsis of this branch of knowledge. We need only
mention the law of definite proportions which fixes the
composition of every body in nature in determinate pro-
portional weights of its ingredients. Indeed, it is a
character of all the higher laws of nature to assume the
form of a precise quantitative statement. Thus the law
of gravitation, the most universal truth at which the
human reason has yet arrived, expresses not merely the
general fact of the mutual attraction of all matter, not
merely the vague statement that its influence decreases
as the distance increases, but the exact numerical rate
at which that increase takes place, so that when the
amount is known at any one distance it may be calcu-
lated exactly for any other."* Similar language is used by
Humboldt : — " The progress of modern physical science
is especially characterized by the attainment and the
rectification of the mean values of certain quantities by
means of the processes of weighing and measuring.
And it may be said that the only remaining and widely
diffused hieroglyphic characters still in our writing —
numbers, appear to us again as powers of the cosmos,
although in a wider sense than that applied to them by
the Italian school."f
In looking at other departments of nature, we find
similar examples of numerical order. Thus, ten is the
typical number of the fingers and toes of man, and,
»
* Herschel's Natural Philosophy, Art. 116.
t Cosmos, translated by Otto, vol. 1. p. 64.
18 ORDER IN RESPECT
indeed, of the digits of all vertebrate animals. It is
also a curious, though perhaps not very significant
circumstance, that in mammalia seven is the number of
vertebrae in the neck,* and this whether it be long as
in the giraffe, or short as in the elephant, whether it be
flexible as in the camel, or firm as in the whale. In
the vegetable kingdom we find that two is the prevailing
number in the lowest division of plants, the acrogenous
or flowerless ; thus, 2, 4, 8, 16, 32, 64, &c, are the num-
ber of teeth in the mouth of the capsule in mosses.
Three, or multiples of three, is the typical number of
the next class of plants, the monocotyledonous or endo-
genous ; and five, with its multiples, is the prevailing
number in the highest class, the dicotyledonous or exo-
genous plants. We shall shew, as we advance, that a
curious series, 1, 2, 3, 5, 8, 13, 21, 34, &c, in which any
two numbers added together give the succeeding one,
regulates the arrangement of the leaf appendages of
plants generally, and in particular of the leaves and the
scales on the cones of firs and pines. In the inflores-
cence of the plant we find that the outer organs, or
sepals, always alternate with the petals which are next
them, and that the whorl of organs further in, namely,
the stamens, is generally either the same in number as
the petals, or some multiple of them. When there is
an exception to this rule there is reason to believe that
there has been some abortion of the stamens ; and the
traces of this abortion are not unfrequently visible in the
rudiments of the organs undeveloped.
Secondly, There is an Order in Nature in respect
of Time. — It is obvious that all such laws can be ex-
pressed in proportional numbers, taking some fixed time
* Apparent or real exceptions will be referred to afterwards.
OF NUMBER AND TIME. 19
as a unit. But we are here introduced to a new funda-
mental power, deserving of being put under a separate
head. For the laws of which we are now to speak imply
a peculiar arrangement in reference to time. We see
the principle most strikingly exhibited in those move-
ments of natural objects which are periodical. No doubt,
there is some disposition of physical forces necessary to
produce this periodicity ; but this just shews all the
more clearly that an arrangement has been made to
produce the regularity. The ancients were much struck
with the order in respect of time of the celestial motions.
The stars, the planets, and even the comets, wero seen to
perform their revolutions in certain fixed times. Some
of them seem to depart from this rule only to exemplify
it the more' strikingly, for their irregularities, which
are periodical, are as methodical as their more uniform
movements. There have been regular epochs, to all
appearance, in the changes on the earth's surface, and in
the succession of plants and animals, as disclosed by
geological science. The variations of magnetism on the
earth's surface seem to be periodical, and attempts have
been made of late to connect this cycle with that which
the spots of the sun are known to follow. There is a
beautiful progression, as shewn by the science of embry-
ology in the growth of the young animal in the womb,
and the whole life of every living creature is for an
allotted period. The plants of the earth have their
seasons for springing up, for coming to maturity, and
bearing flowers and seeds ; and if this order is seriously
interfered with, the plant will sooner or later be incapable
of fulfilling its function. Thus the hyacinth may be
prematurely hastened into flower for one season, but the
next year it will be found impossible to make it flower
or produce seed. In this way great natural events, and
20 ORDER IN RESPECT
especially the life of animals and plants, the movements
of the heavenly bodies, and the eras of geology, become
to us the measurers of time, rearing up prominent land-
marks to guide us as we would make excursions into the
past or future, and dividing it for our benefit into days
and months, and seasons, and years, and epochs.
Thirdly, There is an Order in respect of Colour
running through Nature. — Colour is not without its
significance among the works of man. Every nation,
every regiment has its distinctive colours upon its flags,
which are its visible symbols and representatives. Colour
appears as a peculiar mark on the stamps impressed by
the post-office, and on many of our public conveyances.
It is used as a signal by sea and by land, in our ships
and on our -railways ; it announces danger and proclaims
safety. It has also, we are convinced, a meaning in
nature. It has been far too generally supposed that
colour obeys no laws in natural objects. It has been a
very common impression, that it is spread indiscrimi-
nately over the surface of earth and sky, animal and
plant. We are sure that further research will shew that
this is a mistake. It is true that colour has not so much
value as form and structure in the classification of plants
and animals. Still, we find that some tribes of alga? are
arranged by Harvey according to their colours, and that
some fungi are classified by Berkeley according to the
colours of their minute seeds. We are convinced that,
amidst all the apparent irregularities, there will be found
to be some fixed principles in the distribution of colours
in the animal and vegetable kingdoms, and, indeed, over
the whole surface of nature. Seldom or never, for ex-
ample, are the two primary colours, blue and red, found
on the same organ, or in contact on the same plant
OF COLOUR AND FORM. 21
Liable to certain modifications, which are limited, it is
probable that there is a fixed distribution of colour for
many families of animals and plants, and that this dis-
tribution is fixed within still narrower limits for tho
species. It is certain, whether we are or are not able to
seize it, and turn it to any scientific or practical purpose,
that there are plan and system in the arrangement of
colours throughout both the animal and vegetable worlds.
Every dot in the flower comes in at the proper place,
every tint and shade and hue is in accordance with all
that is contiguous to it. We shall shew at considerable
length as we proceed, that the distribution of colours in
the vegetable kingdom is in beautiful accordance with
the now established laws of harmonious, and especially
of complementary colours. We shall likewise point out
some very curious and interesting relations between the
forms and colours of plants. The eye testifies, too, that
there is an order in respect of colour in the decorations
of insects, in the spots and stripes of wild beasts, and in
the plumage of birds. "He who," says Field, "can
regard nature with the intelligent eye of the colourist,
has a boundless source of never-ceasing gratification
arising from harmonies and accordances which are lost
to the untutored eye."
Fourthly, There is an Order in Nature in respect
of Form. — We use the word form in a large sense, and
as including not only figure, in the narrow sense of the
term, but structure, which is the relation or connexion of
forms. Great attention is evidently paid to this quality
in the construction of natural objects. It appears before
us as a significant element in every department of nature.
The planets, with their satellites, have a definite spher-
oidal shape, and they move in orbits which have a cer-
22 ORDER IN RESPECT
tain outline in space, namely, the elliptic. It is because
strict regard is paid to this principle in the structure of
the universe, that the science which treats of forms, that
is, geometry, admits of an application to so many of the
objects and arrangements of nature. And here it is
worthy of being noted that the ancient geometers, from
a general idea of the importance of forms, had carefully
investigated the properties of those figures called the
Conic Sections, (because capable of being produced by
sections of the cone,) at a time when no very important
application could be made of the propositions established
by them. When Kepler discovered that the planets
moved in elliptic orbits, the properties of the ellipse, un-
folded so many centuries before by Apollonius and others,
were ready to be applied to the solution of a host of im-
portant questions connected with the movements of the
celestial bodies. It is instructive to notice that the
clusters of stars revealed by telescopes of great power,
shew regular forms, some of them being round, and a
number of them having apparently a spiral tendency.
In the mineral kingdom, we find forms playing an
important part. In circumstances admitting of the ope-
ration, most (if not all) minerals crystallize — that is,
assume regular forms. These forms are mathematically
exact in a variety of ways. Every perfect crystal is
bounded by plane surfaces, its sides are parallel to each
other, and the angles made by. its sides are invariable.
Each mineral assumes certain crystalline forms, and no
others. These forms have now an important place
allotted to them in the classification of minerals. They
have been expressively designated the geometry of
nature.
But it is among organized objects that we find form
assuming the highest significance. Every living object
OF FORM. 23
composed though it be of a number, commonly a vast
number and complication of parts, takes, as a whole, a
definite shape, and there is likewise a normal shape for
each of its organs. The general or normal form which
any particular tribe of plants or animals assumes, is
called its type. Animals and vegetables, it is well known,
are classified according to type ; and they can be so
arranged, because types are really found in nature, and
are not the mere creation of human reason or fancy. It
is because attention is paid to type, and because it is so
fixed and universal, that it is possible to arrange into
groups the innumerable natural objects by which we are
surrounded. Without some such principles of unity to
guide him, man would have felt himself lost, as in a
forest, among the works of God, and this because of their
very multiplicity and variety. In some cases the forms
assumed by organic objects are mathematically regular.
A series of beautiful rhomboidal figures, with definite
angles, may be observed on the surface of the cones of
pines and firs. It may be noticed, too, how the leaves
and branches of the plant are placed round the axis in
sets of spirals. The spiral structure is also very evident
both in the turbinated and discoid shells of molluscs.
Mr. Mosely has shewn that the size of the whorls, and
the distance between contiguous whorls, in these shells,
follow a geometrical progression ; and the spiral formed
is the logarithmic, of which it is a property, that it has
everywhere the same geometrical curvature, and is the
only curve, except the circle, which possesses this pro-
perty. Following this law, the animal winds its dwell-
ing in a uniform direction through the space round its
axis. " There is traced/' says Mr. Mosely, "in the shell,
the application of properties of a geometric curve to a
mechauieal purpose, by Him who metes the dimensions
24 ORDER IN RESPECT
of space, and stretches out the forms of matter according
to the rules of a perfect geometry."* We are reminded
of the ancient Platonic maxim, that Deity proceeds by
geometry.
The lower tribes of animals and plants often assume
mathematically regular forms, such as the triangular,
p*olygonal, cylindrical, spherical, and elliptical. It is
seldom, however, that we meet with such rigid mathe-
matical figures in the outline of the higher orders of
organic beings. Those who have any sense of beauty will
be grateful that trees are not triangular, that animals are
not circular in their outline ; in short, that they have not
taken any such painfully exact shape. Still, the forms
of organic objects — such as the sweep of the veins of
leaves and the outline of trees — though more flowing and
waving, are evidently regular curves. There is truth, we
suspect, in a favourite maxim of Oersted, "that inorganic
beings constitute the elementary, and organic the higher
geometry of nature/'
Besides the typical resemblances which enable us to
classify plants and animals, and the beautiful curves
which do so gratify the contemplative intellect, there are
certain correspondences in the structure of organs which
seem to us to be especially illustrative of a plan intelli-
gently devised and systematically pursued. At an early
date, these struck the attention of persons addicted, to
deep reflection, but it is only within these few years that
they have been scientifically investigated and expounded.
Aristotle noticed the correspondence between the hands
of man, the fore-limbs of mammals, and the wings of
birds, and between the limbs of these animals and the
fins of fishes, and spoke of it as an interesting species of
* See Philosophical Transactions for 1838.
t Soul of Nature, IToruer's translations, p. 348
OF FORM. 25
analogy, (*«?' " do not know of any reason
why the sun's attraction and the tangential force co-exist
in the exact proportion they do, but we can trace no co-
incidence between it and the proportions in which any
other elementary powers in the universe are intermingled."
But this we can clearly perceive, that if these proportions
and coincidences had been different, there would have
been confusion throughout the universe ; that if the cen-
tripetal force had been proportionally greater, the earth
and all the planets would have been drawn into the body
of the sun ; and that if the centrifugal force had been
much increased, the earth would have wandered into
regions so far from the sun that all living beings must
have perished. The beauty and fitness of "these coinci-
dences and proportions compel us to see, that though
they do not proceed from natural law, they must proceed
from an Intelligence planning all things, and the rela-
tions of things, from the beginning.
* Mill's Logic, vol. ii. p, 44,
38 NEED Of SPECIAL ADJUSTMENTS
Taking these principles along with us, we are entitled
to say that mutual adjustments are necessary in order
not only to individual effects of a beneficent character,
but also to those general results of an orderly description,
which are very commonly and very properly called laws
of nature. We call the general facts observed by Kepler
laws, but they are evidently the result of the relation of
the planets to the sun, and of their centripetal to their
centrifugal tendency. We talk of the law of the plant
according to which it springs up, assumes certain forms,
bears leaves and seed ; but every one sees that we have
here a complex effect proceeding from a vast number of
arrangements, in which the laws of vitality, whatever
they be, with the laws of moisture, heat, light, and elec-
tricity, are all made to act in unison. It seems to be a
law of the appendages of the plant, of branches, leaves,
and scales, that they are arranged in a spiral manner
round the axis ; but no one looks on this as a simple law ;
it is obviously the result of certain methodical disposi-
tions. We suspect that most of what we call laws of
nature, that most of the principles of order observable in
nature, are of this compound or derivative character.
They are the harmonious result of adjustments many
and varied among a vast number of bodies and of forces,
which, in our present state of knowledge, we must regard
as different from each other, and which at least require
adaptations to be constituted in order to their operation
in a beneficent manner.
If these remarks be just, we are entitled to argue, that
there has been adaptation not only in two or more bodies
being so arranged as to produce an isolated effect of a
benign character, but also in their being so disposed as
to produce general laws or general results, these being
wide-spread and continuous, stretching through extensive
IN ORDER TO THE OPERATIONS OF NATURE. 39
regions of space, and prolonged through many successive
ages, such as the seasons, and the regular forms and
periods of plants and animals. These — indeed all the
principles of order in respect of number, time, colour,
and form — are entitled to be called laws. But they are
not original, they are derivative laws, not simple but
composite, and the result of arrangements. We are thus
enabled to connect the principle of order with the prin-
ciple of special adaptation ; for it is required in order to
the existence of general order, that there should be adap-
tation upon adaptation, and these necessarily of a most
ingenious and far-reaching character.* We shall have
occasion to return, as we proceed, to this subject, as serv-
ing to combine general law and special use in a higher
unity.
SECT. II. THE ADJUSTMENTS ARE DESIGNED, AND NOT CASUAL
NATURE OF CHANCE.
The argument from design in behalf of the Divine
existence, has sometimes been so stated as to make its
main premiss a mere truism, and the whole argument a
begging of the question. It sets out with the maxim,
that whatever exhibits marks of design must have pro-
ceeded from a designing mind ; but by exhibiting marks
of design, is meant proceeding from a designing mind,
and thus the whole ratiocination is nothing but the
pompous repetition of the same proposition. When put
* As the arrangements needful are not only very numerous but very varied, It is pro-*
posed that the word adaptation or adjustment should be substituted for collocation — a
phrase which seems to confine the arrangements to those of place, whereas they may
also include time, number, active property, &c. As these adjustments are necessary
even to the production of those uniform results which we call laws of nature, the proper
distinction is not between the laws of matter aud the collocation of matter, but be-
tween the properties of matter aud the adjustments required in order to thoir beneficent
•ct on. See Method of Divine Government, EoqH II. chaji. j. sect. ji. and iij., 4tl edit.
40 THE ADJUSTMENTS ARE DESIGNED.
in this way, the argument is easily repelled and turned
against him who urges it. But it is not thus that it
has been propounded by any skilful defender of religion.
The argument from final cause, properly understood,
is derived from those concurrences and correspondences
of agents to produce a given end, which everywhere fall
under our notice. These mutual -adaptations of different
and independent powers are so numerous, so curious, and
so beneficent, that they clearly shew that there has been
an Intelligent Being arranging them beforehand. They
cannot proceed from chance, and we therefore conclude
that they must proceed from design.
And this leads us to inquire what is meant by the
word Chance, what is usually meant by it, and what is
the proper meaning of the phrase. A thousand errors
have been lurking in the confused ideas afloat on this
subject, and we must be'" allowed to say that we have
seldom found the nature of chance thoroughly expounded,
or the various meanings of the word distinctly stated.
The ancient atheists argued that there was such a thing
as chance, and ascribed to it the formation of the uni-
verse. Modern materialists and pantheists maintain that
there is no such thing as chance, that there can be no
such thing, and thence argue that there can be no traces
of design, since all things proceed from a chain of phy-
sical or metaphysical causes. We are convinced that
the one as well as the other of these parties is mistaken.
We mean to shew, in opposition to the modern, that
there is such a thing as chance, and, in opposition to the
ancient, that there are adjustments in nature which can-
not proceed from chance.
In maintaining, however, that there is really such a
thing as chance, it is proper to announce that there can-
not be chance in this sense, that there is an event without
NATURE OF CHANCE. 41
a cause. It is not necessary in the present day to insti-
tute any proof of this ; there is no principle more firmly
established or more universally admitted. There may he
a difference of opinion as to the nature of cause and effect,
and a still greater diversity of view as to the nature of
the belief in causation, whether it is derived from interna]
or external sources, but there is none as to the law or the
fact itself. It is admitted that in our world no event
happens without a cause. In this sense chance does not
exist. " There is no such thing as chance," says Hume.
Some would say that it cannot so much as be conceived
to exist.
But still there are senses, and these most important
senses, in which there may be said to be chance in our
world. The word chance, and the corresponding words
accident, casualty, fortuity, may be used, and have an
intelligible meaning when used in two different senses.
First, To use the language of Professor De Morgan,
" the word chance, in the acceptation of probability, refers
to events of which the law or purpose is not visible ;"
and elsewhere, " events do happen by chance, for they cer-
tainly do happen so that we can see no reason why they
should not have been otherwise."* In this sense, whether
looking forward to the future, ever dimly seen, or to the
present or the past as so far unknown, we may speak of
chance, that is, of events of which we do not see the
cause or purpose. As thus used, however, the word is
significant merely of our ignorance, or rather of the ne-
cessary limits set to our knowledge. In this sense it
can have no application to the Divine mind, which is ever
cognizant, of the antecedents and consequents, of the in-
tention and the issue, of all that has occurred, or that is
occurring, or that will occur. As thus employed, the
* De Morgan on Probability, p. 23 ; and Theory of Probabilities in Ency, MetYojv,
42 THE ADJUSTMENTS ARE DESIGNED.
word can have no place for or against us in the argument
which we are now advancing. The limit of our know-
ledge cannot settle the question as to whether the adjust-
ments in nature are or are not designed.
Secondly, Things may be said to be casually related to
each other when the relation between them is not that of
cause arid effect, nor designed by the person producing
them. Every event has a cause, but every event is not
causally connected with every other which may happen
about the same time or place, or have some relation to it
of property or number. This part of the truth is ex-
pressed by Mr. J. S. Mill, — " Facts causally conjoined are
separately the effects of causes, and therefore of laws, but
of different causes, and causes not connected by any law.
It is incorrect, then, to say, that any phenomenon is pro-
duced by chance ; but we may say that two or more phe-
nomena are conjoined by chance, meaning that they are
in no way related through causation, that they are neither
cause and effect, nor effects of the same cause, nor effects
of causes between which there subsists any law of co-ex-
istence, nor even effects of the same original law of colloca-
tion."* The meaning of the phrase, " law of collocation,"
and the use to which it may be turned in the theistic
argument, as pointing to a designed adjustment in the
original constitution of things, have already been noticed.
So much, then, for casual as distinguished from causal
connexion. But casual connexion may also be opposed
to contrived connexion. It is needful to illustrate this,
for it is a position of great importance in our argument.
An agriculturist, let us suppose, is using the means ne-
cessary to secure a crop from his ground. Every step
which he takes must have a causal connexion with some-
thing going before and something coming after ; to this
* Mill's Logic, Book III. chap, xvji.
NATURE OF CHANCE. 43
there can be no exceptions whatsoever. But among the
many agencies he sets a-moving there will be some which
have no discoverable mutual relation, while there will be
others which very visibly have such a relation, which, we
would have it observed, may either be casual or designed.
Thus it may be by accident that he began to plough the
land on the same day as he did the previous year ; by
chance that the two horses in a particular plough are of
the same age ; that his harrows, constructed by different
makers, are painted the same colour ; that the workmen
employed by him have the same Christian name ; and
that he has precisely the same extent of land in crop as
in the previous year. There may be many such relations
and correspondences which persons of a particular turn
of mind find pleasure in noticing, and this because they
are purely casual. But there are other connexions which
are not of this fortuitous diameter. It is not by accident
that he begins his work about the same season as he did
the previous year ; that he has put two horses into his
plough ; that the ploughing has been followed by sowing
and harrowing ; that he has workmen engaged in tilling
his ground, and a certain portion of his whole ground
under cultivation. There is here an evident distinction
between two sets of events, and this distinction does not
arise from the one class having causes, whereas the
others have not, nor from the two proceeding from alto-
gether unconnected laws of collocation, but from the one
being designed as a mean toward an end, and the other
not being so designed, as having no reference to that end.
This distinction between the concurrence of independent
means intended to produce an end, and mere coincidences
which promote no special end, is an all-important one in
the argument from design or final cause.
According to these views we cannot speak of an event
44 THE ADJUSTMENTS ARE DESIGNED.
being produced by chance. Such language has either no
meaning, or a meaning opposed to the universally acknow-
ledged principles of all science and all philosophy. In
respect of causal connexion, chance has and can have no
place ; it is absolutely excluded. But in respect of other
connexions of co-existence or succession, of number and
property, there is room for .chance, and, as opposed to
chance, of designed coincidences and correspondences,
and a co-operation of associated means for the production
of a given end. In respect of production there can be
no such thing as chance, but in respect of disposition
there may. There are mutual relations which are not
designed, even as there are relations which are designed.
We cannot speak of accidental occurrences, but we may
speak of accidental concurrences. We are to shew that
in the place where there is room for chance, there we
have the most striking examples of design.
It may be difficult at times to determine whether
certain events or phenomena are conjoined by chance,
or whether an arrangement has been made to produce
the conjunction. It is no proof of an intended connexion
that they have been conjoined once or twice, or a few
times. Nor can any absolute rule be laid down as to fre-
quency of co-existence, which shall decide every supposable
case that may arise. But there are cases of designed con-
currence so clear that they do not admit of a moment's
hesitation. When we see independent agents all moving
towards one end — when we see stone, lime, wood, glass,
slate, and lead, all combined in a house — when we find
various kinds of metals, and wheels, pulleys, cylinders, of
various shapes and sizes, conjoined to produce a machine,
we at once say the connexion cannot be accidental, but
is the result of arrangements made to secure a contem-
plated end,
NATURE OF CHANCE. 45
Let us suppose that, on entering a room, we discover
on a table before us five or six balls formed into a ring-
like fifrure, we do not allow ourselves for one instant to
imagine that the balls came hither of their own accord,
and without any one placing them there ; but it may be
a question whether the mutual arrangements involved
in the figure are accidental or designed. This question
would at once be settled if we saw other five balls on
the same table formed into a similar figure. We would
then acknowledge at once that there can be as little of
accident in the mutual arrangement of the balls as in
their being brought to this particular place.
These distinctions and explanations enable us to bring
out very distinctly the nature of the argument derived
from adaptation of parts in favour of the existence of God.
In physical nature we have the universal reign of
causation, or every event connected .with at least one
other event as its cause, and yet another event as its con-
sequence. In regard to this point there is no difference
of opinion. But in perfect consistency with this doc-
trine we may find a number of events occurring at the
same time or place, or nearly at the same time or place,
or having some sort of bearing towards each other of a
purely accidental character. In this sense there is no
doubt much of chance in this world, that is, many events
have some sort of discoverable relation, which may yet
have no intended connexion. The year in which a
comet blazes in the heavens may be a year of famine or of
fearful wars and intestine feuds, but this does not go to
prove that the one was meant to forebode the other. We
are quite willing to admit that all these phenomena can
be traced up to God — we are sure that God foreordained
both the comet and the famine ; but it is quite a different
thins; to affirm that the two have a designed connexion
46 THE ADJUSTMENTS AKE DESIGNED.
with each other. Every scar upon the rocks of our
earth may have been produced by causes set in opera-
tion by God, but this will not convince us that there is
deep design on the part of God in presenting to us, here
and there, on these rocks, a figure, which men discover
to bear a rude resemblance to the face of George ifl., of
Nelson, or Napoleon Bonaparte. The fact that there are
accidental concurrences, in the sense now explained, will
not be urged, by any one who seriously reflects upon the
subject, as proving this world is not the product of
design, and that there is not design in every department
of it. In the works of man which exhibit the clearest
signs of contrivance, it is not found that every one part
of the work has an intended relation to every other. In
the construction of the walls of a church there may be
the most careful attention implied in the way in which
the stones are made to fit into each other, but it may all
the while be purely accidental that two stones of much
the same size, weight, or colour, are placed exactly
opposite each other.
But wherever there may be chance, there may surely
be design likewise. If there may be coincidences which
are casual, there may also be concurrences which are
contemplated. It is in the very place where there might be
accident that we discover the clearest and most con-
vincing evidences of design. Upon observing a number
of separate forces acting in union and harmony, we must
believe that there has been a designing mind bringing
them together and causing them to co-operate. When
we see these agencies working in happiest association to
produce innumerable effects of a beneficent character —
when we find them consenting and consorting through-
out thousands or myriads of years or geological ages,
the evidence is felt to be overwhelming beyond the
NATURE OF CHANCE. 47
power of human calculation. Yet this is the sort of
conjunctions and co-operations which is constantly pre-
senting itself to our view. We observe everywhere a
host of separate bodies and powers, all tending towards
a particular end ; — say a number of material substances
with the vital agency, the heat agency, the light agency,
the electric agency, all conspiring to the production of
a living plant or animal ; or bone, nerves, and muscles,
meeting to give an easy motion to a limb. " How often,"
asks Tillotson, " might a man, after he had jumbled a
set of letters in a bag, fling them out upon the ground
before they would fall into an exact poem, yea, or so
much as make a good discourse in prose ? And may not
a little book be as easily made by chance as this great
volume of the world ? How long might a man be
sprinkling colours upon canvas, with a careless hand,
before they would happen to make the exact picture of
a man ? And is a man easier made by chance than this
picture ? How long might twenty thousand blind men,
which should be sent out from the several remote parts
of England, wander up and down before they would all
meet upon Salisbury Plains, and fall into rank and file
in the exact order of an army ? And yet this is much
more easy to be imagined than that the innumerable
blind parts of matter should rendezvous themselves into
a world."
We have the mathematical theory on this subject,
with a most important application, laid down by an
eminent living mathematician. After stating that when
we have a question of pure numbers we can absolutely try
the question with chance in precisely the same manner
in natural theology as we try it in the common affairs
of life, Professor De Morgan thus proceeds :* — "Let us
* De Morgau's Essay on Probability, p. 25.
48 THE ADJUSTMENTS A HE DESIGNED.
assume, as we must, that a number produced by chance
alone, (in the anti-deistical sense of the word,) might as
well have been any other as what it is. And further, let
us require, before we grant intelligence and contrivance,
not merely the presence of an adaptation, which would
have been unlikely from chance alone, but two such
phenomena perfectly distinct from each other considered
as phenomena, each of which might have existed with-
out the other, and both tending to the same object, which
would have been defeated by the absence of either. Let-
it be also granted, to fix our ideas, that we admit as
proved a proposition which has a hundred million to one
in its favour. This being premised, and laying it down
as our object to shew that the necessary result of the
theory of probabilities lead to the conclusion that the
existence of contrivance is made at least as certain, by
means of it, as any other result whicli can come from it,
we proceed to state a consequence. The action of the
planets upon each other, and that of the sun upon all,
(the most certain law of the universe,) would not produce
a permanent system, unless certain other conditions were
fulfilled which do not necessarily follow from the law of
attraction. The latter might have existed without the
former, or the former without the latter, for anything
we know to the contrary.'-'5 Two of these conditions are,
that the orbital motions must be all in the same direc-
tion, and also that the inclinations of the planes of these
orbits must not be considerable. Granting a planetary
system, which is what ours is in every respect, except
either of these two, and it is mathematically shewn that
* An important note is here appended : — "The only way in which we can guess any
two things to be independent. It must be remembered as a result of the theory, that of
things perfectly unknown, the probability of their coming to act, when known, against
an argument is counterbalanced by the equal probability of the future discovery being
on the other side."
NATUKE OF CHANCE. 49
sucli a system must go to ruin ; its planets would not
preserve their distances from the sun. Neither of these
phenomena can be shewn to depend necessarily on the
other, or on any law which regulates the system in
general. For anything we know to the contrary, then,
they are distinct and independent circumstances of the
organization of the whole. Now, let us see what are the
phenomena in question.
" 1. All the eleven planets yet discovered" [that is,
when the work was written] " move in one direction
round the sun. 2. Taking one of them (the earth) as a
standard, the sum of all the angles made by the planes
of the orbits of the remaining ten, with the plane of the
earth's orbit, is less than a right angle, whereas it might
by possibility have been ten right angles.
" Now, it will hereafter be shewn that causes are likely
or unlikely, just in the same proportion that it is likely
or unlikely that observed events should follow from
them. The most probable cause is that from which the
observed event could most easily have arisen. Taking
it, then, as certain that the' preceding phenomena would
have followed from design, if such had existed, seeing
that they are absolutely necessary, ceteris mancntibus, to
the maintenance of a system which that design, if it
exist, actually has organized, we proceed to inquire what
prospect there would have been of such a concurrence of
circumstances if a state of chance had been the only
antecedent. With regard to the sameness of the direc-
tions, either of which might have been from west to east,
or from east to west, the case is precisely similar to the
following : — There is a lottery containing black and
white balls, from each drawing of which it is as likely
a black ball shall arise as a white one, what is the
chance of drawing eleven balls all white ? — Answer, 2047
3
50 THE ADJUSTMENTS ARE DESIGNED.
to one against it. With regard to the other question, our
position is this : — There is a lottery containing an infinite
number of counters, marked with all possible different
angles less than a right angle, in such a manner that
any amrlc is as likely to be drawn as another, so that in
10 drawings the sum of the angles drawn may be any-
thing under 10 right angles. Now, what is the chance
of ten drawings giving collectively less than one right
angle ?— Answer, 10,000,000 to one against it. Now,
what is the chance of both these events coming together ?
— Answer, more than 20,000,000,000 to one against it.
It is consequently of the same degree of probability that
there has been something at work which is not chance in
the formation of the solar system. And the preceding
does not involve a line of argument addressed to our
perception of beauty or utility, but one wdiich is applied
every day, numerically or not, to the common business of
life/'
We have quoted this passage mainly for the mathe-
matical principles which it unfolds. Since the treatise
was written a great number of small planets have been
discovered. These all run in the same direction as the
planets previously discovered, and so add enormously to
the weight of the argument. It is true that the incli-
nation of some of them is considerable, but their mass
is so diminutive that this circumstance is not fitted to
produce any permanent disturbance.*
This is the argument from "Final Cause," as it is
commonly called. At the same time we are inclined to
* See rferschel's " Outlines of Astronomy," p. 453. Should it be said all these con-
ditions can be accounted for by the hypothesis of the cooling and shrinking of a rotating
mass of heated cosmical matter the answer is, that in order to the production of a world
like ours out of such matter, there is need of a whole host of adjustments or collocations.
This subject will be formally taken up in the chapter of next Book which treats of th«
Adjustments of Celestial Phenomena.
NATUKE OF CHANCE. 51
look upon the phrase as rather an unhappy one. The
word, according to the all but invariable usage of our
tongue, points to that which has efficiency ; and there is
nothing of the nature of power implied in the great class
of facts which we are now advancing. In this branch
of investigation we are contemplating not so much a
cause, as an end aimed at, by a combination of means,
by a concurrence of causes. The science which treats of
the relation of means and ends has an unexceptionable
name applied to it, and is called teleology. It would
serve several important ends to have an equally good
phrase to denote the class of facts which it is the business
of that science to explore. As "typos" and "cosmos"
have been naturalized into our language, we wish that
some high authority would introduce " telos" likewise.
In the absence of any such authorized phrase we shall
be obliged to employ final cause, or, in lieu of it, such
terms as aim and purpose, end and special end. We
are to shew that throughout the whole of nature there is
a union and co-operation of means for the production of
what are evidently ends, and such special ends as argue
a living being arranging the means in order to their
accomplishment.
It is not necessary, in order to the conclusiveness of
such an argument, that we should be able to say that
we have discovered the ultimate end aimed at in all
these concurrences of means to produce anterior or inter-
mediate ends. There are persons who seek to cloak the
hicleousness of their atheism under the guise of an
affected humility, urging it is not for them to be so
presumptuous as to pretend to detect the purposes of
Deity. And there might have been some plausibility in
this pretext, provided it had been necessary, in order
to the validity of the argument, to determine the grand
52 THE ADJUSTMENTS ARE DESIGNED.
ultimate design of creation. But it is by no means
requisite in order to prove the existence of design that
we should he able to fathom all the depths of the Divine
counsels, and settle what is the last end of the Creator's
work. On seeing Napoleon Bonaparte gathering his
army to a given point — on finding one battalion coming
from one province and other battalions collecting from
other provinces, distant from the first and from each
other, persons would have been entitled to conclude that
these were means, and well-devised means, to an end,
and this though entirely ignorant of the ultimate pur-
pose to be effected by the subordinate ends ; it would
be enough for them that they discovered the immediate
end and the means employed to accomplish it. On
precisely the same grounds are we justified in maintain-
ing that we observe in nature a singular combination of
means towards the production of an end. This end may
not be the final end of creation, but still it is an end —
a subordinate end, aimed at by a combination of means
arranged by intelligence. Nor can this inference be at
all affected by the circumstance that these ends are com-
monly found to be means towards some other and a higher
end. In God's works all the means are ends, and all the
ends are means, and all means and subordinate ends are
obviously concurring towards a final consummation, which
man can not fully compass, but which he has abundant
reason — from the tendency of the inferior ends — to regard
as at once grand and beneficent.
The argument advanced under this head seems a
complete one in itself. It does not require in order to
its conclusiveness that it should be proven that this
world has had a beginning, nor to look to any physical
facts except those adduced in the premises. The adjust-
ment of the bodies and forces of nature so as to produce
NATURE OF CHANCE, 53
harmonious and useful results, is in itself a proof of an
arrangement not casual but planned by intelligence.
We require not, in order to its conclusiveness, to specify
the time when the adjustments were constituted, nor to
shew that God has created matter as well as arranged
it, nor even so much as that matter has had a beginning.
These other truths may be established more satisfactorily
after it has been demonstrated, from the design mani-
fested in the universe, that there is a God the author of
the design.
The force of the argument now adduced is not to be
turned aside by going back in the chain of causation,
and shewing how each of the combined circumstances,
which form this means towards an end, has proceeded
from a cause. We are not to discard final causes, as
Laplace used to do so summarily, as soon as the physical
cause of the individual circumstance is pointed out.
Nor are we, with Kant, to lay down the principle that
we are at liberty to call in final cause only when mecha-
nical cause fails to account for each particular fact.
The argument which we adduce in favour of final cause
is derived from the wonderful combination of physical
causes. It is freely admitted, that in the material uni-
verse every phenomenon has had a cause, but this does
not weaken the argument founded on the correspondence
between a number of associated phenomena, proceeding
from different and independent causes. No doubt it
forces us to acknowledge that there has been a correspon-
dence in the causes producing such concordant results,
but in carrying us back thus far it only opens up larger
views of the wisdom and foresight involved in a plan
which contemplated such far-reaching consequences. In
Divine workmanship, as also in the higher kinds of
human workmanship, order and utility are commonly
54 THE ADJUSTMENTS ARE DESIGNED.
produced by a long previously arranged consortment of
means or causes. For example, the crop which the
cultivated ground yields is the result of a vast number of
preparations, human and Divine too, made long before.
It is the peculiarity of the Divine workmanship that we
can see in it a set of causes so ordered that they can pro-
duce a series or succession of orderly and benign results
going on from age to age. The plants and animals now
on the earth have all proceeded from progenitors created
many thousand years ago, and which were so constituted
as to produce an offspring after their kind. To argue
from the succession of such effects that they are not
designed, is to make the very beauty and perfection of
the work a proof that it has not proceeded from an intel-
ligent being.
Nor is the force of the argument to be weakened by
the attempt to discover an alleged contradiction ; if
everything, it is said, comes from God, there can be no-
thing casual, there is no room for chance, and therefore
no room for design as distinguished from chance. Now,
it is at once admitted that every physical occurrence may
be regarded as proceeding from God ; at this point, that
is, in regard to the production of the event, there is no
room for accident. But while every event comes from
God, this does not prove that the coincidences between
every two events were designed by Him to produce a
specific end. God has no doubt appointed both the
eclipse and plague which may have happened the same
year, but this does not prove that He designed the one
dark event to foreshadow the other. As there may be
casual relations in nature, so there may be, so there are,
in nature designed concurrences, as distinguished from
accidental coincidences. All that is now occurring is
doubtless the result of collocations previously made; and
NATURE OF CHANCE. 55
m tracing it \>s,ek we must come to certain original col-
locations. At this point physical research stops, but all
inquiry is not arrested. The mind asks, whence this
systematic collocation of agents and forces which has
produced such good and useful results for thousands, or
it may be, millions of years ? The present so full of order
carries us back to the past as also full of order, and shews
that the system now in operation had been planned from
the beginning.
Still less is the force of the argument to be evaded by
the miserable subterfuge of certain French materialists,
who tell us that this consorting of means and end is the
mere condition of existence. When it is found, for ex-
ample, that certain independent members of carnivorous
animals are in admirable harmony, the limbs for running
after the prey, the claws for seizing it, the muscles for
keeping hold of it, the teeth for tearing it, and the sto-
mach for digesting it, an attempt is made to avoid the
force of the appeal by urging that these are the condi-
tions of the existence of the animal. True, we reply, but
the argument is derived from the circumstance that these
independent conditions should meet so as to enable the
animal to exist and to enjoy existence. He who brings
in the principle of the conditions of existence will find
it, if legitimately followed out, landing him in a design-
ing intelligence no less certainly than the principle of
final cause does. The argument, whether for or against
theism, is not to be made to depend on a word or the
shifting of a word. It is not to be established on the one
side by a verbal sophism about design implying a de-
signer, but neither is its overwhelming force to be turned
aside by changing the word final cause for conditions of
existence. It seems that conditions are necessary to cer-
tain existences, audit is the concurrence. of these condi*
56 THE OBVIOUSNESS AND COMPLETENESS
tions, proceeding from various and independent quarters,
which proves so irresistibly that there must have been
design in their arrangement and collocation.
SECT. III. THE OBVIOUSNESS AND COMPLETENESS OP THE
SPECIAL ADAPTATIONS.
The argument from adaptation to a particular end is
one which addresses itself to every human being. It is
suited to every intellect, and comes home to every man's
experience.
1. Every manual labourer may see something analo-
gous to the art by which he earns his livelihood operat-
ing among the natural objects by which he is surrounded.
The sailor may discover the peculiarities of his craft
among marine animals. Thus, among the lower tribes,
he has observed a jelly-fish— called by him the Portu-
guese man-of-war — setting up a sail which consists of a
crest surmounting the bladder. He may notice, too, how
the mussel and pinna anchor themselves by means of
threads of a horny material. The tail of the fish, it is
well known, acts as a scuttle, enabling its possessor to
plough its way through the deep. The web-foot of the
swimmers is an example of what is called " feathering
the oar ;" when advanced forward the web and toes col-
lapse ; the leg (usually so called) of the gillemot and
divers is compressed laterally, presenting a knife edge
before and behind, and thus gives resistance in the fore
and back stroke. It is also worthy of being mentioned,
as illustrating the same point, that the whale's tail col-
lapses in the upward but expands in the downward stroke.
The fisher, as he prepares the bladder to make the
edges of his net float on the water, may observe that the
OF THE SPECIAL ADAPTATIONS. 57
sea-weed, is buoyed on the surface of the deep by a con-
trivance more ingenious than his own, that is, by vesicles
which act as floats. Most fishes have one or more blad-
ders filled with air, the amount of which is regulated by
the will of the animal, so that it can vary its depth, sink
or rise to the surface, as may suit its purposes, The
fisher, too, may see that if he has nets to catch the food
needful for his sustenance, so also have spielers and other
species of animals.
The shepherd knows how much care and watchfulness
are necessary in order to protect his flocks from the wild
beasts which attack them, and is thus led to admii-e the
instincts of those animals, such as the deer, which set a
watch to give a signal of danger. The hunter knows how
much cunning he must exercise in order to come within
reach of the wild animals pursued by him, and should not
withhold a feeling of wonder when he observes how their
instincts lead the brutes to shew such dexterity in avoid-
ing their natural enemies. The weapons with which he
and the fisher attack the animals which they wish to
seize or kill, do not point more clearly to a purpose, than
the instruments, whether claws or teeth, with which they
defend themselves. The Aphrodite hispida, for example,
is furnished with very curious weapons of defence ; they
are harpoons with a double series of barbs, these are re-
tractile, and the animal can draw them into the body by
a muscular apparatus, and in order to prevent them, when
drawn in, from injuring the animal itself, each barbed
spine is furnished with a two-bladed horny sheath, which
closes on the barbs in the act of retraction. Some of these
provisions have a reference to the native instincts of the
animals, others have rather a regard to the position of the
species. Thus we find that those liable to be chased as prey
often take the colour of the ground on which they hahi-
3*
58 THE OBVIOUSNESS AND COMPLETENESS
tually feed. The riflemen of our army are dressed in the
hue which is deemed least conspicuous, and which is best
fitted for concealment ; and is there not an equally clear
proof of design furnished by the circumstance that fishes
are often of the colour of the ground over which they
swim, and that wild animals are not unfrequently of the
colour of the covert in which they hide themselves ?
Thus the back of the young turbot may be seen of the
same colour as the sand on which it lies. The red grouse
and red deer are of the colour of the heath on which they
feed, whereas the lapwing and curlew, themselves and
their eggs, take the grey hue of the pasture among which
they are usually found.
The horticulturist and agriculturist regulate their
plans in accordance with the seasons, and in doing so
they should observe that the plants of the ground suit
themselves in regard to the time of budding, bearing
leaves and fruit, to the same seasons, which are all deter-
mined by the movements of the celestial bodies. The
builder may easily perceive that the woody structure of
plants and the bones of animals are constructed on archi-
tectural principles, being strengthened where weight has
to be supported and pressure resisted, and becoming more
slender where lightness is required. The form of the
bole of a tree, and the manner in which it fixes itself into
the ground, so as to be able to face the storms of a hun-
dred winters, is said to have yielded some suggestions to
the celebrated engineer, Smeaton, in the construction of
the Eddystone Lighthouse. The architect of the Crystal
Palace confesses that he derived some of the ideas em-
bodied in that structure from observing the wonderful
provision made for bearing up the very broad leaf of the
beautiful lily which has been brought within these few
years from the marshes of Gruiana to adorn our conser~
OF THE SPECIAL ADAPTATIONS. 59
vatories The weaver cannot but notice tbat there are
certain tribes of insects which fashion a web of finer tex-
ture than his own. The clothmaker obtains not a little
of the material of the fabrics with which he clothes the
human frame, from the covering provided for the lower
animals, and he derives it all from natural products.
When man wishes to protect his body from severe cold,
he steals their covering from the lower animals, and by
no means of his own devising can he furnish clothing* so
warm as that which has been provided for the brutes in
the Arctic regions. The dyer and calico-printer, with
all the aids of modern chemistry, cannot produce such
rich and agreeable colours as are made to appear for our
gratification in the flowers of plants and the plumage of
birds ; no doubt through the influence of principles which
have not been detected by the very deepest scientific re-
search. Eising higher in the arts we find the painter
taking credit to himself for the beauty of his figures and
colours ; but he cannot, with all his skill and genius,
match those lovely ideal forms and exquisite tints which
everywhere fall under our eye in nature.
" Who can paint
Like nature? Can imagination boast,
Amid her gay creation, hues like these ?
What hand can mix them with that matchless skill,
And lay them on so delicately fine,
And lose them in each other, as appears
In every bud that blows ?':
2. Every kind of contrivance, every principle of me-
chanism used by man, is visibly employed in the opera-
tions of nature. The lamp placed in a window to direct
the benighted traveller, the lighthouse erected on the
harbour to guide the mariner to a place of safety, are not
clearer and more decided illustrations of purpose than.
60 THE OBVIOUSNESS AND COMPLETENESS
the phosphorescent spark by which the glowworm allures
its mate in the darkness of night. What contrivances
does man resort to in order to keep his dwelling warm
and comfortable, but the physiologist will tell him that
there are still more wonderful schemes devised for keep-
ing up the heat of the bodily frame.
Every mechanical power employed by man is at work
in nature. There is as much skilful leverage in the
human frame as in the most ingenious human machine.
The pulleys by which heavy bodies are lifted from the
ground do not give such clear indications of means and
end, as the tendons and muscles by which the bones are
moved. The mechanician has often a large cylinder
running across or through his works, and to this he at-
taches the lesser parts of his machinery. Have we not a
similar contrivance in the backbone of the higher ani-
mals, and the axis of the plant, constituting the support
of all the appendages ? Every one who has seen the
cord of plaited iron by which a carriage is dragged up
an inclined plane, and has noticed how in it strength and
flexibility are combined, should be prepared to admire
the different means by which the same end is effected in
the backbone of all animals, but especially in that of such
animals as the eel and the serpent. The mechanician
who wishes to combine the saving of materials and light-
ness with strength, makes his cylinder a hollow tube : it
is on this principle that Messrs. Stephenson and Fairbairn
have spanned the Mersey by a tubular bridge ; but the
principle was in operation before man adopted it, or was
created to observe it, in many of the bones of animals
which are hollow. Found in the bones of all grades of
living creatures, it is carried out to the greatest extent
where most needed in the bones of birds, so as to allow
them to float in the air, In the case of birds, too, the air
OF THE SPECIAL ADAPTATIONS. 61
from the lungs permeates the larger bones as well as
the smaller parts, the higher temperature of the body
(10S°-112° F.) rarefies it, and imparts an increased
buoyancy to the whole frame.
Every joint in the animal frame can be shewn to be
exactly suited to the function which it has to perform.
Where motion backward and forward in one direction is
all that is required, we have a common joint ; where mo-
tion all round is necessary, we have, as at the shoulder
and hip, the ball and socket-joint admitting of a rotatory
motion round a ball. We have a beautiful example of
ball and socket-joint in the sea-urchin, the spines of
which have a cuplike cavity at the base, which is fitted
to a converse tubercle ^n the shell, fixed by ligaments,
and combining strength and great freedom of motion.
In some parts of the animal frame, a single bone is all
that is required, and more would injure the strength ; in
other parts, as in the fore-arm, a kind of rotatory motion
is furnished by two bones, a radius and an ulna, so ad-
justed as to move to some extent round each other.
Almost every sort of instrument employed by man has
something resembling it in the operations of nature.
The parts of the mouth of insects are made according to
the instincts and habits of the animal, to act now as saws,
now as knives, and, in the case of the leaf-cutting bees,
the mandibles become scissors. The hyena is led by its
instincts to crush the bones of carcases and feed on them ;
and when certain teeth of that animal were shewn by
Professor Owen to an engineer, they were declared by
him to be admirable models of hammers to break stones
for roads. The tongue of many shell-fish, that of the
common limpet for instance, has numerous siliceous
spines, and the organ is used as a rasp or drill. One end
of the shell of Pholas resembles a file, and, by varied
62 THE OBVIOUSNESS AND COMPLETENESS
motions, the animal makes for itself tunnels in clay and
in other substances. The foot of the mole is an admir-
able tunnelling instrument, and enables it to construct
for itself those subterranean passages through which it
is led, by its instincts, to wend its way in search of food.
Instruments of a more peculiar nature, and instru-
ments invented by man only at a late date in the history
of the race, have all along had their analogues in nature.
Millstones are selected because they have gritty mate-
rials in the midst of softer substances ; and we find that,
on a like principle, soft and hard matters are mixed in
the grinding-teeth of mammals. The cupping instru-
ments of surgery were anticipated in the animal king-
dom ; the mouth of the leech combines in itself the offices
of cupping-glass and scarificator ; hence the import-
ance of the animal, as a remedial agent. It is also
worthy of notice in regard to this animal, that the
capacious stomach, with its lateral appendages or reser-
voirs, enables it to extract a very considerable quan-
tity of blood before being detached. Some of the feet
of argulus foliaceus, a parasite on various fresh-water
fishes, are so modified that they act as real suckers or
cupping-glasses ; by a certain arrangement of muscles
the animal can exhaust the cavity of its disc-like feet, and
produce a vacuum, and is thus enabled to stick closely
to the body of the fish.
The tubes and pipes which conduct water and gas
through all the streets and dwellings of a great city, arc
not such ingenious contrivances as the veins and arteries
which convey the blood to every extremity of the frame.
The means by which water is forced to rise in a pump
are not so wonderful as those by which, proceeding on a
different principle, fluid is made to mount in the plant
to the most distant twigc and leaf. We construct valves
OF THE SPECIAL ADAPTATIONS. 63
to allow fluids to pass in one direction, but to prevent
them from flowing back in the opposite direction ; but
before man devised such agency they were already in his
own veins ; and it was upon noticing them that Harvey,
proceeding, as he tells us, on the principle that they were
there to serve a purpose, was led to the discovery of the
circulation of the blood. In the back of the mouth of
the crocodile are two cartilaginous plates or valves, one
above, the other below ; these, acting as floodgates, cut
off communication between the mouth and throat, so
that the animal can hold its prey underneath the water
till dead, and itself continue all the while to breathe by
its nostrils.
3. Among the most curious special modifications are
those in which there is a provision made beforehand for
the support of living creatures not yet in existence.
Every one sees that there is foresight implied in parents
laying up wealth to promote the future comfort of their
children ; but there are equally clear evidences of fore-
thought in the anticipations found among natural objects.
In expectation of the birth of her child the mother makes
preparation for its clothing and comfort ; but there lias
been a preparation by another Designing Mind, so as to
cause the milk to flow at the very time at which it is
required for the sustenance of the infant. In the case
of animals developed from the e\l«
t Fig, 4. Milk-vessels from dandelion.
OF THE CELL. 73
partitions between them become obliterated ; it is obvi-
ous that a continuous tube or duct will be the result.
The pollen or fecundating matter of the plant, so
essential to the continuance of the species, consists of
transformed cells, and the first trace of the new plant is
also a cell, which is stimulated to full development by
the contents of the pollen cell.
These are some of the principal modifications of the
vegetable fabric ; a general plan prevails, which plan is
made to accommodate itself to some particular purpose,
whether this be to produce a tough or elastic fibre, a
hard structure for defence, or a tube required for the
passage of fluids. In the absence of such special and
evidently designed adaptations it seems evident that the
plants which have been so bountifully disseminated over
the surface of our world, would be unavailable for vari-
ous economic purposes, man could not derive from them
food and clothing for his j)erson, nor covering and furni-
ture to his dwelling ; nay more, the very existence of
many vegetable forms is dependent on the special modi-
fications of their simple elements.
2. In Animals. — The higher endowments of the animal
organism imply, in the way of final cause, greater depar-
tures from the primitive cell structure, and, accordingly,
we meet with a greater number of more widely diverg-
ing modifications. The researches of different observers
have, however, tended to shew that a common plan re-
gulates the nature of the primary tissues. We may now
proceed to inquire into modifications bearing a relation
to some necessary end in the economy of the animal.
The thin rjellicle which is separated from the skin in
consequence of a scald, or the application of a blistering
plaster, is called Epidermis ; in its different layers we
can distinctly trace transitional forms of the typical cell.
4
74
SPECIAL MODIFICATIONS
-
©
Fig, 5.*
This part, tliin and delicate although it be, is admirably
fitted to give protection to the tender and sensitive true
skin which lies beneath it, with-
: out at the same time interfering
with the function of sensation or
touch exercised by the latter. It
is specially worthy of notice that
where protection is more essential
than sensation, there is frequently a
very evident increase in the thick-
ness of the cuticle, as in the soles
of the feet and other parts.
Over all the internal free sur-
faces of the animal body, such as
the digestive canal, &c, there is a covering, denominated
Epithelium, essentially of the same nature as the Epider-
mis ; the two are, in fact, continuous, and there is a gradual
transition from the one to the other. There are two
principal forms of epithelium ; the first consists of flat
polygonal cells, the second is composed of others almost
cylindrical, the free surface of the latter often shewing a
fringe of minute filaments, called cilia. Both these kinds
serve to protect the delicate surfaces on which they lie,
and doubtless' act as secreting organs. The cilia of the
second form of epithelium, by their rapid motion, propel
over the surfaces fluids necessaiy for lubrication and other
purposes, and, no doubt, aid in
tlie expulsion of foreign bodies
of small size.
The Adipose Tissue, or Fat, as
Vm. a-t it is commonly called, presents
a very fine example of the cell type. The fat-cells may
* Fig. 5. Oblique section of epidermis, shewing its cellular structure,
t Fig. 0. Cells of fat, or fatty tissue
OF THE CELL. 75
be either spherical or polygonal, the latter being pro-
duced by the mutual pressure of aggregated cells. The
contents consist of oily matter, which each cell has the
peculiar power of forming. The masses of fat thus
constituted are reservoirs of nourishment, to be used
up as occasion requires, and in some cases serve as
a soft bed for delicate organs, such as the eye. The
rounded contour of the body depends in a great mea-
sure on the presence and regular distribution of this
material. Where it is needed we find it, and where
its presence would be inconvenient it is never formed.
Thus, in the palms of the hands, soles of the feet, &c,
it is generally abundant, and serves as a jirotection
against pressure ; it is never deposited in the eyelids,
where its accumulation would undoubtedly be an ob-
stacle to the action of those important appendages of
the organ of vision. Further, being a bad conductor
of caloric, its abundance in certain animals of cold
regions, tends to prevent loss of animal heat ; and in
•some aquatic species, as the seal, its presence diminishes
the specific gravity of the whole body, and thus facilitates
certain movements of the animal.
The Tendons, Ligaments, &c, are examples of fibres
knit together, and occupying certain parts of the body
for the performance of special functions. It is admitted
that all varieties of these may originate from cells, which
are skilful modifications of the type, admirably fitted to
accomplish the end they are made to serve. There are
two different kinds of fibre, the white and yellow. The
white is inelastic, the yellow is highly elastic. The
former is present in the animal body wherever strength
and economy of space are requisite, and wherever im-
portant organs require protection and support. Tendons
and ligaments, the membranes which cover the brain
76 SPECIAL MODIFICATIONS
and soft parts of the eye, &c., consist of inelastic fibre.
By means of the yellow elastic fibre the claws of the
feline tribe are kept retracted when not in use, and a
strong band of the same material, stretching between
the head, neck, and back, and acting as a natural spring,
enables many animals to keep their heads up without
any active effort on their part.
Cartilage, or Gristle, consists mainly of cells, with in-
tervening connecting substance, which may be homoge-
neous, as in the purer forms of cartilage, properly so called,
or the cells may have in the interstices white or yellow
fibre besides. Elasticity, flexibility, as well as solidity,
are properties possessed in an eminent degree by carti-
lage.
The cartilaginous and fibro-cartilaginous modifica-
tions of f he cell type are produced in parts of the body
where a solid material possessed of the properties above
mentioned is required. The flexibility and strength of
the soft part of the nose and of the external ear are ow-
ing to the combination of cartilage and fibre. The ends
of the bones forming the joints, have a covering of carti-
lage, and being thus padded, they are less liable to injury
by sudden shocks. The peculiar properties of the mate-
rials in question perform an all-important function in
the economy of the parts concerned in the formation of
the voice. The strength and elasticity of the entire
spinal column or back-bone depend chiefly on the inter-
vening cartilages by which the entire series of pieces is
connected.
Muscular Tissue or Muscle, constituting the flesh, com-
monly so called, presents, on careful examination, no
veiy remote departure from the cell type ; in fact, the
muscular tissue is essentially composed of modified cells,
which, being first arranged in linear series, with greater
OF THE CELL.
77
Fig. 1*
oi less regularity, subsequently unite to constitute the
elementary fibres. It is unnecessary in such a work as
this, to enter into details regarding the
two varieties of muscular tissue, called
striped and smooth, and their respec-
tive properties ; suffice it to say, that
both perform most important functions
in the animal economy. The active
motions under the control of the will
present the greatest possible variety in
the amount* of force exercised and the
resulting effect. How different the
enormous muscular power exerted by
the whale, when it throws itself entirely
out of the water, from that put forth in the motions of
the eyelids, or of the little muscles which are concerned
in the modulation of the voice, and yet both are formed
by the same tissue ! The giant steam-hammer which can
weld a mass of iron, or simply crack the shell of a nut,
is not more capable of control, and exercises no greater
comparative range of force, than does the muscular ap-
paratus of the animal frame. In singular contrast with
those masses of muscular matter subject to the control
of the will, are those over which we have no control,
such as those of the heart, alimentary canal, &c. But
wherever voluntary or involuntary muscles occur, they
are found precisely where each is most necessary in the
animal economy.
TJie Bones.- — But organs of a harder texture than any
of those already described, are required in the animal
frame, either to protect important paits, or to serve as
levers for the active functions of certain muscles. With-
out the bones the goodly frame of animals would be use-
* Fig. 7. Smooth muscular fibre from the renal vein In man ; shews cell type.
78 SPECIAL MODIFICATIONS
less ; a due combination of soft and of hard parts is
necessary ; active organs of motion must have relation to
others which are passive ; levers must be provided to
sustain and direct the force exercised by the muscular
system.
In a subsequent part of this work we shall have occa-
sion to speak of types and modifications in the general
arrangement of the animal skeleton ; it may be sufficient
to state here that in minute structure the skeleton con-
forms itself to the same type as the soft parts. But
since hardness is requisite, there is superadded to the
cellular element a very large proportion of earthy matter,
consisting chiefly of phosphate of lime.
Nervous Tissue is another modification of the cell
type, for a very important function in the animal frame.
The presence and peculiar functions of the nervous sys-
tem specially distinguished the animal from the vegetable
kingdom. The intercourse of animals with their fellows,
and with the external world, depends on the presence of
a system of nerves, which are necessary to sensation and
to the exercise of every mental endowment. Whatever
may be the form under which nervous matter appears in
the animal body, whether fibres or ganglia, the modi-
fication of the cell type can be traced in course of the
development.
Not only, however, do cells, or their modifications, act
an important part in the protection of surfaces, the sup-
port and strengthening of organs, and the performance
of various active motions, they are also the chief instru-
ments in other functions of the animal economy. With
the exception of the simplest or very lowest tribes, there
is in all animals a system of Vessels for the conveyance of
fluid ; these owe their primary origin to cells arranged in
linear series. In all animals having a true circulationj
OF THE CELL. 79
simple isolated cells form an important part of the circu-
lating fluid ; the Blood-Corpuscles — as they are com-
monly called — to which that fluid owes its colour, are
truly referrihle to the cell type. The food, after under-
going certain changes in the stomach and alimentary
canal, constitutes the fluid called chyle ; it is admitted
that certain Epithelial cells select and absorb the mate-
rials of the chyle, and, becoming turgid with them,
subsequently transfer them to minute vessels — the lac-
teals, which convey them to the blood-vessels. In the
stomach and alimentary canal, certain cells are actively
engaged in pouring out some peculiar and useful secre-
tion. In the stomach such cells are continually form-
ing new broods, which pass out in great numbers, their
contents yielding matters necessary in the process of
digestion.
There are organs whose function it is to separate
matters for some special use, as the milk for the nour-
ishment of the young, or to remove substances whose
presence would be injurious if retained. It is unneces-
sary here to enter into details regarding these various
organs ; suffice it to say that their essentially important
parts belong to the cell type. In short, we find that in
the animal body some special modification of the cell
is concerned in every important function. Cartilage,
bone, muscle, nerve, serve different ends in the animal
economy, but the cell is the essential element in each.
The formation of an image in the eye is mainly effected
by the optical properties of parts having a cellular
origin, and the impression is conveyed by another tissue,
which, as we have already stated, may be referred to the
same general type.
In a subsequent section we shall have ■ occasion to
allude to the general structure as well as modifications
80 SPECIAL MODIFICATIONS OF THE CELL.
of Teeth, Details respecting their mode of development
would be out of place here ; it may he sufficient to state
that cells perform an essential part in the formation of
every tooth.
Nails, Hoofs, Horn, are all essentially Epidermic pro-
ducts, and necessarily partake of the nature of that part
of the skin, that is, are modifications of the cell. Theii
importance in the animal economy is too obvious to re-
quire discussion, serving as they do to protect delicate
parts, and to act as means of defence and offence.
Hairs and Feathers, notwithstanding their variety in
color and texture, have a common origin in cells. The
thick and warm fur of the hare, the smooth and silky
coat of the mole, the spines of the hedgehog, the quills
of the porcupine, and the coarse hairs — resembling split
whalebone — of the elephant and ant-bear, are all mere
modifications of the elemental cell, and each has a re-
ference more or less obvious to the habits of the animal.
The hairs of the mole are closely set, they stand out per-
pendicularly ; in other words, have no particular shed,
and thus present no obstacle to the rapid movements of
this burrower when traversing its narrow and intricate
subterranean tunnels. The spines of the hedgehog and
the sharp quills of the porcupine are respectively admir-
able means of protection to these otherwise defenceless
animals. Feathers, constructed, as we have said, after
the same cellular type as hairs, present similar modifica-
tions in character, varying with the habits of particular
birds. The soft plumage of owls enables them noise-
lessly to steal on their agile prey. The thick-set feathers
and down of divers and other aquatic birds, effectually
repel the water and prevent soiling of surface, as well as
loss of animal heat.
CHAPTER II.
THE FORMS OF PLANTS.
SECT. I. TRACES OF ORDER IX THE ORGANS OF PLANTS.
" When Jupiter," says Herder, "was summoning the
creation, which he meditated in ideal form before him,
he beckoned, and Flora appeared among the rest. Who
can describe her charms, who can image forth her beauty?
Whatever the earth showers from her virgin-lap was
mingled in her shape, her colour, her drapery." We are
to attempt no description of her beauty, which can be
appreciated only by those who look upon her charms di-
rectly, and not through any representation cf them. But
we are to attempt to give something like a scientific ac-
count of that development and structure, of that disposi-
tion of parts and distribution of colours, which mainly
contribute to give to the plant its graceful proportions and
its loveliness. Our present aim is to show that there are
system and design in the progress of the plant, from the.
time it springs from the seed to the time when it yields
seed, and that there are determined types to which all its
organs are made to conform themselves.*
Botanists describe two modifications in the structure
of the seeds of the higher forms of plants. In a pea or
* Wo are to confine our illustrations to flowering plants, partly because the order in
these classes of plants is most easily explained, but mainly because the morphology of
the lower tribes of plants has not been so fully investigated.
4*
82
TRACES Of ORDER
Fig. 8.*
bean, we observe that the principal bulk of the seed con-
sists of two large bodies in close contact ; they are called
seed-lobes, or seed-leaves, and, technically, Cotyledons.
When two are present, the
plant is a Dicotyledon. Be-
tween these organs we ob-
serve the rudiments of the
future stem and leaves. In
other plants, such as the
oat, wheat, Indian corn, etc.,
there appears to be only one
cotyledon, and such plants are called Monocotyledons.
There is a difference between these two kinds of seeds as
regards the process of germination ; but it
will suffice for our purpose, to state that in
both there is a general tendency in one part
to fix itself in the soil, while the other tends
to raise above it into the air ; the former is
the root or descending axis, the latter the stem
or ascending axis. Mere position in reference
to the soil is not, however, an invariable test
of the nature of a part whether stem or root ;
fur there are not a few instances in which
the true permanent stem is underground, as
well as the proper root. But whatever be the position of
these organs, we may see in the plant a continuous prin-
cipal axis, one part of which constitutes the root, and the
other the stem. Attached to the latter there are various
appendages.
* Fia. 8. Embryo of Pea, shewing the point where the young root arises, r; the
young stem or plumule, g ; the stalk, *; connected with the cotyledons, c c, which are
separated and laid open ; f, the depression in which the plumule lay.
t Fir,. 9. Vertical section of grain of oats, shewing the embryo plant at the lower
part, consisting of /, the parts whence the roots proceed; ff, the young stem; c, the
single cotyledon. The covering of the entire grain, o ; covering of the seed proper, t;
the nourishing matter, or albumen, a.
Fig. 9.t
IN THE ORGANS OF PLANTS.
83
On the ascending axis of the plant, we observe two
kinds of appendages, leaves and buds. These last, how-
ever, are mere repetitions of the plant ; each bud con-
sisting of a short axis, and of lateral organs — the young
leaves.
The Leaf, therefore, is the only essential typical appen-
dage of the vegetable organism. It requires no minute
description here ; the most inexperienced observer can
recognize it ; it belongs to the class of " common things."
The study of its many forms lies within the province of
the botanist.
While this typical appendage varies in outline, its
general structure is simple enough. The outer surface
Fig. lu.*
Fro. ll.t
Fis. 124
has a covering called cuticle or skin; the internal portion,
or parenchyma, as it is technically called, has ramifying
* Fjg. 10. To shew curved venation of Endogen.
t Fig. 11. To shew divergent venation of Endogen.
J FaG. 12. To shew netted venation of Exogen — cherry leaf.
84
TKACES OF ORDER
through It the parts called veins.* These different parts,
of which the leaf is made up, are all modifications of the
typical cell, already described. Botanists have described
a difference of the arrangement of the leaf-veins, between
monocotyledons and dicotyledons. In the first of these two
classes, there may be simple veins running more or less
parallel to each other from end to end of the leaf {Fig. 10),
or there may be only one principal vein (midrib) giving off
lateral veins, all of which run parallel to each other {Fig.
11). In dicotyledons, on the other hand, there maybe one
or more principal veins giving off numerous branches and
branchlets on each side, thus constituting a more or less
complicated network {Fig. 12). Lilies, palms, bananas,
&c, present examples of parallel venation; the oak, beech,
&c.j have the netted form. But it may further be observed,
that there is a relation between the structure of the stem
and of the f.eod, and the venation. Dicotyledons have
Fig. 13*
Fig. 14.t
the stem conposed of concentric annual zones, as may be
seen on a transverse section. Monocotyledons present
no such appearance, the vascular parts do not form con-
centric zones, but are broken up into bundles, giving a
* Skeleton leaves, either prepared artificially or found among- fallen leaves after long
exposure to the weather, consist of these alone, skin and parenchyma having dlsap.
peared.
* FlG. 13. Transverse section, stem of oak, an Exogen, or dicotyledon.
t Fig. 14. Transverse section, stem of palm, an Fndogen, or monocotyledon.
IX THE ORGANS OF PLANTS. 85
dotted appearance to the stem when cut across. We may
now proceed to examine certain appendages of the stem,
in order to shew that although they are named as if dif-
ferent in nature from stem and leaf, they are in reality
modifications of one or other of these.
Stipules are leaf-like organs, situated on either side of
the point at which the leaf is attached to the stem, some-
times adhering to the stalk of the leaf, at other times
free. They have various forms, and differ also in size
and texture, according to the plant in which we examine
them. It may he ohserved, however, that they are not
always present, and are not therefore necessary organs.
They are evidently modifications of the leaf, and have
the same general structure and functions.
Pitchers. — These remarkable and beautiful appendages
might afford models to the potter in the construction of
vases for ornamental and useful purposes. Those of the
Sarracenia of North America, usually called Indian-cups,
and the still more remarkable and elegant organs of
Nepenthes, or true pitcher-plant, are examples. They
are all admitted by botanists to be merely modifications
of the leaf type.
Phyllodia, so called from their leaf-like appearance,
are present in not a few plants. In some Australian
acacias they are flattened leaf-stalks ; when young, they
are of narrow dimensions, and actually bear true leaves
of small size ; when the true leaves drop off, these modi-
fied leaf-stalks increase in breadth. In some shrubby
species of wood-sorrel, the transition from leaf-bearing to
leafless flattened stalks can be clearly traced.*
Hairs, scurfs or scales, glands, stings, and prickles, &c,
* The phyllodia of Butcher"s-broom, of Xylophylla, and of Phyllocladus, usually con-
sidered to be flattened and leaf-like branches, may be taken as proof of the relation
be tween branch and leaf-
86 TRACES OF ORDER
are simple prolongations and modifications of the cells
which form the external covering of the leaf or stem. In
dandelion, and numerous other plants of the family
Composite, as well as of some other natural orders, the
divisions of the calyx become transformed into hairs or
hair-like organs. Lenticular glands or lenticels, sup-
posed to he connected with the formation of new or
adventitious roots, and peculiar in their nature, are
now known to be the homologues of cuticular appen-
dages. They present, in different cases, a gradation to
hairs, glands, &c. (See Comptes Bendus, August
1855.)
Spines are abortive branches ending in sharp points.
That this is their nature is evident from such cases
as the following : first, they often produce buds and
leaves, as in the hawthorn ; second, they have the same
general structure as the stem and ordinary branches,
and are therefore not appendages of the surface merely ;
third, they occasionally become branched, as in Gle-
ditschia.
Tendrils are thread-like organs, which have the same
properties as twining stems. They vary in their true
nature. In Gloriosa superba, the midrib or principal
vein of the leaf becomes lengthened, and assumes the
appearance and functions of a tendril. In the Vanilla
plant, the whole leaf sometimes undergoes a similar
transformation. In the pea, vetch, &c, which have com-
pound leaves, the end of the common footstalk forms the
tendril. In Lathyrus aphaca, not merely the end, but
the entire stalk of the compound leaf assumes a similar
form. That such is the nature of the tendril in this
plant is evident from the fact that occasionally a small
leaf is developed upon it. In Smilax, the two tendrils at
the base of the leaf are the homologues of the two sti-
IN THE ORGANS OF PLANTS. 87
pules, and the solitary thread-like appendage or tendril
at the point of attachment of the cucumber leaf, is also
the representative of a stipule. The tendrils in passion-
flower are the homologues of terminal, and in the vine,
of lateral leaf-buds.
The tasteful eye cannot fail to be delighted with the
liveliness and freshness of summer tints, and the gorge-
ousness of autumnal colouring, in the foliage of our forest
trees. Variety of form and diversity of size add to the
aesthetic feelings called forth by the umbrageous canopy
of the vegetable world. Our pleasure and admiration
are greatly enhanced when we proceed to examine more
closely the disposition of the several parts. A casual
glance, indeed, at a tree in full leaf, might leave upon
the mind the impression that its parts were arranged
according to no law, but this arises from the exuberance
of the leafy covering hiding the wonderful method in the
structure. A careful examination will soon reveal to us
that vegetable arrangements are subject to mathema-
tical laws, not less exact in themselves (though ad-
mitting, for special ends, of wider deviations) than those
which regulate the movements of the planets in their
spheres.
The arrangement of the typical appendages has been
fully examined by Braun, Henslow, and others. The
former has endeavoured to shew mathematically, not
only that the spiral regulates the position of the appen-
dages of the stem, but that each species is subject to fixed
laws, by which the nature of the spires, and in many
cases their number, is determined.
The part of the stem or branch from which a leaf ori-
ginates, is called a node, the intervening space an inter-
node. Leaves are said to be alternate, when each node
produces a single leaf, and when the successive leaves
TRACES OF ORDER
A
occupy alternately different sides of the stem. When
there is such an arrangement, a line commencing at the
first leaf, passing round the stem, and touching
the point of attachment of each succeeding leaf,
forms a spiral, the cycle ending with the leaf
placed directly above the one from which we set
out. When two leaves originate from a node,
and are placed face to face, they are called
opposite, and such position has been explained
by some, on the supposition of two spirals pass-
ing simultaneously up the stem. Three or
more leaves springing from a
node form a whorl ; such po-
sition may be owing to the
non-development of the inter-
nodes of an entire cycle, each
spiral being thus reduced to a
circle. In opposite and whorled leaves,
we find not less evident traces of order
as regards the individual leaves of suc-
cessive nodes. In opposite leaves, for
example, the pairs of leaves stand at
right angles to each other ; and in the
whorled, the leaves of each often stand
opposite to the spaces between those of
the next.
The beauty and simplicity of such an arrangement as
the spiral can be clearly seen and appreciated by exa-
mining a branch of an Araucaria, or the cone of any fir.^
Fig. 15*
Fig. 16. -t
* Fig. 15. A stern with alternate leaves arranged in a ouinouncial manner. The sixth
leaf is directly above the first, and commences the second cycle, expressed by fraction f.
t Fig. 16. A stem with opposite leaves. The pairs are placed at Tight angles alter-
nately.
t The spiral twisting of an entire organism, or of some of its parts, is worthy of notice
here: for instance, the stoms of twining plants, as honoysuckle, convolvulus, && The.
IN THE ORGANS OF PLANTS.
89
In the spiral, the number of turns made round the
stem in completing the cycle is different, and we cannot
do better than introduce here the following demonstra-
tions and examples, as given in Professor Balfour's
" Glass-Book of Botany."
Suppose that, commencing with a leaf No. 1, wo reach
leaf No. 8 directly above No. 1, after making three turns
round the stem, the fraction indicating such an arrange-
ment would be ■?. In another case we may reach No. 8
after one turn ; the fraction would then be }. The frac-
tions mark the angular divergence between any two
leaves of the cycle, as represented in the divided circles
Fia. 17.
Tig. 13.
at the upper part of the stems. In Fig. 17, between 1
and 2, the angular divergence is obviously 5 of a circle,
or ^ of 360° = 154;°. In Fig. 18, the divergence is \ of
the circle, or \ of 360° =, 51?°.
leaves of many plants •while in the bud, of banana, for instance, and some modifications
of leaves and brandies, follow the same law, such as tendrils, the flower-stalk of Cycla-
tnon, the seed-vessel of Btreptooarpos, <&c. Among the lower tribes we observe similar
instances, as in the sea-weed Chorda fllum; in species of Desmidium found in fresh,
water; the teeth surrounding the month of the capsule of some mosses, and in others
besides. The "winding of leaves " has been recently examined by Wichu.ra, " Flora*
1852,"
90 TRACES OF ORDER
The following are some of the usual modes of diver-
gence of leaves, and of their modifications : —
Distichous ; -i, as in lime-tree, &c.
Tristichous ; -i, as in Cereus triangularis, &c.
Quincunx ; |, as in apple, cherry, &c„
|, as in holly, laurel, &c.
TS3, as in wormwood, &c.
28T, as in cones of Pinus Pinea, &c.
^f , as in Plaiitago mediea, &c.
f |, as in cones of some pines.
All these fractions embrace common arrangements J
each bears a constant relation to the other ; the numera-
tor of each fraction is equal to the sum of the numerators
of the two preceding fractions, while the denominator is
the sum of the two preceding denominators, and the nu-
merator of each is likewise the denominator of the next
but one preceding.* Such arrangements in regard to
position and number may possess little interest in the
estimation of some, and may seem of minor import, but
they awaken profound reflections in the minds of all
who are disposed to trace the indications of intelligence
in the works of nature.
Organs of Reproduction. — The stem and its append-
ages present us with almost innumerable phases of de-
parture from the primitive type, thus giving a variety of
aspect to the vegetable world, pleasing to the eye and
instructive to the mind. The production of flowers and
fruit, which is the final effort of every plant, immeasur-
ably enhances its value, and adds much to the variety
and the pleasing effects produced by it. In order to the
" herb yielding seed, and the fruit-tree yielding fruit after
his kind," that is, fulfilling one of the very obvious ends
of their existence, there must be superadded an endless
diversity and combination of contrivances. These con-
* Balfour's Class-Book of Botany, p. 99.
IN THE ORGANS OF PLANTS. 91
trivawoes, while they enable the plant to fulfil its func-
tions, are made by Him who accomplishes several ends
by one and the same means, to minister to the pleasure
of man by the aesthetic feelings stirred up. These parts
were described by Linnaeus, in one of his few poetical
fits, as the " nuptial dress :" — they are " In glory gar-
mented, each in its own."
Bracts are parts intervening between the ordinary
leaves and the flower, properly so called. They usually
have a flower in the angle formed between them and the
stem. So closely do they resemble leaves in most plants
that it is not easy to define the difference. In many
cases they have the same colour as leaves, but differ from
them in size and form. In other cases their colour is
materially different ; in certain species of Salvia, for in-
stance, they are as brilliant as the flower. There are
plants in which they exceed the flower itself in size and
beauty, as, for example, Euphorbia splendens. When
very numerous, as in the daisy, the cup of the acorn,
&c, it is very obvious that, like leaves, they obey the law
of the spiral. It is worthy of notice, that in Marc-
graavia and Norantea they resemble pitchers, just as
leaves become transformed into similarly modified ap-
pendages. Bracts, therefore, present us with examples of
transition between true leaves and the parts of the flower.
Inflorescence, or arrangement of flowers on the stem.
There is evidently a plan running through all such
arrangements, just as the spiral law regulates the posi-
tion of the typical appendages. As we shall presently
shew, flowers consist of parts essentially of the same
nature as leaves, and flower-buds may, therefore, be
expected to follow the same law of position as leaf-buds.
The Flower and its parts. — The idea that the leaf is
the type of all the floral organs originated with Linnaeus.
92
TRACES OF ORDER
A clearer enunciation of this theory, and a fuller develop-
ment of the whole, were made by the poefc Goethe. We
now proceed to an examination of this interesting subject.
A complete flower, usually so called, consists of four
series of organs, succeeding each other from below up-
wards, viz., calyx, corolla, stamens, and pistil. The
two first of these, usually so ornamental, are not unfre-
quently absent ; the two last are, properly speaking, the
only essential parts of a flower. If all these organs are
of the same nature as leaves, we ought to find similarity
of general structure, and like obedience to the law of
position.
Calyx. — This, which constitutes the outermost of the
parts which enter into the formation of a complete flower,
consists naturally of separate pieces, called sepals ; these
Fig. 19*
Fig. 20.t
have usually the appearance of leaves, and exactly re-
semble them in structure. In the common bugle (Ajuga
reptans) we find a gradual transition from below upwards,
from leaf to bract, the lower bracts being of the same
colour and form as leaves, while further up they gradu-
* Fk;. 19, Diagram of symmetrical pentamcrous flower, shewing four whorls or con-
centric series of organs, viz. outer row of five sepals, the calyx- ; second row of five
petals, the corolla; third row,-, the five stamens; fourth row, the five pistils. (Flower
of a Dicotyledon.)
+ Fie. 20. Diagram of the symmetrical trimerous flower of Fritillary, having three
divisions of calyx, and three of corolla; six stamens in two rows; and a pistil composed
OS three united. (Flower of a Monocotyledon,)
IN THE ORGANS OF PLANTS. 93
ally assume a bluish purple tint, while their venation is
also modified, in both which respects they resemble the
calyx. But this is only one of the many instances in
which we can trace upon the same plant a transition from
leaf to bract, and from bract to sepal.
Corolla. — The term flower is, in common laneniagre,
employed to express that part which is most brilliant in
colour ; this, in botanical language, is the corolla, and
the pieces of it are called petals. In fuchsia the calyx
and corolla are equally conspicuous in colour ; nay, in
some varieties the former is the more splendid of the
two. In monocotyledons, the two whorls, that of the
calyx and corolla, generally resemble each other both in
form and colour ; thus, in Herb Paris there is a striking
similarity between them. Magnolia, certain species of
water-lily, and other plants, present in the same flower a
decided transition from calyx to corolla, and the converse.
In general structure the two organs are little different.
We have already seen that the transition from leaves to
bracts, and from the latter to sepals, is obvious enough ;
and as the two first are evidently of the same nature, so
it may be inferred that sepals and petals are really con-
structed after the leaf type ; and the highest authorities
are agreed on this point.
Stamens. — These, which form the third series of floral
organs, from without inwards (see Fig. 19), present the
greatest departure from the type of the leaf, there being
a general diminution of superficial extent, with an in-
crease of thickness at the extremity. A perfect stamen
consists of two parts, anther and filament, the latter
corresponding to the leaf-stalk, the former to the blade.
The filament is no more essential to the anther than the
stalk is to the blade of the leaf, and is often absent.
In many double flowers we observe a series of changes.
94 TRACES OF ORDER
which illustrate the true nature of the stamen. It is
clearly demonstrable in some double roses, the stamen
Fig. 21*
passing from its normal condition into a petal, and this
again into a sepal. The common white water-lily pre-
sents us with the same transition as its natural structure.
Since, then, bracts are of the same nature as the leaf,
and bracts are allied to sepals, and sepals to petals, and
all this in more than one particular ; and as petals pass
into stamens, and the converse, all may be regarded as
formed after the same type.
The part of the stamen called anther corresponds to
the blade of a leaf ; its two halves represent the two por-
tions of the leaf divided by the midrib, and the whole
surrounded by cuticle, and containing cellular tissue or
parenchima, some part of which becomes transformed
into the pollen or fecundating powder of the plant, so es-
sential to the formation of the seed.
Pistil. — This central organ of the flower more resem-
bles the typical leaf than the stamen. It differs from
the three outer whorls of the flower in this respect, that
when simple it is generally the representative of a leaf
folded upon itself, and with some of its parts adhering
more or less together ; the same is true of its individual
pieces when it is compound.* This folding and adhe-
sion of the seed-vessel is not always complete, as we may
* Fie. 21. Transformations in the stamens of the rose. The complete stamen is altered,
gradnallv passing through different states, until it becomes a petal, and the petal resem-
bles a si'pal with a midrib.
t Schleiden believes some pistils to be really hollow stems.
IN THE ORGANS OF PLANTS. 95
gee in the common mignonette of gardens. The leaf-
like structure of the seed-vessel or pistil, as its natural
condition, is very obvious in many plants ; fur instance,
in the pod of the pea or bean, and in that of hellebore
and marsh-marigold. In some cases we see it reverting
to the general type ; for example, in the
cherry with double flowers, the fruit of
which is abortive, and in its stead we ob-
serve one or two green leaves, resembling in
miniature those of the tree.
In order to understand the nature of
this part of the flower, let us imagine a
leaf such as that of the cherry or laurel, to «— F>o- 22.*— 5
be so folded that the two edges are brought in contact,
the two halves of the upper surface being opposed to each
other, and the whole in a vertical position ; the lower
surface of such leaf will correspond to the outer surface
of the pistil, and the upper to its lining or inner surface.
Such, in fact, is the real nature of a simple leaf-pistil.
The seed. — We may now inquire into the nature of
the seeds, called, technically, when young, ovules or little
eggs. It is well known that the leaves of some plants
bear buds on their edges ; for example, Malaxis and
Bryophyllum ; the ovules are representatives of such
buds. Suppose a leaf of one of these plants folded on
itself, and the edges also folded inwards and adherent,
we have in this way an exact representative of the seed
vessel and seeds of not a few plants. Sonic abnormal
cases illustrate the same truth ; thus, Professor Henslow
has shewn that in mignonette the ovules sometimes
become transformed into small leaves attached to a short
axis, precisely the structure of a bud. Whether, there-
* Fio. 22. Seed-vessel of double-flowering cherry converted Into a small leaf, in two
states ; unfolded, a; folded, t>.
96 TRACES OF ORDER
fore, we adopt Schimper's view, that " ovules are buds
of a higher order, their integuments leaves, and their
stalk the axis," or Lindley's, that they are " leaf-buds in
a particular state, and their integuments composed of
scales or rudimentary leaves," we are still constrained to
admit that they are formed after the same type as the
other parts.*
But if the different series of organs which we have
been describing as entering into the formation of a perfect
flower, are really of the same nature as the leaves or typical
appendages, they ought to have their position regulated
by the same law. Such, in fact, is the case.
We have already had occasion to allude to the whorlcd
arrangement of leaves on the stem, and in these instances
we have a type of any of the four whorls of the flower
already described. Farther, the relative position of the
leaves in successive whorls, represents also that of the
parts of the flower ; for as the leaves in successive whorls
are, generally speaking, alternate, the same holds in the
flower. This comparison is admitted by all authorities.
The whorled arrangement of leaves is but a modifica-
tion of the spiral, and the same law regulates the position
and mutual relations of sepals, petals, stamens, and pis-
tils. The parts of succeeding whorls, in both cases,
occupy the same relative position as in whorls of leaves —
that is, each is placed opposite the space between two in
the next series ; in other words, the parts of the flower
alternate with each other. (See Fig. 19). But the law
of the spiral extends also to the individual pieces of each
whorl, though it is frequently not very obvious, and is
* Itmay be worthy of notice here, as connected with this subject, that in some plants
called viviparous, we observe mixed up with the flowers and flower-buds small bulbs.
which, when mature, drop off and take root ; they are, in fact, miniature buds. Poly-
gonum viviparum, Sasifraga cernua, and others are examples. In bume of these tho
true flowers are reduced to two or three at the upper part of the stem.
IN THE ORGANS OF PLANTS. 97
liable to be overlooked by a careless observer. In some
species of rock-rose, and in Polygala, as well as in many-
other plants besides, some of the sepals, or pieces of
the calycine whorl, are lower or more external than the
others, which are higher and within the former. This
prevalence of the spiral is especially obvious in the pistil
or central part of the flower. The common strawberry,
when ripe, illustrates this ; the numerous small pistils
(or seeds, as they are commonly but improperly called)
dotted over its surface, will be found, on close examina-
tion, td follow the spiral arrangement. The soft, juicy
part of the strawberry is just an enlarged fruit-stalk,
axis, or stem (receptacle of botanists), and the numerous
minute pistils or seed-vessels which it supports, are ar-
ranged according to the same law which regulates the
position of leaves, of which they are homotypes.
Adolphe Brougniart long since showed, that what are
called floral whorls are not strictly such in many cases, but
merely a series of organs closely approximated, and occu-
pying different heights on the short axis. This, aj we
have shown, is often sufficiently obvious as the na fcural
condition of the parts, but it is at times more palpable
in monstrous flowers.
We have stated that ovules are of the same nature aa
buds. Since these latter, growing usually in the angle
between stem and leaves, necessarily follow the law of the
spiral in regard to position, the same ought to be true of
ovules, and examples of this are easily found. Even
when the ovules or young seeds are very numerous in a
seed-vessel, there is no confusion, but the utmost regu-
larity in their arrangement. Thus in the pod of a pea,
where they form two rows, corresponding to the infolded
edges of the typical leaf, those on one side alternate with
those on the other. In the seed-vessel of the wall-flower,
where they are more numerous, they follow the same law.
5
98 TRACES OF ORDER
The regularity is not less obvious when Ave examine cases
in which they are more abundant still, as, for example,
in the seed-vessel of the common foxglove. It is worthy
of notice, as an illustration of the same law, that the two
seeds usually found at the base of each scale in the cone
of a fir, are often not exactly on the same level, one being
generally a little higher than the other.
Not only are there relations of structure and position
in the parts of the flowers, but we also observe relations
in number.
The typical flower in plants, having the dicotyledon-
ous structure of seed, has its parts regulated by the num-
bers four or five, or some multiples of them ; in flowers
of monocotyledons, the number three, or some multi-
ple of it, prevails.* The fundamental structure in both
may be modified in three ways ; 1st, by lateral adhesion
of the pieces of the same series, or of organs of different
series ; 2c?, by increase or diminution in the number of
the parts ; od, by inequality of size and form, or union
of the different parts, or peculiarities in the development
of the axis which supports them. Some botanical author-
ities admit the existence of nine thousand genera, and
about one hundred thousand species of the higher forms
of plants.f The characters of the former being founded
* Linnreus, in classifying plants according to the number of stamens, attached, prob-
ably without being aware of the importance of the principle, a greater weight to num-
bers than has been assigned to them by more modern observers. In Geraniums we
may often observe five stems, five leaves divided into five parts, five flower-stalks, five
sepals, five petals, and the stamens in multiples of five. In the natural family Umbel-
lifer* (carrot and hemlock arc examples), the number five prevails not only in the
flower, but it also seems to regulate the inflorescence, five or some multiple of it occur-
ing very frequently in that part. The common elder-tree belonging to the Honey-
suckle family, has live leaflets on a common stalk, the inflorescence or flower-stem has
five primary branches, each of these has in turn five secondary, and so on repeatedly ;
five being also the typical number in the flower. In the true heaths, four is the typical
number in the parts of the flower, but it (or its multiples) often appears also to regulate
the number of leaves which appear together, as well as the number of flowers which are
grouped together.
t These numbers are doubtless far above the mark, as regards plants actually discov-
ered.
EST THE ORGANS OF PLANTS. 99
on differences in the organs of reproduction, there are
therefore numerous modifications of the typical flower.
All parts of the plant furnish characters of species, and
there are therefore many thousand modifications of the
typical plant. Amid so much variety, it is pleasing to
contemplate the common plan which regulates all ; and
knowing that plan, we possess a key to explain those re- '
markable forms which are so common in the vegetable
kingdom, whether the coronet-like flower of Napoleona
imperialis, the irregular flower of Aristolochia, or of the
Balsams, or that peculiar slipper-shaped corolla from
which Calceolaria derives its name. The gaping flower
of Mimulus, and the irregular mask-like flower of Lina-
ria, are all referable to a common type. A knowledge
of the typical flower in the Endogens enables us to ascer-
tain the true nature of those modifications which render
the grotesque flowers of the Orchids so remarkable ; in
some it resembles an insect, in others a spider, and in a
third case, a helmet with the visor up, indeed there is
scarcely a common insect or reptile to which some of them
have not been likened."* The flowers of the bee and
spider orchis, the toad-like Megaclinium Bufo, and the
Caleana nigrita of Swan River, whose flowers capture in-
sects, and all the anomalous Cape species, can be inter-
preted when we know the type.
Having gone over the organs of the plant indivi-
dually, we are now to inquire whether there may not be
indications of a unity running through all classes of
plants.
Allusion has been made to two great classes of flower-
ing plants called Monocotyledons and Dicotyledons, each
characterized by peculiarities in the structure of seed, of
stem, and of leaf (and also by a difference in the mode
* Llndley's Vegetable Kingdom, p. 176.
100 TRACES OF ORDER
of germination of the seeds). Each also has, generally
speaking, a certain number or its multiples regulating
the number of parts in the flower. There seem, however,
to bo evidences that these two great classes, thus usually
distinguished, really possess much that is common. Ac-
cording to Mohl, the structure of the stem of an Endo-
gen and of an Exogen, during the first year of their
growth, is altogether the same. Dutrochet indicates the
Bryony as an example of such identity. As to the seed,
Professor Lindley remarks,* " It is apparent that dicoty-
ledons are not absolutely characterized by having two
cotyledons, nor monocotyledons by having only one.
The real distinction between them consists in the mode
of germination, and in the cotyledons of dicotyledons
being opposite or in whorls, while in the monocotyledons
they are solitary or alternate."
The difference in the arrangement of the veins of the
leaves in these two classes present not a few exceptions ;
thus, on the one hand, among monocotyledons we have
examples of netted venation, as in Arum, Calla, Lilium
giganteum, &c, and on the other, examples of parallel
venation among dicotyledons, as in Nerium. There seem,
therefore, to be transitional forms between the two great
classes into which the largest proportion of the highei
plants has been divided by botanists.
There are indications, too, of a unity of structure run-
ning through all the organs of the individual plant. We
think it of importance to illustrate this at considerable
length.
It will not be reckoned by any scientific botanist, in
the present day, as an excess of refinement to represent
the developed organs of the plant as all formed after one
or other of two different types or models, the Stem and
the Leaf.
* Introduction to Botany, vol. ii. p. 267,
IN THE ORGANS OF PLANTS. 101
First, The more solid parts of the plant are composed
of a number of stems, proceeding the one from the other
in linear succession. Springing from the embryo, or
seed, there is the axis mounting upward and becoming
the aerial stem, and growing downward and becoming the
root. From the former of these, or the ascending axis,
there go off lateral stems, which we may call branches,
and from these, other stems, which we may call branch-
lets. There proceed, in like manner, from the descending
axis, or top root, lateral branches which also ramify
through the soil. There are important differences be-
tween the aerial and the subterranean steins to fit them
for their different functions. Boots, for examjxle, have
no pith, no scales or leaves, and, in ordinary circum-
stances, no leaf-buds like the upward axis. Still the two
are alike in the general character ; the branched plant is
found to have a branched root. The tendencies of the
underground ramification have not, so far as we know,
been carefully determined ; but above ground, it is very
evident that the stem, branch;, and branchlet obey the
same laws. u If a thousand branches frond the same
tree," says Lindley, " are compared together, they will be
found to be formed upon the same uniform plan, and to
accord in every essential particular. Each branch is
also, under favorable circumstances, capable of itself
becoming a separate individual, as is found by cuttings,
buddings, graftings, and other horticultural processes.
This being the case, it follows that what is proved of one
branch is true of all the others." We have seen a pear-
tree laid prostrate on the ground by storms, but, with its
roots still fixed in the soil, sending out a branch from its
side, which mounted upward, and took a form precisely
like that of the parent tree. .
The other typical or model form is the leaf. We have
102 TKACES OF ORDEK
shewn that all the appendages of the plant are constructed
on this type. " Linnaeus had a presentiment of some-
thing of this kind, and, in his Prolepsis Plantarum,
carried it out in such a way that, starting from the con-
sideration of a perennial plant with regular periodicity
of vegetation, as in our forest trees, he explained the col-
lective floral plants, from the bracts onward, as the collec-
tive foliar produce of a five-year-old shoot, which, by
anticipation and modification, was developed in one year.
This view is, in the first instance, taken from the most
limited point possible, from the examination of a plant
of our climate ; and secondly, imagined and carried out
with great want of clearness/'* The true doctrine was
first propounded by C. F. Wolff (Theoria Generationis,
1764), but his treatise lay buried in neglect till the doc-
trine became established by the influence of others. It
was first presented to the world by the great German
poet, Goethe, who, though not learned in the artificial
botany at that time taught in the schools, had a fine eye
for the objective world. We are not willing, indeed,
to admit that the form in which Goethe expounded the
doctrine is in every respect correct. It is wrong to repre-
sent floral organs as metamorphosed leaves, for they
never have been leaves in fact ; the accurate statement
is, that these organs and leaves are formed after the same
general plan. Nor are we to represent nature as striving-
after a model form, which she fails to reach, in the va-
rious modifications of organs ; for the modifications are
as much an end and intended, as the parts which may-
be pointed to as patterns. Still, Goethe may be regarded
as having seized the great law of vegetable morphology.
His Versuch die Metamorphose der Pflanzen zu erklaren,
was published in 1790, and has furnished the foundation
* Schleiden's Principles of Scientific Botany, translated by Lanfeester.
IN THE ORGANS OF PLANTS.
103
to scientific botany. But as Goethe had no name among
the initiated, little attention was paid by botanists gene-
rally to his speculations
till long after, when they
were mentioned by Jus-
sieu, and brought into
general notice by De Can-
dolle, in his " Organogra-
phies published in 1827.
The doctrine, somewhat
modified, is now acknow-
ledged by the great doctors,
and has been sanctioned
by the great councils of
science.
According to this idea,
a plant is composed of two
essentially distinct parts,
the stem and leaf. The
leaf is attached to the
ascending stem, and be-
sides its common form, it
takes, while obeying the
same fundamental laws,
certain other forms, as
scales, bracts, sepals, pe-
tals, stamens, and pistils.
Schleiden, who has deve-
loped this view, gives, in
his " Plant, a Biography," a picture of a typical plant con-
structed on this principle. This makes a plant a dual.
* Fig. 23. The typical plant— 1 to T. Axis.
I to VI. Appendages.— I. Cotyledon; II. Leaves; III. Calyx; IV. Corolla; V. Sta«
men; VI. Pistil.
II. Typical appendages. I. III. IV. V VI. Modified appendages.
b b, Buds composed of shortened axes, with rudimentary appendages.
104 TRACES OF ORDER
But it appears to us possible to reduce a plant by a
more enlarged conception of its nature to a unity, that
is, to shew that there is a unity of plan running through-
out the whole.*""
Looking first at the ramification of the stems, we may
observe a central stem, or central stems, sending out
other stems at definite angles, and of a normal length,
and altogether in so regular a manner that the whole
plant is made to take a predetermined form. Looking
next at the venation of the leaf we perceive (see Figs.
12, 24, 25, 26) that it too has a ramified character, that
it has in the centre a main rib, or ribs, from which pro-
ceed other ribs or veins in so definite a manner that the
whole skeleton assumes a regular shape. Now, we main-
tain that a number of correspondences can be detected
between the ramification of the stems and the ramifica-
tion of f he leaf- veins.
In prosecuting this inquiry, let us first inspect, in a
general way, the leaf of a tree, with its central vein, or
veins, and its side veins. (See Figs. 12, 24, 25, 26.)
On the most cursory inspection the impression will be
left on the mind that the central vein, or midrib, as it is
called, corresponds to the central stem or axis of the
tree, and its side veins to the branches. Having seized
the figure of the leaf- venation in the first instance, let us
now look at the skeleton of the tree, say a tree stripped
of its leaves in winter, and we may notice how like it is
in its disc and the arrangement of its parts to the skele-
ton and outline of a leaf. We shall be particularly
struck with this if we view the tree in the dim twilight,
* Br. M'Cosh has here to express his obligations to Professor Balfour of Edinburgh
who, without prematurely committing himself to these views, has kindly helped to give
them publicity and bring them under discussion. See Transactions of Botanical Society
of Edinburgh, July 1851, and Balfour's Class-Book of Botany, 2.1 edit. p. 113; see als«
$ecUon;il Reports, for 1S52 and 1S,jl, of British Association for Promotion of Science,
IN THE OBGANS OF PLANTS. 105
jt " pale moonlight," between us and a clear sky, as we
may conceive Wordsworth to have viewed it.
" Often have I stood,
Foot-bound, uplooking at this lovely tree,
Beneath a frosty moon."
We are quite aware that, in the tree, the branches go off
ail round the axis, and give to the whole figure a sphe-
rical form, whereas in the leaf the fibrous veins all lie
in one plane. But then we have a transition from the
one to the other, and a point of connexion in the branch,
the branchlets of which — as, for example, visibly in the
beech — often lie in one plane, and, if filled up, would
make the figure bear a resemblance to the leaf. The
principal difference between the tree and leaf may pos-
sibly be found to consist in this, that for special ends the
cellular tissue which, in the tree and its branches, is col-
lected into the pith and bark, (which are connected by
the medullary rays,) is in the leaf spread out so as to fill
up the interstices in the fibrous matter which forms the
veins. The general impression produced by the first
thoughtful survey of a morphological correspondence
between stem ramification and leaf ramification will be
confirmed by a more searching and scientific investiga-
tion. In maintaining this, we always assume that in the-
cases subjected to examination both stem and leaf are
fully and fairly developed.
But here it will be necessary to have it settled, at the
outset, that every species of plant tends, if allowed to
grow freely and in favorable circumstances, to take a
particular form, and that the same is also true of the
leaf. This statement will be allowed, after a moment's
recollection and thought, as to the leaf. The cherry
leaf (Fig. 12) obviously assumes one shape, the beech
leaf (Fig. 24) another shape, the lime leaf (Fig. 25) a
5*
106 TRACES OF ORDER
third shape, and the poplar leaf (Fig. 26) yet a different-
shape. Every one who has used his eyes will remember
that the oak leaf has its peculiar figure, and the thorn
leaf its own conformation, and the birch leaf its specific
outline, by which we at once recognise them and distin-
guish them the one from the other. A very little patient
observation of trees growing freely — of lawn-trees, for
example — may satisfy any one, that what is true of the
leaf is also true of the tree. Every species of tree, ac-
cording to naturalists, has its own habit ; and this gives
to it a peculiar physiognomy by which the practised eye
will at once recognise it. We have often found it in-
teresting, (when we had nothing else to interest us,) in
passing along a road, to detect, by their configuration,
the various species of trees which met the eye, and
this when they were bared in winter, and there was no
foliage to aid us. Towards this normal shape of its
species every individual tree tends. No doubt it is
greatly interfered with, and much thwarted in its efforts
by prevailing winds which bend it, or violent storms
which break it, by too much cold at one side, or too
much sunshine at another side, by a niggard soil denying
nourishment, or officious neighbours jostling it, by cattle
browsing on it, or men cutting it ; still we can see the
native tendency in the most unfavourable circumstances,
while, in more favoured positions, we see the tree grow-
ing up to its beau-ideal.
And here it may be laid down as a general rule, that
every plant takes the fairest shape when allowed to
assume its natural form. True, there are trees which
have been rendered picturesque by being torn or twisted
by the storm, or venerable by the marks of age ; but-
being unaided by associated feelings produced by such
causes, the plant is always injured when attempts are
IN THE ORGANS OF PLANTS. 107
made by man to give it an artificial shape. Every tree
should be allowed fairly to develop itself, protected only
from rude winds, and interfering neighbours, and graz-
ings of cattle, and primings of man, who so often mars
in attempting to mend. All ornamental pruning should
aim, not at improving, but aiding nature — nay, not so
much at aiding it, as cutting off unnatural additions and
removing artificial imperfections. Thus left to their
innate tendencies, all plants will grow into a form more
or less beautiful. A tree growing freely and fairly in a
lawn, where it has soil to feed it, and space to develop
itself, and air to breathe in, and sun to warm it, and
fences to shelter it, stands before us a most interesting
object of contemplation. The parallel branches, and
their spiral arrangement round the axis, their sweep of
curve, and the methodical way in which they first
lengthen and then shorten as they ascend the trunk, and
the graceful rotundity and elegant outline of the whole
between us and the sky, all combine to fix the eye, and
unconsciously excite and engage the musing intellect.
And there is another beauty produced by a number of
differently-formed trees standing on the same lawn, and
each shewing its separate mould and features. For as
one star differeth from another in glory, and as one saint
in heaven differeth from another in glory, so one tree
differeth from another in glory. There is one glory of
the oak, which looks as it it had faced a hundred storms,
and having stood them all, were ready to face as many
more ; another glory of the sycamore, that " spreads in
gentle pomp it honeyed shade \" another glory of the
birch, so graceful in the midst of its maiden tresses ; an-
other glory of the elm, throwing out its wide arms as if
rejoicing in its strength ; and another glory of the lime,
with its sheltering shade inviting us to enter and to linger,
108
TRACES OF ORDER
Each lias its own glory, of which it would he shorn were
it to make an ambitious attempt to usurp the glory of
its neighbour.
It being allowed that there is a pattern form for the
whole plant and for its leaf, we are now to trace certain
interesting correspondences which we have noticed be-
tween the two.*
1. In plants with ivoocly structure, there seems to he a
correspondence between the tree and leaf in this respect,
that a leaf without a leaf-stalk implies a trunk naturally
branched from the ground, and a leaf ivith a leaf -stalk
implies that the species of tree on which it grows has
naturcdly a hare sialic. — In order to the settlement of
this point, it is necessary to have it admitted that there
are trees which are naturally feathered from the base,
whereas there are others which
have less or more of an unbranched
trunk. Belonging to the former
class we may name the greater
number of our ornamental lawn
shrubs, as the box, the holly, the
laurels, bay and Portugal, the ar-
butus, the laurustinus, the privet,
the snowberry. All of these cover
the lawn from near the base, and it
may be observed of the leaves of all
of them, that they have no petiole,
or a very short petiole. To this
same class belong many of the common forest trees, such
* Our observations have been extensive and varied, but they are limited when com-
pared with the whole vegetable kingdom, and so we are prepared to expect that curious
modifications and anomalies will cast up, which, while not. setting aside these general
views, will open new views, and enable science, in the end, to rise to a more thorough
conception of the plant.
+ Fig. 24. Beech leaf, as an example of leaf with little or no leaf-stalk ; shewing nearly
parallel reive; angle of venation, 45° to 50°; the midrib zigzag.
Fig. 24. t
IN THE ORGANS OF PLANTS. 109
as the oak, the elm, the beech. The leaves of these
trees have little or no leaf-stalk, and we are able, from a
rather extensive observation, to affirm that these trees
incline to send out branches from the base. At times,
indeed, this tendency is interfered with. In fields, the
lower branches are frequently eaten by cattle, and in
thick woods they often fail from want of air. The lower
branches of the young oak are studiously cut off by
woodmen, in order to get a tall, unbranched trunk for
timber. The beech is not unfrequently cut over before
being planted out in lawns, and a whorl of branches is
made, in consequence, to spring out some few feet above
the ground. In England, the favorite elm is often
pruned near the base, in order to lessen the shade upon
the field or highway. But when allowed to grow unmo-
lested, and in favourable circumstances, these trees arc
all bushy from the base. The very circumstance that
the oak require"] pruning in order to its having a bare
trunk, proves that its own tendency is otherwise. The
beech shews that it is naturally branched from the roots,
by the closeness of the hedges which it forms. The
pruned elm is ever displaying its native disposition, by
the little branches that crop out from its trunk in spite
of all the cutting to which it is subjected.
Other trees, again, have less or more of a bare trunk,
and the leaves of these have less or more of a leaf-stalk.
To this belong the cherry, {Fig. 12,) the lime, {Fig. 25,)
the poplar, {Fig. 26,) the apple, the pear, the birch, the
chestnut, the sycamore. These cannot be induced, except
by constant cutting, to grow bushy, or to afford shelter,
from their base in rows or fences. The thorn may seem
to furnish a disproof, by its being so commonly employed
in hedges. But every one who has bestowed the least
attention upon iiu subject, knows that thorns need con-
110
TRACES OF ORDER
stant cutting to keep them from becoming bare near the
root, and their native habits are seen when they are
planted out in lawns, where they have invariably (as the
beautiful thorns in Phoenix Park, Dublin, can testify) an
unbranched trunk.
2. There is a correspondence between the disposition
and distribution of the branches, and the disposition and
distribution of the leaf veins. — Some trees, such as the
beech, the poplar, the birch, the oak, have one main
axis, from which there proceed comparatively small side
branches, pretty equably along its length ; and it will be
found in such cases that the leaf (see Figs. 12, 25) has one
central vein, with pretty
equally disposed veins on
either side. Other trees
again, incline rather to
send off, at a particular
height, for each species,
a number of branches at
once. This is the case with
the lime, the common
^ -3^f,^%r\t X^SF^ -/ sycamore, and the horse-
chestnut. The lime has
a few feet of unbranched
trunk, and at the place at
which it begins to branch
there will commonly be
noticed a cluster of branches, which, as they droop down
give to that tree its attractive shade, and, in correspond-
ence with this, we may observe that the leaf has a petiole,
Fig. 25.:
* Lime leaf, as example of a leaf with a leaf-stalk, shewing a clustering or whorling
of veins at the point at which the veins begin to come off, and a nearly parallel venation.
The angles made by the lateral veins from the midrib are 42°, those made by the veins
proceeding from these main lateral veins are 50°. The angle a a corresponds to the
angle of the peduncle upon the branch.
IN THE ORGANS OF PLANTS. Ill
at the top of which there is a clustering of lateral veins.
The trunk of the sycamore, about eight or ten feet above
the surface of the ground, commonly divides itself into
four or five large branches, and, in precise analogy, we
find a pretty long leaf-stalk dividing into five midribs.
The horse-chestnut often sends off at the top of its bare
trunk (as may be seen in hundreds of trees in Cushy
Park) a still greater number of branches, and in corre-
spondence with this its leaf is commonly divided into
seven leaflets. This correspondence between branching
and venation is very strikingly displayed in those plants
which have triplet leaves, such as broom and laburnum ;
a careful observation of a number of such will satisfy
any one that the axis, in the one a few inches, and in
the other a few feet, above the ground, inclines to divide
into three main branches. In some plants there is a
whorling (approximately in the sense explained, p. 96) of
leaves at the point at which they issue from the stem ;
this may be seen in rhododendrons, the common barber-
ries, and azelias. In these plants the branches also go
off in whorls, as any one may satisfy himself by the most
cursory inspection.
This morphological correspondence may be seen in her-
baceous plants as well as in plants with woody structure.
We have the triplet leaf and triplet stalk in marsh-trefoil,
in wood-sorrel, and clover, and the wliorled stalks, with
a clustering of leaves or midribs, in lady's-mantle, gera-
nium, mallow, and lupin. In common lady's-mantle there
are several midribs, and, in the mountain species, a
whorling of leaves, and in both a tendency to whorl in the
stalks.
So far as we have been able to generalize a very exten-
sive series of facts before us, we are inclined to lay down
the provisional law, that the whole leafage coming out at
112 TRACES OF ORDER
one place on the stem corresponds to the whole plant,
and that the venation of each single leaf corresponds to
the ramification of a branch. We state the general
axiom in this form, because in many plants more leaves
than one issue from one point, and in such cases it seems
to be not the single leaf but the whole leafage which is
the type of the tree.
3. There is a correspondence bctiveen the angle at
ivhich the branch goes off and that at which the lateral
veins go off. — And here, again, it will be needful to have
it admitted that there is a normal angle both for the
lateral leaf veins and the lateral branches. So far as
the lateral veins are concerned, it will be acknowledged,
by every one who has ever looked with care at the form
of a leaf, that . there is a normal angle for every species
of plant. An inspection of any leaf picked up at random
will shew that the lateral veins run nearly parallel to
each other (see cherry leaf, Fig. 12, beech leaf, Fig. 24,
lime leaf, Fig. 25), and that in certain plants, as at the
base of the poplar leaf, for example (see Fig. 26), the
veins so off at a much more obtuse angle than in certain
other plants, as, for instance, the lime (see Fig. 25). The
leaves of the elm and hazel have some resemblance to
each other, but may at once be distinguished by their
angle of venation, which in the former is 55°, and in the
latter 40°. It will not be so readily allowed that there
is a normal angle at which the branch goes off in every
species of tree. We have heard it maintained that a
branch sets off from the axis as best it can, taking any
empty space available to it, and in search of air to
breathe in and sun to warm it. But a very little careful
observation, with this special object in view, may satisfy
any candid mind that this is not the case, and that the
branches tend to go out very much parallel to each other,
IN THE ORGANS OF PLANTS. 113
and at a normal angle, for every species. Any one may
see at a glance that the elm and oak branch goes off at
a wider angle than that of the birch or beech. No doubt
many external circumstances tend to interfere with this
internal tendency. A branch will often be bent clown
by its own weight, or turned upward by another branch,
or by want of room and air, or spoiled by cattle or chil-
dren, or men, still the tendency will manifest itself, even
when thwarted ; and in every open lawn, duly sheltered,
there will be trees whose skeleton beautifully displays the
native direction of the branches, which will be seen, like
the leaf veins, to run very much parallel to each other,
and within a few degrees — now on the one side and now
on the other side — of an average or normal angle, which
may be ascertained by a number of measurements.
When it is acknowledged that there is a normal
angle both for vein and branch, what we may call the
angle of venation and the angle of ramification, the
cjuestion is started, and admits of being answered, Is
there a correspondence between the two ? We may
satisfy ourselves that there is, in a general way, by a
bare inspection, or by taking a leaf, abstracting its soft
matter, and then looking through the skeleton venation
upon the ramification of the branches, and comparing
the two. Or the point may be settled more scientifically
by using a goniometer, consisting of a graduated semi-
circle with a movable index, and measuring the angle
both of vein and branch. The angle of the vein is easily
ascertained, and the angle of the branch may also be
obtained approximately by taking a series of measure-
ments and striking the average. By such means it can
be proven that there is a correspondence between the
angle of venation and ramification of each species of
plants. In most plants with woody structure the angle
114
TRACES OF ORDER
of both vein and branch ranges between 45° and 60°.
In the greater number of herbaceous plants* the angle
varies from 25° to 40°. But both in trees and herba-
ceous plants there are angles so acute as 10° or 15°, and
so obtuse as 70° or 75°. So far as we have observed
there are no normal angles more obtuse than the number
last named, though branches are often bent down by their
own weight, so as to stand perpendicular to the axis, or
are even inclined at an acute angle to the part of the
trunk below the point from which the branch springs.
For every species of plant which we have examined
there is a definite normal angle or angles. We say
angles, for the angles may
differ at different parts of the
venation and ramification of
the same plant. Thus, in
some plants the angle of ve-
nation is widest at the base,
and gradually narrows as we
ascend. Whenever this is
the case in the leaf .there is
a similar narrowing in the
ans;le of the ramification of
the branches. It is seen in
a marked manner in the pop-
lar and the beech, and helps
to give to the leaves and coma
of these trees that rounded
pyramidal form by which they are distinguished. More
frequently there is a difference between the angle at
* Dr. M'Cosh has here to express his obligation to a most excellent bnt extremely
modest man, Mr. Mitchell, formerly schoolmaster at Edzell, now in the City Mission.
Edinburgh, for help in applying his theory to herbaceous plants.
t Fig. 26. Poplar leaf, as an example of leaf with leaf-stalk. The primary angle of
venation begins at 70°, .and lessens as we ascend the midrib.
Fig. 2G.+
IN THE ORGANS OF PLANTS. 115
which the main lateral veins and main lateral branches
go off from the midrib or axis, and that at which the
lesser veins and branches go off either from the midrib
or axis, or from the main lateral veins and branches.
We may call the former of these, that is, the angle made
by mail veins and branches, the primary angle, and that
made by lesser branches and veins, the secondary angle.
In looking at the lime leaf, {Fig. 25,) we may notice
that the main veins (primary) go off at a much more
acute angle than certain smaller veins (secondary).
Using this nomenclature, we have found that the primary
angle of the venation of the leaf is the same with the
primary angle of the ramification of the stem, and that
the secondary angle of venation is the same as the secon-
dary angle of ramification.
In measuring angles, then, it will be necessary to dis-
tinguish between the primary and secondary angles of
ramification and venation. In applying this distinction
to herbaceous plants, we found that the angle at which
the peduncle, that is, the flower-stalk, goes off, corresponds
not to the primary, but secondary, angle of venation. In
following out this system, however, we often experience
a difficulty in ascertaining whether we are measuring the
angle of a true branch, or peduncle, as botanists do not
seem to have laid down any rules to enable us to distin-
guish between these organs.
It appears, then, that on inspecting the ramification
and venation of any given plant, we may observe a nor-
mal primary angle which is the same both for main
lateral vein and main lateral branch, and also a secondary
normal angle for the lesser veins, (whether proceeding
from the lateral veins or from the midrib,) and for the
lesser branches, including the peduncle. This secondary
angle is in some few plants more acute than the primary,
116
TRACES OF ORDER
Thus in the common dock the primary angle is 60°, and
the secondary angle of flower-stalk and lesser veins about
40°. But in most plants the secondary angle is the more
obtuse, and helps, when it is so, to give to the tree its out-
spreading appearance. Thus in the lime (see Fig. 25)
this primary angle is 42', and the secondary about 50 ,
and in the oak the primary is 50", and the secondary
angle about 65°. The following may serve as examples
of the angles of venation and ramification in some of out
common plants :* —
Plants with Woody Structure.
Deg. Deo.
Alder, . . . .50 Laurustinus, . . . 50-55
Ash 60 Lime, 42
Bay Laurel, . . . 50-60 Small veins and branches, 50-55
Beech, . . . 45-4S Maple, .... 40-45
Birch, . . . . - . 30-48 Lesser veins and branches, 55
Box, .... 60 Mountain Ash, . . .45
Cherry, . . . .50 Oak, large branches, . . 50
Elm, 50-55 Smaller veins and branches, 65-70
Hazel, . . . .42 Rhododendron, . .60
Holly, 55 Rose, 60
Horse Chestnut, . . .50 Sycamore, .... 50-55
Laburnum, . . . .60 Willow, angle varies in each species.
Herbaceous Plants.
Primary Angle.
Deg.
Secondary Angle.
Deg.
Chenopodium glaucum,
urbicum, .
40
40
50
40
Geranium, .
35
1
varies in each species,
in some, 60
Geum intermedium,
30
30
montanum,
35-38
50
Ranunculus,
25-28
differs hi each species.
Scrophularia aquatica, .
nodosa,
40
40
50-60
50
Sinapsis nigra,
Valeriana officinalis,
40-45
30 (vein)
50-55
45
* We are willing to admit that in following out these views, anomalies will present
IN THE OKGANS OF PLANTS. 117
We have found that the angle of the peduncle seems
specially to correspond to the angle made by a vein
coming forth near the top of the main lateral veins.
(See a in Fig. 25.) Let us here recall the doctrine pre-
viously enunciated, (see p. 95,) that the pistil is a leaf
folded on itself, as may be seen very evidently in a pea-
pod.* If we inspect the interior of such a pea-pod, we
shall find the seed coming off. very obviously from the
top of a lateral vein. We now see that in the leaf a
lesser vein, bearing seed in the pea-pod, corresponds in
the ramification to the peduncle bearing the flower.
This completes the correspondence between the leaf and
the plant on which it grows.
4. Tliere is a correspondence between the curve of
the vein and the curve of the corresponding branches. —
Here we must once more insist that every vein and
every branch has its normal curve. We have not been
able to express this curve in a mathematical formula, but
the eye testifies that it has a mathematical regularity,
and that there is a correspondence between the curve of
the vein and that of the branch.
The parts whose disposition and direction we have
been examining, are those which are chiefly instrumental
in giving their normal figure to the plant and its leaf ;
and as the part in the leaf has a correspondence with the
part in the branch, it follows that there may be a certain
correspondence between the form assumed by the leaf
and that towards which the whole tree tends. We use
themselves. Thus, in plants with decurrent leaves, such as thistles, the decurrency of
the leaf seems often to make the angle of the vein wider than that of the branch. In
the Looibardy poplar the angle of the branches seems to correspond not to the primary
but secondary angle of venation. These anomalies will turn out to be as instructive as
the more regular phenomena.
* Schleiden, it may be proper to mention, considers this to be a stem pistil. There is
a point here, a transition point, at which the correspondence between leaf and branch
becomes very close and visible.
118 TRACES OF ORDER
the restricted language, may be a certain correspondence,,
for there are special circumstances which may modify
the forms of leaf or plant, and make them differ from
each other. Where the leaves are pinnate — that is,
arranged like the barbs of a feather along a common
axis, there is no resemblance between the leafage and
whole plant.* This does not prove that leaf venation
and branch are not homotypal, any more than the differ-
ence between the fore and hind limbs in animals shews
that these two parts are not homotypal. And in not a
few cases the general resemblance between plant and
tree is very visible, as — to take the trees whose outline
strikes the eye, and prints itself on the fancy in all our
landscapes — -in the swelling lime, and the spreading elm,
and the heavy-topped oak ; and trees which stand upon
an unbranched stalk, as the sycamore, with sturdy ribs
and swelling chest, and the birch and poplar, with their
coma first rotund, and then tapering gracefully to a
point. In not a few plants the correspondence becomes
minutely, we had almost said ludicrously, exact, and may
be detected in the most trivial particulars. Thus the
stems of some trees, such as the thorn and laburnum, are
not straight, and the branches have a twisted form ; and
in these plants the venation is not straight, and the
leafage is not in one plane — in this respect very unlike
the beech. But in the beech there is a no less curious
correspondence, for the stems take a turn at every
node at which they send off a branch, and the mid-
* We would not speak on this subject with confidence, but it seems to us that when
the leaf is pinnate, the tree is decomposite -that is, instead of sending up one main axis
(like the beech, the poplar, Arc.) from bottom to top, it sends off in a scattered way, as it
ascends branch after branch, till the axis is lost. We have noticed this in ash, mountain
ash, walnut, Mahonias, Acacias, and also Ailanthus glandulosus, Gymnocladus Canad-
ensis, Koelcreuteria, Sophora Japonica, and Robinias, (R. pseudo-acacia and R. viscosa,)
&c. We have also noticed a frequent, though, we suspect, by no means invaiiable,
connexion between the doubly pinnate leaf and the umbelliferous structure.
IN THE ORGANS OF PLANTS. 119
rib of the leaf has a similar zigzag appearance. (See
Fig. 24.)
Such points as these should be carefully noticed and
attended to by the landscape painter and by the pruner.
When the commonwealth of taste is properly constituted,
one of its first laws will be passed against the common
mode of pruning, which cuts trees into all sorts of un-
natural shapes, and in particular, pays no regard to each
plant's peculiarity of beauty. We can excuse the old
Scotch Earl who planted his trees in groups, to represent
the position of the troops which gained a victory under
him, for if he thereby spoiled the beauties of nature, he
at least imparted some knowledge of military art ; but
those who, in ornamental lawns, form spherical yews and
conical laurels — those who force plants to resemble beasts,
birds, or fishes — those who give the oak or elm a bare
stalk — those who cut over a poplar to make it bushy from
the base — those who break off the triplet from the broom
or laburnum, or deprive the lime, or the chestnut, or the
sycamore of its whorl, should themselves, on the prin-
ciple of exacting one member for another, be subjected
to a similar pruning process, and this because of the
offence which they commit against nature without and
nature within them.
These observations apply to plants which have leaves
veined, unfolded, and presenting a surfaco to the eye.
We now turn to plants which have needle-shaped or
linear leaves. Such leaves correspond, we believe, to the
individual stems proceeding from the axis or branches.
But our observations have been confined to the great
family of the Coniferse, so called because their seed-
vessels are cone-shaped. In what follows, our illustra-
tions are to be taken chiefly from pines and firs., the
120
TRACES OF ORDER
only portions of the large family of cone-bearers which
we have had an opportunity of carefully inspecting.
It is obvious, on the most cursory observation, that a
unity runs through the whole of the structure of each of
this tribe of plants. We may notice first, how the ap-
pendages are regularly arranged in a series of whorls
(using this phrase in the loose sense previously explained)
along the whole axis. There is, first of all, a whorling
in the arrangement of the cotyledons, or first springing
leaves. Some botanists have represented the cotyledons
of the Conifene as numerous ; others are inclined to
think that there are only two cotyledons, and that each
of these is cleft into a number of parts ; all agree that
the parts arc whorled. Looking to the axis above ground,
we observe the same arrangement repeated in the
branches, which come out at the nodes in a succession of
whorls from the base to the top cf the axis. Every node
and internode of the pine is of the same construction as
eveiy other.
We may notice further, how the whole tree, composed
of stem and branches, is made, by the evidently prede-
termined arrangement of these parts, to assume in its
outline a most elegant figure. The form is that of the
cone, rounded off gracefully at the base. We are aware
that in many cases the lower branches, especially if eaten
by cattle, fall off as the tree grows old, and show a bare
trunk surmounted by a bushy top : thus, when the lower
branches of the broad-topped stone pine fall away, we
have that picturesque, umbrella-shaped figure, which so
often appears in Italian scenery and Italian paintings.
But, in its natural and normal shape, every cone-bearer
seems to be feathered from near the root. It is interest-
ing to notice, that if we were to intersect the tree hori-
zontally at any one node, the part cut off above would
IN THE ORGANS OF PLANTS. 121
always be a cone, somewhat similar in shape to the whole
tree. This, no doubt, results from the nature of the
cone as a mathematical figure ; but on noticing the fact,
we are the more impressed with the peculiar fitness of
the pre-arrangement which makes stem and branches
produce so perfect a figure. While the whole family
affect this general form, we may observe that every spe-
cies takes a shape of its own, so that we can at once
determine what it is at a considerable distance. Some,
like the common spruce fir, have a sharp apex, and look
as if they pointed to heaven ; while others, like the stone
pine, are broad and bushy, and look as if they delighted
to embrace and shelter the earth. There is one beauty
of the finely-proportioned cluster pine, another beauty of
the sturdy Scotch fir, another beauty of the tapering-
Douglas fir, another beauty of the graceful Weymouth,
another beauty of the shaggy Montezuma, and another
beauty of the brawny Coulter pine, as he flings out his
arms so powerfully. No attempt should be made, by
cutting or bending, to make any one species take the
form of any other ; all such officious meddling, on the
part of man, will only mar the beauty of the Divine
workmanship. A lawn is fairest to look on when differ-
ent species are planted on it, when each is allowed to
grow naturally, and has room allotted to it to shew its
peculiar type and beauty.
Turning now to the inspection of the seed-vessels, we
find them, as their name (cones) denotes, moulded after
the same form ; nay, the very clusters or bunches of
stamens (amenta) are made to assume a conical shape.
It is evident that, in this tribe of plants, there is a sig-
nificancy in this beautiful mathematical figure ; and we
are inclined to ask whether it was not some mystic per-
ception of this which led the ancient Assyrians to assign
6
122
TRACES OF ORDER
so important a place to the cone in those sacred symbols
which have become so familiar to us by the researches of
Dr. Layard ? But without insisting on this, we think
we are justified in affirming that the circumstance that
the cones, formed of scales, which are modified leaves,
and amenta, which are also formed of modified leaves,
taking the same shape as the tree, formed of branches, is
another illustration of the tendency of leaf and branch to
obey the same laws and follow the same dispositions.
Not only is there a general resemblance between cone
and tree ; we are inclined to think, from a pretty exten-
sive observation, that the full-grown and expanded cone
not unfrequently tends to take the shape of the particular
species of tree on which it grows. It would require a
series of measurements, such as we have not had it in our
power to make, to establish this truth scientifically, but a
general correspondence is often obvious
to the eye. Thus, to take some of the
species most marked in themselves, and
best known among us. The common
Norway spruce is tall in proportion to
its width, and so also its cone. (See
Fig. 27.) The same may be said of
Abies Douglasii, which, moreover, has
a sharp apex ; the cone tends to assume
the same shape. In striking contrast
is the stone pine, (Pinus Pinea, see
Fig. 28,) in which both cone and tree
are wide in proportion to their height.
The cluster pine (P. Pinaster) is beau-
tifully proportioned in its length and breadth, both in
Fig. 27.*
* Fia. 27. Cone of Abies excelsa, bearing a resemblance to a tree, and shewing a set
of spirals from right to left, and another set from left to right. These sets of spirals,
crossing each other, produce on tho surface of the cone rho:nbol ':il figures.
IN THE ORGANS OF PLANTS.
123
tree and cone. (See Fig. 29.) Contrasted with this, both
the tree and cone of Pimis pumila have a crushed ap-
Fig. 28*
Fig. 29.+
pearance. Coulter's pine has heavy, wide-spreading
branches, and its cone is of a rotund, bulky shape. The
cone of the Labrador pine (P. Banksiana) is often bulged
at one of the sides, and any of the trees which we have
seen have a straggling, misshapen appearance.
We now proceed to give the result of a series of obser-
vations"]: in regard to the dispositions of the scales of the
cone and the leaves of the tree.
(1.) The scales are arranged along the axis of the cone
* Fig. 2S. Cone of Finns Pinea; clumpy like the tree, shewing the two sets of spirals
crossing each other, and producing rhomboldal figures, whose angles are approximately
above and below 120° and 00° at each of the sides.
+ Fxg. 29. Cone of Pinus Pinaster, shewing the two sets of spirals, and rhomboids
with definite angles.
X In making these observations, Br. M'Cosh has examined, with more or less care,
nearly every cone In the Museums of Kew, of the Linnsean Society, and of the Fdiri
burgh Botanic Garden, all of which have been kindly thrown open to him.
124 TRACES OF ORDER
in a spiral manner. As the basis of the whole, there
seems to be a governing spiral — that is, the scales are
attached to the axis in a regular spiral. This spiral is
at times from right to left, and at other times from left
to right, and we have not been able to discover any law
determining which of these courses it should pursue ; it
certainly is not determined by its position on the tree, or
by the course of the sun in the heavens. The scales in
this spiral being at equal distances, necessitate mathe-
matically other three spirals, or four spirals in all — one
of these, the governing spiral, and another, running in
the same direction, and other two in the opposite direc-
tion. Sometimes all of these spirals can be noticed ; in
all cases two are very visible, one from right to left, the
other from left to right. (See Figs. 27, 28, 29.)
On comparing the cone with the branch, we find a dis-
position in the appendages of the latter similar to those
of the former. The leaves on the young stem and the
scars left when these leaves fall off, form invariably two
sets of visible spirals, one from right to left, and the other
from left to right.
(2.) The two sets of visible spirals form, by their in-
tersection, a series of very beautiful and mathematically
regular rhomboidal figures on the surface of the cone.
(See Figs. 27, 28, 29.)
The elegance of the whole figure, with these spiral
gjTations, which allure on the eye, and these well-defined
lozenge shapes on the surface, form the ground, if we do
not mistake, of children's predilection for cones. When
they gather these so eagerly and industriously, when they
play with them for such a length of time, it must be be-
cause of some unconscious perception of the visible har-
monies— a perception which they could not of couise
scientifically expound, or even express to others. And it
IN THE ORGANS OF PLANTS. 125
would be well for us in this, as in many other cases, not
contemptuously to cast away the simple tastes of our
childhood, but rather to cherish them, and put them
meanwhile under the guidance of a matured understand-
ing. A pine-cone will reward the study for hours toge-
ther of the very highest intellect. Here, as in numerous
other instances, science, in following up our spontaneous
tastes, will unfold wonders on which the reason gazes with
profound interest.
If we measure these rhomboidal figures on the surface
of the cones of pines and firs, we find that the angles are
definite, being approximately 120° above and below, and
60° at the sides. (See Figs. 27, 28, 29.) We use the
language approximately, because there is often, as might
be expected, a departure from the normal angle on the
one side or the other, but the actual angles stick so
closely, on the one side or other, to the numbers given,
that we may regard these as the normal ones. The
eye, or rather the intellect, feels a pleasure in contem-
plating such a figure, made up of two equilateral triangles,
and in every respect so beautifully proportioned, and com-
bining an easily observable unity with an easily observable
variety.
On the stems likewise, the intersection of the two spi-
rals formed by leaves, and the scars of fallen leaves, forms
a series of rhomboids. We cannot speak so confidently
of the angles of these rhomboids as of those of the cone,
but we have found in many cases that when the leaves
have fallen off and the scars are visible, the angles at two
of the opposite corners arc approximately 120°, and at the
other two opposite corners 60°. But there is this differ-
ence between the rhomboids on the cone and the rhom-
boids on the stem, that whereas in the former the angle
is 120° above and below, and 60° at the sides, in the
126 TRACES OF OKDER
latter the angle is 120° at the sides, and 60° above and
below. We have found these numbers very often on the
stem of a few years old ; as it becomes older the rhom-
boid is less elongated, but by this time the scars are be-
ginning to disappear, being covered up with the bark.
This arrangement produces on the surface of the cone
a series of quincunxes, a figure which has long been re-
garded as possessing many virtues. Virgil, in his Georgics,
in giving directions for planting trees, says, " Indulge or
dinibus," and recommends the quincunx.
" Omnia sint paribus numeris dimensa viarum,
Non animum modo uti pascat prospectus inanera,
Sed quia non aliter vires dabit omnibus sequas
Terra."
VlRG. GeORG. II., 284-286.
Speaking of the same figure, Quintilian says, " Quid
quincunce speciosius qui in quamcunque partem specta-
veris rectus est." Sir Thomas Browne, in his ingenious
though fanciful work, entitled " The Quincunx Mystically
Considered," seems to have had pleasant glimpses of the
truths to be discovered by the study of the cone-bearers,
" Now, if for this order we affect coniferous and tapering
trees, particularly the cypress, which grows in a conical
figure, we have found a tree not only of great ornament,
but, in its essentials of affinity unto this order, a solid
rhombus, being made by the conversion of two equicru-
ral cones, as Archimedes hath deponed. But these were
the common trees about Babylon and the East, whereof
the ark was made." " But," he adds, " the Firr and Pine
Tree do naturally dictate this position ; the rhomboidal
protuberances in pine-apples maintaining this quincun-
cial order into each other, and each rhombus in itself.
Thus are also disposed the triangular foliations in the
IN THE ORGANS OF PLANTS. 127
conical fruit of the Firr tree, orderly shadowing and pro-
tecting the winged seeds below them/'
(3.) There is, we have said, a very visible set of spirals
going from right to left, and another very visible set
from left to right on the surface of the cone. In these
sets there is a definite number of spirals. We propose,
in the absence of an authorized word, calling the parts or
numbers of a set of spirals, threads. The number of
threads, in a set of spirals in all coniferte, seems some
one of the following numbers, 1, 2, 3, 5, 8, 13, 21, 34,
&c, in which scale any two contiguous numbers added
together gives the succeeding one. We have already
fallen in with this remarkable series of numbers in leaf
arrangement ; it now casts up once more in a somewhat
different, and probably a more fundamental relation. In
the case before considered, these numbers were merely
the more common ones ; in the case now before us they
seem to be invariable ones. On the supposition that the
spiral, more or less modified, governs the arrangement of
the appendages of all plants, we are inclined to look on
this series as having a deep significancy in the morpho-
logy of the plant.
We have observed that there is a constant relation in
the number of threads in the two sets of spirals. What-
ever the number of threads in the one set of spirals — ■
say that proceeding from right to left — the number in
the other — those proceeding from left to right — is always
one or other of the contiguous ones in the above scale.
The number of spirals in the two intersecting sets are
1 and 2, or 2 and 3, or 5 and 8, or 8 and 13, or 13 and
21, or 21 and 34. Thus, if the number of threads in
the one set of spirals — say that proceeding from right to
left — is 5, the number in the other set — that proceeding
from left to right — will be either 3 or 8. These nume-
128
TRACES OF ORDER
Fig. 30 *
rical relations seem to regulate the sets of spirals in all
coniferee. In pines the number of threads in by far the
greater number of species, is 5 and 8. In a few species
the numbers are 3 and 5, and in a few others 8 and 13
In Araucauria imbricata
the number of threads in
the two sets respectively is
21 and 34.
In the disposition of the
scars on the stems there are
similar numerical relations.
Thus the number of threads
is one or other of the num-
bers in the scale, 1, 2, 3, 5,
8, &c., and the numbers of
the two sets are always contiguous ones in this scale.
Thus, if the number of threads in the one set of spirals
is 3, the number in the other will either be 2 or 5. We
have remarked, however, that the number of threads in
the spirals of the branch is commonly less or lower in
the scale than the number of threads in the spiral of the
cone. In pines the common numbers for the cone are 5
and 8, whereas the numbers for the visible spirals on the
stem are 5 and 3.
(4.) We have found in the cones of pines and firs, (so
far as we have examined them,) that all the spirals in
one of the sets, and this invariably the one which con-
tains the greater number of threads, take approximately
just one turn in going from the base to the top of the
cone, that is, each goes round the axis once, and stops at
the apex perpendicularly over the point from which it
* Fig. 30. Diagram shewing that two sets of spirals set out from the base of a cone,
and that there is a relation between the number of spirals in the two sets. In the dia-
gram the number proceeding from left to right is 5, and the number from right to left
IN THE ORGANS OF PLANTS. 129
started. Thus, if the spirals be 8 and 5, (as in Fig.
30,) then each of the 8 Avill be found to have taken one
complete turn before it reaches the apex, and if, the
numbers be 13 and 8, the 13 will be found to have
twisted themselves once round the axis. This seems to
be the rule followed by the set of spirals containing the
larger number. The other set appears also to have a rule.
In cones with the ordinary relation between the height
and width, that is, where the circumference is greater
than the height, the number of turns made by the set of
spirals of the lower number is 2, that is, the spirals go
twice round the axis before reaching the apex. But in
cones whose height is great in proportion to their width,
whose length is greater than their circumference, as,
for example, Pinus Strobus, Pinaster, excelsa, monticola,
Lambertiana, filifolia, and Abies alba, excelsa, Douglasii,
the number of turns taken by the spirals is 3.
Such co-ordinated facts as these may possess little
interest to the mere technical naturalist, whose sole aim
is to discover new genera and species, or the mere prac-
tical horticulturist or arboriculturist, whose object is to
find plants of commercial value. But they tend to raise
up profound reflections in the truly philosophical mind,
and open up glimpses to the religious mind of the deep
things of God. They shew that the plant, and all its
members, had been before the mind of God prior to the
time when He said, " Let the earth bring forth grass,
the herb yielding seed, and the fruit-tree yielding fruit
after his kind, whose seed is in itself upon the earth, and
it was so ;" " and God saw that it was good." Mathe-
matical figures, more or less modified to suit special
ends, make their appearance everywhere among the
members of the plant. The mathematical spiral regu-
lates the arrangement of all the appendages of the plant.
6*
130 TRACES OF SPECIAL ADAPTATION
Even the lines which man lias not been able to express
in mathematical formula?, such, as the curve of the veins
and branches, and the outline of the coma of a tree, are
evidently regulated by models in the mind of the Divine
Architect. Numerical relations of a most interesting
character cast up among every class of plants, and among
all the organs of every plant. All appendicular organs,
whether belonging to the nutritive or reproductive sys-
tem, are homotypes. Nay, correspondences may be
detected between the disposition and the distribution of
branches and leaf veins, sufficient to entitle us to repre-
sent root, stem, and leaf, as homotypes, and to prove
that there is a unity of composition in the structure of
the whole plant.
SECT. II. TRACES OF SPECIAL ADAPTATION IN THE ORGANS OF
THE PLANT.
Our aim in this chapter is to shew that in the struc-
ture of the plant there are combined simplicity of general
plan and variety of modification, the latter for special
ends. Having endeavoured in the preceding section to
demonstrate the first great truth, we are in this section
to illustrate the second.
It is evident that stem and common leaves would not
suffice to fit the plant for the discharge of all its func-
tions. It needs, among others, organs or appendages for
covering, for support, and for enabling it to propagate
and perpetuate itself. To meet these wants members
are found to spring up at the very place where they are
needed, and at the very time when they are needed ;
and when they appear they come not as absolutely new
organs, but after the old type, modified to serve the
present purpose. Does the plant demand a covering ? —
IN THE ORGANS OF THE PLANT. 131
the leaf becomes a scale, or the cuticle produces hairs for
that purpose. Is defence required against external
attack ? — leaves or branches become sharpened or hard-
ened at the point, and the whole plant, or the more
assailable parts of it, are bristled all over with spines or
prickles. That the species may live on in a new indi-
vidual, the leaf takes a yet greater departure from its
type, and becomes a stamen or pistil. The general plan
of the Great Architect is kept up, and yet every several
member fulfils a purpose. We cannot conceive of stronger
or more convincing evidence of design being supplied to
human intelligence.
1. ORGANS OF VEGETATION.
The general structure of the leaf has been already
described ; we are now to contemplate some well-marked
special modifications. The cuticle, or skin, shews nume-
rous small openings, (the stomata of botanists ;) these,
like the holes in a barn, keep up the communication
between the air and the interior. In the leaves of aerial
plants, which have the usual horizontal position, these
pores are commonly abundant upon the lower surface,
and upon that under surface the skin is also of a more
delicate nature ; on the upper surface the stomata are
usually less numerous, or even, in some cases, wanting,
while the skin is tougher and denser. In leaves, again,
which float on the surface of the water, the openings are
confined to the upper surface, and in submerged leaves
they are wanting altogether. The intervening portion of
the leaf, already described, called parenchyma, presents
some remarkable peculiarities in relation to the pores
we have been describing. Next the upper surface of
the leaf, it consists of compact oblong cells, placed per-
pendicularly and in close contact with each other, the
132
TKACES OF SPECIAL ADAPTATION
layer nearest the lower surface is less dense, and nume-
rous vacant spaces occur between the cells, permitting free
communication, through the stomata or pores, between
the atmosphere and
the interior of the
leaf. We have here,
therefore, a striking
example of harmony
between the struc-
ture of this part of
the plant and its
function and po-
sition. The pores
are exhaling and
absorbing organs ;
where they are most
abundant, there we iind loose texture of the parenchyma,
permitting free communication ; where stomata are not
needed, they are wanting ; when they are required on
a particular part of the leaf, there we find them. Many
species of Utricularise — delicate water-plants — have nu-
merous small sacs connected with the leaves, which are
stated, about the flowering period, to become filled with
air, and to buoy the plant near the surface of the water.
In Pontederia crassipes and Trapa natans, some of the
leaves have the stalk dilated into an air cavity, which
acts as a float. The magnificent Victoria Kegia presents
several interesting features. It is an aquatic belonging to
the water-lily family, and the fully developed leaf reaches
a diameter of five feet or more. In order to give
strength to such a large surface, the veins on the lower
aspect of the leaf are of great depth, acting as so many
* Fig. 31. Perpendicular section of leaf, to shew different structures of upper and
lower portions.
IN THE OKGANS OF THE PLANT. 133
supporting girders. Between them are formed spaces in
which air might accumulate and lead to a rupture of the
parts ; such an occurrence, however, is obviated by the
perforations which constitute one of the peculiarities of
this remarkable plant. By transmitted light the leaf
resembles a sieve, with numerous minute openings.
Stipules. — These appendages assume different forms,
and vary in size and texture, according to the plant in
which we examine them. They are, as already stated,
formed after the leaf type, and although we cannot, in
every case, point out the purposes served by their modi-
fied form, there are, nevertheless, instances in which we
cannot doubt that they are present for a useful object.
In Lathyrus aphaca they are of large size, and supply
the place of the leaves, which are absent in the mature
plant. In not a few plants they perform important
functions as protecting organs, forming a covering to
the young leaves ; this is obvious enough in Magnolia,
in the Indian-rubber fig, and in the submerged Pota-
mogetons. When the leaves expand, these protective
stipules fall off ; their function being performed they are
no longer needed, and so they disappear.
Covering of plants. — The varied aspect of the external
surface of the different organs of plants, so important to
the botanist in the distinction of species, and designated
by the terms downy, silky, scaly, &c., is owing to the
presence of certain minute appendages, the nature of
which has been already described. (See p. 86.) In cer-
tain cases their presence has some relation to the habitat
or dwelling-place of the plant. Those on the upper part
of the pistil of hare-bell are well-known to act in col-
lecting and retaining the pollen grains as they drop
from the anther. There can be no doubt that in many
cases the very minute fibrils on the underground parts
134 TRACES OF SPECIAL ADAPTATION
of plants, which assist in the process of absorption, are
really hairs, and of the same nature of those which cover
aerial organs.
Armature. — Plants, like animals, have been provided
with organs of defence, varying in strength and in the
effects left by them from the simple and almost innocu-
ous prickle of the rose to the formidable sting of Urtica
urentissima, the wounds inflicted by which often lead to
dangerous or even fatal results. We have shewn in last
section that under the term Armature are comprehended
modifications of several parts. The spines of the white
Thorn and the black Thorn, of which every one has had
experience, are branches turned into spear points, to
repel all sensitive assailants. In Barberry certain leaves
have been sharpened into prickles ; in Holly the leaves
have had their secondary veins hardened and pointed
effectually — as the mouth of any animal which may at-
tempt to eat them will testify. In Kobinia the stipules
have undergone a change of condition, to fit them for
a similar defensive function. The armature of Nettle
and Loasa are modified hairs, as are also the prickles of
the rose, and many other plants.
Supports. — This term comprises various modified
organs, supplying instances of design as palpable as any
furnished by the pillars and buttresses of human archi-
tecture. The native tendency of the stem is upwards,
but there are multitudes of plants too weak to retain
their vertical position ; and to aid them in their heaven-
ward inclination various provisions have been made. At
times the stem itself becomes twined round other plants ;
this spiral twining may either be from right to left, as
in the French bean, Dodder, Convolvulus, &c. ; or from
left to right, as in the Honeysuckle and Hop. At other
times the same end is accomplished by the superficial
IN THE OKGANS OF THE PLANT. 13D
appendages of the stem, as, for instance, by the minute
hooks on some species of Galium.
Tendrils, varying, as we have seen, in their nature in
different plants, but all really referrible to a common type,
possess the same properties as twining stems ; they twist
themselves round other plants, and thus support species
too weak to stand in their own strength.
In Dischidia Rafflesiana the pitcher -shaded organs are
leaves whose margins have become adherent. This plant
is a climber, sometimes reaching the top of the loftiest
trees, and generally the pitchers are confined to its upper
part. It is stated that there, small roots are developed,
and these, entering the pitchers, absorb the fluid which is
accumulated from the fall of rain or dew ; the long strag-
gling stems are thus provided with a means of receiving
nourishment at both extremities.
In human architecture we may discover contrivance
in the means taken to retain the general symmetry of
the entire edifice, while at the same time every part of
it is devoted to a useful purpose ; and surely the ex-
amples we have given indicate the same kind of lofty
design, contriving to make organs conform themselves
to a general type while they accomplish particular ends
essential to the wellbeing of the plant. The cases
brought forward belong to the nutritive system of
plants ; similar examples are furnished by the reproduc-
tive economy. But before proceeding to examine these,
Bracts may be alluded to, as forming an evident tran-
sition, as we have already shewn, from leaf to sepals, or
divisions of the calyx. They are leaves specially modified,
and may help the parts of the flower in the performance
of their office. This may be laid down as a general
rule, though we may not be able in every case to specify
with precision their peculiar function. There are very
136 TRACES OF SPECIAL ADAPTATION
numerous cases in which they serve as protecting organs.
In Palms, and other plants, the large sheath which they
form, called technically a sjmthe, encloses numerous
flowers as yet in au early stage of development. In
some Palms it is calculated that there may he thus
protected no fewer than 200,000 flowers. In Narcissus,
Allium, &c, the bract forms a protecting sheath to the
flowers while in a young and tender state, and when
these expand it shrivels and decays. In the daisy, and
others of the family Composite, the numerous florets
are protected by one or more series of overlapping
bracts. The cup of the acorn is a protecting organ,
formed also of numerous overlapping organs of the same
nature. Where these parts present much resemblance
to leaves, they often, as in Anemone and other plants,
serve at first as protecting appendages, and subsequently
they aid the leaves in their all-important functions.
II. REPRODUCTIVE SYSTEM.
Calyx. — It is admitted on all hands that the sepals, or
pieces of the calyx, though not present in every instance,
and therefore not absolutely essential in the economy
of the flower, do, when present, perform some good
offices. This is true, whether the pieces, in consequence
of lateral adhesion, are made to take a tubular shape, or
whether they have some other form. There can be no
doubt that, in numerous instances, this part not only pro-
tects the more internal organs, but likewise assists the
leaves in their function. The remarkable, and often very
beautiful, hair-like appendages of the fruit (part of the
calyx modified), in Composite plants, as the dandelion
and others, assist in the dissemination of the fruit and
seed : acted on by the wind, these pappose fruits are
wafted to a distance from the parent plant, and, when
IN THE ORGANS OF THE PLANT. 137
they fall into a suitable soil, become the parents of fresh
colonies.
Corolla. — The general office of this organ is very
obvious. This whorl of petals serves, in most cases, to
support and protect the more vital organs within ; such,
at least, is one function which it evidently performs. It
is all true, that we cannot in every instance state, with a
well-founded confidence, what connexion there is between
the form and colour of the individual piece or of the
entire corolla, and its use in the economy of the plant.
In not a few vegetable organisms, both calyx and corolla
are wanting, and in such cases, at least, they cannot be
essential organs ; but when present, we may believe that
they serve a purpose. It is supposed by some that there
is a relation between the colour of the petal and the
measure of heat which it absorbs, and which the flower
requires. Possibly there may also be a relation between
the form of the corolla and the process of seed or fruit
production in the species ; but science is not yet prepared
to point it out. The brilliant apparatus of the flower
acts, we are convinced, as an attraction to various kinds
of insects, which, in the act of procuring food for them-
selves, assist also in scattering the fertilizing pollen, and
bringing it into contact with the upper part of the seed-
vessel. If we need to seek for any other final cause, we
shall find it in the shapes and colours of flowers, as ad-
dressed to that love of the beautiful which is one of the
most bountiful parts of the wonderful constitution of our
nature.
Stamens. — These, with the pistil in the centre of the
flower, are the most essential organs of all. Their all-
important office is to produce the pollen or fecundating
powder necessary to the formation of the seed.
We have pointed out, in last section, the homology of
138 TRACES OF SPECIAL ADAPTATION
the stamen and its parts. Its departure from the gene-
ral type is not so great as might at first appear, still it
does deviate widely from the leaf, and all to accomplish
the very special end allotted to it. The filament which
supports the anther is (we have seen) no more essential
to that anther than the stalk is to the leaf. This fila-
ment, however, does at times assume forms which act an
important part in relation to other organs and the gene-
ral mechanism of the plant. Thus in Kalmia, each
anther is kept back by a little hood or hollow in the part
of the corolla opposite to it. A slight force — the touch
of an insect, for example — suffices to release the anther,
when the elastic filament, acting as a spring, brings it
forcibly in contact with the upper part of the pistil, the
pollen meanwhile being freely emitted. The same object
is secured by the elastic filaments of the common nettle,
and the irritable filaments of the barberry. Without
entering more minutely into the subject, it may be ob-
served generally, that the application of a fertilizing
matter being absolutely necessary to the propagation of
the plant, it appears precisely where it is wanted ; the
parts which produce it protect it in the first place, and
aid in the final application of it ; while the whole appa-
ratus is a special modification of the typical member.
Pistil. — The leaf type is here modified to form an all
essential organ. Its functions are to receive and retain
the pollen, and the top of the stigma is admirably fitted
for such purposes. Another part of the same organ con-
ducts the fecundating matter to the seed-buds or ovules,
and affords also to these vital members protection — of a
more temporary or permanent kind — till such time as
they attain maturity, and reach a locality in which they
may germinate into new life. In not a few cases the
seed-vessels have appendages which act as wings, and
IN THE ORGANS OF THE PLANT. 139
waft them by the aid of the wind to distant localities.
In other cases the appendages become floats or protec-
tors, and give us nuts and capsules, which are conveyed
by rivers and ocean currents to establish new colonies far
from the parent stock. The hooks and other appendages
of some fruits make them adhere to the coats of animals,
and thus the plant, stationary itself, has its seed disse-
minated wide as the range of the animal, with its feet
and wings. Other instances are not wanting of evident
adaptations in the general structure of fruits, and in the
properties of their elementary tissues. Thus the ripe
fruits of some species of balsam, when touched, suddenly
burst and scatter the seeds with considerable force ; the
squirting cucumber is a still more remarkable case of
the same description.
Another final cause, different in kind, comes into view
very prominently at this place. He who makes every
organ subserve the welfare of the plant, has also made
the plant, as a whole, and its individual parts, to pro-
mote an ulterior end. It seems very evident to us that
certain modifications of the organ under consideration,
and others contiguous, have a direct reference to the
wants of man and the lower animals. We never can
believe that the sugar, acids, oils, starch, and other pro-
ducts formed so abundantly in the fruits of different
plants, were not meant to serve as food, and afford sen-
tient gratification to the animal creation. The fruit, in
its earlier stages, performs a necessary part in fertiliza-
tion, at every period it yields support and protection to
the young as well as the mature seeds, and when, in ad-
dition to these, it presents its beautiful forms and colours
to the eye of man, and pleases all sentient creation with
its perfumes, and gives satisfaction to the palate, and
nourishment to the frame, we are sure that we have be-
140 TRACES OF SPECIAL ADAPTATION
fore us the modification of an organ for a twofold pur-
pose, the one bearing directly on the economy of the
plant itself, and another, and a further, having a respect
to the wellbeing of the animal world.
And these ends, be it observed, are accomplished in
conformity with the grand regulating principle of type or
pattern. In a ripe cherry the kernel is the seed ; the hard
stone, so admirably fitted for protection, corresponds to
the upper cuticle of the leaf— thus singularly transformed
for a useful end ; the skin of the fruit represents the
cuticle of the lower surface of the leaf ; the intervening
delicious pulp is just an expansion of the cellular sub-
stance previously described as lying between the two
cuticles ; nay, the observant eye will discover that in the
line on one side of the cherry, we have the united edges
of the typical leaf.
Finally, in the Seed. itself, that portable epitome of
the entire vegetable organism, we find differences in the
relative development of different parts, all in decided re-
lation to some special function which has a respect to the
continuance of the species, or the necessities of the ani-
mal creation. We have alluded to the adaptation of
certain parts of fruits to the purpose of protecting the
seed ; but the seed itself has often an independent means
of resisting injury in its hard integument or skin, modi-
fied for that purpose. The wing-like appendages of pine
seeds, and the abundant hairy covering of those of wil-
lows, doubtless aid in their dissemination when they are
committed to aerial currents. It may be added, that man
finds in the covering of the seeds of the cotton plant an
economic product of immense value for his clothing and
comfort.
For the better comprehension of special modifications
in the seed itself, we would refer to previous remarks,
IN THE ORGANS OF THE PLANT. 141
(p. 82.) Generally speaking, we find two obvious con-
trasts in the relative development of the internal parts,
viz., large cotyledons, and the albumen small or absent,
{Fig. S,) and small embryo with copious albumen. In
the economy of each individual seed these differences are
of vital importance. In germination the cotyledons in
some cases (as Lupine) rise above ground, assume a green
colour, and, for a time, perform the functions of leaves,
finally giving place to the true leaves of the new plant,
when these have attained sufficient size ; or the cotyle-
dons may remain under ground during the process of
germination, as in the pea and bean, yielding up to the
young plant the store of nourishment which they contain.
Seeds with copious albumen and small embiyo have, in
like manner, in the former a temporary store of food for
the latter.
But further, those parts of the seed which are of such
importance in the early economy of the young plant, pre-
sent also a new relation, viz., to the existence and well-
being of man and numerous lower animals. Starch, oil,
&c, are products yielded in abundance by seeds ; and
the hard albumen of some, as the ivory-nut, is turned to
a useful purpose in the arts.
To sum up what we have said : — The stem bears up the
whole plant, so that the influence of the sun and atmos-
phere may act through the leaves upon the fluids absorbed
by the roots ; which roots perform the functions of stays,
enabling the whole vegetable organism to resist the action
of such physical agents as wind. The law of the spiral,
which regulates the arrangement of the appendages, seems
to be admirably calculated to expose them to the influences
needful in order to the growth of the whole plant. Cer-
tain of the special modifications are either absolutely
necessary to the existence of the plant, or tend to its
142 TRACES OF SPECIAL ADAPTATION
wellbeing, such as tendrils for support, scales and hairs
for protection, spines and prickles as armature for de-
fence. We also know that some of the varied modifica-
tions of the floral organs, namely, the stamens and pistils,
are essential to the continuance of the race. It is very-
evident, too, that regularity in the arrangement of the
flowers and of their parts will promote the function of
fecundation, and tend to lessen risk of failure in this
important end. Even the more common arrangements
seem as if they were intended to promote the fertilization
of the seed. Thus the stamens of the upper flowers of a
spike, (wheat, for instance,) or of a raceme, (as common
currant,) may not only fecundate the ovules in the flower
to which they belong, hut are also well placed to insure
the fall of the pollen on those which stand below them.
In the spikes of some species of carex or sedge, where
the stamens and pistils are often in separate flowers,
those at the apex of the spike have stamens only, and
those further down pistils only. Again, when species of
carex have some spikes with stamens only, and others
with pistils only, the former often stand highest. These
cases are so uniform and so numerous that we cannot
regard them as mere accidental coincidences. There
are, no doubt, exceptions, but in all such cases the same
end is accomplished by the insects which frequent flowers
in search of food, scattering and conveying from one
flower to another the fecundating pollen. Again, the
regular arrangement of the ovules in the interior of the
seed-vessel, will be more likely to give each a better
chance of receiving the influence of the matter conveyed
by the minute tubes which pass down from the pollen,
than indiscriminate jumbling of the whole.
Thus we observe in the vegetable kingdom that
special ends are served both by the typical organs and
IN" THE OBGANS OF THE PLANT. 143
their distribution, and also by the numerous deviations
from the type, whatever be the nature and extent of
these. It must, however, be frankly acknowledged that
we cannot in every instance discover a final cause for
every particular 'part of every plant, or, at all events,
that our present knowledge does not entitle us to speak
confidently on the subject. But this is not necessary in
order to the validity of our argument. In a building
we may be able to recognise design in its general style,
although not in circumstances to point out the special
purpose which every part of it was intended to serve.
On the same principle we believe that we are entitled to
say that we have discovered marks of design in the
plant as a whole, and in its various modifications, even
when we may not have arrived at a stage of knowledge
which enables us to understand why an organ has
assumed one particular form rather than another. It
would be a very limited range of contemplation if our
attention were confined to the function which individual
parts are intended to perform in the vegetable economy.
We cannot doubt that there is a relation between the
existence of plants and the support of the animal world.
In the grass of the field, and the valuable products
yielded by fruits and seeds, we can see a provision made
by the Creator for supplying the necessities of His
creatures.
We go a step further, and affirm that plants were
meant not only to furnish food to the animal creation,
but were intended to afford them pleasure by their tastes
and by their perfumes. It will surely not be affirmed that
the organs of taste and of smell were given us merely as
means of procuring food, or as sentinels, on guard at the
outposts, to warn us of danger. Plants might have been
less sapid or less odoriferous without any derangement of
144 TRACES OF SPECIAL ADAPTATION
the functions wliicli each part fulfils ; and there is surely
some ground, for concluding that He who planned, and
made them all superadded those qualities, and instituted
a harmony between the sensate and the insensate, for
the gratification of animal tastes. Not only so, we think
there is good ground for affirming that not a few vege-
table forms were meant to gratify the aesthetic feelings of
man. We cannot jdeclare with certainty that the forms
assumed by the flower, by its calyx and corolla, are in
every case necessary to the functions of the plant. We
will not affirm that the beautifully rounded form of the
peach, the delicacy of bloom on the surface, and the
deliciousness of its flavour, are required in order to the
production of the kernel and its hard protecting shell.
We have no reason to think that the brilliant scales on
the wings of the butterfly are necessary to its flight, for
the insect, (as any one may observe) can fly after they
are mostly all rubbed off, and some Lepidoptera have
few or no scales at all ; and just as little ground have
we for affirming that the plant could not fulfil its
functions even though the flower had not been so orna-
mented.
Man has aesthetic tastes implanted in his nature ;
these are gratified to the full by the lovely forms pre-
sented in the vegetable kingdom, and we are convinced
that all this was arranged by Him who conferred on
man his love of the beautiful, and supplied the objects
by which that love is gratified." And here we have
to express our regret that philosophers have not been
able to agree upon a theory of the beautiful. If there
had been any acknowledged doctrine on this subject,
* Some insist tbat there is not only the beautiful in plants, (and in animals as well,)
but also the grotesque. Granted, but surely we have here a further example of final
cause in the relation between the grotesque in the plant and the sense gf the ludicrous
in man.
IN THE ORGANS OF THE PLANT. 145
there would have been little difficulty in shewing that
plants are fashioned in accordance with a very high
style of beauty. In particular, we are as yet without
any generally received principles in regard to what con-
stitutes beauty of form. In such circumstances we can
appeal to no admitted rules, but we can appeal to our
own feelings, which declare that the plant, in its general
form, and in its corolla, exhibits perfect models of
beauty. Here we have an all-sufficient final cause super-
added to all the other final causes, bearing more directly
upon the economy of the plant, and coming in at the
parts, such as the flower and fruit, where these others, to
our eyes, might seem to fail.
CHAPTER III.
THE COLOURS OF PLANTS.
SECT. I. THE RELATIONS OF FORM AND COLOUR IN THE FLOWER.
It is a very common impression that there is no rule,
no law, for the distribution of colours in the vegetable
kingdom.* We are convinced that this is a fundamental
mistake. Little, it is true, has been done to establish
scientific principles as to the colours of plants. Still,
there is reason to believe that system prevails here as in
every other department of nature. Laws in regard to
the form, structure, number, and position of organs, are
familiar to every botanist ; and it is surely not unreason-
able to expect that order may also be found in the placing
of colours. One of us has been able to furnish a con-
tribution to this branch of inquiry, by discovering evi-
dence of a very curious relation between the form and
colour in the corolla in plants.f
In order that this may be understood, it will be neces-
sary at this place to explain certain technical terms used
* We are great admirers of Mr. Buskin's intuitional power, but the following state-
ments in his Lamps of Architecture are too unguarded: — "The natural colour of objects
never follows form, but is arranged on a different principle ;" and again, " Colour is sim-
plified where form is rich, and rice versa;" " In nature," he further says, the "boun-
daries of forms are elegant and precise ; those of colours, though subject to symmetry of
a rude kind, are yet irregular — in blotches."
t See Br. Dickie's Papers in Sectional Reports of Proceedings of British Association,
1854; and Annals of Natural History, Dec. 1854.
COLOUR IN THE FLOWER. 147
by botanists. The terra regular is applied to every calyx
or corolla in which each sepal or petal is of equal size
and of similar form ; in other words, in which all the
divisions (whether they are free or adhere to each other
by their edges*) are equally and uniformly developed.
Every flower in which there is unequal or irregular de-
velopment of sepals and of petals, is called irregular.
It is to the very great difference in these respects that
we owe the variety of aspect in the flowers of different
species. As examples, the following familiar plants
may be adduced ; — the pansy has an irregular flower,
that of wall-flower is regular ; a primrose has a regular
flower ; a snapdragon presents an example of irregu-
larity.
The following conclusions appear generally to hold
good as to the relation of form and of colour in the
flower.
1. In regular corollas the colour is uniformly distri-
buted whatever be the number of colours present. — That
is to say, the pieces of the corolla being all alike in size
and form, have each an equal proportion of colour. The
common primrose is an example where there is only one
colour. In the Chinese primrose the same holds where
two colours (the one the complement of the other) are
present, the eye or centre being yellow, and the margin
purple ; these two colours in this regular flower are uni-
formly diffused, that is, each piece has an equal proportion
of yellow and of purple respectively. In Myosotis, Anagal-
iis, Erica, Gentiana, Pyrola, &c, we have uniform corolla
with uniform distinction of colour. All Corolliflorae
Exogens with regular flowers arc examples ; the same is
true of certain Thalamiflora^, as Papaveraceae, Cruciferas,
* It may be necessary to explain that the terms free or adherent, refer to the condi-
tion oftbo mature flower, and not to the mode of development.
148 THE RELATIONS OF FORM AND
&c. ; Calycifloral* Exogens with regular flowers, as Ro-
saceae, Cactaceaa, &c, illustrate the same principle.
2. Irregularity of corolla is associated with irregular
distribution of colour, whether one or more colours are
present. — In irregular flowers where the number five pre-
vails, the odd piece is most varied in form, size, and colour.
When only one colour is present, it is usually more in-
tense in the odd lobe of the corolla. When there are
two colours, one of them is generally confined to the odd
piece. Sometimes when only one colour is present, and
of uniform intensity in all the pieces, the odd segment
has spots or streaks of white. A few familiar instances
may suffice.
_. T , ( Four petals yellow ; fifth, yellow, with
Conmion Laburnum, i ' J ' J
( purple veins.
Trifolium pratense, (com- ) Odd piece distinguished from the others
mon red-clover,) ) by its darker purple veins.
^r ,. .n ( Four petals yellow; fifth, yellow eye and
Kennedia monophylla, < l J ' J J
' purple margin.
c ( Four petals yellow; fifth, white eye on
Swamsonia purpurea < l J ' ' J
( purple ground.
Ajuga reptans, (common ) Four divisions purple ; fifth, has yellow
bugle,) ) spot on inner surface.
Thymus Serpyllum, (wild ) Corolla generally red purple ; two pale
thyme,) ) spots on the odd piece,
Galeopsis Tetrahit, \Fon? divisions generally -yeUow' ^h
( piece has purple spots on yellow ground.
Euphrasia officinalis, (com- ) Corolla purple generally; odd piece has
mon eyebright,) ) yellow spot.
In those well-known annuals, Collinsia and Schizan-
thus, the prevailing colour is purple ; the primary, yel-
low, appears in the odd lobe.
* ThalamifloruB comprehends plants In which there is no adhesion between the whorls
of the flower. Calyciflora; comprehends those in which there is such adhesion. In
Corolliflone the petals are united hy their edges forming a tubular flower, to the inside
of which the stamens partially adhere.
COLOUR IN THE FLOWER. 149
In some genera with irregularity of flower often less
marked than in previous examples, it is worthy of notice
that the two divisions on each side of the odd lohe fre-
quently partake of its characters as regards colour, half
of each resembling the odd piece, as may be seen in
Viola, Gloxinia, Achimenes, Rhododendron, and other
plants.
3. In certain Tlialamiflorous Exogcns ivith unequal
corolla, arising chiefly from difference in size of the
petals, the largest are most highly coloured. — Common
horse-chestnut may be mentioned as an example ; on
each petal there is usually a crimson spot at the lower
part ; the size of this spot and its intensity are in direct
relation to the size of each petal, the two upper being
largest, and the two lateral smaller, and the odd piece
least of all.
4. Different forms of corolla in the same inflorescence
often present differences of colour, but all of the same
form agree also in colour. — The family of plants called
Composite, comprehending Aster, Cineraria, Daisy, &c.,
&c., presents illustrations of this. When there are two
colours, the flowers of the centre, usually of tubular form,
have generally one colour of uniform intensity ; those of
the circumference, having a different form, agree toge-
ther in colour also. Thus the common daisy has all the
tubular flowers of the centre yellow, and all the ligulate
(strap-like) flowers of the ray or circumference are white,
variegated with purple. A yellow centre with a purple
ray is a common association in Compositse ; for instance,
in species of Aster, Rudbcckia, &c. These principles
or laws prevail as well in monocotyledons as in dicoty-
ledons. In the former, the calyx and corolla generally
resemble each other in structure and shape, and in colour
also. This very close resemblance between the two
150 THE RELATIONS OF FORM AND
whorls has given rise to the idea that there is only one
series of external parts in monocotyledons. Kelative po-
sition must, however, not be overlooked, and hence it is
concluded that both calyx and corolla are present. In
dicotyledons we generally find a greater contrast between
calyx and corolla as regards colour. We may say there-
fore,—
5. The law of the contrasts in the colour of the flower
is simpler in monocotyledons than in dicotyledons. —
The flowers of dicotyledons may be symbolized by the
square or pentagon, four and eight, five and ten being
the prevalent numbers in the different whorls ; whereas
since three and six are generally found in the flowers of
monocotyledons, the triangle may serve to symbolize such
arrangement. Such comparison is not fanciful on our
part, but an actual statement of the mode of illustration
adopted by botanists. Thus, in a work by one of the
highest authorities of the day,* a series of triangles is
used for the purpose of demonstrating, more clearly than
could be done by any other means, the true relations of
the flower in the families of the grasses, j>alms, and or-
chids.
We may state in conclusion, therefore, that simplicity
of figure corresponds ivith simpler contrast of colour in
the monocotyledons, tvhile greater complexity of colour
and greater complexity of structure are in direct relation
in dicotyledons. — In all these remarkable co-existences
there is surely something more than mere casual coinci-
dences. As the laws of the beautiful have not been
detected and unfolded, it is not possible to demonstrate
scientifically that the relations we have been treating of
are in accordance with aesthetic principles. But the eye
at once perceives in regard to some of these arrange-
* Lindley's Vegetable Kingdom, pp. 109, 169, 178.
COLOUR IN THE FLOWER. 151
merits, that they tend to enhance the beauty of the planl;
Would not reason he offended if uniform flowers had not
uniform colouring ? Is there not a propriety, when in
an irregular flower there is one petal standing by itself,
that that petal should have more brilliant colours, that
thus the flower may be tempered together, having more
abundant honour in the parts which lacked, that there be
no schism in the plant ? We are persuaded that were we
to put a flower without any colour into the hands of a
skilful colourist, and ask him to put on the colours, he
would do so on the very principles according to which
plants are coloured in nature.
Proceeding on the principle that since plants of all
epochs of the earth's history have been constructed on the
same general plan, so the same associations of colour,
and of colour and form, must have prevailed also, we may
finally glance at a few conclusions to be derived from this
source.
During the earlier geological periods, when Acrogenous
Cryptogamia (Ferns, &c.) were abundant, the secondary
and tertiary colours, as green, purple, russet, and citrine,
probably prevailed.
During the reign of Gymnosperms, when Cycadece and
Conifers were numerous, the secondary and tertiary colours
must still have given a sombre aspect to the vegetable
world.
From the commencement of the chalk formation there
appears to have been a very marked and progressive in-
crease of Angiospermous dicotyledons, which form the
largest proportion of existing vegetation. Among them
we find the floral organs with greater prominence in size,
form, and colour ; and such prominence of the " nuptial
dress" of the plant is peculiarly a feature of species be-
longing: to natural families which have attained their
152 ADAPTATION OF THE COLOURS OF
maximum in man's epoch, and are characteristic of it.
Brougniart,* one of our highest authorities in this de-
partment, states that a remarkable character of the floras
of the eocene, miocene, and pliocene epochs — which im-
mediately preceded man's epoch — is the absence of the
most numerous and most characteristic families of the
Gamopetake.f Nothing announces the existence of Com-
positce, Personatfe, Labiatae, Solanaceae, Boraginaceae, &c.
Doubtless there were lovely flowers in former periods,
" born to blush unseen," at least by human beings, but
we miss those which are our special favourites, and whose
cultivation is one of the characteristics of civilized man.
We cannot avoid thinking that there was design in al]
this, that the succession of created forms in the vegetable
kingdom had a reference to the epoch of man ; and that
just about the time when there appeared an eye to re-
ceive and convey the impressions of beauty, and an intel-
lect to derive satisfaction from the contemplation of such,
then it was that the most highly adorned productions of
Flora's kingdom were called into existence.
SECT. II. ADAPTATION OF THE COLOURS OF PLANTS TO THE
NATURAL TASTES OF MAN.
Artists lay it down as a maxim that a large portion of
a painting should be of a neutral colour. Our natural
tastes would not tolerate a scarlet or purple ground to a
historical painting. In a skilful piece of art the more
prominent figures are made to rise out of colours which
attract no notice. It is the same in the beautiful canvas
which is spread out before us in earth and sky. The
ground colours of nature, if not all neutral, are at least
* Annates des Sciences Naturelles, 1S49.
t In Game-petal* there ia adhesion of petals ; the flowers are tubular.
PLANTS TO THE NATURAL TASTES OF MAN. 153
all soft and retiring. How grateful should we be that
the sky is not usually dressed in red— that the clouds
are not painted crimson — that the carpet of grass on
which we tread is not yellow, and that the trees are not
decked with orange leaves ! The soil, in most places,
is a sort of brown — the mature trunks of trees commonly
take some kind of neutral hue — the true colour of the
sky is a soft blue, except when coloured with gray clouds,
and the foliage of vegetation is a refreshing green. It
is out from the midst of these that the more regular and
elegant forms, and the gayer colours of nature come
forth to arrest the attention, to excite and dazzle us, not
only by their own splendour, but by comparison and con-
trast.
All the gayer colours of the vegetable kingdom seem
to be beautiful in themselves. The eye needs no asso-
ciated object to lead it to detect a loveliness in the red
rose, and the blue harebell, and the yellow primrose.
But there are associations of colour in art which have a
pleasing effect upon the mind. In our Schools of Design
pains are taken to shew what colours may be placed in
juxtaposition, and what colours may be kept at a dis-
tance from each other. In the construction of tapestry,
and other kinds of higher needlework, in the manufac-
ture of our finer texture of fabrics, and in the staining
of glass for windows, strict attention is now paid to rules
on this subject, prescribed by science and sanctioned by
experience. We proceed to shew that in nature colours
have been associated from the beginning, according to
principles which have become known to man only at a
comparatively late date in the history of human civili-
zation and science. In order to explain this, it will be
needful to begin with a few elementary statements in re-
gard to light and colour.
154 ADAPTATION OF THE COLOURS OF
According to the commonly adopted doctrine, there are
three Primary Colours, Eed, Yellow, and Blue. The
combination of these, in certain proportions, yields White.
The absence of them all is Black. These primaries, mixed
together, two and two, produce what are called Secondary
Colours, viz., Orange from the mixture of red and yellow,
Green from the mixture of yellow and blue, and Purple
from the mixture of red and blue. From the combination
of the secondaries arise three Tertiary Colours : — Citrine
from the mixture of orange and green, Olive from the
mixture of green and purple, and Russet from the mix-
ture of orange and purple.
There are certain other phrases which it maybe needful
to explain in their technical sense, as used by colourists.
Tint is employed to denote the gradations of colour in
lightness and shade ; Shade to express the gradations in
depth from white down to black ; Hue is applied to the
mixtures in compound colours. Thus we talk of a light
tint of red where the red approaches to white, of a dark
shade of purple where the purple inclines to black, and
of hues of orange from the yellowest to the reddest, of
hues of green from the yellowest to the bluest, and of
hues of purple from the bluest to the reddest. When
the orange has more than its proper proportion of red,
we call it a red orange hue ; when in green the yellow
prevails, we call it a yellow green ; and when in purple
the blue predominates, we call it a blue purple. This is
the common doctrine taught in schools of art ; it is correct
enough for the purpose which we have at present in view,
and the nomenclature enables us to express, in a rough
way, the infinitely varied colours in nature.*
* Newton thought that there were seven simple primitive colours, red, orange, yellow
green, blue, indigo, violet. Sir David Brewster has shewn that these can be reduced to
three. Some scientific men seem to reckon all such classifications as in some respects
PLANTS TO THE NATURAL TASTES OF MAN. 155
The language of music has been applied to colours ;
and colourists talk of the Melody of colours and the
Harmony of colours. Colours are said to be in Melody
when two contiguous tints, or shades, or hues, run insen-
sibly into each other, as when red slides into pink and
white, and purple deepens into dark purple or merges
into red purple and red. Two different colours are said
to be in Harmony when their association is felt to be
pleasant to the eye.
Two colours are said to be Complementary when they
together make up the white beam. Thus green and red
are complementary, as also purple and yellow, orange
and blue. The eye feels a pleasure in seeing colours
in melody, or melting into each other. It also feels a
pleasure in contemplating certain associations of different
colours. In particular, the eye is pleased when comple-
mentary colours are beside each other, or under the view
at the same time.* Complementary colours contrast the
one with the other, but are always in harmony. It is
necessary to add that white associates pleasantly with
every other colour, as does also black.
The accompanying diagram {Fig. 32) is constructed
with the view of shewing what colours are complementary
to each other. In this figure we have the three primary
colours, red, yellow, and blue, and the three secondaries,
orange, green, and purple, with the hues of the seconda-
ries on either side. We have also the tertiaries, citrine,
arbitrary, and speak of the solar light as composed of indeterminate numbers of differently
colored rays. We have no opinion to offer on these points, or any other disputed point,
in regard to the nature of colour. But as it is needful to use nomenclature of some
description, we adopt the commonly received doctrine as expressing the actual facts
very clearly, and with sufficient correctness for the purpose which we have in view.
* Divers explanations, physical and physiological, have been given of this. None of
these seems to us to be altogether satisfactory, and it would be beyond our province to
discuss them in such a treatise as this. It is enough for us that the fact be admitted, that
the eye is gratified when it can simultaneously unite two complementary colours.
156
ADAPTATION OF THE COLOURS OF
olive, and russet. The diagram is so constructed that
the colours in corresponding segments of opposite circles
are complementary, and so in harmony. Thus, red and
Fig. 32.
green, blue and orange, yellow and purple, are com-
plementary. According to the hue of any particular
secondary, so is also the hue of its complement. Thus a
pure purple requires a yellow, but a red purple requires
a yellow green, and a blue purple a yellow orange, as
the complementary colour ; and so of all the other
secondaries. The tertiary citrine is in harmony with a
dark purple, olive with a dark orange, and russet with a
dark green.
These principles are taught now in every school of art,
and are attended to in the manufacture of all our finer
PLANTS TO THE NATURAL TASTES OF MAN. 157
fabrics in which colour is an element of beauty, as in
dresses, carpets, hangings, and furnishings of various
descriptions. When two colours not in harmony might
come in contact, the discord is avoided by placing a line
of white or black between them. We are now to shew
that these principles are also attended to in the colouring
of certain departments of nature. Thus, to take up the
three secondaries, green, purple, and orange.
1. Green harmonizing ivith red and russet. — The soft
hue which the Author of nature has been pleased to
give to the leaf of tree and herbage, is by far the most
abundant colour in the vegetable kingdom. Now, where-
ever the flower of a plant is red, it associates agreeably
with the leaf. The flowers of the rose, and of many
pinks, geraniums, pelargoniums, mallows, lychnises, and
dozens of others, contrast strikingly with the foliage of
the plants on which they grow. The eye delights to see
the fruit of the cherry, the rose, and the thorn, and the
berry of the holly, the yew, the common barberry, the
mountain ash, and unnumbered other plants, peeping
forth from the green leaves. It often happens that ac-
cording to the hue of the green so is the hue of the asso-
ciated red. In a vast number of plants, the young stems
and the petioles of the leaves, and in not a few cases the
veins of the leaves, are red purple, contrasting with the
leaves which are yellow green. The young cones of the
larch, in spring, are of a reddish purple, harmonizing
with the yellow green foliage. In other cases we find that
it is a russet, that is, in harmony with a dark green. In
the fir tribe and its allies, the leaves are dark green, and
steins are russet. The same colours are the prevalent
ones among rushes, and, indeed, in most of the juncous
family of plants.
2. Purple harmonizing icith yellow and citron. — This
158 ADAPTATION OF THE COLOUKS OF
is the second most prevalent harmony in the vegetable
kingdom. So far as we have been able to observe,
purple of various tints, shades, and hues, such as red
purple where there is a preponderance of red, and blue
purple where there is a preponderance of blue, is the
most frequent colour of the petals of plants. In beau-
tiful contrast we often find yellow in the centre of the
flower. Thus, in the garden polyanthus, and in many
varieties of auricula, the outer rim of the corolla is
purple, and an inner circle is yellow. More frequently
the complement is found in the yellow anthers or yellow
pollen. It is a remarkable circumstance, that as the
most frequent colour of petals is purple, so the most
common colour of the pollen of plants is yellow. It is
curious to notice, that according to the hue of the purple
so is the hue of the associated yellow. Thus, in potato
and bittersweet (Solanum dulcamara), the corolla is blue
purple, and the anthers are red yellow, whereas in poly-
anthus the outer edge of the flower-cup is red purple,
and the heart greenish yellow. In other plants the
complementary is. not yellow but citrine, a colour not
uncommon in matured and decaying vegetation, where
it contrasts with a dark purple. Purple and citrine are
also commonly associated in the flowers of grasses. The
newly-ripened cone of the cluster-pine is citrine ; when
the scales open, the complementary purple is revealed on
the base of each.
3. Orange harmonizing with bhie and olive. — This
harmony is less frequently met with in the vegetable
kingdom, (it is very common in the sky.) Still, there
are examples to be found. Thus in several species of
Strelitzia, (as S. Regina?, S. juncea, &c.,) the sepals
are orange and the petals blue. A pure blue, however,
is rarely to be met with in the flower of any of the
PLANTS TO THE NATURAL TASTES OF MAN. 159
organs of plants. Most of the flowers called blue have
more or less of a tinge of red. In such flowers the har-
mony is often very evident. Thus, the reddish-blue
petals of blue lupines and Jacob's-ladders are associated
with reddish-yellow anthers. In not a few composite
plants, in some Hieraciums, for example, (such as Hiera-
cium aurantiacum,) we may observe an orange disc sur-
rounded by an olive involucre. The olive in some of these
plants seems to be produced by purple spots on a green
ground.
Not unfrequently the complementary colours may be
found on the same organ. Thus, the side of a young
branch exposed to the sun is often reddish purple, and
the other side yellow green. But it is in the flower that
we most frequently meet with the sister colours. They
may be seen in many of the popular favourites, both
among wild and garden plants. In the flower of the
11 forget-me-not," which ever greets the eye so cheerfully,
there is a border of blue purple, and a centre or throat
of orange yellow. In the pansy, so rich and soft that it
has got the name of " heart\s-ease," we have yellow and
purple of various hues and degrees of intensity, bright-
ened by a mixture of white. Eyebright has a purple and
white corolla, with a sprinkling of yellow on its odd lobe
In many of the universal favourites, harmony of colours
adds at least to the effect produced by beauty of form.
It is probably the elegant shape and the hanging posture
of the flowers of foxglove which allure children to it, but
the interest which they feel in it may be unconsciously
increased by its purple and white petals, and its yellow
anthers adorned with purple spots. The yellow Iris
(I. pseudacorus) has a yellow flower lined with purple,
and it has purple dots on the yellow anthers. In the
daisy, described as " crimson-tipped" by Burns, there is,
160 ADAPTATION OF THE COLOURS OF
ft
the yellow disc harmonizing both with the white ray and
the purple on its tips. These flowers are favourites with
all classes — peer and peasant, old man and young maiden,
countryman and townsman. They pleased us in our
childhood, when we seized them and sought possession
of them so eagerly, but found them fading like all earthly
enjoyments, and they please us still in our advancing
years, as we prefer lazily looking at them, and allowing
them to grow where God has planted them, that they
may gratify us and others as we pass on in the journey
of life. That which has thus endeared them to multi-
tudes is, we believe, to some extent at least, this very
harmony of colours, which all feel, because it is intended
— it is natural, that we should feel it, but which could
not, till within these few years, have been scientifically
expressed. We may also notice that yellow and purple
are found in close contiguity on the flowers of many of
the plants which man has domesticated, and which find
a place in every garden, such as Chinese primrose, auri-
cula, polyanthus, mimulus, calceolaria, Indian cress, snap-
dragon, and marigolds.
But we are not to suppose that the two colours are
always to be found on the same organ, or that this har-
mony is confined to the inflorescence. On the contrary,
it appears in a vast number of situations, and we have
often found pleasure in detecting it under its various
modifications. Frequently one of these colours is on
one organ, and its complement on another organ. Very
commonly (as we have seen) we have purple petals with
yellow anthers or pollen, but at times there is a different
order and relation of colours between these two organs.
Thus, in several species of poppies (e. g. Papaver orien-
tale) the petals are red orange, and the anthers olive.
Usually the anthers, or at least the pollen, of plants is
PLANTS TO THE NATUKAL TASTES OF MAN. 161
yellow ; but in the turn-cap lily, the decidedly red pollen
is associated with the green filaments of the anthers, and
in Hypericum Androsaemum, we meet with purple an-
thers, contrasting with the yellow filaments and yellow
petals. In Amygdalis communis, the yellow anthers
have their complements in the purple filaments. In
wood-sage, the purple filaments contrast with the yellow
petals.* In some syngenesius plants, there is one colour
in the ray, and its complement in the disc ; thus, in
Guillardia pinnatifida and Coreopsis Drummondii, the
ray is yellow and the disc purple. Sometimes the one
colour is in the calyx and its sister colour in the corolla.
Thus, in evening primrose, (Oenothera macrocarpa, and
also in 0. tenuifolia,) the petals are yellow and the sepals
purple. In some species of Kanunculus, (R. repens,
E. bulbosa, R. Flammula,) the yellow flowers have their
complement in purple on calyx, leaf-stalks, or leaf-sheaths
on one or other, at times on all. In certain species of
rushes, (e. g., Juncus cornpressus,) the anthers and pollen
are yellow, the ovary and stigma are purple, and the edge-
of the perianth is russet, and the centre dark green.
In the paper reed of Egypt, we may observe that the
sheaths at the base of the stalks are red purple, while the
stalks themselves are yellow green. In some plants the
stems and leaves have one of the hues of green, and the
spines and prickles the corresponding hue of rod. At
times the leaf or stalk is one colour, and upon it there
are spots of the complementary colour ; thus, on hemlock
we may notice red purple spots on the yellow green
* It may be proper to allude here to Count Euniford's principle, that two colours, to
be in harmony, must both present the respective proportions of the coloured light ne-
cessary to form white. In most of the instances we have adduced, it would not be easy
to prove a conformity to this principle. But Chevreul, one of the highest authorities on
this subject, considers Rumford's statement "as nothing more than an ingenious inven-
tion of fancy." (See Taper in Chern. Rep. of Cavendish Society, p. 1S9:)
162 ADAPTATION OF THE COLOURS OF
stalks. Nay, we have observed that, if there be but a
diseased spot or wart on a leaf produced by an insect, the
colour of the spot will at times be complementary to that
of the leaf, as may be seen in the little galls on the leaves
of willows and roses. The scales of young cones are often
purple, whereas the scales of the old cones, hanging on
the same tree, are citrine. In Victoria regia, we may
notice on the leaf (besides the beautiful mechanism by
which it is supported) red jjurple ribs harmonizing with
the prevailing yellow green, and in the expanding flower
the red purple calyx harmonizing with the yellow green
at the edge of the sepals.*
* This frequent juxtaposition of complementary colours must have a physical as well
as a final cause. If it be asked what this is, we are inclined to answer this question by
asking another, the answer to which may p^sibly open up the way to an answer to the
first question. When a beam of light falls on a green leaf, the green is said to be reflected
and the red absorbed; but what, we ask, becomes of the red? When the beam falls on
a purple petal, the purple is said to be reflected and the yellow absorbed ; but what be-
comes of the yellow? Are the red and the yellow in these cases absolutely lost? If these
constituents of the beam be lost, they are the only powers' in nature which are so. In
this world of ours nothing which has existed at any time is lost, even as nothing abso-
lutely new comes into being. It is now a received doctrine, that the heat absorbed by
plants in the geological era of the coal measures is laid up in fossil deposits, and may
come forth in our epoch when the coal is ignited. May we not suppoes, in like manner,
that the red absorbed by the plant when the green is reflected by its leaves, will come
forth sooner or later, in some form, in young stem, flower, or fruit; and that the yellow
absorbed by the flower when the purple is reflected, will come out in the yellow pollen,
or in some other form? AVe have thought at times that as the pure white beam, when
it reaches the earth with its atmosphere, is divided into several rays, and that no one of
these is lost, and as they all come forth sooner or later, we have thus a harmony of colours
In nature. We h"ve thus the brown earth, the ultimate recipient of the ra\ s which have
passed through the atmosphere, harmonizing with the blue sky, and ligneous substances
become orange when ignited. But we throw out this view as a mere hypothesis in the
absence of a better, and in order, if not to guide, at least to stir up inquiry; and we beg
that it may be carefully separated from the co-ordinated facts presented in the text. In
whatever way we may account for it, there is a most singular succession as well as co-
existence of colours in the vegeiable kingdom. Harmonious colours come out not only
contemporaneously, but consecutively. In several species of Geum, (as G. urbanum and
G. intermedium,) the petals are yellow and the pistils purple, but it Is not till the yellow
petals are falling off that the purple pistils appear. We have the same curious pheno-
menon in some species of Fragraria. In Cytisus Canadensis, the yellow corolla is fol-
lowed by the purple pod. In some Cactacea?, the yellow flower Is succeeded by a purple
fruit. In Taxodium sempervirens, the young shoots are yellow green, those of a year
old are red purple, and those older still, citrine. Generally branches, when young, are
green, as they advance they are purple, at a farther stage they are citrine, and finally
PLANTS TO THE NATURAL TASTES OF MAN. 163
These harmonies are found in plants belonging to all
the principal divisions of the vegetable kingdom. Thus,
among the family of Mosses, the red or red purple teeth
of the peristome are associated with the green or yellow
green capsule ; and the same is true of the different parts
of their stems and leaves. Among Fungi, we have Bo-
letus luridus and Boletus luteus with yellow and purple
stems. In Lycopodiums, the most common colours are
yellow and purple. Among Ferns, we have noticed
Doodia aspera with its young fronds red purple and
yellow green, and Dicksonia adiantoides with yellow
green fronds and red purple stalks. Most exotic Or-
chidese have yellow with purple spots, or yellowish green
with red purple spots on calyx and corolla. In the flower
of grasses, the prevailing colours are purple and citrine,
russet and dark green. We have already detected this
harmony among rushes, among herbaceous plants, among
the cone-bearers, and trees generally.
It is a most interesting occupation to trace it at every
season of the revolving year. In spring it is very obvi-
ous in the contrast between the yellow green leaf and the
red purple of the stalk on which it grows ; thus the
young leaves of the primrose are yellow green, while the
stalks are red purple. At the same season we may no-
tice that the flower of Tussilago is yellow, while the
involucre and scales of the stalks are purple. In the
summer season the powerful beams of the sun bring
forth this harmony in plants of every description. In
russet. Surely these successions are instructive. We have felt a deep interest in no-
ticing how, in a vast number of plants, the colours which make up the full beam do
some time or other, separately or in combination, make their appearance during the
life or at the death of the plant. There are also curious cases, in which one colour
appears in the outside, and its complement in the inside of the fruit. The inside of a
nearly ripe tig is red-purple, the outside yellow-green : the same is true of the pericarp
In some species of Pseony. The skin of the berry of Mahonias is blue, whereas the
Interior is orange.
164 ADAPTATION OF THE COLOURS OF
autumn it is very strikingly exhibited in the contrast
between the leafage and the berry, and other fruits. Nay,
it is often very visible in the fruit itself. Thus in cer-
tain varieties of apple, hues of red and purple are asso-
ciated with hues of green and yellow green, while in
some varieties of pear, yellow green, red purple, and
citrine occur together. The year dies (like the day) in
glory amidst a magnificence of colouring in its phase,
in which prevailing hues are greenish yellow and deep
red purple, and citrine relieved by dark purple spots.
In winter itself, we may see the harmony in those plants
which (like friends in adversity) choose that season to shew
their beauty ; thus the greenish yellow corolla of the ar-
butus harmonizes very beautifully with the red purple of
the anthers, and also of the flower-stalks. The eye is
refreshed in the depth of winter by seeing the red berries
peeping forth from the midst of the green foliage of the
yew and holly. Thus does the harmony run on till the
returning sun of spring calls forth a new cycle.
We may discover in it, if we patiently seek for it, in
every description of natural scene. In the grass of the
fields we may observe it in the stems, which are often
red purple in harmony with the yellow green leafage, and
in the purple and citrine of the flowers. Nor can any
one walk far in the fields without meeting plants which
he has only to examine to discover that they illustrate this
conjunction. If the bird's-foot (Lotus corniculatus)
catch his eye, he may notice that its lively yellow co-
rolla is relieved by purple on the outside of its large lobe.
Or if he pick up the flower of purple clover, he will find
that the anthers are yellow. If he carefully examine the
common buttercups, he will find that as a set-off to the
yellow flower there is purple on the calyx or some othei
organ. The yellow flower of silverweed (Potentilla anse-
PLANTS TO THE NATUKAL TASTES OF MAN. 1Q5
rina) has a visible contrast in the purple stalks and run-
ners. He may notice how the yellow flower of common
hawksLit (Hieracium Pilosella) has purplish tips and
purple on the outside, and how numberless yellow syn-
genesious plants, such as dandelion and Apargia autum-
nalis, grow on purple stalks, and have purple spots on
the involucre. Here and there he will discover Sym-
phytum tuberosum, with dull yellow corolla and dull
purple stem ; or self-heal, (Prunella vulgaris,) presenting
its calyx with russet border and dark green centre, sur-
mounted by blue purple corolla and whitish anthers. Pos-
sibly he may be so fortunate as to fall in with a rock rose,
(Heliantliemum vulgare,) with its yellow petals melodiz-
ing into crimson, and striped with purple. In our drier
meads he cannot but notice yellow rattle, (Khinanthus
Crista-galli,) with yellow corolla tipped with purple, and
Lathyrus pratensis, with purple veins in the large lobe of
its yellow corolla ; and in our watery marshes the lousewort,
(Pedicularis palustris,) with its purple petals and yellow
anthers. In our pools he may meet with the Comarum
palustre, with its dark red purple corolla and its yellow
green heart. If he wander by our rivulets he may fall
in with Geum rivale, with its purple petals, and its abun-
dant and prominent yellow anthers, with its russet calyx,
harmonizing with its dark green leaf. If he go forth
into our wastes, he will meet with our sedges and rushes
with their purple and citrine. In shady and moist
places he may see the common loose-strife, (Lysimachia
nemorum,) with yellow corolla, and stems and leaves
tinged with purple. In our hedges he has the yellow
green leaf of the thorn harmonizing with its red purple
shoots, and growing up in the midst of them the purple
vetch, (Vicia sepium,) with its purple corolla and yellow
anthers ; while in the ditch there may be the lovely
166 ADAPTATION OF THE COLOUKS OF
"forget-me-not," with its reddish blue and yellow
orange. If he enter the wood he may see the com-
mon anemone, with its purple flowers and yellow anthers,
or the leafage of the bush contrasted with its berries,
or the cones of the fir and pine contrasted with one
another, or with the foliage. If he betake himself
to the sea-side, he will fall in with the sea sandwort,
(Arenaria marina,) or the common sea-pink, (Statico
Armeria,) both with purple corolla and prominent yellow
anthers ; or the common sea-radish, (Raphanus mariti-
raus,) whose open yellow corolla harmonizes with the
unexpanded flower-buds, which are purple.*
We are inclined to think, farther, that there is often a
beautiful harmony in the way in which different plants
are associated in nature. It is a curious circumstance
that the colours of some sea-weeds are red of various
hues, and of others are green of various hues, and as these
grow together they help to embellish one another. We
have heard skilful colourists declare that there is a har-
mony in the colours of the plants growing together in our
finest meads, and our own eye testifies to the same effect.
We are quite aware that in our cultivated fields there are
often plants growing together with colours that are discor-
dant. We could never discover any beauty in the yellow
mustard growing among the green stalks of the farmer's
grain. But in nature's own meads, in all places in which
she not only grows but is allowed to sow her own plants,
she commonly distributes her colours very gracefully.
We are not prepared to give the full rationale of this.
So far as the herbage is concerned, it may be partly ac-
counted for by the circumstance that yellow and purple
are the most common associations in the flower of grasses,
* We would i-cfer to the Appendix for additional examples of harmonious colours in
different plants.
PLANTS TO THE NATURAL TASTES OF MAN. 167
and red purpie and yellow green in the stalks and leaf-
age. The green foliage, too, is everywhere relieved by
red fruit and red flowers, such as wild roses, ragged
robins, red campions, and geraniums. In the summer and
early part of autumn, there will be buttercups still lin-
gering, and bird's-foot, and clivers syngenesious plants,
such as ragweed and hawksbit, all yellow or yellow inclin-
ing to orange, and in contrast there will be purple clover
and scabiouses, and self-heal, and harebell, and common
bugle, and thistle, and knapweed, all purple or purple
inclining to blue. We may notice, indeed, that in many
of our fields some of these colours prevail to an unpleasant
extent above the others. Thus in some spots there may
be a disagreeable glare of yellow caused by ragwort and
buttercup ; but we have noticed that if the progress of
agricultural improvement does not interfere with the
natural process, the thistles and knapweeds will soon so
spread themselves as to restore the proper balance of
colour. Nor let it be forgotten that nature lightens the
whole scene, and heightens the effect of every other
colour by her white flowers, by her daisies, her stitch-
worts, her chickweeds, her great white ox-eyes, her mil-
foils, and her meadowsweets. One-reason why man loves
and longs in these times to retreat from our best culti-
vated regions to the wilds of nature, is to be found in
the circumstance that nature, in her own domains, min-
gles so gracefully her forms and colours.
We have thus a frequent harmony in the colouring of
the individual plant, and a not unfrequc-nt. harmony in
the colouring of plants growing contiguous to each other.
When the plant is near, the eye will naturally fix itself
on the complementary colours of the individual plant, and
when we are looking at a lawn at some little distance,
the eye will rather select the harmony presented by dif-
168 ADAPTATION OF THE COLOURS OF
ferent plants. And here it is worthy of being mentioned,
that colourists acknowledge that if there be complemen-
tary colours among objects bofore the eye, it will instinc-
tively fix on them, to the neglect of adjacent colours.
Chevreul, who is the highest authority on the subject
of simultaneous contrast of colours, recommends that in
planting out flowers in gardens, attention be paid to the
rules of complementary colours. " The principal rule to
be observed in the arrangement of flowers, is to place the
blue next the orange, and the violet next the yellow,
while red and pink flowers are never seen to greater ad-
vantage than when surrounded by verdure and by white
flowers ; the latter may also be advantageously dispersed
among groups formed of blue and orange, and of violet
and yellow flowers." * But this eminent chemist does not
seem to have observed that plants in nature are arranged
on these very principles. A skilful colourist, conducted
into a garden, planted out on the plan recommended by
Chevreul, would at once discover that there were plan
and purpose in the distribution of the plants. But there
are no less convincing proofs of design in the way in
which colours are arranged on individual plants, and in
which plants are distributed over our meadows and
mountains.
Though it does not fall within our immediate subject,
we may here be allowed, as an illustration of the general
subject, to remark that traces of harmony of colours
may likewise be found in the plumage of birds. The
following seem to be the most common forms in which
it presents itself. First, We often observe some dark
colour, at times a black, but more commonly a dark blue,
* See Paper by Chevreul, p. 2fiS, in Chemical Reports and Memoirs, 1S4S, of Works of
Cavendish Society. The same views are more fully developed in Chevreul's great work,
"De la loi du Contrasto SimultauB des Couleurs, (1S39.)''
PLANTS TO THE NATURAL TASTES OF MAN". 169
or very blue purple, in harmony with white. Sometimes
the white is on the belly or breast, while the dark hue
is on the back ; at other times there are white spots
relieving the dark shade all over the body. This is a
common association in our birds of plainer plumage. It
may be seen in many web-footed fowls, such as geese,
divers, and gulls. The second most common harmony —
if, indeed, it be the second and not the first — is between
a sort of tawny hue, being a yellow, with more or less of
red, and a dark blue, or rather dark blue purple. This
collocation of colours is very frequent among raptorial
birds, as, for example, many falcons and owls, and is
found among wading birds and many species of thrushes.
TJiirdly, in our more ornamented birds we discover red
associated with green. This congruity appears, and at
once arrests the eye, in a great many parrots, in a num-
ber of todies, and in the Curucuis, a tribe of birds which
live in low damp woods in the tropical parts of America
and Asia, and feed on insects and berries.
These seem to be the more marked associations, but
these three forms run into each other. Thus, some horn-
bills are dark blue and reddish yellow, but others have
white instead of yellow. This is also the case with some
of the raptorial birds. In the plumage of some fowls
the reddish yellow seems to be a pure orange ; this
seems to be the case with some toucans — other toucans
seem more nearly green and red. The same may be said
of many solitary warblers, fly-catchers, and starlings.
In some birds the red yellow is brightened into a scarlet,
harmonizing with a greenish blue ; this is a very com-
mon association among chatterers and finches. The
scarlet ibis has the greater part of its plumage of the
hue which its name denotes, but has a greenish blue on
its wings. Among pheasants we often discover a red
8
170 ADAPTATION OF THE COLOURS OF
orange and a blue green, and the same colours, differently
distributed, appear on our more ornamented ducks. In
reviewing these associations we may notice that we have,
on the one side, white rising into yellow orange and red,
and on the other side blue sliding into purple or green.
We have not paid special attention to the subject, but
similar harmonies prevail, we doubt not, in other depart-
ments of nature, as, for example, among insects. Any
one may notice the yellow and purple on bees and wasps.
The most cursory glance is sufficient to shew that many
shells of mollusca are characterized by a yellow ground
adorned with purple spots. In another department of
nature it has been remarked by Field that the brown
earth harmonizes agreeably with the blue sky.
Surrounded as we are by such harmonies, we are
convinced that whenever the mind seeks for them it will
discover them ; nay, the eye fixes on them when it is not
designedly seeking for them, and rejoices in them when
it can give no account of the cause of its joy. At the
same time, the contemplative intellect experiences a far-
ther pleasure, and a pleasure of its own, when it can
scientifically explain to itself the source of all this enjoy-
ment, and systematically look out for the pleasing asso-
ciations of nature.
The heart, rightly tuned to the praise of its Maker,
will experience a farther pleasure. Present to a skilful
colourist an article of human workmanship, constructed
according to the rules of simultaneous contrast in colour-
ing, and he will at once say, Here are art and design.
Place before him a piece of Gobelin tapestry, one of our
finer carpets, or the stained glass of a window, and he
will perceive at a glance that the associations of colour
are not accidental, but that they are purposely suited to
the physiological and psychical nature in man. We are
PLANTS TO THE NATURAL TASTES OF MAN. 171
convinced that there are equally clear proofs of con-
trivance in the colouring of natural objects, organic and
inorganic. Indeed, colourists, long ago, observed that
there was a beautiful harmony in the colours of nature ;
and within the last age, Field and Hay, and very pos-
sibly others, have stated what is the nature of this har-
mony, though they have not followed it into the various
departments of natural history. He who can trace up
all these adaptations to Him who causes His works to
make sweet music by their harmony, has surely here
a source of higher — we should rather say, of highest
joy-
But the question is here started, Are there no colours
associated in nature except harmonious ones ? This is
a question which we are not prepared dogmatically to
answer, either in the negative or positive. One thing,
however, seems to us very certain, that complementary
colours appear so often in nature, and cast up, under
such different modifications, and in such a variety of
objects and situations, that their conjunction cannot be
the result of mere chance. Besides the generalized facts
of a positive character, we are prepared to say negatively
that we have never observed in- a corolla, or in any one
organ of a plant, pure red and pure yellow, or blue and
red, in contact with each other.* But in making these
affirmations we are, at the same time, prepared to admit
that there are colours in nature in juxtaposition which
are not complementary. This, however, just raises the
question, Can no colours be pleasantly associated except
c< tmplementary colours ? This question must be answered
in the negative, and being so answered, a host of inquiries
* The same statement was made to us by Mr. Wood, an experienced flower painter
And lately assistant master in the Belfast School of Design. He fartuer informed us
that he invariably found associations of harmonious colours in the different parts of
plants, such as we have been describing.
172 ADAPTATION OF THE COLOURS OF
come to be made as to what other associations are
agreeable, and these should be followed by a series of
investigations, having it for their end to discover how far
all the non-complementary associations of nature can be
described as pleasant. Chevreul tells us that we cannot
prescribe arrangements of non-complementary colours, so
as to please the eye, in as positive a manner as may be
done with reference to the assortment of complementary
colours. " This is the reason," he adds, " that in treat-
ing of the distribution of flowers in gardens I have only
recommended an assortment of flowers whose colours are
complementary, at the same time that I admit the exist-
ence of many other assortments productive of a very
agreeable affect."0 This whole subject is just opening
upon us, and we must be satisfied for the present to sub-
stantiate a certain amount of truth, to acknowledge that
there are unsolved points and difficulties, and trust that
these may be cleared up by further investigation.
AVe must here state, however, that many of the seem-
ing exceptions co these general views, are exceptions
merely in appearance.
It not unfrequently happens, in the vegetable kingdom,
that the discord between two contiguous colours is sub-
dued by a patch of white, which, like innocence, (of
which it has always been reckoned an emblem,) has never
occasion to be ashamed of itself, for it may appear any-
where, and is in harmony with every object it can meet
with. In Lycopsis arvensis, in harebell, and speedwells,
the blue of the petals has no complementary orange, but
then it is beautifully relieved by an adjacent white.
It may seem as if the leaves of plants were liable to be
seen simultaneously with every other colour in the vege-
* See in works of Cavendish Society, Chemical Reports and Memoir?, 1848. — Paper by
Chevreul, p. 219.
PLANTS TO THE NATURAL TASTES OF MAN. 173
table kingdom, that there must be discord when the green
leaf is perceived at the same time with the yellow and
blue of the flower. Chevreul, in speaking of the artifi-
cial arrangement of flowers in a garden, lays down a rule
which enables us to escape the difficulty. " I must, how-
ever, reply to the objection that might be made, that the
green of the leaves, which serves, as it werer for a ground
for the flowers, destroys the effect of* the contrast of the
latter. Such, however, is not the case ; and to prove
this, it is only necessary to fix on a screen of green silk
two kinds of flowers, (in the manner pointed out in the
paper,) and to look at them at a distance of ten paces.
This admits of a very simple explanation, for as soon as
the eye distinctly and simultaneously sees two colours,
the attention is so riveted that contiguous objects, espe-
cially when on a receding plane, and where they are of
a sombre colour, and present themselves in a confused
manner to the sight, produce but a very feeble impres-
sion."*
Nor is it to be forgotten, that the coloured flowers of
many plants are raised out from the midst of their leaves,
and are so far above them that the petal and leaf do not
come simultaneously into view in a marked manner,
This is the case very obviously with harebell, dandelion,
hyacinths, and many other plants. In such cases, it
may be found either that the flower has a beauty of its
own independent of any adjunct, or that it has a harmo-
nizing concomitant in some other plant usually growing
in the neighbourhood.
More important than any of these, we find that there
is a physiological provision in the eye itself, which helps
it to overcome any slight defects in the balancings of the
colours in nature. Chevreul lays down the law, that in
* Chevreul's Taper on Chemical Reports, p. 207.
174 ADAPTATION OF THE COLOUKS OF PLANTS, ETC.
the case of the eye seeing at any time two colours which
are in contact, they will appear as dissimilar as possible.
In other words, on two colours being seen simultaneously,
the complementary of the one will be added to the other.
Thus, if a yellowish green leaf and a red flower be under
the view at the same time, the yellow green will thereby
be more inclined to green, and the red will acquire a
slight tinge of blue, and the two will be brought more
nearly into the complementary state. In this way the
eye itself can rectify any slight defect in the harmonies of
adjacent colours.
CHAPTER IV.
THE VERTEBRATE SKELETON.
SECT. I. THE HOMOLOGIES AND HOMOTYPES OF THE
VERTEBRATE SKELETON.
In the last age there raged a famous scientific contro-
versy, which may be summarily represented as a dispute
as to which of the two great principles which we are un-
folding should he detected in the animal frame. The illus-
trious Cuvier, in building up the science of comparative
anatomy, proceeded, in all his investigations, on the prin-
ciple that every particular member of the body had a
special or final cause. On the other hand, the great
Geoffroy St. Hilaire, first the co-operator and then the
rival of Cuvier, delighted to trace a unity of plan running
through the bones of the skeleton. In 1830, this con-
troversy came to a public explosion, which was viewed
with intense anxiety by all interested in natural science,
and in particular by the poet Goethe, who proclaimed it
to be a far more important event than the French Revo-
lution, which was ringing that same year in the ears of
Europe. In conducting the dispute, extreme positions
were taken by both sides. Attached to the principle of
final cause, and having found how prolific it was, in his
hands, of brilliant discoveries, Cuvier was not willing to
admit the theory, (though he helped greatly to establish
the fact,) that there is in the skeleton a general corre-
176 THE HOMOLOGIES AND HOMOTYPES
spondence of parts, which can have no reference to the
wellbeing of the animal, or the special functions of the
organ. Geoffroy St. Hilaire, on the other hand, did not
see that his doctrine of analogy was perfectly consistent
with teleology, and he connected his theory of unity with
the untenable doctrine of the transformation of species.
This dispute should now be regarded as settled, by the
establishment of both doctrines — both that of general
homology and that of special teleology ; and the former,
we are convinced, will be found, when properly interpre-
ted, to yield as rich a contribution to the cause of natural
theology as the latter.
Any one may convince himself, very easily, that in a
general sense there are model forms in the construction
of the skeleton. He will see at a glance that every spe-
cies of animal has its normal shape, and this is, to a
considerable extent, determined by the length, thickness,
and relative position of its bones. In the human frame,
there are organs which have been used as standards of
measurements, which they could not have been unless
their size had been approximately definite. The length
of the arm, from the elbow to the tip of the mid-finger,
furnished the cubit to many nations of antiquity. The
hand-breadth and the span were measures among the
ancient Hebrews. In not a few countries the stretch of
the arms, the pace, the palm, the breadth of the thumb,
have been used to indicate linear measure. Among
artists the human frame has long been known to have
proportions in its members. The visible outline of the
head in front is divided into four equal parts ; — the first,
from the top of the head to the setting of the hair ; the
second, from this to the root of the nose ; the third, the
nose ; and the fourth, from the lower part of the nose to
the chin. The height of the figure is found to be eight
OF THE VERTEBRATE SKELETON. 177
beads ; the first reaching from summit of head to chin,
the second from chin to breast, the third from breast to
navel, the fourth from navel to top of thigh, the fifth to
middle of thigh, the sixth to knee, the seventh to the
calf of the leg, and the last to the heel. The body is
thus divided into two equal parts — one from head to hip,
the. other from hip to heel. The length of the frame is
also known to be equal to the line drawn from finger-tops
to finger-tops of the outstretched arms.
But without dwelling longer on these general topics,
we proceed to shew, in a scientific manner, that the ver-
tebrate skeleton consists of a series of pieces constructed
on a common plan ; and in doing so, we shall largely
avail ourselves of the masterly researches of Professor
Owen, who has done so much towards the completion of
this most interesting subject.
We know that the skeleton is not a peculiarly inter-
esting object to an untutored eye. It has been associated,
in the minds of many, with the grave's mouth and mor-
tality. It possesses in itself no physical beauty ; it is
meant to be wrapt up from the view by a covering of
flesh and muscles, which are made, for our gratification,
to present themselves in full and rounded forms. Still,
to minds which are fitted to penetrate beneath the sur-
face, it has become an object of intense interest, and is
felt to possess not a little beauty. The reason is, that
there has been a perception of the unity of the structure
along its whole length, and from the highest to the low-
est animal in the class, and of the suitableness of the
infinitely varied parts to their infinitely diversified func-
tions.
Each of the series of parts which makes up the verte-
brate skeleton is called a Vertebra. It will be sufficient
for our purpose to indicate here the principal parts of the
178 THE HOMOLOGIES AND HOMOTYPES
typical vertebra, without entering into those more minute
details which are necessary for the purposes of the com-
parative anatomist ; for these details we would refer to
Professor Owen's paper on the Megatherium, in the Phi-
losophical Transactions for 1850.
Typical Vertebra consists of a centre or body, around
which are arranged other pieces,
(called technically apophyses, or
projecting parts,) so as to form
two principal arches, one supe-
rior,* the other inferior. The
upper arch gives protection to
nervous matter, and is hence
called neural : it is bounded on
each side by two principal pieces,
called neurapophyses, and is
closed above by the neural spine,
ks so called from its frequently
Fl0-83-+ pointed form ; (it is, however,
sometimes bifid.) The lower arch, called haemal, pro-
tects blood-vessels, &c, (hence its name, from Greek,
haima, blood ;) it also consists of lateral pieces, called
respectively pleurapophyses and haemapophyses, and is
closed by the haemal spine, which, like the neural spine,
is sometimes cleft. The body of the vertebra may be con-
sidered the foundation of the arches, and the neural and
haemal spines represent, in position, the keystones of each.
Sometimes the upper arch comprehends a pair of bones,
called diapophyses, and the lower an additional pair, call-
ed parapophyses.
* In the erect position of man, these are respectively posterior and anterior.
t Fio. 33. Typical Vertebra; tos, neural spine; n, neurapophysis ; N, neural arch;
e, centrum, or centre piece; pi, pleurapophysis ; h, haemapophysis ; hs, hsemal spine ; II
haemal arch ; d, diapopbysis ; p, parapophysis.
OF THE VERTEBRATE SKELETON.
179
Generally speaking, it is not difficult to demonstrate,
that in the chain of bones extending from the head to
the tail inclusive, we have a series
of pieces partaking of the nature
of the common typical structure
just described. It is true that
some present a near approach to
the model, while in others the
rea. nature of the parts is consid-
erably masked, so that careful ex-
amination is necessary to show the
relation. Knowing the type, how-
ever, we can explain all departures
from it, whether owing to omission
or contraction, adhesion or compli-
cation of pieces.
As there is a model vertebra,
so there is an archetype skeleton,
and we shall transfer to our pages the instructive dia-
gram given by Professor Owen in his work on the
" Homologies of the Vertebrate Skeleton." The elements
of each vertebra are indicated by the peculiar shading, —
n, neurapophyses, . thus
d. diapophyses,
jo, parapophyses, .
c, centre or body, .
pi, pleurapophyses,
thus
thus
thus
thus
a
^s
ns, neural spine, and lis, hsemal spine, are left unshaded,
the appendages are represented by dots.
* Fig. 34. The relations of the parts in Fig. 33 will be rendered more evident by com-
paring it with Fig. 34; the references are the same in both; y, hypapophysis ; «, epapo-
physis.
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ophysis; the lower jaw represents hcemapophysis and
hcemal spine.
Third, or parietal vertebra. — The centre is formed
by the basisphenoid bone ; the alisphenoids are neura-
pojmyses; the parietal bones form the cleft and expanded
neural spine; the styloid pieces of the temporal bone are
pleurapophyses ; the lesser cornua of the hyoid bone
(lying in the upper and fore-part of the neck) are hcema-
OF THE VERTEBRATE SKELETON.
183
popliyscs; the body of the same bone forming the hcemal
spine, and thus completing the arch.
Fourth, or occipital ver-
tebra.— The basilar piece of
anatomists is the centre ; the
sides which bound the open-
ing in the occipital bone,
through which the upper part
of the spinal cord is continu-
ous with a portion of the cere-
bral mass, constitute the neu-
rapophyses surmounted by the
expanded neural spine. The
lower or haemal arch of this vertebra is removed from its
natural position in man, as in most vertebrata — the
reason of this will be discussed in a subsequent para-
graph. The scapulas or shoulder-blades are the pleu-
rapophyses ; their appendages, called coracoid processes,
constitute the haamapophyses ; and the haemal spine is
wanting, f
The seven vertebras of the neck are admitted to pos-
sess the general elements of the typical vertebra, the
parts, however, are generally considerably modified in
relation to the functions of that portion of the frame to
which they belong.
The vertebras of the back are twelve in number ; the
neural canal in each is sufficiently obvious, and of mo-
derate size ; the haemal arch is highly enlarged ; the ribs
* Fio. 30. Parietal segment, or vertebra of man. The neural arch Is ample, (n,) to
protect part of the brain : the hicmal (n) is contracted. In this case the diapophyses,
d, arc wedged between the neurapoph yses, n, and the large neural spine, m ,' e is a piece
called epapophysis, which lies upon the centrum, c ," a, h, and hs represent the parts of
the hyoid bone suspended in the upper and fore-part of the neck, and closing the ha;mal
arch.
t In order to simplify the subject, we have omitted reference to diapophyses and pa-
rapophyses.
184 THE HOMOLOGIES AND HOMOTYPES
are the pleurapophyses, succeeded by the hasmapophyses.
or cartilages of the ribs, and finally closed by the united
hcenial spines, which constitute the sternum, or breast-
bone.
In the five vertebras of the loins, the elements are not
so obvious as in those of the back. The pleurapophyses
are short, and firmly joined to the central portion ; the
hasmal arch is not completed by bony elements.
The five sacral vertebras have their bodies firmly joined
in the adult, and these same elements diminish in size
from the first to the last. The neural arch is complete
only in the first three ; the neural spines of the last two
are absent. The heemal arch of the first sacral vertebra
is usually considered as formed of that part of the pelvis
called ilium ; the portion called pubis is a hasinapophysis;
the ischium is the basin apophysis of the second sacral
vertebra.
The four or five succeeding and terminal pieces of the
back-bone in man, correspond to the tail in the lower
animals, and for the most part consist of the centra
only.
Such is a brief summary of the generally admitted
views held respecting the nature of the human skeleton,
(exclusive of the limbs, which will occupy attention in
subsequent paragraphs ;) and as such have been, in many
instances, arrived at by comparison with the bony frame-
work of animals lower in the scale, it is unnecessary to
allude to these under this department of our subject.
While, therefore, the entire skeleton in every vertebrate
animal is constructed according to a common plan, and
the series of vertebras of which it consists may all be re-
ferred to one model, it appears to us that there is good
reason for proceeding a step farther, and coming to the
conclusion, that unity of form also prevails in the indi-
OF THE VERTEBKATE SKELETON. 185
vidual pieces of the typical vertebra and its appen-
dages.
We may first allude to the appendages or limbs, as
affording the most evident indications of such unity. If
we take, as the typical form, a bone of the hand (meta-
carpal,) or of the foot (metatarsal,) we shall find that
there is a striking resemblance to it in all the elements
of every limb.* This typical bone may be described as
having a nearly cylindrical shaft, dilated towards its two
extremities. The large cannon bone in the foot of the
horse (see Fig. 43) may serve to illustrate the form
alluded to. Now, this is the prevailing shape in all the
principal bones of the limbs. In man, for example, such
general outline exists in the bones of the arm, fore-arm,
hand and fingers ; in thigh, \eg, foot, and toes. The
short and frequently irregular bones of the wrist and
ankle present the greatest departure from the type ; but
in some animals the relation is obvious enough. Thus,
in the common frog, certain of the ankle-bones (calca-
neum and astragalus of anatomists) assume exactly the'
typical form.
In the individual pieces of the vertebra itself, we shall
find evident traces of similarity to a typical form.. The'
centrum, or body of the vertebra, presents a close approach
to the model in the caudal part of the skeleton. This is
evident in a great number of instances. One may suffice:
the bones of the tail, in the young African elephant, con-
sist of centrum only, and each very much resembles in
form a metacarpal or metatarsal bone.
* It is a fact worthy of notice here, that the same form of an organ appears in plants.
For example, the stalk which supports the leaflets of species of ./Eseulus, the horse-
chestnut, exactly resembles a bone of the hand or foot; and in the manna-ash, we have
four or more pieces of like shape forming the main stalk of the compound leaf, separating
at the joints, and resembling a series of phalanges, as in a finger or toe. The same gen-
eral outline is often visible in the bole of well-developed trees.
186 THE HOMOLOGIES AND HOMOTYPES
The elements of the inferior or haemal arch present
very clear examples of conformity to the type. Pleura-
pophyses or ribs are not always curved and fiat bones,
such as we see in Mammalia generally, and in the New
Zealand bird called Apteryx. In not a few instances,
especially certain aquatic birds, (the guillemot, for ex-
ample,) the ribs are narrow and cylindrical, and bear
considerable resemblance to the lengthened bones of the
fingers which form the framework of the bat's wing. The
numerous ribs of the boa and other serpents, differ from
the model only in being curved. The shoulder or sca-
pular is a pleurapophysis, (sometimes with conjoined
hseroapophysis.) In man and mammalia generally it is
broad and flat, but in many birds it is long and narrow,
exactly like a rib ; and since, in some aquatic birds, the
ordinary ribs very much resemble the model shape, we
have thus transitional forms conducting us to the original
type. The pelvis, intended to support and protect im-
portant viscera, and give attachment to powerful muscles,
shews also striking departure from the model. But in
the frog, the iliac bones (pleurapophyses) very much re-
semble the typical form. We have evident examples of
likeness to our assumed model in the other elements of
the lower arch, viz., the rib and its cartilage, (pleurapo-
physis and hasmapophysis.) Mere curvature of the parts,
so as to assist in the formation of an arch, cannot be
considered as very materially affecting the conclusion to
be drawn. As regards the haemal spine, it would not
be easy to recognize any conformity to a primary shape
in the sternum or breast-bone of man or of a bird ;
but in many animals, such as the lion, elephant, walrus,
greyhound, &c, this part of the skeleton consists of
a linear series of pieces, exactly resembling the typical
form.
OF THE VERTEBBATE SKELETON. 187
In the elements of the superior or neural arch, the
departure from the model is generally greater and more
constant than it is in the lower or haemal arch. The flat
bones of the skull deviate widely from the type, but not
more so than the shoulder blade or the pelvis, both of
which, as we have seen, present transitional forms. The
very important functions of the brain-case, as a protector
of the important parts within, necessarily imply a great
and constant deviation from the model form. If we exa-
mine the principal element of the neural arch (neurapo-
physis) of any large vertebra, as in the baleen-whale, or
in the finner, we see that, after all, it may be referred to
the same general form which ribs assume, and they, as
we have seen, can be traced to a model bone. The neural
spine is indirectly referable to the same type, and by
similar steps. We observe it in the dorsal region of
ruminants, and other animals attaining great length, and
resembling a rib, being, however, straight. There is but
little difference in form between the longer neural spines
of the dorsal vertebras in the horse, and the first rib of the
same animal.
On the whole, we think there are evident traces of
community of form in the parts of the typical vertebra.
The subject is interesting, and merits attention and fur-
ther investigation by those favourably situated for oppor-
tunity of examining and studying the forms and transi-
tions in an extensive series of skeletons.
There are not only proofs of general order as regards
reference to a typical bone, vertebra, and archetype
skeleton, but there are some well-established facts
respecting the number of the vertebras themselves.
Those entering into the formation of the brain-case
in mammalia are four, those of the neck are seven,
except in the case of the three-toed sloth, which has
188 THE HOMOLOGIES AND HOMOTYPES
nine, and the manati, in which only six are said to
exist.*
The dorsal vertehra3 are usually considered as charac-
terized by the presence of long, arched, more or less
moveable, pleurapophyses or ribs, and, taking such as a
mark of distinction, we find that their number varies in
different cases.
In most carnivorous or flesh-eating animals, the num-
ber of vertebras of back and loins together is very con-
stant, though the exact number of those called dorsal
presents variations, as the following examples will
shew : f —
Back.
American Black Bear, . . 14
Dog, 13
Panther, 13
Spotted Hyaena, . . . 15
Glutton, 15
According to Professor Owen, all mammiferous ani-
mals, called Artiodactyles, as the ox, &c, having either
two or four toes, agree in having nineteen vertebras be-
tween the neck and the sacrum ; this is remarkable when
compared with the odd-toed group, usually called Peris-
sodactyles, which present great irregularity in the number
of the corresponding vertebras, there being, for example,
twenty-two in Khinoceros ; twenty-three in Tapir and the
Palasotherium ; and twenty-nine in Hyrax.
DIVERGING APPENDAGES OR LIMBS.
These constitute the limbs of animals, which are just
lateral appendages of the typical vertebra. The simplest
* According to Maclise, some of the monkey tribe have only five or six neck vertebrae,
und occasionally also in man the same occurs. — (Medical Times and Gazette, January,
1854.)
t Coote on Homologies of Human Skeleton, p. 26.
Loins.
Total.
6
20
7
20
7
20
5
20
5
20
OF THE VERTEBRATE SKELETON. 189
example of such appendage is very evident on examining
the skeleton of a bird. Attached to its ribs or pleurapo-
physes, there are seen short flat pieces, which, being
directed backwards, overlap the external surface of the
next rib behind. (See a, Fig. 37 ; also a and 65, Fig.
35.) Similar appendages are found, less perfectly de-
veloped, in certain reptiles. They also occur in the
abdominal parts of the most bony fishes, in which their
length is such that they reach even to the skin. They
are considered as parts of the primitive segment or ver-
tebra, though less constant than the arches which support
them. Now, the simplest form of limb is, in its nature,
but very little removed from such diverging appendage ;
in some of the lower vertebrata, as Protopterus, the limbs
are reduced to an unbranched ray.
Through various a
species of Amphiuma,
and in Proteus, we
observe greater com-
plexity, (though still
of low type compared
with the extremities
of man,) and this goes
on step by step in dif- FlG- 37-*
ferent animals, till we reach the arrangements which
characterize the higher forms. The Protopterus, whose
simple limbs afford proof of their identity with the
diverging appendages of the typical vertebra, present
also proofs that the fore and hind limbs are homo-
types, both being in that animal precisely of the same
simple nature. But even in the higher animals, man
* Fio. 3T. Occipital vertebra of Protopterus. The hcemal arch is large, consisting oi
pi, pleurapophysis ; h. hremapophysis ; hremal spine is wanting. The long, simple,
jointed ray, a, 57, is the diverging appendage or rudimentary limb.
190 THE HOMOLOGIES AND HOMOTYPES
for example, the resemblance is sufficiently obvious ;
the arm and thigh, fore-arm and leg, wrist and ankle-
joint, hand and foot, are the corresponding parts of
each limb ; these members are therefore homotypes.
But under whatever forms the limbs exist, they are sup-
ported by inverted arches, the presence of which is more
constant than that of the appendages which they support,
and for an obvious reason — the arch is required to pro-
tect certain important organs which are always present,
as the brain and spinal cord, heart and lungs ; the ap-
pendage of the arch comes in as a secondary instrument,
necessary, doubtless, in the economy of the animal ; but
yet less important in a general sense than the other or-
gans just mentioned.
The parts usually considered as entering into the form-
ation of the upper and lower limbs in man, are the
following : — The scapula, or shoulder-blade, and the
attached process called coracoid, represent respectively
pleurapophysis and haemapophysis of the occipital verte-
bra ; the clavicles, or collar-bones, are the hremapophyses
of the atlas, or first vertebra of the neck ; there is here,
therefore transference of arches (which are also imper-
fect) from their natural position ; — the end of this we
shall afterwards examine. Then follows the arm-bone,
next the two bones of the fore-arm, called radius and
ulna ; then the carpus, or wrist, composed of eight bones
apparently, but really of ten in two rows ; connected to
certain of these, we observe five bones of the hand called
metacarpus, then follow those of the fingers, styled pha-
langes, each digit having three, excepting the thumb which
has two.
The pelvic portion of the skeleton has been already
noticed ; it is in like manner an arch supporting diverg-
ing appendages, the lower limbs. Each of these consists,
OF THE VERTEBRATE SKELETON. 191
first, of thigh-bone, succeeded by the leg-bones, called
tibia and fibula ; then follows those of the ankle, the tar-
sus of anatomists, consisting apparently of seven bones
in two rows, which, however, really represent ten primi-
tively distinct pieces. Then follow five metatarsals, or
bones of the foot, and connected with their lower ends are
the toes, each, with the exception of the great toe having
three bones.
Now, whatever be the functions of the extremities in
any of the higher vertebrata, we find all, whether fore or
hind limbs, constructed on the same plan as that just
described, five being the typical number of digits. It
may be remarked how different is the relative develop-
ment of the digits, of thumb, index, middle, ring, and
little fingers, styled, respectively, 1st, 2d, 3d, 4th, and
5th in the human hand.* The first digit has only two
joints ; the fifth has the usual number, viz., three, but
the whole being short ; the second comes next in length,
then the fourth ; and the third is the most highly deve-
loped of all. These peculiarities have distinct reference
to the general permanence of these digits respectively,
and throw light on certain modifications observed in ani-
mals lower in the scale.
In the typical limb, the shortening of the thumb and
little finger, or the first and fifth digits, is a step towards
their disappearance,f the 2d, 3d, and 4th being more
permanent ; the two last reaching the ground in the ox,
and the longest of the two, namely, the 3d, is the only
one which serves as a jjoint of support in the horse.
Professor Owen remarks, that "a perfect and beautiful
* The same numbers are used to represent the toes ; great toe, number 1; little toe
number 5.
t A similer law reigns in certain plants. In CrucifertB, (cabbage tribe,) the stamens
»re usually six, four of these being longer than the other two. In Cardamine hirsitta
there are usually only four, the two shorter being absent.
192 SPECIAL ADAPTATIONS
parallelism reigns in the order in which the toes succes-
sively disappear in the hind-foot with that of the fore-
foot."*
Commencing with man as possessing the typical num-
ber, and descending to the lower animals, we find that
that digit, (the first, or thumb, viz.,) whose uses, par ex-
cellence, characterize him, is one of the first which disap-
pears. Departure from the typical five is a characteristic
of mammalia lower in the scale, hence the tetra-, tri-, di-,
and mono-dactyle limbs common among them.-j* Descend-
ing lower in the scale to fishes, we find the limbs present-
ing often (with a nearer approach to the simpler diverging
appendages) a less subordination to the typical number,
there being usually an excess. This, however, as Professor
Owen remarks respecting the pectoral fin of the skate and
its numerous digits, is not an example of complex devia-
tion, " true complexity not being shewn in the number,
but in the variety and co-ordination of the parts." In a
word, all diverging appendages or limbs are constructed
on a common plan ; we shall afterwards examine their
numerously diversified modifications for special ends. We
also observe in them evident traces of order as regards a
law of number, and a general rule in accordance with
which they are present or absent, as the necessities of the
animal require them or not.
SECT. II. — SPECIAL ADAPTATIONS IN THE STRUCTUEE O:-
THE SKELETON.
The subject here opened to us is of vast extent, and
even not yet thoroughly exhausted by all that has been
done in human and comparative anatomy. It must be
* On Limbs, p. 23.
t This has reference to digits which attain functional size.
IN THE STRUCTURE OF THE SKELETON. 193
acknowledged that the relation between special modifi-
cations or departures from the general plan, and final
ends of such, have not been determined as to every part
of the animal frame. Nevertheless, so many striking
examples present themselves to the careful and unpre-
judiced observer, that it may be considered a legitimate
conclusion that there is such a general relation, although
the cautious reasoner may hesitate to give a positive
decision in every instance which may come under his
notice.
We can indicate only some of the more obvious cases
illustrative of the coincidence between the principle of
order and that of special adaptation. We may appro-
priately open this part of our subject by glancing at the
modifications observed in the vertebrate series in man.
In the cranial vertebra? we observe two remarkable
contrasts in the development of the neural arches ; which
are more or less extended according to the purpose which
they serve in reference to the particular part of the brain
over which they are situated. The great size of the
nervous centre, that is, the brain, requires a correspond-
ing enlargement in certain neural arches, and this is
found to be actually provided. Each vertebra gives pro-
tection to corresponding parts of the nervous matter ;
thus, the cerebellum is protected by the occipital, the
mesencephalon (or middle portion)"* by the parietal, and
the prosencephalon (fore-part of cerebral mass) by the
frontal vertebra. In all of these the neural arch is
ample, in distinct relation to the size of the part requir-
ing defence. The less development, or rather nearly
complete obliteration, of the neural arch in the first or
nasal vertebra,, is commensurate in man (and other ani-
* Comprehending also Pons Varolii, Corpora quadiigemina, pituitary body, and third
ventricle.
194
SPECIAL ADAPTATIONS
Fig. 38.*
mals besides) with the small size of the remaining portion
of the brain mass represented by the olfactory ganglia.
It is by means of the first and second vertebrae of the
neck that free rotation
of the head is effected.
The anterior part of
the first (forming a
portion of its centre)
is excavated, in order
to receive the tooth-
like projection 0f the
second, or axis, which
is so called because there rises from the upper part of
its body a piece, round which the first, or atlas, plays as
on a pivot, giving rise to the lateral movements of the
head. The base of this pivot is in reality the body of
this second vertebra ; its apex,
however, is formed of part of the
body of the first, removed from
its natural position, and united
to that of the second. Now, we
do not consider it any strained
inference when we affirm, that
there is here presented to us
a notable instance of special
adaptation for a particular function.
Generally speaking, the haomal arch is imperfect in the
vertebras of the neck, because the large size of its ele-
ments (viz., pleurapophyses, hasmapophyses, and hasmal
* Fig. 38 represents the first neck-vertebra in man : it is called atlas, as supporting
the head. A strong ligamentous band stretches across the large central opening, and
divides it Into two. The tooth-like projection of Fig. 39 is received into the fore-part
of this divided ring, the posterior allows passage to the spinal cord.
+ Fig. 39 is the axis or second vertebra of the neck in man. The apex of the tooth-
like projection is part of the centre or body of the atlas, joined to the body or centre of
the axis.
Fio. 39.t
IN THE STRUCTURE OF THE SKELETON. 195
spiii^) would have interfered with free motion in this
region of the body. Nevertheless, certain parts which,
on a cursory glance, appear to be absent, are in reality
present, but are specially modified by decrease and coal-
escence ; thus, the portion of a cervical vertebra project-
ing outwards on each side, and hence called by anatomists
the transverse process, in reality consists of diapophysis,
parapophysis, and a short pleurapophysis or rib, firmly
joined, but together forming a hole or short canal and a
groove, to give protection and support to a blood-vessel
and nerve respectively. The excessive development of
the haemal arch in the dorsal vertebrae, is a provision for
the large and important organs to be protected — the
heart, lungs, &c. The elastic and moveable ribs (pleura-
pophyses) and their cartilages (haemapophyses,) are ad-
mirably adapted to the exhalation and inhalation of at-
mospheric air during the act of breathing.
The vertebras of the loins are large and strong, thus
affording a firm basis of support to the superincumbent
column ; the haemal arch is not completed by bony ele-
ments, but by soft elastic walls, which yield to the vary-
ing expansion of the viscera within.
The union of the sacral vertebras gives additional
strength to this portion of the column, supporting, as it
does, the elastic spine above it. The excessive develop-
ment of that part of the haemal arch — the pelvis of ana-
tomists— is obviously intended to support and protect the
larger viscera, and to present a surface of attachment for
powerful muscles. The united bodies of the coccygeal
series, forming a partial concave floor to the pelvis, afford
additional support to the organs protected by this last.
In short, while the skeleton of man consists throughout
of a series of parts all formed on one model, yet there is
a wide range of difference in most of them, and the special
196 SPECIAL ADAPTATIONS
modifications have in all oases a very decided, and, in
most instances, a very obvious relation to the development
of different organs, without which our goodly frame could
not perform its functions, or even continue to exist.
We may now examine some of the special modifica-
tions of vertebral elements, as exemplified by animals
lower in the scale ; from a multitude of instances, our
limits constrain us to select only a few. Whether we
examine fishes, reptiles, birds, .or mammals, we shall find
obvious illustrations of departure from the model or type
in accordance with some function neebssary to the very
existence of the animal.
In Ophidia, or serpents, certain elements of the two
anterior cranial vertebrae are freely moveable on each
other, instead of being closely joined together, as is
usually the case ; strength and firmness are here sac-
rificed to mobility and expansile power of the parts,
and why ? The arrangement has a clear and express
relation to the mode of feeding ; serpents often swallow
very large prey entire ; but this they could not do were
the parts firmly banded together. As it is, the mouth is
capable of great extension, and the prey is taken in with
ease.
In fishes there exists a remarkable concentration of
important organs in the fore-part of the body. The head
contains, not merely the brain and organs of the senses,
but, in addition, the heart and gills ; we find, accord-
ingly, that the haemal arches are commensurate in size
with the presence of the important parts which they sup-
port and protect. In the words of Professor Owen,
"Brain and sense-organs, jaws and tongue, heart and
gills, arms and legs, may all belong to the head ; and
the disproportionate size of the head, and its firm attach-
ment to the trunk, required by these functions, are pre-
IN THE STRUCTURE OF THE SKELETON. 197
cisely the conditions most favourable for facilitating the
course of the fish through its native element."
In the whale, the vertebrae of the neck are joined into
one solid column. By this arrangement, greater protec-
tion is afforded to the nervous cord, as this large and
heavy animal ploughs its way with rapidity through the
water. Flexibility in the neck, not needed in this case
for other purposes, would have been an inconvenience.
The three-toed sloth presents an example the very
converse of the last; the additional vertebras (we have
already alluded to it as an example of departure from
the typical number) in the neck of this animal are ad-
mitted to have a relation to its habits ; in the words of
Professor Bell, " the object of the increased number of
vertebras is evidently to allow of a more extensive rota-
tion of the head ; for, as each of the bones turns, to a
small extent, upon the succeeding one, it is clear that
the degree of rotation of the extreme point will be in
proportion to the number of pieces in the whole series."*
But, in addition, as this animal spends its whole life on
trees, clinging to the branches with its powerful limbs,
and feeding on the twigs of its arboreal dwelling-place,
the length of its neck gives it an advantage in better en-
abling it to reach the tender and extreme branches.
In carnivorous animals, having four limbs fitted for
seizing and holding their living prey, and a mouth armed
with strong teeth for tearing it, the neural spines and
transverse process of certain neck- vertebras are highly
developed, so as to become commensurate with the power
of the oblique muscles of the head, which are in them of
great strength, to enable them to perforin their impor-
tant functions. In other words, the levers supplied by
certain elements of the neck vertebras are in direct pro-
* Cycloptedia of Anatomy and Physiology, Article Edentata.
198 SPECIAL ADAPTATIONS
portion to the active organs of motion, that is, the muscles,
which require them as mechanical powers.
In birds, the fore-limbs are used in flight, and the
function of the arm is transferred to the neck, that of
prehension to the beak, which supplies the place of the
hand. The neck is the only flexible, part of the verte-
bral series, and motive power is abundantly provided for
on the same principle as we have seen it to be in the
sloth. It is curious to notice that there is a departure
from the number seven, so constant in mammals ; the
vertebras ranging from nine in the sparrow to twenty-
three in the swan. The mode of connexion of the ver-
tebras is also such as to admit of the utmost freedom of
motion.
In the dorsal portion of the vertebral series, we may
also note a few striking adjustments. In certain mam-
malia, as the ox, deer, camel, &c, owing to the weight of
the horns and antlers, or length of the neck, continued
muscular exertion would be necessary, in order to retain
the head in its natural position. Such disadvantages is
obtained by the presence of the part called pax-wax, or
ligament of the neck — composed of yellow elastic fibres —
which acts as a natural spring, and obviates the need of
constant voluntary muscular effort. Accordingly, we
find that certain neural spines in the back (as well as in
the neck) are greatly elongated, to give attachment to
the remarkable organ referred to. In the aurochs, for
example, some of the dorsal vertebras have neural spines
which are actually longer than some of the ribs. Such
modifications are, indeed, generally observed in browsing
animals.
As in the fish, excessive development of certain parts
of the skull is a provision for the forward position of the
heart and sills, so in air-breathing animals the lower
IN THE STRUCTURE OF THE SKELETON.
199
arches of certain other vertebras are highly developed,
forming the ample thorax or chest, for the protection of
their heart and lungs. In the neck of the bird we have
seen that flexibility is necessary; in the back, firmness is
the essential requisite, and we observe there union of ver-
tebras. Further, the hasmapophyses, which in man and
others are cartilaginous, become in the bird converted
into bone, and the united hasmal spines from the keel of
the sternum or breast-bone, the extent of the surface pre-
sented by which is directly as the development of the
powerful muscles which are attached, and directly also,
of necessity, as the powers of flight.
In the ostrich, and cursorial birds
generally, which cannot fly, the
hasmal spines do not form any crest.
In birds, we also observe union be-
tween the vertebrae of the loins, an /#[|
arrangement admirably calculated to
give firm support during the powerful
and rapid movements in flight •
Coalescence of the remaining ver-
tebras, in the adult human subject
we have seen to be the usual arrangement, and this —
together with size particularly in the sacrum — appears
to have relation to the erect posture of the body. In many
mammalia, the sacrum is proportionally narrower than in
man, and coalescence of vertebras is not the law ; but in
certain species, which have the faculty of assuming the
erect or semi-erect posture, as some monkeys, bears, and
certain rodents, the sacral portion of the skeleton is pro-
portionally stronger .than in others which have no such
* Fig. 40. Thoracic segment or vertebra of raven. The ha-mal arch is ample in ac-
cordance with its functions as a protector of heart, lungs, &c, and as furnishing surfaces
for attachment of powerful muscles. References are same as in preceding figures.
200 SPECIAL ADAPTATIONS
faculty. The permanently separate condition of the
sacral elements in the beaver is an arrangement admir-
ably suited to its peculiar habits, "using, as it does, not
only its long and powerful tail, but even the whole pos-
terior half of the trunk, as an" organ of propulsion through
the water."*
In man, as we have already seen, the terminal portion
of the spine, forming the coccyx, consists of a few small
pieces, reduced to little more than the centrum or body of
the vertebra. But in many of the lower animals, the tail
performs important functions, and attains higher develop-
ment. Sir John Richardson, in his account of a journey
through Prince Rupert's Land, mentions a curious case
of departure from the usual type in the bovine family,
which is generally characterized by the high development
of the terminal portion of the vertebral series. He says,
" The musk-ox has the peculiarity, in the bovine tribe,
in the want of an evident tail ; the caudal vertebrae are
only six in number, being very flat, and nearly as short,
in reference to the pelvis, as in the human species. A
tail is not needed by this animal, as, in its elevated sum-
mer haunts, moschetos and other winged pests are com-
paratively few, while its closer woolly and shaggy hair
furnishes its body with sufficient protection from their
assaults."
The special modifications of the elements of the caudal
portion are numerous, and have an obvious reference to
final cause or end to be served. In the human coccyx
there is no hasmal arch. In the tails of not a few ani-
mals, lower in the scale, it is distinctly formed of haama-
pophyses and haemal spine. The prehensile tails of the
spider monkeys, the powerful oar-acting tail of the
beaver, and the supporting pillar-like organ in the kan-
* Cooto oa Homologies of Skeleton, p. 61.
IN THE STRUCTURE OF THE SKELETON. 201
garoo, present individual peculiarities of the vertebral
elements admirably suited to the different uses of the
part. In the kangaroo the lower surface of the tail is
subject to pressure, and the same is true of the prehen-
sile tails in Phalangista and the opossum, and in all these
the haemal arch is well developed, in order to protect the
blood-vessels. " In Pctaurus, Phascogale, and Dasyurus,
the tail acts as a balancing pole, or serves, from the long
and thick hair with which it is clothed, as a portable
blanket, to keep the nose and extremities warm during
sleep. The haemal arches in the tails of these are not so
largely developed as in the kangaroo &c, their mecha-
nical office of defending the blood-vessels of the tail from
pressure not being required."*
It is admitted that the typical structure may be
departed from by excess in the number of the elements ;
if it can be shewn that such departure has decided
relation to the habits and wellbeing of an animal, it
appears to us a powerful argument in favour of com-
bined order and adaptation ; we may here adduce a few
examples.
Seals and penguins are not fitted for general sojourn
and progression on the land, nevertheless they do occa-
sionally frequent the shore, but their movements, under
such circumstances, are peculiar. One of the highest
authorities to which we can refer, specially alludes to
these animals, and to modifications in certain vertebras
related to the habit in question. In the Greenland seal,
Professor Owen describes processes superadded to the
lower surface of the lumbar vertebrae, (hypapophyses,)
"indicating great development of anterior vertebral mus-
cles, relating to peculiar gasteropod progression on land.
In penguins, similar hypapophyses attain their maximum
* Owen, in Proceedings of Zoological Society, 1838.
202 SPECIAL ADAPTATIONS
of development, and have an analogous function to that
in the seals, extending the surface of attachment of the
powerful muscles on the ventral aspect of the vertebral
column, which act in the shuffling gasteropodal move-
ments."*
In the armadillo, whose bony armour (giving to the
animal its name) is of considerable weight in proportion
to the size, and serves as a defence against its powerful
foes, we find two additional spines (metapoplryses) de-
veloped, one on each side of the neural spine, upon the
principle that three points are better fitted than one to
support a superincumbent weight. Certain serpents feed
upon the eggs of birds; their teeth are few and feeble — for
if the shell of the egg had been broken in the mouth, the
want of flexible lips would have occasioned loss of the
nutritious contents. Besides, these serpents follow the
law of their congeners ; loose attachment of cranial ele-
ments, as we have shewn, enables them to take their food
entire. The egg, being thus received, is ripped open as
it passes along the gullet, and this is effected by a con-
trivance no less remarkable for its simplicity than for its
efficiency. Sharp projections (hypapophyses) from cer-
tain vertebras of the neck, perforate the tube of the gul-
let, are capped by hard enamel, and effectually perform
their proper office.
We shall close this part of our subject by alluding to
two notable instances of special modifications pervading
almost the whole skeleton in serpents and tortoises. In
the former we find a long series of vertebras, some of
whose elements supply the place of limbs, which are
generally wanting, or, if present, as in boa, so rudimen-
tary as to be incapable of performing their usual func-
tions. The pleurapophyses have free motion, and act as
* Professor Owen, in Philosophical Transactions. 1851.
IN THE STRUCTURE OF THE SKELETON. 203
efficient organs in progression on a hard surface, by
means of the large scuta or shield-like scales covering
the belly of the animal. These scuta form a number of
movable broad surfaces, bearing the same relation to
the ends of the ribs which hoofs do to the ends of the
toes. In pelagic serpents which swim by lateral motion
of the tail, the pleurapophyses are more freely movable
m a lateral than in a backward direction, progress in the
water being accomplished by rapid lateral curvatures of
the tail and body. The neural spines of the dorsal ver-
tebra? are small, those of the caudal portion large and
compressed, and gradually lessening in size to the point
of the tail — a peculiarity of these vertebral elements in
strict harmony with the general compressed state of the
body, and that of the short but strong and flat tail,
which acts as an oar for propelling, as well as a rudder
for guiding.*
The spinal column of the Ophidia shews the maximum
of number of the different vertebras, and of flexibility as
a whole. In the words of Professor Owen, "At first
view, the principle of vegetative repetition seems to have
exhausted itself, in the long succession of incomplete ver-
tebras which support the trunk of the great constrictors ;
but by the endless combinations and adjustments of the
inflections of their long spine, the absence of locomotive
extremities is so compensated that the degraded and
mutilated serpent can overreach and overcome animals
of far higher organization than itself; it can outswim
the fish, outrun the rat, outclimb the monkey, and out-
vvrestlc the tiger ; crushing the carcase of the great
Carnivore in the embrace of its redoubled coils, and prov-
ing the simple vertebral column to be more effectual in
the stiuggle than the most strongly-developed fore-limbs,
* Dr. Cantor, Transactions of the Zoological Society.
204 SPECIAL ADAPTATIONS
with all their requisite rotatory mechanism for the effective
varied application of the heavy and formidably armed
paws."*
As the serpent shews us the highest possible flexibility,
so does the tortoise exhibit the greatest rigidity and in-
flexibility of vertebral elements, intended also to accom-
plish an end necessary to the wellbeing of the animal.
The carapace or upper arch, and plastron or floor, of the
turtle's or tortoise's shell may be compared to the skull ;
to use the expression of Professor Owen, it is actually
" an abdominal skull, formed of the centra of back,
loins, and pelvis united together, their pleurapophyses,
haemapophyses, and other elements, being expanded and
laterally adherent ; appendages of the skin — the der-
mal bones — are connate with some of the vertebral
elements, the whole forming a defence to a well-deve-
loped system of hiemal organs, heart, lungs, and alimen-
tary canal."
DIVERGING APPENDAGES OR LIMBS.
These assume various forms, from the simple structure
which we have noticed in the thorax of the bird up to
the perfectly developed limbs of man. Among them re-
markable modifications present themselves, having evi-
dent reference to the uses of the member, whether for
grasping, supporting the body, flying, swimming, leaping,
or burrowing. f The inference from all these adapta-
* Owen on Nature of Limbs, p. 90.
1 In a former paragraph (p. 1SB) we have shewn evident traces of community of form
In the elements of the vertebra and its appendages ; in reference to the modifications of
the latter, it will be necessary here to allude to the very ingenious but, we think, over-
strained, views of M. Gervais — (Ann. des. So. Naturelles, 1853.) According to Duges,
there is an arithmetical progression in the number of the parts from arm to fingers,
and from thi'srh to toes, viz., arm and thigh, each of one piece, leg and fore-arm, each of
two pieces; in wrist and ankle, hand and foot, fingers and toes, the number five prevails
we have therefore the progression, 1, 2. 5. M. Gervais thinks he finds proof that in
IN THE STRUCTURE OF THE SKELETON. 205
tions of means to end cannot be explained away "by
affirming that the animal, finding that it has an organ
suited to a certain purpose, uses it for that purpose. For
in the first place, the creature is compelled to a certain
mode of life by its instincts, which are altogether differ-
ent from its limbs or any of its organs ; and, secondly,
its limbs are suited to its other organs, and all its organs
are suited to one another. There is in all this no wis-
dom or foresight on the part of the animal, but there are
arrangements made for its welfare by a Power above it,
causing independent organs and instincts to concur and
co-operate.
It may be laid clown as the common rule that the pec-
toral and ventral limbs are appendages of the fourth and
twenty-six segments of the vetebral series.* The oc-
cipital is always the fourth vertebra, the pelvic may be
less constant in its position. But displacement of verte-
bral appendages from their typical position in the skele-
ton is not uncommon, and will generally be found to be
a provision for some peculiarity of function. In most
fishes, the pectoral fins, which are its arms, occupy the
typical position, being in connexion with the occipital
vertebra, whereas in man, and many other animals, the
same limbs are removed from their natural position, and
are attached to the upper part of the chest. These dif-
ferent dispositions are admitted to be, in the one case as
in the other, admirably adapted to the necessities of the
animal. Professor" Owen, referring to such modifica-
tions, remarks, " Wherever either arch with its appen-
dages may be situated, it is in its best possible place
limbs of vertebrata the number Ave prevails even in the arm and fore-arm, thigh and
leg, and that therefore there is union of bones in these parts. If this view should prove
to be correct, such union may be considered as a special modification of the type in re>
lation to the functions of the parts.
* We adopt here the views of Professor Owen.
206 SPECIAL ADAPTATIONS
in relation to the exigencies and sphere of life of the
species."*
We may next examine some of the principal modifica-
tions of the diverging appendages themselves, and of their
elements, traces of a general plan having already been
pointed out, and proofs adduced that law and order pre-
vail also in departures from the type. Although the
limbs of animals are diverging appendages of the typical
vertebra, all such apjDendages do not necessarily perform
the functions of limbs. Their simplest and most rudi-
mentary condition has been already alluded to as they
are seen in the thorax of the bird, where they appear to
serve merely the purpose of giving additional strength
and firmness to the ribs, (pleurapojihyses,) from which
they originate.
In the head of the fish we observe them offering greater
.advance in development, and in beautiful harmony with
their proper function. Those of the third or parietal
vertebra constitute the parts called, technically branchi-
ostegals, which, in most fishes, support a flap, whose
function is to assist in protecting the gills, and regulat-
ing the admission of fresh currents of water to these
important organs. The diverging appendages of the
second cranial vertebra are modified to form the opercu-
lar bones which together constitute the framework of the
gill-covers, by the movements of which the amount and
direction of the respiratory currents are principally deter-
mined. The corresponding part in the anterior segment
of the head consists of two pieces called pterygoids, the
outer of which serves as a means of connexion between
the haemal arches of the first and second vertebra?.
How different, then, the forms and uses of corre-
sponding appendages in the head of the fish, for, in
* Owen on Limbs, p. 81.
IN THE STRUCTURE OF THE SKELETON. 207
contrast with those just mentioned, we observe the
appendages of the fourth or occipital vertebra forming
the j)ectoral fins, which correspond to the upper limbs in
man, and perform an important, though not the prin-
cipal part in aquatic progression. The beautiful harmony
which subsists between the uses of the pectoral fins and
their peculiar structure, has been so frequently and fully
discussed in works of Natural Theology,'"''5 that it would
be needless to go over the same ground here. In the
frog-fishes, which have the power of moving on the
ground when left by the receding tide, in the expanded
pectorals of the flying-fish, acting as parachutes during
its powerful aerial leaps, in those of the climbing perches
of the tropics, and in the ordinary forms presented by the
fins of most fishes, we observe modifications of parts
constructed after the same model, but each in striking
unison with the habits of the animal. In the fish, then,
the fore-limbs (pectoral fins) are the diverging appen-
dages of the occipital vertebra, and occupy their natural
position as such, (that is, are placed far forwards,) being
attached to the hind-head. In other vertebrata, the arch
which supports them is transferred from its normal place
to the upper part of the trunk, and this transference, and
the structure of each piece, are admitted on all hands to be
in complete harmony with the function of the limbs, and
necessary to the comfort and wellbeing of the animals.
In birds, for example, the parts supporting the ante-
rior limbs are modified, so as to fit the diverging appen-
dage, to become an organ of flight. It has been already
mentioned, that the scapula and coracoid are respectively
pleurapophysis and htemapophysis of the occipital ver-
tebra, and the clavicles or collar-bones the haemapophyses
of the atlas, or first cervical vertebra. The relation of
* See Paley ; also Bell on the Hand, Roget's Bridgewater Treatise, &c.
208 SPECIAL ADAPTATIONS
these to the appendages which they support are such,
that the two can only be instructively studied together.
This is specially true of birds. The great strength of
the coracoid qualifies it for its main function, namely,
to give attachment to the limbs, and afford a strong basis
of support in flight during the quick and powerful strokes
of the wings.
The function of the pectoral wings in the bird being
peculiar, we find corresponding modification in the hard
parts. Appendages of the skin, the feathers namely,
serve the purpose of resistance to the air in flight, and so
the full development of digits, with freedom of motion
in the other bones of the limb, is not needed. The fore-
arm and the wrist-joint are so constructed, that free
motion is sacrificed to firmness, and the bony framework
of the hand is rudimentary ; nevertheless, the parts are
not so obscured that we cannot indicate their relations,
for in every case the general type still prevails, and each
finger corresponding to the second, third, and fourth of
the archetype, is clearly visible." The diverging appen-
dages, suspended to the pelvic arch, and forming the
lower limbs, are equally in harmony with their function
in different birds : this is more specially observable in
the metatarsal portion and toes. The part commonly
called leg in the bird, consists generally of three meta-
tarsal bones, which, in the adult, are so firmly united
as to give the appearance of one only. There is also, in
most instances, another, which is, however, small, and
whose function is to support the inner toe. The posi-
tion of this inner digit has an express adaptation to the
habits of the bird ; being on a level with the other toes
in perching birds, and therefore admirably fitted to give
* We may refer here to a former paragraph (p. 191) respecting the relative lengths of
the digits and their permanence.
IN THE STRUCTURE OF THE SKELETON. 209
increased power of grasping ; whereas it is removed
higher and higher in different waders, and is finally ab-
sent in cursorial birds, as the emeu and others. Still
greater reduction in the number
of the toes takes place in the
ostrich, the third and fourth alone
remaining. There is final cause
in all this ; " whilst unity of de-
sign is clearly manifested, the
wisdom of the Designer is dis-
played, by the greater strength
which results from the minor
degree of subdivision of the part I1&.41.*
which takes the largest share in the support and propul-
sion of the body."f
Among mammalia we find instruments for support,
grasping, climbing, running, leaping, burrowing, flying,
swimming, and diving. Now, it is distinctly observable,
that whatever be the function of the limbs, all are con-
structed after the same plan, but varied to suit the end
which is required by the instincts of the animal and its
allotted sphere. It has been already stated that the arms
and legs of man are homotypes, and that the individual
parts which form them are also homotypes ; yet while so
far corresponding to each other, the two scries are made
to differ in order to suit them to their several uses. The
harmonies of structure and function in each of the limbs
of man, as well as in other animals, have been so fully
discussed in different works, that it is unnecessary for us
to enlarge much on the subject here.
The fore-limbs of that expert tunnel-maker, the mole,
are admirably suited to its habits. The bones of the arm,
* Fio. 41. Foot of Ostrich, consisting only of digits 3 and 4.
t Owen o.i Limbs, p. 105.
210 SPECIAL ADAPTATIONS
fore-arm, wrist, hand, and fingers, are of great strength
in proportion to the size of the animal ; the whole limb
is short and broad ; the fingers are armed with strong
nails ; and the general conformation is such that the
action of the powerful muscles renders the hand a most
effectual instrument for burrowing in the soil, through
which, as Professor Owen remarks, the animal may be
said to swim.
The chief part of the bat's wing consists of a highly
developed hand. The junction of the arm-bone with the
shoulder-blade is in harmony with its function, being
such as to permit upward and downward motion chiefly.
It is interesting to observe that the collar-bone is longer
and broader in those whose flight is more powerful, such
as the insect-eating species, than in those which subsist
on fruit, and which, consequently, do not need swiftness
in pursuit of food. The arm-bone is long and slender in
all bats, the fore-arm is also long, and the two bones
which form it are incapable of rotation on each other, as
such movement would have lessened the impulse of the
wing. The bones of the hand and fingers, excepting
those of the thumb, are of great length, in order to add
to the superficial extent of the part, which is, in fact, a
highly developed hand, the long fingers being connected
by a web to the very tips. The small, but free, thumb,
with its well-developed nail, enables the animal to cling
easily to perpendicular surfaces as well as to climb
them.
When we examine animals remarkable for their powers
of progression on a hard surface, we meet with singular
deviations from the typical limb, in respect not only of
the number, but also the union of parts ; in every in-
stance structure and functions are clearly in harmony
with the kind of life for which the animal is intended.
IN THE STKUCTUKE OF THE SKELETON.
211
In the ox, the parts called, in common language, the knee
and the leg, do not, in fact, correspond to these portions
of the typical member. The so-called fore-knee is the
wrist., with the hones considerably modified ; the so-called
fore-leg is, in reality, part of a hand composed of two
metacarpal bones firmly united together in the adult
state, corresponding to the third and fourth, and together
forming the cannon-bone. On either side are the rudi-
ments of the second and fifth digits ; the thumb has dis-
appeared, and those which attain functional size, in other
words, which reach the ground, are the third and fourth.
" The rudiments of the second and fifth digits
are not without their use. When the elk or
bison treads on swampy ground, the hoofs ex-
pand, the false hoofs are pushed out, and the
resisting surface is increased as the foot sinks ;
but when it is lifted up the small hoofs col-
lapse to the sides of the large ones, which con-
tract, and, by their diminution of size, the act
of withdrawal is facilitated. In ruminants,
confined to arid deserts, we should hardly ex-
pect to meet with the mechanism which seems
expressly adapted to the marsh and the swamp,
and, in fact, every trace of the second and
fifth digits has disappeared from the foot of fig.42*.
the camel and dromedary."f
In the horse, there is still farther reduction in number,
only a single digit, the third, reaching to the ground and
serving for support. The perfect adaptation of the limbs
* Fig. 42. Foot of the ox. At the upper part are seen the bones of the ankle, viz.
fc, the cuboid; oe, ectocuneiform; s, scaphoid; a, astragalus; cl, calcaneum; succeed-
ing which is the cannon-bone, composed of the third and fourth metatarsal joined to-
gether. HI. and IV. are digits ; II. and V. are digits in a rudimentary condition ; 66 and
C7 are bones of the leg.
t Owen on Limbs, p. 34.
212
SPECIAL ADAPTATIONS IN THE SKELETON.
Fig. 434
of the animal to its habits and power of rapid progres-
sion is so generally admitted, and has been so frequently
discussed,* that it would be a work of super-
d( ; erogation to go over the same ground here.
Suffice it to say, the principle of reduction in
number of the parts, which we have already
seen to be associated with swiftness of progres-
sion in the foot of the ostrich, is carried out
fully in that of the horse : "he paweth in the
valley, and rejoiceth in his strength."
In conclusion it may be added, that there is
distinct evidence of arrangement in the height
of the limbs in most mammalia, and this is true
of both fore and hind extremities : their length
is equal to that of the head and neck together ;"f
the browsing species, therefore, can easily reach
the herbage necessary for their subsistence.
Bimanous and Quadramanous animals (man and the
apes) are exceptions ; but then they have limbs whose
digits are fitted to procure food. Certain others may be
noted as exceptions, namely, bats, which feed on wing,
and the elephant, whose short neck and prominent tusks
would prevent browsing, but in compensation, it is pro-
vided with that "grotesque hand/' the trunk, which is
merely a nasal apparatus, having superadded to its ordi-
nary function that of sensation and prehension.
* See Bell on the Hand, etc.
t Straus-Durckheim, Theologie arts are not always in strict accordance with their true
nature, but it so happens in this particular instance, that
the term Cephalopod is homologically correct, for the
appendages which surround the fore-part of the animal
in reality correspond to the
lower surface or foot, being ac-
tually lateral appendages of that
part. These • organs vary in
number ; in some species there
are eight, in others ten. In the
well-known Argonaut, two of
the appendages are webbed, so
as to present considerable ex-
tent of surface. These were
described by Aristotle as the
sails of the animal, which, in
fine weather, and when floating
on the surface, it expanded and raised to catch the
wind — a description which, as it is now well known,
does not indicate the true use of these parts ; for their
function is to form the shell, and progression is accom-
plished by the forcible ejection of water from the funnel,
the animal being urged on its course by the recoil.
* Fig. 48. Plan of cuttle-fish, to shew its relation to the archetype, pp, ms, mt, the
parts of the foot modified to form the arms which surround the head ; ep, epipodium
forming the funnel through which water is discharged. The alimentary canal and heart
will be seen in the middle of the shaded part of this figure.
ARCHETYPE MOLLUSC. 229
In those with ten appendages, two are longer than the
others, and serve as anchors to moor the body, or are
darted out to capture prey beyond reach of the shorter
arms.
Allusion has been made to the functions assigned to
the funnel ; this part, so necessary in the economy of
the animal, may be also referred to its corresponding
part in the archetype. It is derived from 'the epipodium,
upper foot, {Fig. 48. ep,) the posterior part only is con-
sidered by Professor Huxley as contributing to the for-
mation of this important organ. " The mouth is thrust
back between the halves of the mesopodium, the propo-
dium and mesopodium forming a continuous sheath —
bearing tentacles — around the oral aperture. The two
halves of the epip'odium united form the funnel."*
Pteropoda. — The animals so denominated are gene-
rally of small size, but this is compensated for by their
numbers. In the tropics, as well as in the Arctic
seas, they abound, and, with other marine invertebrata,
serve to stock the pasture-grounds of the great whales.
The peculiar appendages, or lateral flaps, from which
they derive their name, {Pteropoda, wing-footed,) are
the principal means of progression by which they flit
hither and thither — whence they have been approjiriately
called the moths and butterflies of the ocean. As littoral
productions they are not generally known, excepting
from the shells of some which are occasionally cast up ;
but in the open sea, far from land, they are sufficiently
familiar to the observant navigator.
In these interesting molluscs, the parts called fore,
middle, and hind foot, are generally in a rudimentary
condition, and the epipodium or upper foot forms the
wing-like appendages so necessary in the act of progres-
* Knight's English Cyclopedia, Art. Mollmoa.
230 MODIFICATIONS OF THE
sion, and giving such a marked character to these ani-
mals. Cleodora, Euribia, Clio, Pneumodermon, and
others, present each peculiar but minor modifications of
the epipodium, doubtless in harmony with the habits of
the respective species, but, nevertheless, essentially of the
same nature, and performing the same general function.
The epipodium, which is but a narrow band in the
archetype, appears, therefore, to attain its maximum of
development in certain Pteropods, and forming wing-like
appendages copiously traversed by strong muscular fibres,
is admirably fitted to be employed as oars, and the testi-
mony of observers confirms such idea respecting its use.
— There are other mollusca not far removed in appearance
from those just described, which also deserve to be no-
ticed here as examples illustrative of the argument. They
have been called Heteropoda. Like the Pteropoda, they
are constituted for free progression in the water. The
relations of their parts have been very fully examined by
Professor Huxley in the Essay already quoted. The body
in one genus, namely, Firola, is clear as crystal, so that all
its internal organs can be distinctly seen, and the author
quoted describes it " as hardly distinguishable in the
water, except by the incessant napping of its flattened
ventral appendage." The shape of this organ, by which
the animal makes progression in the water, is that of a
cheese-cutter ; it is a modification of the propodium or
fore-foot of the archetype, the other parts remaining
rudimentary. In another genus, viz., Atlanta, progres-
sion in the water is accomplished by means of an appen-
dage similar to that of Firola, and a modification of the
same part, thus remarkably constituted to serve an im-
portant end in the economy of the animal. But Atlanta
has the power of attaching itself to marine plants by
means of a sucking disc placed behind the propodium ;
ARCHETYPE MOLLUSC. 231
this part is the inesopodium, which thus presents a mo-
dification different from that of the propodium, the one
as well as the other, however, "being admirably suited to
its function. Moreover, the metapodium, or tail, as it is
sometimes called, hears on its surface the hard body-
called operculum, which serves as a lid to close the mouth
of the shell when the animal retreats into that appen-
dage.
In Aplysia, or sea-hare, the epipodium is highly de-
veloped for a special purpose, namely, to assist in loco-
motion. Professor Huxley describes a tropical Aplysia
as flying through the water in precisely the same way as
a Pteropod would do. In Natica, we observe the rneso-
podium modified, to serve as a disc for locomotion by
creeping ; the metapodium bearing the operculum or lid
which closes the mouth of the shell when the animal
takes refuge in it.
Among Bivalves, as they are called, from the form of
the protecting shell, we find numerous modifications of
the neural surface in evident relation to the wants of the
animal. In the oyster, destined to sedentary life, it is
small ; in Solen or razor-fish it is large, constituting the
foot, which the animal employs as an effective means of
burying itself in the loose sand. According to the views
of some, the same part is actually so modified in its form,
and in the nature of its constituent tissues, that it may
be used as an instrument for perforating wood and rock.
\"Y hatever be the form or function of this necessary organ
of the bivalve mollusc, it is supposed to correspond to
the metapodium of the archetype.
Certain Gasteropodous mollusca are, when young, pro-
tected by a shell resembling that of the nautilus in
miniature. At this stage they do not possess the power
of creeping, but swim freely in the water — a provision
232 MODIFICATIONS OF THE ARCHETYPE MOLLUSC.
which secures their wide distribution, and gives rise to
fresh colonies at a distance from the parent. At this
early period of life they are provided with two wing-like
appendages fringed with cilia ; these are employed as
oars, by which they move from place to place. The ap-
pendages in question are believed to correspond to the
anterior part of the cpipodium. This peculiar modifica-
tion is, however, only a temporary arrangement ; a time
arrives when it is no longer needed ; it then disappears,
and the adult animal accomplishes progression on hard
surfaces by means of the foot proper. The ciliated epi-
podium is provided for a temporary purpose, and when
that is accomplished it disappears, to be superseded by
another part.
It is therefore admitted, that all mollusca present
traces of a common plan ; and although in every instance
it may not be possible to indicate with clearness and pre-
cision the special ends of the many modifications of the
archetype, still, arguing from what we do know, it is not
unreasonable to conclude that we have here independent
members in harmony with each other, and conspiring to
promote the wellbeing of the animal in its destined
sphere of life.
CHAPTER VII.
ARTICTJLATA.
SECT. I. HOMOTYPAL RINGS AND APPENDAGES.
We now pass to the Articulate type of the Inverte-
brata, comprehending crabs, barnacles, insects, spiders,
and others.* These agree in one obvious character —
their body consists of a series of similar or homotypal
rings, which present almost endless variety in size, form,
and other particulars, according to the habits of the spe-
cies. The rings are generally, in the higher kinds at
least, of more or less hard texture, giving support to
appendages, and serving as points of attachment to nu-
merous muscles, as well as protecting various important
organs concerned in the function of sensation, motion,
circulation, &c. They present us with examples of a
highly-developed outside covering, technically called exo-
skeleton, the character of which varies as there is neces-
sity in different parts, for variety of motion, for solidity,
or for simple protection.
The endless diversity in form, and the exquisite beauty
of colour and sculpture, exhibited by certain of the Arti-
culata, have rendered them favourite objects of study,
and their history has been in general very thoroughly
* It is not our intention to discuss here all the classes of the Articulate type ; a selec
tion will suffice for our purpose.
234 HOMOTYPAL KINGS
investigated by observers in different countries. The
fertile results which have accrued from such inquiries pre-
sent admirable examples of what may be expected from
the patient labours of ardent naturalists, guided by care-
ful attention to philosophical methods of investigation.
We have stated that all the pieces in the linear series
of which an articulate animal is made up, are homotypal,
that is, constructed on the same plan. This unity of
composition is not necessarily coincident with any law of
number, viewing the Articulata as a whole ; but in the
higher types, at least, the number of similar pieces of
which the body consists is usually uniform.
We may here introduce the general law announced by
M. Aucloin, in 1820, that the similarity or difference be-
tween the segments, the union or the separation of the
pieces of which they consist, the excessive development
of some and the rudimentary conditions of others, occasion
all those differences observed in the entire series of arti-
culated animals. It is well established that a common
type determines the general organization of the animals
in question, and we may now examine the structure of
the typical ring or segment.
Milne Edwards, in his history of the Crustacea, has
demonstrated very clearly the
composition of this part. It
may be described as consisting
of two arches, a superior and an
inferior. The former consists of
four pieces, arranged in pairs on
each side of the middle line.
FlG- 49* The two upper, occupying a po-
sition on each side of this middle line, are called tergal,
* Fig. 49. Plan of ring of Articulate animal, t, tergals ; ep, epiuieialis; s, sternals; ei
episternals.
AND APPENDAGES. 235
because forming the back, (from tergum, back ;) those
on each side are called epimerals, or flank pieces. The
lower arch has similar composition : the middle pieces
are called sternal, because corresponding in position to
the breast-bone (sternum) in Vertebrata ; the lateral
pieces are called episternals. Instead of the technical
terms epimeral and episternal, we may use the terms
upper and lower flanks. In all this we find some resem-
blance to the neural and haemal arches in the vertebrate
segment, with this difference, that the body of the ver-
tebra serves at once as a foundation and line of demar-
cation between the two arches, each of which is complete
and independent. The typical segment in the Articulata
may be compared to a segment of a tunnel, not merely
arched in the roof, but having also a concave floor. A
series of such rings constitutes the external framework of
the animals under discussion, and protects the nervous
centres, which are placed near to the floor, and also the
hcemal organs, which lie beneath the roof, and therefore
differ in their position from that in the Vertebrata.
The division of the body, in crabs and insects, into
three regions — head, thorax, and abdomen, is generally
obvious enough. There may exist difference of opinion
regarding the number of segments or rings entering into
the formation of each of these, and respecting the number
of pieces constituting the typical ring ; but it is generally
admitted as an established truth, that the entire body is
made up of a number of similar pieces.
In Crustacea, (crabs, &c.,) Milne Edwards and others
believe each region to be made up of seven segments,
making, therefore, twenty-one in all. In insects, the
head is supposed to consist of five, the thorax of three,
and the abdomen of eleven.* Erichson, in his Entomo-
* Newport, Art. Insecta, Cyclopedia of Anatomy and Physiology.
236 HOMOTYPAL KINGS
graphien, has demonstrated that the thoracic portion of
the body in crabs, insects, and spiders, is made up of
three segments.* But, as we have said, whatever differ-
ence of opinion exists regarding the entire number in
any one region, or in the whole body, it is universally
admitted that a uniform plan regulates the construction
of the entire framework ; " the different forms of the
body are invariably the result, not of the introduction of
new elements, but of the greater or less extent to which
the primary parts are developed."f
We have seen that in the vertebrata the typical verte-
bra supports appendages ; so the typical ring in the
articulate invertebrata also gives attachment to lateral
appendages. Their form and function vary according to
the part of the body which supports them. They differ
also in different species, and even at various periods of
the life of the same individual, but they all possess cer-
tain common characters.
M. Audoin, long ago, demonstrated that the appen-
dages in question belong either to the upper or lower arch
of each ring of the body, the first constitute the wings
of insects, and the second their legs ; the same applies
to those of crabs and spiders, which, however, want the
upper appendages. They are, therefore, arranged in
pairs on either side of the middle line, and each ring
supports either two or four such appendages. Those of
the inferior arch are the more important, and are of more
universal occurrence than the others.
In Crustacea the complete appendage is constituted by
three distinct portions, which it will be necessary briefly
* In Dana*s Crustacea of the U. 8. Exploring Expedition, there are some peculiar and
Important views as to the organization of the different groups, and the number of ring*
in the different regions (head, &c.,) of the body; as well as the mean normal length of
lings. It is, however, unnecessary for our purpose to discuss the subject.
t Newport, iu Cyclopaedia of Anatomy and Physiology,
AND APPENDAGES.
237
to describe. The first and most essential of these is the
stem, which gives support to the other two ; it is formed
of a number of pieces attached in
linear series. The second part is
called palp, and is generally attached
near the base of the stem. The third
is called by M. Edwards the fouet,
or flabellum ; it also originates from
the stem, but at a point more exter-
nal than the palp. In conclusion,
it may be remarked that attention
to the number of appendages in any
part sometimes affords a good crite-
rion for deciding its composition,
where, owing to adhesion or other
circumstances, the number of rings may be obscured
Modifications or departures from the general plan nay
arise from several causes ; — as from soldering of two or
more of the elementary pieces ; from confused develop-
ment of parts whose presence may be indicated by the
existence of special centres during the process of harden-
ing ; from wasting of one or more of the elements of the
typical segment ; the abortion of certain parts of the
same ; unequal development ; overlapping of neighbour-
ing parts ; disappearance of typical parts ; and, lastly,
from multiplication by repetition of similar parts.f
Fig. 50*
SECT. II. SPECIAL MODIFICATIONS OF RINGS AND APPENDAGES.
Crustacea. — In the higher forms, usually called Deca-
pods, (ten-footed,) from the number of their chief loco-
* Fig. 50. Appendages of Crustacean, showing its essential parts; rt, stem; J>, palp ;
c, flabellum.
t M. Edwards, Annals des Sciences Naturelles, 1851
238 SPECIAL MODIFICATIONS
motive members, we observe three principal modifications
in the general form of the body. First, there is the Bra-
chyura or short-tailed crabs, (as the common crab,) in
which the abdominal part of the body is of small size,
and usually folded beneath the thorax, (so called,) which
part is generally very highly developed. The second
form comprehends the Anomoura, in which the abdomi-
nal portion of the body is soft and defenceless, as in the
hermit crabs. Under the third head are included all
those called Macroura, (long-tailed,) the posterior extre-
mity of the body being well-developed ; the lobster may
be cited as an example. Details regarding the real na-
ture of the departures from the archetype in each of these
three forms are unnecessary for our purpose ; it is enough
to say that in every case, the structure, habits, and in-
stincts of the animals are all in beautiful harmony with
each other.
Where, as in the first of these, the thorax is well
developed, and usually of great strength, the ambulatory
appendages, in five pairs, are generally of large size, and
constitute very efficient organs for progression as well as
other purposes. The great strength of the general frame-
work i3 in admirable harmony with its function as a
supporter of the powerful limbs, and the protector of
important internal organs. But, since the relations of
the segments, and of the appendages which they support,
are so intimate, the special modifications and functions
of each are best studied in conjunction.
The tabular view which we here submit, of some of the
segments and their appendages, will afford an idea of the
deviations from the common plan which occur in different
parts of the body of the same individual, and shew how
each deviation has reference to some peculiar function of
the part. There is an absence of the centralization and
6f rings and appendages. 239
specialization which characterize animals higher in the
scale ; all the segments and their appendages together
constitute the individual, and each performs its respec-
tive function in order 4o contribute to the wellbeing of
the whole. The following table represents the general
arrangement of most of the rings and appendages in one
of the higher Crustacea, a lobster, for example : — ■
Rings.
1
2
3
Appendages.
Eyes.
1st, pair of Antennae,
2d, do. do.
Functions.
Vision.
■ Touch, &c.*
4
Mandibles.
*
5
1st, Maxillae,
6
2d, do.
Capture and division of
7
1st, Feet-jaws,
food, &c.
8
2d, do.
9
3c?, do.
0
11, 12, 13,
14
Limbs,
do.
■ For progression.
Then follow appendages of abdominal rings, varying in use.
The individual is thus made up of a number of organs,
each of which fulfils a special office ; by this division of
labour each most effectively performs its part in the
general economy, and the wellbeing of the whole is
amply provided for.
The typical appendages of the first and second rings
are modified for the purposes of vision, touch, hearing,
&c. ; then follow organs surrounding the mouth, and
which are employed by the animal when food is required :
the ihibellum of the second pair of foot-jaws assists in respi-
ration ; the thoracic appendages are limbs fur locomotion,
and sometimes for prehension ; those of the abdomen are
either for locomotion or respiration, or are concerned in
the function of reproduction. It is to be observed that
* One or both of these are now believed to perform the function of smelling.
240 SPECIAL MODIFICATIONS
each appendage presents special modifications not only
in its general form, but also in the number of the ele-
ments of which it consists, but in every instance the
departure from the typical apj)endage has a decided rela-
tion to its use and the comfort of the animal.
In Crustacea of lower organization, the king-crab, for
example, the appendages of the head and thorax closely
surround the mouth ; they are nearly all of the same
form, and act not only as limbs for motion, but also as
instruments for the capture of the food, and farther, their
bases act as jaws for dividing that food.
The mandibles corresj3ond to the stem of the typical
appendage, strengthened and usually toothed. In Che-
lura terebrans, whose habits of boring render it so
destructive to wooden piles, the jaws present a file-like
surface, admirably fitted to reduce to powder any such
structure.
Generally speaking, it may be observed that the ap-
pendages of the fourth to the ninth segments inclusive
have forms and dimensions varying in harmony with
their uses. In the words of M. Edwards, " they are so
much the shorter and flatter as they are more peculiarly
apportioned to the oral apparatus, a disposition which is
nowhere more conspicuously displayed than among the
short-tailed Decapods, (common crab, for instance,) in
whicli they resemble horny laminee, armed with teeth of
various sizes, and supporting a jointed palp as well as a
flabellum."*
In the thoracic portion of the body, some of the more
anterior appendages or limbs are, in the higher Crustacea, of
large size and peculiar organization, constituting the pin-
cers, which are very formidable instruments for offence and
defence, and are sometimes used for other purposes. One
* Cyclopa;dia of Anatomy, Art. Crustacea.
OF KINGS AND APPENDAGES.
241
of the most striking examples of such modification in
harmony with function, occurs in the large land-crab
(Birgus Latro) of the Keeling Islands. We shall quote
the description given by Mr. Darwin : — " The first pair of
legs end in strong and heavy pincers, the last pair are
fitted with weaker and narrower. The animal tears off
the cocoa-nut husk, fibre by fibre, and always from that
end under which the three eye-holes are situated ; when
this is completed, the crab commences hammering with
its heavy claws on one of the eye-holes till an opening is
made, then, turning round its body by the aid of its pos-
terior and narrow pincers, it extracts the contents — a
curious instance of instinct and adaptation of structure
between two objects so remote from each other as a crab
and a cocoa-nut. The strength of the fore-pincers is
great : an individual was confined in a tin box, the lid
secured with twine, but the crab turned down the edges
and escaped ; it actually punched many small holes quite
through the tin."*
In the species of Portunus of our own seas, the last
joint of some of the thoracic
members is flattened, and the
limb serves as a paddle for
swimming, or is used by the
animal as a means of scut-
tling itself in soft sand.
In many Crustacea, cer-
tain appendages are modified
to serve as apparatus for re-
spiration, acting, in fact, as fig. en
branchiae or gills. Those called Branchiopods (gill-
* Darwin, Journal of a Naturalist, p. 463.
t Fio. 51. Transformation of appendage of abdomen in a Branchiopodous (gill-footed)
Crustacean, b, fiabellum ; c, palp, which act as respiratory organs.
242
SPECIAL MODIFICATIONS
footed) receive their name from this peculiarity ; the
whole of the thoracic appendages are in the form of
lamellas, and the parts corresponding to palp and fla-
bellum are membranous vesicles highly vascular, and
fitted to expose the circulating fluid to the action of the
air contained -in the surrounding water.
Tn certain others, the Amphipocls, for example, localiza-
tion of function
is more com-
plete, the fiabel-
lum alone acting-
as a gill. In
those called Iso-
pods, the mem-
bers for loco-
motion have no
other function
superadded, re-
spiration being performed by the first five pairs of ab-
dominal appendages, which appear to have no other use.
In the lobster, cray-fish and others, in which the
hinder part of the body is well developed, certain of its
elements are very specially fitted for the progression of
the animal through the water. The last ring, and the
appendages of the one which precedes it, are specially
modified to form their powerful tail fin.
In the soft-tailed hermit crabs, which protect their
tender and defenceless abdomen in empty spiral shells
of Mollusca, certain appendages are modified to act as
hooks by which the animal holds fast to the inside of its
borrowed habitation ; and it is a curious circumstance,
that some of these hooks are wanting on one side, since
Fig. 52*
* Fig. 52. Appendage of Amphipodous Crustacean, the flabellum, c, alone serving
as a gill for respiration.
OF RINGS AND APPENDAGES. 243
they /rould be useless or even an encumbrance to the
animal, owing to the curve of its body corresponding to
that of the shell in which it lives. The instincts of the
hermit crab lead it to seek in an empty shell that pro-
tection which is wanting in the texture of its own body.
The means by which it holds fast are also admirably
fitted by form and position, to the exigency of the case.
Not a few of the Crustacea are parasites, that is, they
attach themselves to other animals, and feed on their
juices ; those called fish-lice are examples. Such habits
require special peculiarities of organization, and we are
constrained to admire the wisdom which foresaw and
provided for all the necessities of these singular beings.
The mouth apparatus in some is fitted at once lor pier-
cing and sucking the juices of the foster-parent ; and cer-
tain of the appendages in other species, corresponding to
those already alluded to under the name of foot-jaws, are
constructed in such a way that they enable the little
animal to keep fast hold of its foster-parent.
In the curious Lerneadee, whose grotesque forms have
puzzled not a few observers, the young are furnished with
a well-developed eye, and are provided with two large
pairs of appendages, which serve as oars. Their peculiar
instincts lead them to fasten themselves to various fishes,
some selecting one part of the fish, others a different part.
Some after they become fixed, the eye, no longer of any
use, is lost, the oar-like appendages either disappear, or
undergo a change of form suited to the new mode of life ;
in a word, there are several independent successional
arrangements concurring to one end. Certain parts are
necessary to the existence and comfort of the animal,
and such are provided, and everything is in conformity
with the position which it occupies in the economy of
nature.
244
SPECIAL MODIFICATIONS
Barnacles. — These remarkable animals, in some one
or other of their forms, are doubtless familiar to our
readers. Many of them are attached, by more or less
flexible stalks, to sea-weeds, to drift-wood, even to quills
shed by sea-birds, or they adhere in countless multitudes
to the bottoms of ships which enter our harbours from
some warmer region ; so abundant are they, in fact, as
sometimes to impede the motion of the vessel in the
water. Other kinds contribute to the formation of that
white line which marks the limit of high-water on our
rocky shores, or give a continuous covering to the exposed
parts of marine piles or stakes of salmon-nets. Others
invariably attach themselves to corals ; not a few find a
suitable dwelling-place in the thick skin of whales, and
Fig. 53*
certain others in the shell of the sea-turtle, and some bury
themselves in sponges. All these curious animals are
* Fig. 53. a, A Stomapod Crustacean of the genus Leucifer. The abdominal portion
is not shaded. The shaded part corresponds with the next, b.
b, Cirriped, or Barnacle— a mature individual. All the parts correspond to shaded
portion of a; the eyes and antenna;, which are distinct in early life, are also represented
here, for the sake of comparison.
OF KINGS AND APPENDAGES. 245
constructed on the same general plan as the Crustacea we
have been examining, and are, in fact, so nearly allied, that
naturalists justly include them in that class. (Fig. 53.)
The archetype has undergone remarkable transforma-
tions in the barnacles, in order to fit them — and how
admirably are they fitted — to that particular part which
the Creator has assigned them in the economy of na-
ture.
In the earlier periods of their life, barnacles are free ;
that is, unattached, are possessed of efficient locomotive
members, and furnished with organs of vision ; in this
condition they very much resemble some of the simpler
forms of Crustacea. Peculiar instincts lead to the choice
of a proper habitat, whether a floating body, or a rock,
a sponge, a whale, or a turtle ; how admirable, therefore,
the harmony between the structure and the instinct !
The voluntary roving animal becomes fixed to some
object, and, after various transformations of its organs,
the adult state is finally assumed, and the change of
form is commensurate with that of its mode of life. The
fixed state of the full-grown animal renders several con-
ditions necessary to its existence and comfort. Having
no power of movement from one place to another, the
barnacle is incapable of voluntarily avoiding injury from
without. The animals require means of attachment, a
shell for protection, and provision for the supply of
their wants. All these points have been attended to in
their structure, and there is remarkable concurrence of
arrangements tending to the well-being of the entire or-
ganism.
The masterly researches of Mr. Darwin, forming two
volumes recently published by the Kay Society, have
fully elucidated the remarkable modifications of the
Crustacean type met with in the animals under discus-
246
SPECIAL MODIFICATIONS
sion. Comparison of the following tabic with that already
given, will show the relation between a barnacle and a
crab : —
Rings. Appendages.
No. 1, Eyes, ) .. ,. _,_. _,_ .
J- quite distinct in early stages.
2 & 3, Antenna?, J l J b
4, Mandibles.
5 & 6, Maxilla;.
7 & 8, Generally coalesce or disappear.
j, 10, 11, 12, 13, 14, Six pairs of limbs.
_ ._ ..* l form three small abdominal segments;
( the last four are wanting.
The appendages of the third ring, or the second pair of
antennae, are the primary means of attachment, the union
being subsequently consummated
by a cementing material, which at
first issues from these appendages,
and finally also, in some, through
special openings in the head. Sucl^
then, is the simple means by which
the attachment of the barnacle is
provided for. In connection with
this part of their history, allusion
may be made to the habits of a
species not uncommon on some of
our coasts. In Lepas fascicularis,
the cement is very copiously given
out, and forms a vesicular ball,
which acts as a float. Mr. Darwin
Btates that sometimes several individuals have their
stalks imbedded in the same ball, which swims like
a cork on the water. As this species grows into a
Fio. 54.*
* Fi«. 54. Lcpas fascicularis. -with its stalk (together with three others, the stalks of
which are alone seen) imbedded in a vesicular ball ^constituting a float) of their otd
formation, of which a slice has been cut off to shew the internal structure.
OF RINGS AND APPENDAGES. 247
bulky animal, we here see a beautiful and unique con-
trivance in the cement formed into a vesicular membra-
neous mass, serving as a buoy to float the individuals,
which, when young and light, were supported on
the small objects to which they originally had been
cemented in the usual manner. We have seen a cluster
composed of at least a dozen large specimens, any one of
which, without the float, would have been sufficient to
sink the small quill-feather of a sea-gull to which they
were attached. It will be remembered that the posi-
tion and production of this singular contrivance depend
on modifications relating to certain appendages of the
body.
As regards means of protection, we may quote Mr.
Darwin, who states, "In the mature animal, the whole
external covering, whether shell and operculum, or capi-
tulum and stalk, is formed of the third segment of the
head."* It consists of distinct plates, which overlap each
other, and are capable of various movements, in which
respect it differs from that of all crustaceans, and farther,
is never moulted or cast off, as is the case in them.
But the animal requires, also, means for procuring
food ; this is provided for, in all common barnacles, by
a special modification of the thoracic limbs, which form
six pairs, and are admirably suited to their intended use.
(See Fig. 53, b.) Each is two-oared and many-jointed ;
" they have a peculiar character, different from the limbs
of other crustaceans, not being natatory, ambulatory, nor
branchial, but ' captorial/ or fitted for sweeping the
water, and thus catching prey/'f Mr. Hancock describes
these appendages as acting like a prehensile net. Is it
possible to conceive any better example of parts con-
structed according to a general model, and yet harmo-
* Darwin, loc. cit. vol. ii., p. 13. + Ibid., vol. il, p. 14,
248 SPECIAL MODIFICATIONS
niously combined and modified in distinct relation to a
special purpose, than that found in the barnacle ? As-
suredly the lately-developed principle of homology does
not set aside, but corroborates the old-established prin-
ciple of final cause ; and it appears to us that the more
intimate our acquaintance with the one, so much clearer
will be our idea and appreciation of .the other.
Insects — The busy bee, that master architect and
builder of its class ; the industrious ants, from some of
which man might derive useful lessons in social economy,
division of labour, and persevering toil ; the locusts, those
rovers and depredators, the Goths and Vandals of the
winged articulata ; the painted butterflies, sipping the
nectar which Flora provides so bountifully ; the mailed
beetles, the athletes of the insect world — notable as
swimmers and divers, as sappers and miners, indeed, as
adepts in various departments of nature's economy too
numerous to be mentioned here ; — all these now invite
our attention. The field is so vast that we can only
glance at a few cases in which we observe modifications
of the archetype, obviously concurring to serve useful
ends in the economy of the animal.
Whatever difference of opinion may exist in regard to
the number of the segments entering into the formation
of the body of the perfect insect, the best authorities are
agreed that the different pieces are homotypes of each
other, and that all modifications and departures less or
greater from a common model. We are now to shew
that these modifications are intended to serve an end
which is more or less obvious. The varied forms of the
whole body, in different insects, depend upon the relative
development of the parts of each segment and appendage,
and the diversities are invariably in direct harmony with
the peculiar function to be performed.
OF RINGS AND APPENDAGES 249
The reader is doubtless familiar with the transforma-
tions, greater or less, through which insects pass before
reaching maturity. How different is the general ap-
pearance of the caterpillar from that of the winged
butterfly — the one incapable of flight, and feeding
upon the solid parts of vegetables, the other possessed
of powerful wings, and having extensive and rapid means
of aerial progression, and feeding on the sweet juices of
flowers ! Both possess the same number of true appen-
dages for walking, namely, three pairs attached to the
segments of the thorax ; those in the caterpillar, or larva,
are nearly of the same size and form. But many larvee,
as requiring efficient means of locomotion on a hard sur-
face, are furnished with additional limbs, usually called
false, because they are not appendages of the archetype,
but only prolongations of the external covering of the
body, and are attached to the abdomen. Without enter-
ing into details respecting the very numerous modifica-
tions of these false appendages, it may be sufficient to state
that whatever their number or form, they are invariably
so constructed as to answer every purpose for which they
may be wanted in the economy of the animal.
It may further be observed that many larvte are des-
titute of feet, and yet possess the power of locomotion.
And here we sec a beautiful compensatory arrangement
in the form of minute hooks, which are prolongations of
the external covering of the body, the position, number,
and forms of which are wonderfully adapted to the pecu-
liar habits of the individual. We may conclude this
part of the subject by quoting a passage from Mr. New-
port:*— " In apodal larvas, endowed with powers of locomo-
tion, the place of the true organs of progression is supplied
by peculiar developments of the cuticular covering of the
* Cyclopaedia of Anatomy and Physiology, Art. Inseota.
11*
250 SPECIAL MODIFICATIONS
body, analogous to the scales on the bodies of Ophidiam
reptiles, and these are employed by the larvce in all their
progressive movements in the same manner as the scales
on the body of the snake. But in those apodal (footless)
larvre, which remain in the same locality until they have
passed through all their changes, as the larvas of the bee
and wasp, these developments of the cuticular surface do
not exist, but the body is perfectly smooth."
If such remarkable conformity exists between the
habits of the immature animal and the development of
certain temporary organs with which it is furnished, we
may be prepared to expect harmonious adaptations of
the archetype all conducing to the existence and comfort
of the perfect insect, suited to its instincts and fitting it
to the position which it is to occupy, in earth, air, or
water. The usual elongated body of the grovelling
larva in general presents evident uniformity in the devel-
opment of the segments as well as of the true appen-
dages when present, in other words, there is a close
approach to the archetype. The new sphere which it is
subsequently designed to occupy, demands corresponding
modifications in the form of the whole body, and in that
of the segments and appendages.
In the perfect insect, division of the body into three
regions, head, thorax, and abdomen, is generally obvious.
Each of these consists of parts adapted to certain ends,
and all concurring to the well-being of the entire organ-
ism. All of them present entire fitness for their respect-
ive functions ; those of the head support certain sensatory
organs and appendages for capture, retention, and reduc-
tion of the food ; those of the thorax afford attachment to
wings and limbs ; the abdominal segments protect certain
viscera, and serve other purposes besides.
The differences to be observed in the hardness of the
OF RINGS AND APPENDAGES. 251
framework are remarkably adapted to the uses of the part
Where close union and density are wanted for strength,
there we find them-; in the head this is specially evident ;
mobility is sacrificed for firmness precisely where such is
necessary. The consistence of the head segments is, as
a general rule, greater than that of any other region of
the body. The head is the part of all others most ex-
posed during progression, whether in air, earth, or water ;
besides, it supports mandibular organs, whose function
frequently is to act upon very hard materials and fit them
for digestion. Owing, in fact, to the close union of the
elements of the typical rings forming the head, there has
been more difference of opinion regarding the number of
its segments than those of any other part of the body.
The muscles of the insect are inserted on the internal
surface of the framework, and we might naturally expect
a relation between the development of the two. Where
strong organs of mastication are needed, the segments of
the head 'are large, being directly proportional to the
power which the mandibular apparatus is fitted to exer-
cise. Mr. Newport remarks, " we invariably find that in
those insects in which the mandibles are large, the whole
head is either short and wide, or its posterior portions, to
which the muscles of the mandibles are attached, greatly
exceed those of the anterior."*
The great extent of surface occupied by the organs of
vision in many insects, has an influence also on the gene-
ral development of the whole head and of its elements.
The rapacious dragon-flics, for example, hunt solely by
sight, and their eyes occupy almost two-thirds of the sur-
face of the head, and we observe corresponding modifica-
tions in the segments. It is unnecessary to enter into
minute details regarding the variously modified appen-
*Cyc1opxdia of Anatomy and Physiology, Art. Tnsecta.
252
SPECIAL MODIFICATIONS
dages of the different segments of the head ; it will "be
sufficient to indicate some of the more obvious adapta-
tions of the elements to their respective functions. The
wide dissemination of insect life implies considerable
range in the instincts and means of existence. The
predatory habits of some constitute them the carnivora
of their class, and others are not less fitted — than rodent
mammals — to gnaw hard vegetable matters. The in-
stinct which leads some to sip the sweet fluids of flowers,
or stimulates others to tap the integuments of animals
or of plants for the purpose of feeding on their juices,
equally require adaptation of the mouth to such purposes.
But whatever the end to be accomplished, and however
great the apparent difference of the organs which mini-
ster to the subsistence of the insect, it was long ago
demonstrated by Savigny that in every case the parts are
fundamentally identical, though varied to suit a purpose.
The study of the mouth-organs in insects has occupied
the attention of numerous observers,
and the results of such researches
have shewn how admirably each piece
is fitted for its function, and at the
same time accommodated to act in
harmony with every other.
In the great water-beetle (Hydrous
piceus) the mandibles are two strong,
arched and toothed jaws moving hori-
zontally in opposition to each other ;
this species is omnivorous. In the
truly carnivorous forms, as the brilliantly-coloured and
active tiger-beetles, the mandibles are acutely pointed,
strongly toothed, and crossing each other like the blades
* Fig. 55. Mandible of a large water-beetle, (Hydrous piceus.) There are two such
which act in opposition to each other, like the blades of scissors. The opposed edge*
are hard and toothed.
Fig. 55*
OF RINGS AND APPENDAGES.
253
of scissors, and are thus admirably fitted for dividing the
prey. Those of Melolontha (the cockchafer) have short
blunt teeth fitted to bruise vegetable matter ; in Cetonia,
which feeds on the pollen of plants, the edges of the man-
dibles are soft and flexible. The mandibles of the locust
are in front so constituted as to form cutting organs, and
behind act as grinders of the vegetable food.
The maxillae, or lesser jaws, are organs of prehension
and retention chiefly, but may aid also in
mastication. Like the organs just de-
scribed, they present differences in form
and texture in direct consistency with the
habits of the insect.
Among Hymenoptera, comprehending
bees, wasps, &c, the mandibles present
very considerable difference in form ; " in
the Vespida), (wasps,) which gather the
materials for their nests by rasping off
little packets of fibres from decaying wood,
they are broad, triangular, and armed along
their edges with strong teeth ; and such is
also their structure in Anthidium manicatum, which
scrapes off the down from the woolly stems and leaves of
plants for the same purpose ; while in the hive-bee, which
employs them in moulding the soft wax in the construc-
tion of the combs, they are shaped at the apex like a
spoon, without indentations ; their form in each instance-
being thus distinctly conformable to the habits of the
insects."f
The highly-developed instincts of bees, which lead to
Fig. 56.*
*Fig. 56.. Maxillr, or smaller jaws of the Hydrous. Thoy act in pairs, but as tholr
function is to hold the food and convey it to the back part if the mouth, they are not so
strong as the mandibles, which divide, and bruise the food ; they, however, hare a gene-
ral resemblance in shape.
t Newport, Cyclopaedia of Anatomy and Thysiology, p. 898.
254 SPECIAL MODIFICATIONS
the formation of very ingeniously constructed nests, imply
the necessity of tools for the work ; these are furnished
by the mandibles, while the maxilhe and another cranial
element termed the labium, are principally concerned in
collecting the food ; the former are elongated, and with
the latter beneath, together constitute a tube by means
of which the honey of flowers is conveyed to the mouth.
But wc must pass on to consider arrangements suitable
to the habits of suctorial insects properly so called, and
here also, while the general plan is evidently adhered to,
the modifications are in strict conformity with the wants
of the animal, and all concurring to a common end.
Hitherto we have seen that the mandibular appen-
dages have occupied either the chief, or at least the pro-
minent place in the operation of feeding ; in the Haus-
tellate insects (those furnished with a proboscis) the
mandibles no longer perform the same important offices,
while the maxillae and -the labium now assume greater
prominence and importance in the economy of the insect.
Every one must be familiar with the habits of moths
and butterflies "hovering around those opening flowers,"
and closer inspection would reveal that the insects carry
with them an apparatus admirably fitted to reach the
sweet juices in parts of the plant, into which the body of
the animals could not possibly find access. The short
mandibles of the voracious vegetable-feeding caterpillar,
though admirably fitted to that stage of life, would be
utterly useless in the new sphere which it occupies, when,
issuing from the mummy-like case of the pupa, it emerges
as a winged imago, endowed with new instincts and new
faculties. The perfect insect carries with it an instrument
admirably fitted for reaching and drawing up the nectar
of flowers. The mandibles are no longer capable of
supplying the wants of the animal, as the sweet fluid
OF RINGS AND APPENDAGES. 255
on which it feeds requires no mastication ; but an organ
is needed to suck it up, and of sufficient length to reach
the parts of the plant where it abounds ; such an organ
is supplied.
It would be difficult to select, in the entire range of
the animal kingdom, such a remarkable example of
special modification of typical organs, as that presented
to us in the proboscis of the butterfly. The problem
is to convert the maxilla3 (which in some insects we
have seen to be organs for prehension and mastica-
tion) into organs adapted to the function they have to
perform in the moth or butterfly ; for, as we have just
said, the portable flexible tube in these animals really
corresponds to the maxillae of a beetle. Sweet juices
abound in flowers, access to the bottom of every floral tube
would be impossible to insects having the large prominent
eyes of those under discussion, and so a peculiar contriv-
ance is necessary under the circumstances — that contriv-
ance is simple, yet efficient for every purpose required.
The structure of the mouth-apparatus in the Lepidop-
tera has been so fully illustrated in other works, that a
summary may be sufficient here.
The appendages called maxillse constitute the sucking
apparatus. In the words of Mr. Newport, " each maxilla
is composed of an immense number of short, transverse,
muscular rings. It is convex on its outer surface, but
concave on its inner, and the tube is formed by the ap-
proximation of the two organs."* But something more
is necessary. By what means are the two opposed chan-
nels to be kept in sufficiently close contact so as to form
a perfect tube ? Reaumur, Kirby, and others, have de-
scribed numerous minute and delicate hooks or teeth,
(for they assume varied forms in different species,) which
* Newport, loc. cit., p. 90.
256
SPECIAL MODIFICATIONS
are arranged in close series along the inner margin of
each maxilla, and the teeth of the one set lock between
the teeth of the other. The animal is now furnished
Fig. 57.*
Fig. 58.+
with a means of searching every crevice of a flower for
the tempting juice which is formed there. But some-
thing more is necessary. Not a few Lepidoptera feed
upon wing, and the act of feeding is very quickly per-
formed ; in the twinkling of an eye the tube is inserted,
and the flower is robbed of its sweets. The act of suc-
tion, by producing a vacuum, which enables the infant
to procure nourishment from the breast, is also brought
into play in order that the fluid may rise in the butter-
fly's proboscis. The peculiar air-tubes which traverse
the bodies of insects, for the purpose of respiration, are
abundantly distributed throughout the maxilla? in the head,
and over the gullet and alimentary canal. Experiments
made by Mr. Newport led to the conclusion that the
* Fig. 57. Head of Noctua libatrix ; m, mandibles, small ; ma, the two maxillae,
large, and forming the proboscis or sucking apparatus,
t Fig. 58. A single maxilla of the same.
OF RINGS AND APPENDAGES. 257
insect first makes a strong effort to expel the air, and
just when the proboscis comes in contact with the fluid, a
powerful inspiratory effort is made, which occasions dila-
tation of the tube, producing a vacuum, and thus causing
the liquid food to rise. There is still, however, another
arrangement necessary in this simple hut efficient appa-
ratus. It must be long enough to reach to the very
bottom of the floral shaft whence the food is to be drawn,
but a long and flexible tube would be liable to injury,
and also inconvenient during progression on the ground
or in the air ; it must, therefore, be portable ; and here
another modification comes in to provide for the comfort
of the insect. The two maxillas, conjoined in the way
we have described, are, when at rest, coiled like the spring
of a watch, but can be extended with ease and surprising
rapidity as required. (See Fig. 58.)
The mandibles — which, as we have seen, are so highly
developed in some insects of prey, and are, on the con-
trary, so useless in the butterfly — assume a new aspect
and function in the blood-sucking Tabanidae. In the
typical genus of that family, they are long and lancet-
shaped ; and Mr. Newport describes them as acting not
from side to side, but with a horizontal movement from
behind forwards, cutting also vertically with a sweeping
stroke, like the lancets of a cupping instrument. We may
add, that the bite of the gnat is effected in the same way.
We may now proceed to examine the modifications pre-
sented by the next region of the body — the thorax, namely.
The three different segments which constitute this part
will also afford means of illustrating the argument.
The first or anterior ring (prothorax) supports the
first pair of legs ; the second or middle portion (meso-
thorax) gives attachment to the first pair of wings and
the second pair of legs ; the third or posterior (meta-
258 SPECIAL MODIFICATIONS
thorax) bears the second pair of wings and third pair of
legs. All these segments and their corresponding ap-
pendages ju-esent notable differences, according to their
relative importance in the same or in different insects.
We have now to examine organs concerned principally
in that faculty which is so eminently characteristic of
the insect tribes. We have seen the very admirable pro-
vision made for enabling each to secure its peculiar food ;
no less remarkable are the modifications of organs in
co-operation for the function of locomotion, so that the
necessary food may be sought after.
• The alar appendages, or wings, are viewed by some
as not constituting a necessary part of the archetype, but
organs superadded, and serving both for flight and respir-
ation.* Their relative development in different species
is accompanied with co-ordinate changes in the segments
which support them, and the other appendages which form
the legs of the insect. Entomologists in treating of this part
of the body, cannot avoid alluding to and enlarging upon
the evident relation between the habits of the insect and
the modifications of the thoracic segments and their ele-
ments, and we cannot do better than introduce an abstract
of Mr. Newport's remarks.f There is wonderful modifi-
cation in shape and variety, in size and position, of the
thoracic elements, in order that the body of the insect
may be in conformity with its mode of life. In the great
water-beetle, (Hydrous piceus,) which burrows deeply in
the mud of stagnant waters, and rises also to the surface
to bask in the sun, the form of the lower surface of the
entire thorax is admirably adapted to its habits. The
sternal elements of the meso-thorax and the meta-thorax
* The wings may, however, he considered as homologous with the upper appendage*
of Annelida; — sea-worms are examples,
f Cyclopaedia of Anatomy and Physiology, Art. Insecta, p. 917.
OF KINGS AND APPENDAGES. 259
are strongly keeled and firmly united together, enabling
the insect to float securely. In others nearly allied, but
of more active aquatic habits, swimming with ease and
quickness, and capable of rapidly turning and following
all the movements of their living prey, there is but a
slight keel below, and the edges of the body are sharp, so
as to oppose little resistance to the water. In beetles
there is always a beautiful relation between the general
structure of the thorax and the habits of the insects,
whether in walking, flying, or in swimming. In those
which pass great part of their lives on the ground, run-
ning or walking, the middle and posterior segments of the
thorax are often firmly joined together, in order to give
greater strength to the whole body. This occurs in all
beetles which require great muscular effort during flight,
and in those accustomed to laborious efforts in tearing,
in burrowing, or in running.
But, without enlarging on this subject, it may be
observed that the size and strength of each segment of
the thorax are in direct proportion to that of the append-
ages which it supports, and the whole structure of rings
and appendages present admirable conformity to the
mode of life. For example, when, as in bees, moths, and
the common fly, the anterior pair of wings are the chief
locomotive organs, the meso-thorax or middle segment is
highly developed, and there is corresponding decrease in
the other two.
The proper appendages of the thorax may now be ex-
amined, and in them we find notable correlation between
the habits of the insect and the modifications of the
parts. Here there is a wide field illustrative of the ar-
gument ; but since this subject has been already so fully
discussed in different treatises on natural theology, it will
be unnecessary to do more than refer to a few examples.
260 SPECIAL MODIFICATIONS
The legs are the proper organs employed in terrestrial
locomotion, and for other purposes besides. As already
stated, there are three j)airs of such appendages attached
to the corresponding segments of the thorax.
We have seen that in the vertebrata the limbs or di-
verging appendages of certain parts of the model frame-
work are variously and suitably modified, according as
they are intended for grasping, walkings swimming, or
flying. The same law of consistency between form and
function prevails among insects, and as in the higher
animals, unity and diversity are singularly combined, the
same is true among the winged Articulata ; in the words
of Professor Ilymer Jones — -"Nothing is, perhaps, better
calculated to excite the admiration of the student of ani-
mated nature, than the amazing results obtained by the
slightest deviations from a common type of organization.
The limbs used in swimming exhibit the same parts, the
same number of joints, and almost the same shape, as those
employed for creeping, climbing, leaping, and numerous
other purposes ; yet how different is the function assigned
to them !"* The predatory tiger-beetles are swift of
foot — freedom of motion and lightness of the organs are
necessary accompaniments, and such is the character of
their thoracic appendages ; it is the same in every in-
stance where the habits are similar. In those which swim
and dive, as the water-beetles, &c, length of lever-power,
breadth of surface, and strength of the parts, ail are neces-
sary— and such we find to be provided in their limbs.
They are not, however, all of equal length, 'nor do all act
equally in aquatic progression. The posterior pair, as
regards position and form, are the chief propellers of the
insect, they are flat like the end of a paddle, and the ex-
tent of surface presented to the water is very much in-
* The Animal Kingdom, p. 245.
OF RINGS AND APPENDAGES. 261
creased by a fringe of hairs, which do not materially
add to the weight of the whole limb. This admirable
contrivance servos another purpose, viz., what is called
feathering the oar, when a new position is necessary for
a fresh impulse ; for in the forward stroke of the limb the
hairs are of such nature and so arranged, that they
change their position and accomplish the object in ques-
tion. Limbs simply intended for walking are usually
equally developed in all respects. Surfaces intended to
act as sucking discs by the pressure of the air, are by no
means uncommon, as in certain water-beetles. In some
instances, flat cushions on the limbs, giving out a clammy
secretion, are provided in order to enable the animal to
climb smooth perpendicular surfaces, or hang with its
body lowest from the ceiling ; such is now generally be-
lieved to be the arrangement in the house-fly. Mr. New-
port remarks, " those insects which support themselves
upon the surface of water, as the common gnat, have the
under surface of each tarsus covered with rows of fine
hairs, which repel the water and support the insect upon
the surface. If the under part of the tarsi be wetted with
spirits of wine, the insect can no longer. support itself
upon the surface, but immediately sinks down."
The powers of the most accomplished vaulter, aided
by mechanical adjuncts, are insignificant in comparison
with those possessed by not a few insects. For the
accomplishment of such mode of progression, we find cor-
responding modifications of the posterior pair of legs,
which are chiefly concerned in this kind of function.
The large and strong coxa or first piece, is received into
a deep depression of the supporting arch ; the piece called
thigh is of great length, and very greatly enlarged in
transverse diameter, so as to furnish attachment to the
powerful internal muscles. The sudden unbending of
262 SPECIAL MODIFICATIONS
the strong limb enables the animal to accomplish its
purpose. An additional arrangement is alluded to by
entomologists as being provided in such cases ; the lower
surface of the tarsus is covered with elastic cushions,
which are supposed to assist in the first effort, and finally
to act in breaking the fall when the insect alights. The
flea, turnip-fly, grasshoppers, &c, present examples of
such limbs.
In the mole-cricket the fore-limbs are used in tunnel-
ling, and admirably suited they are for such purpose,
and the corresponding part of the thorax is of commen-
surate strength. The basal joint of the limb, called
coxa, is of unusual size. The thigh is joined to both
coxa and trochanter — an arrangement which adds ma-
terially to its strength. The succeeding portion, the tibia
or leg, usually so called, is the instrument by which the
soil is penetrated and thrust aside ; it is short and broad,
the outer surface of it also is furnished with several strong,
curved projections, the whole presenting a strong and
broad surface, and therefore becoming an efficient instru-
ment by which the animal burrows in the soil.
In conclusion, it may be observed that the last joint
of the foot in insects is usually furnished with a pair of
strong hooks, which afford important aid in climbing or
clinging to rough surfaces.
In a word, whatever the peculiar habit of the insect,
the elements of the limb are variously modified to minis-
ter to its existence and comfort.
We pass on to examine the last part of the body, and
in it, the abdomen namely, we shall find modifications
of the model not less instructive than those already
brought forward.
Some difference of opinion exists respecting the exact
number of segments entering into the formation of the
OF RINGS AND APPENDAGES. 263
third or abdominal region of the insect ; whatever may
be the normal number, it is nevertheless admitted
that all are homotypes, and each fitted to its respective
function.
Generally speaking, the appendicular elements are
wanting, or, for the most part, of very secondary import-
ance, in the abdomen. This part of the body protects a
large proportion of the organs concerned in nutrition and
reproduction, and, as the space occupied by these is
liable to vary, we generally find considerable capacity of
expansion in the segments of which it consists, and, in-
deed, throughout the whole of this region.
But abdominal appendages are not always wanting,
and sometimes they are of the utmost importance in the
economy of the insect ; they usually belong to some of the
terminal rings. There is an order of insects denomi-
nated Hymenoptera, among which we meet with highly
developed instincts, leading to the performance of various
acts, which could not be accomplished without some cor-
responding adaptations in the frame ; all of these are pro-
vided and are exactly suited to the instincts and to each
other. The saw-fly, the gall-fly, the inchneumon-fly, and
others, in the larva condition, feed upon different parts of
plants, or on the internal organs of other insects. The
female deposits her eggs in suitable localities by "means
of an instrument, the ovipositor, fitted specially for that
purpose. Others are provided with formidable weapons
of defence and offence, in the form of a sting. But what-
ever be the function of the instruments in question, they
are invariably modifications of the same abdominal ele-
ments, and in every instance suited to their end.
It is among Hymenopterous insects that we find the
most perfect forms of an egg-depositing instrument ; and
as the localities in which the eggs are placed differ, the
264 SPECIAL MODIFICATIONS.
modifications of the instruments are of commensurate
import. The leaf-flies, the gall-insects, the saw-flies, and
ichneumons, all present instruments varying in length
and strength, according to the substance which each is
intended to penetrate. Those of the leaf and gall-insects
are just sufficient to allow them to penetrate vegetable
tissues : neither is there any great force requisite to enable
the ichneumon female to deposit her ova in the bodies of
other insects, or in the cocoons of spiders, or in the eggs
of butterflies. The saw-flies, which penetrate hard wood
for a similar purpose, are provided with an apparatus of
great power and admirable construction. In every special
case there is some remarkable harmonious adaptation, so
that by inspection of the apparatus we can ascertain the
way in which the eggs are deposited. The elements con-
cerned in boring are placed in pairs, and furnished with
teeth on the edge and sides, the former serving as a saw,
the latter as a rasp. This delicate instrument requires
support when acting, as well as protection ; and accord-
ingly these desiderata are provided, and the sword does
not more accurately fit the scabbard than are the respec-
tive parts of the ovipositor suited to each other, and to the
habits of the insects.
Certain elements and appendages of the terminal part
of the abdomen are transformed into a saw or file, or both,
as the case may be, and others are fitted to give them
strength and protection.* It matters not what the size
of the insect, whether the comjiaratively large Sirex or
the very minute Ichneumon ovulorum,f the general plan
is the same, but in every instance presents some peculi-
arity adapted to the nidus selected by the species.
* Lacaze Duthiers, on Genital Armature of Insects, Annales des Sciences Naturelles,
1849, 1S52.
t This tiny insect deposits several ova in a single egg of a butterfly, the contents of
ivhicb afford sufficient food, as well as protection, to all the young which are produced.
OF RINGS AND APPENDAGES. 265
The formidable sting of the bee and of the wasp are
examples of other modifications presenting no less beau-
tiful harmony between organ and function. Generally
speaking, the appendages of the abdominal segments are
absent, or if present, very rudimentary, because not re-
quired in the economy of the insect. What we have
stated respecting the ovipositor and sting, affords proof
that when certain appendages are necessary they are
provided.
We find them, however, in other cases, furnished for a
different purpose. The insects called skip-tails present
remarkable examples of this. In the genus Lepisma,
there is a pair of appendages attached to eacb abdominal
segment. In Podura, and others, the singular tail-like
organ consists of an elastic stem ending in two branches,
like a fork and its handle. During repose this instrument
is bent beneath the insect, and is lodged in a groove' ;
when suddenly straightened the animal is thus enabled to
spring a considerable distance. The handle of this fork-
like organ is believed to represent the sternal or lower
part of the abdominal segment, the two prongs are stated
to be the homologues of the lateral appendages.
We may finally, and very briefly, allude to a remark-
able transformation of abdominable appendages in another
class of the jointed invertebrata.
The web of the spider is constructed of delicate
threads, which are given out by parts called spinnerets ;
these are organs consisting of two or more joints. The
end of the spinneret is pierced with, a great number of
small holes, each of which gives out a drop of fluid
which hardens in the air. The minute threads of each
organ are joined to form one, and those of all the spin-
nerets again unite to form the apparently simple thread of
the spider, which is therefore in reality complex. There
12
266 MODIFICATIONS OF RINGS AND APPENDAGES.
appears to be no doubt that 'the organs which produce
the spider's thread are really abdominal appendages, thus
singularly modified for the animal's convenience. They
are composed of several joints, as limbs are, and in some
species one pair of them- — not being perforated nor fur-
nished with an organ to produce the thread, and there-
fore apparently not needed — are nevertheless of interest
to the zoologist, as indicating the real nature of the true
Bpinnerets.
CHAPTER VIII.
RADIATA.
SECT. I. — TTPIOAIj FORMS OF RADIATA
The Radiate type of animal structure, as the name
indicates, is characterized by a tendency to repetition of
parts round a centre. This division of the animal king-
dom comprehends, on the one hand, the minute and soft
hydra of our fresh waters, and, on the other, the hard and
formidably-armed urchins of our seas.
At one time, many of the radiates were supposed to
belong to the vegetable kingdom ; more accurate obser-
vation has resolved the doubts respecting their nature,
and demonstrated that they belong to the animal king-
dom. It may be added however, that still more recent
discoveries have shown that, in the mode of reproduction
by buds and ova, they present a remarkable parallelism
to plants. And here we see evidence that certain ani-
mals and plants have so much of unity of plan, as to
shew that they have been constructed by the same
Architect.
Our aim is to shew that, while there is adherence to a
Radiate plan, there are departures from it on the one
side and on the other — deviations which have reference
to some end in the economy of the animal. We meet
with difficulties in this as in other departments, but we
268 TYPICAL FOKMS
doubt not that as science advances, and our knowledge of
their development, of their structure, and of their habits ,
becomes increased, additional proofs will accumulate in
favour of our argument.
Professor Huxley has done good service in shewing
the relations of certain Kadiata, viz., Medusas, Physo-
phorida?, and Diphvdas, belonging to the Acalepha, or
sea-jellies, and Hydra and Sertulariadaa, placed among
Hydroid Polyps. He considers them " members of one
great group, organized upon one simple and uniform
plan, and, even in their most complex and aberrant forms,
reducible to the same type."* Among Echinodermata,f
there is evident adhesion to a common type, while there
is, at the same time, wide range in their general aspect.
In some of the sea-urchins the body is almost spherical,
in the sea-stars it is angular ; but these extremes pass
into each other by almost insensible gradations. Among
the sea-urchins, Echinocyamus and others present a
pentangular outline ; in Asteriscusj one of the sea-stars,
the general form is similar, the angles, however, being
very indistinct. In Solaster, the angles are more prom-
inent ; in Asteracanthion, Ophidiaster, and Luidia, the
angles are changed into true rays, and become more and
more' distinct from the body. In Ophiura, this separa-
tion into arms and body is complete, and in Eoryale, the
arms become very much branched. The flattening of
the body also differs ; — in Palmipes membranaceus, we
have a good example of extreme depression, while in
some species of Oreaster, the arms are very much dilated;,
so as to present in section the form of an equilateral tri-
angle. Among the sea-urchins we observe similar dif-
* Philosophical Transactions, 1S49.
+ Some hold that the Echlnodermata possess annnloso or articulate characters. W«
bere follow the views usually adopted respecting them.
OF EADIATA. 269
ferences ; the Echinus Sphasra is remarkably in contrast
with the depressed form of the Echinocyamus pusillus.
The soft and vermiform Holothurias are examples of
other sub-types of the Echinoderms ; still, a general plan
can be traced in all.
In star-fishes and urchins, we find copious deposits of
calcareous matter in the skin, in the form of distinct
plates. M. Graudry has very fully illustrated the general
plan which regulates this part of their organization.* He
has shewn that the protecting armour in all may be re-
ferred to three systems of parts — the endodermic or
internal, the dermic or intermediate, and the epidermic
or superficial. The internal system is absent in some.
The dermic consists of four systems in parts — the ambu-
lacra!, so called from the locomotive function of the
soft appendages which pass through them ; the inter-
ambulacra!, placed between the former series, and adding
strength and solidity to the whole framework ; the other
two, ovarian, and ana! or tergal plates, are respectively
connected with the reproductive and digestive systems.
The epidermic part of the armour comprehends all those
appendages called spines, scales, tubercles, &c, which
he shews to be formed after a common plan.
There are, moreover, traces of unity, when we examine
the minute structure of the plates or of the superficial
appendages. The microscope demonstrates that the
hard matter consists of branches disposed vertically, and
connected together by lateral branches, all of which are
referable to a typical form.
But while the Eadiate law generally regulates the
external form (and the general arrangement of certain
internal organs as well) we find that the number of the
radii is also subject to law. However much a sea-star
* Annales des Sciences Naturelles, 1S51.
270 TYPICAL FORMS
seems to differ from a sea-urchin, the number five pre-
vails in both. The question was long ago put by Sir
Thomas Browne, " Why, among sea-stars, Nature chiefly
delighteth in five points ?" and again, " By the same
number (live) doth Nature divide the circle of the sea-
star, and in that order and number disposeth those ele-
gant semicircles or dental sockets and eggs in the sea
hedgehog." "Every plate of the sea-urchin," says Pro-
fessor E. Forbes, "is built up of pentagonal particles.
The skeletons of the digestive, the aquiferous, and tegu-
mentary systems, equally present the quinary arrange-
ment, and even the hard framework of the disc of every
sucker is regulated by this mystic number."*
The same writer remarks, " When the parts of Echino-
derms deviate from it (five) it is always either in conse-
quence of the abortion of certain organs, or it is by a
variation by representation, that is to say, by the
assumption of the regnant number of another class.
Thus do monstrous star-fishes and sea-urchins often ap-
pear quadrate, and have their parts fourfold, assuming
the reigning number of Actinodermat a, consistent with
a law in which I put firm trust, that when parallel
groups vary numerically by representation, they vary by
interchange of their respective numbers."
Four is the number which generally prevails in the
Acalephs or sea-jellies. In Cyanasa, for example, the
stomach is usually subdivided by four ; four aasophageal
tubes are continued to their commencement, which is in
the form of a quadrate mouth, the angles being prolonged
into four tentacles. Sixteen canals radiate from the
central cavities.f
In the charming Cydippe of our own seas, the same
* Forbes1 British Star-fishes ; Introduction,
t Owen's Lectures on Invertebrata, p. 165.
OF RADIATA.
271
quaternary subdivision of the digestive system prevails.
Moreover, the cirri by which it makes progression in the
water, are arranged along eight equidistant bands.
The Actinas or sea-anemones, not merely have some
general resemblance to the well-known flower after which
they are named, but we find remarkable order as regards
number and relative position of organs, such as we have
seen to prevail in plants. M.
Hollard has shewn that the
concentrical series of tentacula
in the sea-anemone are subject
to a law of alternation. This
is well illustrated in the full-
grown Actinia senilis, the four
concentric series of tentacles
alternate with each other, and,
as regards the numbers in each,
the following is the formula : —
10 + 10 + 20 + 40=80
In some others the typical number is six or a multiple of
six. Thus there are twelve tentacula in the first row in
Actinia equina, six in Actinia pedunculata, and there
are four rows in the first species, and five in the second.f
Fig. 59*
SECTION II.-
-ADAPTATION OF RADIATE TYPES TO THE
MODE OF LIFE.
Amid the general adherence to the Radiate type, we
find modifications of parts in reference to locomotion,
prehension, and retention of food, protection from external
injury, and reproduction, all in evident accordance with
the wants of the animals.
The simple Hydra of our fresh waters, consisting as it
* Fin. 59. Plan of Sea-a;iemone, upper surface. The center circle represents the
month. The smaller circles represent the tentacula in concentric and alternating series,
t Annales des Sciences Naturelles, 1851.
272
ADAPTATION OF RADIATE TYPES
does of little more than stomach, has, in the position,
arrangement, and properties of its tentacula, admirable
means of securing its prey. Its habits, and the adapta-
tions of its organs, have been so often and so fully dis-
cussed elsewhere, that we need not dwell on the subject
here.* We merely allude to it in the outset on account
of its relations to certain others of which it may be re-
garded as the type.
The little Hydra propagates both by a process of bud-
ding and by the formation of ova. The buds sprout out
from the body of the parent, and passing through various
stages, finally become detached and independent beings,
each capable of produc-
ing others by the same
process. But sometimes
this mode of reproduc-
tion is so rapid, that
each new Hydra-bud ac-
tually has buds of its own
before it quits the parent
stock. These buds, how-
ever, finally drop off and
become independent, each
forming a fresh colony.
The same mode of
budding takes place in
many others of the Hy-
droida, with this dif-
ference, that the buds
usually remain attached
to the stock or parent.
Fig. 60.t
* See Trembley's Memoirs; Johnston's British Zoophytes,- &c.
t Fig. 60. Hydra fusca propagating by buds, a, mouth ; 5, base or point of attach-
ment, a to &, the original animal or stock from which the young or buds are formed
c, point of origin of one of the buds.
TO THE MODE OF LIFE.
273
But this building up of a tree-polyp could not proceed to
any great extent if all polyps were entirely of the same soft
texture as the Hydra. And here comes in a modification
to which we owe many of
those varied arborescent
forms with which the ocean
abounds. Long regarded
as plants they are now well
known to be compound
Hydroida. The develop-
ment of hard matter on the
outside serves as a means of
protection and support, in
a medium liable as the sea
is to such fluctuations in its
condition. The soft material
which pervades the centre of
the hard covering is just a
continuation of the digestive
system of the polyps, each
of which, protected in its
little cell, captures food
by means of its tentacula. The nourishment thus ob-
tained contributes to the growth of the united colony,
furnishing pabulum for the formation of new cells and
new polyps. But there is another mode of propagation.
There is a limit to the increase of the polyp-tree, and
necessity for the establishment of new colonies at a dis-
tance from the parent. There appear at certain periods
in the life of the Zoophyte, cells differing in form and
size from those which protect the individual polyps.
Fig. 61.*
* Fig. 61. Campanularia gelatinosa. A, fragment natural size; B, portion enlarged;
6, young polype-bud; d, adult polype in its horny cell e; e, transformed branch with
mednsoid buds iu different stages.
12*
274
ADAPTATION OF RADIATE TYPE
These are usually known by the names of ovigerous vesi-
cles. {Fig. 61, e.) The late Professor E. Forbes has de-
monstrated that these vesicles are not new organs differing
in their nature from other parts of the organism, but that
they are really modifications of a part or parts for a special
purpose. " The vesicle is formed from a branch or pinna,
through an arrest of individual development by a short-
ening of the spiral axis, and by a transformation of the
stomachs (individuals) into an ovigerous placenta, the
dermato-skeletons (or cells) uniting to form a projecting
capsule or germen, which metamorphosis is exactly com-
parable with that which occurs in the reproductive organs
of flowering plants, in which the flower-bud (normally a
branch clothed with spirally arranged leaves) is consti-
tuted through the contraction of the axis, and the whirl-
ing of the (individual) appendages borne on that axis,
and by their transformation into the several parts of the
flower (reproductive organisms.)"*
The vesicles are, therefore, branches modified for a
special purpose in the economy
of the animal, in the same way
as we have seen that the parts of
the flower in plants are merely
modifications of the typical ap-
pendage (the leaf ) arranged upon
a shortened axis. In some in-
stances special buds or indivi-
duals issue from the vesicles, be-
coming detached to enjoy for a time an independent
existence, for which they are accordingly fitted by special
Fig. 62.*
* Annals of Natural nistory, vol. xiv. 1844.
+ Fig. 62. Medusoid bud (See Fig. 61, e) of Campanularia ; it swims freely in the water.
a, body; b, mouth; c, upper surface; d, cirri. It is believed to produce ova, which are
developed intr> a Campanularia stock.
TO THE MODE OF LIFE.
275
modification. Through means of these buds the true re-
productive process by ova is effected. These individuals
when fully matured are flat discs, they may be compared
to an expanded umbrella with a short stalk. The margin
of the disc is provided with appendages, by which, and by
its own contractile powers, the organisms move and dis-
perse themselves in the water. They, in fact, correspond
to the flower in plants. Their organization fits them
admirably for an independent existence, and for dispers-
ing the ova at a distance from the parent stock.
The sea-anemones may in general terms be compared
to a jointless cylin-
der, the extremities
of which present
two distinct modifi-
cations in accord-
ance with their func-
tion. The base or
lower end is that by
which the animal
is fixed to a shell or
rock, the part acting
on the principle of
the sucker, but ca-
pable also of becom-
ing detached and
performing lateral progression to a new place at the will
of the animal. The free end of the body presents one or
more radiate series of hollow tentacula capable of pro-
trusion or retraction, and this by a very simple mechan-
ism, the injection or expulsion of water. When fully ex-
* Fig. 63. To show form and structure of Actinia or sea-anemone, a, point of at-
tachment or ba?o; b, mouth, c, tentcula; e, stomach; g k, partitions or vertical plates
ft, passages into tentacula.
Fig. 63.*
270 ADAPTATION OF RADIATE TYPE
paneled, these tentacles are effectual means of capturing
and retaining prey, and of conveying it to the central
mouth. This opening leads by a short canal to the capa-
cious stomach, the outer surface of which is connected
with the walls of the body by a number of radiating ver-
tical plates. The cells formed by the plates, which are
muscular, have a special function as regards the protru-
sion of the tentacula. As there is a certain order in the
arrangement of these organs, we find corresponding dis-
tribution of the vertical plates.
In our native species of Helianthoida, reproduction^
by ova is the most usual mode ; reproduction -by buds is
less common. But in many of the varied and beauti-
ful stone corals of tropical seas, colonies are formed by
the budding process. And here we meet with interest-
ing modifications in harmony with this mode of increase,
and the localities where the animals usually occur. A
colony of soft Actinias could not attain any great size,
and at the same time resist the destructive influence of a
turbulent ocean. The species of Millepora, Madrepora,
3tc., so well known to navigators, are Helianthoida, which
have the peculiar power of separating carbonate of lime
from the sea-water, and building it up in forms which
equally astonish us by their size, and please us by the
beauty of their details. The coral-builders, it is well
known, thrive best in the surf of the breakers, and their
peculiarities of organization fit them admirably for such
localities. In many of them we find calcareous matter
deposited in the interstices of the perpendicular plates
already alluded to, which afford support to the soft parts,
and enable them to resist the action of the surrounding-
medium ; the sea>water, at the same time, yields to them
the material for such purpose. In former epochs of the
earth's history, as well as in our own, the coral-builders
TO THE MODE OF LIFE.
277
have contributed in no small degree to modify the
earth's surface, and prepare it for the abode of higher
animals.
The statements already made regarding the compound
Hydroida and their detached animal-flowers, apply also
to certain of the Acalepha or sea-jellies.
The true Medusae commence
their existence as animals resem-
bling in no small degree the com-
mon fresh-water polyp. They
multiply for a time, by a process
of budding, and the final effort
is to produce other buds which
become developed into the full- fig.64*
grown Medusae. In both the conditions the radiate
type is retained, but in each kind of organism there is a
special modification in accordance
with the mode of life. The ordi-
nary buds are modified in accord-
ance with their sedentary existence;
one end forms a point of attach-
ment, the other is provided with
tentacula for the capture of food.
The other special buds, which pass
off from the common stock, are fitted
for independent existence, and for
progression in the water ; they move
from place to place by the un-
dulations of the umbrella-shaped disc and the action
.^--■'
* Fin. 64. Original stock, or polype condition of Medusa. Shews a group of Ave, four
of which have sprouted as buds from the original stock.
t Fig. 65. Polyp of Medusa, producing young Medusis. a, the stock or body; e, a
bud, as in last figure ; a, tentacula of stock; d, young Medusa, (corresponds to flower-bud
In plants,) with its tentacula and proboscis. Tentacula, c, are a second growth.
278
ADAPTATION OF RADIATE TYPE
of marginal appendages. They are provided with a diges-
tive system, and organs for capturing prey, and, finally,
produce abundance of ova,
each of which becoming fixed
to a rock or shell forms
a polyp stock, and gives
origin to similarly modified
organisms.
Among Ecliinoderms, as
we have already seen, there
is remarkable unity amidst
great diversity of form and
consistence of parts. This
diversity in particular cases
has an evident relation to
the wellbeing of the species.
The hard covering of star-
a mailed defence, combining, in most
instances, strength and flexibility. The many pieces
(thousands) of which it consists in some species, are
evidently suited for both the functions mentioned.
While the ovarian plates, pierced for the passage of the
ova, and the ambulacral, giving exit and support to the
delicate cirri, respectively occupy important relations as
regards the economy of the animal.
The Comatula, or rosy star of our own seas, presents
modifications in conformity with its habits. In its adult
condition it can cling to a rock, a sea-weed, or a coral, by
means of the simple-jointed arms with which it is pro-
vided for that purpose ; while, on the other hand, the
large pinnated arms may be used for free progression in
fishes constitutes
* Fig. 66. Advanced state (Medusa) of d, Fig. 65. A, side view; a, proboscis; &,
lobes, or subdivisions of margin. B, upper view of A, shows quadrilateral mouth in the
sontro.
TO THE MODE OF LIFE. 279
the water. In striking contrast with it are the sluggish
sea-urchins, whose protecting spines serve both for pro-
gression and defence, while the numerous cirri protruded
from the openings in the ambulacral plates, acting on
the principle of the sucker, enable the animal to anchor
itself, or when occasion requires, to move up a perpen-
dicular surface.
The Holotlmrias, while preserving the same general
radiate type in certain organs, differ in this respect that
their body is elongated — approaching the vermiform —
and the integuments are generally soft. They present
us with another modification adapting them for a dif-
ferent mode of life. Now moving by the suckers, which
protrude from the pores of the skin, and again by the
extension and contraction of their soft bodies, they are
fitted for localities inaccessible to their allies, the star-
fishes and sea-urchins.
CHAPTER IX.
NERVOUS, VASCULAR, AND MUSCULAR SYSTEMS.
In these systems of parts so essential to the animal
economy, we may expect also to find examples of types
and special adaptations, and our argument would be
incomplete without some reference to the subject. It
must not he supposed that the brevity with which we
discuss this department is any indication of its inferior
importance. More space has been devoted to types and
modifications in the internal and external skeleton and
appendages, because we believe that the proofs are more
easily accessible to the general reader.
NERVOUS SYSTEM.
The presence of a system of nerves is the most marked
character which separates the animal from the vegetable
kingdom. In some of the lower forms, its existence has
not been clearly demonstrated ; in many it is very rudi-
mentary. But as we rise higher in the scale we find an
evident advance, commensurate with the endowments of
the animal.
The simplest function of this system is that of convey-
ing an impression sufficient to excite the contraction of
muscular tissue, and thus effect some motion in an organ
or its parts. The impression is conveyed by one set of
nervous fibres to a centre — a ganglion, and from this it
NERVOUS SYSTEM. 281
is communicated to the muscle, which is thus stimulated
to contract. This reflex function is not necessarily ac-
companied by sensation, and the movements of the lower
forms of animal creation appear to be of the nature just
mentioned. But when we take a general view of the
animal kingdom, we find other superadded functions
dependent on this system ; " it is the instrument of con-
sensual and instinctive actions, of mental processes, and
of voluntary movements."*
In Mollusca, the typical nervous system is usually
described as consisting of three sets of nervous centres
or ganglia : — 1st, cephalic, supplying the eyes and other
parts about the head and mouth ; 2d, pedal, supplying
principally the foot ; 3d, parieto-splanchnic, supplying
the walls of the body, the heart and gills, &c. (See Fig.
45, parts marked n.) Now, we observe modifications of
this type corresponding to the development of the dif-
ferent organs, and the necessities of the animal. In dif-
ferent mollusca, where the foot is more or less developed,
we observe corresponding development of the pedal gang-
lion.
In the Cephalopoda, or cuttle-fishes, the large organs
of vision, the complicated buccal apparatus, and active
movements, are all in relation to the increase of nervous
matter, and concentration of its parts. Professor Sharpey
has further shewn an interesting modification in the
nerves of the arms in evident harmony with the habits
of these cuttle-fishes. Each sucking disc (on the arms)
is connected by a set of fibres with a ganglionic centre,
while all the ganglia are at the same time brought into
connexion by another fibrous tract with the cephalic
portion of the nervous system. Each sucker can, there-
fore— by reflex action — attach itself to any body which
* Carpenter's Comparative Physiology.
282 NERVOUS SYSTEM.
touches it, while all are also under the control of the
animal.
In Articulata, the typical nervous system consists of
two nervous cords running parallel to each other, and
connected at intervals by dilatations or ganglia in pairs.
(See Fig. 67.) The general arrangement is such that
every part of the body is furnished with two sets of
nervous connexions ; one of these is with the ganglion
of its own segment, and another with the cephalic gang-
lia. The distribution of the nervous system in Articulata
was an obvious relation to the general arrangement of the
hard parts, the body, as we have seen, being composed of
homotypal rings, bearing lateral appendages in pairs.
And as we find various modifications of this type in har-
mony with some important function, we also find corres-
ponding modifications in the nervous system. The late
Mr. Newport, to whose investigations we owe so much in
connexion with this subject, has shewn that in certain
cases there is an enlargement of a portion of the nervous
system at certain points, " corresponding to the apparent
greater necessity for accumulations of nervous matter at
those parts of the cord/' This remark is generally ap-
plicable as regards the ganglia of the head, the arrange-
ment being evidently in direct relation to the functions
of the important appendages of that part. There are,
farther, certain local modifications, having more special
connexion with the appendicular organs. Mr. Newport
states, regarding the nerves which supply the mandibles
and maxillae, that "union of those nerves at their base
is interesting from the circumstance that during mandu-
cation a consentaneous movement of the parts is required,
since, while the mandibles are employed in chewing, the
maxilla? are also employed in turning and assisting to
pass the food into the pharynx."
NERVOUS SYSTEM.
283
The concentration of the nervous matter in the thorax
is evidently a modification of the type in conformity with
the presence of wings and legs, the active appendages of
that part of the body. t
The wings require the
exercise of great mus- 2 '
cular effort in order ^fyl:--'
to support the insect 3 '"""^^^^^^[f^^^^^l^
during flight, and the j' -HHr"-
distribution of the ncr- 4 — .-r^pr^f^^
vous matter is in ac- jjjv
corclance with this ne-
cessity. But there must
also be perfect unison
in their action, and this c^-C2^
is also provided for.
Mr. Newport has de- — <^ •'■'aJjA
monstrated that there I-p^^Pr^E^
is a remarkable pecu- (WWo ^
liarity in the relations
of the thoracic ganglia
V I 1 > I ,
and their connecting
fibres, in those insects
in which both pairs of Fl(,_ 6r *
wings are actively con-
cerned in flight. He remarks, " that this is the reason for
this curious union of the nerves for the wings, seems ap-
parent from the circumstance that it exists in very many
tetrapterous insects of rapid or powerful flight, as in the
* Fio. 67. Nervous system of pupa of Sphinx Ugustvi, composed of two parallel ner-
vous cords, for the most part joined together side by side, and connected knots of ner-
vous matter— ganglions. The two larger masses and branches of nervous matter, or the
upper part, supply organs in the head, as eyes, jaws, &c, viz., 1 and 2; the two nervous
masses and branches succeeding to these supply the wings and legs, 8 and 4. The re*
maining portions are more uniform.
284 NERVOUS SYSTEM.
Apiclse and Ichneumonidas ; while in others, even of the
same order, as in Athalia centifolia?, which is well known
to fly heavily and hut a short distance, there is no such
combination/' In farther proof of the reason for the modi-
fication alluded to, he refers to the Coleoptera, in which
the anterior pair of wings is modified to protect the pos-
terior during repose. These anterior wings are merely
elevated, and nearly motionless during flight. Now, in
these insects " the nerves are derived separately from the
cord, and proceed to their destination without being first
combined in a nexus."*
As regards Articulata generally, the modifications of
the typical nervous system are admitted to be in consis-
tency with the functions of the organs to be supplied.
The larger nerves supply the organs of the senses, those
of secondary size go to the voluntary muscles, and the
smallest are for organs concerned in automatic motions.
In the organs of the senses, the size of the nerves appears
to be in inverse proportion to the density of the agent,
so the eyes receive the largest. The size is also in direct
proportion to the complex nature of homologous organs
in different species.f
The remarks of Dr. Carpenter are so much to the
point in reference to this part of our subject, that we
cannot do better than sum up in his words : — " In Inver-
tebrata, the nervous system consist of a series of isolated
ganglia, connected together by fibrous trunks. The
number of these ganglia, and the variety of their func-
tion, depend upon the number and variety of the organs
to be supplied. In the lowest Mollusca, the regulation
of the ingress and egress of water seems almost the only
function to be performed ; and here we have but a single
* Cyclopifidia of Anatomy, Art. Tnsecta.
t Straus Durckbeim, Comparative Anatomy of Articulata.
NERVOUS SYSTEM. 285
ganglion. In the star-fish wo have five or more gan-
glia ; but they are all repetitions one of another, and are
obviouslv the centres of action to the several segments
to which they respectively belong, neither having a pre-
dominance over the rest. And in the higher Mollusca,
and in Articulata, we have a ganglion, or more com-
monly a pair of ganglia, situated at the anterior extremity
of the body, connected with the organs of special sensa-
tion, and evidently exerting a dominant influence over
the rest. In the lower Mollusca, we have but a single
ganglion for general locomotion ; but this is doubled
laterally and repeated longitudinally in the Articulata,
in accordance with the multiplication of their locomotive
organs, so as to form the ventral cord. In like manner,
the Mollusca possess a single ganglionic centre for the
respiratory movements ; and this is repeated in every
segment of the Articulata, forming a chain of respiratory
ganglia, which regulates the action of the extensively-
diffused respiratory apparatus of these animals. The
acts of mastication and deglutition, again, in both sub-
kingdoms, are immediately dependent upon a distinct set
of ganglionic centres, which are connected, however, like
the preceding, with the cephalic ganglia. And wherever
special organs are developed, whose operations depend on
muscular contraction, ganglionic centres are developed in
immediate relation with them ; so as to enable them to
act by their simple reflex power, as well as under the di-
rection of the cephalic ganglia, as in the case of the
suckers of the cuttle-fish."*
From what has been stated, wc see evidences of a
common plan, with numerous special modifications neces-
sary to some end in the animal economy. In Vertebrate
animals, we find a very obvious correspondence between
* Manual of Physiology, p. 028.
286 NERVOUS SYSTEM.
the arrangement of the bony framework and the distri-
bution of the nervous centres ; skull and spinal column
are respectively constructed to give them support and
protection.
In viewing the entire animal kingdom, we find that
we cannot compare the whole of the well-developed ner-
vous system in the higher forms with that of the lower ;
still we find, in the nervous system of the Vertebrata,
certain parts which are homologous with the whole of
that of Invertebrata. In the higher Articulata, the
cephalic ganglia are considered homologous with a series
of ganglia forming a most important part of the brain-
mass in Vertebrata, and having relation to certain organs
of sense, as the eye, &c. The abdominal nervous cord
in Insects, &c, is homologous with the spinal cord of
Vertebrata, the essential difference being greater con-
densation of parts in the latter. than in the former. The
superadded portions in the nervous system of Vertebrata
have an evident respect to the superior endowments of
the animal. Cerebellum and cerebral hemispheres have
no distinct representatives among the Invertebrata series.
On comparing the very lowest of the Vertebrata with
the highest, we find evident difference in the relative
development of the most characteristic parts of their
cerebral system. The lowest forms of fishes have the
hemispheres of the brain in a very rudimentary condi-
tion, while in man they attain their highest development.
" The size of the Cerebral Hcmisjrfiercs holds a close
relation with the increase of the Intelligence, and with
the predominance of the Will over the involuntary im-
pulses. The increased size of the cerebellum, on the
other hand, seems connected with the necessity which
exists for the adjustment and combination of the loco-
motive powers, when the variety in the movements per-
NERVOUS SYSTEM. 287
formed by the animal is great, and a more perfect har-
mony is required among them."*
Such being the functions of brain and cerebellum, we
may expect to find modifications consistent with the ne-
cessities of the animal. The size of the cerebellum dif-
fers very much in the class of fishes ; but its development
appears generally to be in direct proportion to the active
powers of the animal. "Thus it is very small in the lazy
lump-fish, and extremely large in the active and warm-
blooded Tunny/'f In the Lampreys, whose mouth acts
on the principle of the sucker, so that they can attach
themselves to their prey and devour it at leisure, we find
that the cerebellum is relatively small. Whereas, in the
active and predacious sharks, it is of great size ; these
Felidas of the ocean have no swim -bladder, and their
mouths being placed transversely beneath the snout, they
require peculiar and active movements of the whole body
for securing and overcoming the struggles of the resisting
prey. This conformity of the development of the cere-
bellum to the peculiar habit of the animal is equally
illustrated in the class of Reptiles. Their habits are
generally inert, and the cerebellum is proportionately
small. The very reverse is true of birds — characterized
by the variety and power of their muscular movements.
We have already seen that in Articulata there are
local adaptations of the nervous system, co-ordinate with
the functions of the parts supplied by it. In the homo-
logous part — the spinal cord — of the vertebrata, we find
similar harmony. In certain reptiles we find this corre-
lation very obvious. In the serpent the absence of limbs
is accompanied with a remarkable uniformity of the spinal
cord and the nerves given off from it. On the other
* Carpenters Manual of Physiology, p. 530.
■( Owen's Lectures on Comparative Anatomy, vol. ii., p. 176.
288 VASCULAR SYSTEM.
band, in the frog, whose hind limbs are highly developed
and of great comparative muscular • power, we find cor-
responding enlargement of lower part of the nervous cord.
Two enlargements occur in the spinal cord of birds,
one corresponds to the wings, the other to the legs. As
might be expected, these enlargements generally present
differences of relative size, corresponding to the different
relative development and powers of the anterior and
posterior extremities. The posterior enlargement is
greater than the anterior in the Struthious birds (ostrich,
&c.) in which the whole function of progression is effected
by the posterior extremities. In contrast with this, we
observe that in birds of powerful flight, the greatest en-
largement of the nervous matter corresponds to the posi-
tion of the wings.
VASCULAR SYSTE1I.
Our remarks under this head will be confined to the
highest animals, viz., mammals and birds. In them we
O 7 7
find a highly developed system of vessels for distributing
the products of digestion, removing certain materials
derived from the waste of the animal frame, and supply-
ing the system with oxygen gas. These two latter func-
tions are in intimate relation to a surface or organ for
respiration.
The central organ of circulation — the heart — presents
the same structure in mammals and birds, and, generally
speaking, the blood-vessels are distributed according to
a plan which is common to both. Of the four cavities
of which the heart consists, two are set apart for the
purpose — the one receives, the other propels — of trans-
mitting the dark-coloured or venous blood to the lungs,
for the purpose of respiration.
Such are the functions of the cavities on the righl
VASCULAR SYSTEM 289
side, constituting the respiratory heart. Of the two on
the left side, one receives and the other propels the blood
— arterial — which has been oxygenated in the lungs.
"We have already pointed out the harmony between
the development and distribution of nervous matter, and
the necessity for variety and force of muscular effort.
But nerves and muscles cannot perform their respective
functions without a supply of oxygen. Now such co-ordi-
nation of parts is clearly illustrated by some peculiarities
in the arterial system of birds. The large muscles called
pectoral, (from their position on the breast) which are
chiefly concerned in the movements of the wings, are
supplied by arteries of great magnitude, " which, instead
of being inconsiderable branches of the axillary artery, are
the continuations of the trunk of the subclavian, of which
the humeralis only a branch."*
Another adaptation in the arterial system of birds we
shall allude to in the words of Dr. Carpenter: — "In
most Mammalia, as in Man, the right anterior extremity
is more directly supplied with blood from the aorta than
the left ; so that the superior strength and activity of
this limb would seem to be not altogether the result of
habit and education, as some have supposed. In birds,
however, where any inequality in the powers of the two
wings would have prevented the necessary regularity in
the actions of flight, the aorta gives off its branches to
the two sides with perfect equality."f
Among the mammalia, also, we find singular departures
from type in order to accomplish a special end. We
have already alluded to the habits of the sloth, and the
remarkable provisions in the structure of the skeleton.
The distribution of the vessels in its fore and hind limbs
* Cyclopaedia of Anatomy, Art. Aves, p. .534.
+ Principles of Comparative Physiology, p. 264.
13
290 VASCULAR SYSTEM.
is admitted to be a modification of the general ptan
suited to the habits of the animal. The arteries which
supply the fore and hind limbs are subdivided into a
number of branches, of nearly equal size, which commu-
nicate laterally with each other, and are exclusively dis-
tributed to the muscles. Those which supply the bones
and other parts, present no such peculiarity. The effect
of such distribution of the arteries will be to diminish the
velocity with which the blood flows to the parts. The
peculiar arrangement is admitted to have a relation to
the slow movements of the animals, though it may not
be easy to say " whether such slow movements of the
blood sent to the muscles be a subordinate convenience
to other primary causes of their slow contraction, or
whether it be itself the immediate or principal cause."*
The celebrated John Hunter long ago pointed out a
remarkable distribution of the vascular system in the
whales, an evident provision in conformity with their
power of diving and remaining for a time under water.
Their arterial system is characterized by extensive net-
works of vessels, chiefly distributed over the walls of the
chest. " It is to be presumed that this singular compli-
cation of vessels is caused by the necessity in which the
Cetacea are often placed of suspending their respiration,
and consequently the oxygenation of their blood during
a considerable time. These numerous arteries form,
therefore, a reservoir of oxygenated blood, which, re-
entering the circulation, supports life throughout, where
venous blood would only produce death/'f
We may now briefly allude to adaptations in the ves-
sels which carry dark-coloured or venous blood. The
typical venous system of the mammalia, according to
* Cyclopaedia of Anatomy, Art. Edentata.
t Cyclopaedia of Anatomy, Art. Cetacea.
RESPIRATORY SYSTEM. 291
Rathke, consists of four lateral primitive trunks. We
have just stated a peculiarity in the arterial system of
the whales ; the same animals present also a special
modification of the venous system in evident adaptation
to their habits. The extensive net-Works of veins in the
interior of the chest and abdomen, serve as reservoirs of
blood highly charged with carbonic acid, the accumula-
tion of which in the right side of the heart would occasion
death. The suspension of respiration during the act of
diving, renders such co-ordination of parts absolutely
necessary.
As connected with this subject, we may allude to a
peculiarity of the veins of the bat's wing, described by
Professor Jones. The wall of these vessels are endowed
with a power of rhythmical contraction and dilatation,
which, in the natural state, is continually going on at the
rate of seven to thirteen in a minute. This contractile
power, " supported by the presence of valves, is called
forth to promote the flow of blood in the wings, which,
on account of their extent, are, as regards their circula-
tion, in a considerable degree, though not entirely, be-
yond the sphere of the heart's influence."*
RESPIRATORY SYSTEM.
We shall only briefly refer to type and modifications
in this department. Allusion has already been made to
the necessity for oxygenation of the blood, and the re-
moval of the carbonic acid which accumulates in it dur-
ing its course through the system. In the warm-blooded
animals, whose temperature is generally higher than that
of the medium, air or water, which surrounds them, there
is another requisite, viz., the power of keeping up such
temperature by the combination of oxygen with materials
* Philosophical Transactions, Part I., 1852.
*292 RESPIRATORY SYSTEM.
supplied by the food, a process which is really a kind of
combustion.
In many of the lower animals there are no special or-
gans for respiration, the fluids of the tissues being suffi-
ciently aerated through the medium of their own walls,
or of the general external covering of the body. When,
however, special organs for respiration are provided, they
are admitted to be all mere modifications of one plan,
viz., a portion of the surface of the body of more delicate
texture than the rest, and permitting the atmospheric air
to pervade the parts, and come in contact with the nu-
merous vessels with which the organ is provided. Such
is the common plan on which both lungs for aerial and
gills for aquatic respiration are constructed.
Hitherto, in treating of types and modifications, we
have spoken of homological organs ; but in examining
the respiratory system, wTe have to do with some which
are analogous, but not necessarily homologous, that is to
say, similar in their function, but frequently different in
their nature. Nevertheless, it appears that in this view
also there are arrangements bearing on our subject.
Generally speaking, gills and lungs are respectively in
singular conformity with the different media inhabited
by the animals. Gills are usually extensions of some
part or other of the external surface of the body, and
being necessarily in contact with the water which yields
the air requisite for the performance of the function of
respiration, complex arrangements of organs are less re-
quisite, more especially since the general surface of the
body takes a part likewise in the act of respiration. In
reptiles,* in birds, and mammals, the respiratory surface
is internal, and the whole apparatus is more complicated,
* Certain reptiles begin life with gills, and some, even when mature, have both gills
»nd lungs.
RESPIRATORY 'SYSTEM. 293
and there are adaptations of various organs for perform-
ing the acts of inspiration and expiration.
We have said that respiratory organs are not neces-
sarily homologous, and in connexion with this, we find a
remarkable instance of departure from a plan, in accord-
ance with the necessities of the animal. The gills of fishes
are not of the same nature as the lungs of other Vertebrata,
still the latter organs have their homologues in the fish,
but they assume a new function, and one which is admi-
rably suited to the wants of the animal. The sound-blad-
der, swim-bladder, or air-bladder (for it has all these
names) by which certain fishes can regulate their depth in
the water, is a rudimentary lung turned to a new purpose.
Finally, whatever be the modification of the respira-
tory system, there is general adaptation to the nature of
the medium and the wellbeing of the animal. The gills
of fishes require no powerful efforts to bring fresh sup-
plies of water, and thus there is room for greater expen-
diture of muscular force in swift progression through the
medium they inhabit. Internal extension of respiratory
surface, well protected from external injury, is just such
a provision as is most conducive to the comfort of the
mammal. The whale, living in water, yet breathing by
lungs, has arrangements, in the form of its tail and in
the position of its nostrils, which enable it to rise to the
surface with ease, and get fresh supplies of necessary air.
The wide diffusion of air from the lungs through the soft
parts and bones of the bird, all directly co-operate to
facilitate ascent in the air, by diminishing the relative
weight of the body.
In Articulata, we find homologous parts concerned in
respiration, but acting through different media. In some
of the lower aquatic forms, the water- vascular system is
homologous with the branched vessels of insects, which
294 MUSCULAR SYSTEM.
are adapted for aerial respiration. In both instances the
arrangement is suited to the necessities of the indi-
viduals ; the extensive distribution of the air-vessels in
the perfect insect being in correspondence with the power
of flying, by reason of the diminished specific gravity of
the animal consequent on the very free access which the
air has been to every part.
MUSCULAR SYSTEM.
In this as in other departments, there is still much to be
accomplished as to our knowledge of a plan, and of modi-
fications. The few observations which we have to offer
will be confined to vertebrate animals. The general
arrangement of the muscular system corresponds very
much to the form of the skeleton. The greater or less
flexibility of the vertebral column, the size of the limbs,
the mode of progression, whether in water, in the air, or
on the ground, all imply greater or less peculiarities of
this system. The idea of general correspondence with a
type is clearly indicated by the nomenclature adopted in
describing the muscles of at least the three higher classes
of Vertebrata, viz., mammals, birds, and reptiles. The
following are some of the principal groups admitted by
anatomists : — muscles of the shin, of the spine and head.
of ribs, and ivalls of abdomen and chest, of limbs, of the
lower jaiv, of voice, eye, &c.
Intimately connected with the skin, and lying beneath
it there is a layer of muscular fibre in all Vertebrata —
with the exception of certain reptiles where it is unneces-
sary, owing to the development of hard matter in the
skin, and its consequent inflexibility.
At different parts of the body in the same animal, we
find local modifications evidently suited to some peculiar
end in the economy. Such tegumentary muscles are in-
MUSCULAK SYSTEM. 295
tended to act either on the skin itself, or on some of the
appendages which arise from it.
Among fishes, tegumentary muscles appear in connec-
tion with the dorsal and other fins, which can be thus
elevated or depressed according to necessity, either for
defence or offence, or as balances or partial aids in aquatic
progression.
In the class of birds, the muscles of the integument
frequently attain a high degree of development. In the
Apteryx of New Zealand they are of great strength — a
provision of the utmost importance, because its habits
expose it to accumulation of soil about its feathers, which
must be shaken with some force in order to dislodge it.
But there are bundles of small muscles connected with
the quill portion of the feathers. In some of the web-
footed species each feather has four or five small muscles
specially intended to move it in different directions. In
the Gannet it has been calculated that there are about
3000 feathers provided with such muscles, the total num-
ber of which will therefore not fall short of 10,000 or
12,000. It is by means of such skin-muscles that the
cockatoo elevates or depresses its crest, and the turkey-
cock bristles up his feathers. There are numerous oc-
casions on which these and other special arrangements
of the cutaneous muscles are needful in the economy of
the bird, and essential to its comfort.
The peculiar shield-like scales on the belly of serpents
are put in motion by muscles which belong to the cu-
taneous system, and are thus fitted to aid in progres-
sion over a rough surface.
In Mammalia, we find greater or less modification of
the same system in conformity with the habits of the
animal. The quills of the porcupine and of the hedge-
hog are set in motion by similar cutaneous muscles, and
the latter animal presents an additional arrangement
296
MUSCULAR SYSTEM.
Fie. 6S *
in other muscles of the same part, by which it can roll
itself into a ball, and thus
present a surface bristling
with formidable spines.
In other parts of the
muscular system, while
there is conformity to a
\ general plan, we also meet
I with local modifications.
J The very large proportion
which active voluntary
muscles bear to the whole
body, requires not only
proper adaptation to their
uses, but also such pecu-
liar packing and arrange-
ment as may be most conducive to the comfort and well-
being of the animal. We do not find the same propor-
tional distribution of muscle in whales, in fishes, in
birds, and in swift-footed mammals. In birds, for ex-
ample, "the principal masses of muscle being collected
below the centre of gravity, beneath the sternum, beneath
the pelvis, and upon the thighs, act like the ballast of a
vessel, and assist in maintaining the steadiness of the
body during night ; while, at the same time, the extre-
mities require only long and thin tendons for the com-
munication of the muscular influence to them, and are
thereby rendered light and slender."f
The great importance of the hand in man, and the
special development of his first digit, the thumb, imply
a correspondingly perfect system of muscles in the limb.
In the apes, which approach him nearest, the digits are
endowed with less individual mobility, and there is corre-
* Fig. 68. Shows muscular apparatus by which the hedgehog rolls Itself into a ball,
t Owen, Cyclopedia of Anatomy, Art. Aves.
MUSCULAR SYSTEM. 297
spending departure from the muscular type. These
animals are destitute of the power of appointing or indicat-
ing by means of the fore-finger, and man alone, who has
this faculty, possesses a distinct muscle for its perform-
ance. The muscle homologous with that called latis
simus dorsi in man, gives off in the apes a slip which is
attached to the elbow part of the ulna (the innermost of
the two bones of the fore-arm). This modification is es-
pecially obvious in the long-armed species, and enables
them to sling the arm forwards with great force and
quickness, by which they are better fitted to grasp distant
branches during their more rapid acts of climbing.
The singularly modified nostrils of the elephant, con-
stituting its trunk, present modifications of size, as well
as of relation in homologous muscles which are of less
general importance in other mammalia. The levator
and zygomatic muscles of the upper lip are very large,
and incorporated with those of the prolonged nostrils —
arrangements which are admitted to be commensurate
with the important functions of the parts. In hoofed
animals, fitted for rapid progression, the extensor muscles
of the limbs are more powerful than their opposing
flexors, the tendons also are long, the muscular portion
being short but strong — a modification suited to their
habits. When treating of the typical form of the verte-
bral elements, we have alluded to the singularly modified
arm-bone of the mole, a departure from the type in dis-
tinct relation to the no less remarkable arrangement of
powerful muscles in the limb, so necessary to the burrow-
ing habits of the animal.
The habits of that remarkable bird, the cross-bill,
(Loxia curvirostra) render necessary certain modifica-
tions in the muscles of its jaws. Feeding on the seeds
of firs, it requires an apparatus to extract them from the
hard and tough scales of the cones. The jaws cross each
298 MUSCULAK SYSTEM.
other, and act much in the same way as the blades of
scissors. We find a want of symmetry in the muscles
on the two sides of the jaws, and this has constant re-
lation to the position of parts. In a state of rest, the
lower mandible is drawn to one side or the other — for
there is a difference in this respect in different indi-
viduals. Now, the muscles are strongest upon that side
to which the jaw is so directed, and being of great
strength, the animal is thus provided with an apparatus
which enables it, with case and rapidity, to cut up the
tough cones and disengage the seeds. Perching birds
present a curious adaptation, by which the act of bending
the knee and ankle necessitates also bending of the toes,
independently of any active effort ; they can thus grasp
a branch even while sleeping. They are enabled to do
this by a peculiar disposition of the tendon of a muscle,
which is the homologue of that called gracilis* in mam-
malia. This muscle passes over the convex part of the
knee-joint, and then over the projecting portion of the
heel, and ends by being connected with a flexor muscle
of the toes. It is obvious, therefore, that the digits must
bend simultaneously with the bending of the knee and
ankle when the birds fall into sleep. In other parts of
birds we observe certain muscles differently modified for
different purposes. Thus, generally speaking, those on
the upper part of the tail are more highly developed than
those on the lower surface ; by such arrangement it is
that the peacock expands his gaudy tail feathers. On
the other hand, we find in some species that the muscles
which depress the tail are the strongest. This is well
illustrated in wood-peckers, which use that part as an
additional means of support when climbing, by pressing
it strongly against the bole of the tree.
* Meckel considers it homologous with the muscle called Rectus femoris ; this, how-
ever does not affect the importance and singularity of the modification alluded to.
CHAPTER X.
COMMt N1TY OF PLAN, WITH SPECIAL ADAPTATIONS IN THE
DEVELOPMENT OF ORGANIZED BEINGS.
Preceding observations have been confined to plan
and modifications of perfect adult organisms. It will be
necessary, in order to complete the argument, to glance
at tbe co-existence of the same two principles in the em-
bryonic condition of plants and animals. The nature of
the topic and its extent, obviously preclude its full dis-
cussion in this treatise ; but an epitome of some of its
leading truths will be found to fit in with what has gone
before, and with what is to follow.
If we enter a large ship-yard (to borrow an illustra-
tion from a friend) we may be able to discover a com-
munity of plan in the materials gathered together and
cut out for use, and in the very first blocking out of every
vessel. But it is as the fabric advances that we begin to
detect — what, however, was all along known to the
builder — what is the special purpose for which the ship is
intended ; whether it is to be propelled by sails or by steam ;
whether it is meant for warlike or peaceful occupations ;
whether it is to carry articles of commerce or passengers.
We are to show that it is the same in organic nature.
Every plant and animal is formed after a general plan,
while it is intended all along by its Maker for a special
and, and no other ; but it is only as it advances that we
300 COMMUNITY OF PLAN, WITH SPECIAL ADAPTATIONS.
can discover that end. We are to show that there is a
close resemblance between the foundation structure, or
earliest rudiment, of all plants and animals ; we are to
show that as the structures advance, each takes its peculiar
form to suit it to its evidently predetermined end ; and
we are to show, at the same time, that there is a remark-
able parallelism in the development of organic beings,
and this along the whole separate lines of their progress.
At an early part of our work we pointed out the iden-
tity of cell material in all organized beings. The germs
of every animal and plant present to us unmistakable
evidence of conformity to a model. The embryonal
vesicle of the animal, and the embryonic cell in the
plant, are obviously similar. Our remarks will be chiefly
confined to the former, as best fitted to illustrate the two
principles which are occupying our attention.
In plants, the contents of one peculiar cell (the pollen
grain) applied to the embryonic cell, determine all future
changes of the latter. Subdivision of the above-named
cell (embryonic) takes place ; the same goes on with con-
secutive cells ; these increase at the expense of the pabu-
lum supplied by the ovule, and the result of the process
is the formation of cotyledons, of rudimentary stem, of
root, and of leaves.
The structure of the ovum of the animal is very simple ;
it consists of a sac containing yolk, in the midst of which
there lies the embryonal vesicle which is essentially a
cell. This becomes filled with a mass of smaller cells, to
which it finally gives exit as a preparation for the process
of fecundation : after which a new cell, a primordial,
undergoes a change like that which follows the applica-
tion of the contents of the pollen to the vegetable germ.
This primordial cell begins to multiply by self-division,
until, from a single cell, we find that an aggregate mass
IN THE DEVELOPMENT OF ORGANIZED BEINGS. 301
of minute cells has been produced. It is worthy of
notice here, that as the cells forming the simpler plants
increase by subdivision, according to a fixed law as regards
number, viz., 2, 4, 8, 16. &c, so it has been distinctly es-
tablished that the same law of geometric progression re-
gulates the subdivision of the primordial cell, in the lower
forms of animals at least. In some of the Articulata this
process of increase is confined to the primordial cell and
its progeny, all being nourished at the expense of the
entire yoke, which disappears. But in other cases, the
subdivision of the above-named cell determines also a
similar process in the yolk which surrounds it, each cell
produced from the former attracting to itself a certain
portion of yolk, so that the increase of the original cell
occasions a corresponding- subdivision of the yolk. In
certain animals higher in the scale, it is observed that the
development of the cells (from the primordial one, placed
near the surface of the yolk, and not in the centre as in
lower forms) gives rise to subdivision of the yolk only
which lies near them. And here comes in one of those
adaptations presented to us in this early chapter of the
animal's history. The two portions of yolk are distin-
guished by different names, that which becomes subdi-
vided is termed " germ-yolk ;" the other portion, exempt
from such change, is called "food-yolk." The latter is
looked upon as something superadded to the former, and
is considered as a store of nourishing material to be used
up in the subsequent development of the new being. It
may be remarked that the size of the yolk is generally in
direct proportion to the advance made by the animal be-
fore leaving the egg. Thus in birds it is very large, they
escape from the egg in a well-developed condition. In-
sects quit the egg in an imperfect state, and the yolk of
their ovum is small.
802 COMMUNITY OF PLAN, WITH SPECIAL ADAPTATIONS
Iii Echinodcrms (sea-stars, &c.) there have been dis-
closed some remarkable modifications in the ovum for
special ends. The observations of Sars,* rightly inter-
preted by Professor M tiller, have shewn that soon after
the subdivision of the yolk, the young embryo of Echin-
aster rubens escapes from its membrane, and becoming
free and independent, is able to make progression in the
water. Here the mass of cells provided with numerous
cilia on the surface, assumes a kind of independent life,
and moves from place to place, thus providing for the
wider dispersion of the new being, which is subsequently
formed from it by a kind of budding process. A similar
example occurs, with some modifications, in course of the
development of the sea-urchins. The ovum escapes at
an early stage, and the ciliated surface of the cells is a
provision for locomotion and wide dispersion of the new
generation. Among Mollusca we meet with similar ex-
amples of adaptations for special ends. In the embryo
of certain Tunicata, we observe that a portion of the
yolk separates from the remainder, and is considered as
forming a tail-like organ, (the young animal is, in fact,
very like a tadpole) which is a most effectual means of
progression in the water, and accomplishes its purpose in
the same way as a single oar at the stern of a boat en-
ables us to scull it along. At a subsequent period, the
new animal becomes attached to some fixed object, and
the tail-like appendage disappears after having accom-
plished its temporary function.
In the ova of Vertebrata, we also meet with examples
of arrangements for special ends ; one may suffice. In
the egg of birds, we have concurrence of adaptations
very clearly illustrative of our subject, while, at the
* Sars, Fauna littoralls Norwegian; and MiMer, Uber den allgemeinen Plan in del
Entivickelung der Echinodermen.
IN THE DEVELOPMENT OF ORGANIZED BEINGS. 303
same time, the essential parts of the ovum are con-
structed on the same plan as that of other animals.
Commencing in the centre, we observe the yolk with the
germ-spot or cicatricula enclosed in the yolk-bag ; these
are suspended in the midst of the soft albumen, and
retained in position by means of two clastic chords, the
chalazce, which originate at the broad and narrow ends
of the egg. The albumen is in turn surrounded by a
tough membrane, (lining the shell) consisting of two
layers, which, being separated at the broad end of the
egg, constitute a chamber which serves as a reservoir of
air to be used up in the earlier stages of development.
The hard shell on the outside, and the lining membrane
for protection, the soft bed of albumen in which the yolk
is suspended by the elastic chalazae, and so suspended
that, being lighter than the other parts, the germ-spot
always rises uppermost, and so is nearest the warm body
of the parent during incubation — all constitute a series
of remarkable adaptations for special ends.
But without dwelling longer on the combination of the
two principles in the ova of organized beings, we may
briefly glance at certain conclusions which have been
founded upon the unity of structure which we have been
examining.
Because there is such remarkable similarity in the out-
set of life, it has therefore been rashly asserted that there
is resemblance also in subsequent stages, and that the
higher animals pass through a series of changes, each of
which exactly represents the permanent condition of some
other being, lower in the scale. It is true that in the
earlier stages of development, the ova of the highest and
of the lowest animals are much alike, find both also very
similar to the embryo of the higher plants, and to the
adult forms of some of the lower. Still we are not entitled
304 COMMUNITY OF PLAN, WITH SPECIAL ADAPTATIONS
to conclude that there is absolutely identity. The animal
ovum, removed and transferred to the same medium as the
simple plant, would assuredly perish. The resemhlance
amounts to this, the one is cellular as the other is, and the
cells in both propagate according to the same law. But
there is no ground for asserting more. And, as regards the
advanced stages, we have no foundation for the belief that
there is absolute identity of certain embryonic conditions
with permanent forms of animals lower in the scale. The
embryo of man is never, at any stage, of the same nature
as that of a worm, of a fish, a reptile, or a bird. In loose
terms, the higher foetus at a certain period may be ver-
miform in the sense of oblong, but it is never articulate ;
the relations of its parts are such that it could never by
possibility be declared that the two are absolutely the
same in organization. But as such assertions are not now
supported by authorities of any weight, we deem it un-
necessary to enter into further details. The great gen-
eral principle enunciated by Von Baer gives us the true
explanation of the phases of embryonic life, viz., " a he-
terogeneous or special structure arises out of one more
homogeneous or general ;" which may be simply illus-
trated by saying that the common homogeneous or
general material gives rise in course of development to
other special structures which are heterogeneous.
The idea has also been entertained by some that even
in the full developed or adult state there is unity of
plan in all, that the four types, Vertebrate, Molluscan,
Articulate, and Kadiate, are identical. Thus G-eoffroy St.
Hilaire considered the cuttle-fishes to be doubled up ver-
tebrates. This comparison cannot hold ; for although
Cephalopoda are the most highly developed of the mol-
luscan type, and, in some sense, higher even than that
singular fish called Lancelet, vet we cannot view them as
XS THE i>EVEU>PMENT OF ORGANIZED BEINGS. 305
nimMed vertebrata ; they are essentially molluscan. In-
sects were denominated by St. Hilaire, vertebrata with
free ribs — the legs being so considered. Others have
compared them to a vertebrate animal turned upside
down, the abdominal surface of the insect, next which
its nervous system lies, being considered as representing
the back of the higher animal, and the limbs as the
homologues of the laminre doisales.* That is to say, the
articulaf a were compared to the embryo of vertebrata with
the dorsal lamina? free, not entering into the formation
of the neural arches, but modified for purposes of loco-
motion. There can be no doubt, from what has been
already stated in a previous part of this work, that such
attempts at indicating identity in the four plans of struc-
ture are overstrained and far from representing the truth.
We have seen that in all animals alike there is a com-
mon starting-point, but as the development advances, we
observe that varied structures and arrangements of organs
appear, respectively suited to the sphere to be occupied
by the new being, and assigned to it by Him who is great
in power and excellent in working.
It has been already remarked how close the resem-
blance is between the ovum of the animal and of the
plant in the earlier stages of existence. But farther, there
is very striking similarity between the simpler kinds of
plants (Protophyta) and the lower forms of animals (Pro-
tozoa). This has given rise to the idea, that between
these at least there is no true line of demarcation, and
that therefore there is a merging of the vegetable king-
dom into the animal kingdom at the lowest extremes of
each. In both cases we have beings composed either of
a single independent cell, or of aggregated groups of
* These appear at a very early period of embryonic life, their external portions form
the rudiments of the back-bone and cranium.
306 COMMUNITY OF PLAN, WITH SPECIAL ADAPTATIONS
cells, and this has given rise to a difference of opinion in
defining which is the simple animal and which is the
simple plant. Both may have this in common — they can
propagate by simple subdivision of their parts. A mark of
distinction has been sought for in their mode of nutrition,
and it is probably in this that the true difference lies. The
simple plant is dependent on the presence of carbonic
acid and a sufficient supply of moisture ; the animal organ-
ism receives its supplies from other animals or plants.
In the resemblance which they bear to each other, we
can at least trace an amount of unity which indicates the
Oneness of the Designer.
In still higher forms of both animals and plants, we
have no difficulty as to fixed characteristics ; but even
among such, strange to say, we observe a remarkable
parallelism in the phases of development. In certain
animal organisms, a detached part, bearing the ova, has
been described as an entire and perfectly independent
organism, while the animal stock which produced it may
have been either overlooked or described as something
different. We have already alluded to this subject (see
Radiata) and, without further discussion, we shall now,
in further illustration of it, give a few parallel passages
from the history of the animal and of the plant.
Radiata.
Medusa. — 1. Ovum.
2. This egg becomes fixed, and
grows into a polyp-like animal or
stock, which produces free buds of
the same nature as the stock.
3. The stock gives off free swim-
ming Medusa?, these produce ova,
which repeat the two phases de-
scribed.
* Free buds are produced In not a few plants, as Lilium bulbiferum, Polygonum viv*
parum, Saxifraga cernua, &c., &c.
Plant.
Plant— I. Seed.
2. This seed germinates, and forms
stem and buds ; some of the latter
may drop off, and produce plants
tike the parent stock.*
3. The stock finally produces
flowers, fruit, and seeds.
eliaja:JC.p,301
McCOSH ON TYPICAL FORMS.
p. 307
IN THE DEVELOPMENT OF ORGANIZED BEINGS. 307
Plant.
Plant.— I. Seed.
2. This seed germinates, and pro-
duces stem and buds; some of the
latter may become detached from
the stock.
3. The stock finally produces
flowers, fruit, and seeds.
Plant.
Plant— 1. Seed.
2. This seed germinates, and pro-
duces stem and buds ; some of the
latter may become detached from
the stock.
3. The stock finally produces
flowers, fruit, and seeds.
MOI.LTJSCA.
Salpa. — 1. Ovum.
2. This ovum produces a solitary
Salpa, in the interior of which a
chain of Salpai is produced by a pro-
cess of budding.
3. Each Salpa of the chain pro-
duces an ovum, which attains nearly
full development, as No. 2, before
escaping.
Articulata.
Aphis. — 1. Ovum.
2. This ovum, in spring, produces
an individual which gives origin,
during summer, to several others
like itself, by a process of budding.
3. In autumn, males and females
are produced by the same process ;
the latter deposit ova.*
We have selected an example from each of the three
types, the Radiate, Molluscan, and Articulate, illustrative
of . the complete parallelism which there is between the
phases of their life and those of the plant. May there
not also be traced a parallelism between the plant and
the Vertebrate animal ?
* See Engraving. A. A seed produces a plant, which increases by the formation
of buds, which -are usually fixed. Finally, flowers and seeds are produced. B. An
ovum produced a ciliated organism, which becomes attached, and then gives rise to a
succession of polyps by a process of budding. Certain modified individuals produce
ova. C. An ovum produces a wingless Aphis (green-fly, &c. ;) this gives rise to others
like itself by a process of budding; (these differ from those of B and C in being free.)
Finallj', perfect males and females (winged) are produced, and ova are deposited.
1. Leaf-stalk of Ash, composed of a series of similar pieces.
2. Digit of Pithecus, (a species of Ape,) consisting of metacarpal bone and phalanges,
all presenting similarity of form.
I. Small branch of a species of Bamboo, (Bambusa arundinacea,) consisting of a series
of similar pieces.
II. Part of back-bone of Proteus, (reptile,) composed of a linear series of similar
pieces, (centra.)
1, 2, I., II., illustrate the remarks on typical form at pp. 185, 186.
A, B, and C, are adopted from Professor Owen's work, entitled " Parthenogenesis. "
308 COMMUNITY OF PLAN, ETC., OF ORGANIZED BEINGS,
Plant.
1. Seed.
2. Stem (internodes) and leaf-
stalk. (See p. 1S5 footnote.)
3. Ramified branch and venation,
(pp. 104-119.)
4. Axis, subterranean and aerial^
with appendages.
Vertebrata.
1. Ovum.
2. Typical bone. (See pp. 184-
187.)
3. Typical vertebra.
3. Vertebral column, with appen-
dages.
We can trace throughout organic nature a system of
Homotypes, or serially repeated parts, in the individual
plant and animal. We can also discover, in each of the
great leading divisions both of the vegetable and animal
kingdoms, a system of Homologous or answerable parts.
In very different organic structures we can find Analo-
gies, or different organs fulfilling the same function. But
we can do more : when ice compare the various organic
kingdoms one ivith another, we can detect parallelisms
in development, (Homceophytes.*) These parallelisms
may not be of the same scientific value as homologies
which now enter into the very structure of every. depart-
ment of natural science, but they are of equal, or at least
of similar, value in Natural Theology. The homceophytes
shew fully as clearly as the homotypes, the homologies,
or the analogies, that all organic creation has proceeded
according to a plan devised in eternity, and being realized
in time.
* This phrase has been suggested to us by our friend and colleague, ProC M'Douall.
CHAPTER XI.
GEOLOGY.
SECT. I. — TRACES OF PLAN IN FOSSIL REMAINS.
We have discovered proofs of One Designing Mind in
the organization of plants and animals in the existing
epoch of our earth's history. But our earth has also had
a past history. Could our globe relate the story of the
scenes which have taken place on its surface, what a
thrilling narrative would it furnish ! The dumb earth
possesses no power of detailing its past changes in lan-
guage, but it has carefully prepared in its crust, records,
which man has faculties to decipher, and which he may
succeed in deciphering, provided he proceeds with pains
and caution, and in the spirit and method of the induc-
tion of Bacon. The archaeologist draws conclusions from
the style of the ancient buildings examined by him, and
finds an entire history in the coins which he disinters
from their crumbling ruins ; the geologist can also gather
most important ins! ruction as to the past from the still
more valuable relics which are preserved in the rocks and
dust of our earth. It will be found that geology extends
our argument in respect of time throughout ages which
cannot be numbered, and shews that God has been pro-
ceeding in a pre-arranged system from the commence-
ment of creation.
310 TRACES OF PLAN
First, On examining the deposits of geological eras,
there is little difficulty in shewing that plants and animals
have been constructed on the same general plan from the
beginning.
Secondly, As the organisms of different geological eras,
while formed on a general model, do yet differ widely
from each other, the question is started, Is there a pre-
determined scheme in the successive appearances of ani-
mated beings ; or, in other words, is there plan in the
creation of classes, orders, genera, and species ? It
should be frankly acknowledged that geology is not yet
prepared to give a certain and decided answer to this
question. Still it has revealed phenomena which raise
the question, and supplied some facts which may help to
answer it, and furnishes proofs that there is order in the
succession of animal races, even though it has not yet
entitled us to say with confidence that we have discovered
the plan. Geology thus opens up glimpses not only of
a plan in respect of contemporaneous and existing nature,
but of a plan in respect of past and successive nature.
Thirdly, Geology has a further, and this a most im-
portant principle to reveal. It shews not only a uniform
but an advancing plan. It does more, it unrolls a pro-
phetic scroll, in which • the earlier animated creation
points on to the later, and in which the later comes as a
fulfilment of the anticipation of the earlier. These are
the topics to be discussed in this section.
I. Uniform Plan. — The silex dissolved in the water
of some ancient spring or lake has often entirely replaced
the materials of a stem, taking not merely the place but
assuming the very form and essential character of every
cell and modification of it, so that, when subjected to the
wheel of the lapidary, slices may be cut which, under the
microscope, reveal the most minute structure of the ori-
IN FOSSIL KEMAINS. 311
ginal plant. The elements, the living stones of the
extinct vegetable, have thus been wonderfully preserved for
our examination and instruction. Even when scarcely a
trace of vegetable structure can be detected, the inorganic
material of the earth's crust— the clay or mud of some
ancient lake or swamp, or the sand now forming the
stone of some modern quarry — has come into the place
of the organic framework, or received such an impression
by contact, that we have thus singularly preserved for
our inspection an accurate representation of a fruit or of
the venation of a leaf torn from the parent plant by a
passing hurricane, or shed naturally in the autumn of
some one of those countless years which elapsed before
man appeared. These relics shew that the same system
governed the building up of the ancient tree-ferns, palms,
and pines, as still regulates the formation of those that
surround us with all their symmetry and gracefulness.
How interesting is it to trace on these fossils, as we have
often done, the same crossing or winding spirals, and the
same rhomboidal figures produced by their intersection,
as we have in the tree-ferns and firs still growing on our
earth ; a proof that tire spiral then, as now, regulated the
position of the appendages of the plant.
A similar conclusion may be drawn from the animal
remains imbedded in the crust of the earth. The Uraster
obtusus of the older Silurian rocks has a striking resem-
blance to the Uraster rubens of our own coasts ; the ra-
diate arrangement of parts is identical in both. The
earliest Crustacea known to us, the Trilobites, present
the articulate type so familiar to us in the lobster and
crab. The shell of the little Nucula varicosa, found in
the same old strata, must have given protection to an
animal like that of our living species of that same genus.
The earliest spiral shells which have been discovered are
312 TRACES OF PLAN
governed by the same mathematical principles as those
which the molluscs are following at this day in the con-
struction of their habitations. The vertebrate column
and appendages of fossil fish, bird, and mammal, whether
of the older or more recent geological epochs, were formed
on the same models as those of the same models that still
people our earth and its waters. The teeth of extinct
animals were constructed on the same general plan as
those of existing species, and this,, whether we view them
as regards form, jDOsition, number, or minute structure.
Indeed, the geologist proceeds, and is entitled — by a
large induction of facts, and the verifications which are
ever casting up— to proceed, on the principles which we
have all along been illustrating in this treatise. It is
seldom that he iinds a fossil plant or animal entire ; most
commonly he falls in with only a fragment ; yet this
fragment, if it be a significant one, enables him to recon-
struct the whole. The process of theoretical reconstruc-
tion is conducted on those very principles of homology
and teleology which we have shewn to pervade all orga-
nic nature. The paleontologist supposes that the whole
organism, whether plant or animal, was constructed on a
plan ; that there were answerable parts in the genus or
species, and a series of homotypes in the individual ; and
he goes on confidently to supply the wanting parts on the
principle of homology. He proceeds, too, on the princi-
ple of final cause ; he supposes that the part had an end
to serve, and that there would be a conformity of every
other organ to fulfil that end. By means of these two
principles he can often, when he is in possession of but a
fragment, make the entire organism stand before us with
all its harmonies and its fitnesses. When at any time
he falls in with an entire fossil organism, he finds that
his principles are verified, and that he is entitled to pro-
IN FOSSIL REMAINS. 313
ceed on them. In the next section we shall shew how he
uses the principle of final cause ; in this section we are to
observe him as proceeding upon model forms in his inves-
tigations of the various kingdoms of nature.
Fossil Plants. — Certain vegetable organs have been
imperfectly preserved in the earth's strata, or have under-
gone such changes that it is often difficult to detect their
relations. The paleontologist does not hesitate to trace up
these to one or other of the great leading divisions of the
vegetable kingdom. He may not have before him, or be
able to find, some one part of the organism — say the seed,
so as to ascertain the structure of the embryo ; but he is
not thereby prevented from referring the plant to its pro-
per place, provided he can find out the structure of some
other part — say its stem, or the arrangement of the veins
of its leaves. If the venation of the leaf is netted, he
concludes that the plant proceeded from a seed with two
cotyledons, and was exogenous. Associations of charac-
ter, such as we have described in the chapter on the
Forms of Plants, are all important, not only in the exa-
mination of living, but of fossil plants. Fortunately the
structure, whether exogenous or endogenous, can be de-
tected in most fossil plants, and thus we have a key to
explain other arrangements which must have been asso-
ciated with it, and this holds true, whether we have the
whole stem or merely a fragment. In most cases we have
only a part, but when we do meet with an entire trunk, as
of Mantellia nidiformis in the petrified forest of the Isle
of Portland, we see at once that we have drawn legitimate
conclusions.
The characteristic venation, whether netted or other-
wise, obvious in the impressions of leaves met with in
various geological strata, it is so well preserved, that bota-
nists do not hesitate to refer them to one or other of tho
14
314 TEACES OF PLAN
leading divisions of the vegetable kingdom. The special
modifications of the veins arc also such that we can state
whether the leaf belonged to a plane or a beech ; and
one of the highest authorities, the late Von Buch, has
recommended a close study of the venation of the leaves
of living species as necessary for the discrimination of
vegetable leaf inpressions belonging to extinct forms.
In the remarkable Clathraria Lyellii, found in the chalk
marl of the Isle of Wight, the appearances are such as to
indicate that the leaves were shed naturally, just as in
existing trees ; a proof that the same organic relation of
stem and appendages existed in ancient as in modern
epochs.
Radiata. — The relics of corals, of sea-stars, and sea-
urchins, preserved and handed down to us in the pages
of the Palseographic volume, must be studied according
to the principles which apply to living forms. We can
expect fruitful results only when we proceed on the idea
of a regulating type.
The corals of different periods have in general a
certain plan of structure, but at the same time present
a remarkable contrast as to numerical type. It appears,
from the researches of M. Milne Edwards, that the cup-
shaped corals of the Palceozoic age have the stony
lamelke or rays regulated by the numbers 4, 8, 16, &c. ;
while in those of the Neozoic period, (including all
epochs from the Trias to the present time,) or newest
type, the regulating numbers were 6, 12, 24, &c.* Here
we have a remarkable example of order, enabling the
geologist to arrive at instructive results respecting the
position, in time, of rocks in which corals are pre-
served.
* It Is stated that only two exceptions occur ; one species of the quaternary type being
found in the chalk formation, and one of the ternary type in the Silurian rocks.
IN FOSSIL REMAINS. 315
HIoIIusca.—But few traces remain of the soft bodies of
the Molluscan inhabitants of the ancient world, and these
generally in such a state that we cannot draw any sure
conclusions as to their organization. The perfect condi-
tion of fossil shells, however affords us data from which to
reason as to the ancient modifications of the archetype ;
and we cannot doubt that extinct species were formed
after the same general plan as those which still exist.
The mathematical principles which determined the
forms of the shell in living species, as demonstrated by
Moseley, Naumann, and Elie de Beumont, have been ap-
plied by D'Orbigny in the examination of fossil species.
In his " Palasontologie Frangaise/' he shews that, even
when fragments alone remain, as is often the case in geo-
logical formations, the whole shell can be restored theoret-
ically, provided we have so much as two contiguous turns
of a spiral shell entire.
Articulata. — The Crustacea or crabs, the insects, and
other jointed animals of former ages, present the same
type of structure which prevails among the same families
at present, and this holds true from the very remote
epoch of the earliest Trilobites to the more recent forms
of Crustacea, preserved to us in the Lithographic slate of
Solenhofen. The Astacus Leachii and A. Sussexiensis,
from the chalk of the South Downs, had their rings and
appendages formed on the same general model as their
living allies, the lobster and cray-fish. Professor M'Coy
has shewn, that some disputed points in the characters of
the Trilobites can be interpreted when we proceed on the
principle of a general plan.*
Vertcbrata. — The able investigations of Cuvier, of
Owen, and of numerous other Continental and British
observers, are founded on the existence of a type or model
* British Palieozoic Fossils, Part II.
316 TRACES OF PLAN
in Vertebrata. It matters not how far back we examine
the records of the geological volume, we can see that the
method which regulated the construction of the most
ancient vertebrate animal known, was identical with that
which we can recognise in every being of the vertebrate
sub-kingdom which surrounds us. The well-preserved
jaws with teeth, and other relics, disinterred from the
bone-bed of the upper Ludlow rock, enable us to draw
conclusions as to the nature of the skeleton, and the mo-
difications of the archetype presented by it.
The ancient reptile, Telerpeton Elginense, is, so far as
we know at present, the oldest of its class. Imbedded
in its stone sarcophagus, we can recognise the existence
of a skull, back-bone, ribs, pelvis, and limbs. We can
count the ribs and the pieces of the spine, and sec that
the pelvis is placed after the 24th vertebra, just as in the
living Iguana.
Even " footprints in the sands of time" are capable of
yielding valuable results, where nothing else is left. We
can recognise, in ancient sandstones, the trail of tortoises,
of frogs, of lizards, and of birds. The feet which im-
printed them, and the entire beings, may have decayed
but the impressions left are such, that the nature of the
digits can be made out ; and authorities are agreed as
to the extinct Vertebrata having been respectively fur-
nished with the same kind of limbs which characterize
living forms belonging to the same classes of the verte-
brate type.
II. Progressive Plan. — The inherent desire of our
intellectual nature to discover laws, prompts us to inquire
whether there has not been order in the successive crea-
tions of animals and of plants. The facts already dis-
closed by geology seem to us to show that there was a
IN FOSSIL REMAINS. 317
predetermined plan in the appearance of new species of
organized beings. It is, however, very difficult to enun-
ciate what this order is.
One of our most distinguished geologists holds that
we have not arrived at a stage of knowledge to entitle us
to draw dogmatic conclusions as to the order of the ap-
pearance of animated beings, and his arguments, as well
as his name, must ever carry great weight. " I shall
simply," says Sir Charles Lyell, " express my own con-
viction that we are still on the mere threshold of our in-
quiries ; and that, as in the last fifty years, so in the
next half century, we shall be called upon repeatedly to
modify our first opinions respecting the range in time of
the various classes of fossil vertebrata. It would there-
fore be premature to generalize at present on the non-
existence, or even on the scarcity of vertebrata, whether
terrestrial or aquatic, at periods of high antiquity, such
as the Silurian and Cambrian."* While admitting the
force of this statement, it will, nevertheless, be necessary
briefly to state and examine some other views which have
been advanced.
First, it will be needful to notice the view of those who
maintain that there has been a gradual rise in the type
of animated beings, from the earliest period to the present
epoch. There has been at times associated with this,
another theory which derives all the higher and later
forms by natural law and progressive development from
the lower and earlier. It is proper to state, however,
that these two opinions have no necessary connexion ; the
former may be maintained by persons who deny the
latter ; the former may be true while the latter is false.
The facts revealed by geology seem to point to a begin-
ning of organized life. The lower we descend in the
* Lyell's Manual of Elementary Geology, 5th edit., 1S51, p. 663.
318 TRACES OF PLAN
strata of successive periods, the fewer the remains of living
beings. In passing downwards we reach a point where
there is but a single record preserved of the existence of
any organism ; we refer to the Zoophytes, Oldhamia an-
tiqua, and 0. radiata, found in the lowest Silurian rocks.
If we proceed from this point upwards, we find what
looks at first sight like a rise in type. What we mean
may be made evident without entering upon the con-
sideration of any other fossils than those belonging to the
vertebrate sub-kingdom.
And here we first of all meet with the fact that the
Invertebrata preceded the Vertebrata ; for there are no
traces of the latter till we reach the upper Silurian rocks,
far more recent in time than those which are lower. The
thin bone-bed of the upper Ludlow rock contains frag-
ments of fishes, relics of the most ancient beings of their
class. If we continue our examination, we next meet
with remains of reptiles in the upper Devonian strata.
The quarry of Cummingston, near Elgin, has yielded the
earliest reptilian relic known to us, and so well preserved
that the ribs and most of the skeleton can be distinctly
seen. It appears to have combined in itself the charac-
ters of the lizard and of the frog. Next in order of time,
birds and mammals appear in the Trias formation. The
Connecticut sandstone, which bears well-marked impres-
sions of footprints of birds, would seem to present the
earliest indications of that class. * And in the upper Trias,
Professor Plieninger has found molar teeth of an insect-
eating quadruped. Now we have here an evident pro-
gression in one sense ; first, invertebrata alone present
* In birds, every toe has the number of its hones remarkably constant, and each hav-
ing a characteristic number, it is obvious that we can by such marks distinguish the
foot-print of a bird. The outermost toe has always five phalanges, the fourth has four,
the third has three, the second lias two, and the spur or inner toe, has only one piece.
IN FOSSIL REMAINS. 310
themselves ; next, and after a long period, vertebrata
appear, beginning with the lowest, viz., fishes, next rep-
tiles, then birds and mammalia, in the inverse order
which they occupy as regards organization. But then
another question comes in, Is the first fish the lowest of
its class ? A similar question has to be asked of reptiles,
birds, and mammals.
The fish relics of the Ludlow bone-bed are sufficiently
well preserved to enable us to judge of the characters of
the beings of which they are the remains. Their jaws
and teeth are very perfect, and they give indications that
they were not the lowest of their class. The Onchus of
the upper Silurian rock " was a fish of the highest and
most composite order ; and it exhibits no symptom
whatever of transition from a lower to a higher grade of
the family, any more than the crustaceans, cephalopods,
and other shells of the lowest fossiliferous rocks. The
first created fish was as marvellously constructed as the
last which made its appearance, or is now living in our
seas."*
But it may be inquired whether the ancient Silurian
ocean was stocked only with fishes of high organization.
Suppose a sea with its scaly inhabitants, comprehending
sharks with hard teeth and shagreen skin, and also soft
lampreys and hags ; it is obvious that the relics of the two
former are more likely to be preserved to us than those
of the two last. This may be admitted, without, how-
ever, improving the argument as to a progression in type.
For although all are comprehended in Cuvier's division
characterized by soft skeleton, the sharks rank much the
higher — they are, in fact, the highest of their class ; the
highly developed brain, their organs of sense, &c., prove
them to hold the rank we have stated, the lampreys and
* Murchison's Siluria, p. 239.
820 TKACES OF PLAN
hags being far lower in type. The unequal development
of the tail (heterocercal) in the full-grown shark is the
only remaining argument in favour of their being per-
manent representatives of an embryonic state, and, there-
fore, low in type. But this also must fall to the ground
as an argument, because founded on an erroneous or
mistaken view of the case ; for the symmetrical develop-
ment of the tail actually precedes the unsymmetrical, in
certain fishes. The observations of M. Vogt, in reference
to this matter, has been either misunderstood or misre-
presented. The young Coregonus, (one of the salmon
family,) on which his investigations were made, has
actually at the first rays of the tail-fin arranged sym-
metrically above and below the end of the spinal column,
and therefore homocercal ; the unequal development of
the tail-fin (heterocercal) is the final condition, as, in-
deed, it is in the Salmonidas, contrary to the usual
opinion.* The earliest fishes known to us were not
the lowest of their class, but actually among the highest.
Evidence of a similar tendency is derived from a con-
sideration of some of the earlier invertebrata. One of
the most ancient Crustacea yet discovered, Hymenocaris
vermicauda, found in the Bangor slate, is not of low type,
it is among the highest of the Phyllopod order, which is
not very far removed in structure from the very highest
of the Crustacean class. It is not true, as has been
affirmed, that man and the higher animals, in their dif-
ferent stages of embryonic life, represent some permanent
forms of organisms lower in the scale ; nor can any proof
be adduced of an analogous progress in the womb of
time. Even if it were strictly true that there was a gra-
dual improvement in type as time rolled on, it would
* For additional remarks on this subject, we would refer to a Lecture by Professoi
Uuxley at the Eoyal Institution, April 1855. Annals Nat. History, July 1855.
IN FOSSIL REMAINS. 321
still be necessary that those who adopt the "development
hypothesis," should prove that transmutation of a low
into a high grade had been accomplished. Allowing
that the first position had been established, the question
remains, whether this might not have been the plan of
the Creator in bringing forward the beings which live on
our earth.
The supporters of the idea of progressive development
and transmutation of species in a long series of ages, be-
lieve also in a progression of life from sea to land, and
that this explains what they denominate " the barrenness
of Creation ;" that is to say, that certain conditions of the
earth's surface, favourable to the support of animals, long
preceded their appearance, inasmuch as time was required
for the necessary transformations of marine animals into
others fitted to live upon the land. It may be true that
uninhabited dry land existed at periods when the sea
was the abode of many invertebrata, and so may have
continued for a time previous to the appearance of ter-
restrial beings. But all this does not prove transforma-
tion of one animal into another, nor the progression of
life from sea to land. It remains to be proved — and the
onus probandi lies with those who make the assertion —
that marine animals can, by any force of circumstances,
or in any course of time, however long, become converted
into beings fitted to a new sphere of life on land.
If certain terrestrial conditions have preceded the ap-
pearance of animals suited to them, we have in all this
a manifestation of the foresight and beneficence of the
Author of all, and proof of a method which pervades all
creation. The bird constructs its nest before the callow
brood appears ; the bee lays in a store of food when the
flowers yield their sweet juices in abundance, and long-
before winter arrives ; an internal instinct leads to innu-
14*
322 TRACES OF PLAN
merable acts on the part of animals for the preservation
of their own lives, and for their young. In a word, there
are acts of anticipation flowing from instinct, which have
a special relation to some important end as yet in the
womb of time. And when we attribute foresight and
work of anticipation to Him " Avho knows the end from the
beginning," we do not consider such as derogating from
the infinitude of the wisdom of the Great Creator.
We find so many remarkable relations between the
physical condition of our earth and the wellbeing of its
races, that we cannot avoid seeing in the historical evi-
dence of geology some traces of order, a winter, a spring,
the seed-time, and a harvest of creation ; a winter when
life was absent ; a spring when preparation for it was
accomplished, and an era when it was called into being ;
and so sucessively to the time when the highest created
intelligence of our earth was brought forward to take pos-
session and occupy the earth now prepared for him. As
taking this view, Ave think that the argument in favour
of progressive development and transmutation of species,
founded on the pre-existence of conditions fitted for or-
ganic life — before that life appeared — is of no value.
The late Professor E. Forbes, by whose researches
geology has been so much enriched, has propounded an
ingenious theory on this subject.* In order to charac-
terize it he uses the term, " Polarity in Time," as expres-
sive of a law which corresponds to the primitive plan of the
Divine creation, but which, as being Divine, is completely
independent of the notion of time, although only com-
prehensible by us in relation to time. The different
geological epochs he comprehends under two heads, the
Pakeozoie, or most ancient, and the more modern, styled
Neozoic. On comparing these he finds that " the mani-
* Royal Institution, Evening Meeting, April 28, 1854.
IN FOSSIL REMAINS.
323
festations of generic types during each exhibit striking
and contrasting phenomena. The maximum development
of generic types during the Palaeozoic period was during
its earlier epochs ; that during the Neozoic period towards
its later epochs."
The following table renders the meaning more evi-
dent : —
rr> t J* „.• i. I Epoch of maximum develop-
Present and tertiary epochs, I * r
Neozoic
period,
Cretaceous epochs,
period.
Oolitic epochs, .
> Triassic epochs, . .
Permian epochs, . .
Palaeozoic J Carboniferous epochs,
Devonian epochs,
Silurian epochs, .
•| ment of Neozoic generic
( types.
Intermediate.
\ Epochs of poverty of production
( of generic types in time.
Intermediate.
! Epoch of maximum develop-
ment of Palaeozoic generic
types.
But besides the concentration of a maximum of gene-
ric types toward the earlier stages of one and the later
of the other great period, he thinks also there is a sub-
stitution of group for group during the contrasting epochs,
as shewn by the following comparison : —
Neozoic.
Cycloid and Ctenoid fishes.
Malacostraeous Crustacea.
Dibranchiate Cephalopoda.
Lamellibranchiate Acephala.
Echinoidea.
Six-starred Corals.
Palaeozoic.
Ganoid and Placoid fishes.
Entomostracous Crustacea.
Tetrabranchiate Cephalopoda.
Palliobranchiate Acephala
Crinoidea.
Four-starred Corals.
Some objections have been made to the general classi-
fication of geological epochs adopted by the author of
these views.* Where experienced and professed geolo-
* The objections refer to the position of the Permian and Triassic epochs in the tabular
view, and the propriety of comparing the primary period with the Jurassic, Chalk, and
Tertiary formations.
324 TRACES OF PLAN
gists arc at issue, it would be presumption in us to offer
any dogmatic decision ; but we cannot belp thinking that
an obvious objection applies here — and indeed, more or
less to every theory — it seems to be taken for granted
that we have almost, if not altogether, attained a suffi-
ciently complete knowledge of extinct forms. This is
surely far from being the case, and the lamented author
of the theory of Polarity had himself, in his comparatively
brief career, contributed so largely to our records of extinct
beings, that there is room for expectation that very much
still remains to be done, and that more information must
flow in as time rolls on.
But we pass on to another opinion, which seems, upon
the whole, very consistent with facts hitherto revealed by
the observations of palaeontologists.
As there is a certain law of progress in the develop-
ment of the young animal to the day of its birth, so there
seem to be some traces of parallelism to this in the order
of creation — a progress in uterine life, and a parallel
march in the womb of time, from the beginning of the
Creation to the day when man was ushered into existence.
In the development of the animal, Von Baer has shewn
that " the more special type is developed from the more
general" There seem to be proofs of similar progress in
time.
The subject has been very fully illustrated bjT Professor
Owen in his various writings. He remarks, " As we ad-
vance in our survey of the organization and metamor-
phoses of animals, we shall meet with many examples
in which the embryonic forms and conditions of struc-
ture of existing species have, at former periods, been
persistent and common, and represented by mature and
procreative species, sometimes upon a gigantic scale/'0
* Lectures on the Invertebrate.
IN FOSSIL REMAINS. 325
The common crab, in the different periods of its life, re-
presents conditions which resemble those met with in the
Crustacea of succeeding geological epochs.
2. Macrourous. (Tail long,) . . j
DEVELOPMENT OP THE COMMON CRAB. EXTINCT CRUSTACEA.
1. Entomostracous, .... Trilobites of the Palaeozoic age.
Crustacea of the Oolite forma-
tion.
3. Anomourous. (Tail moderately de- j Crustacea of the Chalk forma-
veloped, and of soft consistence,) l tion.
4. Brachyourous, the adult condition. \
(Tail short, and turned in beneath I Crustacea of the Tertiary epoch,
the thorax,) )
Other examples might be cited ; the above is suffi-
cient for our purpose. It must, however, be specially
observed, that " no extinct species could be reproduced
by arresting the development of any known existing
species of Crustacea ; and every species of every period
was created most perfect in relation to the circumstances
and sphere in which it was destined to exist."*
But extinct forms are not always the representatives
merely of the earlier stages of higher forms in the earlier
periods of creation. We find another principle illus-
trated : in some instances it is very evident that the
earlier forms " present in combination those characters
which are found to be separately distributed, and more
distinctly manifested among groups that have subse-
quently made their appearance/'f
A remarkable extinct order of Echinodermata has
been very fully examined and described by the late
Professor E. Forbes — the Cystidea : it illustrates the
point alluded to.
* Owen's Lectures ou Invertebrata.
+ Carpenter's Principles of Comparative Physiology, p. 112, 4th edition. In this ad
mirable work the reader will find a very lucid demonstration of the subject
326 TRACES OF PLAN
Paleozoic. Recext.
Order Cystidea. A stem, and intes- ) Order Crinoidea. A stem, and in-
tine with two openings. ) testine with two openings.
Order Cystidea. Certain species have } Order Ophiurida. Rays or arms
arms like those of Ophiurida. I snake-like, spines for locomotion.
Order Cystidea. In certain genera ) Order Asteriada. Body lobed, that
the body is lobed. ) is, angular or rayed.
Order Cystidea. Body enclosed in [ Order Echinida. Spherical or de-
a shell of polygonal plates. ) pressed shell, of polygonal plates.
Order Cystidea. Ovarian opening ) Order Holothuriada. Ovarian open-
single. > iug single.
From the above comparison, it will be seen that the
single extinct order Cystidea comprehended in itself
characters which are, so to speak, divided among five
orders at the present day. We have here, therefore, a
very notable instance of a progress from the more
general character to the more special in the lapse of
time — for the orders in the right hand column were very
partially represented in earlier epochs, and some of them
did not exist at all. Other illustrations might be
brought forward among Vertebrata ; we shall only allude
to one, as regards dentition. Professor Owen remarks
that the typical character of the dentition was more
closely and generally adhered to in genera than existed
during the oldest tertiary epochs in -geology, Hum in
their actual sticcessors. The earlier forms of mammals,
whether herbivorous or carnivorous, very generally pre-
sented the typical number of teeth, (p. 215,) whereas, in
the present day, such dental character is the exception
and not the rule.
It would be presumptuous in any one, at the present
stage of science, to suppose that he had been able
adequately to apprehend the plan in the Divine mind ;
but these facts seem to show that there has been an
IN FOSSIL REMAINS. 327
advancing series of some kind, proceeding all the while
on a uniform plan.
III. Peophetic Plan. — We are next to inquire
whether the earlier books of the stone-volume present
any records of organic forms, which point to higher forms
to come forth in later epochs; whether it discloses any-
foreshadowing of beings that were to follow ; and espe-
cially of man, the consummation of all.
The nature of the divine and creative act by which the
earliest of earth's creatures were summoned into being
must ever remain unknown to us. But it is allowable to
examine the aspect of these early organisms, and inquire
into the relations which they bear to the succeeding series
of animated beings. Our position in time, and the van-
tage ground on which natural science enables us to take
our stand, admit of our drawing an instructive comparison
between the forms of the Fauna in earlier epochs, and
those that appeared in later times. We confine our at-
tention, in what follows, to the Vertebrate type.
Siluria, rendered notable by the resistance of Carac-
tacus to the invaders of his country, is as famous in geo-
logy, as its former people are in the history of ancient
Britain. In its rocks are a succession of strata which
reveals to us what seems the dawn of creation in our
world. Its signatures appear to be the most ancient re-
cords of organic life. Those beautiful organisms, the
Graptolites, are not found in any palaeozoic rock younger
than the Silurian ;* and only one — the Graptolites prio-
don — is plentiful in the upper divisions of that system,
Grap. Flemingii of the Wenlock rock being rare.f We
have, therefore, a mark by which to determine the re-
lative age of the upper Ludlow bone-bed, in which the
* Murchisons Siluiia, p. 4T. U»id.v p. 208,
328 TRACES OF PLAN
earliest vertebrate remains occur. There is clear evidence
that they belonged to fishes, and, consequently, animals
formed after the vertebrate model. This is enough ;
here we find at a very early period, a plan of structure
which has appeared under various modifications in every
subsequent era.
Those few species of the upper Silurian period were
but the herald to indicate the subsequent advent of those
of the old red sandstone, remarkable not only for their
numbers, and their singularly bizarre forms, but some of
them especially interesting in relation to this head of our
subject. The highest authorities are agreed as to their
general place in the class of fishes, and the names of
Agassiz, of Professor Fleming, of Mr. Hugh Miller, and
others, must ever remain associated with the elucidation
of the history of these singular beings.
As the Onchus of the Ludlow rock announced, as it
were, the dawn of vertebrate life, and foreshadowed also
others of its own class that were to follow, so the Holopty-
chus, and others of the old red sandstone, in turn pointed
forwards to the Reptilian class. The term Sauroid (lizard-
like) has been applied to many extinct, and a few living
forms, in order to indicate their relationship to the
reptiles. The still existing Lepidosteus of America, and
Polypterus of the Nile and of Senegal, present a combin-
ation of characters eminently developed in not a few of
those found in the rocks of the Devonian epoch, (Old Red
Sandstone.) We can here take shelter under the high
authority of Agassiz, who remarks, " In Lepidosteus the
articulation of the vertebras differs from that of the verte-
brae of all other fishes, no less than the structure of their
scales. The extremities, especially the pectoral limbs,
assume a higher development than in fishes generally.
The jaws, also, and the structure of the teeth, are
IN FOSSIL REMAINS. 329
equally peculiar. Hence it is plain that before the class
of reptiles was introduced upon our globe, the fishea
being then the only representatives of the type of verte-
brata, were invested with the character of a higher order,
embodying, as it were, a prospective view of a higher de-
velopment in another class, which was introduced as a
distinct type only at a later period ; and from that time
the reptilian character which had been so permanent in
the oldest fishes was gradually reduced, till in more re-
cent periods, and in the present creation, the fishes lost
all their herpetological relationship, and were at last en-
dowed with characters which contrast as much with those
of Eeptiles as they agreed closely in the beginning."*
In a few existing forms, (Lepidosteus of America, and
Polypterus of the Nile,) and in all primeval fishes, the
pelvis and posterior limbs retained their position in con-
nexion with the point of junction of trunk and tail,
a character indicating superiority of type. This does
not apply to the fishes of subsequent epochs, for, from the
period of the chalk formation down to our own day, a
large proportion of them have the ventral or hind fins re-
moved from the typical position and placed far forwards,
near the head.
Such position of posterior limbs in the very dawn of ver-
tebrate creation, indicates an arrangement which was large-
ly to prevail in the vertebrata of subsequent epochs.
The Telerpeton of the Elgin sandstone ushered in the
dawn of reptilian life ; it is the earliest of its class yet
known to us. Fitted for a sphere of existence different
from that proper to fishes, it presents to our view a new
modification of the vertebrate plan. Its well developed
limbs point to a character which was to come forth more
prominently in succeeding periods.
* Natural History of Lake Superior.
330 TRACES OF PLAN
In 1726, Scheuchzer detected, in the comparatively re-
cent rock of (Eningen, a fossil which he set down as hu-
man, styling it " homo diluvii testis/' (man a witness of
the flood.)"5 This opinion did not stand the test of com-
parative anatomy, and the supposed human relic turned
out to be that of a large salamander. The time had not
yet arrived for the advent of man ; long ages had yet to
roll on before the consummation of the vertebrate type ;
the preparations for man's appearance were not yet com-
pleted. Nevertheless, in this fossil of Scheuchzer's, there
was a prefiguration of the more perfect type which man's
bony framework presents.
In 1847, Professor Plieninger of Stutgardt found two
fossil molar teeth, which must have belonged to a warm-
blooded quadruped ; they were disinterred from a bone-
bed in Wurtemburg, lying between the Lias and Keuper
formations. The original owner of these interesting relics
is supposed to have been an insect-feeder. A well-marked
tooth, pronounced on the highest authority, to have been
that of a warm-blooded quadruped, implies adaptations
of the vertebrate archetype of a far higher character than
any yet indicated in previous geological records. Such a
relic indicates associations of structure which are found in
man himself; and at this point in the earth's history, we
have the herald of the great mammalian class at the head
of which man is placed — the first in nature, though the
last in time.
Certain bipedal footsteps in the new red sandstone of
Connecticut, are recognized as those of birds. Man, the
true biped, was to appear in a subsequent and still distant
epoch.
But such early impressions and remains are not with-
* It is agreed on all hands that the origin of the human species is of comparatively
modern date. All fossil human remains, those of Guadaloupe, for oxamplo, are withia
the historical epoch.
IN FOSSIL REMAINS. 331
out their instruction ; we may recognise in all these pre-
existcnt beings the same type of skeleton, the beau ideal
of which was to come forward in the time appointed, after
the lapse of long ages.
Fishes, reptiles, birds, and mammals, predecessors of
man, presented in their frames anticipations of more
perfect structures which characterize Mm. They had
arrangements to protect the eye and the organ of hearing,
a bony vault to contain the brain, and limbs for various
functions necessary to their wellbeing.
The Supreme could foresee that which was to come,
and which He had pre-ordained ; the revelations of geo-
logy enables us to take a retrospective view. But they do
more ; they afford us the means of exercising a reflex
faculty ; we can examine the first figure in the vertebrate
series, and from that point look down the long vistas
which are opened, to the period when man appears as the
final and foreseen product of the one mighty plan — the
last in time, but the first in the contemplation of Him
who called them all into being. Precedent vertebrata
snadowed forth certain peculiarities of frame and of
psychical powers, which have their full, and evidently in-
tended, significance brought out and manifested only in
man. When he appears on the scene which had been so
long prepared, and, as it were, waiting for him, the con-
summation of the earthly type comes out ; — in a goodly
frame, with gait erect ; in eyes to contemplate, and men-
tal faculties to appreciate, the beauty of the objects
around him ; in limbs to bear that frame upright, and
carry it on in the fulfillment of its high sphere of duties ;
and in hands to minister to the wants of the individual
and of his fellows. Doubtless the structure of his body
binds him to the earth's surface, but he has mental
powers which enables him to soar from earth to heaven,
33- TRACES OF PLAN
to penetrate far into the regions of space, and throw back
a reflective glance upon the remotest points of time.
In the exercise of these mental faculties, it is expected
of him that he should contemplate with wonder and ado-
ration the wondrous scene spread out before him ; and
in the survey of the past he can discover that the earliest
fishes of the palaeozoic age pointed onwards to a higher
realization of the vertebrate plan ; that the plan has
never in any succeeding age been departed from ; that
it was at last perfected in his own wonderful frame ;
and that all this had been from eternity in the counsel of
Him who worketh in the whole from the beginning unto
the end.
We are happy to be able to adduce, in favour of this
general view, the testimony of the two greatest living
comparative anatomists. "It is evident," says Agassiz,*
" that there is a manifest progress in the succession of
beings on the surface of the earth. This progress con-
sists in an increasing similarity to the living fauna, and
among the vertebrata, especially in their increasing
resemblance to man. But this connection is not the
consequence of a direct lineage between the faunas of
different ages. There is nothing like parental descent
connecting them. The fishes of the Palaeozoic age are
in no respect the ancestors of the reptiles of the secon-
dary age, nor does man descend from the mammals which
preceded him in the tertiary age. The link by which
they are connected is of a higher and immaterial nature ;
and their connexion is to be sought in the view of the
Creator Himself, whose aim in forming the earth, in
allowing it to undergo the successive changes which geo-
logy has pointed out, and in creating successively all the
different types of animals which have passed away, was
* Agassiz and Gould's Comparative Physiology, p. 41T.
IN FOSSIL REMAINS. 333
to introduce man upon its surface. Man is the end to-
wards which all the animal creation has tended from the
first appearance of the* first Palaeozoic fishes." The lan-
guage of Owen is equally explicit.* " The recognition of
an ideal exemplar in the vertebrated animals proves that
the knowledge of such a being as man must have existed
before man appeared ; for the Divine Mind which
planned the archetype also foreknew all its modifications.
The archetype idea was manifested in the flesh long
prior to the existence of those animal species that actually
exemplify it. To what natural laws or secondary causes
the orderly succession and progression of such organic
phenomena may have been committed, we are as yet igno-
rant. But if, without derogation of the Divine power, we
may conceive of the existence of such ministers, and per-
sonify them by the term ' Nature,' we learn from the
past history of our globe that she has advanced with slow
and stately steps, guided by the archetypal light amidst
the wreck of worlds, from the first embodiment of the
vertebrate idea under its old ichthyic vestment, until it
became arrayed in the glorious garb of the human
form."
SECT. II. ADAPTATIONS OF FOSSIL ORGANISMS TO THEIR
FUSTCTIOXS. PREPARATIONS FOR MAN.
Plants. — The stem of the extinct plant (now converted
into stone) must have been as well fitted to sustain itself
erect, to receive and convey the fluids taken in by the
roots, and to support leaves for the elaboration of these
fluids, as the axis of any of our living trees. If we meet
with but the impression of a leaf, we cannot avoid draw-
ing the conclusion that the original, now lost to us, must
* On Limbs, p. 86.
334 ADAPTATIONS OF FOSSIL ORGANISMS
have had a framework of veins and an arrangement of
the softer tissues to enable the organ to fulfil its func-
tions. But any doubt existing on this point is removed
by the investigations of Groppert, who has found, in the
coal of Silesia and other countries, vegetable remains in
such a state of preservation that he could point out the
structure of the cuticle, and of its numerous stomata or
pores. He has also fallen in with a fossil plant, nearly
allied to the birch, with its branches bearing flowers.
And as, in our day, pine forests emit clouds of yellow
pollen, (giving rise to reports of showers of sulphur,) so
the giant pines of the ancient world have left proofs of
their existence, in abundant deposits of the same material,
characterizing certain strata in Bohemia and other local-
ities. On finding, in a geological formation, any remain
of what bears evidence of having been a fruit, the prin-
ciple of concurrence between structure and function leads
us to infer with confidence that the said fruit must have
been fitted to receive the pollen, and transmit its fertiliz-
ing principle to the ovule or ovules, and subsequently to
protect them during the process of ripening.
Radiata. — In fossil Radiata, the original hard material
of the body may remain, or silex has taken its place, (as
in some flints,) or merely casts of the organism may have
been preserved for our inspection ; but in whatever shape
presented, palaeontologists invariably proceed in their ex-
amination, whether consciously or unconsciously, from the
two principles of a plan and modifications.
Corals are abundant, even from the most ancient fossili-
ferous strata to the present epoch. In the seas of the
primeval earth, they were effectual agents in bringing
about changes in the contour of the land surface not less
important than those which are but too familiar to our
navigators in the form of coral reefs. The same modifi-
TO THEIR FUNCTIONS. 335
cations of the Kadiate structure which fit our modern
coral-builders for the part they are to play in the eco-
nomy of nature, must have existed in species long since
extinct.
The Wenlock limestone of Siluria abounds in remains
which afford unmistakable proofs of the agency of coral-
builders even in a very remote epoch. Species of Favo-
sites, of Stenopora, of Heliolites, (one of which is said
to resemble the Heliopora caerulca of the Australian
reefs,) were silently at work in former times, abstracting
from the sea-water its calcareous matter, and transform-
ing it into shapes which now delight us by their regu-
larity, while at the same time, they aided in adding to the
solid part of our earth's crust. Proofs of similar agency
occur in the carboniferous limestone, the reefs of which,
now rearing their crests far above the level of the existing
ocean, present us with evidences of some of the scenes and
changes through which our world has passed during its
eventful history.
In different geological strata, we meet with very per-
fect relics of Echinodermata, shewing modifications of
the type to which they belong, similar to the star-fishes
and urchins of our seas. Different adaptations for de-
fence, for capture of food, and for locomotion, present
themselves to us in species which have long since perished,
as in those with which we are so familiar. They give
evidence of relations of hard parts, and modifications of
form, and relations of form and function, similar to those
which we can read in a relic of any existing species cast
up by the tide, or put to the test, if we choose, in the
living animals themselves.
The Pentacrinus Briareus of the Lias is sometimes
found attached to fossil wood, which must have belonged
to some ancient tree, whose fragments formed drift-wood
336 ADAPTATIONS OF FOSSIL ORGANISMS
in the sea in which this singular Echinoderm lived. It was
a stalked species, characterized by excessive repetition
and subdivision of the radiate arms, ever ready to secure
the prey, as the animal was borne along on its wooden
float. If any doubt could exist respecting the modifica-
tions of the radiate type in this extinct Pentacrinite, it
must be dispelled when we compare it with the singularly
organized species (Pentacrinus caput-Medusse) which still
lives in the Gulf of Mexico.
In the upper Silurian rocks, we find preserved to us an
extinct form, which must have perished in the very act
of feeding. The Marsupiocrinites cselatus is frequently
found with its proboscis inserted into the shell of a mol-
lusc, (Acroculia Haliotis,) — both alike extinct.
In various strata we meet with abundance of animals
allied to the sea-urchins of our coasts. We can recog-
nise in the one, as in the other, some adaptations of the
hard parts to form a strongly arched shell for protection,
pierced with holes for the protrusion of the suckers, and
presenting the same arrangement of spines moving by
ball-and-socket joints.
Articulata. — Animals constructed after the Articulate
type had their representatives from the most ancient
periods in which traces of organized beings appear,
(Lower Silurian,) down to the most recent epoch which
preceded our own.
The Crustacean sub-type was a characteristic feature
of the Lower Silurian fauna : the singular Trilobites
must have swarmed in those early periods, and the re-
mains handed down to us, while they shew conformity to
a general plan, present also an almost endless variety in
the sculpture of their exoskeleton and the nature of its
contour. The admirable investigations of Burmeister
have thrown great light on the organization and habits
TO THEIR FUNCTIONS. 337
of these remarkable Crustaceans. They were nearly
allied to the Phyllopoda, characterized by the bladder-
like gills — the modified palp and fiabellum of the appen-
dage. The Silurian strata yield them in great numbers,
and their bodies are often found rolled up, so that the
head and tail are in contact. The best authorities seem
to be agreed as to the adaptations of the type, in these
ancient Crustaceans, to fit them for the kind of life
assigned to them. They constituted a remarkable feature
3f the Fauna of the Silurian Ocean. The soft abdomen,
and its delicate appendages, were liable to injury, and
by way of compensation, they possessed the power, when
alarmed, of doubling up the body, so as to bring the tail
under the head — the hard covering of the back thus serv-
ing to protect the more delicate under parts. The sud-
den catastrophe, which in some instances must have oc-
casioned their destruction and their imbedding in the
mud of the primeval ocean, induced also that change in
position to which we have alluded, and hence the occur-
rence of rolled-up Trilobites in the Silurian rocks.
In some parts of the -Old Red Sandstone formations,
fragments of a giant crustacean have been occasionally
met with. Being allied to the existing Limulus, or
king-crab, of warmer regions, the extinct species must
have presented similar adaptations ; — the limbs differing
little from each other ; the more anterior serving for
capture, retention, and mastication of the food, as well
as for locomotion.
Other articulata of the primeval world have been found
in excellent preservation. In the gypseous marl of Aix,
spiders are not unfrequently found. And in some speci-
mens, the spinnerets are distinctly perceptible. These
species, now lost to us, were, therefore, like our living
forms, provided with similar modifications of abdominal
15
338 ADAPTATIONS OF FOSSIL ORGANISMS
appendages for spinning the delicate web to ensnare
their victims.
Fossil insects, belonging to different orders, are not
uncommon in certain strata. We can recognize Neu-
roptera, Coleoptera, Diptera, and others, all implying
well-known adaptation of the articulate type. In the
Lacustrine deposits of (Eningen, a species of dragon-fly is
found in its different stages of larva, of pupa, and of
perfect insect.
MoUusca. — A skilled conchologist, finding a bivalve or
a spiral shell on a sea-beach, has little difficulty in form-
ing an opinion as to the general characters of the being
which reared such a habitation for itself. And so it is
when similar remains are disinterred from some stratum
of the earth's crust.
There seems to exist no doubt respecting the nature
of those fossils called Gomphoceras, Orthoceras, the
Clymenia of the Devonian epoch, the Ammonite, the
Hamite, and the Baculite. They were the Cephalopods
of the primeval seas, and, in general organization, were
allied to the cuttle-fishes with which we are familiar.
They must have been distinguished by their voracious
habits, and were provided with the necessary means of
securing and resisting the struggles of their prey. Shells,
which were built up by the ancient cuttle-fishes, abound
in various strata, and enable us to form some opinion
respecting the animal which they protected and sup-
ported. We can recognize an apparatus like that of the
living Nautilus. Compartments of the shell, not occu-
pied by the body of the animal, served as air-chambers,
giving buoyancy to the whole, and, by greater or less
compression of the air so enclosed, afforded a simple
means of rising or sinking in the water. It is no ro-
mance when we picture to ourselves the same modifica-
TO THEIR FUNCTIONS. 339
tions of the archetype in the extinct Cephalopoda, which
we have already seen to characterize those which are our
cotemporaries. The rapidity of their varied movements,
and their powerful arms, provided with sucking discs,
must have rendered them formidable enemies to their
fellow-inhabitants of the primeval oceans. But if any
doubt could exist respecting the general organization of
the beings about which we cannot help speculating, such
must vanish on examining the relics, or at least one spe-
cies, which have been presented for our inspection. The
Oxford clay of Chippenham has yielded the Belemnoteu-
this antiquus, with shell, mantle, fins, ink-bag, funnel,
eyes, and tentacula covered with sucking discs and hooks.
We have here, therefore, a complete epitome of structures
which we find in specjes which are our cotemporaries, and
a complete confirmation of all our conjectures.
In certain of the older Silurian rocks there have been
found relics which must have belonged to species of
Pteropoda ; Conularia, Theca, &c„ are examples. As
Creseis, Cleodora, and others of our own time, flit from
place to place in their ocean element — in a habitation of
their own building — so the extinct species have made
progression by wing-like appendages, a modification of
the epipodium of the archetype. But there were giants
on the earth in those days. Judging from their shells,
the Pteropods of the Silurian ocean greatly exceeded in
dimensions the species which swarm in some of our seas.
Vertcbrata. — Not only do we find in fossil remains
evidence of the first great law we have been illustrating,
there are equally clear proofs that the different organs
preserved for our examination had a final cause, and it
is impossible to avoid the conclusion that there must have
been a concurrence and co-operation of other' parts to ac-
2omplish the end in view. The statement of Cuvier on
340 ADAPTATIONS OF FOSSIL ORGANISMS
this point can never bs set aside ; " every organized in-
dividual," says he, " forms an entire system of its own,
all the parts of which mentally correspond and concur to
produce a certain definite purpose by reciprocal reaction,
or by combining towards the same end." " If the viscera
of any animal are so organized as only to be fit for the
digestion of recent flesh, it is also requisite that the jaws
should be so constructed for seizing and retaining it ; the
teeth for cutting and dividing its flesh ; the entire sys-
tem of the limbs, or the organs of motion for pursuing it
and overtaking it, and the organs of sense for discovering
it at a distance. The animal must also have been en-
dowed with instinct enough sufficient for concealing itself,
and for laying plans to catch its necessary victims."
The giant Megatherium of the new world presents, in
itself, an epitome of departures from the archetype skele-
ton for special ends. Its comparatively light skull was
supported by neck-vertebraa, small when compared with
their homotypes in other parts of the body. Those of
the loins are largely developed in harmony with the great
size and strength of the hind limbs ; and for the purpose
of additional strength, the sacral portion is united in a
peculiar way to the pelvis. The vertebrae of the tail are
of large dimensions, commensurate with the functions
of this part, which was used as an additional supporting
pillar, just as the same part is employed by the living Ar-
madillo in certain of its movements. The high develop-
ment of the hamial arches of the Megatherium's tail, in-
dicates that the blood-vessels supplying it were duly pro-1
tected from risk of injury from pressure. Its powerful
arms were so formed as to allow free rotation when root-
ing up the plants necessary for subsistence, the strong
hind limbs and tail together forming firm pillars of sup-
port during the process.
TO THEIK FUNCTIONS. 341
In certain geological epochs, the earth had also
its feathered inhabitants. In the remains which have
been preserved for our inspection, we find special adap-
tations in the skeleton such as occur in the class of bird;-;
generally, and also local modifications in harmony with
the habits of the particular species. The giant Dinornis
of New Zealand doubtless employed its beak as a kind of
pick-axe (which it resembles in form) for digging the
farinaceous roots on which it fed. The peculiarities of
the neck-vertebras, and the strong ridges and processes of
the occipital part of the head, all indicate the presence of
powerful muscles necessary for the exercise of such a habit
as that mentioned.*
A period was, when numerous reptiles, of varied form
and habits, constituted a leading feature of the Fauna in
the primeval world. The waters swarmed with species
fitted for aquatic life ; others roamed on the dry land ;
and not a few, possessed of the power of flight, obtained
sustenance by pursuit and capture of insects — pursuers
and prey being now alike extinct. In all cases the relics
which have been discovered present such marked modi-
fications, that anatomists are agreed as to the habits
of the species, so that the restoration of- their forms
and descriptions of their habits with which we have been
furnished from the ready pencil and graphic pen of
palaeontologists, however romantic they may seem, are,
we believe, nearer the truth than the accounts which
have sometimes been given even of certain animals which
still exist. The Pterodactyles (wing-fingered) were en-
abled to support themselves in the air by means of mem-
branous expansions, supported principally by the fifth
digits of their fore-limbs, each of which exceeded in
length the whole vertebral column of the animal, and
* Professor Owen, Zoological Proceedings, 18-18.
342 ADAPTATIONS OF FOSSIL ORGANISMS
was therefore not a little finger, though the homologue of
the smallest in the hand of man.
In the British Museum there is a model of a young in-
dividual of an extinct colossal tortoise (Collossochselys
Atlas) from India. This model is ten feet in length,
tvveny-five in horizontal circumference, and fifteen in ver-
tical girth, a third less than in the full-grown animal. In
this giant of former days there existed the same singular
modifications of skeleton (p. 204) which we have already
alluded to as a characteristic of tortoises which still exist.
The fishes which glided through the seas of the primeval
earth have left behind them such well-marked relics that
we can see pectoral and ventral fins, and a well-developed
tail for aquatic progression. We observe that the same
modifications of skeleton and appendages had the same
relation to the wants of the animals which we find in the
scaly inhabitants of our own waters.
The teeth of extinct vertebrata are found in such abun-
dance, and in such a state of preservation as to afford in-
disputable evidence of special adaptations, whether we
examine them in mammals, reptiles, or fishes.
The gigantic Iguanodon, of the Wealdcn formation,
presents singular adaptations in the form and structure
of the dental apparatus with which it was provided. " To
preserve a trenchant edge, a partial coating of enamel is
applied, so that the thick body of the tooth might be
worn away in a more regularly oblique plane ; the dentine
diminishes in density as it recedes from the enamel.
Finally, when the enamel is worn away by constant use,
and the tooth from a kind of cutter becomes a grinder, a
third substance of a different density from the dentine,
viz., the ossified pulp, adds to the efficiency of the tooth
in its final capacity."*
* Professor Owen's Odontography, p. 283.
TO THEIR FUNCTIONS. 343
In extinct fishes, adaptations of teeth are equally ob-
vious. The Rhizodes had long and powerful teeth, fitted
for overcoming the struggles of its prey. In order the
better to fit them for holding fast, " the teeth have a
broad base divided into a number of long and slender
cylindrical processes, implanted like piles in the coarse
bony 'substance of the jaw."*
When peculiar modifications have enabled us to ascer-
tain that a fossil bone belonged to fish, reptile, bird, or
mammal, we hesitate not to conclude that scales, feathers,
hairs, &c, must have been respectively the external cover-
ing of the animal. The extinct Glyptodon of South
America has left behind it relics of a tessellated, bony
cuirass, very much resembling that of living armadillos.
The carcase of the mammoth, embalmed in the frozen
soil on the banks of the river Lena, has supplied our
museums with samples of hair and wool, which must
have assisted to protect the animal from the cold blasts
of the region it frequented. And, in fact, the presence
of such covering affords us no insignificant evidence of the
probable nature of the climate in that remote epoch and
region, when those northern elephants formed a chief cha-
racteristic of a fauna which is now so changed.
Finally, the history of our earth's crust cannot be pro-
fitably examined apart from the different plants which
have at various periods clothed its surface, or the animals
which were successively brought into being. A palaeon-
tologist must also be a zoologist and botanist, for we
cannot view living and extinct forms as essentially differ-
ent embodiments of the Divine Counsel, but rather as
manifestations of the same Supreme Wisdom in various
consecutive ages. Respecting the unity of plan in all
epochs, there seems to be no difference of opinion. The.
* Odontography, p, 63.
344 ADAPTATIONS OF FOSSIL ORGANISMS
universality of the second principle — adaptations for spe-
cial ends — may not be equally capable of demonstration
in extinct forms. It is scarcely to be expected that we
should be able, in every instance, to prove a relation be-
tween means and end in the economy of every extinct
animal or plant, seeing that in general we have only
fragments to deal with. Such expectation would be,
besides, presumptuous on our part ; for while our finite
understandings can comprehend so much, they cannot
fathom the full depths of the Infinite Mind. As science
advances, however, we may expect that obscure points
will be rendered more clear, our doubts dispelled, and
proofs of special ends increased. The admission of
the first principle — of type, namely — will greatly aid
as a means of multiplying examples illustrative of the
second, and simplify the study of the organic beings of
every epoch. We shall close this part of our subject
with a quotation from Professor Owen :* — " Of the nature
of the creative acts by which the successive races of ani-
mals were called into being we are ignorant. But this
we know, that a-s the evidence of unity of plan testifies to
the oneness of the Creator, so the modifications of the
plan, for different modes of existence, illustrate the bene-
ficence of the Designer. Those structures, moreover,
which are at present incomprehensible as adaptations to
a special end, are made comprehensible on a higher prin-
ciple, and a final purpose is gained in relation to human
intelligence. For in the instances where the analogy of
humanly-invented machines fails to explain the structure
of a divinely-created organ, such organ does not exist in
vain, if its truer comprehension,, in relation to the Divine
* Orr's Circle of the Science?, Treatise, No 2 ; a work which contains an admirable,
summary of facts regarding typo and modifications in skeleton of vortebrata. The low
price of the work brings it within reach of all.
PREPARATIONS TOR MAN. 345
idea or prime Exemplar, lead rational beings to a better
conception of their own origin and Creator."
Turning to a somewhat different branch of the same
general subject, we find that throughout the whole series
of geological ages there has been an adaptation, one to
another, of the animals and plants on the one hand, and
of the state of the earth, its atmosphere and climate, on
the other. There has also been a preparation going on
all along for the appearance of a higher being on our
earth's surface. The comfort of man is dependent on
the condition of the earth— the place of his temporary
abode and probation, and this is the result of methodical
operations going on for long successive ages.' Man's life,
too, is inseparably linked with the plants and animals
which coexist with him, and these are also the issue of
long anticipations and preparations. The eternal Logos
— himself in due time to become flesh — had contemplated
all this from the beginning. " The Lord possessed me
IN THE BEGINNING OF HlS WAY, BEFORE HlS WORKS OF OLD.
I WAS SET UP FROM EVERLASTING, FROM THE BEGINNING,
OR EVER THE EARTH WAS. WHEN THERE WERE NO
DEPTHS, I WAS BROUGHT FORTH ; WHEN THERE WERE NO
FOUNTAINS ABOUNDING WITH WATER. BEFORE THE
MOUNTAINS WERE SETTLED ; BEFORE THE HILLS WAS I
BROUGHT FORTH : WHILE AS YET He HAD NOT MADE THE
EARTH, NOR THE FIELDS, NOR THE HIGHEST PART OF THE
DUST OF THE WORLD. WlIEN He PREPARED THE HEAVENS,
I WAS THERE : WHEN He SET A COMPASS UPON THE FACE
OF THE DEPTH : WHEN He ESTABLISHED THE CLOUDS
ABOVE : WHEN He STRENGTHENED THE FOUNTAINS OF THE
DEEP : WHEN HE GAVE TO THE SEA HlS DECREE, THAT
THE WATERS SHOULD NOT PASS HlS COMMANDMENT : WHEN
HE APPOINTED THE FOUNDATIONS OF THE EARTH : THEN
was I by Him, as one brought up with Him : and I
15*
346 preparations for man.
was daily hls delight, rejoicing always refore
Him; rejoicing in the habitable part of His earth;
and my delights were with the sons of men."
Since the remote period when dry land first appeared,
the different substances entering into the formation of
the crust of the globe have been continually subjected to
a process of decomposition brought about mainly by the
influence of heat, and moisture, and by the action of the
atmosphere. The same moisture which aided in this
process has been the means by which the products of
such decomposition have been carried off and re-arranged
in some neAv form. Eivers and their tributaries have
served to convey the debris of the rocks to the ocean,
there to be deposited in the form of a fine sediment,
which enriched, perhaps, by the decay of marine organisms,
and, after various changes, has been finally upheaved
above the surface of the waters. The extent and great-
ness of those operations by which the dry land has been
fitted for the growth of land vegetation, and prepared
for the reception of animals, may startle us by their vast-
ness ; but there are abundant proofs of such great changes
— the records of geology indeed teem with them. The
gigantic scale on which operations, which may be styled
the husbandry of nature, have been conducted, may well
surprise us ; but we cannot withhold our belief as to such
processes, and the important results which have followed
in their train. Subsoil ploughing, mixing and re-mixing
of soils, have been going on in all ages. Man is but the
unwitting copyist, on a small scale, of actions which
have been conducted on a far greater scale, and ap-
parently with his benefit in view. Those very qualities
which a good soil ought to possess, have been induced in
course of time by various chemical and physical agencies,
which have been in continual operation. The debris
PREPARATIONS FOR MAN. 347
of rocks yielding calcareous, silicious, aluminous, and
other mineral ingredients, have been brought together, and
mixed in a way which the husbandman imitates when
necessity demands. The furrows drawn by our plough-
shares are but scratches on the surface of the soil, com-
pared with the changes to which that same soil has been
subjected in former ages, and to which it owes its varied
capabilities of supporting plants, and yielding subsistence
to the animal kingdom.
• The respiration of animals, the decay of certain organ-
ized substances, the act of combustion, and emanations
from volcanic foci, add to the atmosphere a gas which is
not chemically a necessary ingredient of that atmosphere.
The gas referred to is carbonic acid, which at the present
day forms about one thousandth part, by weight, of the
air which surrounds us. It is one of the chief sources
from which plants derive their more solid ingredients.
They are continually taking it in, and storing it up by
moulding it into shapes and qualities, of which we con-
tinually avail ourselves for different necessary purposes.
An excess of carbonic acid (the miner and well-digger call
it choke-damp) would render our wTorld unfit for animal
life. It does not accumulate in our atmosphere, because
every plant is busily, yet silently, absorbing it, and under
the stimulating influence of light and heat, selecting,
so to speak, the carbon ; while the remaining ingre-
dient, the oxygen, is given out for the behoof of animal
life.
The earliest traces of terrestrial plants were about co-
eval with the first appearance of vertebrate life in the
form of fishes. These ancient land plants were the fore-
runners of a vegetation which gradually advanced in
richness to the carboniferous epoch. The fragmentary
samples preserved in the upper Ludlow rocks appear to
348 PREPARATIONS FOR MAN.
have been of the club-moss family, (Lycopodiacea3.)a
They ushered in the Flora of the succeeding or Devo-
nian epoch, richer than its predecessor, but of minor
importance when compared with that garb and stature
which characterized the rich vegetation of the carboni-
ferous period. The flora of the coal formation must have
equalled, perhaps even far excelled, the most luxuriant
vegetation of tropical lands at the present clay. Dense
forests of tall Sigillarias, with their scar-marked and
fluted stems ; furrowed and jointed Calamites ; giant pines,
allied to our Eutassa and Araucaria, with an undergrowth
of graceful ferns, the delicacy of whose forms cannot be
excelled by any of the fern beauties of our own day ; these
constituted some of the principal features of a Flora which
has left us abundant and imperishable records of its
character, and has enriched our country with its valuable
relics in the shape of coal.
We have the high authority of M. Brongniart for the
belief that carbonic acid was far more abundant in the
air during certain epochs than it is at present. The at-
mosphere of the Pala30zoic period was warm, moist, and
highly charged with the gas mentioned. These condi-
tions have been shewn by Professor Daubeny to be pecu-
liarly favourable to the development of a rank vegetation,f
such as certainly prevailed during the later Palaeozoic
ages. We cannot doubt that the plants composing the
ancient Flora required supplies of carbonic acid for their
full development. In the vegetables of the carboni-
ferous epoch we can recognise the existence of agents
destined to perform an important part in the economy
of those days. While able to obtain abundance of
necessary pabulum to build up their organs and add
* Murchison's Stluria, p. 238.
fr Professor Daubeny, in Proceedings of British Association. •
PREPARATIONS FOR MAN. 349
to their carbonaceous ingredients, they were, at the
same time, preparing the way for the advent of animals
by subtracting the excess of a gas noxious to animal
life*
We have reason to believe that living beings must
have required supplies of pure air, in whatever epoch they
lived. Only a few vertebrate remains, those of reptiles,
have been found in the rocks of the carboniferous era.
Animals of this class are capable of surviving under less
favourable conditions of atmosphere than birds or mam-
mals. These facts are adduced by geologistsf as at least
some confirmation of the theory they hold regarding the
atmospheric features of the coal epoch. At all events,
there is something more than mere accidental coincidence
in all such relations of organized beings at the time al-
luded to ; we could scarcely expect aerial breathers to
abound at a period when the air was charged with choke-
damp.
But the plants of the epoch succeeding that of the De-
vonian, were undoubtedly the sources whence supplies of
coal have been derived, and deposited in the storehouses
of the earth for the benefit of mankind.
It has been calculated that the coal of Great Britain
alone, counting only what comes to the surface, contains
carbon enough to add one-fifth to that now existing in
the whole atmosphere. The plants of the coal period
have left us obvious proofs of their existence in the shale
of our coal mines ; and in the coal itself, we can find
some data from which to estimate the vast amount of
carbonic acid which (from whatever source derived) was
abstracted by the plants of that remarkable epoch, and
then stored up in a convenient form for our use. In our
coal fields, we have a rich deposit of material available
* Phillip's Manual of Geology p. 613; 1S55. t Ibid.
350 PREPARATIONS FOR MAN.
for various purposes, and among others, as a generator of
mechanical force — in fact, a latent power — which, in its
proper application with its usually associated ironstone
and limestone, has contributed largely to give to the pre-
sent age " its form and pressure."
All this invaluable treasure — in the form of deeply-
seated coal basins — must, however, have 'remained buried
for ever in the bowels of the earth, beyond our reach,
had it retained its primary geological conditions and re-
lations. But, by various and gigantic appliances of sub-
terranean force, the original deposits of coal have been
broken up, changed in position, and brought nearer the
surface, and thus placed more within the reach of man.
Upheavings have followed the internal throes of mother
earth ; elevating forces have brought about dislocations
of strata ; and if the deposits of subterranean fuel have
not in every case been brought to the very surface, those
still remaining in a buried condition have had their ori-
ginal relations so altered that they can be subjected to
the explorations of the adventurous miner with greater
prospect of success.
The regularity in the succession of different strata, and
the peculiarity of the organic remains by which these are
characterized and become capable of recognition, afford
man important assistance in his search after the coal which
lies under his feet. He has some more trustworthy guide
than the divining rod, to indicate the locality in which
he may expect the subterranean treasures, as well as cal-
culate their probable extent. A knowledge of geology
affords invaluable aid in the discovery of coal where it
has never previously been wrought, and we may trust to
the same sure guide in forbidding the waste of time and
capital in searching for it where it cannot possibly be
found. But when infallible marks have brought us to
PREPARATIONS FOR MAN. 351
the spot where the coal lies, the same principles aid us
in getting at it, or (as the miners say) " winning it."
Geology affords a sure guide in the development of those
commercial and industrial resources which the different
regions of the earth possess.
The carboniferous epoch, constituting the " reign of the
Acrogens," whose characteristic features have been briefly
alluded to, was followed by times when Pines and Cycases
prevailed — the " reign of the Grymnosperrns." The pines
had the predominance in the dawn of that kingdom ; they
were finally almost entirely supplanted by the Cycadeas —
a family probably familiar to our readers in one of their
aspects, under a form which maybe compared to a gigan-
tic fir-cone, with a tuft of wiry, fern-like leaves growing
from the top.
But we pass on to a new epoch of terrestrial vegetation.
With the Chalk period there began a fresh era, the " king-
dom of the Angiosperms." In its dawn there was a kind
of transition from the previous dynasty to the new one
which we are now to consider, and which reached the zen-
ith of its strength in our own time. Through the Eocene,
Miocene, and Pliocene epochs, the Angiosperms increased
in numbers, and some families which characterize our own
days became more and more numerous.
We have, in a previous part of this work, endeavoured
to shew that those species which delight us by their forms
and colours are peculiarly characteristic of existing
Floras. And if we seek for more substantial properties,
we still miss, in the species of the primeval epochs, those
distinguished for their utility at the present day. Doubt-
less the earth formerly yielded Ferns, Firs, Cycases, and
Palms, and plants of the same families to supply useful
products. The New Zealunder and Tasmanian derive
some sustenance from the subterranean stems of a fern ;
352 PREPARATIONS FOR MAN.
we ourselves owe nmcli to the firs of our own forests ; and
the natives of Northern Europe sometimes use the ground
bark of a pine (as well as of other trees) to eke out their
scanty meal ; some of the human family can, by a
troublesome process, extract nourishing matter from the
stems or seeds of a Cycas ; and certain Palms do furnish
valuable products — constituting, in fact, a vegetable ba-
zaar, yielding food and clothing, and luxuries besides ;
but how small a part, after all, of the millions of men in
our world do the foresaid plants support, and that part is
the least civilized and intellectual !
While it must be acknowledged that the researches of
the Paleontologist have not yet exhausted our informa-
tion as to the plants which clothed the earth before man
was called into being, wo cannot but remark the almost
total absence of families whose products minister to his
wants and comfort. We find few traces in the tertiary
epoch, which immediately preceded man's, of plants be-
longing to families from which he derives his necessary
food. In that tertiary age there were, so far as geology
reveals to us, few or no spices yielding " cinnamon, and
odours, and frankincense, and wine, and oil, and fine flour
and wheat. There are few evident indications of any
vegetables from which man derives food and valuable
fibre, and, in a word, of species which support and clothe
by far the largest proportion of the human race. Scarcely
any Grammes (grasses appear in the lists of extinct
forms ; may we not conclude that the principal cereal
plants are characteristic of man's epoch, that barley, oats,
rye, wheat, millet, Indian corn, and rice, were special pro-
visions in order to man's appearance ? From the lists
of Pliocene vegetation we miss the Labiate plants, which
so charm us by the beauty of their flowers, and which
yield essential oils to regale us by their perfumes. Of
PREPARATIONS FOR MAN. 353
Rosacea there are few traces, and in the list of finally-
added species, we must include the roses which yield us
their precious " Attar," and the delicious fruits which
characterize our more temperate climes.
There were thus long preparations made both in the
crust of the earth and in its living organisms for the sup-
port of man, who appears when all is prepared for him.
" The earth hath He given to the children of men."
"as for the earth, out of it cometh bread, and
under it is turned up, as it were, fire."
CHAPTER XII.
INORGANIC OBJECTS ON THE EARTH'S SURFACE.
SECT. I. CRYSTALLINE FORMS AND CHEMICAL PROPORTIONS.
In surveying, in a superficial manner, the mineral
substances which compose the crust of the earth, the
impression might be left that they are irregular through-
out, and that they can exhibit no marks of intelligence.
Paley, in the opening of his beautiful work, refers to a
stone, as, in this respect, in striking contrast to a watch.
The meaning of Paley is obvious and correct, but science
will not admit, in the present day, that there are no traces
of order and design in the stone. Isaac Barrow, two cen-
turies ago, spoke, in one of his Sermons, of stones, metals,
minerals, as " probably furnishing obvious proofs of the
Divine Wisdom, provided our senses were able to dis-
cover their constitution and texture." What the senses
cannot do has been done by the penetrating intellect of
man, following the principles of inductive science. In
dead minerals we observe — under a simpler, but, at the
same time, more unbending aspect — the commencement
of those wondrous forms which, under wider and more
accommodating modifications, play so important a part in
the economy of living beings.
No man can look on the columnar structure of Staffa
or the Giant's Causeway without having a sort of vague
AND CHEMICAL PROPORTIONS. 355
notion, that here there is method and intelligence too.
We feel as if we have traces of these in this architecture
of nature, just as we have them in those objects of rnan's
construction which they so much resemble — the columns
of a temple, the pipes of an organ, or the oaken ribs of a
ship. We are not sure that the impression is altogether
a mistaken one. There is here, it is true, no special
adaptation of parts to produce a useful end : this indi-
cation of design is altogether wanting ; but there is
general law operating here as in every other part of
inorganic nature, producing an orderly result, such as is
characteristic of intelligence. How often, too, may we
see the coal on our fires tending towards a definite ma-
thematical shape, and, on breaking up the pieces we find
the fragments not less regular ! When we open up the
stones of the ground, we may discover, in the component
parts of most of them, a regular structure, and the im-
pression is left on the mind that they are crystalline
throughout. In short, minerals are found everywhere
in nature in very beautiful symmetrical forms, and most
minerals assume regular forms in circumstances which
admit of this operation being formed — that is, when
there is a slow and gradual change of fluid into solid, and
the arrangement of the particles is undisturbed by motion.
It was a happy misfortune which befell Abbd Haiiy,
when a fine group of calcareous spar which he was ex-
amining fell from his hands and was shivered into frag-
ments. The original crystals were of a prismatic shape,
but as he gathered up the broken pieces of one of the
prisms in sadness, he observed that, while not less regu-
lar in shape than the original crystal, they were all
rhomboidal ; and the thought flashed on him — bright as
the lustre from the mineral — that all the varied crystal-
line forms in nature micdit be derived, according to fixed
356
CRYSTALLINE FORMS
laws, from a few primitive forms. He felt as if a new
world had opened upon him, and he exclaimed, " All is
found I" In prosecuting his investigations, he did not
scruple to break his whole collection of crystals to pieces,
and he succeeded in discovering certain laws, the deter-
mination of which has turned out to be a more precious
acquisition to human science than all the crystals in all
the museums of the world.
The regular forms assumed by minerals has now been
carefully examined, and the science of Crystallography is
the result. The figures of crystals found in nature and
formed artificially are very diversified, but any given sub-
stance is limited in the number of crystalline forms which
it takes : thus, fluor spar crystallizes in cubes, but never
in six-sided pyramids. The forms are all in conformity
with a beautiful law of symmetry. In every one of them
there will be found a line passing through the centre of
the crystal, round which all different parts of the crystal
are symmetrically grouped ; this is called the crystalline
axis. The numerous crystalline forms which exist in
nature can all be reduced, on rigidly scientific principles,
to a few primitive forms. There
are Six Primitive Forms, ac-
cording to Professor Weiss of
Berlin, whose views seem to be
generally adopted.*
First, The Octohcdral System.
It has three equal axes at right
angles to each other. (See
Fig. 69.) Owing to the perfect
symmetry by which it is cha-
racterized, it has been called the regular system of crystal ■
lization.
* The views of Professor Miller of Cambridge are substantially the same.
AND CHEMICAL PROPORTIONS.
357
Fig. 70.
Second, The Square Prismatic System. In this, as
in the former, there are three axes,
which intersect each other at right
angles ; but in this system only two
of the three are equal. (See Fig. 70.)
Third, The Bight Prismatic Sys-
tem. Here, as in the
two preceding, there are
two rectangular axes,
but no two of the axes
are equal. (See Fig.
71.)
In these three sys-
tems the axes of the crystals are all at
right angles to each other. In those which
follow, there is the same symmetry round
the axes, but the axes are not rectan-
gular.
Fourth, The Bhombohedral System. Here, as in the
first group, the axes are equal. They cross each other
at equal angles, but not at right angles.
(See Fig. 72.) The most simple form is
the rhombohedron,
which is bounded by
six equal and similar
rhombic faces.
The Oblique Prismatic
In this system, two of
the axes intersect each other ob-
liquely, while the third is perpen-
dicular to both. The axes are
unequal in length. (See Fig. 73.)
Sixth, The Double Oblique System. The three axes
intersect each other obliquely, and are unequal. Much
Fig. 71.
Fig. 72.
Fifth,
System.
Fig. 73.
358
CRYSTALLINE FORMS
Fig. 74.
of the symmetry of form observable in the others disap-
pears in this system. Still, the faces which are diagonally
opposed are parallel to each other. (See
Fig. 74.)
Out of these primary forms, other and
derivatory ones are fashioned, according
to principles which are of a mathema-
tical character. Thus, the forms which
the octohedral system takes are seven,
being limited to the number of ways in
which a plane can intersect the three
axes. The most prevalent are the octohedrbn, the cube,
and the rhombie dodecahedron, which are frequently met
with in nature.
Simple forms of different systems are never combined,*
but the mineral which assumes one form of a sj^stem
may, and often does, take other forms of the same system.
Thus the salt alum may be obtained in the form both of
the octohedron and the cube. During the process of
crystallization it will often happen that the faces of seve-
ral of these forms are simultaneously developed, furnishing
crystals of the greatest diversity of appearance. Thus,
in the crystallizatian of alum, the faces both of the cube
and the octohedron may be produced, and the feces of the
cube will be seen truncating the angles of the octohedron.
It is an instructive illustration of the means by which
infinite variety is produced in nature, in accordance with
a rigid unity. The transition from the more rigid crys-
tal to the freer forms of organic nature, may be seen in
the beautifully-ramified figures made by the frost on our
flag-stones and windows, and also in the branchings of
coralline structures.
* It should be stated, however, that the mineral may crystallize in a different system
when in an allolropic condition.
AND CHEMICAL PROPORTIONS. 359
An interesting connexion had been traced, by Mitscher-
fichof Berlin; between the crystalline form and the com-
position of bodies. Substances which take the same
crystalline form may be substituted for each other in
combination, without affecting the external character of
the compound. Thus, sulphate of potassa and peroxide
of iron may be made to take the form and aspect of alum
without the presence of any aluminous earth. Substances
which assume the same crystalline form are called Iso-
morphous. Isomorphous bodies have been distributed
into distinct groups ; — thus, oxygen, sulphur, chlorine,
belong to one group ; potassium, sodium, lithium, cal-
cium, zinc, lead, silver, are associated in another group ;
while arsenic, antimony, phosphorus, and tellurium, form
a third group. Bodies belonging to the same group may
be substituted for each other in the composition of salts,
or of minerals, without the external qualities of the bodies
being affected. Isomorphous bodies have often very
close points of resemblance in physical properties as well
as in form. Thus, arsenic and phosphorus have nearly
the same odour ; they both form gaseous compounds with
hydrogen ; they differ from nearly all other bodies in
their mode of combining with oxygen, and yet agree with
one another, and their salts are disposed to combine with
the same water of crystallization. Isomorphous sub-
stances, owing, doubtless, to the various points of analogy
which have thus been traced, crystallize together with
great readiness, and are separated from each other with
difficulty.0 These researches are not yet carried so far
as to entitle us to lay much weight on them in our pre-
sent argument, but, even at this stage, they furnish
glimpses of depths which have not yet been explored.
That which has only been imperfectly ascertained, points.
* Tomer's Chemistry, edited by Liebig and Gregory.
360 CRYSTALLINE FORMS
equally with that which has been more fully determined,
to designed connexions and parallelisms running through
the whole of nature.
When we go still farther down towards the very ele-
mentary constitution of bodies, we find indications of
what must be an order in respect either of form or
number. From a very early data there was a vague
impression that there must be something definite in the
way in which bodies chemically unite. But the law was
not scientifically evolved till within the last age, when
Dalton propounded his atomic theory. Proceeding on
the view commonly adopted in modern times, that mat-
ter is composed of atoms, he supposes that all atoms are
of the same form, that the atom of each element has a
specific weight, and that when bodies combine, it must
either be by one atom of one body with one atom of an-
other, or by one atom of the one element with two atoms,
three atoms, or four atoms of the other element. It was
thus that he gave expression to the law discovered by
him, and accounted for the relation between the weights
of the combining proportions of bodies. There has been
a difference of opinion as to the atomic theory which
Dalton employed to explain the laws of chemical combi-
nation, but there has been none as to the laws themselves,
In order to chemical combinations between bodies, we
must have a certain proportional weight of the one and a
certain proportional weight of the other, and if an excess
of either ingredient of the compound be present, it re-
mains uncombined, and with its properties unchanged.
Thus, in order to form water, it is necessary to have one
part by weight of hydrogen and eight parts of oxygen ;
and if there be a different proportion, say one part of
hydrogen and eleven parts of oxygen, then there will only
be eight of the oxygen absorbed in joining one of the
AND CHEMICAL PROPORTIONS. 361
hydrogen to make water, and three will remain free and
unchanged.
This, then, is the first part of the law of chemical
equivalents or definite proportions ; — bodies combine in
certain numerical proportions by weight, and in no others.
As the result of the united labours of Thomson, Berze-
lius, and a host of other chemists, the equivalent number
of the elementary bodies (about sixty in number) has
been approximately determined. To give a few ex-
amples : —
Hydrogen, 1. Chlorine, 35-4.
Oxygen, 8. Potassium, 39-2.
Carbon, 6. Copper, 3P8.
Nitrogen, 14. Lead, 103-8.
Sulphur, 16. Quicksilver, 100.
It is another part of the same law, following from that
which has been explained, that in entering into other
chemical compositions, the constituents of any chemical
compound replace each other exactly in the proportion in
which they combine. Thus it is found that one part of
hydrogen combines with eight of oxygen to form water,
with six of carbon to form carburetted hydrogen, and with
35*4 of chlorine to ■ form hydrochloric acid. But these
numbers express not only the proportions in which the
last-named bodies unite with hydrogen, but the propor-
tions in which they combine with each other. It follows
that if we know the proportion in which one body com-
bines with a number of others, we know also the propor-
tions in which all these bodies combine with each other,
and replace each other in new compositions.*
In order fully to understand this truth, which has re-
duced the science of chemistry to the most rigid law, it
is further to be taken into account, that when two bodies
* See Liebig's Letters on Chemistry.
362 CRYSTALLINE FORMS
combine with each other in two or more proportions, the
higher proportions bear a very simple ratio to the lower.
Thus there will unite with 14 parts of nitrogen the follow-
ing parts of oxygen, but no intermediate numbers : —
Nitrogen.
Oxygen.
14
8
Protoxide of nitrogen.
14
16
Deutoside of nitrogen,
14
24
Hyponitrous acid.
14
32
Nitrous acid.
14
40
Nitric acid.
It has been ascertained that a similar multiple relation,
capable of being numerically expressed, exists between
the proportions higher and lower in which all bodies com-
bine with each other.
In consequence of the discovery of these great truths,
which constitute the fundamental laws of chemistry, it
has been found possible to employ symbolical language
in that science, so as to enable chemists to express in the
simplest manner the constitution of every compound body
and indicate the way in which its elements maybe replaced.
Each elementary substance is designated by the first
letter of its name, compounds by the combination of the
initial letters of their elements, and the number of simple
equivalents of each element by their attached numbers.
The memory which would otherwise be burdened by the
number of particulars, (described by Plato as infinite,)
is able by their being thus bound into bundles, to use a
phrase of Locke's, and happily labelled, to bear its know-
ledge about with it, and the whole doctrine of the com-
position of bodies becomes comprehensible by the human
intellect. But chemists in setting forth their own dis-
coveries, and in giving due praise to one another, are
apt to forget that they have been able to accomplish their
work through the simplicity and numerical regularity of
AND CHEMICAL PEOPORTIONS. 363
the laws operating in nature. Human science is possible
because there has been the strictest attention paid to or-
der in the objects which it could arrange and classify.
It is also worthy of being mentioned that the volumes
of compound gases always stand in a very simple ratio to
the volumes of the elements thus : —
50 oxygen -f- 100 nitrogen yield 100 protoxide of nitrogen.
100 oxygen -f- 100 nitrogen " 200 binoxide of nitrogen.
50 oxygen -f- 100 hydrogen " 100 water.
100 nitrogen -f- 300 hydrogen " 200 ammonia.
Some curious discoveries seem to have been made in re-
gard to the connexion between chemical equivalents, and
volumes ; but they are not so perfected as to allow of their
introduction in such a treatise as this.
Attempts have been made in the same science to form
bodies into groups or congeners. M. Dumas, in particu-
lar, has detected a number of triads, or series of three
bodies, which have analogous properties, and showing a
singular numerical progression in their equivalent weights;
the equivalents of two of these added together, and divided
by two, giving approximately the equivalent of the third
thus : —
Chlorine, 35 J Potassium, 40
Bromine, >- 80 Sodium, \- 24
Iodine, 125 ) Lithium, 1
Calcium, 20 ) Sulphur, 16
Strontium, V 44 Silenium, V 40
Barium, 69 ) Tellurium, 64
" Regarding," says Faraday, " chlorine, bromine, and
iodine, as one triad, it will be seen that between the first
and the last there is recognisable a well-marked progres-
sion of qualities. Thus chlorine is a gas, under ordin-
364 CRYSTALLINE FORMS
ary temperatures and pressures ; bromine, a fluid ; and
iodine, a solid ; in this manner displaying a progression
in the difference of cohesive force. Again, chlorine is
yellow ; bromine, red ; iodine, black, or, in vapour, a red-
dish violet."*
In the higher chemistry of organized bodies we meet
with another kind of organic groups ; " these are named
Organic Types, the meaning of which is, that the atoms
are grouped together in a certain mode, on which the
properties or the compound so entirely depend, that pro-
vided this grouping or arrangement be retained, great
changes may be made in regard to individual elements,
without changing the general character of the compound.
This leads us to the very remarkable and important law
of substitution, which has become so fertile in discoveries
of late years."f In organic chemistry every compound
represents a type, and all chemical changes are substitu-
tions, but only like for like can be substituted, one metal
for another, or chlorine for iodine, &c.
In organic chemistry, the arrangement of the atoms
determines the character of the type. A certain arrange-
ment gives acids, another ethers, and so on. As an ex-
ample of such an organic type, Dr. Gregory gives the
case of Naphthaline C20 H3, the character of which is that
it is volatile and combustible. Now the hydrogen in this
compound may be replaced, atom by atom, by chlorine,
yielding a compound C2l) Cla, which still retains the gen-
eral characters of the type.
But there is another form of substitution giving rise to >
homologous series, in which hydrogen is replaced by cer-
tain compound radicals which are themselves homologous
and give origin, when substituted for hydrogen, to other
* Faraday's Lectures on Non-Metallic Elements, pp. 158, 159.
t Elementary Treatise of Chemistry, p. 265.
AND CHEMICAL PROPOKTIONS. 365
homologous series. The following tabular view will ren-
der this plain : —
Hydrogen H.
Water HO.
' Methyle, C2 H3
r
Oxide of Methyle
C2 H3 0
Ethyle, C4 H5
£
do. Ethyle,
C4 H5 0
It
Propyle, C6 H7
si
■j
do. Perpyle,
C6 H- 0
w
Butyle, C8 H9
m
do. Butyle,
C8 H9 0
^ Amyle, Ci0Hu
I
do. Amyle,
CioHnO
Taking methyle, the first in the series, we can see the
simple relation which exists between it and all the mem- .
bers of the series. The second, ethyle, is derived from
the first by adding two atoms of carbon, and two of hy-
drogen. The third bears a like relation to the second, and
so on throughout. The carbon equivalents form an even
number, those of the hydrogen are odd numbers. It is far-
ther worthy of notice, that the volatility of each is inversely
as the amount of carbon and hydrogen, and, consequently,
the density is in direct proportion to the amount of car-
bon and hydrogen. The density and boiling point in-
crease from the top to the bottom of the scale, in the
order in which they stand in the above table. Methyle
is a gas like hydrogen, requiring twenty atmospheres to
reduce it to a fluid state, amyle is an oily fluid boiling
at 311° Fahr.
The radicles in the first part of the table are all homo-
logous with, and analogous to, hydrogen. And as hydro-
gen, H, was the starting point in the series of radicles,
(Ethyles,) so water HO is the starting-point of a new ho-
mologous series formed from these radicles, forming
ethers, as represented in the second series in the table.
The first of these, oxide of methyle, Ca Ha 0, is a gas at
ordinary temperatures, the others are liquids less volatile
than ether, and so on.
From this second series a third homologous series is
366 CRYSTALLINE FORMS
formed, viz., the alcohols, by the addition of two equiva-
lents of water ; one example may suffice : C2 H, 0, HO
give methylic alcohol, &c. It may be observed that these
series, ethers and alcohols, are also analogous as well
as homologous, that is, their general characters are the
same.
" We can now see," says Dr. Gregory, " that the pro-
gress of science must inevitably reduce the whole of or-
ganic chemistry, in which we must remember only the
same three or four elements are perpetually met with,
to a collection of homologous series, in which every com-
pound will have its natural place, indicative at once of
its origin, its immediate derivation, and its properties
both physical and chemical."*
It is not necessary to maintain that all the laws re-
ferred to in this section, or in any of the sections, are
simple and original ; it is not necessary that we should
regard any one of them as being so. We are at liberty
to suppose that the very law of gravitation itself is de-
rived from a simpler law, as is maintained by some in
our day ; still the order in the derivative law would be a
proof of order in the original law itself, and in the ar-
rangements made in order to its operations ; thus, upon
the discovery of the law of gravitation, the laws of Kep-
ler were accounted for, but by a law orderly in itself, and
having beautiful arrangements made in order to its bene-
ficent action. Most of the forms of crystals found in
nature are derivative, but when we go back to the origi-
nal forms, we find them, like the derivative, distinguished
by the most methodical symmetry. On the same prin-
ciple we may argue that should the laws at present
acknowledged in science be resolved into simpler ones,
it would still be found that the original laws, with the
* Gregory's Elementary Treatise on Chemistry, pp. 264, 269, 272.
AND CHEMICAL PROPORTIONS. 367
adjustments made in order to their operation, are of a
regular and mutually adaptive character. The forms of
crystals, and the relations of chemical equivalents, if not
simple, must, just because they are regular, proceed from
forms or from forces, one or both, which are also charac-
terized by regularity. From disorder there can flow onty
confusion ; order can proceed only from order.
SECT. II. ADAPTATIONS OP INORGANIC OBJECTS TO ANIMALS
AND PLANTS.
Many of the adjustments which might be adduced
under this head are so obvious that it is not necessary to
dilate on them ; indeed, they can scarcely be made more
impressive by any scientific treatment. While the ele-
ments of nature obey their own methodical laws, they are
so arranged as to form living organisms, and supply
them with the needful sustenance. Each agent has its
rule of action, but is made to co-operate with every other.
Law is suited to law, property fits into property, colloca-
tion is adapted to collocation, and the result is harmony
and beneficence. The whole is dependent on every one
of its parts, and the parts all lend their aid to the pro-
duction of the whole. A break in a single thread of the
complicated network would occasion the failure of the
whole design.
There are upwards of sixty substances, which, in our
present state of knowledge, we must regard as uncom-
pounded. Each of these has its own properties, and the
system is sustained by the joint action of all. Very pos-
sibly the absence of any one of the elements, certainly
the absence of any one of the thirteen more universally
diffused, would throw the mundane system into confusion.
Each has a purpose to serve which could be served by no
368 ADAPTATION OF INORGANIC OBJECTS
other. Oxygen, so essential to animal breath and life,
is the most largely distributed of them all, composing
more than one half of the whole inorganic objects known
to us. Hydrogen, the other element of water, no less
necessary to living bsings, seems to have a relation to
every living organism. Carbon is a main source to us
of artificial light and heat. In order that it should fulfil
this end, it is necessary that it should be a solid while
evolving its light and heat, (a gas has little, and this
only a momentary, power of illumination) ; this is pro-
vided for by carbon being in itself always solid. But if
the result of combustion had been also a solid, then the
world would have been buried in its own ashes ; this evil
is avoided by the carbon going off in carbonic acid, which
is volatile. The mass is all glowing one instant, the next
it is dissipated into air. " Carbon," says Faraday, "posses-
ses every quality to render it adapted to its intended uses;
not one property, however seemingly unimportant, could
"be added or taken away without destroying the whole
harmonious scheme of nature, devised with such wisdom,
maintained with such care."*
Each of the powers and elements of nature is in itself
potent, and capable of working destructive effects, but is
checked and balanced by nice adjustments. What tre-
mendous energies does oxygen display in the phenomena
of combustion, and when in the condition of ozone ; yet
how tranquil and passive as one of the elements of water,
and as locked up in so many of the constituents of the
earth's crust. The electric force held in balance in a
single drop of water would, if let loose, exceed in energy
the electricity of a thunder-storm. Man is placed in a
state of things in which, as he is dependent, he is made
every instant to feel his dependence.
* Letters on Non-Metallic Elements, p. 2TT.
TO ANIMALS AND PLANTS. 369
What a vast number of independent agencies must
combine and co-operate in order to the life of organized
beings ! It is wrong to talk of an organism developing
itself by its simple and independent energy. Whatever
be its internal nature — in which also, in our opinion,
there is complexity and combination — it requires exter-
nal agents in exact adaptation to it. All plants need
nourishment, and this is supplied by inorganic matter ;
all animals need nourishment, and can be nourished only
by matter that has been organized, and this is furnished
directly or indirectly by the plant. How beautiful that
adjustment by which animals breathe of the oxygen of
the atmosphere, and set carbonic acid free for the use of
plants, while plants absorb carbonic acid, and set oxygen
free for the benefit of animals ! Then all animated beings
need moisture, which depends on the chemical laws unit-
ing oxygen and hydrogen to form water, and also on heat
to retain it in a state of vapour in the air, and on certain
adjusted relations, in respect of quantity and weight, to
the atmosphere in which it floats. All organized beings,
too, depend on light coming in the needed proportion
from a distant body, and on heat, the measure of which
depends on the state of the central part of the earth, on
the radiations of the sun, and on the temperature of the
regions of space. A considerable change in any one of
these essential conditions would be fatal to the whole an-
imated beings on the earth's surface.
But instead of dwelling on these familiar topics, we
shall turn to, perhaps not so conclusive, but still to a less
known set of facts, in which it has been supposed that
disorder reigns.
We have, in a previous chapter, brought forward some
evidences of adaptation in the march of events which
preceded man's epoch, and which have given rise to im-
10*
370 ADAPTATION OF INORGANIC OBJECTS
portant changes on the earth's surface*, to fit it as the
dwelling-place of animals and plants, and apparently ef-
fected with a view more especially to the advent of man.
In the development of this scheme, a suitable vegetation
was called into being, animal tribes were introduced, with
the command to multiply, and finally, to man was com-
mitted a power over every living thing.
Our aim, in the present section, is to show that there
are traces of fitness in the general aspect of the earth's
contour, in the arrangement of its dry land and waters,
and in the relations of its surface to temperature and
moisture ; and that these, in turn, have some connexion,
more or less evident, with the distribution of animal and
vegetable life, and also with the wellbeing of the human
family.
The study of Physical Geography, which has of late
years come into prominence, has little or no reference to
those arbitrary divisions of the world which occupy the
attention of the mere geographer. In examining the
structure of the earth's surface physically, attention is
rather directed to the valleys and elevations which diver-
sify its surface — those furrows drawn by the hand of
time, and the mountains which, by their upheaval, have
so remarkably diversified it, and indirectly have such im-
portant bearing on the existence and wellbeing of ani-
mals and plants. Those deep furrows and prominent
ridges, constituting so remarkable a feature of the earth,
are lasting records of the great changes to which it has
been subjected : we cannot suppose them to have been
fixed by mere chance ; they bear distinct traces of sub-
jection to those great principles which regulate all the
plans of Him, every part of whose works is adajDted to
every other.
The investigations of observers in different aires have
TO ANIMALS AND PLANTS. 371
established the following leading truths in regard to this
subject.
Land predominates in the northern hemisphere, water
in the southern ; the lands comprising the old and new
worlds stand at right angles to each other ; the new
world is perpendicular to the equator, the old parallel to
it. In reference to the contour of the dry land, it has
been observed, that the southern ends of the old and new
worlds terminate in a point, while they widen toward the
north ; that the southern points are high and rocky;
that the continents present, to the east of their southern
extremities, a large island or group of islands ; and that
each continent has a large gulf to the west. Humboldt
has indicated the parallelism of the two sides of the
Atlantic ; the projecting parts of the one correspond to
the gulfs of the other. Steffens has remarked, that not
only do the great continents expand towards the north,
and become narrower toward the south, but that the
same is true of their peninsulas also. He speaks, like-
wise, of the grouping of masses of land two and two
together, and points out an isthmus or chain of islands
uniting them.
G-uyot, in his " Earth and Man," enunciates the follow-
ing great laws, which apply to all continents in regard
to their relief or elevation : — All increase gradually in
height from the shore to the interior ; in all the conti-
nents the maximum of elevation is not in the centre —
hence there are two slopes of unequal length, and in the
mean, one of these slopes is always at least four or five
times greater than the other ; and the height of the
plains and of the table-lands increases at the same time
with the absolute elevation of the mountains. In the
old world, though the principal slope is toward the north,
we still observe a gradual decrease of the reliefs from
372 ADAPTATION OF INORGANIC OBJECTS
east to west : in the same manner, in the new world,
while the principal slope is from the west to the east, it
can be shewn that the reliefs go on gradually increasing
from, north to south, as in the old world.* Generally
speaking, although the mountains increase in elevation
from the poles to the tropical regions, the greatest heights
are not exactly at the equator ; in the old world they
occupy the vicinity of the tropic of Cancer, and in
the new, are near the tropic of Capricorn. To use the
words of Guyot, — "A great law, a general law unites
all the various systems of mountains and of table-lands
which cover the surface of our globe, and arranges them
in a vast and regular system of slopes and counter-
slopes."
From all this it is evident that the position of the
great masses of land, the forms of their coasts, the situa-
tions and relations of their mountains, table-lands, and
plains, have not been left to chance. A casual glance at
a map, or a cursory examination of an individual country,
may leave the impression that there is a want of definite
order, that all is in inextricable confusion ; but careful
examination of the entire wide surface of our globe, and
of the relations of its various parts, conclusively demon-
strates that He who commanded the dry land to appear,
accomplished His purpose according to a predetermined
plan, the issues of which must have been foreseen by Him,
even as they can now be seen by us. An inquiry into the
special modifications, in their relations to climate and the
distribution of living objects, enables us to see what fatal
consequences must have resulted, so far as the present
economy of things is concerned, if the plan and modifica-
tions had been different.
A water surface is slowly heated, and the consequent
* Guyot, Earth and Man, p 50
TO ANIMALS AND PLANTS. 373
evaporation produced has an additional retarding effect ;
such surface is also slowly cooled by radiation. A land
surface, on the other hand, becomes rapidly heated, and
as rapidly parts with its heat. Change of temperature
in water occasions a change of position in its particles ,
no such effect is produced on the land surface. The
sun's rays are weakened in their passage through the
atmosphere, owing to the presence of clouds and mists,
and the increased density of the lower strata ; the por-
tion of that medium nearest the earth, however, receives
its temperature principally through radiation from the
soil. The variety of surface, whether in respect of
smoothness or irregularity, elevation or depression, water
or dry land, necessarily occasions also a corresponding
difference in the amount of heat received by different
countries ; from which it appears that terrestrial as well
as atmospheric conditions modify the distribution of the
heat which we derive from the great central luminary of
our system.
The processes of heating and those of cooling are
slower and less sensible on water than on land, and the
portions of air in contact with these surfaces respectively,
are affected by the peculiarities of each : over the former
the atmosphere contains more moisture, and is of more
uniform temperature, than over the latter. Lands far
from the influence of the sea have great extremes of heat
and cold, whereas maritime districts have a more uniform
temperature throughout the year. The division into
torrid, temperate, and polar zones, though generally ap-
plicable as regards the climates of our earth, is greatly
modified by local configurations of surface, so that there
are no exact lines of demarcation separating the torrid
from the temperate zone. In the words of Humboldt,
" The temperature is raised by the proximity of a western
374 ADAPTATION OF INORGANIC OBJECTS
coast in the temperate zones ; by the divided configura-
tion of a continent into peninsulas with deeply-indented
bays and inland seas ; the prevalence of southerly or
westerly winds ; chains of mountains acting as protect-
ing walls against winds coming from colder regions ;
the vicinity of the ocean current, and the serenity of the
sky in summer : it is lowered by elevation above the seas,
when not forming part of a plain ; the compact configura-
tion of a continent having no littoral curvatures nor bays;
the vicinity of isolated peaks ; mountain chains whose
mural form and direction impede the access of warm
winds ; and a cloudy summer sky, which weakens the ef-
fects of the solar rays."
That comparative sameness which would result from
uniformity of surface, exposed to regular amount of solar
radiation at different seasons of the varying year, is coun-
teracted by special modifications of our dry land and ocean;
and hence the variety of climate, and corresponding diver-
sity in the vegetation clothing the earth, and in the living
beings that people it.
The atmosphere near the earth's surface, or in contact
with it, is, as a whole, much warmer in the vicinity of
the equator than at the extreme polar regions. This
hotter and higher air has a tendency to ascend, and the
colder and heavier, an equal tendency to rush in from all
sides and supply the place of the former. To such dif-
ferences of temperature may generally be referred all
those atmospheric currents which constitute the different
winds.* The direction of these currents towards the
equator might be uniform if the earth did not rotate,
and if its surface were level. The currents from the
vicinity of the pole have little rotatory motion, but in
* For some ingenious speculations on this subject, we would refer to Lieut. Maury'i
recent and excellent work on the Physical Geography of the Sea.
TO ANIMALS AND PLANTS. 375
their progress toward the equator they reach successively
portions of the earth's surface, which revolve more and
more rapidly, and thus leave them behind, (if we may so
speak,) and then they appear to blow in a direction con-
trary to that of the earth's rotation. Hence the cur-
rent from the north becomes converted into a north-east,
and that from the south into a south-east wind. Such is
the origin of those regular winds called the Trades.
The land and sea breezes of warm climates depend on
the same general cause ; the cold air from the sea during
the day flows in to supply the place of the air which is
more rapidly heated over the land, (and consequently
ascends,) the converse happening at night.
The aerial currents spoken of, and their more regular
modifications, also exercise a greater or less influence upon
the ocean surface, giving rise to interchanges in its parts.
There is a similarity in the effects produced by heat upon
the sea, to those produced by the same agent on the aerial
ocean. The warmer waters of the equatorial sea have a
tendency to flow towards the poles, the colder and heavier
portions forming an under current toward the equator.
Differences in the amount of saline matter, occasioned by
excessive evaporation at certain points, or by the influx
of large rivers, producing differences in the specific gravity
of the ocean water,* necessarily also give rise to currents.
Whatever power, however, sets the water in motion, the
direction of the current is variously modified by the con-
tour of the land.
Moisture is being continually evaporated in an invi-
sible state, and mixes with the atmosphere ; an abso-
lutely dry air is therefore almost an impossible occur-
rence, though there may be endless modifications in the
amount of moisture depending on various causes. The
• We would again refer to Lieut. Maury's work for details on this interesting subject,
376 ADAPTATION OF INORGANIC OBJECTS
quantity differs according to elevation above the earth's
surface ; the diminished density of the air upwards is
accompanied also with decrease of the absolute amount
of vapour of water contained in it. In the vicinity of
the equator the suspended vapour is abundant, owing to
the excessive heat, and the extent of water surface ; it
diminishes toward the poles ; it is generally greatest over
the open sea, and decreases from the coast to the interior
of continents.
From this brief account some idea may be entertained
of the remarkable relations between the gonial beams of
the sun reaching us from a distant point in space, and the
atmosphere, dry land, and water of our world. These,
again, have a connexion with the distribution and well-
being of the animals and plants which have been distri-
buted over its surface with bountiful hand.
We may now briefly examine some of the consequences
of the present arrangements of the earth's surface. Not
the least remarkable of these is determined by the gene-
ral position of the highest elevations. The concentration
and grouping of the high and extensive mountain sys-
tems towards the equator tend to reduce the temperature
of that region ; and the great extent of its water-surface
contributes to the same effect. If the equatorial surface
had been all land, and that land all plain, it is obvious,
for reasons already stated, that the whole of that part
would have presented the character of a parched desert,
and, in reference to animal and vegetable life, would have
been a dead waste. One of the agents necessary to the
development of living organisms, namely, heat, would have
been supplied in excess, and another, no less essential,
namely, moisture, would have been withheld. By the
complicated, but nicely adjusted, arrangements we have
already described, the present constitution of the earth's
TO ANIMALS AND PLANTS. 377
surface determines enough of the agents in question for
the wellbeing of vegetable and animal life.
If the elevated mountain ranges had been all grouped
toward the higher latitudes, eternal snows and ice would
have debarred animals and plants from a large extent of
surface at present occupied by both. A combination of
all these arrangements, namely, extensive flat land at the
equator, and high land at the poles, would have neces-
sarily limited the range of living forms, which must have
been chiefly confined to a comparatively narrow zone be-
tween the two extremes.
Even when we consider the present aspect of limited
portions of the earth's surface, we are struck with the
beauty of the adaptations. Mixtures of two masses of
air of different temperatures, and with dissimilar amount
of moisture, will occasion condensation of that moisture
in the form of mist or rain. The ascent of the hot and
humid air of the equator brings it in contact with strata
lower in temperature, and condensation of moisture is the
result. The same may happen when it moves on as an
overflowing current towards the north and south. Nor
is it to be forgotten that there is another condensing
agent ; we refer to land of high elevation. Moist winds
meeting with such an obstacle to their flow, have their
onward progress arrested, and their horizontal changed
into an ascending course ; the consequence is, that the
mass of air, becoming cooled by contact with other and
colder air, and with the land surface, loses its power of
retaining the same amount of vapour, and condensation
of moisture is the result. In the words of Guyot, — " The
mountain chains are the great condensers, placed here
and there along the continents, to rob the winds of their
treasures, to serve as reservoirs for the rain waters, and
to distribute them afterwards, as they are needed, over
6iiS ADAPTATION OF INORGANIC OBJECTS
the surrounding plains. Their wet and cloudy summits
seem to be untiringly occupied with this important work.
From their sides flow numberless torrents and rivers,
carrying in all directions wealth and life."*
In the new world, the chain of the Andes — its "great
backbone" — is situated not far from the western border ;
to the east of this vast range are extensive plains, with
interspersed secondary mountain ranges ; and this pecu-
liarity of conformation has a most important and neces-
sary relation to its climatic peculiarities. The trade-
winds from the Atlantic, in their progress first reach the
eastern slope, where the secondary chains of mountains
condense part of the moisture in refreshing showers, and,
finally coming in contact with the great and elevated
principal range, the air is robbed of most of the vapour
which remains. Hence a continual flow of water down
the eastern slope, clothing that fertile region with the
richest vegetation, and giving it the largest river sys-
tems in the world. A necessary result of this influence
exerted on the moist trade-wind in its progress to the
west, is, that by the time it reaches the western side of
the Andes nearly all its moisture has been lost, and a
line of coast on the Pacific presents the character of an
arid desert. The extent, however, of this region of
draught is very small, compared with that which profits
at its expense. The advantage derived from the arrange-
ment on the one side of the Andes, far more than com-
pensates for the disadvantage, and then this latter is still
farther lessened by local peculiarities, for the Chilian de-
sert would have presented greater latitudinal extent, if
the Cordilleras toward the north had been higher, or the
continent of greater breadth.
Imagine a different arrangement of surface ; the great
* Guyot's Earth and Man, p. 144.
TO ANIMALS AND PLANTS. 379
mountain chain, for example, transferred to the eastern,
instead of occupying the western side ; the consequence
would have been that the Atlantic trade-wind must
have had its progress arrested, and its vapour condensed,
at a comparatively early part of its course ; the ocean
giving up a portion of its waters to the passing wind,
would have received them hack again at no great dis-
tance in space, and after a short lapse of time ; no ex-
tensive river systems could have possibly existed as at
present ; — in a word, the whole influence of the genial
wind would have been lost, the descent to the west would
have been far more extensive, and the change in the
land surface, and the resulting effects on climatic pecu-
liarity, would have resulted in a very different distribu-
tion of organic forms, would have given rise to new fea-
tures in the zones of animal and vegetable life, and
changed the habitations of man, and the relation of one
part of mankind to another.
We may now direct our attention to another part of
this wide subject, to modifications which have respect
to the waters of the ocean and their currents. It has
been already stated that there is a tendency to a general
transference of the warmer equatorial waters to the north
and south, and of the colder polar waters towards the
equator, subject to modification in consequence of the
earth's rotation. Now the configuration of the land sur-
face determines peculiarities in the distribution of the
great currents, which exert no mean influence on the
distribution of organic forms. The great equatorial
current of the Atlantic has the largest mass of its waters
bifurcated by the projecting point of Cape San Roque,
one part being deflected to the south, along the coast of
Brazil, hence called the Brazil current ; while the re-
maining and largest passes into the Gulf of Mexico, and
380 ADAPTATION OF INOKGANIC OBJECTS
then issues from the north-east extremity of the same,
under the name of the Gulf Stream, and at a tempera-
ture exceeding 80° of Fahrenheit's thermometer. At its
exit from the narrow passage between the point of Florida
and the island of Cuba, and for some distance beyond,
its flow is comparatively rapid and northwards, till the
cold currents from the north, and the change in the con-
tour of the coast line, produce such an influence that its
direction becomes north-east.* Nevertheless it still re-
tains a high temperature ; in lat. 41° N. it is at 72*5° F.,
and 63-5° F. on the outer border of the stream. Its in-
fluence is admitted to extend to a large part ■ of north-
western Europe, and it bears with it evidence of its pre-
sence, and of the regions whence it flows, in the form of
tropical seeds and fruits, &c, which are stranded on the
shores bathed by its waters, and this even as far as North
Cape.
The effect of such a body of warm water (at Cape
Hatteras, Professor Bache found its temperature little
altered at a depth of 3000 feet) upon the distribution of
marine animals and plants might be expected ; but this
influence extends also to the lands along whose shores it
moves. The late Professor E. Forbes has shown its
effect as regards the distribution of animal forms on the
British coasts, the general Fauna of the German Ocean
being different from that of the Atlantic border-line.
This difference we have shown to be not less marked in
regard to marine vegetation ; certain species of sea-plants
abundant on the Devonshire coast, range also along the
Atlantic border as far as the Shetland Islands, while
most of them are wanting over a large proportion of the
* This explanation is not deemed sufficiently satisfactory by some, and we would re-
fer to the remarks on this subject, in Lieut. Maury's work already quoted. We have
chiefly to do with the course of the current.
TO ANIMALS AND PLANTS. 381
coasts washed by the German Ocean.* The general
mildness of the western coast of Britain, as compared
with the eastern, is mainly to be attributed to the com<
paratively warm water of the Atlantic. The influence
extends to the land vegetation of the continent ; the con-
sequence being that the line of cultivation extends nearly
to North Cape, and barley may be grown as far as 70° N.
latitude.
It appears that while there is a general plan regulating
the relations between our earth's surface and the influence
of the central luminary of our system, there are modifi-
cations affecting the more local distribution of heat and
moisture ; and these are associated with certain features
of organic life, inasmuch as there is a relation between
the amounts of the necessary agents and the constitution
of animals and plants. We cannot avoid coming to the
conclusion that there are indications — at least in our
hemisphere, that great centre whence civilization has ex-
tended— of suitable physical conditions, which were not
brought about by mere chance.
Knowing the connection which exists between the na-
ture of the surface, whether land or water, and the influ-
ence of the sun's rays on the temperature of our atmos-
phere, is is quite legitimate to speculate regarding altera-
tions of climate as related to changes of surface.
Those great revolutions which have taken place at dif-
ferent epochs of our earth's history, and the corresponding
phases which have occurred in animal and vegetable life,
are among the more interesting points which occupy the
attention, and are revealed by the investigations of the
geologist and of the palaeontologist. In man's compara-
tively brief period, such have not been distinctly exhibi-
ted on any great scale ; nevertheless with no inconsider-
* See Dr. Dickie's Taper in Proceedings of British Association for 1852
382 ADAPTATION OF INORGANIC OBJECTS
able degree of certainty, the physicist can show what
general climatic changes would follow the submergence
of a continent, and the increase of water surface, or the
converse. Farther, the. average height of any portion of
land above the level of the sea, can be shewn to exercise
a distinct influence on the climate of the region, and con-
sequently, on the beings which inhabit it. Sir Charles
Lyell in his " Principles of Geology," has shewn how the
numbers and distribution of animals and plants are
affected by changes in the physical geography of the
earth, and that these changes may also promote or retard
migrations of species, or alter the physical conditions of
the localities which they inhabit. " There are always,"
says he, " some peculiar and characteristic features in the
physical geography of each large division of the globe, and
on these peculiarities the state of animal and vegetable
life is dependent."
Mr. Hopkins, in his introductory address to the meet-
ing of the British Association at Hull, has very clearly
shewn the relation between the climate of northern
and western Europe, and the present configuration of the
American coast line, in reference to the direction of the
great Gulf Stream. He remarks — " It is to the enormous
mass of heated water thus poured into the colder seas of
our own latitudes, that we owe the temperate character
of our climate and not only do the maps of M. Dove
enable us to assert distinctly this general fact, but also
make an approximate calculation of the amount to which .
the temperature of these regions is thus affected. If a
change were to take place in the configuration of the
surface of the globe, so as to admit the passage of this
current directly into the Pacific, across the existing
Isthmus of Panama, or along the base of the Kocky
Mountains of North America into the North Sea — a
TO ANIMALS AND PLANTS. 383
change indefinitely small in comparison to those which
have heretofore taken place — our mountains, which now
present to us the ever-varying heauties of successive sea-
sons, would hecome the unvarying abodes of the glacier,
and regions of the snow-storm ; the cultivation of our soil
could be no longer maintained, and civilisation itself must
retreat before the invasion of such physical barbarism."
We are anxious not to stretch the argument beyond
what it can bear. Where the relations are so many and
complicated, we are not entitled to say that no other sys-
tem could have served the same ends ; but we think that
we can discover proof that there is a system. We can
see that certain changes unless counterbalanced by other
changes, would have been fatal to many of the animated
beings on the earth's surface. We can see, too, that the
present condition of the globe is, in fact, suited to the
existing distribution of organized beings ; and we know
of no means by which plants and brutes could have
adapted themselves to an essentially different state of the
earth. It is evident that every part is suited to every
other. " The mind/' says Lieut. Maury, " is delighted,
and the imagination charmed, by contemplating the
physical arrangements of the earth from such points of
view as this which we have now before us ; from it, the
sea, and the air, and the land appear each as a part of
that grand machinery upon which the wellbeing of all
inhabitants of earth, sea, and air, depend ; and which, in
their beautiful adaptations, afford new and striking evi-
dence that they all have their origin in one omniscient
idea, just as the different parts of a watch may be consi-
dered to have been constructed and arranged according to
one human design."
We fully acknowledge, in regard to man, that he is
capable of suiting himself to a variety of conditions and
384 ADAPTATION OF INORGANIC OBJECTS
circumstances, but, in this respect, he stands almost alone
in creation ; and we cannot view him apart from animals
and plants, for his existence is intimately linked to theirs.
His range in latitude is certainly very extensive, from
the snows of the Artie lands — where those outposts of
humanity, the Esquimaux, pass their lives between the
extremes of satiety and starvation — to the tropical zones,
where their swarthier brothers are exposed to the heat of
a meridian sun. But it is not to be forgotten that while
he can exist in such widely different circumstances, there
are certain terrestrial conditions necessary to the develop-
ment of his higher nature and qualities. u The distri-
bution of man," says Guyot, " over the surface of the
globe, and that of other organized beings, are not founded
on the same principle. There is a particular law which
presides over the distribution of the human races, and of
civilized communities, taken at their cradle in their in-
fancy ; a different law from that which governs the dis-
tribution of plants and animals. In the latter, the
degree of perfection of the type is proportional to the in-
tensity of heat, and of other agents which stimulate the
display of material life. The law is of physical order.
In man the degree of the perfection of the types is in
proportion to the degree of intellectual and moral im-
provement. The law is of moral order. . . . Here is
the reason that the Creator has placed the cradle of man-
kind in the midst of the continents of the north, so well
made, by their forms, by their structure, by their climate,
to stimulate and hasten individual development, and that
of human societies."
When God gave the earth to the children, He meant
it to be to them a source of something more than mere
sustenance. There are scenes spread all over its surface
which have delighted or roused the soul of man, and
TO ANIMALS AND PLANTS. 385
helped to shape his character and his history. The fer-
tile field, the pleasant dale, the murmuring rill, the
gently-flowing stream, the rugged mountain, the bold
headland, the thundering cataracts, these have all been
7 O 7
the means of soothing, of exciting, or awing the spirit of
man. The vegetable productions enhance and vary the
effect by the lightness and gracefulness of their forms
and harmony of their colours, by their tangled luxuriance
in our meadows and by our rivers' banks, or by the
sombreness of their hue and depth of shade which they
furnish. These aspects of nature have all had their in-
fluence in raising up new ideas and fresh feelings in
man's soul. The physical characters of a region, the na-
ture of its surface whether flat or hilly, its soil and
minerals, the size and flow of its rivers, the mountain
chains which cross it, and the bays of the sea which in-
dent it, the clearness or cloudiness of its atmosphere — all
these have moulded to some extent the psychical pecu-
liarities of man, and determined his tastes, his pursuits,
and his destiny.
And there are still higher views to be taken of human-
ity. " God hath made of one blood all nations of men for
to dwell on all the face of the earth, and hath determined
the times before aj)pointed, and the bounds of their habi-
tation." As the drama of our race's history is only being
acted, we cannot see the issue ; but we are convinced
that in this allotment there was a reference to the devel-
opment of man's mental faculties, and ultimately to his
moral and religious elevation.
We should leave a wrong impression if we did not
here state our belief that our earth, while adapted to man,
is adapted to him as a being fallen, frail and depraved.
Our earth had a paradise upon it only for a brief period,
and within a narrow range ; and, truly, an Eden would
386 ADAPTATION OF INORGANIC OBJECTS
not be suited to man with his present character. We
frankly acknowledge that we could not comprehend the
suitableness of many of the physical conditions and ac-
tions of our globe, of its waters and its vapours, if we
regarded man as a pure and holy being, who did not re-
quire to be restrained from evil by physical barriers, who
needed not suffering to punish and to purify. Our earth,
while it affords nourishment to man, yields it in such a
manner that man must toil for it, and, in toiling for it,
is kept from much sin. Physical geography announces,
as clearly as Scripture, that man must eat bread in the
sweat of his face. Not only so, our soil and atmosphere
have chilling damps, and unwholesome heats, and dele-
terious ingredients, which breed and cherish disease, and
help to bring man to the grave. These are essential
parts of the economy of things in which man is placed.
In short, our eartli was prepared for man as possessing
sinful inclinations, and needing to be exposed to suffering.
Let us add, that it has been prepared as the scene of the
action and passion of Him who must " needs suffer many
things," and who had to say, " the foxes have holes, and
the birds of the air have nests, but the Son of man hath
not where to lay his head."
But " unto us a child is born, unto us a son is given,"
and " this same shall comfort us concerning our work
and the toil of our hands." We are convinced that not
a few of the conditions of the earth have a reference,
more or less direct, to the diffusion of Christianity as the
only element fitted to regenerate our world. As the
leaven is only yet leavening the mass, we cannot discover
its full relations to the agents among which it is placed.
But as the past condition of the earth was an anticipa-
tion of the present, so the present points on to the future.
We do not believe that the present is the consummated
TO ANIMALS AND PLANTS. 387
state of the earth. Just as among the old geological
vertebrates, there were members which had not unfolded
all their capabilities, so, in our present earth, there are
agencies at work which have not completed their office.
A grand plan of prophecy is advancing both in the phy-
sical and moral world, and we live in the expectation of
a coming era, when the streams which have run for ages
alongside of each other will unite, and yield, at the same
time, a nobler condition of the earth's surface, and of the
spiritual character of its human inhabitants. " They
shall not labour in vain, nor bring forth for trouble/'
" Instead of the thorn shall come up the fir-tree, and in-
stead of the brier shall come up the myrtle-tree." "The
child shall die an hundred years old."
CHAPTER XIII.
THE HEAVENS.
SECT. I. ORDER IN THE MOVEMENTS OF THE HEAVENLY BODIES,
The ancients appealed with great confidence and evi-
dent delight, to the heavens, as fitted above almost
everything else, to prove that there is in nature, or above
it, a presiding Intelligence. The spectres of which they
stood in awe were either a grim fate or an unsteady
chance, and from these they felt that they could be most
readily delivered by the light which shone from the
heavenly bodies. The argument was perfectly conclu-
sive of the end proposed by those who advanced it, and
it was so, notwithstanding that they were not able to
shew whence the order to which they pointed proceeded.
They observed that the movements of the celestial bodies
were harmonious ; that there was, in consequence, a
beneficent succession of day and night, and of seed-time
and harvest, summer and winter, cold and hot ; that the
motions of the very stars, which they styled planetary or
wandering, were orderly — their apparent regularities
obeying a higher law of order ; that there was a cycle
for eclipses, whose return, therefore, could be predicted ; —
and they argued, we believe legitimately, that the " music
of the spheres" had been arranged as certainly as the
ORDER IN THE HEAVENLY BODIES. 389
concord which comes from a concert of musical instru-
ments. "We are justified in inferring that there has been
intelligence exercised in the production of the harmonies
of music, whether we are or are not able to shew how
the tones are produced ; and, on a like principle, we are
entitled to conclude that the harmony of the heavens does
not arise from the concurrences of chance, even when we
cannot unfold its nature with perfect accuracy. Theo-
logy has not been employing this argument so frequently
for the last age or two, but it is because the old spectre,
raised in the darkness of heathenism, has disappeared,
and it is now more terrified by another delusion, that of
pantheism, which has originated in the deception of the
eye when gazing on a brighter light. But the argument
drawn from the heavens is as conclusive as it ever was,
and can now be expounded more fully and satisfactorily.
We mean, in the brief survey which follows, to begin
with the Solar System, and thence rise to the region of
the Sidereal Heavens.
In all, about seventy planetary bodies — planets,
planetoids, and satellites — are moving round the sun,
or round each other in the most regular manner. Each
of them is of an oblate spheroid shape ; rotates round its
own axis ; moves in an elliptic orbit, with a sun or a
planet in one of the foci ; has a fixed length of day, that
is, time of rotation on its axis ; and a fixed length of
year, that is, time of making a revolution round its pri-
mary. The rotatory motions and the revolutionary mo-
tions of the planets round the sun, and of the primaries
round their secondaries, are all, with the exception of
those of the satellites of Uranus, in one and the same
direction, from west to east. All these bodies are held
in their spheres by a central force, of which Newton gave
us the proportional expression. These may not be the
390 ORDER IN THE MOVEMENTS
ultimate expression of the laws of nature, but thoy are
the obvious forms in which they present themselves to
human observation.
We have spoken of the orbits and movements of the
planets as being regular, but this is true only approxi-
mately. There are irregularities in them all, and appre-
hensions were at one time entertained that these might
go on increasing, till the whole system became hope-
lessly deranged. But it was shewn, by the eminent con-
tinental philosophers of last century, that all these are
periodical, or balanced one by another. The earth's
orbit, at this present time, is approaching nearer the
circular figure ; but it has been demonstrated, that after
a time it will become more elongated, leaving the length
of the year and the mean temperature of the earth un-
changed. The obliquity of the ecliptic — that is, the
inclination of the earth's axis to the plane of its orbit —
is at present lessening ; and as the seasons depend on
this obliquity, which allows the sun to shed his full ra-
diance on different portions of the earth at different
times, it was feared that they, and all on the earth which
depends on them, might be seriously affected by the
change; but it can be shewn that the obliquity will, in
course of time, begin to increase, and that the variation,
whether of increase or decrease, cannot sensibly affect
the seasons. The moon's mean motion has for some
time been increasing ; this is due to the diminishing
eccentricity of the earth's orbit ; but in the course of
time the eccentricity of the earth will begin to increase,
and the moon's mean motion to diminish. The planes
of the planetary orbits vary in their positions, but all the
variations are periodical, and can lead to no inconveni-
ence. When these questions were still unsettled, the
apprehensions of derangement arose chiefly from the
OF THE HEAVENLY BODIES. 391
perturbations of Jupiter and Saturn, each of which is as
largo as all the planetary bodies then discovered put to-
gether. Long and anxious calculations were instituted
on this subject by Lagrange and Laplace : these cannot
be detailed without the aid of the highest mathema-
tical analysis, but the result may be given. Laplace
found that " there existed in the motion of Saturn an
inequality, the period of which is 029 years, and in the
motion of Jupiter a corresponding inequality, which is
affected with a contrary sign, and whose period is nearly
the same — the difference between the two scarcely
amounting to a degree in a thousand years. This was
balm to the apprehensions of philosophers, for all fears
as to the probable disorganization of the frame of nature
evaporated, and the explanation of Laplace produced the
true fcioxaiiioTuatc, by which astronomers signified the
restitution of things to their former state."* It is thus
proven that, looking to the law of gravitation, and the
disposition of the various planetary bodies in reference to
the sun and each other, the solar system has a remark-
able principle of stability in the midst of constant
change.
Connected with the Solar System there is a still greater
number of comets. These used to be regarded even by
astronomers with feelings of alarm, as apparently disturb-
ing rather than harmonizing agents. Byron speaks of
"A pathless comet and a curse,
The menace of the universe."
The impression was that they appeared and disappeared
iu the most capricious manner, and that the earth might
regard itself as fortunate if it did not come within the
sweep of their tails, which at times spread themselves
* Smyth's Celestial Cycle, vol L, p. 284
392 , OEDEK IX THE MOVEMENTS.
through a space of 180,000,000 miles. But it has now
been demonstrated of some of them, and may be inferred
of all, that they obey laws as constant as those of the
planets themselves. They seem to consist of floating
vaporous matter through which the stars can easily be
seen. It has been ascertained that forty of them move
in elliptic orbits. Some of these are comparatively small,
being within the orbits of the known planets ; others
extend much farther into sjiace. Neptune revolves round
the sun at a distance thirty times that of the earth ; but
the great comet of 1680 moves in an orbit exceeding that
of Neptune nearly as much as it exceeds that of the
earth — the distance of the comet being 853 mean distances
of the earth. The period of the revolution of a number
of them has been ascertained, and the time of their re-
turn can be predicted. It should be added that a few of
them seem to move in hyperbolic curves, while a large
number are said to have curves sensibly parabolic.
Though we do not know the ends contemplated by these
wanderers into space, nor, indeed, by comets generally,
yet we know that they obey the same law as the planet-
ary bodies, and reasoning from analogy we may conclude
with Newton, that they carry with them, and dispense
through wide regions a beneficial influence.
We are now to pass on from the sun and planets to
the contemplation of the stars. The distances of some
of the nearest of these stars has been ascertained, and
shew us that in going from the outer planet to the nearest
body of the sidereal regions, we have leapt across an in-
conceivable void of twenty-one billions of miles. Others
are supposed to be so far distant, that light, which travels
from the sun in eight minutes, would require millions
or even thousands of millions of minutes to come from
them to our earth. It follows that the stars which we
OF THE HEAVENLY BODIES, 393
now see are stars as they existed many long ages ago.
There is thus opened to us a glimpse not only of
regions of space, but of periods of time stretching far
into infinity. The telescope shows within its range one
hundred millions of self-luminous bodies like our own
sun. These are collected in many cases into groups
with regular shapes, and, in not a few cases, are in
binary, or ternary, or multiple combination with each
other.
We can discover even by the naked eye that the stars
in some places are gathered into clusters. Thus six or
seven stars are seen by the naked eye as forming the
Pleiades. The telescope shows that in this constellation
there are nine or ten times the number of stars collected
together and separated from the rest of the heavens.
The number of such clusters is very great, and they may
be discovered by artificial glasses, here and there, over
the whole surface of the heavens — but more numerous
in some places than in others, more numerous in par-
ticular in the northern than in the southern hemisphere.
The stars in so many of these clusters are so many that
they cannot be counted ; but on a rough calculation it
would appear that many of them must contain ten or
twenty thousand stars, in an area not more than the tenth
part of the moon's apparent disc. Some of these groups
are of an irregular shape, which it is difficult to classify,
or even to describe ; but a large number of them assume
such regular forms, as to show that there is some princi-
ple of order or combination among them.
The most common form is stated by Sir J. Herschel
to be the circular, or the elliptic of various degrees of ec-
centricity, from moderately oval forms to ellipses so elon-
gated as to be almost linear.* Dr. Robinson, in describ-
+ Outlines of Astronomy.
394
ORDEIt IN THK MOVEMENTS
ing the discoveries made by Lord Bosse's telescope, says
they may be separated into three classes ; those which
Fig. 75.*
are round, of nearly uniform brightness ; those Avmch
are round, but appear to have
one or more nuclei ; and those
which are extended in ouo di-
rection, so as to become long
stripes or rays.f It should be
added, that although there can
be no doubt as to the regular
character of the forms assumed
by distant groups, yet as won-
derful changes are made in their
appearance by higher optical pow-
ers, we are not at liberty to
assume that we have ascertained their forms with per-
fect accuracy. Thus some of the nebulas which presented
Fig. 764
* Fig. 75. Cluster in Hercules. t Transactions of Royal Irish Academy, 184T.
% Fig. 70. Annular Ketmla in Lyra.
OF THE HEAVENLY BODIES. 395
the appearance of a spherical body to Sir John Herschel's
eighteen-inch reflector, have been transformed by Lord
Rosse's six-feet speculum into a luminous spiral of
unequal convolutions, which are prolonged at both ex-
tremities into granular globules. " Almost every new
observation appears to confirm the fact of that curious
tendency to spiral arrangement in these nebulous masses,
of which mention has been so frequently made."*
Sir John Herschel discovers in these aggregations of
stars the operations of physical laws. " Their round
form clearly indicates the existence of some general bond
of union of the nature of an attractive force, and in many
of them there is an evident acceleration in the rate of
condensation as we approach the centre which is not re-
ferable to a merely uniform distribution of equidistant
stars through a globular space, but marks an intrinsic
density in their state of aggregation greater in the centre
than at the surface of the mass."f The same distinguished
astronomer J regards it as a general law in the constitution
of extended nebulae, that their interior or brighter strata
are more nearly spherical than their exterior or fainter,
their ellipticity diminishing as we proceed from without
inwards, a character which he represents as favouring,
though not conclusively, " the idea of rotation on an axis,,
in the manner of a body whose component parts have such
an amount of mutual connexion as to admit of such a mode
of rotation, and of the exertion of some degree of pressure
one on another." Some of the late disclosures of Lord
Rosse's telescope, in regard to the prevalence of the spiral
form in nebular groups, may so far effect these specula-
tions, but in doing so they open to our view a more won-
derful harmony, the law of which has not been determined.
* President's address to British Association, 1853.
i Outlines of Astronomy, p. 593. J Observations at Cape, p. 8.
39G ORDER IN THE MOVEMENTS
The Milky Way, which spans our heavens so conspicu-
ously, is not a cluster of stars, but a succession of clusters.
Our sun is one of the stars composing this system, and
is supposed to be placed not far from the centre, but
nearer the one side than the other, and in one of the
poorer or almost vacant parts of its general mass. Sir
Fig. 77.*
W. Herschel thought he was able to number eighteen
million stars in this girdle, and his son speaks of it aa
consisting entirely of stars, scattered by millions like
glittering dust on the background of the general hea-
vens. That there is some sort of concentration in this
zone is evident from the statement of Struve, that there
are nearly thirty times as many stars in the centre of the
stratum as in the regions near the extremities.
On looking into the concave of the heavens, there are
perceived at unmeasurable distances, luminous masses
which look like fleecy clouds, and have been called ne-
bulee by astronomers. Upwards of two thousand of these
" world islands" have been discovered in the northern,
and upwards of a thousand in the southern hemisphere,
by the telescope employed by the Herschels. According
to Sir W. Herschel's estimate in 1811, they cover ojoth
part of the whole heavens. They were at one time sup-
* Fig. 77. Herschel's section of the Milky Way. The Milky Way appears more
brilliant in the direction ofF, of D, of B, than in that of E, C, and A.
OF THE HEAVENLY BODIES. 397
posed by certain speculators to consist of a sort of lumi-
nous matter, or star-dust, out of which worlds are being
made even now by general law. This supposition has
not been confirmed by later speculation. Within these
few years, not a few of these nebulas which were regarded
as being most certainly luminous vapours, have been
shewn to be stars. The magnificent telescope of Lord
Eosse, not long after it began to be used, shewed that
the great nebula in Orion, which was supposed to be one
of the most unresolvable of them all, consisted of clusters
of distinct stellar bodies. Since that time, nebula after
nebula has been resolved by Lord Eosse's telescope, and
another of less power but in a finer climate, at Cam-
bridge, in the United States. In 1850, Sir J. Herschel
was prepared to declare it as being almost certain, since
Lord Eosse's telescope had resolved, or rendered resolv-
able, multitudes of nebulae, that all the rest could be
resolved by a farther increase of optical power, and the
language might be made still stronger and more decisive,
in consequence of what has been accomplished by that
magnificent telescope since that date. The nebula? may
now be confidently regarded as clusters of stars, and
give evidence of order, combination, and law in the ex-
treme boundary of that sphere of immeasurable magni-
tude which constitutes the universe as knowable by us.
It is worthy of being mentioned, as illustrative of order
and law, that there are to be seen in the expanse of hea-
ven, in many places two or more stars which are appa-
rently near each other, and which have been shown to be
mutually connected as part of one system. It not unfre-
quently happens that a centre of light, which appears as
only one star to the naked eye, is turned into two or more
stars by a telescope of very ordinary power. Sometimes
the relation is merely optical, and not real, that is, stars
398 ORDER OF THE MOVEMENTS
at a great distance from each other may seem near, be-
cause though the one he far behind the other, they lie
nearly in the same line of vision to the eye. But the
number of double stars in the heavens, being about 6000
in all, is far too numerous to be referred to any such
cause. Among these, according to a table published in
1849, 650 are known in which a change of position
can be incontestably proved.* Besides it has been as-
certained in regard to considerably more than 100 double
stars, that they revolve about each other in regular or-
bits. In some cases there is a smaller star joined to a
large one, in other cases there are two or more stars of
Fig. 78.
nearly equal size revolving round a common centre of
gravity. The orbit in which these connected stars move
is ascertained to be elliptical. These phenomena lead
Sir J. Herschel unhesitatingly to declare the stars to be
subject to the same dynamical laws, and obedient to the
same power of gravitation, which govern our sys-
* Humboldt's Cosmos, vol. iii. p. 280, Otte's Translation ; additions being made every
year by the labours of Argelander, Sturve, &c.
t Fig. 78. Binary star, that is, two stars rovolving round a common centre.
OF THE HEAVENLY BODIES. 399
tern. The period of revolution of some of these combined
stars has been determined, and is found to vary in differ-
ent binary and multiple systems from 30 to upwards of
700 years. We have thus glimpses opened up to us in the
depths of the sky, not of planet revolving round sun, but
of sun moving round sun.* In the solar system we have
satellite rolling round planet, and planet around sun, and
double, triple, and multiple stars revolving round each
other, and thousands of millions of stars grouped together
in a common system.
In consequence of having ascertained, as is supposed,
the distance of some of these binary stars from the
earth, it is not difficult to calculate, with an approach to
certainty, what are the dimensions of their orbits. These
combined stars seem to be at a much greater distance
from each other than the farthest planet of our system is
from the sun. The distance of the two stars of 61 Cygni
from each other, is 44 times the distance of the sun from
the earth. The distance of these double and triple suns
from each other is thus greater than the distance of the
planets from the sun, in nearly as high a proportion as
the distance of the planets from the sun exceeds that of
the satellites from their primaries. All this gives the
appearance of a regulated order in the relative distance
of satellite from planet, of planet from sun, and of sun
from sun, so as to allow them to move freely, each in its
own sphere, whether a wider or narrower.
The region wrhich we have been surveying used to be
called that of the fixed stars ; but it has been shewn that
the language is inapplicable. Every star is in motion :
* It is most interesting to notice that many of the double stars have colours which
are complementary the one of the other. The larger slaris commonly of a ruddy or
orange tinge, and the smaller one appears blue or green. "No green or blue star of
any decided hue," says Sir J. Ilerschel, "has ever, we believe, been noticed unassociated
with a companion brighter than itself."
400 ORDER IX THE MOVEMENTS
absolute rest is unknown in the material universe. Our
sun, with its retinue of planets, is travelling through
space at the rate of 422,000 miles a day, towards a point
near the constellation of Hercules. The mind grows
dizzy in contemplating such velocity, but everything,
meanwhile, is as stable as if all were at rest. It is evident
that arrangements have been made to produce equilibrium
among powers, each of which, acting alone, might work
only destruction, and stability among objects which are
never for one instant at rest.
Even before the construction of Lord Kosse's telescope,
it was thought that astronomers had sounded space to
nearly 500 times the distance of Sirius, that is, ten thou-
sand billions of miles. " Hence it seems as if, were the
world island, in which our system is placed, viewed from
the cluster in the hand of Perseus, it would probably
appear as an assemblage of telescopic stars, ranged behind
each other in boundless perspective ; from that of An-
dromeda, it would diminish to a milky way, or pure
nebulosity." It may be asserted, without any risk of
contradiction, that nowhere within this wide knowable
space, do we discover even the semblance of chance, con-
fusion, lawlessness, or oversight. Nay, it may now be
most confidently affirmed, that nowhere within this ex-
tensive region, or in the long ages opened up to us by
the time which light requires to travel from different
stars, do we discover any traces of a chaos now existing,
or ever having existed, or of worlds being formed by na-
tural law, or of worlds only half formed or in the course
of formation, or of any object overlooked, or out of place,
or not in harmony with all the rest. As far as the tele-
scope can carry our vision, or enable thought to carry out
its calculations, we find all the bodies already formed,
already in harmony, moving on in their spheres as if per-
OF THE HEAVENLY BODIES. 401
forming some great and good office, and all so perfect,
that our feelings are in harmony with the declaration of
their Maker, when He is re23resented as proclaiming them
"to be all very good."
SECT. II. SPECIAL ADJUSTMENTS NEEDED IN ORDER TO THE
HARMONY OP COSMICAL BODIES.
It is very manifest that every one part of the visible
universe is intimately connected with every other. There
are certain agents which seem to operate through the
whole of it ; — there is gravitation attracting all the
bodies to each other ; there is light flowing from mil-
lions of luminaries ; there is heat radiating everywhere
from the warmer to the colder regions ; there is probably,
also, a universally diffused ether ; and possibly, also,
some others of no less extensive influence, such as elec-
tricity and magnetism. We are now to shew that all
these require an adjustment in order to their beneficial
operation.
First, Gravitation. — The planets move in nearly cir-
cular orbits round the sun, and the satellites round their
primaries, and binary and multiple stars round each
other, in consequence of the balanced adjustment of the
velocity of the moving body and the central attractive
force. Without a nice adaptation of the one to the
other, two opposite but equally deleterious results might
have followed. Had the velocity been beyond its proper
proportion, the body would have rushed away in a
hyperbolic curve into space, to run the risk of collision
with other bodies, and certainly to derange every other
well-arranged system into which it might intrude. On
the other hand, had the centripetal force been in excess,
the separate existence of the bodies would have been lost
402 SPECIAL ADJUSTMENTS NEEDED IN ORDER
in a mutual collapse and embrace, which must have de-
stroyed every existing arrangement upon their surface.
In a calculation of probability in a previous section,
(pp. 48-50,) we have referred to two circumstances as
needful to the stability of the mundane system : first, that
the planets have a motion round the sun in the same di-
rection ; and have orbits with very little and scarcely-vary-
ing eccentricity, in planes with very moderate differences
of inclination. There are conditions absolutely necessary
to the continuance of the system ; — as the invariability of
the major axis of the orbits of the planets, proved by
lengthened investigations, in which the highest powers
of the infinitesimal calculus were employed, by the most
distinguished mathematicians of the latter half of last
century ; as the long periodic change of the eccentricity
of Jupiter and Saturn, which together amount to nearly
a thousandth part of the mass of the sun, and which
might have deranged the whole system under a different
arrangement ; and there is the farther circumstance, that
the planetary revolutions have among each other no com-
mon measure. Change any one of these essential condi-
tions, and the issue, sooner or later, would be a fearful
conflict, in which every existing cosmical arrangement,
with the planetary inhabitants, such as animals and
plants, would inevitably be destroyed.
But here we must allude to the attempt which has
been made to turn aside the force of this argument, by a
scheme of ingenious cosmogony suggested by Laplace.
According to this hypothesis, the whole solar system has
been formed out of floating matter rotating round an
axis, and which, being at first greatly heated, has, in
the process of cooling and condensation, given off the
planets one by one, beginning with the outer ; which
planets, again, being thrown off in the form of rings, have,
TO THE HARMONY OF COSMICAL BODIES. 403
in their condensation, given off the satellites. We do
not mean to enter upon a minute examination of this
hypothesis. It was connected with, and received much
of its support from, the supposed existence of unformed
nebulous matter floating in space. Lord Rosse's telescope
has dispelled these clouds, and the theories, light as
clouds, which were built on them.* It may be acknow-
ledged that there are some of the peculiar phenomena
of planetary movements which can thus be accounted for.
But there are other facts beyond its power to explain, as
that the satellites of Uranus should move in a direction
opposite to that of all the other planetary bodies. " The
satellites," says Professor Nichol, an ardent supporter of
the hypothesis, " present farther a curious anomaly, or
rather peculiarity. So far as we know, they all rotate
on their axis, like our moon, in the exact period of a
revolution in their orbits. This mode of rotation is evi-
dently that of the original ring, but why the satellites
have preserved that period is a mystery."f This theory
has been subjected to a searching examination by Sir J.
Herschel. " If," says he, " it is to be regarded as de-
monstrated truth, or as receiving the smallest support
from any observed numerical relations which actually
hold good among the elements of the planetary orbits, I
beg leave to demur. Assuredly, it receives no support
from the observation of the effects of sidereal aggrega-
* Some may urge that the hypothesis has heen corroborated by certain experiments of
Plateau as to the phenomena of a free liquid mass withdrawn from the action of gravity.
In speaking of the division of liquid masses into parts, Plateau had compared the minute
masses to satellites; but in a subsequent paper he corrects the misapprehensions to which
his language had given rise, as if it favoured Laplace's cosmogony. " It is. clear," he
eays, "that this mode of formation is entirely foreign to Laplace's Cosmogenic Hypothe-
sis; therefore, we have no idea of deducing from this little experiment, which only re-
fers to the effects of molecular attraction, and not to those of gravitation, any argument
in favour of the hypothesis in question, an hypothesis which, in other respects, we do not
auopt." — Taylor's Scientific Memoirs vol. v.
+ Planetary System, p. 241.
404 SPECIAL ADJUSTMENTS NEEDED IN ORDER
tion, as exemplified in the formation of globular and
elliptic clusters, supposing them to have resulted from
such aggregation. For we see this cause, working out
in thousands of instances, to have resulted, not in the
formation of a single large central "body, surrounded by
a few smaller attendants, disposed in one plane around
it, but in systems of infinitely greater complexity, con-
sisting of multitudes of nearly equal' luminaries, grouped
together in a solid elliptic or globular form. So far,
then, as any conclusion from our observations of nebulae
can go, the result of agglomerative tendencies may, in-
deed, be the formation of families of stars of a general and
very striking character, but we see nothing to lead us to
presume its farther result to be the surrounding of those
stars with planetary attendants."'*
But let us admit, for argument's sake, the truth of this
hypothesis, and we still urge that numberless adaptations,
and these of a very remarkable description, are needed in
order to admit of this loose floating matter being formed
into the harmonious and beneficent results which fall
* Opening Address, British Association, 1845. There follows a severe criticism of the
pretended verification of that hypothesis by M. Comte, -which had been quoted with
approbation by the author of the Vestiges of Creation, and by J. S. Mill in his Logic.
"If, in pursuit of this idea, we find the author first computing the time of rotation the
sun must have had about its axis, so that a planet situate on its surface, and forming
part of it, should not press on that surface, and should therefore be in a state of indif-
ference as to its adhesion or detachment; if we find him, in this computation, throwing
overboard, as troublesome, all those essential considerations of the law of cooling, the
change of spheroidal form, the internal distribution of density, the probable non-circula-
tion of the internal and external shells in the same periodic time, on which alone it is
possible to execute such a calculation correctly, and avowedly, as a short cut to a result,
using, as the bijsis of his calculation, ' the elementary Huygcnian theorems for the
evaluation of centrifugal forces in combination with the law of gravitation,' — a combina-
tion which, I need not explain to those who have read the first book of Newton, leads
direct to Kepler's law ; and if we find him then gravely turning round upon us, and ad-
ducing the coincidence of the resulting periods, compared with the distances of the plan-
ets, with this law of Kepler, as being the numerical verification in question ;— where, I
would ask, is there a student to be found, who has graduated as a Senior Optime in this
University, who will not at once lay his finger on the fallacy of such an argument, and
declare it a vicious circle ?" &c.
TO THE HARMONY OF COSMICAL BODIES. 405
under our notice on the earth, and which maybe presumed
to exist also in the other planets. Whence, for example,
the striking adaptation of the gravitating, chemical, gal-
vanic, and electric powers to each other ? Whence the
plants and animals which cover the face of the earth ?
Whence animal instincts and the human soul ? Whence
the correspondence between all these, and the atmosphere,
and the light of the sun ? All this is wonderful on
any system, but becomes vastly more incomprehensible
when it is supposed that it originated in certain nebulous
matter. The cosmogony referred to has never been car-
ried out into details ; but if it had, we could have taken
these up, and have proved that every one of them implies
an adjustment. But dealing with it in its present vague
form, it may be maintained that either the properties of
this cosmical matter must have been such as in their own
nature to imply a designing mind in the formation of
them, or adjustments must have been made in order to
their beneficent operation ; and on either supposition we
have evidence of intelligence, and the hypothesis leaves
the theistic argument where it found it.
We go on to mention another beautiful arrangement
which should be regarded as equally striking, whether
we adopt or reject the hypothesis of Laplace. In the
annual motion of the earth round the sun, its axis is in-
clined from the perpendicular to its orbit at an angle of
twenty-three degrees, and remains constantly parallel to
this direction. By this arrangement the changes of
temperature on the earth's surface, and of the seasons, are
produced. Had the axis of the earth, instead of being so
inclined, been perpendicular to the plane of its orbit, as
is the case in Jupiter, the sun would always have been
vertical to the same line of places, the equatorial regions
would have been parched by the heat, while the regions
406 SPECIAL ADJUSTMENTS NEEDED IN ORDER
called teemprate in the present arrangement, would have
been consigned to utter desolation. By the existing dis-
position, the various parts of the earth are brought more
fully under the solar influence, and we have all the de-
lightful and beneficent effects which flow from the variety
of climates.
Again, the earth is nearer the sun at one season than at
another, and without some counteracting influence there
would be an inconvenient increase both of the cold of
winter and the heat of summer in the southern hemi-
sphere, and the climates of the two hemispheres would be
rendered altogether unlike each other. But any injury
which might arise from this cause is made to disappear
chiefly by means of the circumstance that the point of
the earth's orbit which is nearest the sun is that over
which it moves with the greatest speed. " It follows,"
says Poisson, "from the theory of Lambert, that the
quantity of heat which is conveyed by the sun to the
earth, is the same during the passage from the vernal to
the autumnal equinox, as in returning from the latter to
the former. The much longer time which the sun takes
in the first part of his course is exactly compensated by
its proportionably greater distance, and the quantities of
heat which is conveyed to the earth is the same, whether
in the one hemisphere or in the other, north or south."®
Second, Tlie Universal Diffusion of Light. — Under
this head we are called first to admire the wisdom of the
arrangement by which a luminous body is placed in the
centre of a solar system ; there being no physical ne-
cessity, so far as we can discover, for such a disposition.
Some astronomers have supposed (it has not been con-
firmed by later investigation) that there are binary and
multiple stars moving round central bodies which are
* Humboldt's Cosmos, vol. iv. p, 460.
TO THE HARMONY OF COSMICAL BODIES. 407
not luminous ; it is evident that if our earth had been
made to circle round such a body, or round a body simi-
lar in constitution to itself, most of the living objects
upon its surface would have become extinct. There is
an evident harmony between the force or amount of
light coming from the sun and the organism of plants
and animals, for the life of both of which light, and this
in a certain measure, is requisite. Had the light been
much stronger than it is, it would have dazzled and
blinded the eyes of animals, and stimulated to an exces-
sive extent the growth of certain plants, while it would
have utterly destroyed others. On the other hand, a
diminution to any great extent of the luminiferous power
of the sun would have imparted to our earth a dull and
murky appearance, and have rendered it impossible for
the plants of the earth, deprived of their needful stimulus,
to subsist. If our earth, with its present vegetable cover-
ing and animal tenants, had been as far removed from the
sun as Uranus or Neptune, or even Jupiter, it is certain
that a large portion of the species of living beings would
long before this have ceased to exist. Taking the inten-
sity of light upon the earth as one, the proportions in the
other planets will be as follows : —
Mercury,
. 6-674.
Jupiter, .
. 0-036.
Venus, .
. 1-911.
Saturn, . .
. o-oii.
Mars, . .
. 0-431.
Uranus, .
. 0-003.
Pallas, .
, 0-130.
Neptune, .
. o-oov
It is very evident that the earth could not have been
placed in the room of any one of the other planets with-
out endangering the existence of the greater number of
the organized objects upon its surface. It may also be
mentioned here that there is a beautiful harmony insti-
tuted between light and the gaseous envelope surround-
* Cosmos, vol. iv. p. 4(U
408 SPECIAL ADJUSTMENTS NEEDED IN OEDER
ing our earth whereby the sun's rays are diffused through
the atmosphere, and are reflected upon us from every
point of the concave heavens, in the infinitely varied
hues of sky and cloud, instead of all streaming with burn-
ing power from the sun alone, and leaving the rest of the
hemisphere black as if it had been clothed in mourning
attire.
It is also worthy of being noticed, that in consequence
of the comparatively small eccentricity of its orbit, much
the same quantity of light falls upon the earth at all
times. In this respect it may be compared with some of
the other planets.
While the earth, in perihelion, is T034, it is in aphelion 0-967.
Mercury, 10-58, 4'59.
Mars, 0-52, 0'36.
Juno, 0-25, 0-09.
If the earth's eccentricity had been as great as that of
Mercury or Juno, it is certain that not a few of our most
useful and beautiful plants would have altogether disap-
peared, or rather could never have existed.
TJiirdly, The Universal Diffusion of Heat. — There is
need of a number of harmonious adjustments in order to
the beneficent operation of this agent so powerful for good
but also for evil.
It will be readily acknowledged that there must be a
uniform temperature on the surface of the ground in
order to the continuance of organized beings upon it.
We know, as a matter of fact, that the earth's surface
has had much the same temperature throughout histori-
cal ages. The paintings and inscriptions on the monu-
ments of Egypt shew that in that country much the
same plants were cultivated, and that they ripened about
the same season between 3000 and 4000 years ago, in
the ages of the Pharaohs, as at this day. The plants of
TO THE HARMONY OF COSMIOAL BODIES. 409
Canaan at the time of Moses and Joshua were not dif-
ferent from what they are now. But the sustaining of
this equable temperature depends on a combination of
circumstances. First, there are various sources of heat,
and, in particular, there is the internal heat of the earth,
which is known to be much greater than that of the ex-
terior, and increasing as we go farther down, and there
are the beams of the sun daily taking the circuit of the
earth. Were these influences operating alone, the
temperature of the earth's surface would soon be in-
conveniently or rather destructively heated. But to
counterbalance them, we have the earth's surface and its
atmosphere . radiating heat into the circumambient re-
gions of space, which are ascertained to have a very low
temperature, being lower than the freezing point of
mercury. Our earth has thus, on the one hand, power-
ful fires to heat it, and, on the other hand, an extensive
reservoir of cold to keep it cool ; its surface is warmed by
the internal heat, and by the heat of the sun ; and its
temperature being thus rendered higher than that of the
vault of heaven, it is ever radiating heat towards the re-
gions of space according to the beautiful law of the
universe, whereby all things tend towards an equilibrium.
The uniform temperature of the earth from year to year,
and from age to age, necessary to the continuance of the
races of plants and animals, is sustained by the harmoni-
ous adjustment of agents which seem to be distinct from,
and independent of, each other, except in the original
collocation of all things. An increase in the internal
heat, or in the heat streaming from the sun, would
speedily scorch the ground, and burn up the plants
which grow upon it. The same dire effects would fol-
low, were the cool celestial regions not ready to receive
the heat from the sun-warmed face of our earth and at-
18
410 SPECIAL ADJUSTMENTS NEEDED IN ORDER
mosphere. On the other hand, were there not sources
of heat within or without, the temperature of the earth
would speedily sink below zero, and the whole globe
be as much ice-bound as the north or south poles.
It is to be remembered that the temperature of the ce-
lestial regions is dependent, if not in whole, at least in
part, on the temperature of the innumerable bodies which
move in them. We are thus led to see that we are de-
pendent for our continued existence, and our everyday
comforts at home and abroad, on the disposition through
millions of years of millions of bodies, removed from us
millions of miles.
On the earth we are dependent for our very artificial
fires, and for the mechanical power which can be gene-
rated by them, upon influences which have descended
from heaven in ages long past. We are using coal
formed of vegetables fostered in former geological eras
by the sun's rays. Allusion is made to these and to
some other beneficial effects of the solar rays in the fol-
lowing passage from Sir John Herschers Treatise on
Astronomy : — " B}r the vivifying action of the sun's rays,
vegetables are enabled to draw support from inorganic
matter, and become in their turn the support of animals
and of man, and the sources of those great deposits of
dynamical efficiency which are laid up for human use in
our coal strata. By them the waters of the sea are made
to circulate in vapours through the air, and irrigate the
land, producing springs and rivers. By them are pro-
duced all disturbances of the chemical equilibrium of the
elements of nature which, by a series of compositions and
decompositions, give rise to new products, and originate a
transfer of materials."
The far-reaching truth here enunciated has opened
the way to experiments, calculations, and speculations,
TO THE HAKMONY OF COSMICAL BODIES. 411
which all tend to shew how intimately connected every
one part of the visible universe is with every other. " We
must look, then, to the sun," says Professor W. Thomson,
" as the source from which the mechanical energy of all
the motions and heat of living creatures, and all the
motion, heat, and life derived from fires and artificial
flames, is supplied. The natural motions of air and water
derive their energy partly, no doubt, from the sun's heat,
but partly also from the earth's rotatory motion, and the
relative motions and mutual forces between the earth,
moon, and sun. If we except the heat derivable from the
combustion of native sulphur and of meteoric iron, every
kind of motion (heat and light included) that takes place
naturally, or that can be called into existence through
man's directing powers on this earth, derives its mechani-
cal energy either from the sun's heat, or from motions
and forces among bodies of the solar system."
Such results having been attained in regard to the
source of the heat and mechanical energy called forth on
the earth, the question is started, Whence does the sun
set the heat and light which he sheds ? There are insu-
perable scientific difficulties in the way of supposing that
the sun is a heated body losing heat, or that the sun is a
great fire emitting heat due to chemical action ; and it
has been surmised that " the sun's heat is probably due
to friction in his atmosphere between his surface and a
vortex of vapours, fed externally by the evaporation of
small planets in a surrounding region of very high tem-
perature, which they reach by gradual spiral paths, and
falling inwards, in torrents of meteoric rain, form the
luminous atmosphere of intense resistance to his sur-
face."*
* See Professor W. Thomson's Paper In Trans, of Fvoyal Society of Edin., 1854, and
abstract of same in Edin. New Phil. Jour., January, 1855.
412 SPECIAL ADJUSTMENTS NEEDED IN OKDER
Fourthly, Indications of some other Universally Ope-
rative Agents. Possibly all those wo have been consi-
dering and those we are now to contemplate, may be
modifications of one and the same force : this is a
favourite idea of not a few living men of the very highest,
scientific eminence, and it may be granted without affect-
ing our argument. For if there be only one force, what
a variety of adjustments must have been made in order
to its producing such a number of results, so different
from each other and so beneficent in their character !
Our conclusion follows equally from the admission of a
number of forces suited to each other, or one force with
an infinite number of adjusted collocations. But at the
present stage of science, we are not entitled to say that
all the forces of nature are one ; they present themselves
to us as diverse, but all correlated, and capable of excit-
ing each other. Meanwhile, we must look at them in the
forms which they assume, and besides those which have
been already before us, there are the magnetic and chemi-
cal powers.
It has been ascertained that there are periodical varia-
tions in the magnetic forces on the earth depending on
the solar day and the time of the year, and pointing to
the sun as the cause. It has also been discovered that
there is a variation in the direction of the magnetic
needle, going through all its changes exactly in each
lunar day. cc It would seem, therefore, that some of tbe
curious phenomena of magnetism, which have hitherto
been regarded as strictly terrestrial, are really due to
solar and lunar as much as terrestrial magnetism.*"
It has also been supposed that there is a connexion be-
tween the period of the recurrence of the sun's spots and
* President's (Mr, HopkiDs) Address to British Association 1853.
TO THE HARMONY OF COSMICAL BODIES. 413
the period of the variation of magnetism on the earth's
surface. The maxima of the sun's spots occurred m
1828, 1837, and 1848, the minima in 1833 and 1843;
and it has been shown that the cycle of the variations in
the earth's magnetic intensity is also about ten years, and
bears a relation to the other cycle. These discoveries open
up curious glimpses of relations between things on the
earth and things in the sky, such as men have not been
inclined to believe in since science expelled astrology from
human credence.
We know further, that in the sun's rays there is a che-
mical (actinic) as well as a luminiferous and calorific
potency. These principles have each, in its own way, an
influence on the germination and growth of the plant;
and it is affirmed that all are. in harmony with the
seasons, and that each is strongest relatively at the time
when most needed for the function which it has to dis-
charge in fostering the vegetable. Actinism is needed
in order to the healthful germination of seed ; light is
required to excite the plant to decompose carbonic acid;
and caloric is necessary in order to develop and carry
out the reproductive energies of the plant. " It is now,"
says Mr. Hunt, " an ascertained fact, that the solar
beam, during spring, contains a large amount of actinic
principle, so necessary at that season for the germination
of seeds and the development of buds. In summer, there
is a large proportion of the light-giving principle neces-
sary to the formation of the wooden parts of plants. As
autumn approaches, the calorific or heat-giving principles
of the solar rays increase. This is necessary to harden
the woody parts and prepare them for the approach of
winter. It is thus that the proportions are changed with
the seasons, and thus that vegetation is germinated,
grown, and hardened by them, We have these state-
414 SPECIAL ADJUSTMENTS NEEDED IN ORDER
ments on the authority of Mr. Hunt.* It is affirmed
that every flower has its own peculiar power in reference
to heat, and that different plants take the different tem-
peratures needed in order to their health, by reason of
their different colours, which also determine the relative
amount of dew deposited on the leaves.
Fifthly, Traces of an all-pervading Ether. The ex-
istence of such a medium between the various cosmical
bodies had long been suspected, and has now been estab-
lished to the satisfaction of most scientific men. The
resistance offered by it to the comet of Encke is the
cause, it is believed, of the acceleration of the period
of the revolution of that body, by causing it to fall
nearer the sun. The acceleration is appreciable, being
about two days in each revolution, which occupies about
3T3o years, and it has been observed during a number of
revolutions. But we know too little of the nature of
this ether to admit of its being turned to much use in
such a treatise. It may be legitimately argued, however,
that if light — according to the prevailing theory in the
present day — consists in vibrations in an ether, we must
call in an important class of adaptations, the absence or
alteration of any one of which would disturb the economy
of the universe. The three rays, the violet, the yellow,
and the red, must each have ether waves of different
lengths ; and they must each make a different number
of vibrations in a second, upon which circumstance the
character of the coloured rays depends. The number of
vibrations in the second is approximately as folows : —
Violet, 699 billions. Yellow, 535 billions. Red, 477 billions.
It needs no lengthened statement to show how liable
* Report on Chemical Action of Solav Radiations, British Association, 1850. We are
by no means at the bottom of this subject. Farther investigation is evidently needed
In order to put us in possession of the exact facts, and we are not yet within sight of the
rationale of them.
TO THE HARMONY OF C03MICAL BODIES. 415
such a complicated system is to go wrong, and how nice
must be the continued adjustments so as to admit of oui
distinguishing stars and the colours of stars, so distant
that light must require thousands of years to travel
from them to us. For, on looking abroad on the face of
the sky, we cannot be said to be looking on the stars as
they now exist, but on these stars as they existed many
years, it may be thousands of years ago. We have per-
fect confidence that there is no deception in all this, but
in order to our trust being well founded, it is needful to
suppose, if there be any truth in the prevailing scientific
theory, that the ether has retained its laws and colloca-
tions through both immeasurable ages of time and regions
of space.
Before closing this subject, we must refer to a most
important class of facts, and speculations founded upon
them, which have come into great prominence in the
present day. The calculations of Lagrange and Laplace
in regard to the stability of the solar system, (see p. 390,)
proceeded on assumptions which later science has shewn
not to be warranted. In particular, they pre-supposed
that the planets moved in vacuo. But the prevalent
opinion at this stage of advancing science is, that they
move in an ether, the effect of which must be to lessen
the velocity, and bring all the planetary bodies nearer
and nearer the sun. The influence thus exercised in
a brief period must be very small, but acting constantly,
as it does, it must, in the course of ages, produce appre-
ciable effects, and tend to break up the solar system.*
Other facts, not reconcilable with absolute stability
* The demonstration of the French mathematicians proceeded on the further assump-
tion that the planets are solid throughout, and not fluid. But our earth, whatever may
be the case with the other planets, has the largest portion of its surface covered with
waters ever agitated by tides produced by the gravitation of the moon. Now, it is well
known that when there is water in a boat, the motion of the boat is retarded by the agi*
416 SPECIAL ADJUSTMENTS NEEDED IN ORDER
have come into view. Sir J. Herschel says that the
breaking up of the Milky Way affords proof that it can-
not last for ever, and equally bears witness that its past
duration cannot be admitted to be infinite.
Certain very important conclusions, tending in the
same direction, have been established on following out
the modern doctrine in regard to heat. Heat is now
regarded as, if not identical with mechanical power,
at least the means of producing it. As has been already
stated, we are at present taking advantage of the mecha-
nical energy excited on our earth, and laid up in store
for us during the age of the coal formation. As this
dynamical agency is being dissipated and wasted, we
have here another disturbing element. The following
are the conclusions drawn by Professor W. Thomson,
who has deeply studied this subject : — " I. There is at
present in the material world a universal tendency to the
dissipations of mechanical energy. II. Any restoration
of mechanical energy without more than equivalent dis-
sipation is impossible in inanimate material processes,
and is probably never affected by means of organized
matter, either endowed with vegetable life, or subjected
to the will of an animated creature. III. Within a
finite period of time past, the earth must have been, and
within a finite period of time to come, the earth must
again be unfit for the habitation of man, as at present
constituted, unless operations have been, or are to 'be,
performed which are impossible under the laws to which
the known operations going on at present in the mate-
rial world are subject."'"'
All this does not in the least detract from the skill
tation of the water ; and on the same principle (a scientific friend assures us) the tidal
agitation of the waters on the surfac.e of the earth exercises a disturbing influence on tho
movements of the earth in its orbit. Can any counteracting power be detected ?
t Transactions of Royal Society of Edinburgh, 1S52.
TO TIIE HARMONY OF COSMICAL BODIES. 417
displayed in those wonderful adjustments and counter-
poises which were brought to light by the analytic dex-
terity of the French mathematicians ; but it brings into
view an overlooked set of agencies which must, in the
course of ages, change the present system of things,
provided always that they are not corrected by some
well-adjusted counterbalancing arrangements. Professor
Thompson says that there is not in nature any counter-
acting agency. Without dogmatizing on so difficult a
subject, it may be confidently asserted that science at its
present stage cannot point out any means of restoring
the lost energy. Even though it could be restored by
natural means beyond the ken of man, it must be in con-
sequence of a wonderful adjustment planned by intelli-
gence. In either case, we are made to feel the depen-
dence of all physical nature upon a higher power either
to keep things in their present stable condition, or, in
the event of some great change, such as seems not ob-
scurely pointed to in the Word of God, to render that
change beneficent. Doubtless the world is stable, (for
" the earth abideth forever,") but it is by means of forces,
each of which would make it very unstable, and which
are made to produce stability by counteracting each
other, so that there is a truth in that part of the theogony
of Hesiod which represents- Eros, the healer of divisions,
as the world-forming principle. All this balancing is
fitted, we should say intended, to carry up our minds to
Him who holds the balances in His hands. Our confi-
dence in the permanence of things must be made to rest,
after all, on the purposes of a G-od who has ordained all
things from the beginning, and who, when He changes
any existing state of things, changes them in the deve-
lopment of one and the same mighty plan.
We are now in circumstances to estimate the amount
418 SPECIAL ADAPTATIONS IN ORDER
of truth in a statement of Paley, which has been quoted
with approbation by others, — " My opinion of astronomy/
says he, " has all along been that it is not the best
medium through which to prove the agency of an intelli-
gent Creator ; but that this being proved, it shews beyond
all other sciences the magnificence of his operations/'*
Now, it may be admitted that astronomy does not dis-
play so many cases of special adjustment as the animal
kingdom, so beautifully illustrated by Paley. The reasons
are not difficult to find : First, we do not know so much of
celestial bodies as of objects on the surface of the earth ;
we know little or nothing of the internal structure of the
planets ; we know absolutely nothing of the composition
of the sun or stars ; we do not know for certain whether
any one of them, is inhabited ; and so we cannot expect
to be able to unfold such adaptations among them as
among the objects with which we are familiar. Then,
secondly, and more especially, there is no necessity for
such special adaptation in the case of inorganic bodies as
is required for living bodies, and more particularly for
animals requiring provision to be made not only for their
existence, but for their comfort. It will be found as a
general rule, that we discover the clearest examples of
special adaptation where our knowledge is most extensive
and minute, and that they are more abundant where we
see they are most required, in the frames of organized
existences, especially of animated beings capable of plea-
sure and pain, and the most abundant of all in the
frame of man, the being who needs the greatest number
and complication of organs to enable him to fufill his
high destiny. But it is satisfactory to observe, that we are
able to detect a number of striking examples of special
adaptation among the celestial bodies in general, and the
* Paley's Natural Theology, chap. xxii.
TO THE HARMONY OF COSMICAL BODIES. 419
planetary bodies in particular, and it is an instructive cir-
cumstance, that in consequence of the advance of know-
ledge, we are able to unfold a greater number than could
be developed in the days of Paley. There is no just
ground, then, for the scoffing remark of the haughty
and eccentric Frenchman, (who denies that he is an
atheist, seeing that he adores himself and has set up a
formal worship of his system,) that the heavens cannot
now be appealed to as a proof of the existence of Deity,
or for the inference drawn by him, that the time will
speedily arrive when organized objects will be in the
same condition ; for while, for the reasons stated, animals
and plants must ever furnish the most striking examples
of design, it is still true that " the heavens declaee
THE GLORY OF GOD, AND THE FIRMAMENT SHOWETH HIS
HANDIWORK."
BOOK THIRD.
THE INTERPRETATION OF THE FACTS.
CHAPTER I.
THE ARGUMENT FROM COMBINED ORDER AND ADAPTATION.
We are now to estimate the force of the influence of
two streams, which we have hitherto been contemplating
as flowing in parallel channels.
The principle of Order has been scientifically ex-
pounded only in modern times, and in regard to the
animal and vegetable kingdoms only within these few
years. But it existed from the creation of the world,
and had been noticed in a general way since the creation
of intelligent being. Science in its latest advances is
simply coming up to, and explaining, the spontaneous
suggestions of human thought, which as it muses upon
the universe is at once struck with the model forms and
correspondences which everywhere prevail. The late
discoveries in regard to homotypes, homologies, and we
may add homceophytes, or parallel developments in ani-
mal and vegetable structures, is but the scientific exposi-
tion of what all along impressed intelligent observers,
without their being able to give an account of it. Nor
COMBINED ORDER AND ADAPTATION. 421
are these remarkable facts of an isolated or exceptionable
character ; on the contrary, they are merely striking ex-
amples of what is universal, and they have their homo-
types, analogues, homologues, and parallels, in every
department of nature.
The principle of Special Adaptation, or that of par-
ticular conformity to the position of the object and func-
tion of the organ, has also been noticed all along by minds
addicted to reflection. Socrates is represented by Xeno-
phon as delighting to dwell upon it. So strong, indeed,
was this tendency in the ancient world, and in the middle
ages, that Bacon felt himself called on to remove the
inquiry from physical science, where it hindered the dis-
covery of physical agents, to metaphysics, where it might
have a legitimate scope. Bacon was right in saying, that
the propensity to discover final cause had sometimes
come in the way of the discovery of physical cause ; but
he is altogether wrong in affirming that it is. barren of
results in scientific inquiry, for in certain departments
of natural science, such as physiology and comparative
anatomy, it is a most powerful instrument of discovery,
and such eminent men as Cuvier and Sir Charles Bell
delight to inform us that they have proceeded on the
principle of final cause in all their researches.
It is not difficult to discover the beauty and the ap-
propriateness of both these principles.
On the one hand, the mind discerns the need and ap-
preciates the propriety of the principle of Order. Without
some such governing principle, nature would be incom-
prehensible by human intelligence, and this because of
the very number and multiplicity of the objects of which it
presents, each eager to catch our notice ; and the mind
in trying to apprehend them would have felt itself lost,
as in a forest through which there is no pathway, or as
422 THE ARGUMENT FROM
in a vast storehouse, where the seeds of every species of
plant on the earth's surface, are mixed in hopeless con-
fusion. By what means is it that man is enabled to ar-
range into groups the objects by which he is surrounded,
and thus acquire a scientific knowledge of them, and turn
them to practical purposes ? Plainly, by reason of the
circumstance that there are numberless points of resem-
blance and correspondence between them. Scientific
men have so long been familiar with this process that
they are not impressed by it as they ought, and seldom
do they inquire into the ground on which it proceeds.
It is only when something new, such as the discovery of
homologies in the animal kingdom, comes to light, that
they are led to reflect on what has been too common to be
specially noticed. But if they but seriously reflect on
the subject, they will find that it is because of the uni-
versal prevalence of points of resemblance and corre-
spondence that man is enabled to grasp the infinity of
objects which fall under his view, into classes and sub-
classes, which can be comprehended by the intellect, and
treasured up in the memory.
No doubt the mind has in itself a power of forming
classes altogether independent of any special arrange-
ment in order to aid it ; but such groupings, though they
may at times help the memory, are of no intellectual or
scientific value. But there are means in nature of guid-
ing the mind to the formation of classes which have
a deep and far-reaching significance. It is true, in an
important sense, that classes are already formed for us
in nature. Man will find it expedient, in all cases,
to attend to these arrangements made to his hand, and
he must attend to them, provided he represent his
classification as a natural one. It may illustrate our
general subject to show what are the distinctive marks
COMBINED ORDER AND ADAPTATION. 423
of natural classes, that is, of classes having the sanction
of nature.
And first, we may take a classification which is not
of this description. It is conceivable that a person
might arrange all animated beings according to their
size. He might put all animals of a certain height in one
class, and all animals below that in another class. Every
one sees how arbitrary, in short, how contrary to nature,
such a distribution would be. It would often separate
animals belonging to the very same species, while it would
put in one confused group bird and fish, mammal and
insect. And why, it may be asked, does the naturalist at
once reject such a classification ? Perhaps it is answered,
because he is seeking a natural arrangement. But this
answer, though correct so far as it goes, does not go down
to the depths of the subject, for we immediately ask, Is
not the distinction of size a natural one ? He who would
really sound the depths of this subject, and not skim over
it, must be prepared to state what is the difference between
an artificial and a natural classification.
All natural classes will be found to have not merely
one, but an aggregate of common attributes. It follows
that, when objects are classified according to a natural
arrangement, the possession of any one characteristic is a
mark of a great many others. Thus, when an animal is
described as a reptile, we know that its blood is cold, that
its heart consists of three cavities, and that its young are
produced from eggs ; and when we hear an animal called
by the name of mammal, we know not only that it
suckles its young, but that it breathes by lungs, that its
blood is warm, and that its heart consists of four com-
partments. In short, when we have fixed on a truly
natural arrangement, the presence of any one character-
istic becomes a sign of others, commonly of very many
424 THE ARGUMENT FROM
others, at times of an inexhaustible number of others.
The co-existence of these characteristics in one object, and
their invariable co-existence in all objects possessing any-
one of them, is a clear evidence that such an arrangement
has b*een purposely made. A class with such an aggregate
of qualities as its ground, is said to be one of " Kinds."
There are some valuable remarks on this subject in the
Logic of Mr. J. S. Mill. " There are some classes, the
things contained in which differ from other things only
in certain particulars, which can be numbered, while
others differ in more than can be numbered, more than
even we need ever expect to know. Some classes have
little or nothing in common to characterize them by,
except precisely what is connoted by the name ; white
things, for example, are not distinguished by any com-
mon properties except whiteness, or if they are it is only
by such as are in some way dependent upon or connected
with whiteness. But a hundred generations have not
exhausted the common properties of animals or of plants,
or of sulphur, or of phosphorus ; nor do we suppose them
to be exhausted, but proceed to new observations and
experiments, in the full confidence of discovering new
properties, which were by no means implied in those we
previously knew. It appears, therefore, that the pro-
perties on which we ground our classes sometimes
exhaust all that the class has in common, or contain it
all by some mode of implication ; but in other instances,
we make a selection of a few properties from among not
only a greater number, but a number inexhaustible by
us, and to which, as wc know no bounds, they may, so
far as we are concerned, be regarded as infinite."*
We now see wherein lies the essential distinction be-
tween an artificial and a natural class, and the superiority
* Mill's Logic, B. 1. c. vii. 4
COMBINED ORDER AND ADAPTATION. 425
of the one to the other. In an artificial arrangement, we
seize on a quality — not arbitrarily, it may be, but still for
mere convenience' sake — and our arrangement does not
yield us any farther information on the subject. In a
natural classification, on the other hand, we fix on qua-
lities which are invariably accompanied with certain
other qualities, and which are, therefore, signs of them.
All that an artificial class can do is to aid the memory,
by having the innumerable objects put into a convenient
number of groups. Even for this purpose a natural
arrangement, if we can seize it, will be vastly more useful
than an artificial one, as it will be found, in fact, that no
artificial arrangement can embrace all the facts, and
enable us to carry them about with us in convenient
groups. But a natural classification does more than
help the memory, it imparts positive knowledge, inas-
much as one property is a sign of the presence of a vast
number of others. The most fundamental of all groups
in Natural History, that of species, is always one of Kinds.
It is formed on the principle that all the animals in-
cluded in it might have proceeded from a common pa-
rentage ; but all animals belonging to the same species
are found to have a great many other points of resem-
blance besides their belonging to one stock. The same
is true, to a greater or less extent, of all other natural
groups, such as genera, orders, and kingdoms. In all
such natural classes, the presence of some one attribute is
a means of informing us of the presence of others. Thus,
the power of speech is one of the characteristics of huma-
nity ; but there are many others, so many others, that
physical and metaphysical science cannot be said to have
fully exhausted them, and the presence of the power of
speech is a sign of all these others. A traveller has lost
himself in a deep forest, amidst wild birds and beasts,
426 THE ARGUMENT FROM
whose cries all raise within him feelings of alarm ; sud-
denly he hears a human voice, and that sound at once
announces that there is intelligence at hand, and pro-
bably also a compassionate heart, and the power and
disposition to aid him. All the marks of a natural class
are significent, in the same way, of an indefinite number
of other attributes.
This invariable collocation of characteristic qualities
in certain objects, so that the one is a sign of all the rest,
is a clear proof that classes do exist in nature — that is,
that objects are gathered into classes. This, we doubt
not, was at least one of the truths which led to the mystic
doctrine of Plato about Ideas or Types, above individual
things and prior to individual things, and in mediaeval
times to the doctrine of Realism, according to which,
universals or classes have an existence as well as indivi-
duals. There is a great truth at the basis of these
theories, now exploded, but entertained in former ages by
some of the deepest thinkers which our world has pro-
duced. This truth was not correctly seized, was very
imperfectly, indeed, often very erroneously represented,
but still it is deep in the constitution of things. All
natural objects, and especially all organic objects, are
fashioned according to type, and operate according to
unchanging laws. The individuals all die, shewing how
insignificant they are, whereas the genus and species
survive. The flowers of last summer are all faded, but
in the coming summer, other flowers will spring forth to
continue the same form. Amidst the flux and reflux of
all individual existences, the laws which they obey are
permanent. In particular, classes, genera, and species,
have as certain an existence in nature as the objects
which are classified.
There is no new thing under the sun. The modern
COMBINED ORDER AND ADAPTATION. 427
doctrine of organic correspondences is but the distinct
articulation of what thinking minds have ever felt — but
the scientific interpretation of the unconscious musings
of deeper thinkers, as they have been gazing on the cryp-
tic symbols of nature, ever since the time when inward
reflection was awakened by outward objects. Nay, it is,
after all, but the extension, into a new field, of the prin-
ciples on which scientific classification has all along been
resting. The facts on which the new doctrines are
founded are the homotypes and homologues of the facts
on which ordinary classification proceeds. The classifi-
cation into genera and species proceeds on corresponden-
ces among a vast number of individuals. The doctrine
of homotypes takes its rise from the correspondences in
many parts of the same individual. Homologues are
corresponding members in different individuals. We
may add that homceophytes are corresponding stages in
the development of different organic kingdoms.
So' much, then, in regard to the fitness of the one
principle — it enables mankind to make a practical and
scientific use of the objects by which they are surrounded;
and, as some one remarks, nature was made to be enjoyed
by brutes, but to be contemplated by man. It is still
more necessary that the other principle, that of Special
Adaptation, be attended to ; for if the comprehensibility
and beauty of the universe depend on the one, the very
existence of the objects in it, and especially of animated
beings, depends on the other.
And here it may be important to remark, that the
principle of special adaptation assumes two distinct
forms. So far as the efficient powers, the dynamical ener-
gies, the active properties of matter, are concerned, the
adaptation consists in their adjustment so as to produce
a general law, or it may be also an individual effect of a
428 THE ARGUMENT FROM
beneficent character. It is thus that the centripetal and
centrifugal forces are adjusted to yield the harmonies of
the planetary system ; thus that the relation between
the earth's orbit and the sun are arranged to yield the
seasons of the year. In organic bodies, again, where the
law is one of type or structure, we find the special adap-
tation taking a somewhat different form. We now meet
not with an adjustment of forces to produce a law, but a
modification of a general type, or a departure from it on
one side or other, and this obviously to enable the part
to execute its office. Under the first of these forms the
adaptation is necessary in order to the very existence
of general order ; under the second, it bends the general
order to the accomplishment of sj)ecial ends.
It is in this second form that adaptation appears in
the structure of animated beings. Not only the comfort
of the animal, but its very continuance upon the earth,
depends on every organ being made to serve its special
function. And here it is satisfactory to find, that while
attention is paid both to order and special end, the most
uniform regard is had to the latter. There are cases, as
we have seen, in which the general plan, if not sacrificed,
is at least kept in abeyance, so that it is very difficult to
detect it. It is of all pretensions the most absurd, in
certain naturalists to profess to be able to see the general
homologies, which are often very obscure, and yet regret
that they can never discover special modifications to serve
a given end, which are often so very clear. It is satis-
factory to find that the wellbeing of the plant and the
happiness of the animal are never sacrificed in following
out the typical form. The general often gives way to
the special, but the special never gives way to the gene-
ral. It cannot be said of any animated being, that its
individual comfort has been sacrificed in the attention
COMBINED ORDER AND ADAPTATION. 429
paid to some general law or model shape. It is a cir-
cumstance worthy of being noted, that the typical form
is most clearly exhibited in the lower animals, whose
wants and functions are fewest ; and that the principle of
teleology is carried out to the farthest extent in animals
higher in the scale, whose organism is the most compli-
cated, and has the most numerous and varied functions
to perform ; and, farthest of all, in man, whose frame is
so fearfully and wonderfully made to enable it to become
the fit instrument of that spiritual nature to which it is
united.
When we take an enlarged view of these two princi-
ples, we shall find that they are not inconsistent with
each other, but rather that they depend on each other.
There is an adaptation necessary in order to those regu-
lar successions of events and model forms which come so
frequently before us. The regular flow or periodic re-
currence of such phenomena as the tides, the seasons, is
the result of arrangements many and varied. The forms
assumed by plants and animals is evidently the contem-
plated issue of a multitude of forces made to combine to
this end.* On the other hand, the general order, in some
* When the action of the combination of powers necessary to the development of an
organ is interfered with, we have a Monster. In monstrosities the principle of order is
not accommodated to the usual special end. They are always comparatively few in
number — in short, the exception. But we are not to conclude that they are faihires,
or that they have no end to serve. A world in which they were the rule would certainly
be a failure; but, as exceptions, they are as instructive as the rule. They help man to
discover the nature of those agencies which combine to form typical organs, and they
shew how derangements which, when few, work no evil, would have been fearful if
they had been frequent. Teratology, which treats of natural monstrosities, has now a
place among acknowledged sciences. Single monsters are produced by arrest of develop-
ment; double by the union of homologous parts, as of veins to veins, and arteries to
arteries. The aberrations of monstrosity do not exceed certain limits. They have their
distinctive characters, and long ago there were noticed five orders, twenty-three fami-
lies, and eighty-three genera. So far as these monstrosities do not produce pain, they
are not evils any more than an irregularly-formed crystal is. So far as they are the
means of entailing suffering and humiliation among mankind, thev carry us into the
profouudest of all mysteries (which we cannot here discuss) — the existence of evil.
430 THE ARGUMENT FROM
cases, accomplishes very useful purposes ; as when the
mathematical law of the increment of the shell enables
certain molluscs to ascend and descend the water at
will ; and when the spiral arrangement of the leaves
and buds all round the axis exposes them equally to the
light and to the air. In all cases the general order is
adapted to the intellect of those who are expected to
contemplate it.
Everything has, after all, a final cause. The general
order pervading nature is just a final cause of a higher
and more archetypal character. In the special principle,
we have every organ suited to its function ; in the more
general principle, we find all the objects in nature suited
to man, who has to study and to use them. Professor
Owen has declared, that his practical assistant found
himself greatly aided, in setting up the bones of the skull,
by proceeding on the principle that they were constructed
on the vertebrate type. Lecturers on anatomy find their
students following them much more readily when they
expound the skeleton on the archetypal idea. It is only
by proceeding on some such method that the nomen-
clature of comparative anatomy can be retained by the
memory. Without ■ some such principle, there would
require to be one set of names for the bones in man,
another set for the bones in quadrupeds, and a third and
a fourth set for the bones of birds and fishes. By the
discovery of homologous parts running through all, it
has been found possible to devise a common nomencla-
ture, admitting of application to all vertebrate animals.
But let it be observed, that it is not the unity of the
nomenclature which gives the unity to nature, but it is
the unity of nature which has given a unity to human
science, and the nomenclature which science employs.
These correspondences are admirably fitted to make
COMBINED OKDER AND ADAPTATION. 431
creation comprehensible by the human faculties. The
more obvious points of resemblance enable man to recog-
nise the nature and end of the objects by which he is
surrounded. The more fixed points allow him to arrange
them into classes in due subordination. The repetition
of parts permit of his at once taking an intelligent glance
along the whole length, and over the whole frame, of the
animal and plant. The answerable parts permit of his
discovering unity among organs that serve very diverse
purposes. The members with similar functions invite
him to observe a universal final cause. The parallel de-
velopment points to a unity of arrangement in the forces
by which all these correspondences are produced. The
prophetic system of geology entitles him to look on the
earliest past as a foreshadowing of the future, and on the
present as the fulfilment of what has gone before.
Before the time of GeofTroy St. Hilaire, the undeve-
loped rudimentary organs were frequently thrown away
as useless in the Museum of Comparative Anatomy in
Paris. But it is rash, it is wrong to declare that any
part of nature is useless. GeofTroy restored these organs,
and thus led the way to grander generalizations of organic
objects than had ever been formed before. We have now
before us a sufficient final cause of typical forms. We
may rise above a special adaptation of parts to an arche-
typal adaptation of the whole to the constitution of
intellectual beings. We have here a most beautiful cor-
respondence between the laws of external nature and the
laws of the mind, between the laws of things and the laws
of thought.* While the special modifications or adapta-
tions investigated so carefully by Cuvicr, are intended to
promote the wellbeing of the particular species of ani-
mal, the archetypal plan investigated by Owen is fitted
* This is so interesting a topic, that we ave to devote the next chapter to it.
4:32 THE ARGUMENT FROM
to make the animal intelligible by the intelligent crea-
tion. Owen has developed — to some extent perhaps un-
consciously, but to a far greater extent consciously — a
teleology of a higher order than Cuvier.
Viewed in this light, the two principles, though evi-
dently differing from each other in many respects, and
requiring to be separately treated, come to be very much
alike, may be seen to be analogous — that is, different
organs fulfilling a similar function. The special adap-
tation proceeds on a general principle of beneficence, and
the general principle is an example of adaptation to a
special end. There is a general plan in the purpose,
and a purpose in the general plan. The teleology is a
homology, and the homology is an example of teleology.
There are some who prefer a somewhat different re-
ligious interpretation of the model forms of nature.
Order and law, they say, are the natural methods of the
Divine procedure, the ways in which God's nature and
character spontaneously exhibit themselves. We need
seek, they say, no other explanation than this of the
typical forms in heaven and earth, they are just the
manifestation of the divine ideas. And, as to man's re-
cognition and appreciation of these laws and models, it
is to be accounted for by the circumstance that he was
made in his Maker's image. We are indisposed to
advance a single word against this view ; possibly it
may be as true, as it is certainly striking and sublime.
It is certainly a doctrine which cannot be disproven : we
may venture to doubt whether it admits of absolute
proof. Do we know so much of the Divine nature as, a
priori, to be able to affirm with certainty, how that na-
ture must manifest itself in creation ? There may even
be presumption implied in declaring, in some cases, that
the harmonies of nature arc after the taste or character of
COMBINED ORDER AND ADAPTATION. 433
God ; for example, that complementary colours are more
beautiful to His eye, as they are to ours, when seen in
collocation, than non-complementary colours. But. while
we cannot predicate much, a priori, of the character of
God, there is much that we can affirm, a posteriori, of
the character of man, of his intellectual aptitudes and
his tastes. We do know that correlations among objects
are needful in order to his being able scientifically to
arrange them, and practically to use them, and that he
has tastes implanted within him, which are gratified by
objects without him ; for example, the ability to receive
gratification from the complementary colours of animals
and plants. We have here a firm ground to stand on,
in reasoning from "what we know," and as there is a
correspondence between man's constitution and the scenes
in which he is placed, we cannot be wrong in inferring
that God, by His nature and character, is led to accom-
modate the external world created by Him, to the intel-
lectual nature of man, also created by Him. There is
sense, then, and this a sense as grand as it is true, in
which we are justified in representing these types as
proceeding from the very ideas of God, from His eternal
wisdom, impelled by His eternal love. Nay, we are
inclined to think that as there are homologies anions
organic structures, so there may also be correspondences
among spiritual natures, and that other intelligences,
differing in many respects from man, may resemble him
in this, that they also delight in these laws and patterns;
while God, over all, may be conceived as rejoicing in all
His works together.
As taking this view, we are not inclined to admit that
the doctrine of final cause has been set aside, or shaken,
or even damaged, by late discoveries in natural history. It
is true that some of those engaged in making these dis-
19
434 THE ARGUMENT FROM
coveries did not see their consistency with teleology.
Oken, as a pantheist, admitted, so far as we know, no final
cause into his system. Geoffroy St. Hilaire reckoned it
presumptuous in man to discover any end designed by
the Creator. Cuvier was led to reject the doctrine of the
unity of the vertebrate skeleton, partly by the practical
turn of his mind, partly by the fear that it would inter-
fere with the doctrine of final cause. Some, we suspect,
have supported the doctrine of a physical uniformity of
parts, because it seemed to deliver them from the ne-
cessity of calling in a personal God to account for the
economy of nature ; while not a few have regarded
it with suspicion, because it seemed to be atheistic or
pantheistic in its tendency. But amidst all these exhi-
bitions of presumption and of fear, the doctrine of final
cause stands as firm and as impregnable as ever, assail-
able by no known fact, consistent with every established
truth.
Physiological research has, we admit, established a
truth which cannot be reduced to final cause in the nar-
row sense of the term, but that truth is not inconsistent
with final cause — it is an illustration of a higher form
of final cause. We blame Cuvier because he would not
attend to the evidence which his own discoveries supplied
in favor of unity of composition. Not being of a specu-
lative turn of mind, he would attend, he said, to nothing
but facts, and content himself with classifying them.
But we must also blame Geoffroy St. Hilaire, when, after
condemning Cuvier for narrowing the field of science, he
professed to be incapable of discovering final cause, and
bids us remain " historiaus of what is." * Final cause is,
to say the least of it, as certain as unity of composition.
It is surely as certain that the eye was made to see, as
* Vie, Travaux ct Doc. Scien. de Geoffroy St. Hilaire, par son Fils, p. 304.
COMBINED OEDER AND ADAPTATION. 435
that it is the homologue of the whisker of a cat.* "We
give little credit for sincerity to those who acknowledge
that they have overwhelming evidence in favour of the
former truth, but no convincing proof in behalf of the
latter.
Again, there are metaphysicians who think that they
have undermined the whole doctrine. We must reserve
to a separate section the examination of any plausible
considerations which they can urge. Meanwhile, let it
be observed that their objections proceed on principles
which would undermine all other objective truth, and
leave us only a series of connected mental processes. The
principles by which they would set final cause aside have
not half the evidence in their favour which the doctrine
of final cause has. We are sorry to find an accomplished
and devout writer saying, " The argument from first or
final cause will not bear the tests of modern metaphysi-
cal inquirers. The most highly educated minds are above
them, the uneducated cannot be made to comprehend
them."f The modern metaphysical speculators who
have rejected final cause, have great need to review their
own principles when they are opposed to a truth so obvious
and so supported by scientific research. The argument
from final cause is one which the uneducated universally
feel, though they are incapable of explicating it logically,
or illustrating it scientifically. The educated can
feel that they are above it only in so far as they are
elevated by the intoxicating fumes of German specula-
tion, which would make man believe that he is a god,
and that he creates from the stores of his own mind the
final cause, which he simply discovers. Verily there are
* In tho animal body the following parts are admitted to be all homologous;— Tactile
Corpuscles, Pacinian bodies, Savian bodies, Muciparous ducts of fishes, Vibrissas (whis-
kers) of cat and others, the eye, the ear.
t Jowctt on Epistle to the Romans; Natural Religion, p. 410.
436 THE ARGUMENT FROM
metaphysicians whose heads have been so dizzied with
the turnings and windings of their own cogitations, that
realities swim before them and they cannot distinguish
between them and phantoms. Living forever in a region
of pure, or rather very impure and cloudy speculation,
they do not, as the physical investigator is ever doing,
meet with stringent facts to restrain and control them ;
they have become utterly incapable of weighing ordinary
evidence, probable and moral ; they cannot see that the
thoroughly established truths of inductive science are in
the least degree more certain than the last spawned, a
priori, theory of the universe ; nay, some of them (such
as Hegel) are prepared to deny the doctrine of gravi-
tation itself, because it will not fall in with their theory.
No wonder that final cause cannot stand the tests of
such inquirers, for these tests need themselves to be tested.
As taking so enlarged a view of final cause, we have
no objection to the general statement laid down by some
eminent scientific men, that there are parts of the vege-
table and animal frame which have no respect to the
functions of the plant and animal. " There is yet an-
other law," says De Candolle, "to be understood to enable
us to judge properly respecting the nature of organs. In
innumerable instances there appear forms similar to
those which are connected with a definite function, but
which do not fulfil that function, and nature, in these
instances, as in the animal kingdom., seems to produce
forms which are completely useless, merely for the sake
of a harmonious and symmetrical structure. The ap-
pearance of filaments with empty anthers in flowers
which are altogether female, and of female parts in
flowers wholly male, the structure of filaments in other
forms where they resemble nectaries, the false nectaro-
thecse in such orchidge as have no nectaries, these are
COMBINED ORDER AND ADAPTATION. 437
all formations which can only be explained by the law
of nature we are now illustrating." Professor Owen
uses similar language : — " I think it will be obvious that
the principle of final adaptation fails to satisfy all the
conditions of the problem. That every segment and al-
most every bone which is present in the human hand and
arm should exist in the fin of the whale, solely because
it is assumed they were required in such number and
collocation for the movement of that undivided and in-
flexible paddle, squares as little with our idea of the
simplest mode of effecting the purpose, as the reason
which might be assigned for the greater number of bones
in the cranium of the chick, viz., to allow the safe com-
pression of the brain-case during the act of extrusion,
squares with the requirements of that act."* And again,
" The attempt to explain by the Cuvierian principles the
facts of special homology on the hypothesis of the sub-
servience of the parts so determined to similar ends in
different animals — to say that the same or answerable
bones occur in them because they have to perform simi-
lar functions — involves many difficulties, and is opposed
by numerous phenomena. We may admit that the
multiplied points of ossification in the skull of the human
foetus facilitate, and were designed to facilitate, child-
birth ; yet something more than such a final pnrpose lies
beneath the fact, that most of these osseous centres repre-
sent permanently distinct bones in the cold-blooded ver-
tebrates. The cranium of the bird, which is composed
in the adult of a single bone, is ossified from the same
number of points as in the human embryo, without the
possibility of a similar purpose being subserved thereby
in the extrication of the chick from the fractured egg-
shell. The composite structure is repeated in the minute
* On Limbs, p. 40.
438 THE ARGUMENT FROM
and, prematurely-born embryo of the marsupial quadru-
peds. Moreover, in the bird and marsupial, as in the
human subject, the different points of ossification have
the same relative position and plan of arrrangement as in
the skull of the young crocodile, in which, as in most
other reptiles, and in most fishes, the bones, so commen-
cing, maintain throughout life their .primitive distinct-
ness. These, and a hundred such facts, force upon the
contemplative anatomist the inadequacy of the teleo-
logical hypothesis."*
It might be argued, if not with truth, at least with
considerable plausibility, that some of these statements
go farther than science warrants. It might be main-
tained that we are not entit]ed to affirm that an organ
has no use, merely because we are not able to detect it.
Science, as it advances, is ever shewing that organs which
were at one time regarded as useless, have most impor-
tant uses in the animal and vegetable economy. Who
will venture to affirm that the bones of the skull of the
young chick, have no reference, directly or indirectly, to
animal instincts ? or that the division in the parts of the
fin of the whale do not the better enable the female to
carry the cub under her arm when she is pursued by an
enemy ?f But, while we throw out this caution, we are
inclined to admit that certainly in the vegetable, and
probably in the animal kingdom, there are parts retained
for the sake of symmetry which are not necessary to the
mere function of the organ. In making such an admis-
sion, we are not, so far as we can judge, weakening the
great principle of final cause, so long as we call in a
higher final cause, and affirm that these part are fitted,
in some cases, to give instruction to mankind, and. in
3ther cases, to gratify their higher tastes.
* Homologies, p. 75. + See Sooresby, Arctic Regions, vol. i p. 471
COMBINED OKDEK AND ADAPTATION. 439
In Civil Architecture there are four principles, it is
said,* to be attended to :— 1st, Convenience ; 2c?, Sym-
metry ; 3d, Eurythma, or such a balance and disposi-
tion of parts as evidences design ; and, 4th, Ornament.
It is pleasant to notice that not one of these is wanting
in the architecture of nature. The presence of any of
them might be sufficient to prove design ; the presence
and concurrance of them all furnishes the most over-
whelming evidence. Upon taking a combined view of
the whole, we feel as if we have proof of much more than
of the existence of law or a pinciple of order ; we feel
as if we have distinct traces of a personal God planning
minute and specific ends. We do not know whether to
admire most the all-pervading order which runs through
the whole of nature, through all the parts of the plant
and animal, and through the hundreds of thousands of
different species of plants and animals, or the skillful ac-
commodation of every part, and of every organ, in every
species, to the purpose which it is meant to serve. The
one leads us to discover the lofty wisdom which planned
all things from the beginning, and the enlarged bene-
ficence reaching over all without respect of persons;
whereas the other impresses us more with the providen-
tial care and special beneficence which, in attending to
the whole, has not overlooked any part, but has made
provision for every individual member of the myriads of
animated beings.
* See Lectures in connexion with opening of Great Exibition.
CHAPTER II.
CORRESPONDENCE BETWEEN THE LAWS OF THE MATERIAL
WORLD AND THE FACULTIES OF THE HUMAN MIND. '
SECT. I. THE FANTASY, OR IMAGING POWER OF THE MIND.
It is Mind that is to be the special object of contem-
plation in this chapter ; — not mind in its essence, of
which we can know but little, but mind in its actual
operations ; mind looking out by the senses on the .world
without, and studying and admiring it ; mind making
the past to reappear, and imagine the absent as if pre-
sent ; mind analyzing the complex structure of nature
into its elements, and discovering resemblances which
group all nature into a few grand systems ; mind rising
from the effect to the remote and unseen cause, arguing
from the known past to the unknown future, and disco-
vering, by cogitation, new planets before the far-pene-
trating telescope had detected them : it is this mind
which is to exhibit a few of its varied powers and move-
ments to our view. Natural philosophy does not unfold
laws of a wider sweej), chemistry does not disclose more
curious combinations, nor natural history a more wonder-
ful organization, than this ever active and living mind.
There is a gradation from the inanimate, up through the
plant and the animal, to mind, as the crowning object.
IMAGINING POWER OF THE MIND. 441
The motion of the planet in its elliptic orbit is no doubt
beautiful to contemplate, but having enjoyed a higher
existence, we would not choose to run, year after year, in
that one unvarying orbit. If the choice were given us,
we would rather be a plant than a planet — we would
rather be a lily, expanding its petals in the sunshine —
we would rather be the oak, shooting out and ramifying
at will, and facing the buffetings of the storm. If the
option were allowed us, there is a higher life that we
would prefer. An eminent man, on seeing the sea-fowl
career from the wave to the cliff, and sweeping from the
cliff to the wave, expressed the momentary feeling, " Well,
I should not dislike to be a sea-bird — I would have such
a variety of life in water, in air, and on land." But hav-
ing enjoyed by our Maker's beneficence, a still higher
life, we would not descend to these lower states of exist-
ence. For this mind with which we are endowed, or
rather, which constitutes our true self, can, in its thoughts,
run a wider orbit than the planets, and wander into in-
finity ; it can, in the midst of sunshine and of storm,
grow on and on in knowledge and in love, and in all that
is great and good throughout eternity ; it can take in
more than earth and sea and air and all the elements,
and rise, by contemplation and purification, to gaze on
infinite perfection embodied in the character of God.
Surely this mind, with its laws and operation, is worthy
of our careful study. We are to shew that, while it is
vastly above them all, it is yet suited, by its structure and
its organs, to all the objects by which it is surrounded,
and which it is expected to contemplate and to use.
When man appears on the earth, which had been so long
in preparation for him, he comes writh powers and apti-
tudes fitted to the scene in which he is placed. We have
now before us a correspondence of a higher kind than
19*
442 THE FANTASY, OR
any previously contemplated. It may be called the
Archetypal correspondence connecting Homology with
Teleology.
In illustrating this subject, we are to use mental facul-
ties and laws, which, under one name or other, are treated,
or at least referred to and incidentally sanctioned, in every
system of mental science. There are, doubtless, differ-
ences of opinion as to the nomenclature best fitted to set
forth these laws and powers ; we are to avoid the diffi-
culties arising from this source, by employing as little
technical language as possible. Even those who regard
our classification as not the best, and our analysis too
refined or not sufficiently refined, will yet be prepared
to acknowledge, that the powers of which we treat are
in the mind of man, either as original or derived ; and
this is all that needs to be admitted in order to our being-
entitled to use them as we do in this chapter. We begin
with the Imaging or Pictorial Power of the Mind.
The reader will be able to discover what is meant by
this power, if he but observe, that whatever is recalled
or imagined by the mind comes with an image more or
less distinct. We call up, let me suppose, some incident
of our childhood. We remember the clay on which we
were sent to school, and how we set out from our parents'
roof with strangely mingled feelings of hope and appre-
hension. As we bring back these scenes, mark how every-
thing appears with a pictorial power. We have a vivid
picture, it may be, of the road along which we passed ;
we see, as it were, the school-house both externally and
internally ; we hear, as it were, the master addressing us,
and the remarks which the children made upon us. Or
more pleasant still, we remember a holiday trip under-
taken by us, in the company of a pleasant companion or
kind relative, to a scene interesting in itself, or made
IMAGING POWER OF THE MIND. 443
interesting by its historical associations ; or, what we felt
to be' still more agreeable, the visit was paid to the house
of a kind friend, who had a thousand contrivances to
please and entertain us. How vivid the representation
before us of the events of the journey, of the little inci-
dents which befell us, of the amusements which were
provided for us, and of the persons, the countenances, the
voice and words of those who joined us in our mirth, or
ministered to our gratification ! We not only remember
that there were such events, but we, as it were, perceive
them before us ; this imaging of them is, as it were, an
essential element of our remembrance. Wordsworth is
painting from the life when he speaks of
" Those recollected hours that have the charm
Of visionary things ; those lovely forms
And sweet sensations that throw back our life,
And almost make remotest infancy
A visible scene on which the sun is shining."
Or possibly there may be scenes which have imprinted
themselves still more deeply upon our minds, which
have, as it were, burned their image into our souls. Let
us cast back our mind upon the time when death, as an
unwelcome intruder, first entered our dwelling. We re-
member ourselves standing by the dying bed of a father
or mother, or sister or brother, and then we recollect how
a few days after we saw the lifeless body put into the
coffin, and, within a brief period after, saw it borne away
to the tomb. How terribly vivid and distinct do all
these scenes stand before us at this instant ! We, as it
were, see that pallid countenance looking forth from the
couch upon us ; we, as it were, hear that voice becoming
feeblei and yet feebler ; and then we feel as if we were
looking at that fixed gaze which the countenance assumed
444 THE VANTASY, OK
after the spirit had fled ; we follow the long funeral as
it winds away to the place of the dead, and we hear the
earth falling on the coffin as the dust is committed to
its kindred dust.
And we would have it remarked, that not only are we
able to represent these sensible scenes, we are farther able
to picture the thoughts and feelings which passed through
our minds as we mingled in them. Not only do we re-
member the road along which we travelled, and the
building into which we entered, we can recall the feel-
ings with which we set out from our parents' house, and
those with which we walked into the school. Not only
do we recollect the amusements which so interested us,
but the feelings of interest with which we engaged in
them. Not only can we picture the chamber in which
a relative breathed his last, we can call up the mingled
feelings of anxiety, of fear, and of hope with which we
watched by his dying bed, and the emotions of grief
which overwhelmed us as we endeavoured to realize the
loss which we had suffered. We can set before us the
feelings which passed through our minds as we sat by
his corpse, or when we returned to our dwelling and
found all so blank and melancholy. We are obliged to
use metaphorical language in describing these recollec-
tions, but it is language which embodies .and expresses
important truth : — we speak of being able to image, to
picture to ourselves not only the outward events which
called forth the feelings, but the very feelings themselves.
This mental power we are disposed to call the Fan-
tasy. It is a phrase used by Aristotle, and explained by
Quintilian — "Quas tpuvmokas Graeci vocant nos sane
visiones appellamus : per quas imagines rerum absentium
ita representantur animo ut eas cernere oculis ac prsesen-
tes habere videamur." Lord Monboddo defines it, in his
IMAGING POWER OF THE MIND. 445
Ancient Metaphysics, the power " by which the images
of things presented to the mind "by the senses are pre-
served." But this definition is too narrow for our purpose,
for the mind can represent not only what has been pre-
sented by the senses, but all that has been before the
consciousness, all that has been under the eye of reflection.
We think it of moment to make this remark, because
the grand object of higher education, and especially of
religious discipline, is to lift the mind above material to
the contemplation of spiritual images.
Every one sees how these mental pictures are fitted to
enliven existence and increase enjoyment. They help us,
too, by their vividness, to carry on trains of thought.
Those nominalists are altogether mistaken who suppose
that man reasons solely by means of words, or artificial
signs of any description. We are far, indeed, from deny-
ing the utility of language as an instrument of thought.
Language is a sort of stenography, by which we can ab-
breviate thought, and it helps us especially in those more
recondite processes, in which our more refined abstrac-
tions or wider generalizations could be represented by no
fantasy, or where images could mislead by their fulness
of detail, or their vividness. But man thinks primarily
by mental symbols, by pictures remembered or created
by the image-fcrming capacity of the mind. So far from
oral or written signs being primarily the object of
thought, the first artificial signs are commonly outward
pictures of the inward image. The earliest words and
writings coined by man were hieroglyphic, and it was by
degrees that they were refined into the highly analytic
expressions furnished by our more advanced languages,
such as those of ancient Greece, or those modern ones
formed out of the debris of old tongues. But language,
if used as the sole representative sign, has its defects as
446 THE FANTASY, OK
well as its excellences. The thoughts thus represented
have, on account of their remoteness from reality, no
interest to vast multitudes ; these dried plants do not
excite half the amount of emotion which collects around
the natural ones with the life circulating in them. The
most popular employers of words are those who use them
to set before lis vivid pictures. In the ages and nations
in which dead symbols are most resorted to, and serve
the highest purposes, we must still go to nature for our
fresh and living symbols. Need we say that nature is
ever presenting them to us in infinite number and variety,
in the forms of the animal and plant, in the mountains
and plains of the earth, in the clouds and stars of the
sky.
It is, indeed, of vast moment to have the mind stored
with a variety of noble images to enliven and elevate it,
to act as Quintilian says, "incitamenta mentis." This
end is much promoted by an early training among natu-
ral objects which are picturesque ; by travelling at a
later period of life into foreign countries, and by the
opportunity thus afforded of holding communion with
nature in her grander forms, and of inspecting the noblest
products of the fine arts. But, while gathering these
material pictures, let the young man and the old man
not forget that there are others which he should not be
losing, and which, if he part with, his gain will be more
than counterbalanced by his loss. For there are images
which it is still more important to have treasured up in
his mind ; they are the images of domestic peace, the
images of home and friends, of the affectionate mother
(we can never have more than one mother) and devoted
wife, of kind sisters and smiling children, and to these
let us acid, by personal intercourse with them, or by ele-
vated reading, the images of the great and good, of
IMAGING POWER OF THE MIND. 447
heroic in en, who toiled and bled for noble ends, and of
equally heroic women, who' lost sight of themselves in
works of disinterested love and sacrifice. These are in
themselves vastly more exalted, and ten thousand times
more exalting, than all your statues, draped and un-
draped, about which connoisseurs so talk and rave ; they
are fitted to become incitements to all excellence, and he
who has been at the pains to collect them and hang them
round the chamber of his mind, is like one dwelling in a
portrait gallery, from which the forms of ancestors are
looking down upon him with a smile, and exhorting him
to all that is great and good.
And there is one other object of which it is more im-
portant still that we have a noble image. The funda-
mental evil of images, as used in the worship of God,
does not lie in their being pictures, but in their incapa-
city to act as pictures. " To whom will ye liken God ?
or what likeness will ye compare unto Him ?" The stars
in their purity are not suitable emblems of His holiness;
nor the moon, shining in beauty, of His loveliness ; the
sun in all his splendour has his beams paled in the
dazzling brightness of His glory. There can be no cor-
poreal image of God, who is a spirit. One grand aim of
Eevelation is to lift us above such gross representations,
and to lead us to worship a spiritual God in " spirit and
in truth." Man in his first estate, not his body but his
soul, was a sort of image of Him ; but man in his fallen
state is a caricature of Him. But we have one perfect
image of God set before us in His Word, as in a glass,
(2 Cor. iii. 18,) in Him who is the brightness of the
Father's glory — only seen under a milder lustre — and
the express image of His person. By such a mediate
representation, aided by the types and figures which the
Old Testament supplies, our minds may rise to a some-
148 THE FANTASY, OR
what adequate idea of a spiritual God, even as, by the
redemption purchased by that same Mediator, we hope
at last to mount to the immediate presence of God. " No
man hath seen God at any time ; the only-begotten Son,
which is in the bosom of the Father, He hath declared
Him." We shall return to this subject before we close
the treatise.
But speaking of the connexion pre-established between
the laws of mind and those of matter, it is most inter-
esting to notice, that the most correct memory, in recalling
an object, seldom reproduces it with all its individualities.
In coming up before the mind as a picture, it appears
with only the more prominent qualities, features, and
colours — only with those which most vividly impressed
the senses, or which were most noticed at the time. The
consequence is, that the recollection appears very much
as a type of the object. In representing, for example,
some animal that we have seen, say a deer, we drop from
our view not a few specialties of the individual, and form
a sort of general picture, which might stand for any other
deer. There may be cases indeed in which we were so
deeply impressed with every part of the object, that we
see it as it were before us, with all its peculiarities ; but
in most instances we so far generalize or idealize it.
That this should be the law of the reproduction of what
we have experienced, we cannot but regard as, in a ne-
gative sense, a most merciful dispensation, as it saves the
mind from the distraction which would be produced by
numberless minutise ever floating before it. But there is
another and more positive advantage arising from this
tendency of the mind to generalize its representations —
the mental image of natural objects becomes a type of the
species or genus. After we have looked at a number of
natural, especially organized objects, the recollection will
IMAGING POWER OF THE MIND. 449.
be found, in fact, to be not far from the type constituted
in nature as the model after which objects are formed.
With this generalized representation in our minds, we
are the better prepared at once to refer the individual
before us to its genus or species, and at the same time
to notice the specialties of the new individuals which
may come before us. There are thus preparations made,
in the very structure of the mind, for the contemplation
and recognition of natural substances and beings. The
very mind and memory supplies a series of typical models,
and he who has his mind furnished with such images, is
like one walking in a museum filled with specimens to?
illustrate the natural orders. The mind is disposed, on
the one hand, to give to every object a typical form in
its representations ; and on the other hand, it finds, in
its actual experience, that types run through nature.
We might almost say, that there are types in nature and
types in the mind corresponding to each other, as an
object does to its image in a mirror.
sect. ii. the faculties whictt discover relatioks
(correlative.)
The soul is endowed with powers called sense-percep-
tion and self-consciousness, by which it is enabled to
know the material objects presented to it through the-;
senses, and also to know self in its shifting moods &Bf\
states. These simple cognitive powers supply us with
the raw elements of our knowledge. The mh\d has also
a set of powers which enable it to retain and reproduce
the past. To this class belong the memory, which re-
tains and recalls the past in the form which it assumed
when it was previously before the mind ; and the imagin
ation, which brings up the past in new shapes and com,-
450 THE FACULTIES
binations. Both of these are reflective of objects ; but
the one may be compared to the mirror which reflects
whatever has been before it, in its proper form and
colour ; the other may be likened to the kaleidoscope,
which reflects what is before it in an infinite variety of
new forms and dispositions. The knowledge thus ac-
quired and reproduced, though furnishing the materials
of all that follows, would, however, be very valueless un-
less there were a higher set of faculties to work upon it.
But the mind has a class of powers which elaborate the
materials thus acquired, by discovering relations among
the objects which have become known to it. By these
faculties, the materials, all but useless in themselves, are
turned into an infinite variety of cognitions and judg-
ments. Nor is there a greater difference between the
wool when stript from the sheep, and the beautiful gar-
ment into which it is woven ; between the flax in its raw
state, and the fine linen of exquisite pattern constructed
from it ; between the stone when taken from the quarry,
and the marble statue into which it is wrought — than
there is between man's primary knowledge through the
senses and the consciousness, and those lofty compari-
sons, and refined abstractions, and linked ratiocinations,
which he is able to construct by his higher intellectual
faculties. There must be a correspondence between our
simplest knowing powers and the objects known ; but these
other, as the scientific faculties, are the powers which
fall more especially under our notice in tracing the cor-
respondence between the laws of the external world and
the laws of human intelligence.
The relations which the human mind is capable of
discovering are very many and very varied : Locke de-
scribes them as infinite — they are certainly innumerable.
It is necessary, in consequence, to classify them. We
WHICH DISCOVER RELATIONS (CORRELATIVE). 451
are far from thinking that the arrangement which we are
about to submit is perfect. It is possible that a better
division might be made ; but it is sufficient for our
purpose that the powers of which we are to treat, by
whatever name they may be called, and however they may
he arranged, have actually a place in the mind. The mind
is able and disposed to discover at least three distinct
classes of relations : — First, that of Whole and Parts;
secondly, that of Kesemblance and Difference ; thirdly,
that of Cause and Effect. Every one who has ever
seriously reflected on the operations of his own mind,
will be prepared to acknowledge that it has the power
and the inclination to notice these various relations.
We could show that the faculties which discover them
may be found, under one name or other, in almost every
treatise on mental science written in modern times. By
the first class of faculties we are able to separate the com-
plex objects which fall under our notice into parts ; by
the second, we discover the varied points in respect of
which the objects around us correspond ; by the third, we
can connect the present with the past and the future.
By the first, we can, in some measure, penetrate into the
composition of the objects by which we are surrounded ;
by the second, we see how objects are related to others
existing at the same time — how plant, for example, is
related to plant, and animal to animal ; by the third, how
the past has produced the present, and how the present
will produce the future. By the first we have our ab-
stract notions ; by the second, our general notions ; by
the third, our notions of causal relations.
Before proceeding to illustrate them individually, we
would have it observed regarding them generally, that
each has an aptitude and a tendency to seek and to find
the relations which it is its function to discover. We
452 THE FACULTIES
believe that there is a tendency in every faculty, with .
which man is endowed, to operate, and that there is a
pleasure attached to the exercise of it. The eye having
the power to see, delights to he employed in seeing,
and light is pleasant to the eyes. There is a similar
enjoyment felt in the action of all the mental powers.
In particular, there is a tendency on the part of all the
faculties under consideration, to exercise themselves, and
an enjoyment in their exercise. We have not only a
desire to know individual things as they present them-
selves, we have a propensity to discover relations subsist-
ing between them. When any new object falls under our
view, the question forthwith presents itself, How is it re-
lated to other objects known to us ? On noticing any
concrete or complex object, there is a strong intellectual
tendency in our minds to analyze it, to take it to pieces.
If it be a city or island that is brought under our notice,
we immediately ask in what part of the world, in what
country or ocean it is situated. If it be a new plant or
animal that is submitted to us, we ask what is its genus or
species. As strong as any of these, is that which we feel
on witnessing a strange phenomenon, to ascertain its
cause. Let us look at these faculties with the view of
ascertaining how far they are fitted to enable us to com-
prehend the laws of nature.
I. The Faculty which discovers the relation of
Whole and Parts ; in other words, the Faculty of Ab-
straction and Analysis.
When we iook abroad on this world, we find it, as a
whole, presenting a very complicated appearance ; it is a
mighty maze, though not without a plan. When we in-
spect individual objects, we find them all more or less
complex. Almost all the natural substances we meet
WHICH DISCOVER RELATIONS (CORRELATIVE). 453
with in the world arc compound. Air, water, earth, and
fire, which were regarded by the ancients as elements,
have been shown to he composites. This piece of mag-
netized iron has a magnetic property, hence it will turn
to the pole ; it has a gravitating power, hence it falls to
the ground if unsupported ; it reflects certain rays of
light, hence its colour ; it has certain chemical proper-
ties, and hence it will chemically combine with one sub-
stance and not with another. What a vast number of
powers of attraction, of chemical affinity, of electricity
and vitality, are in action in every organism that falls
under our eye !
As the objects which thus press themselves upon our
observation are so complex, we see how needful it is
to have a power of separating a part from a whole in
mental contemplation. But this is a power possessed in
a lower or a higher degree by every human being. On
a complex whole being brought before the mind, it feels
a pleasure in dividing it into its parts, and tracing the
relation of the parts to the whole. It is to this principle,
in part, that we must refer the tendency of children to
take their toys to pieces ; it is in order to discover all
the parts, and how they are connected with one another.
On seeing an ingenious machine, we have a strong inch-
nation all our lives to have its parts taken asunder, that
we may see how they co-operate. We feel it to be pain-
ful to stop in the midst of an important problem, or
theorem, or discussion, or process ; we are anxious to
know how it may issue. We feel, indeed, as if our
knowledge of objects must be very obscure till we have
taken it down and resolved it into its elements, till we
have logically divided it, or physically partitioned it.
We feel as if we required to count over our wealth in
order to estimate its value aright, to travel over our pro-
454 THE FACULTIES
perty, field by field, in order to know how much is com-
prised in it.
This mental power deserves to be noticed by us, be-
cause it furnishes an example of the adaptation of the
mind to the objects by which it is surrounded, and which
it is called to investigate. In consequence of the com-
plication of nature, all science must begin with analysis.
" But induction," says Bacon, " which will be useful in
the invention and demonstration of arts and sciences,
ought to divide nature by proper rejections and exclu-
sions." " Analysis," says Whately, " is the form in which
the first invention or discovery of any kind of system
must originally have taken place." We have thus, on
the one hand, the need of such an aptitude, and, on the
other hand, the tendency working strongly and sponta-
neously. The retort in the laboratory of the chemist is
not more obviously an instrument for decomposing the
substances lying around, than the faculty under conside-
ration is for decomposing the complex structure of the
world in its parts, so as to bring them under scientific
observation and experiment, and thus render their rela-
tion intelligible by the intelligent nature of man.
II. The Faculties which discover the relations
or Resemblance and Difference ; in other words, the
Comparative Faculties.
When a resemblance is discovered, it is between two
or more objects in respect of certain attributes. This
class of faculties may be subdivided according to the
qualities in respect of which the agreement is noticed,
whether they be those of Space, of Time, of Quality, or
Active Property.
(1.) The Faculty which discovers the Belations of
Space, or, in other words, of Locality and Form. —
WHICH DISCOVER RELATIONS (CORRELATIVE) 455
There is a tendency in all minds, and a very strong ten-
dency in some minds, to discover spatial relations. The
commander of an exploring expedition sent to the Arctic
regions, reports that he has seen a hitherto undiscovered
portion of the ocean stretching away in a particular di-
rection, and the question is immediately discussed, How
does it stand related to the parts of the ocean previously
known and described ? A star-gazer reports a new planet
detected by the telescope, and the eager question is put,
What is its orbit, and what its relation to the orbit of the
known planets ? We at times experience a painful feel-
ing because we cannot discover the connexion between
two localities. We are carried over night, let us sup-
pose, from a district of country which is known, to
another which is entirely unknown to us. When morn-
ing dawns, and we go forth to survey the new region
our first inquiry will be, How is it located in reference to
the region which we left, and with which we are ac-
quainted ? We know that some persons have been posi-
tively distressed till they found out the relation of the
two localities, that into which they have been carried, and
that which they had left. The naturalist experiences a
similar feeling of pain mingled with his joy, on discovering
a new animal or plant which he cannot refer to its typi-
cal species or family. There is the plant before him, he
sees its form and all its parts ; and what more, we might
be tempted to ask, could he wish to know of it ? But
the naturalist is not satisfied, he feels as if he wanted
something, till such times as he has discovered its relation
in respect of shape and structure to other natural objects,
and has been able to allot to it its proper place in the clas-
sification of organic objects.
We have shewn, in the second book of this Treatise,
that the most careful regard has been paid to the rela-
456 THE FACULTIES
tions of space in the structure of the universe. The
heavenly bodies have definite shapes and move in definite
orbits. Most inorganic objects on the earth's surface
assume, in certain circumstances, a regular mathema-
tical form. Every organic object has a typical shape
Every kind of bird builds its nest according to a plan of
its own, and lays an egg of a peculiar size and shape.
We have found it interesting to notice, that the horns
which adorn the heads of certain animals have a sweep
which differs in every family, and that every kind of tree
has its own curve for its branch and leaf- vein, and the
outline of its coma and leaf. Animals have been arranged
according to type ever since the days of Aristotle, and
the latest investigations have been disclosing new relations
of form, which are scientifically named homotypes and
homologues. Morphology is now acknowledged to be
the fundamental department of botany, and opens the
way to every other. Locality is the principle to be
attended to by those who would study the geography of
plants and animals. Kelative position is the governing
principle in the stratification of the earth and the bear-
ing of mountain chains, as investigated in geology and
physical geography. But we have now seen that the
mind has a native aptitude to observe such relations as
these. The two thus correspond, as a formed substance
to its mould, as a portrait to its original. It is an eminent
example of those striking adaptations between two things
having no necessary connexion, which shew that both
have been formed by an Intelligent Being, who fashioned
the one to be contemplated by the other.
(2.) The Faculty which discovers the Relations of
Time. — There is a natural inclination among all men to
notice how events are connected in respect of time, and
this becomes, in the case of many, a strong and vehement
WHICH DISCOVER RELATIONS (CORRELATIVE). 457
passion. On hearing an incident related, When did it
happen ? is the question on every one's lips. On some
historical event being disclosed by the casting up of a
long-lost record, the inquiry is instantly made, In what
age did it occur ? Hence, when Lavard dug from the
mounds in which they had long been concealed, the
marble slabs which lined the palaces of ancient Nineveh,
there was instantly awakened an intense desire to know
the age at which these palaces were built, and the
connexion of the historical events represented on them
with the known events of Jewish and Egyptian history.
The mind of the historical narrator feels in a state of
painful anxiety till such time as his relation shall have
been discovered. To aid this faculty, chronology has
fixed on certain great leading events, and set them up as
landmarks. Thus, in Sacred History, we fix on the Flood,
on the call of Abraham, the Exodus from Egypt, the
Eeign of David, and the Babylonish Captivity, and dis-
tribute all other incidents in the intervening periods. To
aid this same aptitude, we have artificial chronometers,
which we set up in our dwellings, or carry about our
persons.
Such circumstances as these prove that there is a
strong intellectual tendency on the part of mankind, to
observe the relations of time. But we have seen in pre-
vious portions of this Essay, that attention is evidently
paid to Time in the economy of natural objects and the
occurrence of natural events. The heavenly bodies have
their definite times of rotation and revolution. Every
organized object has a normal age allotted to it for its
existence on earth. There is a periodical return of days,
and months, and years, which admits of our systemati-
cally arranging our plans and anticipating the future.
We measure the ages of the past by the movements of
20
458 THE FACULTIES
the heavenly bodies and the epochs of geology. Time is
thus divided for us, by great physical events, into re-
gular seasons, and all that we may number our days and
apply our hearts unto wisdom. The connexion between
the timepiece on earth and the motion of the sun in the
heavens, is not more clear than is the relation between
man's capacity and disposition to observe time, and the
wonderful periodical arrangements which everywhere fall
under our eye in nature.
(3.) The Faculty which discovers the Relations of
Quantity. — These are equivalent to the relations of pro-
portion mentioned by Locke, and those of proportion and
degree mentioned by Brown ; they are the relations of
less and more. The faculty which discovers them pro-
ceeds upon the knowledge previously acquired by the
mind of individual objects ; and very frequently, also,
upon the judgments pronounced by the other faculties of
comparison. Upon discovering that objects resemble
each other in respect of space, time, and property, we
may proceed to notice how they have less or more of the
common quality in respect of which they are related.
There is an aptitude in all minds, and a very strong
aptitude in certain minds, of a mathematical turn, to
observe, to search for, and prosecute these relations. We
feel as if our ideas of objects were very loose and inade-
quate, till we have made some sort of calculation as to
their number. The mind delights to discover numerical
repetitions, or proportions, or cycles among the objects
foiling under its notice ; hence the propensity among all
nations to trace significant numbers among natural phe-
nomena, and to group historical events into periods of
three, or four, or seven, or ten, or forty, or a hundred
years. This talent, running waste, has wrought out the
most fanciful and extravagant theories as to the power of
WHICH DISCOVER RELATIONS (CORRELATIVE). 459
numbers ; this talent, used as it ought, has constructed
branches of mathematics, often long before they could be
turned to much practical account.
But we have shewn, in earlier parts of this work, how
much attention is paid throughout the whole of the phy-
sical universe to the relations of number. So far as we
can go down to the elementary construction of matter, we
find numerical proportions appearing, and, as we ascend
upwards to compound and organic bodies, we still find a
significance in numbers, and it is the ambition of physi-
cal science to reduce all its laws to a quantitative expres-
sion. The circumstance that arithmetical calculations
and geometrical propositions admit of such an extensive
application to it the laws and structure of the universe, is
a clear proof that quantity is one of the principles which
impart to its order and stability. It is pleasant to
notice that He who hath given to quantity so important
a place in the structure of His works, hath also allotted
to the faculty which takes cognizance of it an equally
high place in the constitution of man. There is not a
more obvious correspondence between a weighing ma-
chine and the goods to be weighed out by it, between a
measuring vessel and the articles to be measured by it,
than there is between the mental capacity to discover the
relations of quantity, and the significant numbers and
proportions which everywhere occur in nature.
(4.) The Faculty which discovers the Relations of
Active Property. — We cannot, as it appears to us, know
either mind or matter, except as exercising properties.
Mind exists " only as it energizes/' In looking into the
soul at any given time, we find it ever changing, ever busy.
In all our apprehensions of matter, whether original or
acquired, it is known as moving or as exercising some
active quality in reference to us or to other objects.
460 THE FACULTIES
Proceeding on this original knowledge, we are impelled
by a native faculty to compare the various active opera-
tions of material substance, and are thus enabled to dis-
cover what its properties are, what is their nature, and
their rule. As we detect the relations between the vari-
ous actions, we refer one set of them to the law of gravi-
tation, another set to the laws of chemical affinity, and a
third set to the vital forces. Taking some one of these,
say the law of chemical affinity, we proceed to farther
distinctions and classifications, and we arrange substances
into groups according to their more prominent properties.
It is interesting to notice that we have now types and
homologies of a deep meaning in chemistry as well as in
natural history. The importance of the mental capacity
under consideration is greatly magnified by the discovery,
in our day, by Mr. Grove and others, that all the physical
forces, light, heat, chemical action, electricity, galvanism,
and magnetism, are correlated, and have mutual actions
and re-actions.
As the result of the exercise of these faculties of com-
parison, we have —
Generalization. — The number of particulars pre-
sented to our notice in the world, if they cannot be
described, with Plato, as infinite, may at least be said to
be innumerable, and the mind would feel itself distracted
were it obliged to carry them about with it ; and so says
Locke — "To shorten its way to knowledge, and make
<3ach perception more comprehensive, the mind binds
them into bundles." In doing so, it notices how certain
objects are alike in this respect, that they possess certain
attributes in common ; — they are of the same shape, or
they are spread over the same time, or they are alike in
respect of number, or they are of the same colour, or
have some other property in common. The things thus
WHICH DISCOVER RELATIONS (CORRELATIVE). 461
resembling each other, thus correlated, are put into a
group or class, which will include an indefinite number
of other objects, indeed all others possessing the common
attribute or attributes. "To be of a sort," says Dr.
Thomas Keid, "implies having those attributes which
characterize the sort, and are common to all the indivi-
duals which belong to it. There cannot, therefore, be a
sort without general attributes, nor can there be any con-
ception of a sort without a conception of those general
attributes which distinguish it."
There is a strong disposition in all minds to notice
the agreement of objects, and to give a unity to the
many, by assorting them into groups ; and in the case
of some, and these usually the minds of noblest mould,
it becomes a strong passion. " This impulse of the
human mind to generalize," this " inductive propensity,"
as Sir John Herschel calls it, is a characteristic of the
higher scientific intellects which often, indeed, carry it
too far; still, as Bacon, who warns them against these
excesses, remarks, " those who are sublime and discursive
put together even the most subtle and general resem-
blances."
There is thus, on the one hand, a tendency in the
human mind to observe relations, and especially resem-
blances, and by them to group objects into classes. But,
on the other hand, the phenomena around us have many
and comprehensive relations one towards another, afford-
ing befitting exercise to the intellectual faculty, and invit-
ing it to dispose all individuals into systems, and connect
all nature into series. Among all natural, but especially
amono- all organic objects, there are groups or classes
formed, altogether independent of a mind to observe
them. There arc species and' genera, and orders and
kingdoms — there are homotypes and homologues in
462 THE FACULTIES
nature, whether we take notice of them or no. In con-
structing natural science, we are not to create classes by
an exercise of our own ingenuity ; classes are already
formed, and we are to discover and not invent them. In
every department of natural science, it is imperative on
us to look to the natural grouping. An arrangement
which does not proceed upon it, however ingeniously
contrived, may be characterized as artificial, even when
it is not denounced as arbitrary and capricious, and will
seldom turn out to be of much scientific or practical value.
But when the naturalist has been able to seize the dispo-
sitions made for him by nature, or rather by the Glod of
nature, his classifications being natural, will also turn
out to be available for the accomplishment of a great
number and variety of ends. Every character in such
an arrangement will be significant, that is, the sign of a
great many other qualities with which it invariably co-
exists ; and the arrangement will be found not only to
be convenient, but instructive ; not only aiding the me-
mory in retaining what we know, but disclosing other
truths, and widening immeasurably the boundaries of our
knowledge.
This account of the correspondence between the classi-
fying aptitude of the mind and the classes in nature, is
fitted to save us from both of two opposite extremes. It
shews us, on the one hand, that the mind is not a mere
mirror, reflecting the objects passing before it simply as
they pass before it. The mind brings with it to the
investigation high capacities, a power of separating the
most complex objects into parts for more especial con-
templation, of discovering resemblances among objects
very dissimilar in most respects, and of devising hypo-
theses to account for the phenomena which present
themselves, usually in the most scattered manner or in
WHICH DISCOVER RELATIONS (CORRELATIVE). 463
most singular combinations. The relations which unite
the objects in nature are often of the most recondite cha-
racter, and it requires the very sharpest subtlety to bring
them forth to view, and the highest invention to propose
the truth which is to solve the enigma. But, on the other
hand, we are. never to look on the mind, in the con-
struction of science, as creating laws which are not in
nature itself. Dr. Whewell everywhere speaks of the
mind, in scientific inquiry, as " superinducing" upon the
facts, " from its own ideas," something that is not in the
facts. " The facts are known, but they are insulated and
unconnected, till the discoverer supplies, from his own
stores, a Principle of Connexion. The pearls are there,
but they will not hang together till some one provides
the string."* To us it appears that the true statement
rather is, that the mind is so constituted as to be able — ■
which is often very difficult — to discern all that is in the
facts. The law is in the facts, whether we observe it or
no, but it often requires much trained sagacity to detect
it. True, the class cannot with propriety be said to be
in the individual phenomena ; it is the law of a large
body of phenomena which have an aggregate of common
qualities, each one of which is a sign of all the others.
We have in nature not only the " pearls," but the
" string," otherwise they would not hang together as they
do ; but the string is often of a very subtle nature, and only
to be discovered by the most penetrating intellect. The
account which we have given shews us, on the other hand,
how vain all attempts must be to reach the secrets of na-
ture by a priori cogitation. The mind, in its widest range,
is a creature, not a creator ; it is cognitive, and not crea-
tive. It has an eye fitted to see ; but if that eye will go
beyond its office, and produce what is not to be seen, that
* "Whewell's Philosophy of the Inductive Sciences, vol. ii. p. 4S. Sec also Aph. xi.
4:64 THE FACULTIES
which is thus conjured up will he a phantom, an illusion,
deceiving the eye which created it. True, it can devise,
and ought to devise hypotheses, hut it should only he to
Drin determine very precisely the nature of the mental ca-
pacity which prompts us, on discovering an event, to look
for its cause. All that is necessary for our argument is,
that the talent and inclination be regarded as native, and
this it shows itself to he by its universal operation, and
its constant craving. The majority of thinkers deserving
the name of philosophers, have regarded the mental
principle as not only an original capacity and disposition
of the mind, but as a fundamental law of the intelligence,
which insists not only that all effects known to us have in
fact had a cause, but that every given effect must have had
a cause. This view seems to us to be the correct repre-
sentation. On the discovery of any particular effect the
mind is led intuitively to look for a cause.* This is not a
principle gathered from experience, it is rather the prin-
ciple on which we proceed in gathering experience. Some
may say that having invariably observed that every event
has had a cause, we generalize our experience, and con-
clude that every effect has had a cause. But the infer-
ence would by no means be legitimate. Suppose our
experience to be that we had seen a spark ignite gun-
powder one hundred times, there would be a mighty gap
between this and the conclusion that it must do so the one
hundred and first time, and the one thousandth time, and
so on for ever. A finite, though it be a uniform experi-
ence, cannot authorize us to rise to a universal and ne-
cessary truth. The experience of all civilized men for
ages, that swans are white, did not entitle them to argue
* We put the axiom In tins form, because we do not believe that causation vises up
Instinctively in the mind as an abstract or general notion, or that it is consciously before
(ho mind as a general axiom or principle; it is in the mind simply as a law of its opera-
tion leading it, on an individual eifect being presented, to seek a cause. (See Method of
Divine Government, 4th edit. p. 508, and Appendix.) The objections current in England
against original mental principles, apply merely to certain extravagant doctrines about
lunate, or a priori ideas.
470 THE FACULTIES
that all swans are white, and must he white, and accord-
ingly there was nothing inconsistent with previous ex-
perience in the discovery of black swans in Australia.
All human experience shows that crows are black, yet
there is no law of our mental nature leading us to be-
lieve that crows must be black in the planets Juno or
Jupiter. But it is very different with the belief in causa-
tion, (as we have explained it above ;) there is something
in our very intelligence which prevents us from believing,
or so much as thinking, that anywhere, in any planet, or
sun, or star, or nebulous matter, there can be an event
without a cause. We have only carefully to notice the
operations of this native principle, to find that there is a
feeling of universality and necessity attached to all its
exercises. And as the mind, on the one hand anticipates
and expects that every effect must have a cause, so it
finds on the other hand, in its experience, that all things
in earth and heaven are in unision with the internal
principles. The intuitive expectation has ever a corre-
spondence in the external reality.
The account which we have given of the intuitive
belief, shews us at once that the internal principle does
not entitle us to proceed in the investigation of nature
by a priori speculation. For while intuition impels us
on the discovery of an effect to anticipate a cause, it does
not reveal to us what that cause is. The actual cause
must be detected by experience, and thus we are thrown
back upon induction as the only means, after all, of
penetrating the secrets of nature.
Such, then, is the account which we are disposed to
give of the relation between the laws of our intellectual
nature and the laws of the external world. The German
metaphysicians have discovered this correspondence be-
tween the subject and object, as they express it, and they
WHICH DISCOVER RELATIONS (CORRELATIVE). 471
have often dwelt upon it, but they have given, as it ap-
pears to us, a mistaken representation of it. Some of
them are accustomed to speak of it as an antithesis, an
antithesis between subject and object, between matter
and form, between ideas and experience. They are very
fund of comparing it (I think very unhappily, as the two
are very different) to the polar forces which are found
to operate in the material world, and they call the one
the positive and the other the negative pole. When
thus stated, we have a dualistic view of nature. But a
bolder set of thinkers, following out the same method,
have found out a synthesis to reconcile this antithesis.
Discovering a relation between the two poles, they have
reduced the duality to a unity, and resolved all things up
into one Absolute Existence. The general result of all
this heterogeneous combination proceeding from a con-
fusion of thought, is a hideous pantheism, in which the
existence of God is affirmed, but His existence separate
from the universe is denied. These speculators would
account for the correspondence between the internal ope-
ration and the outward objects by supposing them to be
correlated parts of one whole. Fichte represents the
internal power as creating the external object, which,
according to our view, it simply observes. Schelling
conceives them to be necessarily parallel developments of
one ethereal essence developing itself, whereas they ap-
pear to us to be parellelisms produced by Him who hath
instituted both. Hegel resolved them into a unity of
logical forms, whereas they are one simply by reason of
the unity of the Divine Counsel. We must return to
this subject, and devote a separate section to it. These
views, under whatever form they may appear, and how-
ever imposing the nomenclature in which they are
clothed, and however formidable the array of logical
472 THE FACULTIES
forms in which they may be set forth, are the wandering's
of great minds, which will not condescend to proceed in
the method of induction, and, having set out in the
wrong way, and with principles not carefully inducted,
are going the faster and the further wrong, the quicker
and more vigorous their march.
There is, undoubtedly, a relation between the internal
and external, between the subjective and objective, but
it is not a relation of antithesis but correspondence ; and
this correspondence is to be traced, not to any identity,
not to any connexion in the order of things, not to any
logical connexion, but to the adaptation of the one to
the other by Him who hath created both, and, in creat-
ing both hath suited the one to the other. The mind,
as the contemplator, is so constituted as to be able to
attain a knowledge of the thing contemplated, and the
thing contemplated is so formed as to suit itself to the
intellectual nature of the being who has to contemplate it.
There is here a correspondence between two things, so
far independent in themselves, which I can ascribe only
to the unity of design on the part of Him who hath
created both, and given to each its nature and its laws,
and these in exquisite adaptation the one to the other.
We can conceive a world without any such correspon-
dence, a world in which the intellects of the inhabitants
might have no capacity to discover relations among the
objects falling under their notice, or in which relations
among the objects might in no way correspond to their
intellectual aptitudes. It is conceivable, on the one
hand, that the relations might have existed in all their
significance, but have remained unknown characters —
like the mysterious wTritings on the rocks of some eastern
countries, which no living man can read — in consequence
of no one having the capacity to decipher them ; it is
WHICH DISCOVER RELATIONS (CORRELATIVE). 473
conceivable, on the other hand, there might have been
the intellectual power and inclinations, and yet that such
relations might not have been found in nature, or found
only to show that they are of no significance. In the
one case, there would have been an inscription without
the means of deciphering it, in the other, a key with
nothing to interpret by it. In the co-existence of the
two, we have, on the one hand, a power of reading the
symbols, and, on the other hand, a wondrous book spread
out before us full of the highest instruction. The con-
sequence to man is, that instead of being a stranger, a
wanderer, and an outcast, as he .must have been in a
world in which there was no such correspondence, he
feels himself to be so far at home in every domain of
nature, with faculties fitted, if only he exercises them pro-
perly, to discover those laws which give its unity and
connexion to the Cosmos, and help him, if he have faith,
upward to the contemplation of Him who hath insti-
tuted them in an all-comprehensive wisdom. " This
also cometh forth from the Lord of hosts, who is wonder-
ful in counsel, and excellent in working."
SECT. III. THE ASSOCIATION OF IDEAS.
Every one has an easy mode of satisfyiug himself that
his thoughts do not succeed each other at random. Let
him, by an act of reflective memory, go back upon the
ideas which have passed through the mind in any given
period : he may take the time when they seem the most
desultory and unconnected, and he will find that the one
has led on the other, like a string of birds floating through
the air. Or let him, by self-consciousness, watch the
train as it moves along, and he will find that every
thought is related to that which precedes it, not by a
474 THE ASSOCIATION OF IDEAS.
material bond, like the carriages on a railway, but still
by ties which can be discovered. A few minutes ago he
may have been musing on home, and friends, and com-
forts, but now his thoughts are in a far distant land,
wandering amidst extended swamps, and burning heat,
and fearful malaria. At first sight it might seem as if
there could be no possible relation between the two men-
tal states : it might look as if the mind had leapt from
the one region to the other without an intermediate step.
But he has only to recall the whole train to discover that
there has been a continuous transition from the one to
the other. He was meditating on home and friends ;
but one of those friends has been called away from this
world, he went to a distant land to earn an honourable
independence, and there he fell a prey to an unwholesome
climate produced by heat and damps. And suppose that
he allow this last thought to run on in its natural course,
he may find it carrying him up to the heavens, there to
indulge in meteorological speculations, and these sug-
gesting scientific principles, which bring him back to his
own land and to his younger years, when he was first
made acquainted with these principles, and to the very
friends of his youth, and the home whence he started on
his wide excursion. Throughout the whole of this cir-
cuit, every thought has been in some way related to that
which has gone before, and to that which has come after.
We owe to Aristotle the first attempt to classify the
relations according to which our mental states succeed
each other. According to the usual interpretation of his
language, he represents our thoughts as associated by
similarity, by contrast, and by contiguity.* To discuss
* According to Sir William Hamilton, (see Note D** appended to his edition of Eeid.)
Aristotle first announces one universal law, and then three subordinate ones. The one
universal law is : Thoughts which have at one time, recent or remote, stood io each othei
THE ASSOCIATION OF IDEAS. 475
the various theories which have been propounded since
his days, would carry us into very irrelevant matter.
Without entering upon any subtleties or disputed points,
we take up the associations of thought in the two forms
in which they present themselves most obviously to our
view, that of Repetition and that of Correlation.
In Repetition, a thought is followed by the very same
thought with which it was previously associated. Thus,
on the first line of a song with which we are familiar
being recited, the mind is apt to run through the whole.
This is the simplest and lowest form of the associative
power. It is apt to be strongest in children, who are
able, in consecruence, to repeat what they have heard or
read more readily than persons farther advanced in life,
and whose thoughts are disposed to obey a higher law of
succession. For there is a higher form assumed by men-
tal association, less or more, in all minds, but most of all
in minds possessed of firmer intellectual grasp. Things
between which there has been a relation discovered may
suggest each other. No matter what the relation has
been, whether one of those mentioned by Aristotle or any
others, it ever afterwards combines the things correlated
in our minds, and the one tends to bring up the other.
This is law of Correlation.
We have already given what appears to us to be, upon
the whole, the best classification of the relations which
the human mind can discover. It can discover, wre have
said, the relation of comprehension — that is, of the whole
to its parts, and of the parts to the whole. Now, when-
ever such a relation has been noticed, the part will sug-
in the relation of co-existence or immediate consecution, ht the enemies of
his country. There are children, whose first lessons in
480 THE ASSOCIATION OF IDEAS.
geography, learned from a mother's lips, will be about
these wild heights, and the blasting storms which raged
around them — for there it was that the father breathed
his last. And why do men's minds wander so often to
these scenes ? it is because their feelings have become
interested in them, and emotion has the power of
preserving, as in amber, whatever has been imbedded
in it.
Now, let us mark how these two laws aid sciencific
men in their pursuits. The attention which they have
given to the subjects which engross them ; their fixed de-
terminations regarding them. ; the efforts which they have
made to master the difficulties ; their very disappoint-
ments and failures — all these tend to bring the objects
more constantly before them, that they may fully exhibit
themselves, and reveal all their truth. Then, their ori-
ginal tastes, and their acquired habits, the result of associa-
tion, cause them to warm as they advance, and now their
hearts are as much interested as their heads in their pur-
suits. The botanist comes to love the plants, the zoolo-
gist the animals, and the astronomer the stars, which he
has often and anxiously watched, and scientific men gen-
erally feel, when engaged with their favourite pursuits,
as if they were surrounded by friends and companions.
But as, when we truly love our friends, we find ourselves
frequently thinking of them, so, those who are engaged
in the study of nature dwell habitually among their
cherished objects, and the images of them start up every-
where to delight and instruct, to furnish new examples
of old laws, and suggest new laws not previously dis-
covered.
THE .ESTHETIC SENTIMENTS. 481
SECT. IV. TUTS. .ESTHETIC SENTIMENTS.
It may be safely affirmed that no one has been able
to give a complete account of the nature of Beauty.
Pleasant are the glimpses which not a few have had, but
to no one has she fully revealed her charms. We have
many valuable contributions towards a correct theory, but
we are yet without a thorough analysis or a full exposi-
tion. ' We are to attempt no systematic discussion of a
subject so interesting from the nature of the objects at
which it looks, and yet shewing itself to be so subtle
and retiring when we would advance towards it. It is
very obvious that, in the judicious treatment of the sub-
ject, there should be a distinction drawn between the
object which calls forth the feeling and the feeling called
forth. We are to content ourselves with shewing that
there is a correspondence between the two, and the com-
ponent parts of each. Here, as in every other province
of God's works, we find the confluence of a number of
streams ; only, in the case of beauty, they are so blended
that it is impossible to trace each to its source.
I. Vigorous efforts are being made, in the present day,
to find out in what physical beauty consists. These at-
tempts have so far been successful. It has been demon-
strated that there are certain distributions of colours
which are more agreeable than others. Certain colours,
if placed alongside of each other in the decoration of a
house, or a piece of dress, are felt to produce a pleasant
impression. But we have shewn that these juxtaposi-
tions of colours are frequently met with in the plant, in
the plumage of birds, and in the sky. There is here a
correspondence between the external world on the one
hand, and our organization bodily, and probably mental
also, on the other.
21
482 THE ESTHETIC SENTIMENTS.
Endeavours are also being made to find out the law
of harmonious forms. Not having fully examined the
subject, we are not prepared to say how far they have
been successful. But we are persuaded that such in-
quirers as Dr. M'Vicar and Mr. Hay are on the proper
route, and that, sooner or later, there will be detected
certain laws of beauty in form, capable of mathematical
expression. But it is to be carefully noticed, that even
when scientific research shall have established all this, it
has not fully explained the phenomena of beauty. For
the mental sentiment, of which we are conscious, corre-
sponding to the physical object which excites it, is as
wonderful as the object which calls it forth ; indeed, the
most remarkable feature of the whole phenomenon is the
adaptation of the one to the other.
II. We are not to speak confidently on so intricate a
subject, but it appears to us that there is a feeling of
beauty resulting from certain exercises of the intelligence,
(we are sure that there is a feeling of beauty awakened
by certain moral ideas.) This emotion issues when the
mind, in contemplating objects, discovers spontaneously,
without will and without effort, a number of seemingly
intended relations of one thing to another. There has
been a striving after the expression of this truth by deep
thinkers in different ages. According to Augustine,
beauty consists in order and design ; according to Hut-
cheson, in unity with variety ; according to Diderot, in
relations. Glimpses of the same doctrine appear and
disappear in the writings of Cousin, M'Vicar, and Rus-
kin.* There is a sort of beauty in a large combination
of independent means to accomplish one end, and in the
co-agency of numberless causes to work one effect, —
* Cousin on the True, the Beautiful, the Good; M'Vicar on the Beautiful, &c, (1837;)
Raskin's Modern Painters, vol. ii. sect. i. chaps, v. v'.
THE ESTHETIC SENTIMENTS. 483
provided always that the end be not malevolent, or the
effect trivial. There is a beauty in certain well-arranged
forms, perhaps also in certain recurrences and propor-
tions. It is said that there is a beauty in certain regular
rectilinear figures, such as the triangle, the parallelogram
and square, and it has been shown that these regulate
not a few forms of beauty. This seems to us, however,
to be only a partial expression of the truth ; we think
that it needs a complementary truth to be added. The
feeling of beauty is called forth only when, along with an
observable regularity of figure, there is something to in-
dicate that there has been more than mechanism at work.
If the form be too evidently regular, there is little or no
emotion excited. On the other hand, if the figure be
irregular throughout, there is no feeling of beauty. But
if there be a regular figure, such as a triangle, at the
basis of the whole, with curvilinear departures to set it
off, or if there be rhomboids set in spirals, as on the sur-
face of cones, then the a?sthetic sentiment is called forth.
This general view is illustrated and confirmed by the
pleasure which is felt in rhyme and in verse of every
description, indeed, in all forms of poetry, ancient or mo-
dem, eastern or western. All kinds of poetry agree in
presenting repetitions, parallelisms, balancings, corre-
spondences of some description. The mind is excited,
and its admiration is called forth, when it finds the varied
thoughts and feelings grouped under correlations of sound
or sentiment, which exercise the intellect, and aid the
natural flow of association, which proceeds, we have
shewn, according to correlation. There is a similar plea-
sure excited by the tropes, figures, apposite allusions,
comparisons, metaphors, contrasts, which are ever ad-
dressing themselves to us in more adorned prose, such as
that of Plato, of Jeremy Taylor, and of Edmund Burke.
484 THE .ESTHETIC SENTIMENTS.
But the correlations of poetry are limited in range
compared with those which meet us everywhere in the
kingdoms of nature. In all organic bodies there is, along
with more or less variety, a symmetry or likeness of side
to side, and also a repetition of similar parts ; and in
higher organisms, there are more complex and recon-
dite correspondences.
In plants, there are regular lines and definite angles
in the framework, but meanwhile the bounding lines are
always curves, all the more beautiful that they are not
the more regular curves, but curves of great freedom of
sweep. We have found it interesting to notice, that in
the leaves of many plants there is a series of visible tri-
angles. These triangles are formed in the upper part
of the leaf by the midrib, by the lateral vein, and by a
line drawn from the apex of the leaf to the top of the
midrib. It has been affirmed that there is a peculiar
significancy in the right-angled triangle ; it exhibits most
observably a unity with variety ; and we have noticed
that in many plants the angles formed by the lateral veins
in the upper part of the leaf, and a line drawn from their
apex to the apex of the leaf, is a right angle. There is
a series of similar triangles in the upper part of the coma
of many trees. Yet every vein and branch, and the out-
line of every leaf, and of the coma of every tree, is not a
straight line, but a curve with a graceful sweep, that is,
a sweep which still maintains an observable regularity.
The triangle would be stiff and formal without the curve,
and the curve would be eccentric without the triangle ;
the beauty arises from the union of the two.
There is beauty in the spiral arrangement of the appen-
dages of the plant, and in the crossing of the spiral lines
on the surface of the stem, and of many fruits. Of the
more regular curves, the spiral combines in itself most
THE ESTHETIC SENTIMENTS. 485
evidently the two principles of unity with variety, that
is. the greatest number of visible correlations ; and it is
interesting to notice, that this figure is perhaps the most
universally prevalent regular figure in nature, being seen
in shell-fish, in plants, and in the starry heavens. There
is a visible beauty, too, in the regular flower-cup, with
the petals all alike, surrounding and guarding a common
centre, and each with curvilinear outline. There is no
less beauty in the irregular flower, with one of its petals
standing out from the rest, but this not by chance or by
oversight, for in order to enable it to counterbalance the
others, it has, we have shewn, a richer colouring.
In the animal frame the relations are more numerous,
but at the same time, as becomes the higher subject,
more manifold, and not so easily noticed. Whatever
disadvantage might arise, from the latter source, to minds
of limited intelligence, is counterpoised by the Life which
distinguishes the animal from the plant. The plant
being soulless, must have a meaning given to it by its
regular shape and regular divergences. The soul of the
animal, on the other hand, is sufficient to impart to it a
concentration of purpose, with a never-ceasing activity
and change.
Still there is a beauty in the forms of the animal.
Mr. Hay thinks he has found triangles regulating the
framework of the human body. But it should be
carefully noted that no such angular figures strike the
eye in the rounded body of man or woman. There are
indeed ratios and proportions carefully attended to in
the construction of the human frame, and perceived un-
consciously by the mind, and these doubtless give the
unity to the body. But these would not kindle a feeling
of beauty, (they excite no such feeling in the skeleton or
in Mr. Hay's plates,) unless they were relieved by rounded
486 THE .ESTHETIC SENTIMENTS.
forms and flowing curves. The feeling of beauty is
raised neither by the one nor the other, but by the
happy marriage-union of the stronger with the more
flexible.
It will not be understood from this statement that we
look upon the perception of beauty as an intellectual ex-
ercise ; what we mean is, that the intellectual exercise
may lead on to it. That there is need of some intellec-
tual perception in order to the sense of beauty, is evident
from the circumstance that nations and persons low in the
scale of intelligence have little sense of beauty, and what
little they have is awakened by the simplest forms of
beauty. The sense of beauty is a sentiment, and not an
act of the understanding, but it is the reward which
God give's to the intelligence when contemplating the
noblest of His works. Not even that it issues simply
from the intellectual act, it proceeds from the intelligence
contemplating those designed relations which appear in
the objects.
If there be any truth in these views, they lend an em-
phasis and significance to much that we have been esta-
blishing throughout this volume. The sense of beauty
in the case of a vast number of organic objects is called
forth by the very union of typical form and intended
modifications ; by the special end being in conformity
with a general plan. Every one of the correspondences
we have been tracing in the plant and animal may, when
taken along with the designed departures, be the means
of exciting admiration and a sense of beauty. Those
who experienced the feeling, may not be able to lay bare
the principle on which it proceeds, but nevertheless they
perceived the plan and the end, and the emotion sprang
up spontaneously.
And here it is instructive to notice how the class of
THE .ESTHETIC SENTIMENTS. 487
sesthetic emotions are meant to lead on our minds from
creation to the Creator. For it is only when there is such
a correspondence among objects as might be designed
that the emotions are awakened. The whole exercise of
mind is thus fitted, and we believe intended, to draw us
on to the perception of design. It is too true that the
thoughts of many are arrested when they would run in
this direction. The assthetic emotions are cherished and
cultivated by many who spurn away every sentiment of
godliness. Alas, it is because a deeply seated ungodliness
is staying the proper outflowing of the soul ! But were
it not that men " restrain prayer," every perception of
the beauty of natural objects would express itself in a
hymn of praise to the Maker of them all. The feeling
excited by the beautiful is the fire which should kindle
the sacrifice into a flame rising to heaven.
III. The theory of Alison, followed out and illustrated
by the late Lord Jeffrey, which refers all beauty to asso-
ciation of ideas, was never favourably received by artists,
and is now abandoned by all metaphysicians. But while
the doctrine of association cannot explain every phenom-
enon connected with the perception of beauty, there is
much that it can account for, and which can be accounted
for in no other way. When living in a rural district, we
hear on the Sabbath the sound of a bell rising in the
midst of the stillness, and we say how beautiful ; but we
feel in this way not so much because of any pleasure
which the sound may give to the bodily organism, (for
the sound may rather be grating in itself,) but because
it is associated with the idea of Sabbath peace, and the
blessing which the Sabbath diffuses. The association of
ideas alone can explain such a phenomenon as this, a
sound or sight rendered pleasant by reason of the delight-
ful feelings which cluster around it. There is a still
488 THE .ESTHETIC SENTIMENTS.
more important part of the complex state of mind which
can be accounted for in no other way ; we allude to the
prolonging of the pleasure, and the variety of the plea-
sure communicated by the image upon image, the feel-
ing upon feeling, all agreeable and exciting, raised by
certain objects, such as a cheerful countenance, a plain
covered with grain, and a river rolling along amid fer-
tile banks. It is in the union of the two, the original
feeling of beauty, and the association with it of other
pleasant feelings, that we are to find the full explana-
tion of the phenomenon. By the one we account for
what is fixed in aesthetics, for the uniformity of men's
judgments in matter of taste ; by the other, that is, by
the difference of the associated feelings, we can account
for what is variable, for what differs, in the case of differ-
ent individuals. And wre do not know whether to admire
most that constitution of our nature, by which there are
certain points of agreement in all men's tastes, which
renders it possible for them to sympathize with each
other, and by which a science of aesthetics is rendered
possible, or that variety of tastes which gives to every
man his individuality, which secures that all do not run
after the same object, and that there is scarcely an object
which may not be made attractive to certain minds.
But then, this very association of ideas in its special
connexion with the beautiful, requires itself to be ac-
counted for. The views which we have propounded may
aid us in doing so. The feeling of beauty is awakened
by means of discovered correlations, and each of these
ramifies into collateral topics. Then all the correlations
point to Design, and Design is a mental quality alluring
on the mind to a thousand pleasant topics. Hence the
retinue of thoughts ready to rise up, and prolonging the
feeling as by answering echoes, and calling in images to
THE ESTHETIC SENTIMENTS. 489
aid it from every object in nature or in art, which may
have fallen under the notice.
IV. We may notice some other and allied feelings, so
far as they are awakened by objects in nature. Some
maintain that there are plants and animals which may
be described as ludicrous. If there be, it is because they
are addressed to the sense of the ludicrous in us. The
feeling of the ludicrous seems to us to be awakened by
the discovery of an unexpected relation between objects
in other respects totally dissimilar. This, too, seems to
point, but in an opposite way, from the sense of beauty
to design, to design in bringing things unlike into one
category. There are, certainly, grotesque and fantastic
objects, both in the animal and vegetable kingdoms,
which call up the feelings of the ludicrous. We smile
when we observe how like the owl is to an old man or
woman with excessive pretensions to wisdom ; how like
certain orchids are to beasts, birds, or insects ; and how
admirably the monkey mimics the movements of hu-
manity.
V. There are scenes met with on our earth which are
expressively called picturesque. They seem to be pecu-
liarly addressed to the imagining power of the mind ; they
are picture-like, and raise a vivid picture of themselves
in the mind ; such as the jagged mountain ridges, the
peaked promontories, the perpendicular rocks. The
mass of objects on the earth are not of this exciting
character. Just as the ground colours of nature are soft,
or neutral, so the earth's common scenes are irregular, or
simply rounded in their outline. Yet here and there
there arise picture-like objects from the midst of them,
to arrest the eye and print themselves on the fancy. It
may be noticed, that the grass and grain of the earth
raise up their sharp points from the surface to catcli our
21*
490 THE .ESTHETIC SENTIMENT3.
eye. A still larger proportion of objects above us, and
standing between us and the sky, have a clear outline or
vivid points. This is the case with the leaves and the
coma of trees, and with not a few rocks and mountains.
Rising out from quieter scenes, they enliven without
exciting the mind, and tend to raise that earthward
look of ours, and direct it to heaven, to which they
point. >
VI. Before closing this paragraph, we must allude to
another kindred subject, the Sublime, so far as natural
objects are fitted to raise the feeling. Visible things can
here do nothing more than aid the mind, which uses them
merely to pass beyond them.
The feeling of the Sublime is acknowledged on all
hands to be intimately connected with the Idea of the
Infinite. In the formation — or rather, in the attempt at
the formation — of this idea, the mind shews, in a very
striking manner, both its strength and its weakness. In
expanding any image. spatially, it finds itself incapable
of doing anything more than representing to itself a vo-
lume with a spherical boundary. In following out its
contemplation in respect of time, the image is of a line
of great length, but terminating in a point at each end.
But where the mind shews its weakness, there it also
exhibits its strength. It can only imagine this bounded
sphere and outline, but it is led to believe in vastly more.
It strives to conceive the Infinite, but ever feels as if it
were baffled and thrown back. But while the mind
cannot embrace the infinite, it feels, at the place where
it is arrested by its own impotency, that there is an in-
finite beyond. Looking forth, as it were, on the sky, it
can see only a certain distance, but is constrained to be-
lieve that there is much more beyond the range of the
vision — nay, that to whatever point it might go, there
THE ESTHETIC SENTIMENTS. 491
would still be a something farther on. " If the mind,"
says John Foster, " were to arrive at the solemn ridge
of mountains which we may fancy to bound creation, it
would eagerly ask, Why no farther ? — what is beyond ?"
It is here that we find the origin and genesis of such idea
as the mind can form of the infinite, and of the belief, to
which it ever clings, in the boundless and eternal.
Now, whatever calls forth this exercise of mind and
the feeling of awe awakened by it, may be described as
sublime. So far as picturesque objects are concerned, the
imaging power of the mind rejoices to find that it can
print them upon its surface. But there are objects which
it tries in vain to picture or represent ; the imaging
power is filled, but they -will not be compressed within
it. Everywhere in nature are there scenes which are
..." Like an invitation in space
Boundless, a guide into Eternity."
A vast height, such as a lofty mountain, is a step to help
us to this elevation of thought and emotion. The reve-
lations of astronomy awaken the feeling, because they
carry out the soul into far depths of space, but without
carrying it to the verge of space. The discoveries in
geology extend the mind in much the same way, by the
long vistas opened of ages — which yet do not go back to
the beginning. Every vast display of power evokes this
overawing sentiment ; we see effects which are great,
arguing a power which is greater. The howl of the
tempest, the ceaseless lashing of the ocean, the roar of
the waterfall, the crash of the avalanche, the growl of the
thunder, the shaking of the very foundation on which we
stand when the earth trembles— all these fill the ima-
gination, but are suggestive of something more tremen-
dous behind and beyond. For a similar reason the vault
492 THEORIES OF CONTINENTAL PHILOSOPHERS AS TO
of heaven is always a sublime object when serene ; we
feel, in looking into it, as if we were looking into immen-
sity. Hence it is that a clear bright space in the sky or
in a painting, always allures the eye towards it ; it is an
outlet by which the mind may, as it were, go out into
infinity.
But whatever may suggest the infinite, there is, after
all,- but one Infinite. The grandest objects presented to
our view in earth or sky, the most towering heights, the
vastest depths, the most resistless agencies — these are
but means to help us to the contemplation of Him who
is "high-throned above all height," whose counsels reach
from eternity to eternity, and who is the Almighty unto
perfection. They are fulfilling their highest end when
they lift us above this cold earth, and above our narrow
selves, to revel and lose ourselves in the height and depth,
the length and breadth, of an Infinite Wisdom, lightened
and warmed by an Infinite Love.
SUPPLEMENTARY SEOT. THEORIES OF THE CONTINENTAL PHILO-
SOPHERS AS TO THE RELATION OF THE LAWS OF NATURE TO
THE LAWS OF INTELLIGENCE.
"We have illustrated, to as large an extent as our plan allows, the facts
which bear upon the relation of the subjective mind to the objective world.
After such a survey, we are in circumstances to examine the theories of
this relation which have been propounded by some of the deeper thinkers
on the Continent of Europe, and especially by some of the German meta-
physicians. It should be frankly acknowledged that we have derived
much new material for thought from the importation into our land of the
loftier speculations of German Philosophy ; but it is not to be forgotten,
at the same time, that there are principles lying at the basis of some of
their systems which would go far to undermine, not only revealed, but
natural religion in all its beneficent forms. Some of the gigantic systems,
which are being eagerly studied by the ardent youth of our land, consti-
tute the chief supports of a pretending pantheism which it is proposed to
substitute for the doctrine of a God possessed of personality, that is, of a
RELATION OF LAWS OF NATURE TO INTELLIGENCE. 493
separate consciousness and an independent wilL Before entering upon
the discussion of these systems, it is proper to state that we are to exa-
mine them only so far as they relate to our own subject, and as they pro-
fess to adduce facts external or internal as evidence in their favour.
In order to understand these theories, it will be needful to trace them
historically from their origin. It was a fundamental principle of Descartes
— so distinguished for the originality and the independence of his thinking
— that tnere existed in the universe two entirely distinct substances, spirit,
whose essence is thought, and matter, whose essence is extension. In his
days, it was a universally acknowledged principle that things which were
like, and they only, could influence each other. This seems to be an un-
founded, or rather a false principle. In this universe, things very unlike
affect each other ; in polar action, like repels like, and things unlike are
attracted to each other. But, being then a universally recognized princi-
ple, we find it acting an important part, in the philosophy of the seven-
teenth century. In particular, it suggested a difficulty which greatly
puzzled the school of Descartes; — How can mind influence matter, and
matter mind ? How does an object, presented to the senses, give rise to
an apprehension of it in the mind ? How is it, that when we will to move
the arm, the arm moves ? It does not appear that Descartes uttered a
very clear or explicit answer to this question, but the reply was given, and
this quite in the spirit of the master, by the disciples, and, in particular, by
the ingenious and devout Malebranche. According to him, matter does
not influence mind, nor mind matter ; the action of matter in reference to
mind, and of mind in reference to matter, is the mere occasion of the forth-
putting of the Divine power, which is the true cause of the effects which
follow. Thus, when we will to move the arm, a present Deity, the source
of all power, actually makes the arm to move. This is the famous doc-
trince of Occasional Causes, as maintained by Malebranche. To us it
appears that God has been pleased to give a delegated power both to mind
and matter, and that there is no greater difficulty in supposing mind to
act on matter, than in supposing matter to act on matter.
These principles and speculations, floating among the reading and
thinking minds of that era, took a deep hold on the meditative spirit of a
glass-grinder at Amsterdam, who had been brought up in the Jewish faith.
The influence exercised by this man — despised and persecuted in his owa
day — upon the whole of the future history of speculation, is one of the
most curious incidents in the whole history of modern philosophy. It is
acknowledged, he argues, that if mind and matter are totally different
Bubstances, they cannot influence each other; but it is very evident, mean-
while, that they have innumerable points of connexion. It is not necessary
to suppose them to be separate substances, they are to bo regarded as
modes of one and the same substance, a substance possessed both of
494 THEORIES OF CONTINENTAL PHILOSOPHERS AS TO
thought and extension. In itself this one substance is Natura Naturans,
possessing all power, and ever developing itself; in the universe it is
Nalura Naturata. This system recommended itself to the mind of Spi-
noza by its simplicity ; it seemed to follow from the acknowledged princi-
ples of the day as to the nature of substance ; and it accounted for the
unity of operation which everywhere runs through nature. It is probable
that Spinoza did not allow himself seriously to contemplate the fatal na-
ture of the consequences flowing from a system which makes evil, even
moral evil, a development or mode of the Divine Being, and denies to man
all free-will, all personality, and accountability to a being different from
himself. Still, he saw and avowed that such consequences did follow, and
was willing to take them as the logical results of a system which had so
much to recommend it to his reason, and which represented the universe
as full of Deity.
Passing over inferior names, we now find the lofty genius of Leibnitz
devoting itself to the solution of the same problems. Proceeding on the
principle that mind could not influence matter, nor matter mind, he sup-
posed that they co-operated so beautifully in consequence of a Harmony
Pre-established between them. Kejecting the atomic theory of matter,
which had found such favour with Bacon, Gassendi, and the majority of
modern physicists, he substituted a theory of Monads. The ultimate prin-
ciples of matter were not, according to him, sluggish atoms, but active
powers. There are two distinct kinds of monads, the one unconscious, the
elements of matter — the other conscious, the elements of mind. These
could not be thought to operate causally upon each other, but still they
acted in unison by reason of relations pre-established by God, the Supreme
and Eternal Monad, between the inferior monads, whereby each monad
acts according to its own principle, and yet acts in harmony with all
around it. Some of these speculations of Leibnitz carry us into regions
where we have really no light to g-uidc us. We are far, at this day, from
being able to determine what are the ultimate elements either of mind or
matter. Some of the principles laid down by him are evidently wrong, as
when he says that the monads, or powers of nature, cannot influence each
other. The agencies of nature, whatever they be, are so constituted as to
be able to operate upon, to affect, and modify each other. But, in these
lofty discussions, there is a truth propounded which can never be set aside,
but which will, on the contrary, appear more and more significant in
every succeeding age. The principle in which we refer is that of pre-
established harmony. We must, indeed, in the first place, affirm, contrary
to the theory of Leibnitz, that the powers of nature, whatever they be, do
stimulate and influence the one the other ; but we must also, if we would
account for the phenomena which present themselves, take along with
us the other doctrine, that they are so constituted and collocated as to affect
RELATION OF LAWS OF NATURE TO INTELLIGENCE. 495
each other, not in a destructive, but in a beneficial manner. Their mutual
action, which Leibnitz denied, implies in itself an adjustment, a pre-estab-
lished harmony; and there are, besides, harmonies proceeding from a con-
currence of independent causes ; and both one and other carry up our
minds to an Intelligent Being appointing, from the beginning, all things
to act in concert.
Such was the state of speculation on this all-important subject when
the profound intellect of Kant was led to meditate upon it. The relation
between the internal and the external, between the subjective and ob-
jective, was his great theme round which his philosophy moved. In
studying his system we have never been able to say whether we should
yield to the feeling of admiration which the logical powers manifested
everywhere, and the important truths unfolded in many places, naturally
call forth, or whether we should not restrain all sucli sentiments as we
deplore the erroneous and dangerous principles which he has been the
means of introducing not only into German, but into European specula-
tion.
Kant saw clearly that we cannot account for human knowledge by
mere impressions from without, that it was needful to have a subjective
power as well as an objective influence. It is his grand aim to unfold
these subjective principles, and in particular, the synthetic judgments d
priori, the judgments pronounced independently of experience, on objects
known by experience. In all knowledge, he says, there is, on the one
hand, an external impression, aud on the other a subjective form; that the
external thing which produces the impression exists, he acknowledges,
but then it is an unknown something. It is at this point that his error
begins. According to our natural cognitions or beliefs, as it appears to
us, the mind is so constituted as to be able to attain a limited knowledge
of the external thing as it is ; but according to Kant, the external thing
is unknown, and there is much in our cognition of it which is given to it
by the mind as it contemplates it. Thus the mind, in looking upon the
external world, perceives it always as in Space or in Time, which have no
objective reality, but are mere forms of the Sense or Sensibility. Again,
the understanding, in judging of the matters of Sense, unites them under
such categories as Quantity, Quality, Relation, Modality, which are not
to be understood as having any external or objective realitj-.
The relation between the subjective and objective may, wo find him
arguing, be conceived to spring from one or other of three causes: — First,
From the objective determining the subjective; Secondly, From the sub-
jective determining the objective; or, TJiirdly, From a pre-established
relation or connexion between them. He then shows how the first suppo-
sition, that of the school of Locke, (as he represents it, confounding it every-
where with the French school of Condillac,) which derives all our know-
496 THEOEIES OF CONTINENTAL PHILOSOPHERS AS TO
ledge from sensation, cannot account for the internal facts. ITe dismisses
the third doctrine, that of a pre-ordained adjustment, in a very summary
manner, neither stating it accurately, nor examining it carefully. His
objections to this middle way are, first, that no end can be seen to such
an hypothesis, and secondly, that necessity would be wanting to the cate-
gories which belongs essentially to the conception of them.0 In reply to
the first, we maintain that the limit to the relation of the objective to the
subjective, is to be ascertained and determined, like its existence," by induc-
tive investigation ; that is, we believe in the relation only so far as we prove
it to exist, that is, find a natural aptitude in the mind on the one hand,
and a corresponding operation of nature on the other. In reply to the
second, we urge, that when there is a feeling of necessity in the internal
principle, there is a universality in the external relation. This is one of
the correspondences which we have traced. But having dismissed the
other two, Kant finds himself shut up into that theory which makes the
mind give its own laws and relations to the objective world. It is thus
that he accounts for the relation of cause and effect, and the harmonies in
the universe ; they are not in the universe itself, they are merely in the
mind, and are thence, as the forms, or categories, or ideas under which the
mind knows all things, projected upon the world. In this system the doc-
trine of final cause, as founded upon the correspondence between the miud
and nature, and upon the harmonies of nature, must necessarily disappear,
for these are not correlations between independent things, but the result
of one principle in the mind itself. It is not difficult, as it appears to us,
to meet this subjective idealism. We are led by the very constitution of
our minds to believe in the reality of external objects ; and to believe in
them not as things unknown, but as things so far known, and known as
professing certain properties; and further to believe, that such relations as
those of quality, and cause and effect, are relations in the things them-
selves. Kant acknowledges that the mind does not create the things, and
on the same ground we maintain that it does not create the properties, the
relations of things; it has a set of powers by which it is enabled to know
them. Deny this, and we deny the very truths of consciousness — the
truths sanctioned by the very constitution of our minds ; and after denying
these, we have no principle left on which to proceed on our specula-
tions, no truth so certain as that which we have set aside. But if we
believe in the existence of external things, and on the same ground in the
reality of the relations of external things, we are obliged as Kant
clearly saw, to believe also, in some "Preformation," between them.
Not that we are therefore to set aside the influence of mind on matter;
for matter is so constituted as to influence the nervous system, and tho
nervous system is so constituted as to excite mental action. Not that we
* Critick of Pure Reason. Analytic of Principles, close of Book I.
KELATION OF LAWS OF NATURE TO INTELLIGENCE. 497
are to deny the separate potency of mind — all that nervous agency does ia
to caU forth the activity of mind, which is so constituted as to know mat-
ter, and the relations of matter. Not that wc are to deny the influence
of matter upon mind in putting it in motion. We must admit, we think,
all these agencies, the action of mind, the action of matter, and their re
ciprocal action. And in order to account for their harmonious action, we
must call in a divinely-appointed adjustment of the two — an adjustment
net independent, as Leibnitz supposed, of their mutual action, but an ad-
justment enabling them to act upon, as well as with each other, so as to
produce consistent and beneficial results. We must, as most important
of all, suppose that there has been a pre-ordained adjustment between the
intuitive laws and beliefs of the mind on the one hand, and the actual re-
lations instituted in the external world on the other.
There was but a step between the doctrine of Kant and that of Fichte,
which professed to carry out the principles of Kant to their legitimate
consequences. Kant admitted that there was an external world; but then
he supposed that the mind gave to it its qualities and relations of Space,
Time, Cause and Effect. It was no violent step in advance which was
taken by Fichte when he alleged that the mind, which was capable of .
creating all the relations of matter, might form matter itself. The whole
external world became in this philosophy a production of the Ego, all its
laws, its order, its harmonies, being given it by the mind itself. In the
progress of these speculations of Fichte, the Ego became expanded, in an
unintelligible, inconceivable manner, into a kind of universal Ego, which
constituted the Moral Order of the universe, and went by the name of
God. Here there was an end, as in the system of Kant, to all final
cause ; but let it be observed that final cause was discarded on grounds
which also set aside all objective truth. Here, too, there was an end to
what had been carefully preserved in the philosophy of Kant, to the per-
sonality of man and to the separate immortality of the soul. It may have
been a discovery of the connexion of this system with his own, which led
Kant in his late years to pray to be protected from his friends. It is not
necessary to subject this system to a critical examination ; in doing so
we should only be wrestling with a shadow. It sets itself against the
fundamental principles of the mind, which announce to us that there is
a reality, independent of ourselves, in the external world.
There was now in the system of Fichte a scheme of pantheism, with
lofty pretensions, and enforced by great beauty of sentiment, set before the
German mind. About this period, certain occurrences which arose out of
a conversation of Jacobi with Lessing, who had a great admiration of
Spinoza, and reported by the former, brought the system of the Dutch
Jew also before the thinking mind of Germany. It was while the Ger-
man philosophic mind was being fermented by the two systems of Fichte
498 THEORIES OF CONTINENTAL PHILOSOPHERS AS TO
and Spinoza, thnt Schelling produced his theory, and irradiated it with
the fascinations of a poetical imagination. According to him, it was
absurd to suppose that the Ego could create all the harmonies of things;
we must go farther back if we would account for the correspondences be-
tween the external and internal. Neither is this relation to be explained,
as Spinoza supposes, by a universal substance possessed at one and the
same time of thought and extension, for this would not account for the
very diverse experiences of subject and object. We must, therefore, go
a step higher, we must go back to the origin both of the subjective and
objective, and there we shall find them identical and flowing out of one
original, living essence, called by the name of God. This self-existent
essence or being develops itself according to a law, and becomes on the
one side the Ego, and on the other the Non-Ego; on the one side the
subject, and on the other the object ; on the one side mind, and on the other
nature. Hence the harmony of the two; it arises from their identity.
The subjective and objective are in such visible correspondence, because
the developments of one and the same principle. Hence the statement
that nature is petrified intelligence, and that mind is conscious reflective
nature. The feeling of beauty in the mind corresponds to beauty in the
world, because both are the unfolding of one eternal power which is at
one and the same time God and the universe. God is lovely, the universe
is lovely, man's soul loves the lovely in nature and creates the lovely in
art, because all are manifestations of the One who is infinitely lovely. It
would be a waste of thought to institute a serious refutation of this specu-
lation, which, taken as a whole, is to be treated as a picture drawn by a
brilliant fancy. In its fundamental truths it is inconsistent with our in-
tuitive knowledge and belief, which announces to us distinctly that we
have a separate personality, that we are not the same with God on the
one hand, or nature on the other. Schelling appeals to an intellectual in-
tuition, which is one with the Divine Intelligence, as capable of gazing on
this identity of existence. But this intuition is acknowledged by him to be
above consciousness; that is, as we reckon it, above the region to which
man's knowledge can reach, that is. in a cloud-land where irradiated mists
may be mistaken for solid bodies. Certain it is, that all the intuitions
which we can discover by consciousness, set themselves against this iden-
tification of ourselves either with the Divine Intelligence or with nature,
this identification of subject and object, of man and God. But with all
its superlative extravagance it contains a truth, a truth not in the systems
of Kant or Fichte ; this is the correspondence between the subjective and
objective, both being represented as real though not independent. Never
was there so beautiful, and, let us add, so true a picture drawn of the
harmony between the beautiful in the mind and the beautiful in nature, as
that which we find in the writings of Schelling and of the disciples of hia
RELATION OF LAWS OF NATURE TO INTELLIGENCE. 499
school. We have here a style of speculation to which the native British
philosophy is a stranger, and which appears irresistibly attractive when
presented to British youth of fervent intellect ; and they are too often
prepared, in their admiration of the mixture of truth contained in the sys-
tem, to embrace the error 'with which it is associated.
It was felt in Germany that the system of Schell'mg, though exquisitely
beautiful, was little better than a speculative rhapsody, when his friend
Hegel, with a much more logical mind, set about amending and systema-
tizing it. We do not propose to give an account of his system. We do
not attempt to fathom its depths or expose its shallows ; for it has depths
in which the tallest intellects would lose themselves if allured into them,
and it has shallows which the most superficial can see and point out. He
is reported to have said, " There is only one person who understands me,
and he does not understand me." Not having the honour to be this per-
son, we make no pretensions to a thorough understanding of Hegel. For-
tunately, we have to consider his system only in one aspect.
In the systems of Kant and Fichte, the relation between the subjective
and the objective lias disappeared, fir the whole is the creation of the sub-
■ective. In the system of Schelling, the relation has reappeared, but has
been accounted for in a most unsatisfactory manner. In the system of
Hegel, the relation is all in all. The subjective has no separate existence,
on the one hand, nor the objective on the other hand ; they exist only in
relation to each other. The relation is here acknowledged, but it is a
relation which does away with the independent existence of the things
related.
Abandoning the intellectual intention of Schelling as a mere gratuitous
assumption, he attempts to show how all things are developed necessarily
by a logical process which is not assumed, but is, in its development, a
proof of its own reality. In following out this process, be begins with the
most general and abstract notions, such as " Idea" and " Being," and
thence develops nature and mind. In all this he reverses the natural order
followed by intelligence, which begins with things individual and concrete
as they present themselves, and thence rises to the general and the abstract.
In doing so, it never for one instant supposes that the abstract or general,
such as " Being," can exist independent of individual things. The abstract
is a part, separately considered, of the concrete whole. The general is the
aggregate of qualities in which individual things agree. It is to reverse
the proper process of thought, to begin, as Hegel does, with the abstract,
the general. It is to contradict the clearest declarations of thought to
deny the existence of the individual, whether subject or object, and resolve
all into a relation. The relations which the mind discovers are relations
among individual things.
According to this system, the All presents a constant evolution of
500 THEOKIES OF CONTINENTAL PHILOSOPHERS AS TO
nothing becoming something, and we have to add, of something falling
back into nothing. In the unfolding of this theory, he represents God
as attaining to consciousness in man, and the whole history of the human
race as a succession of incarnations. At his death, which was occasioned
by cholera, some of his pupils apotheosized him as the noblest of all the
self-conscious developments of Deity. It is easy to see how he accounted
for the harmonies which the mind discovers in the universe. To philoso-
phize on nature, he says, is to rethink the grand thought of creation — it
is to reproduce, from the depths of the soul, the creative ideas of nature.
In a journey which he made to Paris, he was greatly entertained, as ha
discovered everywhere — in nature and in art, in man as an individual, and
in man united in society — confirmations of his system, which widened,
like vapours, to embrace all the agreements and disagreements in exist-
ence.
It might easily be shown that this ambitious and arrogant system de-
stroys all personality — that is, separate consciousness and will — in God,
all personality in man, and that it is inconsistent with human responsi-
bility, and the immortality of the soul as a separate existence. All this
has been dwelt on by the schools which have sot themselves in opposition
to it in Germany. And this argument from consequences should have its
weight, for any s^ystem which sets itself against these truths, cannot be
supported by such evidence as they can adduce in their favour. Again,
various gaps and inconsistencies have been pointed out in it, showing that
it is not so solid a structure as it professes to be. But its fundamental
error lies in this, that it denies the separate existence of individual things
— of the subject on the one hand, and the object on the other. In pro-
fessing to proceed according to the laws of thought, it begins with set-
ting the clearest laws of thought at defiance, and must wander the more
the farther it advances. It is acknowledged, even in Germany, to be a
failure. It fails, in particular, to account for the correspondence between
mind and matter regarded as separate existences.
It should be added, that Herbart met these idealistic views of Kant,
Fichte, Schelling, and Hegel, with great vigour by a realistic scheme, in
which final causes once more have their proper place. But his realism
is professedly a rational system erected on certain philosophic principles,
which may be assailed equally with the grounds taken by those whom he
opposes, and will not find much favour among persons in our country who
have become imbued with the spirit recommended by Lord Bacon, and
followed out, though with but imperfect success, by Locke and Reid. It
is only by proceeding in the inductive method that we can expect fairly
to unfold the subjective laws of mind on the one hand, and the objective
laws of nature on the other, and then discover the relation between them.
In the speculations of all these philosophers, notice is taken of a most
RELATION OF LAWS OF NATURE TO INTELLIGENCE. 501
important class of facts, which have very much escaped the attention of
British writers. But while we acknowledge this, we are convinced, at the
same time, that the correct explanation has not been given by the conti-
nental speculatists. In the days of Descartes and Spinoza, the questions
discussed turned round the action of mind upon material objects, and the
action of material objects upon mind. But from the time of Leibnitz, and
still more from the time of Kant, a new set of questions came to be agi-
tated in regard to the accordance between the laws of the mind within,
and the harmonies of external nature without. Kant and Fiehte referred
this to the mind creating the order which it contemplated, Schelling and
Hegel to the identity of subject and object, of the world within and the
world without. But none of these hypotheses meets the full facts of the
case, nor explains the whole phenomena. Leibnitz, indeed, had a glimpse
of the truth, but failed to represent it fully and correctly. The facts admit
of only oue satisfactory explanation, and this an explanation in strict ac-
cordance with the doctrine of final cause, and implying a specific plan on
the part of an intelligent being.
For mark, that we have, first, a set of internal fads. "We have in the
mind perceptions through the senses ; we have certain intelligent aptitudes,
such as the generalizing propensity, ever seeking to group the objects it
meets with into classes, and causality, anticipating nature, and confidently
looking for certain effects to follow agencies now in operation ; we have
instincts and affections craving for external objects on which to lavish
themselves ; and we have a sense of beauty, longing for scenes of loveli-
ness and sublimity. We insist that these internal facts be not set aside,
but that they be embraced and accounted for in any explanation which
may be offered. It will not do to refer them, with certain French and
British writers, to sensations and impressions from without. They are
evidently powers, instincts, affections, fundamental laws in the mind
itself, making their own use of the influences which may come in from the
external world.
But, secondly, there is a set of external facts. As little are we at liberty
to overlook them. In denying them, we are, in the very act, discarding
the dicta of consciousness, and the very constitutional principles of intel-
ligence in the mind ; and after we have done so, there remains no ground
on which we can reason on this or on any other subject. Here, in this
world which we perceive, are bodies endowed with wonderful properties ;
are objects grouped into classes, and with interesting correlations subsist-
ing between them ; are events causally connected together ; and scenes of
beauty and grandeur. All this must be explained by a hypothesis ade-
quate to meet the case obviously presented, and no part of all this can be
accounted for merely by the inward principles of the mind, except on tho
ground which would make these very principles delusive and a delusion.
502 THEORIES OF CONTINENTAL PHILOSOPHERS AS TO
We have thus a series of facts in congruity with each other within the
mind. We have also a series of facts in beautiful harmony with each
other without the mind. But we have more, there is an accordance be-
tween the internal and external facts. "We have the perceptions of one
sense confirmed by those of another sense. "We have instincts impelling
to certain operations, and we find ourselves placed in a state of things in
which these instincts are gratified, and in being so, perform acts necessary
to our welfare and our very existence. "We have affections general and
special, and we fall in with objects to call them forth, and on which to
lavish 'them. We proceed spontaneously to classify objects according to
certain relations of shape, quantity, and proportion, and actually find them
distributed into groups according to these very principle?. We anticipate
that the same cause will ever produce the same effect, and find our expec-
tations realized every waking hour of our existence. We have aesthetic
tastes, and everywhere there are harmonious colours, and graceful forms,
and lovely scenes to gratify them. As it is not the sound which produces
the ear to receive it, as it is not the eye which creates the light that falls
upon it. so it cannot be the outward harmonies which create the inward
desire to discover them, and the capacity to observe them ; nor the internal
faculties and feelings which create the outward harmonies. Vie are ob-
liged, if we would account for the whole phenomena, to account for both
classes of facts, and the relation between them. This can only bo done
by supposing that one Intelligent Being instituted both series of facts and
their mutual accordance. The facts round which the German philosophy
has been moving, are thus seen to bring us back to the old doctrine of our
British Theology. It is only by calling in a God who designs and exe-
cutes, that we can fully or rationally account for facts, which we cannot
deny without denying our intelligence.
It thus appears that the doctrine of Final Cause, so far from being un-
dermined or shaken by these speculations, is as secure as ever — nay, it
stands forth with new illustrations and confirmations. We are brought
back to what observant minds have noticed from the first, (though they
had not always expressed it correctly.) a concurrence of independent
agencies towards the production of a given end. Hegel is laying down
an utterly mistaken doctrine when (not in words denying final cause) he
speaks of the final cause of a thing being the inward nature of a thing, or
a tiling following its inward nature ; final cause is the co-operation of a
number of independent things to accomplish what is evidently an end.
In particular, there is need of a correspondence of the external and inter-
nal in order to our inward knowledge, and to our experience of the outwad
world. The phenomenon cannot be explained by an internal order pro-
jecting itself upon the external world ; for, as Herbart asks, if it be by
some necessary form of the understanding that final cause is imposed on
RELATION OF LAWS OF NATURE TO INTELLIGENCE. 503
things, how are we to account for the fact that we do not see the final
cause in regard to every occurrence ? How is it, in particular, that we
discover it only in those cases in which we notice a concurrence of agen-
cies acting independently of the laws of thought within ourselves? All
this can easily be accounted for on the supposition that it needs objective
evidence to lead us to discover final cause ; but is inexplicable if the pro-
cess proceeds from a merely subjective principle. But, without pressing
this difficulty, we plant ourselves on ground from which we can never be
dislodged, when we maintain that the outward is real and that the inward
is real, and that there is proof of plan and intelligence in the correspon-
dence instituted between them.
At the close of this review, we find ourselves shut up into a Pre-Estab-
lished Harmony. But it is not the fanciful doctrine of Leibnitz. Accord-
ing to him, no one power or monad can operate upon any other, but each
fulfils its function independent of all others, and yet in harmony with all
others. This seems to us quite inconsistent with what we see everywhere,
the action of objects on each other. The Pre-Established Harmony which
we advocate, pre-supposes the action of matter on matter, of matter on
mind, and mind on matter, and the harmony is manifested in the benefi-
cence of their mutual operation.
This Pre-Established Harmony manifests itself in two forms.
First, Agents mental and material have powers or properties which fit
into each other, and enable them to co-operate in producing consistent and
bountiful results. So far from supposing that they do not act on each
other, we affirm, that they do act, but act in harmony.
Secondly, There has been an original collocation of agents, whereby
concordant results are produced without any reciprocal action. The lily
that grows in one garden, assumes the same forms and colours as the lily
which grows in another garden. The fish of the Old Red Sandstone
epoch had the same general form as the fish which still swims in our seas.
But these correspondences do not arise from any mystic or magnetic influ-
ence of the one upon the other, but because causes have been instituted
and arrangements made, which produce the one in unison with the other.
The comparison of Leibnitz here applies ; the two correspond as two time-
pieces, not because of any mutual influence, but because each has been so
constituted, that it moves in harmony with the other.
We cannot comprehend the harmonies of the universe without admit-
ting and calling in both these principles.
CHAPTER III.
TYPICAL SYSTEMS OF NATURE AND REVELATION.
SECT. I. THE OLD TESTAMENT TYPES.
In looking at any one department of conternporaneciua
nature, we discover that all objects and events are con-
formed to a plan. Organisms differing from each other
in their constituent elements have the same relations of
parts, and differing from each other in use, are cast in
the same general mould. Again, looking at certain de-
partments of successive nature, we find that objects in one
epoch are an anticipation ox prediction of objects to ap-
pear at a later epoch. The science of embryology shows
that there are systematic stages of progression in the for-
mation of the young of all animals. In the Psalm which
celebrates the omniscience of God, this remarkable lan-
guage is employed : — " I will praise Thee ; for I am
fearfully and wonderfully made : marvellous are
Thy works ; and that my soul knowetii right well.
My substance was not hid from Thee, when I was
made in secret, and curiously wrought in the lowest
parts of the earth. thine eyes did see my substance,
yet being unperfect; and in thy book all my membees
were written, which in continuance were fashioned,
WHEN AS YET THERE WAS NONE OF THEM." These two
great truths are seen in beautiful combination in geology,
THE OLD TESTAMENT TYPES. 505
which reveals a typical system, that is, all things formed
after a type, in every age, and also a grand system of
prophecy, in which the past ever points to the future,
and' the future appears as the accomplishment of the
presentiments of the past. Lower animals appear as a
prognostication of higher, and the higher come as the
fulfilment of the prediction set forth in the lower, and
this not hy any physical emanation of the one from the
other, but according to the eternal plan of Him who
hath therein showed the immutability of His counsel.
There is an order in successive, even as there is an order
in contemporaneous nature ; but as the one plant does
not produce the other plant, which in the same type may
be growing alongside of it, so neither does a species of
animal in one age produce the homologous species in a
succeeding age. In this divinely-predetermined progres-
sion man stands as the end or consummation of a process
which had been going on since the dawn of creation.
Views like these have been floating before the minds
of deep thinkers and large-minded observers for the last
two or three ages, and were expressed by some who did
not discover their true meaning. We find Herder writ-
ing, at the end of last century, " See how the different
classes of creation run into each other ! How do the
organizations ascend and struggle upward from all points
on all sides ! And then, again, what a close resemblance
between them ! Precisely as if, on all our earth, the
form-abounding mother had proposed to herself but one
type, one proto-plasma, according to which, and for which,
she formed them all. Know thou what this form is. It
is the identical one which man also wears. It is more
evident internally than it is externally. Even in insecta
an analogon of the human anatomy has been discovered,
though, compared with ours, enveloped and seemingly
506 THE OLD TESTAMENT TYPES.
disproportionate. The different members, and conse-
quently also the powers which work in them, are yet
undeveloped, not organized to our fulness of life. It
seems to me that throughout creation this finger-mark of
nature is the Ariadne thread that conducts through the
labyrinth of animal forms, ascending and descending."*
A similar passage, very probably suggested by that quoted
from Herder, (but without any acknowledgement to this
effect,) is found in Coleridge's Aids to Rerlection.f " The
metal at its height seems a mute prophecy of the coming
vegetation, into a mimic resemblance of which it crystal-
izes. The blossom and flower, the acme of vegetable
life, divides into component organs with reciprocal func-
tions, and by instinctive motions and approximations
seems impatient of that figure by which it is differenced
in kind from the flower-shaped Psyche that flutters with
free wing above it. And wonderfully in the insect realm
doth the irritability, the proper seat of instinct, while
yet the nascent sensibility is subordinate thereto — most
wonderfully, I say, doth the muscular life in the insect,
and the musculo-arteria in the bird, imitate and typi-
cally rehearse the adaptive understanding, yea, and the
moral affections and charities of man. Let us carry our-
selves back in spirit to the mysterious week, the teeming
work-days of the Creator, as they rose in vision before
the eye of the inspired historian of the operations of the
heavens and of the earth, in the day that the Lord Grod
made the earth and the heavens. And who that watched
their ways with an understanding heart could, as the
vision evolved still advanced towards him, contemplate
the filial and loyal bee, the home building, wedded and
divorceless swallow, and above all the manifoldly intel-
ligent ant tribes, with their commonwealths and con-
* Metempsychosis. t Apb. xxxvL
THE OLD TESTAMENT TYPES. 507
fedcracies, their warriors and miners, the husband folk
that fold in their tiny flocks on the honey's leaf, and the
virgin sisters with the holy instinots of maternal love
detached, and in selfless purity, and not say in himself,
Behold the shadow of approaching humanity, the sun
rising from behind in the kindling morn of creation !"
Nor are these the visionary dreams of a poet or a mystic,
clothing nature in forms devised by his own fantasy ;
they are (after deducting one or two slight misappre-
hensions of fact) the results reached by the profoundest
inductive science of our times.
Between this typical system in nature and our powers
of intelligence, there is a beautiful correspondence.
First, there is in the human mind an imagining faculty,
which experiences a strange delight in reproducing what
it has perceived under a kind of ideal or pattern form.
We have seen, let us suppose, a particular plant, say the
Victoria Begina, we cannot remember every insignificant
particular connected with the number of its ribs or veins,
but there is laid up in our minds a general outline of
its shape, colour, and structure, which enables us on any
other plant of the sort falling under our notice, at once
to recognise it as belonging to the same species. The
mind seems thus to idealize to some extent its very re-
collections. And then in the higher intellectual processes
of abstraction and generalization, it abstracts the indif-
ferent and retains the essential, and strives to group the
innumerable objects which it meets with under a few
heads, by means of their common qualities. The rela-
tions thus discovered, cause the classes and individuals
to recur again and again to the mind according to the law
of association, and even aid the mind in the perception
of certain kinds of beauty.
These are the topics which have passed before us m
508 THE OLD TESTAMENT TYPES.
the previous part of this Work. We are now to trace
another correspondence — it is equally wonderful — between
the typical system of nature, and the typical system of
revelation, and to show that this second is as admirably
suited as the first to the native capacity and tendencies
of the mind.
It has long been known to divines, that the Word of
God has a typical system. The types have not been
always expounded in the exercise of a sound judgment,
or in accordance with the principles which should govern
all Scripture interpretation. Not unfrequently imagin-
ation has been allowed unreined to career in this field
at will, and in all treatises of divinity, the word type has
been changed from its scriptural to a theological sense.
In other cases, the fanciful interpretations which have
collected around the types of Scripture have led men of
severe critical taste to reject the whole system as visionary.
Still it is obvious that types run through the whole
Word of God, and cannot be excluded from it without
mutilating the Old Testament, and even parts of the
New Testament, so as to deprive them of some of their
most marked features. But now where such curious
harmonies and prefigurations have been detected in the
organic world, we may be able to show that no one is
entitled summarily to reject Scripture types as being
contrary to reason, or the analogy of things, and even to
trace an analogy between the types of the Works and
of the Word of God. Not that the two systems are the
same ; they are not identical, but homologous or analo-
gous. If the principles which we have been unfolding
are well founded, there should, with the uniformity, be
also a diversity. The typical system of the animal king-
dom is of a different order from the typical system of
the vegetable kingdom ; and when we rise from matter to
THE OLD TESTAMENT TYPES. 509
mind, from nature to revelation, we may expect the typical
system to be of a higher kind than that which pervades
the organic world. There is the same method in all, and
this suited to the intellectual tendencies of mankind, but
it is varied to suit the end which each has to accomplish.
In the theological use of the phrase, the word type is
confined to the prefigurations of Christ set forth in the
Old Testament. In books of divinity we read of certain
ordinances as the type, and Christ as the antitype. But
this is not the sense in which the term is used in Scrip-
ture. Mr. Fairbairn, in his able work on Typology, says,
that he understands the word in the theological sense,
but adds, " as employed in Scripture it is used with
greater latitude, as may be seen by consulting in the
original the passages referred to," (Heb. viii. 5 ; 1 Cor.
x. 6 ; Phil. iii. 17 ; 1 Thess. i. 7 ; 1 Peter v. 3 ; Kom.
vi. 17.) But " the foolishness of God is wiser than men."
We do not know what right divines have to construct a
system of theological types, instead of a system of Scrip-
ture types. We are sure that had they kept to the
Scripture use of the term instead of devising a theolo-
gical sense, they would have been saved from much ex-
travagance, and have evolved much more truth. The
words employed in Scripture (ivno,, dnod^h/fiun^ stand
for pattern-figures, or examples ; and persons living in
ages widely different from each other, and events having
no natural connexion, are represented as being constituted
after the same type or model. There are typical occur-
rences mentioned in Scripture, and full of instruction,
which have no immediate connexion with the person of
Christ, and are in no way prefigurative of Him. Thus
the judgments of God fell on the children of Israel in
the wilderness as types or " examples" (1 Cor. x. 11) of
a method of procedure which is for ever the same, and
510 THE OLD TESTAMENT TYPES.
recorded "for our admonition," in order to shew that it
will be put in execution whenever men commit similar
deeds. Types did not cease when Christ appeared ; there
are types in the New Testament dispensation (Phil. iii.
17 ; 1 Thess. i. 7 ; Horn. vi. 17) as well as in the Old
Testament dispensation. The typical system of the
kingdom of grace is meant fundamentally and primarily
to shew that God proceeds according to one counsel and
purpose from age to age. In this respect there is an
exact correspondence between the typical systems of re-
velation and nature. But as in nature there are fore-
shadowings revealed by embryology and geology, so in
revelation there is also a scheme, and this a very grand
scheme, of prefiguration. In the natural kingdom all
inferior organisms point onward and upward to man ; in
the spiritual kingdom all life points onward and upward
to Christ. Theologians, in discussing types, have con-
fined their attention exclusively to these prefigurations ;
but in neglecting the other and wider view, they have
not only missed much instruction, but have not been
able to estimate precisely the meaning of the important
truths to which their attention has been called.
It strikes us that a typical system runs through the
whole Divine economy revealed in the Word. First,
Adam is the type of Man. He and his posterity are all
of the same essential nature, possessing similar powers
of intuition and understanding, of will and emotion, of
conscience and free agency, and Grod acts towards them
in the dispensations of grace as in the dispensations of
nature, as being one. Then, from the time of the Fall,
we have two different typical forms, the one after the
seed of the serpent, the other after the seed of the woman.
Henceforth there is a contest between the serpent and
Him who is to destroy the power of the serpent, between
THE OLD TESTAMENT TYPES. 511
the flesh and the Sjririt, between the world and the
Church. Two manner of people are now seen struggling
in the womb of time — a Cain and an Abel, an Ishmael
and an Isaac, an Esau and a Jacob, an Absalom and a
Solomon, the elder born after the flesh, and the younger
born after the Spirit. It is this unity of figure fully as
much as the " type" of sound doctrine which gives a con-
sistency, in the minds of believers, to our religion in all
ages ; which enables the Christian to profit, to this clay, by
the teaching of the Old Testament ; to sing, to this day,
the song of Moses and the Psalms of David ; and to per-
ceive and feel that there are the same contests now as
then, the same contests in the heart, the same contests
in the world, between the evil and the good principle,
between the first, or nature-born, and the second, or
grace-born. In short, there are now, as there have ever
been, but two men on our earth typical or representa-
tive ; the first man, which is Adam, the second, which is
Christ. " And so it is written, The first man Adam
was made a living soul ; the last Adam was made a
quickening spirit. Howbeit that was not first which is
spiritual, but that which is natural ; and afterward that
which is spiritual. The first man is of the earth earthy ;
the second man is the Lord from heaven."
There appear from age to age certain great leading
powers of the first or earthy form, distinguished by their
audacity and the oppression which they exercise over the
Church, such as Cain and Lamech, Ham and Nimrod,
Egypt and Babylon. " They have consulted together
with one consent ; they are confederates against thee ;
the tabernacles of Edom and the Ishmaelites, of Moab
and the Hagarenes, Gebal and Amnion and Amelek, with
the inhabitants of Tyre : Assur also is joined with them,
they bave holpen the children of Lot." These are repre-
512 THE OLD TESTAMENT TYPES.
sented in Christian times by Gog and Magog and Babylon
But we must confine our attention here to the examples
of the better type, which appear and reappear throughout
successive ages, and chiefly, in this section, to what is,
after all, the most important, to the prefigurations of
Christ.
It had been determined, in eternity, that, " He whose
delights were with the children of men," should come to
our earth in the fulness of time. He is called "the
Lamb slain from the foundation of the world •" and as
soon as man falls, there are symbols of Him : " Lo, I
come, in the volume of the book it is written of me."
The prefigurations of Christ may be divided into three
classes : — typical ordinances, personages, and events.
First, There is a number of ordinances, more or less of
the same general mould, all imparting substantially the
same instruction, all pointing to guilt contracted, to God
offended, to a propitiation provided, and to acceptance
secured through this propitiation ; — the four great car-
dinal truths of revealed religion, as addressed to fallen
man. There were sacrifices, in which the offerer, placing
his hand on the head of the animal, and devoting it to
destruction in his room and stead, expressed symbolically
his belief in those great saving truths. There was the
tabernacle, with the people worshipping outside, and the
Shechinah, which had to be sprinkled with blood, in its
innermost recesses, pointing to an offended1 God, but a
God who was to be propitiated through the shedding of
blood. There was the ark of the covenant, with the tables
of the law inside, and the pot of manna, and the rod that
budded, and, over all, the cherubim shadowing the mercy-
seat — fit symbol of an arrangement by which the law is
fulfilled, and provision made for a revival of life, and a
supply of spiritual food by a God ready to meet with,
THE OLD TESTAMENT TYPES. 513
and to commune with us on the mercy-seat. There is
the scape-goat, with the sins of the people laid upon it,
pointing, as clearly as the Baptist did, to " the Lamb of
God, which taketh away the sins of the world."
Secondly, There are typical persons, such as Abel and
Enoch, Noah and Abraham, Moses and Aaron, Samuel
and David, Elijah and EKsha, shadowing the prophetical,
priestly, and kingly offices of Christ. From the fall
downwards there is a succession of such personages, with
their individual differences, but all after a pre-determined
model, exhibiting certain features of character in as
marked a manner as the Jewish race show certain fea-
tures of countenance. As the clouds reflect the rays of
the sun before he appears above the horizon, so each of
these, though dark in himself — alas ! at times, shewing
his native darkness, reflects certain of the beams — most
commonly coloured, of the Sun of Kighteousness, and
shows that he is about to shine upon our world.
Thirdly, There are typical events, exhibiting the same
truths in a still more impressive form. There is the
flood, in which many perish, but a few, that is, eight
souls, are saved by an ark symbolical of the Saviour.
There is the destruction of Sodom, in Avhich the inhabi-
tants perish, while Lot and his family are rescued by hea-
venly interposition. Most instructive of all, and, there-
fore, occupying the most important place, there is the
deliverance from Egypt. The state of the Hebrews as
bondsmen, the deliverer prepared for his work by suffer-
ing, the method of the deliverence in the midst of con-
tests and judgments, the wonderfully instructive journey
through the wilderness, with the provision made for the
sustenance of the people, and the statutes delivered are
as certainly anticipations of a higher redemption to fol-
low, as the fish and reptiles' digits are anticipations of
22*
514 THE OLD TESTAMENT TYPES.
the fingers of men. It is all true history, and yet it
looks as if it were a parable written by some man of Grocl
for our instruction. We are trained in this training of
the children of Israel ; and by means of the discipline
through which they were put, our imagining faculty has
acquired some of our clearest and liveliest, some of our
most profound and comforting representations of the
method of redemption.
But we cannot understand the meaning of these ordi-
nances, personages, and events, unless we take along with
us both of the two grand principles which we have been
unfolding in this volume. We must not confine our
attention to their general homology, we must take into
account also their special adaptations. We must not
look upon them merely as prefigurations, we must look
upon them as also " a figure for the time then present."
(Heb. ix. 9.) These typical ordinances, persons, and
events, are all after the same general plan, and exhibit in
shadow the truths which the sinner most requires to know,
and especially the person and work of the expected One
unjder interesting and instructive aspects. But they were
all at the same time adapted with exquisite skill to their
own particular age, and the circumstances of which
they formed a part. The ordinances, for instance, were
appropriate worship on the part of those who were re-
quired to observe them, and, in some cases, they subserved
important national and civil purposes. The persons who
figure as types were all the while doing a work for their
own day, and were, in most cases, we believe, unconscious
that they bore a representative character — they were
conscious only of looking onward to the light, and they
wist not that their face was shining with the reflection of
that light. The events, too, did, in most cases, impart a
special lesson of their own, and, in all cases, were most
THE OLD TESTAMENT TYPES. . 515
important links in the chain of Providence. But just as
the paddle of the whale serves a special purpose, but
contains divisions not needful to its special purpose;
just as the chick's head contains typical bones not re-
quired in order to its extrusion from the egg ; — so the
Old Testament types, while each accomplishes an end
of its own, have all, at the same time, certain common
features of a prefigurative character. Like the different
species in the vegetable and animal kingdoms, like the
answerable organs in different species, they diverge on
either side in order to suit a purpose ; but, meanwhile,
they are all after one pattern. In human architecture,
we are pleased to see that the portico and the passage
leading from it have often a homology to the temple it-
self. It is the same in the temple of God. The gate-
way, and the pillars and avenues of approach, present the
same general outline as the temple to which they form an
entrance.
The whole of this method of procedure is in beautiful
adaptation to the native tendencies and acquired habits
of the mind of man. The skilful teacher is accustomed
to instruct his younger pupils by means of signs, and
pictures, and comparisons ; it is thus that he conveys
the ideas of remote objects and abstract truths. In the
simpler stages of society, mankind can be taught general
truths only by symbols and parables. Hence we find
most heathen religions becoming mythic, or explaining
their mysteries by allegories or national incidents. The
great exemplar of the ancient philosophy, and the grand
archetype of modern science, were alike distinguished by
their possessing the power of comparison in a high de-
gree, and both have told us that man is best instructed
by similitudes. " It is difficult/' says the Guest in the
Statesman of Plato, "fully to exhibit greater things
516 THE OLD TESTAMENT TYPES.
without the use of patterns," (.T«^«&/j^«nif.) Lord
Bacon, in more than one place, has declared, "As hiero-
glyphics preceded letters, so parables are older than argu-
ments. And, even now, if any one wishes to pour new
light into any human intellect, and to do so expediently
and pleasantly, he must proceed in the same way, and
call in the assistance of parables." It appears, then, that
God was acting in accordance with the nature which He
had given us, in His method of instructing the early
Church. In Bible history there are no myths, but real
events are made as lively as myths, and convey far more
important instruction. And, even in Christian, times,
this representative system has been felt by all, but espe-
cially by the simple and unlettered, to be a powerful
means of imparting great vividness and picturesqucness
to the inspired teaching. The truth is exhibited ; not, as
in systems of divinity, as a bare abstraction ; not, as in
the words of Scripture, by a phrase expressive enough,
but still a mere counter, bearing no resemblance to that
which it represents ; but by a picture which the mind,
as it were, sees before it. With such lively images before
us, we feel as if we were walking amid living realities.
We find, in particular, that the types of the Bible have
ever been especial favourites with the " common people,"
who experience a difficulty in seizing an abstraction, or
in grasping a generalization, but feel none in compre-
hending truths which are embodied in an incident, a
person, or an ordinance. Take away the typical repre-
sentations of the deeper doctrines of the Word of God,
take away such figures as sacrifices, as the brazen ser-
pent, as the scape-goat, the city of refuge, the sprinklings
and ablutions under the law — abstract these from the
apprehensions of the Christian who moves in the lower
walks of life, and there would remain, we suspect, scarcely
THE OLD TESTAMENT TYPES. 517
any idea — there would certainly be nothing remaining to
enliven and engage the mind. It was in gracious accom-
modation to the same peculiarities of our nature, that
the greatest of all teachers, " He who knew what was in
man/' taught the people by parables.
The right conclusion has been drawn by one in whose
history difficulties have merely been " Schools and School-
masters" to strengthen his native genius. " As the veil
slowly rises," says Dr. Hugh Miller, " a new significancy
seems to attach to all creation. The Creator, in the first
ages of his workings, appears to have been associated
with what he wrought simply as the producer or author
of all things ; but even in these ages, as scene rose after
scene, and one dynasty of the inferior animals succeeded
another, there was strange typical indications, which
pre-Adamic students of prophecy, among the spiritual
existences of the universe, might possibly have aspired
to read — symbolical indications to the effect that the
Creator was, in the future, to be more intimately con-
nected with his material works than in these ages, through
a glorious creature made in his own image and likeness.
And to this semblance and portraiture of the Deity —
the first Adam — all the merely natural symbols seem to
refer. But in the eternal decrees, it had been for ever
determined that the union of the Creator with creation
was not to be a mere union by proxy or semblance ; and
no sooner had the first Adam appeared and fallen, than
a new school of prophecy began, in which type and
symbol were mingled with what had now its first exist-
ence on earth ; and all pointed to the second Adam, ' the
Lord from Heaven.'* In Him creation and the Creator
* This extract is from a notice by Dr. Miller of the Article in the North British Review
previously referred to. In the same article he shews wherein Oken had erred. " Hence
the remark of Oken, that 'man is the sum total of all the animals.' Hence, too, but
■with a still broader appreciation of the homologies which bear upon the lord of creation
518 TYPICAL NUMBEKS.
met in reality, and not in semblance ; on the very apex
of the finished pyramid of being sits the adorable Mon-
arch of all ; — as the son of Mary, of David, of the first
Adam, the created of God — as God and the Son of God,
the eternal Creator of the universe. And these — the two
Adams — form the main theme of all prophecy, natural
and revealed. And that type and symbol should refer
not only to the second, but, as held by such men as
Agassiz and Owen, to the first Adam also, exemplifies,
we are disposed to hold, the unity of the style of Deity,
and serves to shew that it was He who created the worlds,
that dictated the Scriptures."
SECT. II. TYPICAL XUMBERS.
There is no object on which a greater amount of
extravagant statement has been made, both in ancient
and modern times, than the significance of numbers.
The Pythagoreans, and later Platonists, evidently sought
for some inherent power in numbers to account for the
numeral relations that appear in nature. In the pages
of Philo-Juda3iis and Josephus, numbers have a theoso-
phic signification. In more than one country, certain
as their central type, his essentially profane and erroneous remark, that ' man is God
manifest in flesh.' Let the reader, however, observe in what the error and profantity
consists. There is a loose sense in which man is God manifest in the flesh ;— he is God's
image manifested in the flesh ; and an image or likeness is a manifestation, or making
evident, of that which it represents, whether it be an image or likeness of body or mind.
Originally, at least in moral character, man was a manifestation of his Maker, and in
intellect he is a manifestation of his Maker still. But the ciror and profanity of Oken
consists in applying that to man, the image — man, the being in whom merely the hoino-
logues or natural prophecies converge — which is exclusively applied, in revelation, to a
higher and more real manifestation of God in the flesh— that manifestation of very God
himself which has formed the subject, not of natural, but of the revealed prophecies rl he
transcendentalist has gone, in his irreverent ignorance, a step too far; and yet his mean-
ing seems real, though he himself mistook its nature, and employed improper language
to convey it.'' — Witness, Aug. 1851. We may here be permitted to express a wish that
the author will some time or other republish a selection from the articles in the Wttnest
newspaper; they would be acknowledged not to be inferior to the republicatlcns from
any of the periodicals of our age.
TYPICAL NUMBERS. 519
numbers have been supposed to have a magical power.
Commentators have discovered a mystical meaning in
the special numbers which appear and reappear so con-
stantly in the Word of G-od. Others have not known
what to make of Scripture numbers, while not a few have
looked with suspicion upon the passages which contain
them, or the Bible, because it is so full of them.
The train of observation and reflection followed in this
treatise, may help us to discover what is the true signifi-
cancy of such numbers.
In comprehending and recollecting the isolated and
scattered phenomena of nature, and in the scientific con-
struction of them, in order to these ends, man's intellect
needs such recurring numbers, and when he does not find
them in nature, he places them there. Man seeks them,
too, in chronology, as an aid at once to the memory,
which calls up events by the law of correlation, and the
contemplative intellect, which loves to collect objects into
groups. So strong is this tendency, that when such re-
lations are not found among events, mankind will create
them from the stores of their own ingenuity, and will
lengthen or shorten periods to suit them to the measure
of their Procrustes' bed. Hence it is, that in the specu-
lations of early philosophers, in history handed down by
popular tradition, and in all mythic systems of religion,
we have recurrent numbers, such as three and five, seven
and ten. The existence of this mystical tendency in pre-
mature scientific speculation, should not lead us, by an
extreme reaction, to affirm that numbers have no signj-
ficancy in nature ; it should merely guard us from
adopting them too readily — that is, it should prevent us
from receiving them without inductive evidence, which
is now, however, superabundant. On a like principle,
the numeral relations of mythic religions should not be
520 TYPICAL NUMBERS.
held as proving that biblical institutions and narratives
are fabulous, simply because they contain recurrent num-
bers. It has been far too readily assumed, by certain
neological critics in Germany and their followers in this
/'m^ country, we have shewn their dissecting acuteness by
pruning away — on the pretence of improving it — the
tree of life, till they have destroyed not only its lovely
form, but its very vital principle, that every portion of
the Old and New Testament is to be regarded' as fabulous
which contains a repetition of numbers.
Physical science shews that numbers have a signi-
ficancy in every department of nature. Two appears
as the typical number in the lowest class of plants, and
regulates that pairing or marriage of plants and animals
which is one of the fundamental laws of the organic
kingdoms. Three is the characteristic number of that
class of plants which have parallel veined leaves, and is
the number of joints in the typical digit. Four is a sig-
nificent number in those beautiful crystals which show
that minerals (as well as stars) have their geometry.
Five is the model number of the highest class of plants —
those with reticulated veins and branches, is the typical
number of the fingers and toes of vertebrate animals, and
is of frequent occurrence among star-fishes. Six is the
proportional number of carbon in chemistry, and 3x2
is a common number in the floral organs of monoctyle-
donous plants, such as the lilies of the field, which we are
exhorted to consider. Seven appears as significant only
in a single order of plants, (Heptandria,) but has an im-
portance in the animal kingdom, where it is the number
of vertebra3 in the neck of mammalia, and according to
M. Edwards, the typical number of rings in the head,
in the thorax, and in the abdomen of Crustacea. Eight
is the definite number in chemical composition for
TYPICAL NUMBERS. 521
oxygen, the most useful element in nature, and is
very common in the organs of sea-jellies. Nine seems
to be rare in the organic kingdoms. Ten or 5 x 2 is
found in star-fishes, and is the number of digits on the
fore and hind limbs of animals. Without going over
any more individual numbers, we find multiple numbers
acting an important part in chemical compositions, and
in the organs of flowers ; for the elements unite in multi-
ple relations, and the stamens are often the multiples
of the petals. In the arrangement of the appendages
of the plant we have a strange series, 1, 2, 3, 5, 8, 13, 21,
34, which was supposed to possess virtues of an old date,
and before it was discovered in the plant. In natural
philosophy the highest law, that of forces acting from a
centre, proceeds according to the square of numbers. In
the curves and relative lengths of branches of plants,
there are evidently quantitative relations which mathe-
matics have not been able to seize and express.
He must be a bold man who will insist, that should
the God who fashioned nature be pleased to give to man
a revelation of His will, in order to solve certain great
problems started by the existence of sin in the world, He
shall not be at liberty to make His dispensations of pro-
vidence, and His institutions for instruction and worship,
bear a certain relation to each other. It is presumptuous,
above all things, in any one to condemn as mythic every
part of the Bible narrative which contains a recurrent
number. This principle would turn the discoveries of
the most eminent scientific men in modern times — the
discoveries of Kepler, of Newton, of Decandolle, and
Dalton into myths. The constant recurrence of certain
numbers in the self-devised history of tradition, and the
self-formed religions embodied in myths, is an acknow-
ledgment on the part of man, that he needs such relations
522 TYPICAL NUMBERS.
to enable him to follow history and comprehend doctrine.
And may not He who knows what is the nature of man,
suit Himself to the creatures fashioned by Him, by insti-
tuting, in the realities of His dispensations and His ordi-
nances, those very numerical relations which man will
feign by his imagination, where the actual state of things
does not present them ?
We certainly do meet in Bible narrative with a re-
currence of certain numbers, and these not unlike the
numbers which recent science has disclosed in nature.
The beasts were gathered into the ark, even as they
are assorted in nature, in pairs ; and our Lord sent out
His disciples, as the fowls of the air are sent out, two
and two, to support and comfort each other. Three de-
rives its significancy from the very nature of God, and
appears in the triple sacerdotal blessing of Jacob, (Gen.
xlviii. 16 ;) in the thrice holy of Isaiah, (vi. 3 ;) in the
three great religious festivals ; in Jonah being three days
in the whale's belly ; in our Lord being three days in the
grave ; and in the threefold judgments denounced in
the Book of Revelation, where the tail of the great red
dragon draws the third of the stars, and three unclean
spirits issue from the mouth of the dragon, the beast, and
the false prophet. Let us not forget that the triad is the
representative of Deity in many religions, and appears in
the three-forked lightning of Jupiter, the trident of Nep-
tune, the three-headed dog of Pluto, the tripod^f Apollo,
the three Fates, three Furies, three Graces, and thrice
three Muses.* Four appears in Scripture in the altars,
and sanctuary, and holy of holies, which was a cube ; and
groups of four are found in Revelation, such as, heaven,
earth, sea, and fountains of waters ; kindred, and tongue,
* See Article in American Biblical Repertory, republished in British and Foreign
Evangelical Review, June 1855.
TYPICAL NUMBERS. 523
and people, and nation. Five is found in the pillars of
the courts of the temple, which were five cubits high,
and live cubits apart ; and in the ten virgins, five of
whom were wise, and five foolish. Six is once or twice
mentioned as a significant number in Ezekiel. Seven is
the most frequently repeated number in the Bible. We
have first the seven days of creation ; then the seven days
of the week ; then a series of seasons regulated by seven ;
the seventh year was a Sabbatical year, and 7X7 gave
a year of jubilee ; and at the close of the canon there are
the seven spirits before the throne, the seven churches of
Asia, the seven branches of the candlestick, seven angels
with seven trumpets, and seven vials with the seven last
plagues. The number ten ajjpears in the tithes devoted
to God, in the plagues which devasted Egypt, and in
the commandments delivered amidst the thunders of
Sinai. Twelve was the number of the sons of Joseph, of
the tribes of God's people, and of the Apostles ; the holy
city measured twelve thousand cubits in length, breadth,
and height, and had twelve foundations, twelve gates, and
a tree of life which bears twelve manner of fruits. Multi-
ple numbers are very frequent. Forty days, or 4 X by 10,
was the time of Moses' sojourn on the mount with God, of
Elijah's journey to Horeb, and of our Lord's temptation
in the wilderness. There were 7 x 7 days between the
passover and pentecost, and 7X7 years between the
times of jubilee. The Tabernacle measured ten cubits in
breadth and length, and 3 x 10 cubits in length ; there
were 4 X 12 boards in its frame, anil the court was 10 X
10 cubits long, and the sacrifices on certain occasions were
multiples of seven. We read of 7 X 10 disciples ; Peter
was exhorted to forgive his brother not seven times, but
seventy times seven, and the redeemed on Mount Zion
are 12 x 12 thousand.
524 TYPICAL NUMBERS.
This method of instruction is in admirable adaptation
to the constitution of man's mind. It lends distinctness
to the incident, it helps the intellect to grasp the truth,
and the memory to retain it. It is one of many circum-
stances which adapt the Word like the Works of God to
every capacity, to persons of all ages and sexes, times
and countries. By these and similar means, the greatest
of all Teachers still encourages little children to come
unto Him, when other teachers would forbid them. Its
institutions and its incidents strike the fancy and are
fixed in the memory of youth ; they interest by their cor-
respondences the understanding of the mature man, and
are found wrapt round the decaying memory of old age,
the burden of which they serve to lighten, and the gloom
of which they irradiate.
And in all this, whether in nature or in the Word, we
are not inclined to find anything mystical or even mys-
terious. We are not disposed to believe it to proceed
from any inherent power of numbers, as certain mathe-
maticians and philosophists have imagined. Nor does it
imply that any one number has a special significance.
We have quoted so many cases of numeral relation in
order to show that all, or nearly all, the lower numbers,
odd and even, appear as principles of co-ordination both
in nature and in the Word. No doubt, there are circum-
stances which have determined the use of certain num-
bers. ThuSj the nature of the plant, as having an axis,
with symmetrical sides, may have determinted the selec-
tion of the odd numbers 3 and 5 in the organs of the
vegetable kingdom. The recurrences of 5 and 10 in
human enumeration probably originated in the number
of the digits. The triune nature of Grod, and the divine
institution of the Sabbath, must have given rise to the
frequent use of the numbers 3 and 7 ; and the circum-
TYPICAL SYSTEM OF THE NEW TESTAMENT. 525
stance that the patriarchs were 12 in number must have
brought series of twelves in its train. Still, in all this
there is no evidence of there being any power, virtue, or
significance in any one number considered in itself.
We are not even inclined to look upon these recurrent
numbers as implying any mysterious connexion, as theo-
sophists have supposed, between objects which have the
same number attached to them. We do not conclude
that there is a connexion between the typical organs of
dicotyledonous plants and the digits of animals, because
they both range round the number 5. Nor are we to
look upon biblical events as related, solely because they
appear under the same number. It is possible, indeed,
that the events may have a connexion in themselves,
and have both appeared under the same number because
of this connexion ; but the evidence of their relation
must be sought otherwise than in their numerical corre-
spondence. In vindicating the existence of these nume-
rical relations, we are thus, at the same time, laying an
effectual arrest on the abuse of them. We do not admit
them in natural science, except on evidence which can
stand the rules of inductive logic ; and we should not
allow them in theology, except on grounds which can
stand the tests of sound biblical interpretation.
SECT. III. TYPICAL SYSTEM OF THE NEW TESTAMENT.
In looking at any one epoch of our world's history, we
find traces of contemporaneous order and fitness. In
comparing any one epoch with the preceding one, we
find traces of a progression. It should be admitted,
however, that we are not altogether in circumstances to
determine the character of that progression. In physical
and organic nature, it seems, so far as we can discover,
526 TYPICAL SYSTEM
to be an advance from the simple to the manifold ; from
the more general to the more special ; from the type to
the archetype. It rises from the crystal to the plant and
the animal. Its foundation shows right lines and regu-
lar figures, while the superstructure sweeps out into
varied curves. There is first the simple capacity in the
germ, the bud, and then the unfolding of all the capabi-
lities in distinct members.
Is not this the very law of the advance of the human
mind ? It begins with the simple and goes on to the
multiple. It craves first for mere milk, and then ac-
quires a relish for strong meat and varied dainties. In
their literary tastes, men like first very easy and trans-
parent narrative in prose, and songs with the simplest
cadences ; then more elaborate prose and more adorned
poetry ; and finally, perhaps, a style, to use the language
of Burke, between prose and poetry, and better than
either. Is not the history of human civilisation an ad-
vance from a union of labour to a division of labour ; from
few and simple to many and complicated relations ? Is
not the advance in physical science (as M. Comte has
shown) from pure space to body, to bodies with chemical
affinities, on to bodies organized ?
It is evident that there is some kind of progress in the
history of religion, though we are not in a position to
apprehend it closely, or unfold it fully. All the essen-
tial and saving truths are embraced in the earliest reve-
lation of Grod, but they are in the bud, they are hopeful
and prophetic ; it is only as ages advance that they are
expanded to the view. Under the Old Testament the
shadow becomes more and more defined as the substance
draws nigh ; but it is only in the later prophets that we
discover distinct lineaments. The figure .presented in
the first prediction is as large as it ever is afterwards,
OF THE NEW TESTAMENT. 527
but its lines come out more and more distantly as the
light approaches nearer and nearer. The doctrine
which we are expounding, be it observed, is not the vul-
gar one of type and antitype, but that of typical forms,
serving immediate and important ends in the age in
which they appeared, but, at the same time, epitomes of
an archetype, to appear. When the Archetype presents
Himself, what had before been dim is now distinct. In
the scene on Calvary, in particular, we have the truths
which the sinner is most concerned to know, of sin and
salvation, of God offended and God pacified, set forth in
the most awfully, and yet most winningly, impressive
manner.
There seems to be the very same order in the mode of
communicating the truth. First, there are symbols of
various kinds, then prose narrative and poetry with
simple correlations, then richer and more varied ptoetry,
and, when Ave come on to the New Testament, interesting
narrative, with interspersed spoken discourses with num-
berless parallelisms and most graceful curvatures, leading
on to more elaborate and logically constructed prose, and
the whole closing with a book in which prose, poetry, and
symbol are combined.
And we are not to understand that the scheme of
types, as we have explained it, disappears on the appear-
ance of Christ, or with the close of the inspired canon.
The continued operations of Christ in the Church are
typical. Let us compare what He did when He walked
on the earth in His human person, with what He is still
doing on our earth as He walks spiritually in the midst
of the lamps which He has kindled. His miracles, which
attested His divinity, did not consist in ostentatious dis-
plays of power ; in meteors flashing across the sky ; in
rivers, running backward to their source ; in mountains
528 TYPICAL SYSTEM
being shaken tc their centre ; or in the moon wandering
from her orbit. The testimony was to His love and
compassion as well as His power. " The blind receive
their sight, and the lame walk ; the lepers are cleansed,
and the deaf hear ; the dead are raised up, and the poor
have the Gospel preached unto them." Of a like type
are His operations still. They clo not consist in displays
of physical power ; — but in opening the eyes of the spi-
ritually blind ; in putting activity into those in no way
disposed to the service of godliness ; in curing all man-
ner of soul-maladies ; in gaining access to the most ob-
durate hearts by the power of His Spirit. We do not
affirm that the one set of acts predicted the other — we
venture on no such statement ; but we maintain that
both are after the same figure, that both originate in the
same peculiarities of character, and that both are ad-
dressed to us as homotyj)al correspondences.
We still live under a system of types. Just as all the
figures in the Old Testament look forward to Him who
is the principal figure, so do the figures in the New Tes-
tament look back to Him. But there is this difference
between the former and the latter types, that the latter,
as becometh the dispensation, are not so much outward
and ceremonial as inward and spiritual. There is a close
mystical union between Him and each of His people ;
He and they are said to be one. They are one in respect
of their human nature ; "It behoved Him to be made
like unto His brethren, and, forasmuch as the children
are partakers of flesh and blood, He also likewise took
part of the same ;" — " He took on Him, not the nature
of angels, but the seed of Abraham/' and " was made
in the likeness of man." He has become, too, the " head
of the body, the Church," " the beginning, the first-
born from the dead," and is the "first-born among
OF THE NEW TESTAMENT. 529
many brethren." They are priests under Him as Chief
Priest, kings under Him as Sovereign. By his appoint-
ment they are " predestinated to be conformed to His
image." The Godhead once more issues the decree
in reference to this man and to that man, " Let us
make man in our image, after our likeness," — so " God
creates man in His own image, in the likeness of God
creates He him." In the performance of this decree,
they "suffer with Him," are "crucified together with
Him," are " dead with Him," " buried with Him," and as
they die with Him, so they are " quickened with Him ;"
they " rise with Him," and " reign with Him." In this
household there are many children, and there are differ-
ences between them of gift and taste to suit them to
their different heaven-allotted employments ; but still we
may discern in them all a family likeness, and the image
of Him who hath begotten them. In this perfect system
of types, the whole has a representative in every part,
and every part is a symbol of the whole. Each living
stone in this temple is carved after the similitude of the
whole temple. He is the body, and every member in
particular is after the pattern of the whole body. Each
branch, each leaf of this Tree of Life, is an image of the
entire tree.
With such patterns in the past and in the present, the
disciple may everywhere be instructed. But let him
remember, meanwhile, what is the object to which he
should chiefly look. It is pleasant to see the path in
which we walk trodden by the footsteps of the flock, but
we are to follow the flock only so far as they follow the
shepherd. In Him we have the image of the invisible
God set before us in such a way that we can rise to a
somewhat clear, and an altogether satisfactory apprehen-
sion of His character. It is when the soul is spread out
23
530 • TYPICAL SYSTEM.
before Him, and directed towards Him, that His likeness
is imprinted — as by a sunbeam process, upon the tablet
of the heart. Looking to Him, as lifted up upon the
cross, we learn of Him the lessons which we have most
need to learn— we learn of Him to be meek and lowly.
A similar change, but differently produced, shall take
place in heaven. In the Old Testament Church they
had the shadow ; in the New Testament Church we have
the image ; in heaven we shall have the substance — ■
which as we behold, we shall be brightened into a likeness
to His glory — " we shall be like Him, for we shall see
Him as He is."
It appears, then, that in the New Testament Church
there are post-figurations ; but there are also pre-figura-
tions. In spite of many partial relapses, the Church, as a
whole, is advancing, and its past progress is but an ear-
nest of its future progress till it cover the earth. As it
advances, if not so simple, or perhaps so pure, yet it is
richer and fuller, and shall inspire and fashion a greater
diversity of character and of phases of life, and, at the last,
all the accumulated stores of wealth, civilisation, and in-
tellect, shall be cast into the treasury of the Lord.
When objects lie far distant from us, we must be on
our guard against taking brightened clouds for sunlit
lands ; but we think we see some real truths, lying, we
grant, on the very horizon of our vision. All animal
bodies point to man as the apex of the earthly hierarchy.
Professor Owen tells us that "all the parts and organs
of man had been sketched out, in anticipation, so to
speak, in the inferior animals." But may not this highest
form on earth point to a still higher form ? Man's body
on earth may be but a preriguration of his body in hea-
ven. "But some will say, How are the dead raised up,
and with what body do they come ? The Apostle does
OF THE NEW TESTAMENT. 531
not give a direct answer to this question, but he points to
certain analogies, or rather homceophytes, which shew that
while the body preserves its identity, it will be changed
into a nobler form, as the seed is changed when it springs
up as a plant. "It is sown a natural body, it is raised a
spiritual body ; for there is a natural body and a spiritual
body," and we read of bodies " terrestrial," and of bodies
" celestial." In heaven, then, our bodies are to be after
a higher model, "spiritual" and " celestial." It doth not
yet appear what we shall be, but being planted in the like-
ness of His death, we shall also be planted in the likeness
of His resurrection, and when He appears we shall be
like Him. Our bodies shall then be fashioned like unto
His glorious body, which we may conceive to be the most
sublimated and obedient form and modification of mate-
rial agency ; — and modern science, while it cannot efface
the indelible distinction between mind and matter, is
every day enlarging our conceptions of the capacities of
matter. Thus, the simplest organism points by its struc-
ture upward to man, and man's earthly frame points to
his heavenly frame, and his heavenly frame to Christ's
spiritual body — and we see that all animated things on
earth point onward to His Glorified Humanity as the
grand Archetype of all that has life.
Professor Owen has another idea. He supposes that
in other worlds, as there are the same laws of light and
gravitation as on our earth, there may also be like organic
structures: "And the inference as the possibility of
this vertebrate type being the basis of the organization of
some of the inhabitants of other planets, will not appear
so hazardous when it is remembered that the orbits or
protective cavities of the eyes of the vertebrata of this
planet are constructed of modified vertebra?. Our
thoughts are free to soar as far as any legitimate analogy
532 TYPICAL SYSTEM OF THE NEW TESTAMENT.
may seem to guide them rightly in the boundless ocean
of unknown truth. But if censure be merited for here
indulging, even for a moment, in pure speculation, it
may, perhaps, be disarmed by the reflection that the dis-
covery of the vertebrate archetype could not fail to sug-
gest to the anatomist many possible modifications of it
beyond those we know to have been realized in this little
orb of ours."
If there be any truth in this idea, then the animated
matter of other worlds may point to the same Archetype
as the animated matter of this world. And on the sup-
position, what a significancy would be given to the hu-
manity of our Lord ! When the Word became flesh,
the Divinity was in a sense humbled ; and when the In-
carnate Word ascended into heaven, flesh or matter was
exalted, and made to serve the most glorious ends. We
thus obtain a glimpse of a way in which matter, through-
out all its domains, may be exalted by its association with
the Son of God taking our likeness ; and of a way, too, in
which other worlds, or all worlds, and other creatures,
even principalities and powers in heavenly places, may
be instructed by this " manifold wisdom," and by which
God may " by Him reconcile all things unto Himself—
by Him, I say, whether they be things in earth, or things
in heaven."
But as we stand gazing on our ascending Lord, a
cloud wraps Him from our view, and we hear, as it were,
a voice saying, " Why stand ye here gazing ?" and bid-
ding us return to the observation of things clearly within
the range of our vision.
APPENDIX.
SELECTED LIST OF PLANTS,
ILLUSTRATING ASSOCIATIONS OF COLOURS, AND RELATIONS
OF FORM AND COLOUR. — (p. 166.)
Ranunculus repens,
R. bulbosus, .
R. netnorosus,
Nasturtium Indicum,
Cheiranthus alpinus,
Viola Curtisii,
V. lutea,
Saxifraga ligulata,
S. sarmentosa,
S. Aizoon,
Cytisus Laburnum,
Anthyllis vulneraria,
Lathyrus pratensis,
Kennedya monophylla,
Cytisus scoparius, .
Lotus corniculatus,
Dicotyledons.
) Corolla yellow ; purple on calyx, on leafc
) stalks, and on leaf-sheaths.
Corolla yellow ; tips of young sepals purple.
| Corolla yellow ; tips of sepals purple.
[ Flowers yellow and purple.
Corolla white, with purple spots; the yellow
anthers lie on the purple spots.
Two petals white ; three are spotted with
purple, and are yellow at the base.
Leaves yellow-green and red-purple; corolla
yellow and purple ; ovary first yellow-
green, then red-purple.
Four petals yellow ; the fifth is yellow, with
a purple spot,
Corolla yellow ; tips of calyx purple.
Corolla yellow; the odd lobe with purple
veins.
Four petals purple ; the odd petal has a yellow
spot.
Flower yellow; the odd piece has purple
streaks on the inside.
Calyx yellow-green, and red-purple streaks;
odd lobe of corolla yellow, with purple on
the outside.
534
APPENDIX.
Swainsonia purpurea, .
Hieracium Pilosella,
Aster acris, A. spectabilis,
A. cordifolius, A. Novae An-
glias, ....
Rudbeckia fulgida,
Corvisartia Helenium, .
Gloxinia grandis, .
Ajuga Chamsepitys,
A. pyramidalia,
Galeopsis Tetrahit,
G. versicolor,
Melittis grandiflora,
Antirrhinum Orontium,
Euphrasia officinalis,
Linaria Cymbalaria,
Schizanthus purpureus,
Sarracenia purpurea,
Rurnex pulcher, .
R. Acetosa, .
R. aquaticus,
Pinus sylvestris, and other
Coniferse, .
Ficus elastica,
Drimys Winteri, .
Taxodium sempervirens,
Lycaste Skinneri, .
Corolla red-purple ; odd lobe with a white eye.
Flower yellow, outer surface of ray purple.
Centre yellow, circumference purple.
Centre purple, circumference yellow.
Circumference and centre yellow ; inner scales
of involucre red-purple, outer scales yellow-
green; stems red-purple, foliage yellow-
green.
Odd lobe of corolla red-purple inside; calyx
yellow-green ; stalks red-purple.
Flower yellow, with purple on odd lobe.
Flower purple, yellow spot on odd lobe.
Odd lobe yellow and purple.
Odd lobe yellow and purple.
Flower yellow, purple on the odd lobe.
Flower purple, yellow on the odd lobe.
Corolla purple streaked, yellow on the odd
lobe.
Corolla purple, odd lobe with yellow spots
and yellow hairs.
Odd lobe of corolla yellow and purple.
Pitcher red-purple and yellow-green.
Anthers purple below, j^ellow above.
Perianth red-purple and yellow-green.
Stem red-purple ; dense masses of yellow-
green flowers ; the latter have sometimes
red-purple streaks.
■ Young cones purple and citrine.
Buds red-purple, leaves yellow-green.
Young shoot red-purple, young leaf yellow-
green.
Young shoot yellow-green; more advanced,
red-purple ; when older, it is citrine.
Monocotyledons.
Sepals, outside yellow-green, inside red-
purple ; two upper petals white, or yellow
" with purple spots; third petal yellow and
purple spots.
APPENDIX.
535
Brassia verrucosa,
Oncidium Cavendishii, .
Epidendrum cochleatum,
Lycaste aromatica,
Cattleya Loddigesii,
Oncidium Papilio,
Cypripedium venustum,
Listera cordata,
Iris pseudacorus, .
I. Germanica,
I. Tricolor, .
Pandanus odoratissimus,
Caladium pictum, .
Strelitzia, several species,
Curcuma cordata, and
C. ovata,
Juncus compressus,
Aveua pratensis, .
Papyrus antiquorum, .
Sepals and two upper petals yellow-green and
red-purple ; third petal white, with green
warts and yellowish eye; flower-stalks
purple.
- Third petal yellow and purple.
Leaves red-purple and yellow-green.
Petals yellow-green and red-purple; bract
yellow-green, with red-purple line on the
midrib, and one near each margin ; ovary
yellow-green, with red-purple lines corre-
sponding to the adherent edges of the
pieces of which it consists.
Flowers red-purple and yellow-green.
Flower yellow, with purple streaks ; stamens
variegated with purple.
Calyx yellow and purple ; corolla purple ;
pollen yellow.
Petals yellow; sepals yellow and purple.
Teeth and edge of leaf red-purple, centre
yellow-green.
Centre of leef red-purple, edge of leaf yellow-
green.
Leaf yellow-green, leaf-stalk red-purple; se-
pals orange ; petals blue.
Tip of bracts red-purple, base of bracts yel-
low-green ; flower yellow.
Flower russet and green.
Glumes citrine, with purple streaks and
purple awn ; anthers yellow and purple.
Sheaths red-purple ; stalks yellow-green.
INDEX.
Actinia, 271-275.
Adaptation, principle of, 1, 30-40, 421,
427-438, 514.
^Esthetic feelings, 137, 139, 145, 150, 151,
153, 481-492.
Amphipoda, 242.
Analogues, 25, 292, 293, 308.
Analysis, 452-454.
Angles of leaf-veins and branches, 112-
116.
A priori speculation, 463-470.
Archetype skeleton, 180.
Armature of plants, 134.
Aristotle, 11, 465, 474.
Articulata, 233, 266, 336-338.
Association of ideas, 473-480.
Back, vertebras of, 195.
Bacon, 13, 421, 466.
Barnacles, 245, 246.
Bat's wing, 210.
Bats, teeth of, 216.
Beauty, 481-488.
Birds, vertebriB of, 198, 199; diverging
appendages of, 207, 208 ; cutaneous mus-
cles of, 295.
Blood-corpuscles, 79.
Bone, structure of, 77; typical form of,
185, 307.
Bracts, 91, 135.
Branches, 112-119.
Branchiopoda, 242.
Buds, 83.
Butterfly, mouth of, 254-257.
Calyx, 92, 136.
Camel, 66.
Campari ularia, 273.
Carnivora, teeth of, 218.
Cartilage, 76.
Caterpillar, 249.
Cause and effect, 467-470.
Cell, 70-71.
Centrum of vertebra, 178.
Cephalopoda, 227.
Cerebellum, 287.
Chance, 40-54.
Chemical groups, 363.
Cicero, 8, 9.
Classification, 422-426, 451, 460-46&
Coal epoch, flora of, 348.
Collocation, principle of, 34.
Colours, 20-21, 53-58, 481.
Colours of plants (and birds), 146-174
Comets, 391, 392.
Complementary colours, 154-156.
Cones and Coniferse, 120-128.
Corolla, 93, 137.
Correlation of physical forces, 32, 460.
Cosmology, 30.
Covering of plants, 133.
Cotyledons, 82, 141.
Crustacea, 237-243.
Crystals, 354-358.
Curves, 24, 117, 483.
Cuttle fishes, nerves of, 281.
Cuvier, 421, 431, 432, 434, 439.
Development, community of plan in, SOJ
Development, progressive, 817-822.
Descartes, 493.
Division of labour in science, 12, 13.
Egg, 63 ; of bird, 308.
Elephant, 66, 67 ; teeth of, 219.
Endogenous stem, 84.
Epidermis, 73.
Epithelium, 74.
Equivalents, chemical, 360-862.
Exogenous stem, 84.
i38
INDEX.
Fat, 74.
Feathers, SO.
Fichte, 4T1, 49T.
Final cause, 50-5(5, 427-438.
Firs, morphology of, 119-129.
Fishes, skeleton of, 196; diverging appen-
dages of, 200, 2f)7; teeth of, 221.
Form, as a principle of order, 11, 21-28.
Form, the faculty which discovers the re-
lations of, 454-456.
Form and colour, relation of, 147.
Fossil animals and plants, 309-353.
Fruit, 139.
Generalization, 460-467.
Geology, 309-353.
Gills, 292.
Goethe, 26, 102, 103.
Gravitation, 401-406.
Gulf stream, 380.
Hairs, 80-86.
Haemal spine, 178.
H.i mapophysis, 178.
Heat, 408-411.
Hedgehog, cutaneous muscle of, 296.
Hegel, 471, 499, 500.
Hcrbart, 500.
Herbivora, teeth of, 219.
Homoeopbytes, 308, 420, 427, 431, 531.
Homologies, 25, 176, 308, 420, 422, 427,
430-432.
Homotypes, 25, SOS, 420, 427, 430, 431.
Homologous series in chemistry, 364.
Horn, 80.
Horse, foot of, 212.
Hydra, 272.
Hymcnoptera, 263.
Imaging power of miud, 442-448, 507, 516.
Inflorescence, 91.
Insects, 243-265.
Isomorphism, 359.
Kant, 53, 466, 495-499.
Laplace, 53; his cosmogony, 402-405.
Laws of Nature, 14, 15.
Leaves, arrangement of, 88, 89 ; leaf-veins.
angles of, 112-116; structure of, 131,
132.
Leech, 62.
Leibnitz, 494, 495, 501-503.
Ligaments, 75.
Light, 406-408.
Limbs, nature of, 189-192, 208-212.
Loins, vertebrae of, 195.
Ludicrous, sense of, 144 note, 489.
Magnetism, 19, 412, 413.
Malebranche, 493.
Man, preparations for, 346-353 ; distribu-
tion of, 384.
Mandibles of insects, 61, 252.
Maxillae of insects, 258.
Mechanical power, 32, 349, 350, 410, 411
416, 417.
Medus e, 277.
Mollusca, archetype, 223-226; modiflca
tions of, 227-232; fossil, 338.
Monsters, 429 note.
Mountains, 371, 372.
Muscle, 76, 294-298.
Nails, 80.
Neck vertebrae, 194.
Nervous tissue, 78; nervous system, 280-
2S8.
Neural spine, 178.
Neurapophysis, 178.
Number as a principle of order, 11, 15-18,
98, 1SS, 191, 192, 208, 215, 246, 270, 271,
814, 318, 518-525.
Number, faculty which discovers rela-
tions of, 458, 459.
Ocean. 373; currents of, 379.
Oken, 27, 181, 182, 434, 517 note.
Ophidia, skeleton of, 196-203; teeth of,
221.
Order, principle of, 1, 420-427, 429-489i
464, 467, 514.
Ostrich, foot of, 209.
Ovipositors, 264.
Ox. foot of, 211.
Phyllodla, 85.
Physical geography, 870-382.
Picturesque, 489, 490.
Pines, morphology of, 119-129.
Pistil, 94.
Pitchers, 85.
Planets, 389-391.
Plant, typical, 103.
Plato, 7. 11, 426, 465.
Pleurapophysis, 178.
Podura, 263.
Pollen, 78.
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INDEX.
539
Pre-established harmony, 431, 441, 442,
4G2-40T, 470-473, 494, 495, 501-503.
Prcfururations of Christ, 512-514.
Progression, theories of, 317-326.
Progressive plan in geology, 317-327.
Progressive plan of Providence, 525-527.
Property, faculty which discovers rela-
tions of, 459, 460.
Prophetic plan in geology, 327-333.
Pteropoda, 339.
Pythagorean system, 10, 465, 518.
Quincunx, 126.
Radiata, 267-279; fossil, 314, 334-836.
Pain, 377, 378.
Relations observed by mind, 450-452.
Respiratory system, 291-294.
Rodents, teeth of, 217.
Sacral vertebr.e, 195.
Schelling, 471, 497-499.
Seed, structure of, 82; nature of, 95,
140.
Serpents, 60, 196-203.
Shells of moliusca, 23, 65, 66, 815.
Skeleton, vertebrate, 175-212.
Sloth, 67, 68, 197.
Spinal cord, 288.
Spines, 86.
Spinnerets, 266, S37.
Spinoza, 493, 494
Spinal arrangement of appendages of
plant, 88-90, 96, 97; of conifera, 124-
128; of fossils, 311.
St. Ililaire, Geoffioy, 27, 431, 434
Stamens, 93, 137.
Stars, 392-400.
Stem, structure of, 84; typical form of,
187, 307.
Stipules, 85, 133.
Sublime, 490-492.
Sun's rays, 154, 411, 413, 414
Supports of plants, 134.
Tail offish, 56, 320.
Tail vertebr.i-, 200, 201.
Teeth, 213-222; fossil, 842,343.
Teleology, 30, 51, 428-43S, 514
Tendons, 75.
Time, as a principle of Order, 18-20.
Time, faculty which discovers relations
of, 456-458.
Tortoise, 204.
Types, 1, 23, 401, 456.
Types, organic, 364-366.
Typical appendage of plant, 83.
Typical bone, 1S5, 307.
Typical cell, 69, 70.
Typical limb, 192.
Typical numbers, 518-525.
Typical plant, 108.
Typical skeleton, 180.
Typical system of New Testament, 527-
532.
Typical system of Old Testament, 508-
518.
Typical tooth, 214, 215.
Typical vertebra, 178.
Universals, 466.
Vascular system, 288-291.
Vertebra, typical, 178.
Vessels, 72.
Whole and Parts, faculty whicb disoov
ers relations of, 452-454.
Winds, trade, 374, 375.
Wood, 72.
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