/. ^ EXPLANATION OP FIGURE I. OSSEOUS STRUCTURE. 1. Frontal bone. 2. Parietal. 3. Occipital. 4. Temporal. 5. Nasal. 6. Lachrymal. 7. Malar. 8. Superior maxillary. 9. Anterior " 10. Inferior " 11. Cervical vertebrae. 33. Scapula. 34. Humerus. From 4 to 7 is the zjgomatic arch. MUSCLES OF THE HEiiD, NECK, AlTD SHOULDER. *. Ligamentum colli. a". Trapezius. 6". Rhomboideus longus. c". Scalenus. e". Stcrno scapulari — pectoralis transversus. f". Antea spinatus. g". Postea spinatus. h". Teres. c. Dilator naris lateralis. d. " " anterior. e. Orbicularis oris. f. NasaKs longus. ff. Levator labii superioris. h. Buccinator. i. Zygomaticus. j. Retractor labii inferiorus. k. Masseter. I. Abducens aiu:em. m. Attolentes et adducens aiu-em. n. Retrahentes aurem. o. Adducens vel deprimens aurem. p. r. Tendon of the splenius and complexus major. q. ObUquus capitis superior. s. Splenius. t. Obliquus capitis inferiorus, u. Levator humeri. V. Sterno maxillaris. X. Subscapulo-hyoideus. 1. Temporal vein. 2. Facial vein. 3. Jugular vein. 10. Parotid gland. THE A.I:^^TOMY ^NT> FHYSIOLOaY THE HORSE: ANATOMICAL AND QUESTIONAL ILLUSTRATIONS. CONTAINING, ALSO, A SERIES OF EXAMINATIONS ON EQUINE ANATOMY AND PHYSIOLOGY, WITH INSTRUC- TIONS m REFERENCE TO DISSECTION, AND THE MODE OF MAILING ANATOMICAL PREPARATIONS. TO WHICH IS ADDED, #lossarg of ycttrinarjt Eethnicaliiits, oTovuological (fbart, anb giritonarg of ©tftrhiarn ^rirnre. GEORGE H. DADD, M. D., V. S., AnTHOK OP " THB MODERN HORSE DOCTOR," "CATTLE DOCTOR," ETC., ETO . BOSTON: PUBLISHED BY JOHN P. JEWETT AND COMPANY. CLEVELAND, OHIO : H. P. B. JEWETT. NEW YORK: SHELDON, BLAKEMAN AND COJIPANY. 1857. ST76ir Entered accoriliug to Act of Congress, in the year 1866, by JOHN 1". JEWETT AND COMPANY, In the Clerk's Office of the District Court of the District of Massachusetts. UinOTYPED BV THE AJIERICAN STEREOITPE COMPANY, PnOSNIX BUILDING, BOSTON. PREFACE. American Veterinary Literature has hitherto possessed no work devoted to the anatomy and physiology of the Horse ; consequently such subjects are either discussed theoretically and imperfectly, or else fail to be noticed. But a new era is now dawning upon Veterinary Science ; a spirit of inquiry is abroad; and the people of this Republic find themselves in possession of some of the most magnificent specimens of "Uve stock" to be found in the world. The natural inquiry is, "How shall we protect our property?" And the conclusion arrived at is, "Veterinary science offers the only probable and practicable security against the numerous casualties incidental to the halitats of the barn and stable." Hitherto, much indifference has been manifested regarding this science, in consequence of the difficulty encountered in its study, for want of proper text-books and teachers ; and its unsatisfactory results when tested by men unacquainted with its fundamental principles. The well-known works of Enghsh and French authors furnish all the necessary information, yet their cost is beyond the means of many, and, therefore, their circulation is very limited. In view, therefore, of supplying the above deficiency, which is disclosed in the barrenness of our anatomical and 23hysiological knowledge, and for the purpose of furnishing a work that shall come wuthm the reach and financial means of all men, the author has undertaken the double task ; and it is hoped that the effort will not be thought untimely. There are a vast number of highly educated physicians in this country who are often urged by their employers to give advice in the management, medical and surgical treatment, of the inferior orders of creation ; yet decline to do so, in consequence of a lack of authoritative knowledge regarding anatomy, physiology, therapeutics, and pathology. To such, whose sympathies lean in the right direction, and who are wiUing to give counsel, and lend a helping hand in the restoration of a sick or d3dng animal, this work is offered, and the author, therefore, submits it to their candid perusal and criticism. The work, however, is principally intended for veterinary surgeons, teachers of the art, and students of veterinary medicine, whose wants the author professes to have some knowledge of; and he has endeavored, to the best of his abiUty, to cater to the same. The necessity for such a work, at the present time, is evident from the facts, that three veterinary colleges have lately come into legislative existence, and (in) it is very uatui-al to suppose that, ere long, many candidates for the honors of these institutions will knock at the door of science, and seek admittance ; they must then need fe.ii-ho6ks ; and, m view of furnishing a part of what the author foresees every teacher and student must necessarily need, he oifers this, not as a work pregnant with his own ideas, for that were presumptuous, when anat- omy and physiology are the texts ; but, as a work carefully prepared from the writings of our best authorities, the work may be considered as the legitimate offspring of scientific observation and experience. Another argument in favor of the necessity that will soon exist for a text- book of anatomy and physiology is founded on the fact, that agricultural colleges will soon be endowed in every State of the Union ; many already exist; and each mil, probably, endow a professorship of veterinary science. With such, and among the young and aged men that may seek for knowledge^ the author hopes that his work may find favor; and, if such should be the result, he will have the satisfaction of knowmg that he has not labored in vain. There are other classes of men that need a work of this description; namely, the husbandman, the horse-owner, and the horse-lover, as well as the purely scientific man. The three first, incited by laudable sentiments, or pecuniary motives, will read the following pages, and study the anatomical illustrations ; some with veneration of that wonderful piece of mechanism, a horse's structure ; others for the purpose of making themselves acquainted with the form, action, and capacities of the same. The purely scientific man, who desires to inform himself how veterinary science is to be studied, — what are its legitimate objects, and its appropriate sphere, — will read these pages with considerable profit. GEORGE H. DADD. Boston, January, 1857. REMARKS IN REGARD TO THE COMPOSITION OF THIS WORK. The plau of the author, in the commencement of this work, was to prepare a complete manual of examinations on the Anatomy and Physiology of the Horse ; but findino- that he could not bring the mutter withm the prescril^ed limits, the plan was speedily abandoned. The examinations, exceptmg those descnptive of the osseous structure, are intended, either to elicit some physiological tact, or to introduce topics that have not been treated of in the body of the work. In attempting to furnish the public with a systematic treatise on Anatomy and Physiolocrj" it wiU be obvious that the author must necessarily avail himself of the labor of others; for, as regards the science of anatomy, no one has anythino- new to offer. The industrious anatomists and dissectors ot early times have°borne off all the laurels, and there remains but little, if anything, for future discovery. As regards physiology, also, there are very few tacts to discover: we now allude," however, to practical physiology— that science which teaches us the functions of the animal body, or the uses of its parts, ihe author has, however, occasionally stepped beyond the details of practical physiolo^ry, and has endeavored to throw some light on the complex combina- tions in which vital phenomena present themselves, and the nature ot their dependencies one upon another. _ Matter of this kind he has thought best to introduce in the form of examinations. ■, , ^ x In preparincr this work, the author has endeavored to select the most recent and rehable information. The following list of authors consulted and com- pUed from, together with the foot notes and the writers' names appended will serve to indicate the principal sources on which the author has rehed tor information. ^ , , , . Mr PercivaU's Aiiatom>/ of the Horse has been freely employed m composing the anatomical part of the work. The description of the abdommal viscera is from the pen of Mr. Gamgee, and was witten as a pme cssai/, and pub- lished in the London Veterinarian. Carpenter's Physiology, general and compar- ative, is also quoted. Liebig's Chenmtry, Hooper's Dictionary, Percivalls Ucppa- pathology, Eoget's Outlines of Physiology, have also been consulted, and extracts made from the same. The iUustrations, not otherwise indicated, are by Inrard; explanations translated by the author. For the loan of the French plates, the author is indebted to C. C. Grice, V. S., of New York City. ^ ^^ , .. , The plan of the examinations was suggested to the author |jy Ludlow s Manual, of Examinations,— ii work which he formerly, while studying medicine, had occasion to use. The subject matter, in this work, of course diflers kom that ot "fcliG foniiGr In preparino- the ''Definitions of Veterinary Technicalities," and "Diction- ary" the author has availed himself of the works of Cooper, Hooper, Cleve- land, Blaine, Mahew, and White ; and, regarding the method of making ana- tomical preparations, etc., the works of Parsons, Pope, and Swan, have been consulted. ^- ^- ^- CONTENTS. FAas Preface, - 3 Remarks regarding the composition of the work, .-.-..--.. 6 Remarks on the osseous, cartilaginous, and ligamentous structures, .-..-. 11 TeGUMEXTARY System. — On the hair of horses ; examinations on the common integument; physiol- ogy of the skin, of the cellular membrane, of the adipose tissue ; examinations on the same, 14-17 Of the External Parts. — The hoof; its form, spread, color, magnitude ; the wall ; its situation and relation, connection, figure, di\'ision, solar border, laminae, quarters, heels, coronary border and bars, 17-23 The Sole. — Situation and connection, figure, arch, di\ision, surfaces, and thickness, - - 23-25 The Frog. — Its situation and connection, figm-e, division, surfaces; the cleft of the frog, its supe- rior sm-face, the sides, the commissures, toe, heels or bulb, coronary frog band ; development of hoof; structure of the hoof; production of the hoof; propftties of horn, .... 25-30 Intern.u- Parts of the Hoof. — The coronary substance ; its situation, connection, structure, and organization, ..-.-...-....-. 30-31 The C.\RTn,AGES. — Their situation, attachment, and form ; the false cartilages, and their use ; the sensitive laminae ; division of the same ; elasticity, number, dimensions, and organization ; the sensitive sole ; its structure, connection, thickness, and organization ; the sensitive frog ; its situation, division, sti-uctm-e, and organization, ...-..--- 31-34 A tabular \iew of the bones of the horse, -...-...-- 35 Anatomy of the skeleton, introduced in the form of questions and answers, embracing a complete system of osteology, -.--...------- 36-54 Remarks on the changes which horses' teeth undergo, with examinations on the same, - - 54-56 Myology. — A table of the names and number of muscles, divided into regions, ... 57-60 A tabular s}'nopsis of the number, name, region, situation, insertion, and action of all the muscles, 61-78 Ox DissECTlox. — Dissecting instruments; subjects suitable for dissection; rides in reference to dissection of muscles, .......-..-.- 79-80 Anatomical Prep.aratioxs. — Injecting instruments; directions for using the sjTinge, - 80-81 On THE DiFFEREXT KiXDS OF INJECTIONS. — FormultB for coarse warm injections ; fine injections ; minute do; plaster mjection; cold injection; as regards the course of injections ; quicksilver injections ; mode of injecting the Ij-mphatics with quicksilver ; method of injecting the lac- teals, and parotid gland ; wet preparations ; preparations by distension ; method of preparing and distending the limgs ; menstrua for preserving specimens ; method of preserving the brain and lungs ; method of macerating and cleaning bones ; to render bones flexible and transpa- rent; method of cleanmg and separating the bones of the cranium; exposition of Mr. Swan's new method of making dry anatomical preparations, -------- 81-87 Digestive System. — The mouth, lips, cheeks, gums, palate, tongue, salivary glands, pharj-nx, oesophagus, and nasal fossa; cavity of the cranium; the orbits and cavities of the nose ; the mouth, peritoneum, stomach, intestines ; the vessels, nerves, and IjTnphatios of the intestines ; the spleen, liver, pancreas, kidneys, supra, renal capsules, ureters, bladder, urethra, - - 87-119 GENER.vTrvE Org.vns OF THE M.ALE. — Vasa defercutia, vesiculae seminales, ejaculatory ducts, pros- trate gland, Cowper's glands, -......-•..- 119-121 0RG.4NS OF Generation Continded. — Testicles and scrotum, spermatic cord, epididjTnis, penis, and urethra, - 121-125 Fem.\le Organs of Generation, 125-128 Physiological considerations on the reproduction of organized beings, ..... 128-136 Examination on the digestive system, 136-138 Remarks and examinations on the eye, - 139-143 (vii) vni CONTENTS. Respiratory System. — Observation on the same; the larynx, glottis, epiglottis, trachea, hron- chial tubes, pleura, mediastinum, lungs, bronchial glands, ------- 144-154 Circulatory System. — Remarks on the blood ; examinations resumed on the blood, pericardium, and heart, ------- 155-167 Arterial System. — Distribution of the arteries, 158-163 A table showing the modes of the distribution of the arteries, - 164^166 Distribution of the veins, ...-.--.-.--- 166-168 A table showuig the mode of distribution*bf the veins, -- 169-170 The brain and its appendages ; the nervous system, - 171-177 Examinations on neurology, .--...---.-- 177-180 Distribution of the lymphatics, 181-184 A glossary of veterinary technicalities, - 185-193 Toxicological chart, - - - - 195-209 A dictionary of veterinary science, - 211-287 Appendix. — Ligamcntary mechanism of articulations and joints, ----- 289-291 INDEX OF THE ILLUSTRATIONS. FIGUKE I. Presents two ^iews : one of a portion of the osseous structure, showing the head, neck, and shoulders ; and the other is composed of the supei-ficial muscles, covering the above parts ; precedes the title page. -------------- FIGURE II. Is a section of the osseous structui'e, giving a side of the spinal column, ribs, and a part of the rear, anterior, and posterior extremities. 10 FIGURE III. — Is a representation of the superficial muscles of the body, of a part of the neck, and of the extremities, ...------------'■0 FIGURE rS'. — Has fom- illustrations of the hind extremities, as follows : No. 1 is a side -view of the bones of the oft-hind leg ; No. 2 shows the muscles and tendons of the oiT-hind leg ; No. 3 is a front view of the bones of the same ; No. 4 shows the muscles and tendons in the anterior region, or front part, of the off-hind extremity. .---- 30 FIGURE V. — Presents Ueo illustrations : the first shows the superficial muscles in the region of the head, neck, and shoulders, on the near side ; and the other is a corresponding section of the osse- ous structm-e, on wliich the insertions of the ligamentum colli into the occiput, cervical vertebrae, and dorsal spines, are sho\ra. ...----------40 FIGURE VI. — Presents four views of the forward extremities : No. 1 shows the bones which enter into the composition of the near fore-leg ; No. 2 is a side view of the muscles and tendons of the near fore-leg ; No. 3 is an anterior \iew of No. 1 ; No. 4 is an anterior view of No. 2. - - 50 FIGURE Vn. Presents four views of the near fore-extremity : Nos. 1 and 3 are side and posterior views of the bones of the near fore-limb ; Nos. 2 and 4 show the muscles and tendons belonging to the above regions. ..-.---------- oO FIGURE Vin. — Has four views of the off-hind extremity: Nos. 1 and 3 are side and posterior views of the bones entering into the composition of the Hmb ; Nos. 2 and 4 show the muscles and ten- dons of the same. .-.----------- 70 FIGURE IX. — Presents two views : one, of the bones ; the other shows the superficial muscles of the head, neck, shoulders, and breast, viewed in an anterior direction. ----- 80 FIGURE X. — Has two cuts : one of wliich shows a portion of the osseous framework ; the other shows the superficial muscles belonging thereto. --- 90 FIGURE XI. — Is illustrated by two cuts : one of which shows a portion of the muscles of the body, neck, and Umhs ; it is a sort of anterior side riew ; the second cut shows the bones which enter into the composition of these parts. --.---------100 FIGURE Xn. — Has two illusti-ations, which are the counterpart of Fig. XI., as seen from the oppo- site or posterior dii'ection. .-.-.-------- 119 FIGURE XHI. — Presents a side view of the deep-seated muscles : it is talien from Mr. Blaine's " Out- Knes," and is one of the most magnificent plates ever presented to the pubKc. - - - - 120 FIGURE XrV. — Is a riew of the mu.scles and tendons of the fore and hind extremities. - - 140 FIGURE XV. —Is illustrated with five views of the off and rear fore extremity : Nos. 1, 2, 3 show very distinctly the action of the flexors of the Umb, as well as their location, and that of the extensor tendons and muscles. The triceps extensor brachii, and pectoral muscles, are also quite prominent and easily recognized ; No. 4 is the same hmb divested of the soft parts ; No. 5 is an interior view of the near fore-leg, and shows some of the tendons and muscles wliich are not seen in the other cuts. ----- 150 FIGURE XVI. — Presents five views of the hind extremities, in wliich the use and action of several very important muscles and tendons are accurately delineated : Nos. 1, 2, 3, 4 compose the bones, muscles, and tendons of the near-hind extremity ; No. 5 shows the muscles and tendons on the inside of the near-hind leg. ------------- 160 2 (ix) X INDEX TO THE ILLUSTRATIONS. FIGURE XVn. — Presents two views (as seen from a posterior direction) : one contains a great portion of the superficial muscles of the body and limbs, and the other shows the basis of their super- structure. --.-.--...------ 170 FIGURE XVm. — Is the skeleton of a horse, for which the author is indebted to Blaine's " OutUnes of the Veterinary Art." ..-- 180 FIGURE XIX. — Is a counterpart of Fig. XVn., as seen from an opposite direction. ... 175 FIGURE XX. — Is an excellent representation of the muscles of one side of the head, neck, body, and Kmbs. 211 [The author considers it due to himself to remark, that, in consequence of not seouiing from the engraver good proofs of the plates, there will occur a few inaccuracies between the lettering ou the cuts and explanations accompanying them. These, however, are not of material consequence ; yet, if necessary, the reader can, from analogy, — by comparing one plate with another, — correct the errors with a pen.] EXPLANATION OF FIGURE II. OSSEOUS STRUCTURE. 11. Cervical vertebra. 12. Dorsal " 13. Lumbar " 14. Sacrum. 15. Coccygeal bones. 16. True ribs. 17. False " 18. Sternum. 19. Pelvis. 20. Posterior part of the pelvis, or ischiatic spines. 21. Inferior, or pubic region. 22. Femur. 23. Patella. 24. Tibia. 33. Scapula. 34. Humerus. 35. Radius. e. Fibula. /. Ulnar. REMARKS OSSEOUS, CARTILAGINOUS, AND LIGAMENTOUS STRUCTURES. The bones are the solid framework which fibrous arrangement is more irregular and gives stability to the whole fabric, and af- ford fixed bearings upon which the powers regulating the varied movements operate. The bones, then, are considered as the most dense and solid structures of the animal frame : affording support, and in many parts protection, to some of the softer parts ; at the same time, the leverage which regulates the action of a Kmb is derived from the osseous structure. On making an examination of a bone, we find that its external surface is the hard- est part, and it differs very much in thick- ness in different bones, and in different animals. The long bones (or cylmdrical) of the horse contain less marrow, and are more cancellated within, than the bones of the human subject: in many of the former the whole arena is occupied by cancelli. The bones of the ribs have an osseous plat- ing differing in thickness in various sub- jects, and within is a cellular structure which may be termed diploe. The marrow, as it is termed, is a soft substance of an oleaginous character, con- tained in an infinite number of sacs, depos- ited and suspended in the cavities of bones and in the canceUi. The marrow sacs are composed of a delicate vascular membrane, which isolates them from each other, and prevents the marrow from gravitating or passing into the osseous structure. Bones present the appearance of lamella, yet they are fibrous ; the fibres of the cylin- drical bones are longitudinal; in the flat bones they have a radiated appearance, and in the short and peculiar shaped bones, the difficult to trace. The basis of the osseous structure is nearly the same as the membranous parts,* being composed of fibrous laminae or plates, which are connected together so as to form, by their intersection, a series of cells anal- agous to those of the cellular structure. This theory has been disputed by some distinguished physiologists; the moderns contend that the osseous fabric is ceUular.f Bones are invested, on their exterior, ex- cept those parts plated with cartilage, with a membrane termed periosteum. Through this medium an arterial and venous com- * " The analysis of a bone into its two constituent parts is easily eftected by the agency either of acids or of heat. By macerating a full-gro-mi bone for a sufficient time in diluted muriatic acid, the earthy portion of the bone, amounting to nearly one-thu-d of its weight, is dissolved by the acid; the animal portion only remaining. This animal basis retains the bulk and shape of the original bone, but is soft, flexible, and clastic ; possessing, in a word, all the properties of membranous parts, and corres- ponding in its chemical character to condensed albumen. A portion of this solid animal substance affords gelatin by long boiling in water, especially under the pressure, ad- mitting of a high temperature, to which it may be si'.b- jeeted in Papin's digester. On the other hand, by sub- jecting a bone to the action of fire, the animal part alone will be consumed, and the earth left untouched, preserv- ing, as before, the form of the bone, but having lost the material which united the particles, presenting a fragile mass which easily crumbles into powder. Tliis earthy basis, when chemically examined, is found to consist prin- cipally of phosphate of lime, which composes eighty -two hundredths of its weight ; and to contain also, according to Berzelius, minute portions of fluate and carbonate of lime, together with the phosphates of magnesia and of soda." — Eoget. t The best authority in support of the cellular theory is Scarpa. Percivall advocates the laminated and fibrous theories. (11) 12 ANATOMY AND PHYSIOLOGY OP munication is established between the dense and soft parts. The periosteum is anal- agous to the fibrous textures, being com- posed of numerous inelastic fibres of great strengtJi and density. The inner surface of the periosteum is connected with the bone by the vessels pass- ing from the one to the other, and also by numerous prolongations, which pervade the osseous substance. The blood-vessels of the periosteum are numerous, and are easily demonstrated by injection. CARTILAGE.* The structure which appears most inti- mately connected with the osseous is carti- lage. It is a firm and dense substance, apparently homogeneous in its texture, semi- pellucid, and of a miUc-white or pearly color. The surface of cartilage is smooth and uniform, presenting neither eminences nor cavities, pores nor inequalities. It has, however, minute capillary vessels, the diam- eters of which are too small for ocular demonstration. Notwithstanding its den- sity, it has a minute circulating apparatus, wliich is demonstrated in diseases known as spavin and ringbone, in which absorp- tion of cartilage occurs. Cartilaginous structures are chiefly com- * " The mechanical property which particularly distin- guishes cartilage is elasticity, a quality wliich it possesses in a greater degree than any other animal structure, and which adapts it to many useful purposes in the economy. Hence it forms the basis of many parts where, contrary to the purposes answered by the bones, pliancy and resistance as well as firmness are required ; and hence cartilage is employed when a certain shape is to be preserved, to- gether with a capability of yielding to an external force. The flexibility of cartilage, however, does not extend beyond certain limits ; if these be exceeded, fracture takes place. Great density bestowed upon an animal stnictiu'e, indeed, ajipears to be in all cases attended with a propor- tionate degree of brittleness. These mechanical proper- ties of cartilages, as well as their intimate structure, although nearly homogenous in all, are subject to modifi- cation in different kinds of cartilage. Cartilages are covered with a fine membrane, termed the perichondrium, analogous in its structure and office to the periosteum, which we have already had occasion to point out among the fibrous membranes, as investing the bones." — Ror/et. posed of gelatin, albumen, and phosphate of calcis. Cartilage occurs in two forms, temporary and permanent. The former prevails pre- vious to adult life ; the latter are identical with the permanent structures after the animal has migrated from colthood. There are three or four different forms of cartilages, viz : the membriform, interosseal, articular, and inter-articular. Tlie membriforni are fibro-cartilaginous ; they furnish a basis of support to the softer parts, supply the place of bone, and give form, shape, and firmness, to parts miossi- fied. By then- elasticity, they admit of con- siderable variation of figure and form, yield to external pressure, and recover their proper shape as soon as pressure is removed. This kind of cartilage is found in the nostrils, ears, larynx, and trachea. T/ie interosseal cartilages pass from one bone to another, adhering firmly by their extremities to each. They permit of an increase of extent or motion, as observed between the ribs ; when macerated, they are divisable into lamina of an oval shape, which are united by fibres passing obliquely between them. The articular cartilages are those plates of articular substance which adhere firmly and inseparably to t'lie surfaces of bones which arc opposed to each other in the joints, or over which tendons and ligaments play. The elastic resistance of this carti- lage has a powerful tendency to lessen the shocks incident to sudden and violent actions. The inter-articular cartilages do not dif- fer in composition from the preceding. They are attached to the inside of the cap- sular ligament, by which they arc rendered somewhat movable ; and, being interposed between the bones of the knee and hock, allow them a greater latitude of motion, while at the same time they contribute to adapt their siu-faces more perfectly to each other. The structure of these cartilages is laminated. THE HOESE. 13 FIBRO-CARTILAGINOUS STRUCTURES. Fibro-cartilage appears to be of an in- termediate nature between ligament and cartilage. Having a fibrous texture united to a cartilaginous basis, it combines the characteristic properties of both of the above textures. Fibro-cartilaginous sti-uctiures are found to unite the bodies of the bones of the ver- tebrae ; they then get the name of inter- verte- bral svibstance. They impart great elasticity to the spine, and also diminish the effects of concussion. LIGAMENTOUS STRUCTURES. The ligamentous structures are dense; possess a considerable degree of solidity in some parts, while in others they are modi- fications of fibrous membrane. The liga- mentous system includes a number of parts which have received different names, such as ligaments, tendons, faschia aponeurosis, capsiiles or bursse mucosa; and fibres of ligamentous matter also enter into the com- position of other organs, imparting to them different degrees of mechanical strength. The ligamentous structures vary; we find that in some places they are expanded into faschia, etc., at others they collect into dense, enlonsated cords. The first division in- cludes fibrous membranes, fibrous capsules, tendinous sheaths, and aponsurosis. Fibrous membranes: these resemble or- dinary membranes, only that their fibres are denser. The periosteum is a membrane of this description, and the dxira mater has a similar structure. Fibrous capsules are presented in the form of sacs, which surround various ten- dons and joints. These capsules are also lined by a synovial membrane, which secretes the synovia. Tendinous sheaths are formed by fibrous membranes which surround the tendons, in those parts that are subjected to fi-iction, or liable to displacement, during the action of the muscles which move the joint. Aponeurosis are those extended sheets of fibrous texture wliich in some instances form coverings of parts, while in others they constitute points of attachment to muscles. In the former case they are termed faschiae, and either surround the muscles of a limb, forming a sheath for it, or else invest or confine some particular muscle. In the latter case the aponeurosis presents broad, or narrow, surfaces and fibres which give attachment to particular portions of muscle. ANATOMY AND PHYSIOLOGY. TEGUMENTARY SYSTEM. ON THE HAIR OF HORSES. Hair is a peculiar tegumentary appendage, characteristic of the horse and other mam- mals. It is developed on the interior of oUicles which are formed by a depression of the true skin. These folUcles are lined by a continuation of the epidermis, the cells of which are developed in peculiar abundance from a spot at its deepest portion ; the dense exterior of the cluster thus formed being known as the bulb of the hak, while the softer interior is termed its pulp. The elementary parts of hair are : a cortical or investing substance of a dense horny tex- ture; and a medullary or pith-like sub- stance, of a mvich softer character, occupy- ing the interior. The cortical envelope of hairs is a continuation of the outer scaly layers of the epidermis that lines the follicle ; whilst the medullary is derived from the deeper stratum whose cells are produced in unusual abundance at the .coecal extremity ; and it is by the constant development of new cells at this point, that the continual gi-owth of the hair is kept up. An excoriation or moulting of the hair, which falls off, is replaced by a new gro^vth, which as it comes to maturity assumes the original color. This change in the covering with which nature has so wisely clothed the horse, usually takes place either in spring or autumn, or at both periods. The hair of the mane and tail, however, is not sub- jected to these periodical changes; hence, it acquires considerable calibre and length. By analysis, the hair yields carbon, hy- drogen, nitrogen, oxygen, and sulphur, and its variation in color is due to the presence of dilTerent shades of matter which infil- trates the cortical substance. EXAMINATIONS ON THE COMMON INTEGUMENTS. ^. Of how many parts do the common integuments consist ? — A. Three : cuticle, cutis, and rete mueosum. Q. Describe the cuticle or epidermis? — A. It is a thin, transparent, tough, and elastic porous membrane, serving as an envelope to the cutis, or true skin. It is composed of flexible lamellae, so arranged as to bear some analogy to the scales of fish ; it pervades the whole body, and insinuates itself into porous struc- tures and folHcular passages, inlets, and outlets of the system ; it is supposed to be continous from the mouth to the anus. Q. Describe the pores or perforations. — A. There are tliree. First, those surrounding the hair. Sec- ondly, exhalent pores. Tliirdly, absorbent pores. Q. How is the cutis designated? — A. As the cutis vera, or true skin. Q. '\Vhat is the structure of the cutis? — A. It is of a fibrous texture, tough, clastic, vascular, and highly sensitive, and is what we commonly denominate leather. Q. What are the attachments of the cutis? — A. The cutis is attached to the subjacent parts by cellu- lar membrane, in some places so tensely that Uttle or no motion is admitted of; in others so loosely that it admits of being thro-mi into folds. About the forehead, upon the back, around the tail, and upon the pasterns, it can scarcely be pinched up ; but upon the sides of the face and neck, upon the ribs, along the flanks, and upon the anus and thighs, it wQl easily admit of duplication. Q. "What varieties are there in the density of the cutis ? — A. It varies in density, not only where it covers difierent parts in the same animal ; but in horses of difierent breeds and temperaments, it varies very essen- tially. Q. What is the organization of the rete mucosimi ? A. It is composed of a fine, deHcate, laminated tissue, interposed between the cuticle and cutis, and serves as their connecting medium, and is supposed to secrete the coloring matter of the external surface and haii-. (14) ANATOMY AND PHYSIOLOGY OF THE HORSE. 15 PHYSIOLOGY OF THE SKIN. The sldn is highly sensitive ; yet those persons who are in the habit of making free use of the whip scarcely ever realize the fact. The author has an impression that the sliin of a horse is more sensitive than that of man ; for example, let a small quan- tity of turpentine be applied to a horse's back, — very soon he evinces signs of pain, which cannot be elicited when a 7nan be- comes the subject of the same experiment. Every horse-owner, also, must have noticed the uneasiness a horse manifests when a common fly, or gad-fly, alights on him; and in a variety of other ways the highly sensi- tive state of a horse's skin admits of dem- onstration. -The skin is one of the principal emunc- tories of the body, from the surface of which passes ofl" a large quantity of morbid fluid in the form of perspiration, sensible or in- sensible, as the case may be. The skiii is the great external outlet ; and, should the kidneys or any other organ fail to play their part in eliminating useless fluids, the skin opens its flood-gates, and thus purifies the body. The amount of fluid exhaled from the external siurface has been the subject of some very interesting experiments, and the results are truly astonisliing. OF THE CELLULAR MEMBRANE BENEATH THE SKIN. This tissue abounds in almost every part of the body; thus, says Carpenter, "it binds together the ultimate fibres of the muscles into minute fasciculi, unites this fasciculi into larger ones, these again into still larger, which are obvious to the eye, and these into the entire muscle ; and also forms the mem- branous divisions between distinct muscles. In like manner it unites the elements of nerves, glands, etc., binds together the fat- cells into minute masses, these into larger ones, and so on ; and in this manner pene- trates and forms a considerable part of all the softer tissues of the body. It also serves as the bed in which blood-vessels, nerves, and lymphatics may be carried into the substance of the different organs." This tissue consists of a net-work of minute fibres and bands, which are inter- woven in every direction, so as to leave in- numerable areola, or spaces, which commu- nicate fi-eely with one another. Of these fibres, some are of the yellow or elastic kind, but the majority are composed of the white fibrous tissue ; and, as in the other form of elementary structure, they frequently present the form of broad flat- tened bands, or membranous shreds, in which no distinct fibrous arrangement is visible. The proportion of the two forms varies, according to the amount of elasticity or simple resisting power which the en- doA\Tnents of the part require. The inter- stices or areolEB are fiUed, during life, with a fluid which resembles very dilute serum of the blood ; consisting chiefly of water, but containing a sensible quantity of common salt and albumen. It is the undue accumu- lation of this fluid wliich constitutes drop- sical effiision, the influence of gravity upon the seat of which, shows the free communi- cation that exists among the interstices. This freedom of eommunication is still more shown, however, by the fact, that either air or water may be made to pass, by a moderate continued pressure, into almost every part of the body containing cellular or areolar tissue, although introduced only at a single point. Li this manner it is the habit of butchers to inflate veal; and impostors have thus blown up the scalps and faces of their children, in order to ex- cite commiseration. The whole body has been thus spontaneously distended with air by emphysema in the lungs; the air having escaped from the air-cells into the surround- ing areolar tissue, and thence, by the con- tinuity of this tissue with that of the body in general at the root or apex of the lungs, into the entire fabric. The structure of the serous and synovial membranes is essentially the same as the above. The true cellular membrane is sometimes termed reticular, while that con- taining fat is called adipose . 16 ANATOilY AND PHYSIOLOGY OF ADIPOSE OR FATTY TISSUE. The adipose tissue is composed of iso- lated cells, which appropriate fatty matter from the blood after the same manner as the secreting cells appropriate the elements of bile, urine, and milk. " The portion of fatty matter separated from the circulating fluid to form adipose tissue, is only that which can be spared from the other pur- poses to which they have to be applied ; and hence the production of this tissue depends, in part, upon the amount of fatty matter taken in as food.* This is not entirely the * " Deposition. — In almost all animals that are healthy, copious food of a nutritive kind, combined with little labor, will increase the deposition of fat ; but in the human sub- ject, and, indeed in many quadrupeds, the animal spirits appear to hare very considerable influence over this secre- tion. We see numberless examples of people, wlio appear to enjoy the best bodily liealth, and yet are constantly meagre, though their food and habits of life tend to an op- posite state ; and we may occasionally obsei-ve horses and dogs, particularly circumstanced, in which, from their natural leanness, or poorness upon the rib, something of a mental nature would appear to be operating ; indeed, it is a well known truth, that if you separate a horse of an irritable disposition from others with whom he is accus- tomed to be stalled, he wUl fall away in condition, in con- sequence of (to use the vulgar expression) fretting from beinrj alone; and so much does tliis act of segregation affect some, that I have known them even refuse then- food. Those horses are commonly the fattest that are fed on easily digestible food — such as braised or scalded corn, roots of a nutritive kind, chopped hay, etc., and that have little or no exercise : a fact well appreciated by the horse-dealer, whose horses are^fne and Jit for sale, but in- capable of fatigue. Absorption. — Constitutional diseases, generally speak- ing, extenuate the body, and more particularly such as are of the acute or painful description ; lience, tlie irritation caused by a simple puncture in the foot, will, if it be of long duration, induce a state of emaciation : under which circumstances, the absorbents are supposed to act with more than ordinary effect, and to take up the adeps from the interior of its cells — Percivall. case, however, as some have maintained ; for there is sufficient evidence that animals may produce fatty matter by a process of chemical transformation, from the starch or sugar of their food, when there is an unusual deficiency of it in the aliment." Liebig writes : " Whatever views we may entertain regarding the origin of the fatty constitu- ents of the body, this much, at least, is un- deniable, that the herbs and roots consumed by the cow contain no butter ; that, in the hay or other fodder of oxen, no beef-suet exists ; that no hog's-lard can be found in the potato refuse given to swine ; and that the food of geese or fowls contains no goose nor capon fat. The masses of fat found in the bodies of these animals are formed in their organism; and, when the full value of tliis fact is recognized,it entitles us to conclude, that a certain quantity of oxygen, in sorrie form or other, separates from the constituents of their food, for without such a separation of oxygen, no fat could possibly be formed from any one of these substances." The chemical analysis of the constituents of the food of the graminivora shows in the clearest manner that they contain carbon and oxygen in certain proportions; which, when reduced to equivalents, yield the fol- lowing series : " In vegetable fibrine, albumen, and cas- eine, there are contained, for — 120 eq. carbon, 36 eq. oxygen. In starch, 120 " " 100 " " cane sugar, 120 " " 110 " " " gum, 120 " " 110 " " " sugar of milk, 120 " " 120 " " " grape sugar, 120 " " 140 " " EXAMINATIONS RESUMED. CELLULAR MEMBILVNE. Q. What is the principal use of cellular membrane ? — A. It is employed in uniting, covering, and defending various parts of the body. Q. Does cellular differ from serous or nervous mem- branes ? — A. No, they are all resolvable into the same constituents. Q. How does the periosteum differ from the above ? — A. It presents itself in a more condensed form. Q. How do capsules of joints differ from common cellidar membrane .' — A. They are a modification of it, under a condensed form. Q. In what part of the animal does cellular mem- brane exist in greatest abundance? — A. Immediately beneath the skin ; upon the ribs, and about the breast, and under the jaws, in the scrotum, on the inside of the elbow and thigh. THE HORSE. 17 " Now in all fatty bodies there are con- tained, on an average : " For 120 cq. carbon, only 10 cq. oxj-gen. " Since the carbon of the fatty constit- uents of the animal body is derived from the food, seeing that there is no other source from whence it can be derived, it is obvious, if we suppose fat to be formed from albumen, fibrine, or casein, that for every 120 equivalents of carbon deposited as fat, 26 equivalents of oxygen must be separated from the elements of these sub- stances ; and, further, if we conceive fat to be formed from starch, sugar, or sugar of mUk, that for the same amount of carbon there must be separated 90, 100, and 110 equivalents of oxygen from these com- pounds respectively. " There is therefore but one way in which the formation of fat in the animal body is possible, and this is absolutely the same in which its formation in plants takes place ; it is a separation of oxygen from the ele- ments of food." OF THE EXTERNAL PARTS. THE HOOF.* " The hoof is the horny case or covering nature has pro\Tided for the protection of the sensitive parts of the foot. It may be said of itself to constitute such a shoe or defence, as enables the animal in his wild state to travel about in quest of food, not only without injury to the structures under- neath it, but with a degree of elasticity that preserves his whole frame from concussion. Were one forced into any comparison of the sort, it must be admitted that the hoofs of animals bear some anatomical affinity to the human nails, or claws, of other animals ; though they are vastly superior in physiological importance to any such appendages as these. * Percivall's Anatomy. EXAIVIINATIONS RESUMED. ADIPOSE TISSUE. Q. What is the fatty matter contained in the adipose cells composed of? — A. Stearine, margarine, and oleine. Q. How do they appear when isolated ? — A. The two former are solid, and the latter is fluid. Q. How are they preserved in a fluid state in the animal body ? — A. By the ordinary temperature of the body. Q. What are the observable differences in color occurring in different parts of the body, and in animals of diverse temperaments ? — A. In some parts of the body it is white, in others it has a yellow tinge ; in animals of IjTnphatic and ner\-ous temperaments it is white; in the sanguine it has somewhat of a red tinge ; in the bilious, it presents a yellow appearance. Q. The fat at the ordinary temperature of the li\-ing body being fluid, how is it retained in the fat cells Tvith- out transudation ? — A. The mtervals of the fat cells are traversed by a minute net-work of blood vessels, from wliich they derive their secretion ; and it is prob- ably by the constant moistening of then- walls with a watery fluid, that theu- contents are retained. Q. What are uses of the adipose tissue? — A. It is intended to fill up spaces ; fomis a sort of cushion or pad for the support of movable parts. It also acts as a non-conductor of heat, thus preserring the animal temperature ; it serves as a reservoir of combustible matter, at the expense of which the respiration may be maintained when other materials are deficient. Q. Suppose you desired to fatten a horse or an ox, what method should you adopt ? * — A. I should keep the animal at rest, and furnish him with an abundance of nitrogenized food. Q. In what vegetable constituents does nitrogen abound? — A. In vegetable fibrine, albumen, and caseine. * Experience teaches us that, in poultry, the maximum of fet is obtained by tying the feet, anil by a medium temperature. These animals in such circumstances may be compared to a plant possessing ia the highest degree tlie power of converting all the food into parts of its own structure. The excess of the constituents of blood forms flesh and other organized tissues, while that of starch, sugar, etc., is converted into fat. When animals are Cittened on food destitute of nitrogen, only certain parts of then- structure increase in size. Thus, in a goose, fattened in the method above aUuded to, the liver becomes three or four times larger than in the same animal, when well fed with free motion, while we cannot say that the organized structure of the liver is thereby increased. The liver of a goose fed in the ordinary way is firm and elastic ; that of the imprisoned animal is soft and spongy. The difference consists in a greater or less expan- sion of its cells which are filled with fat. Insome diseases, the starch, sugar.etc., of the food obviously do not undergo the changes which enable them to assist in respiration, and consequently to be converted into fat. Thus, in diabetes mellitus, the starch is only converted into grape sugar, which is expelled from the body without further change. In other diseases, as for example in inflammation of the liver, we find the blood loaded with fat and oil ; and in the composition of the bile there is nothing at all inconsistent with the supposition that some of its constituents may be transformed into fat. 18 ANATOMY AND PHYSIOLOGY OP " Form. — Sainbel viewed the foot as 'the segment of an oval, opened at the back, and nearly round in front.' To a common observer, the hoof exhibits a conoid form ; the part resting upon the ground being the basis, the vacuity above, the ob- truncated apex. Mr. Bracy Clark asserts that this -view is incorrect, and that the general figure of the hoof is a cylinder, very obliquely truncated upon its ground surface. This he demonstrates in two ways ; either by rolling up a piece of paper into the shape of a cylinder, and afterwards cutting one of its ends in a very slanting direction ; or by taking a carpenter's square, and placing one limb beneath the foot across the quarters, then sloping the other backward against the side of the quarters, parallel to the front, when the edge of the iron will be found parallel to the wall of the hoof. This corrected view of its figure will serve to account for the general equi- formity manifest in the hoof, and also for the undeviating correspondence found to exist between its slope or slant, as well in front as behind, which in an ordinary or healthy foot may be estimated at an angle of 45°. Around the coronet, where the hoof unites with the skin, the cylinder is cut directly across its perpendicular — at right angles with it : it is the oblique trun- cation of its ground-surface that occasions the slant, which latter we may consequently increase at pleasure by any means that augment the former, viz. i by lowering the heels ; by cutting away a prominent frog ; or by putting on thin-heeled shoes. At the same time that we increase the slant of the hoof, we increase the obliquity of the pas- terns, and liliewise proportionately augment the ground-surface of the hoof, from heel to toe, the breadth remaining unaltered ; and in the same ratio, consequently, extend the surface of tread.* " Spread. — By the spread, is meant the inclination the hoof manifests, when left unshod, around the toe and sides, to bulge * For further elucidation on the cylindrical form of the foot, consult Mr. Bracy Clark's works on the Foot of the Horse. or protrude at bottom, whereby its ground- surface becomes augmented, particularly around the outer quarter. To a certain ex- tent tliis is worthy of observation ; although, in my opinion, it is to be regarded rather as an effect of pressure than one of abstract growth. The surface of inclination upon which the horn is produced has no such spread, nor can the hoof itself be said, /row growth alone, to have any such natural ten- dency; but, as it continues to grow and shoot beyond the inner foot that produced it, and to which it was so intimately united, it yields to the pressure of the animal's weight, and bulges or spreads out, and more at the outer side than the inner, in consequence of the pressure tending more in that direction. If we examine a num- ber of hoofs of neglected growth, and con- sequent exuberance and deformity, of va- rious descriptions, we may discover that, in them all, the spread seems to have been the first or incipient deviation from that line of growth viewed as consistent with the health and well-doing of the foot. It is only in the unshod hoof that any spread is found : as soon as the ground-surface comes to be confined by a shoe, pressure can no longer exert its influence to produce such consequences. " Mjc. Goodwin aptly observes, that ' to take the form of the hoof correctly, we must strip it of its exuberant or superfluous parts, the same as one would pare the su- perabundant growth off" our own nails. The neglect of this necessary preparative has led to a considerable difl'erence of opinion about the natin-al, healthy, or true form of the ground-surface of the foot. Mx. Bracy Clark, I conceive, has inclined to the side of error in this particular ; though, in the substitution of the cylin- drical for the conical figure of the entire hoof, he has certainly the advantage of other writers. His natural foot is one with great spread to it, much of which the smith would find it necessary to de- prive it of, even on the first shoeing ; and the protuberance of the outer quarter (which Mr. C. points out as an attribute THE HORSE. 19 of health) being wholly owing to the spread, will, of course, disappear with the annihi- lation of the spread.' * " Although Mr. Goodwin has not here explained what he conceives to be the origin or cause of the spread, it is evident we both concur in viewing it rather as a deviation from health or nature than a cir- cumstance worthy of the consideration it has been accounted of by Mr. Clark. " Color. — Hoofs are black or white, or some intermediate shade, or they may ex- hibit a black and white striped or marbly aspect. It is an old observation, and one that passes current among us at the present day, that black or dark-shaded hoofs pos- sess greater strength and durability, and indicate less proneness in the feet to dis- ease, than such as are composed of white or striped horn. The rationale of which appears to be, that white horn (the same as white hair) is the product of parts weaker by nature than such as produce dark or black horn, and, being weaker, consequently are more liable to disease, less able to resist those impressions that tend to disorder. White hoofs are more porous than black ones, and consequently absorb moisture and lose it again by evaporation with more fa- cility : a fact tlfet may probably aid us in accounting for the failures attributed to them. '■'■Magnitude. — It requires no veterinary skill to discover any very material dispro- portion in the magnitude of the foot : it wUl strike us at once as being large or small, in comparison to the limb or the size of the animal. A foot of any description that is out of proportion is to the horse possessing it more or less objectionable : but, for all that, these out-of-proportion feet, abstractedly considered, have their advan- tages as well a& their disadvantages. Sain- bel tells us, that a large wide hoof, by ex- tending the surface of tread, ' will increase the stability and firmness of the fabric ; ' but then, he adds, ' this partial advantage grows into an evil when it becomes applied * Goodwin's New System of Shoeing, edit, second, page 33. to a body capable of translation, and con- sidered in a state of actual motion; be- cause, then, the mass and weight of the foot overburthen the muscles of the ex- tremity.' And because, I would add, the surfaces of contact being greater, the attrac- tion of cohesion becomes greater, and so much the more muscular force is required to raise the foot (particularly in moist ground) from the earth. Besides which, a large foot is apt to become objectionable from its striliing, during action, the opposite leg. On the other hand, it is contended, that a large foot will not sink so deep into soft ground as a small one, and conse- quently will not demand so great an effort of strength to draw it out. This is an argument, however, that can only hold good under the supposition, that in both cases the muscular strength is equal, which we know but rarely to happen, — in general, broad or flat-footed horses possessing supe- rior strength ; small, nan'ow-footed ones, superior speed. There cannot be a doubt about a large foot being unfavorable for speed, a small one for stability; neither one nor the other can be indiscriminately found fault with ; both within certain limits possess their respective advantages ; though to turn out as such, they each of them re- quire to be combined with suitable confor- mation and action. " Large bulky hoofs are found to be mechanically weaker than others, in conse- quence of being composed of a thin, soft, porous description of horn. Sainbel ascribes all this to ' a relaxation of the fibres com- posing the hoof: in which case, the diame- ters of the vessels are increased, the porosi- ties are multiplied, and the fluids abound in them in too great quantities ; conse- quently this kind of foot is soft, tender, and sensible.' Small feet, on the contrary, in general possess a close -woven horn, thick in substance, and consequently prove strong: they are rather oval than circular in figure, with great depth of substance, and are found to be of a durable nature. ' In feet of this description,' says Sainbel, ' from the too close union and too close tension of 20 ANATOMY AND PHYSIOLOGY OP their fibres, the vessels destined to conduct the nutritious fluid are contracted and obliterated ; whence proceeds that dryness of the part which renders the horn brittle and liable to split.' * " Division. — To the common observer the hoof appears to consist of one entire or indivisible case ; but the anatomist finds, by subjecting it to maceration, or coction, or even to putrefaction, that it resolves itself into three separate pieces : still, so long as the hoof maintains its integrity, such is the force of cohesion existing be- tween these three parts, that we as easily rend it in any other place as dissever one of its jointures. These constituent parts are the wall, the sole, and the frog: THE AVALL. " The wall or crust is the part of the hoof which is visible while the foot stands upon the ground. It forms a circular boundary wall or fence inclosing the inter- nal structures. On taking up the foot, we find the wall prominent all round beyond the other parts, making the first impression upon the ground, and evidently taking the largest share of bearing. It is the part to which the shoe is nailed. It is, in fact, the most important division of the hoof; ap- pearing to form (in the words of Mr. Clark) 'the basis or first principle in the mechan- ism of the hoof, the other parts being all subordinate to this.' " Situations and Relations. — The wall takes its beginning at the coronet, from the terminating circular border of the skin, with which it is intimately united ; their line of union being concealed by a row of overhanging hairs. From the coronet the wall descends in an oblique direction to the bottom of the foot, where it em- braces the sole, and terminates in a cir- cular projecting border. The anterior and lateral parts of the hoof are formed entirely by the wall ; but at the posterior part, in- stead of the heels of the wall being con- tinued one into the other so as to complete the circle, they become inflected, first down- * Sainbel's Lectures on the Elements of Farriery. ward, afterwards forward and inward, and are elongated in the latter direction untU they reach the centre of the bottom of the foot, where they terminate: these inflec- tions or processes of the wall constitute the bars. Altogether, the wall may be said to form about two-thirds of the entire hoof. " Connection. — Superiorly, around the . coronet, the waU is united with the sldin; interiorly, within its circumferent border, with the sole ; posteriorly, between its heels, with the heels of the frog; interiorly, be- tween the bars, with the sides of the frog ; and internally, with the sensitive laminEB. Let us now consider the waU in its detached or separate state. " Figure. — That of a hollow cylinder, having the sides presented to the ground cut much aslant, and whose circle exhibits a hiatus or deficiency behind, from the latetEil boundaries of which issue two narrow pro- cesses or appendages. Taking a lateral view, the wall assumes a conical shape, being broad and deep in front, and gradually narrowing as it stretches backward. '■^Division. — For facility of reference, and in aid of our descriptions, we distinguish in the wall. First, the toe; secondly, the quarters; thirdly, the heels; fourthly, the superior or coronary bordevg fifthly, the in- ferior or solar border ; sixthly, the lamincB or lamellcB; lastly, the bars or appendages. " The Toe forms the bow or front of the hoof, and comprehends about two-thirds of the superfices of the waU. It is the deep- est, broadest, and thickest part of the wall ; for reasons that wUl appear hereafter. It exhibits a degree of slant about equal, nat- urally, to an angle of forty-five degrees; though there are variations from this which (as was explained before) will be found, in a measiure, to be dependent upon the oblique truncation of the cylinder. When we come to understand the physiology of this part, however, a more operative and efficient cause for this variation wUl be found in the weight the wall has to sustain, and in its own mechanical strength or force of resist- ance : on which principle it is that light horses, thorough-breds, and ponies, as well EXPLANATION OF FIGURE III. MUSCULAR STRUCTURE. o". Trapezius. 6. Rhomboideus longus. c". Scalenus; and Z. Splenius. e". Pectoralis par\-us. y". Antea spinatus. g". Postea spinatus. a and D. Serratus magnus. 6'. Intercostales. c'. ObUquus externus abdominis. d'. ObUquus internus abdominis. e'. Erector coccygis. f. Depressor coccygis. g': Compressor coccygis. K. i'. Gluteii muscles. J'. Triceps abductors. K.. Biceps abductor femoris. m\ Tensor vaginae. n'. Rectus. o'. Vastus externus. r". s'. Gastrocnemius externus, and plantaris. »'. Flexor pedis accessorius ; its fleshy belly. y\ Peroneus. x\ Extensor pedis. 7»". Teres major. i". Latissimus dorsi. J. Pectoralis magnus. K. Humero cubital. (Pectoral region.) V. Scapulo-ulnaris. m". n". Triceps extensor brachii. o". Pectoralis transversalis. p". Flexor metacarpi, externus. u. V. Levator humeri, and Sterno maxillaris. s". Extensor metacarpi magnus. x". Extensor pedis. At the inferior part of the abdomen the letter h. occurs; it should be 4. 4 The subcutaneous thoracic vein A., hdwever, serves to indicate the region of the rectus abdominalis. 7. The sheath. *. Ligamentum colli. THE HORSE. 21 belonging to the inner part, which is the beginning of the wall itself; the external edge to the white band by which the other is embraced, and to which Mr. Clark has in particular drawn our attention, under the appellation of Coronary Frog-band. This covers the proper or veritable coronary bor- der of the hoof; having, tlnrough its fibres, which are very fine, a sort of dove-tailed connection with it. As it recedes backward, it grows broader to that degree, that its breadth at length becomes doubled ; being about half an inch broad in front, and one inch behind. It is thickest around its mid- dle parts ; its inferior edge, like the superior, becoming attenuated, until it grows so fine as to end in imperceptible union with the substance of the wall, giving it its beauti- fully polished surface : from the heat, how- ever, to which the hoof is artificially exposed, the thin part below the coronet often grows arid, splits from the crust, and becomes everted ; turning, at the same time, in con- sequence of dryness, of a whitish complex- ion. Posteriorly, we find it continued round the heels of the wall and frog, and from thence across the back of the cleft, forming altogether a complete circle, and everywhere showing itself to be the medium of connec- tion between the skin and the hoof. It has been already stated that the cutis terminates in a circular border, let into a groove around the summit of the wall : the cuticle, how- ever, does not end here — it is continued down; in fact, we trace it to the horny band we have been describing, the one being continuous in substance with the other. In- deed, the only detectible diflferences in them are, that one is thicker than the other, and grows hard, and dry, and white, from the effects of heat upon it from without, and the want of moisture from within. This cuticular origin and assimilation may be demonstrated in the putrefied foot ; or, bet- ter still, in the foot of the foetus. The band is broader at the heels than elsewhere, in con- sequence of the greater breadth of exposed cutis at those parts. In its texture it is fibrous, and its fibres pursue the same direc- tion as those of the wall, from which they differ only in being of a finer texture. Mr. Bracy Clark appears to entertain some sin- gular notions in regard to the structure, but more particularly the uses, of this part; which, in the respect I bear for their author, I shall consider, when the time may arrive for me to treat of the physiology of the foot. " The Inferior or Solar Border offers but little worthy of observation. It constitutes the ground or wearing surface of the wall, and is the part to wliich we nail the shoe. It grows thicker and more exuberant around the toe than in other places, and, from its pro- jecting beyond the sole, presents a conven- ient and suitable hold for the naUs of the shoe. Around the anterior and lateral parts, it embraces the sole ; behind, it joins the bars, which two points of union form two principal bearing places for the shoe. The inferior border possesses a larger circumfer- ence than the superior, in consequence of the oblique detruncation of the hoof. " This is a part that requires paring down every time the horse is shod. Such is its exuberating nature, that (like the human nail), were it not continually kept worn down, or broken, or cut off, it would elon- gate very considerably, and gradually turn up, exliibiting forms not only of the most unsightly but even grotesque description, and proving incommodious to a degree to be almost entirely destructive of progres- sion. " The LamincB (better named lamellts) consist of numerous narrow^ thin plates or processes, arranged with the nicest order and mathematical precision upon the inter- nal surface of the wall. They extend, in uniform parallels, in a perpendicular direc- tion from the lower edge of the superior border down to the line of junction of the wall with the sole ; and are so thickly set that no part of the superfices remains un- occupied by them. They are likewise con- tinued upon the surfaces of the bars. In the recent subject they are found soft, yield- ing, and elastic; but from exposure they become dry and rigid. " Every lamellEe exhibits two edges and 22 ANATOMY AND PHYSIOLOGY OF as mules and asses, have vpright or strong- feet (i. e.), walls but moderately sloped; whereas heavy horses, cart-horses, and coach-horses, have commonly flat or weak feet (i. e.), walls that slant immoderately. And (as was before observed) upon the degree of obliquity of the wall must very much depend that of the pasterns. In esti- mating the slant or slope of the wall, it is proper to distinguish between that which is consequent on the detruncation of the hoof, and such as is the effect of a biu-then under which the wall succumbs. The depth of horn in front of the toe, measuring from the termination of the sldn to the most promi- nent point below (and supposing the hoof to be cut and ready to receive the shoe), may be rated at about three and a half inches. The bow or degree of convexity of the toe in front must depend upon its obliquity as well as upon the circularity of the foot. The thickness of the horn com- posing the toe may be estimated at three- eighths of an inch, or from that to half an inch, and this substance is the same from immediately beneath the coronary circle to the junction of the wall with the sole ; at which part there is an accession of horny matter to block up the interstices between the laminffi, and also to fill the angular vacuity that would otherwise exist here between the wall and sole. In the fore-feet, the toe is thicker in substance than either the quarters or heels ; but (we have it from Sainbel) ' in the hind, on the contrary, the heels and quarters are generally thicker than the toe.' " The Quarters are the portions of the wall intermediate between the toe and the heels. They are commonly described as standing vpright, and, according to a car- penter's square set against the wall, so they appear to do ; this is not, however, the view the anatomist ought "to take of their posi- tion : to him the oblique course of their component fibres, together with the slant of their laminae, demonstrate that they slope in the same manner and degree as the toe does. The quarters do not run in straight lines from before backward, but by their prominence describe gentle curves, the outer making a wider sweep than the inner. This gives the hoof altogether a sort of twisted appearance, and makes the inner part of the toe look more projecting than the outer; a deviation that seems principally to have originated in the spread, and one, methinks, that has had more attention given it than any consequences attachable to it render it deserving of. The quarters range in depth from two to three inches; and measure in thickness from one-fourth to three-eighths of an inch. " The Heels are the two protuberant por- tions of the wall by which it is terminated posteriorly. They are the shallowest, and thinnest, and {in connection) only flexible parts of the wall. Though their surfaces recede from the perpendicular, they main- tain the same slope as the toe and quarters. At their angles of inflection, from which are continued the bars, they form (in con- junction with the heels of the sole) pouches or sockets into which are received the heels of the sensitive foot. In depth they range from one and a half to two inches. In sub- stance they do not exceed a quarter of an inch, the outer heel being rather thicker than the inner. " The Superior or Coronary Border is the circular, attenuated, concavo-convex part entering into the composition of the coronet. Its extent is marked exteriorly by the whitish aspect it exhibits, and also by some partial separation and eversion of the outer flakes of horn around its junction with the wall below. Externally, it assumes the same character as the wall below it ; but its in- ternal surface is altogether different. In- stead of possessing laminae, the surface is smooth and uniformly excavated, being moulded to the form of the sensitive coro- net, and everywhere presenting numerous pores for the purpose of receiving the secret- ing villi. Superiorly, the coronary border presents two edges, having a groove be- tween them for the reception of the termi- nating border of the cutis. It is this groove that marks the reception of the coronary border into two parts: the internal edge THE HORSE. 23 two surfaces. By one edge it grows to the wall ; the other, which is somewhat attenu- ated, hangs loose and floating within the cavity of the hoof. The surfaces, which are two lateral, are smooth, and, considering the magnitude of the lamella itself, of enor- mous extent ; so much so that it might be said almost to be constituted entirely of superficies. And this leads us to the con- templation of the great and magniiicent design which Nature evidently had in view in their formation, viz., the production of ample surface mthin a small space, an end that has been obtained tlirough the means of multiplication. Mr. Bracy Clark pro- cured fi'om the late Thos. Evans, L.L. D., mathematical teacher of Christ's Hos- pital, a calculation of what their united superficies amounted to ; and it appeared to aflbrd an increase of actual surface more than the simple internal area of the hoof would give of about twelve times, or about 212 square inches, or nearly one square foot and a half. " The lamellae exhibit no differences but in their dimensions. In length they corres- pond to the respective depths of the wall ; being longest, and likewise broadest, around the toe, and gradually decreasing towards the liinder parts. " In composition they are horny. Viewed through a microscope, ]\Ir. Clark discovered in their substance two planes of fibres, ' the one running in parallel lines to the axis of the hoof, the other obliquely intersecting these.' When stretched, they exhibit signs of elasticity ; but this appears greater in the transverse than in the perpendicular direction. " By means of its lamellae, the wall presents a superficies of extraordinary am- plitude for the attachment of the cofiin- bone. A structure consisting of similarly formed lamellae envelops the bone, and these are dovetailed in such a manner wdth the horny lamellae, as to complete a union which, for concentrated strength, combinin elasticity, may vie with any piece of animal mechanism at present known to us. " TJie Bars are processes of the wall, in fleeted from its heels obliquely across the bottom of the foot. For a long time, by farriers, they were confounded with the sub- stance of the sole, an error that owed its origin and perpetuation to the malpractice they exercised in paring the foot — in cut- ting both bars and sole down, without any distinction, to a common level. In the natural healthy foot the bars appear, exter- nally, as elongated sharpened prominences, extending from the bases of the heels into the centre of the foot, between the sole and the frog ; posteriorly, they are continuous in substance with the wall, with which they form acute angles ; anteriorly, they stretch as far as the point of the frog, constituting two inner walls or lateral fences between that body and the sole. Sainbel conceives, from their position, that they offer resistance to the conti'action of the heels. Their in- ternal surfaces exhibit rows of lamellEe, continued from those lining the wall, but which are here shoi't, and in their direction transverse, two circumstances referable to the narrowness and inflection of the bar. Towards the extremity of the bar they gradually grow shorter, and less distinctly marked, until we at length lose aU vestige of any more of them. While the promi- nence of the bars is such as to give them a secondary bearing upon the ground, their sharpened forms wiU sinlt them more or less deeply into every impressible surface. " THE SOLE. " The sole is the arched plate entering into the formation (as its name implies) of the bottom of the hoof: or, to adopt Sain- bel's definition, ' it is that part which covers the whole inferior surface of the foot, ex- cepting the frog.' It is a very just practical observation of Mr. Coleman's, that although a knowledge of every part of the foot is in- dispensably necessary to render us scientific overseers of the farrier's art, no individual part requires such undivided attention, as regards shoeing, as the sole, since the suc- cess of this mechanical operation mainly depends on the paring and defence of this arched horny plate. 24 ANATOMY AND PHYSIOLOGY OP " Situation and Connection. — It fills up the interspaces between the outer and inner walls (or bars) of the crust. I differ in opinion from those who describe it to sur- round the toe of the frog. I hold its circum- ferent support and connection to be the wall of the hoof, to which it is firmly cemented by an interstitial horny matter, filling the crevices between the laminEe. " Figure. — The circumferent outline of the sole meastires about two-thirds of a cir- cle, the remaining third being omitted to form a triangular-shaped hiatus or opening for the reception of the frog and bars. This circular form, however, is by no means true, or even invariably the same, in its dimen- sions. Generally, the longitudinal exceeds the transverse diameter. Its greatest diam- eter is shown by a line extended from either heel across its middle to the opposite point of the toe. " Arch. — Commonly, the sole presents an arch of more or less concavity inferiorly, and convexity superiorly. But it is not a regular or uniform arch, being one that rather waves or undulates, so as to bear a comparison, made of it by Mr. Clark, ' to the mouth of a bell extremely extended or flattened.' Like that of the bell, the arch is highest in the middle, from which it slopes, laterally, down to a flat, subsequently to rise again around its border, in order to present a dilated surface for attachment towards the wall. There is, however, vast variety in the degree of arch of the sole : in some feet it is of surprising depth; in others, the arch is converted into a flattened svirface ; and yet both seem to perform equally well. In the hind feet the sole is generally more arched than in the fore, and approaches in figure nearer to the oval than the circle. " Division. — In the sole we distinguish an anterior part or toe ; a middle or central part ; two points or heels ; and two surfaces. These divisions are not very well defined : but they prove serviceable in aid of our de- scriptions. The toe of the sole is the part encircled by the toe of the wall, against which it abuts, and to which it is intimately united by horny matter, the two together forming a stout bulwark of defence to those parts of the internal foot included between them. The points or heels are the two pos- terior salient angles received into the angu- lar intervals between the outer and inner walls or bars. Although naturally the least exposed, these are the parts most sub- ject to injury or pressure from the shoe, being the seat of that disease mistakenly called corn. The middle or centre of the sole is the portion more immediately sur- rounding the fore parts of the frog, and would (were the sole a regular arch) be the most elevated part ; but, in general, we find the sole flattened hereabouts ; the highest parts of the arch being the angles alongside of the bars; the lowermost, those around the toe. " Surfaces. — Of the surfaces, the supe- rior (as was mentioned before) is unevenly convex ; the inferior, correspondently con- cave. The former is everywhere pitted, particularly about the heels, with numerous circular pores, running in an oblique direc- tion, the marks of which remain evident upon the inferior smrface likewise. These pores are the impressions made in the soft horn by the villi of the sensitive sole, from whose orifices the horny matter is produced. They also form the bond of union between the horny and the sensitive soles : wliich is of a nature so strong and resisting, that it requires the whole strength of a man's arm to effect their separation — an operation of a cruel description that was wont to be practised in times past, under th& fallacious notion that ' drawing the sole ' was extir- pating the malady. " Thickness. — The natural thickness of the sole may be estimated at about one- sLxth of an inch. There will be found, however, variations from this standard in different horses ; and it will also very much depend on the part selected for measure- ment. The portion of the sole most ele- vated from the ground — that which forms a union with the bars — is nearly double the thickness of the central or circumferent parts ; and next to this, in substance, comes THE HORSE. 25 the heel. I do not find that the sole ' grows thinner from the circumference to the centre,' as has been stated by an author of celebrity. 'THE FROG. " The frog is the prominent, triangular, spongy body, occupying the chasm left by the inflection of the bars. " Situation and Connection. — The frog is fitted into the interval between the bars ; the three, altogether, filling up the vacuity in the sole, and thereby completing the circle, and estabhshing the solidungulous character of the foot. The frog extends forward, towards the toe, about two-thirds of the longitudinal diameter of the ground- surface of the hoof, terminating a little be- yond the central point (or what would be the central point) of the sole — or rather shooting directly through it, so as to anni- hilate the spot. Posteriorly, it is embraced by the heels of the wall; laterally, it pos- sesses firm and solid junctions with the bars, and through their medium with the sole : and these unions are effected not by simple apposition and cohesion of surface, but by a lamellated structure, apparent on the sides both of the frog and bars, by which the parts are reciprocally dovetailed into each other. LameUse are discoverable upon its sides, even all round the toe of the frog ; and this is a circumstance that confirms me in my belief that the bars reach thus far. " Figure. — The fi-og may be called pyra- midal, or cuneiform, or triangular in figure; its outline forming the geometrical figure denominated an isosceles triangle. I know of no comparison so familiarly apt as that of resembling it to a ploughshare : not only do they both correspond, as near as such comparisons can be expected to do, in out- line and make, but they likewise exhibit a singular coincidence in function ; the frog, like the ploughshare, being intended by its point to plough or divide the surface of the earth, and in that manner serve as a stay or stop to the foot. " Division. — We distinguish in the frog two surfaces, an inferior and a superior; two sides ; a point or toe ; and two bulbs or heels. '■'■Surfaces. — Both surfaces of the frog manifest striking irregularities, and these are respectively reversed, making one sur- face the exact counterpart of the other. In other respects, the only difference they ex- liibit, is, that the superior exceeds the in- ferior both in length and breadth. " The inferior sm-face presents to our view a remarkable cavity, broad, deep, and triangular in its shape, bounded on the sides by two sloping prominences, which divaricate from the convexity forming the toe of the frog, and terminate, after a short divergent course, at the heels. This cavity or hollow is denominated " The Cleft of the Frog: with seeming reference to the relationship existing, through its presence, between the horse's foot and the cloven one of the ox, deer, sheep, etc. In consequence of its sides sloping inward, the cleft at bottom gapes wide open ; but along the top is roofed by a simple linear mark running from before backward. The horn is kept continually soft and pliant within the cleft by a pecu- liar secretion from the sensitive parts it covers, the odor of which is notorious. " The solid wedge-like portion of horn in front of the cleft, extending from it to the point of the toe, has been observed by Mr. Clark to exhibit, in the natural foot at its fuU growth, ' a considerable bulbous en- largement,' which, by way of distinction, he caUs the cushion of the frog. On making a perpendicular section of the foot, Mr. C. finds tliis part is situated ' nearly opposite or under the navicular bone.' And it would appear (according to tliis author) that this 'rotundity, or swell of the frog,' is never reproduced after it has once been annihi- lated by the knife of the smith. " The superior surface of the frog, every- where continuous, uniform, and porous, b^ng the counterpart in form of the infe- rior, presents us with nothing but reverses : where the one is hoUow or depressed the other rises into swells and eminences, and vice versa. ■ This accounts for our finding 26 ANATOMY AND PHYSIOLOGY OP the part opposite to the cleft elevated into a conspicuous eminence, bounded on its sides by two deep channels, and a hollow of broader but shallower dimensions in the front. To this central conical elevation Mr. Clark has given the name of frog-stay, from some novel notions he entertains of its physiology. Such a bold promontory of horn rising in the middle of broad and deep channels is well calculated to form that dovetailed sort of connection with the sensitive foot, which greatly augments their surfaces of apposition, and establishes their union beyond all risk or possibility of dislo- cation. It is a part which (as far as my observations on it have extended) grows and becomes developed together with other parts of the foot ; and one that is apt to vary in its relative volume in different feet. In front of the frog-stay, the lateral borders, bounding the hollow in the middle, describe a waving line, which, near half-way to the point of the toe, exhibits a dip or impres- sion : this marks the impression of the navicular bone, and is the part immediately opposite to the ' cushion of the frog,' — a coincidence important to be borne in mind, as tending to throw some light on the na- ture of this new-christened structure.* " The Sides are the parts by which the frog establishes its union with the borders of the triangular vacuity in the hoof into which it is admitted. Along their superior borders they are transversely lamellated, or rather indentated, in order that they may be fitted to the internal surfaces of the bars, which exhibit a similar structure. " The Commissures are the two deep triangular-shaped hollows between the bars and the sides of the frog. It being only the superior borders of these parts that are engaged in their union, their broad, unat- tached parts, below, form the boundary walls of the commissures. Looking into the interior of the hoof, we discover that the commissures, internally, are converted into rounded promontories, similar in ap- * In fact, the cushion of the frog appears to be nothing more than a bulge of the part produced by the superin- cumbent pressure of the navicular bone. pearance and texture to the one in the mid- dle — the frog-stay — on the sides of which they are rising. In the natural state, the commissures must unavoidably get plugged with dnt, or whatever the animal may happen to tread upon ; a circumstance from which some far-fetched notions have been extracted concerning their use. " The Toe or point of the frog is the ante- rior, undivided, elongated portion; that which forms the apex of the pyramid or wedge — the acute or extended angle of the triangle — the only part displaying that prominent or rounded form that would warrant us in using the epithet ' conical ' to the frog. It possesses solidity of substance, firmness of texture, and luxuriance of growth in an eminent degree ; facts well known to the farrier, who, in paring the foot, seldom fails to make more free with this than any other part of the frog. " The Heels or bulbs of the frog are the posterior protuberant parts embraced by the heels of the wall, and separated from each other by the cleft, forming, together, the base of the wedge or triangle. They pre- sent greater depth of substance than the toe, but are of a softer, more spongy tex- ture, and are less resisting and stable, in consequence of being deprived of mutual support by the division of the cleft. Ante- riorly, the heels unite with the lateral promi- nences bounding the cleft ; interiorly, they present two surfaces of tread to the ground, eivdently designed to take a share in the bearing of the foot ; posteriorly and supe- riorly, they exhibit a bulbous fulness, in consequence of receiving at this part a sup- plementary covering from a production which has been (in the description of the wall) adverted to, under the appellation given it by Mr. Clark, of " Coronary Frog-band. — It was there stated, that the coronary groove (the groove or canal in the coronary border of the cutis) broadened considerably as it descended to and turned round upon the heels ; in like manner does the horny band produced by it broaden, and not only grow broader but thicker in substance, and consequently in THE HORSE. 27 the same degree augments the substance of the heels, occasioning that swell of them which has suggested the appellation ' bulb.' The horny band itself is everywhere lamel- lated upon its internal surface ; but these broadened parts of it display lamellae of a much bolder character, and consequently render their union with the heels so much the more intimate and enduring. The in- ferior edge of the band is denticulated, and the denticulations become so interlaced with the lamellated fibres of the wall, that their union is rendered, in the ordinary state of the hoof, altogether imperceptible. For drawing our attention to this part-, we are indebted to Mr. Clark ; and, insomuch as he considers it to be a production of the cutis (not having any connection with the glandular circle that secretes the wall), and to serve the purpose of ' uniting the sensible parts with the insensible,' I agree with him. I find something very similar to this grow- ing upon the human nail, issuing from the superior edge of the terminating border of the cutis, and continued from the cuticle, which proceeds for some way upon the naU, uniting it more closely and firmly with the cutis, and protecting the latter from exter- nal injury. This production is no more the beginning of the nail itself than is the so- called /rog'-band the commencement of the wall : they are both distinct parts, though but supplementary ones, and seem to be of a nature partaking both of horn and cuti- cle. It has no more important relation to the frog, in my opinion, than it has to the wall : it serves the same piirpose to both, — that of strapping up the heels of the frog and binding them in closer and more intimate connection with the neighboring parts. Were I asked what other use it ap- peared to have, I should say, that it was formed to cover and protect from injury the new-formed horn of the hoof, guarding it in its passage downward, until it has acquired substance and hardness sufficient to resist external impressions of itself. "development of the hoof. " During the early months of fcEtality, no horn or hoof is to be found. The foot is covered with a substance, white, firm, and elastic, resembling cartilage in its appear- ance, but proving more of the nature of cuticle on examination, which supplies the place of hoof. At the coronet this substance takes its origin from the cutis, being found to be continuous with the cuticle ; but that which covers the bottom of the foot is a production from the sensitive sole and frog. Altogether, it possesses the general form and appearance of the hoof, differing how- ever in these particulars — that the sub- stitute for the wall is comparatively thin in its substance ; while that which grows from the bottom of the foot is enormously thick, and, instead of being shaped into sole and frog, exuberates to a degree to constitute club-footedness. About the same period at which the pastern and coffin-bones take on ossification, horn makes its appearance underneath this cuticular wall, in the form of plates descending from the coronet, ex- liibiting with peculiar distinctness the lamel- lated structure. The horny wall becomes considerably advanced before we perceive any change in the bottom of the foot. At length, horn is detected forming underneath the cuticular substance, which, increasing in thickness, gradually represents sole and frog. Not, however, in an undeveloped state ; for even at birth these parts are yet concealed by the exuberant cuticular covering, now become loose in its textiue, and shaggy and ragged, in consequence of not receiving any further supply from the parts that produced it, and of being near its decadence ; for it not long after falls off, disclosing sole and frog both ready formed. "structure of the hoof. " Horn is found to differ in its texture or quality, not only in the many animals in which it is met with, but in different parts, and even in the same part of the body of the same animal. That which composes the hoof of the horse is a remarkable ex- ample of this. How different is the horn of the frog from the horn of the wall ; and yet neither of them agree in texture with 28 ANATOMY AND PHYSIOLOGY OF the sole. The horny substance of the wall is resolvable into fibres, bearing a resem- blance to thick or coarse hairs, wliich in the entire hoof are so intimately matted and glued together, as to have the appearance and strength of solidity. By close and ac- curate inspection these fibres may be seen descending in parallel lines, taking the obliquity of the wall, from the coronet to the inferior or solar border ; they do not run promiscuously, but are arranged in rows, forming sorts of beds or strata, lying one upon another — a disposition made manifest in the foot of the fcetus. A clean-cut trans- verse section of the wall exhibits upon its surface numerous minute, circular, whitish spots, which grow larger and more distinct towards the internal part, and through a glass appear to be hollow or tubular. These spots I take to be produced by sec- tions of the horny tubes, apparently contain- ing a whitish matter, a sort of pith, or pulp, or gelatinous instillation which pervades them from then- origin from the villi of the coronary circle ; the same as hairs derive their unctuous matter from the bulbs pro- ducing them, and (as this matter does the hair) renders the horny fibre tough and elastic — in fact, imbues it with the peculiar attributes so well known to smiths by the appellation of living- horn ; the epithet " living " being here used to denote the ob- vious differences the hoof of a living animal evinces from one that has been long detached from the body, or that is dead. We are too apt to believe that the various agents known to act upon the dead hoof or horn must take similar effect on the living ; and upon this erroneous belief we employ hot and cold water, etc., etc., in treating disease of the feet, forgetting that we have opposed to our remedies the resisting or self-preserving properties of living horn. " The sole, as well as the wall, is fibrous in its structure ; but its fibres appear to be of a finer quality, and, in course, are very much shorter : they, however, take an ob- lique direction, from behind forwards, fol- lowing the same degree of slope as those of the wall. They issue from the viUi penetrating the superior surface. To the fineness of its fibres, combined with the rel- ative magnitude of the tubular canals, and consequent proportions of horny and gela- tinous substances, may be ascribed the comparative softness and elasticity of the sole. " The frog, however, displays these qualities in such a remarkable degree as to appear, in fact, to be composed of quite another kind of horn ; though, on examina- tion, we find it to evince the same fibrous structure, the only perceivable differences being the comparative fineness of the fibres and their proportionably greater tubularity : their direction is oblique, correspondent with those of the wall. " PRODUCTION OF THE HOOF. " The wall is produced by the coronary substance, a sensitive and glandular part we shall have occasion soon to examine. Its villi, by some peculiar, mysterious, secretory process, convert the blood circulating through them into a soft pulpy gelatinous matter, which by exposure becomes hard horn, descending from the villous point that produced it, in the form of a tubular fibre, down to the sole. The fibres are united together at their very origin, but their tubes or canals diminish, the lower they descend; which accounts for the porous or honey- comb-like structure of the interior of the coronary border and the comparative solidity of the parts below. The outer layers or strata of fibres are found to be more com- pact and of closer texture than the inner; which arises, in part, from the viUi produc- ing them being removed to a greater dis- tance, and to the comparative smaUness of their canals, and which, consequently, the sooner become obliterated. The use of Mr. Clark's coronary frog-band becomes now more apparent, serving, as it evidently does, to cover and protect these external fibres until they grow sufficiently firm and soUd of themselves to bear exposure and resist casualties. " The sensitive laminas make no addition to the substance or tliickness of the wall : THE HORSE. 29 they simply produce the horny lamella arran!?ecl along its interior; as one proof of which, the wall measures as nmch in thickness at the place where it quits the coronet as it does at any point lower down. Other demonstrations of this fact come every day before such practitioners as have to treat canker, quittor, sandcrack, and other diseases of the feet. " The horny sole is a production from the villi of the sensitive sole ; after the same process as that by which the horny frog is secreted from the villi of the sensitive frog. " In a state of health of the foot, the se- cretion of horn is unceasingly going on. Disease or injury of the glandular parts may diminish or altogether suspend the process ; disease, under certain other forms, appears also to have the effect of increasing it ; but whether we have any artificial means of effecting this, seems questionable. The wall grows from above downwards. If a mark be made in any part of the wall, it will remain until it grows down and be- comes cut off below, at the inferior border ; and by observations made on the gradual descent and disappearance of these marks, calculations may be formed of the period of time required for the renewal or restora^ tion of the wall. " PROPERTIES OF HORN. " Horn is a tough, flexible, elastic sub stance, consisting of tubular fibres, more or less intimately connected together, takin the direction from the surface of the body on which it grows. Its property of tough- ness or resistance much depends on its con- dition in regard to moisture ; for if it is exposed to a degree of heat sufficient to abstract much of its natural juice or imbibed moisture, it loses its flexibility and tough- ness, and becomes brittle. On the other hand, saturated with moisture, it is con- verted into a soft and highly flexible sub- stance, but at the same time becomes weak and unresisting. This known eflect aids us to account for the flat-footedness of horses reared in low, fenny, or marshy situations ; the hoof being constantly in a state of saturation with moisture, the wall and sole yield to the superincumbent burthen of the body, and the latter grows flat (instead of remaining concave or arched), and even in some instances bulges. If oily or unctuous applications have any effect in softening the hoof, they appear to do so by filling the crevices and interstices between the fibres on the surface, and in this manner checking or suppressing evaporation. Horn takes a high and beautiful polish. Although much inferior in transparency to tortoise-shell, it may be worked up to bear so near a resem- blance to it as to be often, in manufactures, substituted for it, as in combs, etc. The hoof admits of an elegant polish ; and in that altered and improved state has been manufactured into articles no less useful than valuable and ornamental : * even the hoofs of the living animal may, by being kept clean, and when dry rubbed with lin- seed oil, be numbered among the ornamen- tal beauties Nature has bestowed upon quadrupeds. " By chemical analysis horn has been found to consist of membranous substance, having the properties of coagulated albumen, and of some gelatine. The horns of some animals, the deer species, from containing bone, become exceptions to this. Mr. Hatchett burnt five hundi-ed grains of ox's horn, and the residuum proved only one and a half grain, not half of which was phosphate of lime. "Shavings of hoof thrown into nitric acid become soft, and speedily melt into a yellow mass, which in about eight hours disappear in complete solution. " The same thrown into sulphuric acid turn black, in becoming soft, and require thrice the time for their solution. Muriatic acid also turns horn black, and corrodes it, but has so little effect towards its solu- tion, that after ten days a piece of hoof soaked in it was found to have become only more brittle or rotten. Common vinegar will turn horn dark-colored, but does not * The Eclipse hoof, presented by his Majesty at Ascot Races, as the reward of the best horse on the turf, forms a notable illustration of this. 30 ANATOMY AND PHYSIOLOGY OF appear to have any power in impairing its texture, or, at least, in dissolving it. Liquor potassae will not only tiu-n it black, but will corrode the horn of the hoof. Ammonia does not change its color, but slowly destroys its texture, rendering it brittle and rotten. "INTERNAL PARTS OF THE FOOT. " The internal, sensitive, organic parts of the foot, comprise the bones, ligaments, ten- dons, coronary substance, cartilages, sensitive lamina, sensitive sole, and sensitive frog. " The bones entering into the composition of the foot are the coffin and navicular bones : to which may be added (as forming part of the coffin-joint, and consequently having intimate relation to them), the coro- net bone. " The tendons immediately connected with the foot are those of the extensor pedis and the flexor pedis perforans : the former being inserted into the coronal process ; the latter into the posterior concavity of the coffin-bone. "the coronary substance. " A less inappropriate name for the part commonly called the coronary ligament.* " To revert, for the sake of elucidation here, to former description — after the hoof has been detached by a process of macera- tion or putrefaction, in a perfectly entire, uninjured condition, it presents around its summit a circular gi-oove, bounded in front by a soft whitish substance, having a thin edge, and being of a nature between horn and cuticle ; and behind, by an attenuated margin, more horny in its character, whose thin edging is denticulated or serrated. Into this circular groove or canal is received the terminating margin of the cutis : the cuti- culo-horny layer of the hoof, in front of it, having every appearance of being a continu- ation of the cuticle. " Situation — Dimension. — The coronary * Averse as I am to changing or altering names, noth- ing less than a palpable contradiction, in regard both to structure and function, would have induced me to do so in the present instance. substance occupies the concavity formed upon the inside of the superior or coronary border of the wall of the hoof: it is the part constituting the basis of the circular prominence commonly distinguished in the living animal as the coronet. It is broadest around the toe of the wall, diminishing in breadth towards the quarters and heels, and being somewhat broader around the outer than the inner side. It is thickest in sub- stance around its middle and most promi- nent parts, gro%ving gradually thinner both above and below. " Connection. — Externally, the coronary substance is connected with the hoof; and the connection appears to be principally, if not entirely, of a vascular nature : the sur- face of the waU presenting a porous honey- comb-like texture, and the villi or vessels issuing from the coronary substance enter- ing the pores, and thus establishing an inti- mate and extensive vascular union between these organic and inorganic parts. Inter- nally, the coronary substance is connected with the coffin-bone, the extensor tendon, and the cartilages, by a fine, dense, copious cellular tissue, which at the same time forms a bed for the assemblage and ramification of the blood-vessels concerned in the secretion of the wall of the hoof. Superiorly, its union with the skin is so intimate and com-- plete, that one has been thought to be a continuation of the other ; and, so far as meets the eye of a common observer, they might be taken as such ; but, when we come to examine them by anatomical tests, we not only find a line of external demar- cation between them, but discover such difference of internal structure as forbids the adoption of this delusive notion. As it descends upon the coffin-bone, the coronary substance not only grows thinner, but in growing attenuated becomes imperceptibly gathered or puckered into numerous points, from which issue a like number of plaits or folds, which afterwards form the sensitive laminae. It is worthy of remark, that the part of the bone upon which this transfor- mation takes place is smaller in circumfer- ence than the coronet; consequently the EXPLANATION OF FIGURE IV. NO. 1. — OSSEOUS STRUCTURE. OFF-HIND ESTREMTTT. 22. Femur or thigh bone. , 23. Patella. 24. Tibia. 25. Os calcis. 26. Astragalus. 27. One of the tarsal bones. 28. Metatarsus magnum. 29. The sessamoids. 30. Os sufeaginis. 31. Os corona. 32. Os pedis. e. The fibula. The above explanation wUl answer for " Xo. 3," of this plate. NO. 2. — MUSCULAR STRUCTURE. SIDE VIEW OF THE OFF-HIND EXTREMITY. j. Triceps. n. Rectus. o'. Vastus. r. s'. Gastrocnemi, and perforans. V. i). Flexor pedis accessorius. u'. (At the hock.) The insertion of the gastrocnemius into the point of the hock. x'. Extensor pedis. y. y'. Peroneus. V. (Beneath the pastern.) Flexor perforatus and perforans. z. Bifiu-cation of the suspensory ligament. if. The hoof. 8. One tendon of the suspensory ligaments. NO. 4. MUSCULAR STRUCTURE. ANTERIOR \TEW OF THE OFF HIND EXTREMITY. n. Rectus. , o'. Vastus extemus. J. Triceps abductor tibialis. y. y\ Peroneus. x'. x'. Extensor pedis. g. Flexor pedis accessorius. 8. Bifurcation of the suspensory ligament. 5. Saphena vein. S,-. The hoof. THE HOESE. 31 same measure of coronary substance which but tensely and smoothly covered the latter, admitted of being disposed in gathers or folds so soon as it reached the former. Pos- teriorly, the coronary substance' forms a junction, indeed becomes continuous in substance, with the heels of the sensitive frog. " Structure. — The coronary substance discloses three different parts in its com- position : 1. A fibro-cartilaguious circling band, forming the substratum and basis of the entire structure. 2. A cuticular cover- ing, so called from its resemblance in tex- ture to the cutis. 3. A network of blood- vessels, reposing upon the former, and covered by the latter. The cartilaginous structure, freed from its vascular connec- tions, is found to be \\Tought in the form of a coarse, open, irregiUar network, and appears designed mainly for the purpose of aflbrding a bed for the lodgment and rami- fication of the blood-vessels destined to pror duce the wall. The looseness of its con- nection, added to its own elasticity, renders this substance peculiarly adapted to accom- modate itself to the motions of the coffin- joint, and thus preventing those movements from operating prejudicially to the super- imposed glandular sti-uctm'e. " Organization. — The coronary sub- stance may be ranked among the most vas- cular parts of the body : no gland even possesses, for its magnitude, a greater abun- dance of blood-vessels, or of blood-vessels (taking them generally) of larger size ; nor does there exist any part in wliich greater care appears to have been taken to arrange its vessels so as to insure an uninterrupted supply of blood. These vessels it is that produce the wall : and there is every reason to believe that they perform this office without any assistance from the vessels of the laminas. " THE CARTILAGES " Are tvvo broad, scabrous, concavo-con- vex, cartilaginous plates, erected upon the sides and wings of the coffin-bone. Pro- fessor Coleman calls them ' the lateral car- tilages,' in contradistinction to two others he has named ' the inferior cartilages.' " Situation. — The cartilages form the postero-lateral parts of the sensitive foot, extending the siurface considerably in both these directions. " Attachment. — The cartilages are fixed into fossse excavated in the supero-lateral borders of the coffin-bone. Their anterior parts become united, on each side, with descending lateral expansions from the ex- tensor tendon, and are also attached to the coronet bone by cellular membrane. Their posterior parts surmount the aim or wings of the bone, to which they are firmly fixed, and from which they project backwards, beyond the bone, giving form and substance to the heel. Supposing one of the carti- lages to be divided into two equal parts by a line drawn horizontally across its middle, the superior half, which extends as high as the pastern-joint, is covered by skin only; and on that account is quite perceptible to the feel, and (in form) to the sight, as the animal stands with liis side towards us. The lower half is covered, superiorly, by the encfrcling coronary substance; inferiorly, by sensitive laminas : consequently, over all by the hoof, which envelopes both the coro- nary substance and the laminae. The ex- treme posterior ends of the cartilages inciu*- vate downward and backward ; but, being overreached by the heels of the sensitive frog, any abrupt or exposed termination of them is prevented. 'Around these points also the coronary substance makes its in- flections upon the sensitive frog, thereby giving them additional substance and sup- port. " Form. — Considered in the detached state, the cartilage in its general figure de- scribes an irregular quadrangle, of which the supero-anterior and infero-posterior an- gles are the most projecting ; the latter at the same time being incurvated inwards. Externally, the cartilage is pretty regularly convex ; internally, it is unevenly concave, the surrounding border turning inwards into the substance of the sensitive frog. The posterior part of the cartilage is somewhat 32 ANATOMY AND PHYSIOLOGY OF thinner than the anterior, and has several foramina through it — three or four of large size — which transmit vessels to the frog. " The False Cartilages. — From the in- ferior and posterior sides of the true carti- lages, proceed in a direction forward — towards the heels of the coffin-bone — two fibro-cartUaginous productions, to which IVIr. Coleman has given the name of ' infe- rior cartilages.' K they are to be consid- ered as cartilages at all, I prefer denomi- nating them false ; they being, as well in structure as in use, different from the true or lateral cartilages. They spread inwards upon the surface of the tendo perforans ; become united at their inner sides with the superior margin of the sensitive fi-og ; are covered inferiorly by the sensitive sole ; and at the same time assist in the support of the sensitive frog. They are triangular in their figure, and are arched in the same manner as the sole. " Use. — Their use appears to me to be, to fill up the triangular vacant spaces left bet^veen the tendo perforans and heels of the coffin-bone, thereby completing the sur- face of support for the sensitive frog, and extending that for the expansion of the sensitive sole. Bone in these places must have proved inconvenient by more or less impeding the impression upon, and con- sequent reaction of, the sensitive frog. "the sensitive lamix^ or lamella. " So is denominated the laminated, mem- branous, vascular structure clothing the wall of the coffin-bone. " Production. — The sensitive laminae appear to be derived from the coronary sub- stance — the one, in fact, seems to be a con- tinuation from the other ; for if, in a foot in a putrid condition, we attempt to part them by force, we may make an artfficial rent somewhere, but can find no natural separa- tion between them. The cnticular covering of the coronary substance having descended upon the coffin-bone, the circumference of which is less than that of the coronet, be- cause thereupon gathered into numerous little plaits or folds, which proceed in paral- lel slanting lines down the wall of the bone : a transformation it may be difficult to ex- plain, since the lamina; unfolded would occupy a much larger surface than the coronet ; at the same time, it is one that has its parallels in the animal constitution, and a remarkable one in the instance of the ciUary processes. " Division. — According to this mode of derivation, every lamina consists of one entire plait or duplication of substance, having its inward sides intimately and in- separably united; its outward sides being the surfaces of attachment for the homy laminas. It has also tw'o borders : one op- posed to the coffin-bone, the other to the hoof ; and two ends or extremities, one issu- ing out of the coronary substance, the other vanishing in the sensitive sole. " Structure. — The substance of the la- minse when held to the light evinces a degree of transparency ; although its nature is extremely dense, and it possesses extra- ordinary toughness and tenacity. Veteri- nary writers and lecturers have endowed the laminse with a high degree of elasticity : but it appears to me that the property is referable to their connections, and not one that is inherent in their own sub- stance. " Elastic Structure. — This is a substra- tum of a fibrous periosteum-fike texture, attaching the laminse to the coffin-bone, in which it is that the property of elasticity resides to that remarkable extent usually ascribed to the laminae themselves : indeed, so elastic is it found to be, that it can be made to stretch and recede the same as a piece of India rubber. Its fibres take a direction downward and backward. At the same time, it affords a commodious bed for the ramification of blood-vessels issuing from the substance of the bone, in which they are (particularly in the stretched con- dition of the substance) protected from in- jurious compression and consequent inter- ruption to their cfrculation. " Number. — In round numbers we may estimate the laminae at about 500 ; not in- THE HORSE. 33 eluding those of the bars. They vary, however, in number : I have reckoned up- wards of 600. ^'Dimensions. — In length they decrease from around the toe towards the sides and heels in a corresponding ratio with the wall ; those in front, the longest, being rather more than two inches in extent ; the shortest, those at the heels, being rather less than one inch. In breadth there is no vari- ation : all measure alike, one-tenth of an inch. " Organization. — The laminse are highly .-^ganized, though they are not equally so with either the sensitive sole or sensitive frog; nor are they so red as those parts: and the obvious explanation of this is, that (over and above what is requisite for their own nutrition) all the blood they have occasion for is only that which is suffi- cient for the secretion of the horny la- minse. "the sensitive sole. " The sensitive sole, or (as Sainbel calls it) the fleshy sole, is the fibro-vascular sub- stance covering the arched concave, or ground surface, of the coffin-bone ; in fact, is the part corresponding to the horny sole. " Structure. — The same land of elastic fibrous structiue that sustains the laminse is found constituting the groundwork of the sensitive sole ; only that in the latter case it is closer, denser, and firmer in its texture. Upon this is spread a remarkably beautiful venous network. And the whole is en- veloped in an outer cuticular covering, derived from the heels and frog, from which are sent villous processes, loaded with the points of arteries into the porosities of the horny sole : not, however, perpendicularly downward, but in an oblique direction — downward and forward — the same in which the horny fibres grow. " Connection. — Around the circumfer- ence of the coffin-bone, the sensitive sole is connected with the fibrous substance de- scending from the wall, together with the tapering, vanishing points of the laminae. In the cenfre, it is united %vith the bars and frog. But its principal attachment consists in its being firmly rooted into the sole of the coffin-bone ; a connection that receives considerable addition from the blood-vessels issuing out of the substance of the bone. " Thickness. — The sensitive sole varies in thickness at difterent places. On an average, it may be said to measure one- eighth of an inch in thickness. In the vi- cinity of the frog, it is something less than this. At the heels, it possesses double that thickness. " Organization. — This is one of the most vascular and sensitive parts in the body. Indepeiadently of the much ad- mired venous network expanded over the fibrous substance of the sole, arteries enter it issuing from the substance of the bone, and penetrate its villi, which, by talung this course, elude aU compression and obstruc- tion : there are also others — the nutrient arteries ; but these have an external origin, from the inferior coronary artery. The chief assemblage of arteries takes place within the villi, upon the cuticular surface — those issuing out of the interior of the bone simply passing through (without ram- ifying within) the fibrous substance : so that, if the substance of the sole is laid open by transverse section, the incised edge, near the surface, exhibits a deep red tint ; while the interior, nearer the bone, has a pinkish or pale red aspect. " THE SENSITIVE FEOG. " Under this head is included the cleft, cuneiform body, projecting from the bottom of the foot, together with the substance continued from it and filling the interval between the cartilages. Sainbel calls it ' the fleshy frog.' " Division. — We distinguish, in the sensitive as in the horny frog, an apex or toe ; two heels, separated by the cleft; and a portion intermediate between these, which is the bodij. " Situation and Connection. — The sensi- tive frog occupies the posterior and central parts of the bottom of the foot, forming in the tread a firm and secure point d'appin. 34 ANATOMY AND PHTSIOLOGT OF THE HORSE. Being in the hoofless foot equally promi- nent with the projecting edge of the coffin- bone, one might be led to infer that the horny frog should take the same line of bearing with the crust. The frog, alto- gether, is lodged in a capacious irregular space, bounded superiorly by the tendo- perforans and common skin, laterally by the cartilages, and inferiorly by the horny frog : with all which parts it has connections ; besides being continuous with the sensitive bars and sole, and at the heels with the coronary substance. On its sides are two shallow, ill-defined hollows, corresponding to the commissures of the horny frog, into which are received the horny prominences opposed to them. " Structure. — Entering into the com- position of this body we distinguish four parts : An exterior or cuticular covering ; a congeries or network of blood-vessels ; a fibro-cartHaginous texture ; and an elastic interstitial matter. " The exterior or cuficnlar covering in- vests the prominent bulbous portion of the frog, and also gives a lining to the cleft. Superiorly, it is continuous with the skin descending upon the heels ; anteriorly, with the cuticular covering of the coronet ; infe- riorly, with that of the sole. Numerous villous processes sprout from its surface, and enter the porosities in the interior of the horny frog, taking a direction down- ward and forward, the same as that in which the fibres of the horn grow. " The vascular covering succeeds the cuticular, lying immediately underneath it. It consists of a network of blood-vessels, principally veins, but which are not so thickly set as upon the sole. " The fibro-cartilagiiwus case comes next. We find it spread over those parts most subjected to pressure, and to be, in many places, one-fourth of an inch in thickness. From its interior are sent off numerous processes, pervading the elastic matter of the frog, forming so many septa intercross- ing one another, and dividing it without any notable regularity into many unequal c^S- partments. Li the posterior and bulbous parts, the septa exist in greater numbers, and are closer arranged than in the middle parts. The fibres of this vaginal substance run obliquely downward and forward, and become intermixed around the borders with those of the bars and sole. " The elastic interstitial matter, however, composes the bulk of the sensitive frog. It consists of a pale yellowish soft sub- stance, which has been mistaken for fat or oil, and hence has been named ' the fatty frog.' When cut deeply into, it exhibits a granulated appearance, and the fibrous in- tersecting chords become apparent, putting on the ramous arrangement of a shrub or tree. Altogether, the sensitive frog forms a peculiar, spongy, elastic body, for which we lack some more appropriate name." A TABULAR VIEW OF THE BONES OF THE HORSE. BOXES OF THE CRAXICM. Komber. Frontal 1 Parietal, 2 Temporal, two pairs, 4 Occipital, ...... 1 Ethmoid, 1 Sphenoid, 1 BOXES OF TILE FACE. Nasal 2 Superior and anterior maxiUar)', . . 4 Malar, . . ' . . ' . ' . . 2 LacrjTnal, 2 Palatine 2 Superior and inferior turbinated, . . 4 Vomer, ....... 1 Lower jaw 1 TEETH. Incisors, ....... 12 Canine, 4 Molars ' . . .24 BOXE OF THE TOXGUE. Os Hjoideus, 1 BOXES OF THE 'EAR. ISIalleus, 2 Incus, 2 Stapes, 2 Orbiculare 2 BOXES OF THE SPIXE. Cerncal, 7 Dorsal, 18 Lumbar (sometimes 6 are found), . . 5 BOXES OF THE SACRUM AXD TALL. Sacral, 1 Coccygeal (tail), about . . . .15 BOXES OF THE CHEST. Ribs, on each side 18, . . . . 36 Sternum, .... . . 1 PELVIS. Innominata (or bones without a name), . 2 BOX-ES OF THE SHOIXDEK. Scapular, . . .... 2 BOXES OF THE ARM. Humeral, 2 BOXTiS OF THE FORE-AEM. Radial and ulnar. The ulnar being, in the adults, connected mth the radius, we shall con- sider them as one bone. Radial, . . 2 BOXES OF THE KXEE. The carpal bones are thus named : ^ ["Scaphoid, | T Pisiform, i§ J Lunai', ^ J Trapezoid, £ 1 Cuneiform, g 1 Magnum, S \ Trapezium. ^ { Unciibrm. Eight bones to each knee, ... 16 BOXES BELOW THE KXEE. Metacarpal, 2 Splents, 4 Pastern, 2 Coronet, 2 Sessamoid, 4 Navicular, 2 Pedal or foot bones, 2 BOXXS OF THE HIXD EXTKEMITT. Femirr, 2 Stifle, 2 BOX-ES OF THE LEG. Tibia and fibula. These we shall consider as one to each extremity, .... 2 BOXES OF THE HOCK. Astragalus, 2 Os Calcis, 2 Cuboid, 2 Cuneiform, ...... 6 BOX-ES OF THE LEG. Two cannons and four splents, ... 6 BOXES BEX-E.\TH THE CAXXOX. Pastern, 2 Coronet, ....... 2 Sessamoids, 4 Na%-icular, 2 Pedal or foot bones, .... 2 Total number of bones, . . 238 The correct technical nomenclature of the above bones will be found in " Osteolog)-," which see. (35) ANATOMY OF THE SKELETON. -OSTEOLOGY. OSSEOUS SYSTEM OF THE HORSE. In the form of answers to a series of questions, the student will become ac- quainted with the name, location, form, use, and general peculiarities of the various bones composing the horse's skeleton. Q. "What is understood by the natural skeleton ? — A. The term is applied when the whole bones are held together by then- natural attachments : ligaments, cartilages, and synovial membranes. Q. Why is the term, artificial, sometimes applied to the skeleton ? — A. Because the bones, having been divested, by maceration or otherwise, of their connectmg ligaments, etc., are united artificially, by wire and plates of metal. BONES OF THE CRANIUM. Q. Enumerate the cranial bones. — A. Frontal, two parietal, occipital, foiu tempo- ral, ethmoid, sphenoid : ten. FRONTAL BONE (oS FRONTIS). Q. Describe the situation of the frontal bone. — A. It occupies the antero-superior part of the cranium in the region known as the forehead. Q. What are its peculiarities ? — A. In form it is irregular, having two surfaces and four borders. Its surfaces are flat externally, concave internally. Its internal surface is divided by a septum into anterior and posterior concavities. The posterior one is occupied by a portion of the anterior lobe of the cerebrum ; the anterior consti- tutes the frontal sinuses, they being sepa- rated from each other by the nasal spine. The concavity is further divided into shal- low chambers by imperfect septa. Q. Describe the borders of the os frontis. — A. They are denticulated and squamous. The posterior is arched, describing segments of two circles. The anterior or nasal is waving, inchnes backwards and outwards. The frontal border is straight, anteriorly broad and triangular. The ethmoidal or outer border is irregular, and unites with the lachrymal, sphenoid, and ethmoid bones. PARIETAL BONES (OSSA PARIETALa). Supposing the horse to be an adult, we shall consider these bones as one. Q. What is the situation of the parietal bone ? — A. It occupies the mesio-superior part of the cranium. Q. Describe the same. — A. Its form is quadrilateral : vaulted, concave internally, and convex externally. It has two sur- faces and four borders, denticulated and squamous. Q. What is observable on the convex sur- face? — A. A longitudinal messian crest, bifurcating anteriorly ; which indicates the location of the sutm-es, now obliterated by age. Between the bifurcatures arises an eminence above the cranial surface. Q. Describe the appearance of the inter- nal surface ? — A. It is indented by, and receives, the lobular eminences of the cere- brum, and it is also furrowed by arterial ramifications which supply the dura mater. TEMPORAL BONES (oSSA TEMPORUm). Q. What portion of the cranium do the ossa temporum occupy ? — A. Its sides and base. Q. How do these bones differ from those in man ? — A. In man they are divided into three portions, squamous, petrous, mastoid ; yet in reality they are united. In the horse (36) ANATOMY AND r PHYSIOLOGY OP- THE .HORSE. 37 they constitute four distinct bones, two on each side. Q. Name them. — A. Two ossa tempo- nun, pars squamosa, pars petrosa : four. Q. Describe their appearance. — One pair is composed of laminae, vaulted ; form ovoid, siumomited by irregular projections ; the other pair are solid and convex. OCCIPITAL BONE (oS OCCIPITEs). Q. What is the situation of the os occi- pites? — A. It is located in the postero- superior and inferior parts of the cranium. Q. What is its form ? — A. Convex externally, irregular, having an occipital tuberosity and condyles. Q. What are the connections of this bone ? — A. It unites, superiorly, with the parietal bones ; inferiorly and anteriorly with the sphenoid ; laterally, ■\\-itli the temporal, and it articulates posteriorly with the atlas. Q. What is the foetal state of the bone ? A. It is easily separable into four portions. Q. State its use. — A. It forms the pos- terior and inferior parts of the cranium, protects this portion of the braiji, and gives exit to the spinal cord. SPHENOID BONE (os SPHENOIDEs). Q. What is the situation of the os sphenoides ? — A. It passes from one tem- poral region to the other, across the antero- inferior part of the brain. Q. What are its general divisions ? — A. It is divided into body, situated in the mid- dle, alse or wings, on each side, and two pterygoid processes, considered as legs. Q. To what bones is it connected? — A. Occipital, ethmoid, squamous-temporal, pal- ate, and vomer. ETHMOID BONE (os ^THMOIDES). Q. What part of the cranium does the OS sethmoides occupy ? — A. Anterior to the sphenoid, and is the boundary of the cranial, and commencement of the nasal, cavities. Q. Describe its form. — .4. The posterior portion bears resemblance to a bird with its wings extended, having no legs, but a long erected neck and a small round head ; the anterior part consists of a slim, brittle, porous, spongy sti-ucture of considerable volume. Q. What are its connections? — A. With the sphenoid, frontal, vomer, and superior turbenated bones; and with the cartilaginous septum of the nose. BONES OF THE FACE. Under this head we shall consider the Ossa nasi, . . . ... 2 " raaxillaria superiora, . ... 2 " maxillaria anteriora, . ... 2 " malaa-um, . . ... 2 " lacrj-malia, . . ... 2 " palati, ... ... 2 " tiirbinati, superior et inferiora, . . 4 " vomer, ... ... 1 Os maxillarc inferus, (lower jaw,) . . 1 Total, IS We shall now consider these bones in the above order. NASAL BONES (oSSA NASi). Q. How many nasal bones are there ? — A. Two. Q. Where are they situated? — A. In the superior part of the face. Q. Describe their form? — A. They re- semble the form of a pear ; are broad pos- teriorly, pointed anteriorly ; they are convex externally and concave internally. Q. To what bones are they connected ? — A. To the frontal, superior and anterior maxiUaria, and laclirymal. Q. What is their use?— .4. To defend the nares, and retain in position the septuni nasi. SUPERIOR MAXILLARY BONES (oSSA MAXIL- LAKIA superiora). Q. Where are they situated? — A. In the supero-lateral parts of the face. Q. Describe their form ? — A. They are somewhat irregular — trilateral; from the centre (wliich is tliickest) they taper, the anterior part being much thinner than the posterior. Q. How are they divided? — A. Each 38 ANATOMY AND PHYSIOLOGY OF bone has a facial, palatine, and nasal sur- face. It has also nasal, alveolar, and pala- tine borders, and two extremities : posterior, which forms the maxillary tuberosity ; anterior, or dental extremity. Q. What are the connections of the ossa maxilaria? — A. With the squamous tem- poral, nasal, anterior maxillary, malar, lachrymal, palate, and inferior turbinated bones. ANTERIOR MAXILLARY BONES (oSSA MAXIL- LARIA ANTERIORa). Q. What is the situation of these bones? — A. They are placed in the supero-anterior and antero-lateral parts of the face. Q. What is the general form of tliese bones ? — A. Very irregular ; consisting of a broad, thick base, turned forwards, from which is sent off a thin flexible plate ; and a narrow, elongated, tapering portion turned backwards. Q. How is each bone divided ? — A. Into three surfaces and tliree borders. Q. Describe the surfaces. — A. The superior or nasal smface is smooth, con- vex, and oblong. The inferior or palatine is vaulted, it contributing to the formation of the palate ; within it, of an oval form, is the interdental space, which is occupied by two thin, flexible plates, the palatine pro- cesses, denticulating along the sides with each other. In the side of the bone is a deep hollow, for the reception of that por- tion of the superior maxillary bone which holds the tusk ; and tlie remainder of the surface, posteriorly, is articulated with the same. The anterior or labial surface is broad, smooth, and convex, and gives at- tachment to the depressor labii superioris, and gums. Q. Describe the borders. — A. The anterior border is broad and curved, and is composed of two laminse, formed apart and divided into septa for the insertion of six incisors. The posterior border is naiTow and sloped, and denticulates with the nasal bone. The internal border is broad, quadrilateral, curved, and denticulates with its fellow, forming thereby the superior maxillary symphysis, through which runs the foramen incisivum, for the transmission of the palatine arteries. Q. How is this bone connected? — A. It connects with the superior maxillary and nasal bones, and with its feUow. MALAR BONES (oSSA MALARUm). Q. What is the situation of the ossa malarum? — A. They occupy the antero- external part of the orbit. Q. Describe their form. — A. Irregularly triangular, presenting a broad basis forw^ards. Q. How is the bone divided ? — A. Into three surfaces, three angles, a basiform and an apiform extremity. Q. Name the surfaces. — A. Facial, maxiUarj-, and orbital. Q. Describe the same. — A. The facial surface is divided into two portions by the zygomatic spine ; the upper division is smooth and nearly flat ; the lower part is narrow and roughened, for the insertion of the masseter muscle. From this surface, posteriorly, arises the zygomatic process, which is very obliquely sloped off, and laminated for adaptation to the process of the same name, meeting it from the tem- poral bone, the tw^o together forming the zygomatic arch. The maxillary surface is concave. The orbital surface has a smooth concavity which forms the infero-external part of the orbit. Q. Describe the angles. — A. There are three, superior, inferior, and posterior. The superior constitutes the external portion of the orbital circumference. The inferior forms the zygomatic spine. The posterior is not so prominent nor defined, but forms an irregular link with the superior maxillary bone. Q. What of the extremities ? — A. The anterior extremity is broad, irregular, and denticulated, and articulates with the supe- rior maxillary and lachrymal bones. The posterior or apiform extremity forms the zygomatic process. Q. With what bones do the ossa mala- rum connect? — A. With the temporal, superior, maxillary, and lachrymal bones. THE HORSE. 39 LACHRYMAL BONES (OSSA LACHRYMALIa). There are two lachmyral bones: we shall describe but one, considering that they are both alike, as indeed are those already refer- red to, in a plm-al sense. Q. What is the situation of the lachry- mal bone ? — A. It occupies the antero- external part of the orbit. Q. How is it divided ? — A. Into three surfaces and five borders. Q. Name the surfaces. — A. Internal, external, and orbital. Q. Name the borders. — A. External and internal facial, nasal, and external and internal orbital. Q. What is observable in the orbital ex- cavation of this bone? — A. The lachrymal fossa. Q. What occupies this fossa or groove ? — A. The lachrymal vessels, sac, and duct. Q. With what bones is it connected ? — A. With the frontal, nasal, malar, and supe- rior maxillary bones. PALATE BONES (oSSA PALATi). Q. What is the situation of the palate bones ? — A. They are placed in the infe- rior posterior part of the face, adjoining the base of the cranium. Q. What does the palatine surface form ? — A. The palatine arch or roof of the mouth. Q. What of the nasal surface ? — A. It forms the posterior surface of the nasal outlet. Q. What other surfaces do these bones present ? — A. Ethmoidal and orbital. Q. To what part of the bone is the vel- lum palatei attached ? — A. To the palatine. Q. How are the palate bones united to the superior maxillary? — A. By their supero and infero lateral borders ; each being den- ticulated. Q. What other connections have the pal- ate bones? — A. They are joined to the frontal, ethmoid, sphenoid, vomer, and in- ferior turbinated bones. TURBINATED BONES (oSSA TURBINATA SUPE- RIORA ET INFERIORa). Q. Where are the ossa tmrbinata located ? — A. Within the nasal cavity : the superior above, and the inferior below. Q. What is their form, and how are they divided ? — A. In form they are oblong, thin, foliated, convoluted, scroll-like, and cavern- ous. They are divided into external and internal sm-faces ; superior and inferior ex- tremities. Q. How many bones are there? — A. Four. Q. Describe the bones. — A. Their ex- ternal surface is convex, and presents series of longitudinal grooves which mark the ramifications of small blood-vessels. The internal surface is cellular, being unequally divided by transverse septa. Their interior is capacious ; they are open superiorly and closed anteriorly. They are porous and elastic. The superior bone exceeds in volume the inferior, and makes its convolu- tion from below, its superior border being attached ; whereas, the reverse is the case with the inferior one. ' Q. What are their connections ? — A. The turbinated bone is connected above with the ethmoid ; and laterally, with the nasal bone. VOMER. Q. From what does the name of this bone arise ? — A. From its resemblance to a ploughshare. Q. What are its uses ? — To divide the nasal chambers and permit the expansion of olfactory nerves. Q. What is inserted into its superior groove? — A. The septum narium. Q. What are its connections ? — A. It unites with the ethmoid, sphenoid, superior and anterior maxiUary, and palate bones. LOWER JAW. INFERIOR MAXILLARY BONE (oS MAXILLARE INFERIUs). Q. What is the situation of tliis bone ? — A. It composes the inferior and posterior parts of the face. Q. What is the foetal state of the bone ? — A. In the fcetal state it is divided, at its inferior junction, by a connecting cartilage, hence the part has been called its symphysis. Q. How is the bone divided ? — A. Into 40 ANATOMY AND PHYSIOLOGY OF body, neck, sides, and branches ; external and internal surfaces, and corresponding borders. Q. What do you understand by these terms?- — A. Body signifies the anterior part reaching posterior to the tusks ; neck signifies the contracted jjart, immediately posterior to the body; sides are the parts comprehended between the neck and the branches ; the branches are the parts poste- rior to the neck, which terminate in the con- dyles. As regards surfaces, the external is convex, rounded, rough, and porous, and affords attachment for muscle and gum. The internal sxirface is concave, rough, and porous, and answers for the attachment of muscles and gum, and as a channel for the tongue, and attachment for the froenum lingua;. Q. "What do you understand by borders ? — A. Each superior border exhibits six alveolar cavities for the molar teeth; the septum are composed of osseous lamiiiEe. The inferior border is tliin and irregular. The posterior border is broad and roughened for the insertion of muscles. GENERAL INQUIRIES. It is now presumed that we understand the location and names of the different bones composing the cranium and face ; and, be- fore we proceed further, it may be profitable to make some general inquiries regarding the bony structure. Q. Is not the number of bones greater during colthood than at mature life ? — A. Yes, many of the bones separable at that period become united in the adult. Q. How are bones divided l^A. They are divided into long or cylindrical, broad or flat, and thick. Q. What do you understand by cpiphy- sisis of bones? — A. The region where car- tilage is interposed between bones that finally become ossified. Q. What is the structure of bones? — A. They consist of a cellular, reticular, and vascular parenchyma, and of osseous mat- ter deposited in it : their base, therefore, is the same as that of the soft parts. Q. Are bones vasciilar ? — A. Yes. Q. How can you demonstrate their vas- cularity ? — A. By numerous small fora- minas and by the tinge they receive from the coloring matter of the food. Q. Name the investing membrane of bones? — A. Periosteum. Q. What is its organization ? — A. Fi- brous. Q. Of what use is this periosteum? — A. It limits the growth of bones, is the medium of circulation and nutrition, and affords attachment for ligaments and mus- cles, and favors the free articulation of the latter. Q. What does its internal surface se- crete? — A. An oleaginous fluid, depos- ited in the cellular structure and cavity of bones. Q. What are foramina? — A. Holes perforating the substance of bones. Q. What are sinuses? — A. Occun-ing in bones, they are large cavities with small openings. Q, What are sinuosities? — A. Superfi- cial but broad irregular depressions. Q. What are furrows ? — A. Long, nar- row, and superficial canals. Q. What are notches ? — A. Cavities in the margin of bones. Q. What are fossae ? — A. Deep and large cavities on the surface of bones. Q. What are glenoid cavities? — A. Cavities for articulation. Q. What are tubercles? — A. Small em- inences. Q. What are tuberosities ? — A. Rough elevations. Q. What are spines? — A. Long pro- jections upon a bone. Q. What are heads? — A. The round tops of bones. Q. What are necks ? — The narrow por- tion of bones beneath their heads.. Q. What are processes? — A. Short pro- jecting portions of bones. THE TRUNK (REMARKS ON THE SAMe). We shall now consider the peculiarities of the trunk; which comprehends the verte- EXPLANATIONS OF FIGUEE V. MUSCULAR STRUCTUKE. LATERAL VIEW OF THE HEAD, NECK, AND SHOULDEK. — THE HEAD. a. Orbicularis palpebrarum. h. Levator palpebroe. c. Dilatator naris lateralis. d. " " anterior. e. Orbicularis oris. /• Nasalis longus. f Levator labii superioris. i. Zygomaticus. J- Retractor labii inferioris. k. Buccinator. K. Masseter. h: Temporalis. m. Attolentes et abducens aurem. /• Facial Teins. THE N*ECK. •. Ligamentum colli. 6". Rhomboideus longus. s. A portion of the splenius. e". Scalenus. e". Pectoralis parvus. 0. Abducens vel deprimens aurem. r. Tendon of the splenius and complexus major. t. Obhquus capitis inferiorus. u. Levator hiuneri. V. Sterno maxillaris. X. Subscapulo hyoideus. 3. Jugular vein. KEGION OF THE SHOULDEE. a". . Trapezius. f" '. Antea spinatus. ^'■ , Postea spinatus. h". . Teres major. OSSEOUS STRUCTURE. * Ligamenture coUi, or subflaviimi. 1. Temporal bone. 2. Parietal bone. 4. Zygomatic arch. 5. Nasal bone. 6. Lachry-mal bone. 7. Malar. 8. Superior maxilla. 9. Anterior maxilla. 10. Inferior maxilla. b. The neck of the same. 11. Cervical vertebree. 33. Scapula. 34. Humerus. c. The incisors. *. The lining membrane of the ear. THE HORSE. 41 bral chain, thorax, and pelvis. It is gener- ally called the spine, or back bone, and extends from the occipital bone to the sac- rum. The spine is divided into three regions, denominated cervical, dorsal, and lumbar. The spine, as a whole, exhibits three surfaces and two extremities. The surfaces are named superior, inferior, and lateral. The superior surface is flat in the region of the neck ; in the back and loins it offers a series of projections. The infe- rior surface is more uniform, and the lateral is very irregular. CERVICAL VERTEBRJ2. Q. How many cervical vertebrae are there ? — A. Seven. Q. What is the name of the first ? — A. It is called atlas. Q. How does it differ from the rest? — A. It has no superior spinous process nor body ; the vertebral hole is larger than in the others, and its transverse processes are very broad. It has three pairs of foraminas : one posteriorly, through which run the verte- bral arteries ; and two anteriorly. Q. What is the name of the second cer- \dcal vertebra ? — A. It is named dentata. Q. How is it recognized from the rest ? — A. By its anterior projection, which in the human subject resembles a tooth. Q. With what does this tooth-like pro- cess articulate ? — A. It articulates with the infero-posterior part of the ring of the atlas. Q. Describe the third, fourth, and ffth cervical vertebra;. — A. They possess the genuine characters of cervical vertebrm, and closely resemble each other; the third, how- ever, has commonly a more elevated supe- rior spine than either of the others, and is narrower across the mesio-siiperior part of the bodi/ (measuring from the roots of the articular processes), which dimension in- creases in the fourth, but is greatest in the fifth. Q. What of the sLxth vertebral — A. It has no inferior spine ; and its transverse pro- cesses are trifid, consisting each of three eminences. Q. What of the seventh l — A. It is the shortest, and in its general conformation re- sembles the first dorsal. Its body, pos- teriorly, presents two semilunar articular depressions, constituting a part of the socket for the first rib. DORSAL VERTEBRJE. Q. How many dorsal vertebrce are there? — A. Eighteen. Q. What is peculiar to the dorsal ver- tebras ? — A. They have each a bodi/, spinous process, and transverse process, and are generally distinguished by the length, form, and direction of their spines. Q. How is the fii-st dorsal vertebrce dis- tinguished from the rest? — A. By the sharpness of its spinous, and singleness of transverse, processes, and by the breadth of its articulatory sm-faces. Q. How do the articular depressions for the insertion of the ribs differ in each bone? — A. They are less deeply marked, as we proceed posteriorly. Q. How are the seventeenth and eight- eenth distinguished from the rest? — A. They have perfect articulatory depressions on the bodies for the insertion of ribs. LUMBAR VERTEBRJ3. Q. How many lumbar vertebrEB are there?— A Five. Q. How are the bodies of the lumbar vertebra distinguished from the dorsal? — A. They are larger, conti-acted in the centre, and their edges are more prominent. VERTEBRAL CANAL. Q. What is the form of the vertebral canal? — A. In the cervical region it is ca- pacious and semi-oval ; through the dorsal, transversely oval and smaller. In the lum- bar it is semi-circular, of less diameter than the cervical and greater than the dorsal. Q. With what does the spinal canal con- nect ? — A. Anteriorly, with the cranial cavity ; posteriorly, with the sacral canal. PELVIS, SACRUM, AND TAIL BONES. We shall now consider the posterior boundary of the trunk. 42 ANATOMY AND PHYSIOLOGY OP OS SACRUM. Q. What is the popular name of this bone ? — A. It is called the " rtimp bone." Q. Where is it located? — A At the superior part of the pelvis, between the ossa illia. Q. What is the popular name of the ossa illia ? — A. They are called the haunch bones. Q. How many pieces enter into the com- position of the sacral bone, in the foal? — A. It is composed of five pieces. Q. How are they uiaited ? — A. By fibro- cartilaginous substance. Q. What ultimate change takes place in tills substance? — A. It becomes ossified, and hence the solid bone. Q. How is the sacral bone divided ? — A Into three sm-faces, two borders, base, and apex. Q. Describe its surfaces. — A. They are named superior, inferior, and lateral. The superior is convex, very irregular; on its central line are five eminences, and laterally are superficial grooves pierced by the four sacral foraminae. The inferior surface is smooth and slightly concave. The lateral sm-face is thick anteriorly, gradually tapering posteriorly ; they are roughened for the re- ception of the sacro-iliac ligament. Q. Describe, briefly, the base and apex. — A. The base is composed of a central and two lateral parts. The apex is oval, and articulates with the anterior bones of the taU. BONES OF THE TAIL (oSSA COCCYGIs). Q. What is the situation of the ossa coccygis ? — A. Posterior to the sacrum. Q. Of how many bones is the taU com- posed? — A. Fifteen. PELVIS OR HAUNCH BONES (oSSA INNOMINATa). Q. How do anatomists divide these bones, in the fostal state? — A. Into ilium, ischium, and pubes. Q. Li the adult horse are there more than two bones ? — A. They are considered as two, yet in reality they are united at the pubes so as to constitute but one bone. In this state, however, they are denominated ossa innoniinata — unnamed bones. Q. What is the situation of the iliatic, ischiatic, and pubic portions? — A. They are in the anterior, superior, and lateral parts of the pelvic region. The ischiatic extends posteriorly and the pubic interiorly. Q. What are the connections of the ossa innominata ? — A. They are connected, anteriorly and inferiorly, to the os sacrum ; posteriorly and inferiorly, to each other, forming the symphysis pubis ; laterally, with the thigh bones. Q. What are the uses of the pelvis? — A. It affords an arch for supporting the posterior parts. It contains the urinary or- gans, rectum, etc., gives protection to blood- vessels and nerves, and origin and insertion to various muscles and ligaments. CHEST OR THORAX. The thorax or chest is formed by the dorsal vertebrae, superiorly ; ribs, laterally ; and sternum, inferiorly. It also affords pro- tection to the principal organs of circula- tion and respiration. Q. State the number of ribs and their arrangement? — A. Their number is gener- ally thirty-sbc ; eighteen on each side, eight of which are termed ti'ue, and the remainder false, ribs. Q. Why are the anterior eight called true ribs ? — A. Because they have a direct cartilaginous insertion into the breast bone or sternum. Q. Why are the posterior ten termed false ribs ? — A. Because they are indi- rectly connected with the sternum. Q. What is the general conformation of a rib ? — A. It is lengthy, curved ; con- vex outwardly or laterally ; terminating in a sharp border posteriorly, which forms a posterior convexity. On the inner surfaces it is concave, and of course the reverse of the external. Q. What are the variations in ribs ? — A. They vary in length, degree of curva- ture, and obliquity of direction. Q. How shall we divide each rib ? — A. THE HORSE. 43 Into a body, external convexity ; internal concavity, a superior and inferior termina- tion ; anterior and posterior edges. Q. What do you understand when the term, head, is applied to a rib ? — A. It signifies its protuberance — its superior portion ; presenting a smooth convexity for articulation with the bodies of vertabras. ' Q. Where is the neck of a rib situated 1 A. Immediately below the head. Q. What is the difTerence between the anterior and posterior edges of the ribs ? — A. The anterior edge is circular and the posterior is sharp. Q. Where is the tubercle of the rib situated ? — A. Posterior to the head ; at the root of the neck. Q. How is the first rib distinguished from the rest ? — A. It is the shortest and thickest, and is almost straight. Q. How does the second rib differ from the first? — A. It is longer, less dense, and has a greater curvature in the region of its neck. Q. How do the ribs differ from the second to the seventh ? — A. They increase in breadth. Q. How do they differ in length ? — A. Up to the ninth. Q. How do they differ in curvatinre ? — A. Gradually up to the eighteenth, which is the most curved of all. BREAST BONE (sTERNUm). Q. What is the situation of the sternum ? — A. It occupies the anterior and inferior portion of the thorax. Q. How does it differ from the human sternum ? — A. In the human subject it is composed of three pieces ; in the adult horse it is considered as a single bone. It is made up, however, of seven irregularly formed bones. Q. What is the structure of the ster- num ? — A. It is composed of an osseaus cellular substance and cartilages. Q. Name the cartilages ? — A. Ensi- form and cariniform. Q. What is the use of the cariniform cartilage ? — A. It affords attachment to the sterno-maxillares and sterno-thyro-hy oidei muscles. Q. To what part of the sternum is the ensiform cartilage inserted ? — A. To its in- ferior and posterior part. Having now considered the bones of the head (with the exception of the teeth), and hyoides (appendages), spine, thorax, and pelvis, we now commence on the bones composing the extremities. These are four in number, disposed in pairs, and known as the fore and hind extremities. Our examination will be conducted with refer- ence only to one fore, and one hind, ex- tremity ; presuming that a description of the bones on one side will suffice for those on the other. FORE EXTREMITIES. Q. What is the situation of the fore extremities ? — A. They occupy the antero- lateral parts of the trunk, from which they proceed inferiorly. Q. How are the bones divided? — A. Into shoulder, arm, knee, leg, pastern, coro- net, and foot. Q. Name the bones composing each region ? — A. SHOCXDER BONES. Scapijla, Humerus. AKM BOXES. Radius, Ulnar. BONES OF THE KNEE. ^ f Scaphoid, § J Lunar, « I Cuneiform, S i^Trapeziun. i ["Pisiform, « J Trapezoid, I 1 Os Magnum, S 1 Unciform. BONES OF THE LEG. Large Metacarpal. Two small Metacarpal (splents). IN THE REGION OF THE FETLOCK. Two Sessamoid Bones. PASTERN BON'S. Os Suf&aginis. COKON'ET BONE. Os Corona. FOO^ BON-ES. Navicular and Coffin-bones. 44 ANATOMY AND PHYSIOLOGY OF Q. How many bones compose one of the fore extremities ? — A. Twenty-one. OF THE SHOULDER. Q. The shoulder being composed of the scapula and humerus, what portion of the thorax do they occupy ? — A. They occupy its antero-lateral region. SCAPULA, (shoulder blade). Q. What is the position of the scap- ula ? — A. It occupies the antero-lateral parts of the thorax. Q. Describe the bone. — A. It is trian- gular, broad, and thin superiorly ; narrower and thicker inferiorly ; its external surface is unequally divided into two superficial concavities, named fossae antea et postea spinata;. Its internal surface is smooth, yet excavated. Q. Describe the borders. — A. The su- perior has a thin, roughened summit for the insertion of the cartilage of the scap- ula ; the anterior is thin in its upper half, yet below it becomes rounded ; the posterior is obtuse and rounded. Q. How does the scapula terminate in- feriorly ? — A. By a glenoid cavity. Q. What are the connections of the scapula ? — A. It has a ligamentous con- nection with the spines of some of the dorsal vertebras ; to the thorax, it is connected by muscular faschia; and its inferior connec- tion is by means of the glenoid cavity, to the head of the humerus ; this latter forms the shoulder joint. Q. Is there anything remarkable about the shoulder joint? — A. Its most remark- able feature is, the great disproportion in size between the head of the humerus and the glenoid cavity. Q. How is this disproportion in magni- tude compensated for ? — A. By an exten- sive capsular membrane, wliich admits of extensive motion. Q. What are the insertions of tliis cap- sular membrane ? — A. It has a circular insertion into the rough margin of the glenoid cavity, and also around the neck of the humerus. Q. How is this membrane protected in- ternally and externally ? — A. Internally it is clothed with a synovial membrane ; ex- ternally by adherent muscles. humerus. Q. What is the situation of the humerus ? — A. It is situated beneath the scapula, occupying a diverse direction, viz., downwards and backwards, and is in con- tiguity with the lateral parts of the thorax. Q. Describe the form of the humerus. — A. It is irregular, cylindroid, having a convoluted appearance, and its superior extremity is much larger than the inferior. Q. How is this bone divided? — A. Into a body, superior and inferior extremi- ties. Q. Describe the body. — A. It is angu- lar, with sides, contracted superiorly, and flattened and rounded inferiorly. From its superior-anterior-lateral margin projects a roughened tuberosity, into which the leva- tor humeri is inserted. The lateral part of the body is hollow or excavated. The in- ner side is somewhat roughened and promi- nent. Q. Describe the superior extremity. — A. The superior extremity being much larger than the inferior, presents a head and several tubercles : it has a projecting, hemispherical surface, designed for exten- sive articulation. It presents a smooth surface, yet has an irregular, indented groove for the insertion of a capsular liga- ment. Q. What is the use of the tubercles ? — A. The anterior, three in number, serve as articulations for the flexor brachii to tra- verse. The fourth serves as a protection against dislocation. Q. Describe the inferior extremity. — A. It consists of two heads or condyles, sepa- rated by deep ovoid fossa, into which is received the olecranon of the ulna. Q. What are the connections of the humerus ? — A. Superiorly, it connects with the scapula ; inferiorly, it articulates with the radial and ulnar extremity of the Os Brachii. THE HORSE. 45 OS BRACHII (arm BONe). Q. Describe the location of the os brachii. — A. It is located beneath the tho- rax, in the inferior region of the humerus. Q. How does this bone differ from those of the human subject ? — A. By being con- solidated into a single bone. Q. How is it divided ? — A. Into radial and ulnar portions. Q. Descrihe ihe radial. — A. It consists of a body, superior and inferior extremi- ties. The body is lengthy, compared with other bones of the fore extremity ; posteri- orly it is excavated and roughened ; ante- riorly it projects with a smooth, cylindrical surface. Q. Describe the superior extremity. — A. The superior extremity presents an in- terrupted articulatory surface, having a central eminence, with two cavities, which correspond to the articulations of the os humerii. Q. Describe the inferior exti'emity. — A. It appears to consist of three articulatory sturfaces, which correspond with those of the bones of the carpus. Q. Describe the ulnar portion of the os brachii. — A. It presents a tapering trian- gular projection, firmly connected with the radius; at its junction with the same, it presents a semilunar concavity ; this, with the articulatory surface of the radius, forms the humero-brachial articulation. Q. Name the projection of the ulnar, commonly termed point of the elbow. — A. Olecranon. Q. What muscle is inserted into the ole- cranon ? — A. The triceps extensor braehii. Q. What is the state of this bone in early colthood ? — A. It is composed of two pieces named radius and ulnar, wliicli after- wards become consolidated. Q. With what bones does the inferior portion of the os brachii articulate ? — A. With the scaphoid, lunar, and cuneiform bones. BONES OF THE KNEE (CARPUs). The bones of the knee correspond to the wrist, or carpus, of man. Q. How are these bones arranged ? — A. They are ranged in tvo rows, or tiers ; one of the number, trapezium, is located in the posterior part of the carpus. Q. Name the bones of the first row ? — A. Scaphoid, lunar, cuneiforme, trapezium. Q. Name the bones crossing the second row? — A. Pisiform, trapezoid, magnum, unciform. Q. What is the general form and situa- tion of each of the bones of the first row? — A. The OS scaphoides is semi-ovoid in form, its superior surface is sigmoid and smooth, the inferior surface is somewhat oval, and rests upon the trapezoides and magnum of the second row. Its internal surface comes in contact with the os lunare. The OS lunare is the second bone of the first row; it articulates superiorly with the brachii ; inferiorly, with the ossa magnum and unciforme ; its superior surface is trian- gular ; inferior, oblong ; on one side, inter- nally, it articulates with the scaphoid, on the other with the cuneiforme. The cunei- forme is known as the external, yet smallest bone of the knee. Its superior surface is concave ; inferior, smooth ; its internal sur- face articulates with the os lunare, and pos- teriorly it unites with the trapezium. Q. What is the general form and situa- tion of each of the bones of the second row ? — A. The os trapezoides is situated on the inner side of the knee, resting on the inner splent bone, and articulating with the OS magnum ; its form is that of an iiTegular, curvated, flattened cone ; its superior surface is convex, and its inferior flat. The os magnum is the middle bone of the second row, and is known as the largest bone of the knee. Its superior surface presents two articvilatory surfaces, one sigmoid and ob- long for the OS lunare, and the other ovoid and flat, to correspond with the surface of OS scaphoides ; its interior surface is flat, and articulates with large metacarpal bone. The OS unciform is situated on the outer side of the second row, and in form resem- bles a blunt hook; its superior surface is convex; its inferior irregular, articulating with the outer splent and cannon. The os 46 ANATOMY AND PHYSIOLOGY OP trapezium is situated in the posterior part of the carpus, and presents two smooth sur- faces for articulation with the ossa cunei- forme and brachii. Its external, lateral sur- face is convex; its internal concave; its superior border gives attachment to the flexores metacarpi ; and into the inferior is inserted a ligament. The ossa pisaforms — for sometimes there are two present — is situated posterior to the trapezoides; its form is orbicular or pea-shape. METACARPAL BONES. The metacarpal bones are three in num- ber, viz : metacarpus magnum, 1 ; metacar- pus parvum, 2. There seems, however, so great a disproportion between the os mag- num and ossa parva, that the former may be considered as the principal support of the fore extremities. Q. What is the situation of the metacar- pus ? — A. Immediately beneath the carpias. Q. Describe the form of the metacarpi magnum. — A. It is a long cylindrical bone, presenting on its anterior surface a circular, smooth appearance; its posterior surface is somewhat flattened and depressed. Q. How is the bone divided ? — A. Into a body, and two extremities. Q. Describe the extremities. — A. The superior presents a smooth articulatory sur- face, tapering towards its outer edges, yet more depressed on its inner and posterior part ; in the anterior region is a roughened prominence, for the insertion of the extensor metacarpi, and on the lateral side of the bone are eminences which afford insertion for the lateral ligaments. The inferior ex- tremity presents a pulley-like surface, with two unequal condyloid surfaces, separated by a semi-cLrcular eminence, which corre- sponds to a counterpart found on the supe- rior end of the suffraginis. Q. What are the articulations of the metacarpi magnum ? — A. It articulates su- periorly with the carpus ; interiorly, with the OS suffragmis ; posteriorly and laterally, with the ossa sepamoidea, and metacarpi parva. OSSA METACARPI PARVA (SPLENT BONEs). Q. How many bones compose the ossa metacarpi parva ? — A. Two : external and internal. Q. Describe their situation. — A. They are attached to the lateral and posterior parts of the metacarpi magnum. Q. How do you divide them ? — A. Into bases, middles, and apices. Q. Describe the base. — A. It is sur- mounted by a smooth articulatory surface, corresponding to the inferior portion of a part of the knee joint. Q. Describe the middle. — A. It is tri- facial: the anterior surface is roughened for the insertion of inter-articular tissue, which connects it with the cannon ; the inner surface is excavated; the outer surface is rounding, and terminates, posteriorly, acu- minately. Q. Describe the apex. — A. It tapers, and ends in a tubercle, which curvates in an inferior and superior direction. Q. How do the ossa metacarpi parva differ ? — A. The external is generally larger than the internal, and has a broader articu- latory surface. Q. What bone does the external splent articulate with ? — A. The unciform. Q. What bone does the internal splent articulate with ? — A. The trapezoid. Q. How are the splents connected to the cannon ? — A. By cartilago-ligamentous tis- sue. Q. What changes does this cartilaginous tissue undergo, subsequent to adult life ? — A. In a majority of cases it becomes ossified. PASTERN BONE (oS SUFFRAGINIs). Q. Describe the location of this bone. — A. It is located beneath the cannon, and takes an oblique direction from the same ; it articulates superiorly with the cannon; posteriorly with the ossa sesamoidea. Q. Describe the form of the os suffra- ginis. — A. It is a flattened cylinder, yet its superior portion is more bullvy than the in- ferior ; it is generally considered as being about one-third the length of the cannon, and is divided into a body, superior and inferior extremities. Q. Describe the body of the os suf&a- THE HORSE. 47 ginis. — A. The body presents two surfaces, anterior and posterior ; the anterior is con- vex, the posterior flattened and uneven ; it lessens in bulk in an inferior direction. Q. How is the superior extremity of the pastern bone recognized from the inferior ? — A. The superior is the largest, and pre- sents two shallow articular cavities ; between them is a groove, which receives the central eminence of the inferior extremity of the cannon bone. The inferior extremity is much smaller than the superior; it is bi- convex, and consists of two articular con- vexities, separated by a transverse shallow depression. SESSAIIOID BONES (OSSA SESSAM0ID.E). Q. Where are the two sessamoids situ- ated ? — A. At the posterior part of the articulation formed by the cannon and pastern bones. Q. What is the form of these bones ? — A. Trapezoid: three sides present triangular faces, whose apices unite in one point, which is directed upwards ; the bases of the same form a fourth side, which is turned down- wards ; and are therefore divided into three sides, base, and apex. Q. Give a general description of the faces or surfaces of these bones. — A. They are known as anterior, posterior, and lateral faces ; the anterior are excavated, smooth, and articulatory, and along their inward borders — which are opposed to each other — are levelled off, so that the two form a groove for the reception of the central eminence of the inferior portion of the can- non. The posterior surfaces are convex and rougli ; the lateral surfaces are grooved and roughened; the bases are narrow and uneven. Q. What appears to be the object in ex- cavating the anterior surfaces of these bones? — A. To extend the articulatory surface of the pastern joint, and admit of extensive anterior and posterior motion. Q. For what purposes are the posterior surfaces roughened ? — A. For the insertion of the suspensory Ligaments. Q. What occupies the cavity which occurs in consequence of uniting the inter- nal surfaces of these bones ? — A. The flexor tendons. Q. What ligaments are inserted into the bases of these bones ? — A. The long, short, and crucial ligaments. CORONET BONE (oS COEON^). The OS coronce is situated beneath, or rather inferiorly, to the pastern, and may therefore be termed the inferior, pastern; it occupies a location between the superior pastern and coffin bone. Q. Describe the os corona, or inferior pastern. — A. It presents a square body; its breadth, however, somewhat exceeds its longitudinal measurement. It has four sur- faces, viz., superior, inferior, anterior, and posterior ; the superior surface is bi-concave, corresponding to the projections of the superior pastern ; the inferior surface is bi- convex, consisting of two condyloid prom- inences, separated by a slight transverse depression, corresponding to the articulatory surface of the coffin-bone ; the anterior sur- face is convex, yet rough and irregular ; the posterior surface is quite smooth, yet exca- vated. Q. What are the conn? ctions of the os corouEe ? — A. It connects with the pastern, cofSn, and navicular bones. BONES OF THE FOOT. THE COFFIN BONE (oS PEDIs). The coffin bone is considered as the base of the osseous structure of the fore extremity. Q. What is the form of the coffin bone 1 — A. It presents a semilunar outline ; an- teriorly and superiorly it is convex ; pos- teriorly and inferiorly it is concave ; it is divided into wall, sole, tendinous surface, articulatory surface, and wings. Q. Describe the wall. — A. It is a miniature of the form of the hoof; it exhibits a porous and furrowed surface, and has in- numerable perforations, varying in size and form ; its superior part is surmounted by the coronal process; the inferior edge of the waH is somewhat oval, and is notched and serrated. 48 ANATOMY AND PHYSIOLOGY OP Q. What are the uses of the porosities and furrows ? — A. They serve as so many attachments for the fibrous tunic of the sensible laminae. Q. What name is generally applied to the largest of the perforations found in the coffin bone ? — A. They are termed for- ammsB. Q. What occupies these forarainae ? — A. Blood-vessels and nerves. Q. Describe the sole of the coffin bone ? — A. The sole exhibits a broad, uniform, concave surface, resembling in most cases the figure of the inferior part of the hoof; it has porosities similar to those formed on the wall ; it is bounded anteriorly and later- ally by the circumferent edge of the wall ; posteriorly, by a sharp, uneven, semi-circular edge, which divides it from the tendinous surface. Q. What do we find on the tendinous surface? — A. 1st, a rough depression in its fore and middle part, marking the inser- tion of the tendo perforans. 2ndly, two lateral grooves, passing obliquely inwards, and terminating each in a large foramen. 3rdly, a porous space intermediate between the two former divisions, into wliich is fixed the inferior navicular ligament. Q. What occupies the lateral grooves? — A. The trunks of the arteries and nerves which occupy the interior of the coffin bone. Q. What are the pecuharities of the ar- ticulatory surface of the coffin bone ? — A. It has two lateral depressions, which extend posteriorly to the alae ; a broad eminence runs transversely between them ; this emi- nence is terminated in front by the coronal processes, having an incurvation backwards ; behind it, the surface is bevelled off", to which part is opposed the navicular bone ; the depressions alluded to are deepened by the prominent edge running around the an- terior and lateral parts. Q. What portion of the coffin bone does the articulatory surface occupy? — A. The superior part. Q. What is the form of this surface ? — A. It is half-moon shaped. Q. Describe the alffi, or wings ? — They consist of a protuberance on the posterior part of each side of the coffin bone ; the protuberance, however, is generally bifid; the lower portion which is the largest, is irregular and asperous, and projects in a posterior direction ; the upper portion is tu- bercular, yet smooth ; between the divisions of the alae is a notch, which, at a certain period in the life of the animal becomes a perfect foramen. Q. What is attached to the irregular sur- face of the larger division of the alee ? — A. The cartilage of the foot. Q. What is affixed to the tuberculated portion of the als ? — A. The coffin liga- ments. Q. What vessel passes through the notch ? — A. The lateral artery. Q. What is there remarkable about the structure of the coffin bone ? — A. It has a spongy, fragile texture, pervaded in every direction by minute canals for the trans- mission of blood-vessels and nerves ; it dif- fers very essentially from many bones of the body, which, in healthy subjects, are remarkable for compactness and sofidity. NAVICULAR BONE (oS NAVICULARe). Q. What is the general form and division of the navicular bone ? — A. It is semi-lu- nar : its lunated border, however, only forms about one third the circle of its dimensions ; it is divided into two sm-faces, two borders, and two extremities. Q. Where is tliis bone situated? — A. At the posterior part of the coffin joint. Q. Describe the superior and inferior sur- faces of the navicular bone. — A. The superior surface bears a corresponding aspect to the articulating surface of the coffin bone, having two superficial lateral depressions, with an eminence betw^een them. The inferior surface is also articu- latory ; and exhibits lateral depressions yet more superficial than the superior; it has also an eminence across the middle, nar- rower yet more prominent than the former. Q. Name the tendon which articulates over the inferior surface. — A. Tendo per- forans. THE HOESE. 49 Q. Describe the borders. — A. The bor- ders are lunated and straight : the lunated is broadest in the centre, and narrows to- wards the extremities ; superiorly it has a smooth narrow strip of surface along the middle, which is adapted to the bevelled portion of the articulatory surface of the coffin bone ; the part beneath is fluted and porous, into which is inserted a ligament which connects it with the coffin bone. The straight border is thinner than the opposite one ; superiorly it is rough and porous ; inferiorly it is smooth and lipped. Q. What is the form and direction of the extremities of the navicular bone ? — A. They are obtusely pointed, one directed outward and the other inward. Q. What ligaments are inserted into the extremities ? — A. The lateral ligaments. HIND EXTREMITIES. Q. What is the situation of the hind extremities? — A. They occupy the inferior and posterior parts of the pelvis, and sup- port the posterior parts of the trunk. Q. How are the bones of the hind ex- tremities divided ? — A. They are thus divided : Femur, stifle, thigh, hock, leg, pas- tern, coronet, and foot. Q. Name the bones comprising these parts. — A. FEMUR. STIFLE BONE. Patella. THIGH BONES. Tibia and Fibula. HOCK BONES. Astragalus, Os calcis. Cuboid bone, Three Cuneiform bones. BONES OF THE LEG. Metatarsi Magnum. Metatarsi Parvieum, two bones (splents). PASTERN JODfT. Ossa SessamoidiE (two bones), Os Suffraginis (pastern). CORONET. Os Corona. BONES OF THE FOOT. Os Pedis and Os Navieulare. Q. Where is the femur situated ? — A. Between the pelvis and thigh bones. Q. How is it divided? — A. Into a body and two extremities. Q. What are the peculiarities of the superior extremity? — A. It consists of two parts : a hemispherical, smooth, articulatory head, directed upwards and inwards, and joined to the body by a flattened neck, and exhibiting on its inner side a fissure, into which is fixed the teres or round liga- ment. The other part is a large irregular projection at the base, and posterior to the same is a deep oval cavity ; at the superior part is a roughened crest; inwardly it pre- sents a concave, smooth surface. Q. What is the proper name of the pro- jection? — A. The great external tro- chanter. Q. What muscles are inserted into the the same ? — A. The gluteii. Q. What is inserted into the concave smooth surface? — A. The capsular liga- ment. Q. What is the form of the body of the OS femoris ? — A. It is cylindrical. Q. How does it correspond in size and weight with other bones of the body ? — A. It is the longest and weightiest. Q. What is the form of inferior extrem- ity ? — A. It is broad and thick, and has a trochleal prominence and two condyles. Q. Give a description of the same. — A. The articular or puUy-Uke surface anteriorly consists of a broad, semi-circular groove bounded on either side by a prominence ; the condyles much resemble each other, excepting that the external is the thickest, and the internal most projecting; they exhibit prominent, convex, articulatory sur- faces ; on their sides are rough eminences ; between them is a deep fossa ; at the base of the external condyle is a pit. Q. What articulates over the pulley-like surface ? — A. The patella or stifle bone. Q. What is inserted into the rough emi- nences ? — A. The lateral ligaments. Q. What occupies the fossa ? — A. The inter-articular ligament. 50 ANATOMY AND PHYSIOLOGY OF Q. What is inserted into the pit ? — A. The tendon of the extensor pedis. Q. What is the state of this bone during colthood ? — A. Exti-emities are attached to the body of the bone by means of cartUage. Q. What changes do the extremities undergo just prior to adult life ? — A. They become consolidated with the body of the bone. STIFLE BONE (paTELLa). Q. What is the situation of the patella ? — A. It is situated on the anterior and infe- rior extremity of the femur. Q. What is its general form ? — A. Quadrangular, convex externally, irregularly concave internally. Q. How is it divided ? — A. Into three surfaces and four angles. Q. Describe the surfaces. — A. The an- terior surface is convex, yet quite prominent in the centre ; it has a roughened surface, and is porous. The superior surface is angular, uneven, and roughened. The pos- terior surface is articulatory, and unequally divided by an eminence running across it into two shallow concavities, which are adapted to the condyles of the inferior ex- tremity of the femur. Q. Describe the form of the angles of the patella. — A. They are obtuse. Q. Why is the anterior surface of the bone roughened ? — A. For the insertion of tendinous and Ugamentary attachments. Q. What is implanted into the uneven and roughened part of the superior border ? — A. The tendons of the rectus and vasti muscles. Q. What is inserted into the inferior and lateral angle? — A. The Kgamentum pa- tella. Q. What are the connections of this bone ? — A. It is connected to the inferior portion of the femur by tendinous and capsular ligaments; to the tibia it is con- nected by similar ligaments. THIGH BOXES (tIBIA AND FIBULa). In consequence of a horse having a very- large femur, and that bone appearing to enter into the composition of the haunch, the tibia and fibula are termed thigh bones, although in man they are termed bones of the leg ; the fibula of the horse, however, is a very small, slender bone, affixed to the superior part of the external side of the tibia. Q. What is the situation of the thigh bone ? — A. It is situated between tiie stifle and hock. Q. What is the form of this bone ? — A. It is long, straight, prismatic ; its superior extremity is larger than the inferior. Q. What is its direction ? — A. Oblique in a contrary direction to the femur. Q. How is the tibia divided ? — A. Into a body, superior and inferior extremities. Q. What is the general form of the body ? — A. It is irregularly triangular, the posterior face is broadest, the anterior angle is rounded, and the sides are roughened. Q. What is peculiar to the superior ex- tremity of the bone ? — A. We find tn^o irregular ovoid articulatory surfaces, corre- sponding to the eminences on the inferior extremity of the femur ; these are separated by an acute elevation, and two fossa, into which is inserted the lateral ligament. Q. Describe the inferior extremity. — A. It is flattened, and has two deep articular grooves running in an anterior and posterior direction ; its exterior margin is roughened. Q. What are its connections ? — A. It connects with the femur and patella supe- riorly ; interiorly, with the bones of the hock. fibula. Q. What is the situation of the fibula? — ^4. At the posterior part of the tibia. Q. How is it connected to the tibia ? — A. By cartilago-ligamentous substance. Q. What is the form of the two ends of the bone ? — A. The superior is bulky, flat- tened from side to side, and roughened. The inferior is slender and tapering, and extends about half way down the tibia. BOXES OF THE HOCK (taRSUs). The tarsus, or liock, comprises a part of the osseous structure of the horse, that <<5 f^»%^ EXPLAJS^ATIONS OF FIGURE VI. NO. 1. — POKE EXTREMn'IES. f. The ulnar. 34. Humerus. 3q. Kadius. 36. Carpus. 37. Metacarpus. 38. Sessamoids. s. or 39. Os suf&aginis. 40. Os corona. 41. Os pedis. The above description also answers for No. 3, — the bony structure. NO. 2. — MUSCULiVR STRUCTURE. L.iTEIUL ■iTEW OF THE NE.1R-F0IIE EXTEEMITY. s". Extensor metacarpi magnus. H' . Humero cubital. n". Levator htunero. p". Flexor metacarpi estemus. i". x". Extensor pedis. jj". j<". v. Flexor tendons. v". Flexor tendons. z. Suspensory ligament. i{. The hoof. NO. 4. ANTEEIOR VIEW OF THE NEAX-FOEE EXTEEMITT, s. Extensor metacarpi magnus. t. Extensor metacarjri obHquus. a;". Extensor pedis. y". Extensor suffiraginis. ^•. The hoof. 8. Bifurcation of the suspensory Ugament^ THE HOBBE. H every veterinary student should aim to be well acquainted with ; it is a part that seems to be, in this country, more liable to anchy- losis and exostosis than any other region ; here is the seat of spavin, and no one can possibly understand the nature of such dis- ease unless he be conversant with the ana- tomical mechanism of the hock. The hock corresponds to the tarsus or instep of man, and is composed of six bones, viz., os calcis, astragalus, os cuboides, ossa cuneiformis ; which comprise three small bones, viz., ex- ternal, internal, and middle cuneiforme. We shall first consider the os calcis. Q. What is the situation of the os cal- cis ? — A. It forms the posterior projec- tion known as the point of the hock — ■ the superior and posterior bone of the tarsus. Q. Give a general description of the bone. — A. Its figm-e is irregular ; presents a body, tuberosity, posterior surface, and base ; the body is most bulky at its inferior part ; as a whole, it is irregularly convex ; concave and expanded at its base, where it presents four surfaces for articulation with the astragalus ; the tuberosity is ob- long, flattened on each side, and terminates in a rough tubercle, into which is inserted the tendons of the gastrocnemii. It is sit- uated on the superior part of the hock. THE KNUCKLE BONE (aSTRAGALUs). Q. What is the situation of the asti-a- galus ? — A. It is situated in the superior part of the hock, and is the principal sup- port of the tibia. Q. How do you distinguish it from other bones ? — A. It is readily distin- guished by its double pulley-like articula- tory surfaces, which consist of two semi- circular prominences, having between them a deep groove, well adapted to receive the projection found on the inferior extremity of the tibia. Q. What is the appearance of the pos- terior surface ? — A. It has four articulatory surfaces, corresponding to those of the os calcis. Q. What is the appearance of the base or inferior extremity ? — A. It has an irre- gularly flattened articulatory surface, which comes in contact with the large cuneiform bone. CUBOID BONE (oS CUBOIDES). Q. What is the situation of the cuboid bone ? — A. On the outer part of the hock. Q. How is the bone divided ? — A. Into four surfaces, viz., external, internal, supe- rior, and inferior. Q. How do you distinguish the external from the internal surface ? — A. The exter- nal siu-face is broad, kregular, curved, and roughened ; on the other hand, the internal is excavated, and has three articulatory surfaces. Q. How does the superior surface difler from the inferior ? — A. The superior sur- face has two articulations, with a fossa between them ; the inferior surfaces are smaller, and correspond, one to the articula- tory head of the splent bone, and the other to the cannon. large cuneiform bone (os cuneiforme magnum). Q. What is the situation of the cunei- form bone ? — A. Directly beneath the astragalus. Q. What is the appearance of this bone ? — A. It presents a triangular form; its acvite termination being in a posterior du-ec- tion, it has superior and inferior sm'faces, sides, and angles. Q. How is the superior surface distin- guished from the inferior l — A. The supe- rior surface has a uniform articulatory surface, with the exception of a small, rough grove running to its centre, from the outer side, which terminates in a central pit. The inferior surface is rather convex, yet presenting a flat appearance ; its poste- rior an^le has an articulatory surface, cor- responding to that of the cuboid bone. Q. What are the articulations of tliis bone ? — A. It articulates with the astra- galus, cuboid, middle and small cunei- form bones. 52 ANATOMY AND PHYSIOLOGY OF middle cuneiform bone (os cuneiforme medium). Q. What is the situation of the middle cuneiform bone ? — A. It is situated be- neath the large cuneiform. Q. What is the relative size of the ossa cuneiformis ? — A. The one beneath the astragalus is the largest ; the middle is the medium ; and that at the posterior part of the hock is the smallest. small cuneiform bone (os cuneiforme parvum). Q. What is the situation of the small cuneiform bone ? — A. It is situated at the posterior part of the hock. Q. What are the articulations of this bone ? — A. It articidates superiorly with the internal angle of the large cuneiform ; anteriorly, with the same angle of the mid- dle cuneiform ; posteriorly, with the inter- nal splent bone and cannon. HIND CANNON (oS METATARSI MAGNUm). Q. What is the popular name of the hind cannon ? — A. Shank-bone. Q. How does it compare in length with the cannon of the fore extremities ? — A. It is about one-sixth part longer than the fore cannon. Q. Is there any difference in the supe- rior surfaces of the fore and hind cannons ? — A. Yes ; the superior surface of the fore cannon corresponds to the surfaces of the inferior bones of the carpus ; the superior extremity of the hind cannon closely resem- bles the surfaces of the middle and small cuneiform bones, and also that of the cuboid. Q. How do the hind and fore cannons differ in conformation ? — A. The bone of the Iiind extremity is more circular and prominent, anteriorly, than the forward one. METATARSI PARVIUM (hIND SPLENTS.) Q. What is the situation of the metatarsi parvium? — A. They are situated at the posterior part of the liind cannon. Q. How are the hind splents recognized from those of fore limbs ? — A. The hind splents are longer than the fore ; their bodies are more circular and prominent forward, and the superior extremities correspond to a part of the cuneiform and cuboid bones ; while the superior extremities of the forward splents correspond to a portion of the in- ferior row of the bones of the knee. We now come to the bones articulating beneath the inferior extremity of the hind cannon, viz., pastern, sessamoid, coronet, coffin, and navicular bones. These, according to the opinion of Mi-. Percivall, " so closely resemble their fellows of the fore extremity " that we shall dispense with examinations regarding them, merely remarldng that the bones of the hind feet are generally broader in a lateral and posterior direction than those of the fore ; the pastern and coronet bones are somewhat longer than their fellows forward. BONES OF THE EAR. Q. Name the bones of the ear. — A. Malleus, incus, stapes, and orbiculare. Q. What is the form of the malleus? — A. It appears to resemble a mallet. Q. Name the long process or handle. — A. Manubrium. Q. To what is the manubrium attached? — A. To the membrana tympani. Q. Describe the form of the incus. — A. It is said to resemble a blacksmith's anvil, but, probably, approaches nearer to the figure of a molar tooth ; it has a depression on its surface, which receives the head of the malleus. Q. Describe the stapes. — A. It resem- bles in form a common iron stirrup, yet has a more triangular appearance. Q. With what bone does it articulate? — A. The OS orbiculare. Q. Describe the OS orbiculare. — A. It is the smallest bone of the body, not exceeding in size a grain of mustard-seed. Q. What is its use in the mechanism of the ear? — A. It forms the medium of junction and communication between the incus and stapes, and facilitates the motions of the latter bones. THE HORSE. 53 03 HYOIDES (bone OF THE TONGUe). Q. Wliat is the situation of the os hyoides ? — A. It is located at the root of the tongue, at the anterior part of the larynx. Q. How is the bone divided ? — A. Into a body and four horns. Q. What is the form of the body ? — A. In shape, it resembles a spur, consisting of neck and branches ; the neck is inserted into the root of the tongue, and the branches are in a posterior direction, embracing the superior border of the thyroid cartilage. Q. What is the appearance of the horns ? — A. There are two long and two short horns; the short, or inferior, ascend oblique- ly from their articulations with the body of the bone, and terminate in oblong, smooth extremities. The long or superior horns constitute two long, flattened, thin bones, extending backward in a horizontal direc- tion from the summits of the inferior horns. Q. What are the connections of the os hyoides ? — A. It is connected with the temporal bone, larynx, pharynx, tongue, and some of the muscles of the neck. OF THE TEETH. Q. How many teeth do we find in the jaws of the adult horse? — A. Forty. In the mare, however, the canine teeth are generally imperfect or undeveloped. Q. How are the teeth divided ? — A. Into tlaree classes, viz.: incisors, or nippers; molars, or grinders ; canini, or tusks. A. Enumerate each class. — A. There are twelve incisors, twenty-four molars, and four canine. Q. Is there anything peculiar about the development of horses' teeth ? — A. Yes ; the teeth with which the animal is furnished during colthood are termed temporary, and are generally shed ere the animal arrives at the age of five ; the temporary teeth are twenty-four in number, twelve incisors and t^velve molars ; they differ from what is termed the "permanent set," in being small- er and whiter, and in having necks or con- tractions at the superior part of the fang, and the eminences on their face are quite sharp. The converse is the case with regard to the permanent teeth. Q. What is the popular theory regarding the periods of cutting the teeth? — A. A foal is said, at birth, to be in the act of cut- ting twelve molars, three on each side of the jaw bone ; at this time, there is no ap- pearance of incisors ; and when they do appear, which period will be about the second or third week from birth, sometimes sooner, the front incisors of the upper jaw are the first to show themselves, and be- t^veen the fourth and fifth week, they are succeeded by the middle incisors ; the side or lateral incisors make their appearance between the sixth and tenth month. The animal is then said to have a full set of temporary teeth. After the animal has at- tained hisfu-st year, the fourth molars malie their appearance. Between the period of the first and second years, the fifth molars, in each side of the jaw, are apparent. Be- tween the second and third years, the front permanent incisors displace the temporary, and, at the same time, the first temporary molars are shed, and replaced by the perma- nent. Between the third and fourth years, the middle temporary incisors are succeeded by the permanent, and about the same time the second temporary molars are shed. During the interval of the fourth and fifth years, the lateral permanent incisors appear ; the sixth and last, permanent molars are up, and then the tusks also appear. At this period the horse is said to have a fuU mouth; a complete set of permanent teeth.* We have now arrived at an era (or re- * On this side the Atlantic we are not in possession of any reliable information as regards the periods of cutting and shedding teeth ; we have to depend entirely on English authority. Their theory is, that the age of a race-horse shall be reckoned from the month of May in the year of his birth, without any inquii-y whatever as to the season, month, or day of foaling ; so that the produce of January are actually four months older than by reckoning, or as their ages appear on the calendar, and these are called early foals ; whereas those foaled in March are denorai- nated-/ate. These data are more arbitrary than truthful ; may suit the convenience of English turfmen, but will not pass current among our breeders, — who, generally, pay particular attention to the time of foaling, and date the birth of the colt accordingly. 54 ANATOMY AND PHYSIOLOGY OF markable period) in the age of the horse ; have briefly considered a series of changes which the teeth of a colt undergo, up to the period of maturity, and shall now turn our attention to the changes observed in the process of wear and tear of the perma- nent teeth. REMARKS ON THE CHANGES WHICH A, HORSE'S TEETH UNDERGO. The nippers or front teeth of a fiiU- mouthed horse, just having shed all the temporary ones, present a beautiful ap- peai-ance : the contrast bet\veen the lily whiteness of the teeth, and the rose-tinted color of the gums and their membranes, are never so much the subject of admiration as at this period. Teeth, when first cut, present a sharp border externally, from which a gradual depression commences until the internal border is reached ; in the course of about a year, in consequence of friction on the external, and growth of the internal, the surface presents two elliptical enamelled rims, one of which borders the face of the tooth, the other encircles the depression or pit. Within this pit is a black incrustation, which is denominated " bean " or " mark ; " at a period of about ttaee years from the time of cutting the permanent teeth, the pit or cavity is consolidated or fiUed up, and the surface of the tooth is worn down so as to present a comparatively smooth one. We must not expect, however, to find the face of the teeth uniform ; for cribbers, and voracious feeders, deface the surfaces very much, which gives to the teeth the appear- ance of age. Still, a good judge, who takes into consideration not only the appearances of surfaces, but also the form and direction of the teeth themselves, is not apt to be deceived regarding the age of a full-mouthed horse. Pessina, from whose work ]\Ir. Percivall quotes, concludes that — " At the age of eight (in most horses). the disappearance of the marks is perfect: the teeth are all oval, the central enamel upon the face is triangular, and nearer to the outward than the inward border, and the cavity of the tooth appears within the outward border like a yellowish band carried from one side to the other. " At nine years, the front teeth appear round, the middle and the lateral contract their oval faces, and the central enamel di- minishes and approaches the inward border. " At ten, the middle teeth become round, and the central enamel has approximated the inward border and is rounded. " At eleven, the middle teeth are rounded, and the central enamel is almost worn off the posterior incisors. " At twelve, the lateral teeth are rounded, the central enamel has quite disappeared : the yellow band has grown wider, occupies the centre of the face of the tooth, and the central enamel continues in the teeth of the upper jaw. " At thirteen, all the incisors are rounded, the sides of the front teeth spread out, and the central enamel continues in the upper jaw, but is rovmd and approaches the inward border. " At fourteen, the faces of the front inci- sors put on a triangular appearance, the middle grow out at their sides, and the cen- tral enamel of the upper teeth diminishes, but still exists. " At fifteen, the front teeth have become triangular, the middle enter upon that figure, and the central enamel of the upper jaw is still visible. " At sixteen, the middle are triangular, the lateral commence that shape, and the enamel of the upper teeth has disappeared. " At seventeen the triangular figures of Ihe posterior jaw are completed ; but their triangles are equilateral until the eighteenth year. Then their sides lengthen in succes- sion from the front to the lateral teeth, in such a manner that — " At nineteen, the front teeth are flat- tened from side to side ; " At twenty, the middle incisors have taken on the same shape ; lastly — THE HORSE. 55 " At h\'enty-one the lateral teetli are also flattened." Professor Passim* "systematically di- vides the lifetime of the horse, wliich he computes at thirty years, into six periods, that take their rise from and are determined by an equal number of changes the teeth naturally undergo, in regular succession. " The first period is that during which the animal retains aU or any of his milli teeth ; it extends from birth to the fifth year. " The second period includes the sixth year, and continues so long as the marks remain visible upon the faces of the pos- terior incisors; which is generally about tliree years. " In many instances, however, and espe- cially among horses that have been kept at pasture, the faces of the front teeth, and sometimes those of the middle, are worn off earlier. " The third period is that during which the teeth retain the oval form. As the pits and marks degenerate, the face of the tooth slowly and gradually undergoes a de\dation of figm-e, from that of a pretty regular ellip- sis, whose long to its short axis bears the proportion of six to three, to an irregular one, in which these proportions are as five to four. This period requires, on an aver- age, the space of sis years for its comple- tion ; the front teeth enter it in the seventh and conclude it at the expiration of the tn^elfth; the middle pass through it one year later; and the lateral, cr side teeth, one year later still. " In the fourth period the faces of the teeth assume a circular figiu-e, and hence have been denominated round. At the commencement of this period, the breadth of the face to its thickness is as 5 to 4 ; at the conclusion, it measures in an inverse ratio, as 4 to 5 ; about the middle of it, the diameters are equal. Tliis period also endures six years ; so that the front teeth, which enter it in the thirteenth year, complete it by the expiration of the eighteenth ; the middle follow one year later ; the lateral, one year later still. * See Percivall's Lectures. k " During the fifth period, the face of the teeth deviates by slow degrees from the round, and passes into the triangular state. In the beginning, its thickness exceeds its breadth as 5 does 4 ; in the end, as 6 does 3. It is the professor's opinion, yet uncon- firmed by experience, that this period, lilte- wise, on an average, includes a space of six years; the front teeth, therefore, complete it with the twenty-fourth, the middle with the twenty-fifth, and the lateral with the twenty-sixth years. " The sixth and last period is one, in the course of which an additional angle is projected from the anterior or inferior part of the tooth ; Pessina distinguishes it by the epithet biangular ; he has never met with a horse that had lost his teeth from age ; but he has seen their faces elliptrical con- trariwise, looking outwards or forwards. This period is milimited. " In the anterior, or upper jaw, the marks disappear from the front teeth in the course of the ninth year ; from the middle in the tenth; and from the lateral in the elev- enth. " What progress these upper teeth have not made in transformation diuring the second period, equivalent with the poste- rior, they gain it in the third; notwith- standing the depth of pit, their proportions are then the same. They continue three years longer in the second, and consequently are only three in the thfrd period ; so that, by the twelfth year, the third period is completed by the front upper teeth, and so on. During the ioxvcih, fifth, and last periods, the changes are alike, and equally perceptible in either jaw. " So far, the upper teeth are entitled to an equal share of our regard; though, in the generality of cases, they need not be inspected. In such a remarkable man- ner the lateral teeth of the upper jaw wear away so that they often appear as if notched or indented. " In regard to the tusk or tush, Pessina remarks that he has found the least regu- larity in its changes of any tooth. The very facts that the tushes are not in all 56 ANATOMY AND PHYSIOLOGY OF THE HOESE. horses cut at the same age, that they have little or no attrition against each other, and that they are worn by the tongue and food, sometimes more, at others less, should lead us to draw conclusions from them with great caution ; in fact, as indications of age, they can only be trusted to when they accord with the incisors. The tush or tusk makes its appearance by the fifth, and is completely evolved by the sixth year. In the seventh, the apex of the cone is worn off. In the eighth, its furrows grow shallow ; in the ninth they are obliterated. Then the apex gradually wears away , in the twelfth year it becomes round ; from which time, though it gradually becomes shorter, its shape varies but little. But it is not uncommon to see the tush blunted like an acorn in the ninth year, nor to find it still pointed in the sixteenth year. " Pessina concludes his account of the changes to which the teeth are subject, by observing, that, as they are dependent on wear, which is no law of nature, but an effect of mechanical and accidental causes, they cannot, but under certain lim- itations, be implicitly reUed on." We are now supposed to be in posses- sion of some of the most important facts tending to elucidate the changes which the teeth vindergo ; and, in view of maldng our- selves more conversant with this subject, we shall re-commence our examinations, for it is a matter of the highest importance that a veterinary surgeon shall understand the method of ascertaining a horse's age. EXAIMINATIONS ON THE TEETH. Q. Does the evolution of the tush always indicate that the animal is five years of age? — A. No. It has been seen between the tliii-d and foiu'th years. Q. Which teeth do you place the most reliance on in ascertaining the age of a horse ? — A. The side or lateral of the lower jaw. They make their appearance last ; their pits are the last to disappear ; after the age of eight or nine, however, the pits in the incisors of the upper jaw are also indicative of age ; they, being deeper, of course remain some time after all vestiges of the same have disappeared in the lower jaw. Q. In adult life is there any continued accretion or after-growth of the teeth ? — A. Yes. If it were not so, the animal would, in course of time, have to gather food, and grind the same with his gums ; for, acccord- ing to the law of icear and tear, destruction of the in- struments — -grinders of food — must more or less reg- ularly take place. Q. AVhat changes talic place as the horse advances in age, in the incUnation of the incisors? — A. They acquire a horizontal direction. Q. How is this change of dhection compensated for in the grinders ? — A. The faces of the latter are worn down by friction, and thus the nippers come in contact. I Q. Are there not times when the consumption of the faces of the teeth, by ii-iction, is not in proportion to growth, in issue from the socket ? — A. Yes. I Q. What is the result ? — A. The faces of the grind- ers do not come in contact, and the food is, conse- quently, imperfectly masticated. » Q. How is this rectified ? — A. By sawing off the nippers to their natural length. Q. Talcing it for granted that there is a time when ' the teeth cease to grow, how do you account for the lengthy teeth observed in aged horses ? — A. The fang shrinks, and is carried upward in the lower and down- . ward in the upper jaw, and the gums also shrink; thus we get length of teeth. Q. What are the general appearances of age, uncon- nected with the teeth ? — A. The muscles of the head and face condense, and give to the same a lean appear- ance ; the canities above the eyes are deep ; the gums and palate become pale and callous ; the submaxillary space is capacious, and gray hairs make their appear- ance in various jjlaces ; the neck appears small and n-iry, the withers sharp, the back ciu-ves, and the hmbs appear sinewy. MYOLOGY. PRELIMINARY REMARKS ON THE MUSCLES. To the naked eye, the muscles appear to be composed of fasciculi, or bundles of fibres, which are arranged side by side in the direction in wliich the muscle is to act, and which are united by areolar tissue. These fasciculi when separated appear like simple fibres, but when examined under a microscope are found to be themselves fas- ciculi, composed of minuter fibres, bound together by delicate filaments of areolar tis- sue. By carefully separating these, we may obtain the ultimate muscular fibre. This fibre exists under two forms, the striated and non-striated. The former is chiefly distinguished by the transversely-striated appearance which it presents. The non- striated consist of a series of filaments which do not present transverse markings. At an early stage of the development of muscular fibre, however, there is no differ- ence in the forms of either striated or non- ' striated. Both are simple tubes, containing a granular matter in which no definite arrangement can be traced, yet presenting enlargements occasioned by the presence of nuclei. But, whilst the striated fibre goes on in its development, until the cells of the fibrillas are fully produced, the non-striated fibre retains throughout life its originsil embryonic condition; the contents of the tube remaining granular. The non-striated muscular fibre is the kind of structure proper to the muscular coat of the alimentary canal, bladder, uterus, trachea, bronchial tubes, etc. They seem to be arranged in a parallel manner into bands or fasciculi, without any very definite points of attach- ment. On the other hand, striated muscular fibre has attachments to its extremities of fibrous tissue, through the medium of which it exerts its contractile power on the part it is destined to move. At the truncated extremity of the striated muscles we find tendons. To the ordinary observer, tendons appear to unite abruptly with muscular fibre ; but this is not the case, for tendinous fibres are distributed over the whole muscle, crossing it diagonally in both directions, so as to form a double-spiraUy extensible sheath ; the tendinous fibre finally collects at the extremity of a muscle, and forms the tendon. Each muscle is smrounded by cellular membrane, which dips into its substance, and, by means of the fat which its cells con- tain, lubricates the parts, and thus guards against friction. A TABLE OF THE NAMES AND NUMBER OF MUSCLES, DIVIDED INTO REGIONS. SUBCUTANEOUS REGION (BENEATH THE SKIN)! 1. Panniculus carnosus. AURICULAE REGION (MUSCLES OF THE EAR). 2. Attollentes maximus. 3. Attollentes anterior. 4. Attollentes posterior. 5. Anterior conchje. 6. Posterior conchae. 7. Retrahentes extemus. 8. Eetrehentes internus. 9. Abduoens vel deprimens aurem. PALPEBRAL REGION (MUSCLES OF THE EYELIDS). 10. Levator palpebra; superioris. 11. Orbicularis palpebrarum. OCLXAR REGION (MUSCLES OF THE EYE). 12. Levator palpebrae superiorus internus. 13. Levator oculi. (57) 58 ANATOMY AND PHYSIOLOGY OP 14. Depressor oculi internus. 15. Abductor oculi externus. 16. Adductor oculi internus. 17. Olibquus superioris. 18. Obli(|uus inl'erioris. 19. Retractor oculi. ANTERIOR MAXILLARY REGION (MUSCLES OF THE NOSE AM) FACE). 20. Zygomaticus. 21. Levator labii .superioris aliquae nasi. 22. Dilator naris lateralis. 23. Nasalis longus labii su])erioris. 24. Caninus vel levator anguli oris. 25. Buccinator. 26. Depressor labii inferioris. 27. Levator menti. 28. Dilator narium anterior. 29. Nasalis brevis labii su])erioris. 30. De])ressor labii superioris. 31. Orbicularis oris. POSTERIOR MAXILLARY REGION (lIUSCLES OF THE TTF4T) AND CHEEKS). 32. Temporalis. 33. JIasseter. 34. Stylo-maxillaris. 35. Pterygoideus internus. 36. Pterygoideus externus. HYOIDEAL REGION (MUSCLES BETWEEN THE BRANCHES OF THE LOWER JAW). 37. Digastricus. 38. Mylo-hyoideus. 39. Gcnio-hyoideus. 40. Plyoideus magnus. 41. Hyoideus parvus. 42. Stylo-hyoideus. GLOSSAL REGION (MUSCLES OF THE TONGUE). 43. Hyo-glossus longus. 44. Hyo-glossus brevis. 45. Genio-hyo-glossus. 46. Lingua lis. PH\RYNGL\L REGION (MUSCLES ABOUT THE PHiRYNX). 47. Hyo-pharyngeus. 48. Palato-pharyngeus. 49. Stylo-pharyngeus. 494. Constrictor phaiTngis, anterior. 50. Constrictor pharyngis, medius. 51. Constrictor pharyngis, posterior. L.iRYNGEAL REGION (MUSCLES ABOUT THE L-tRYNX). 52. Hyo-th)Toidcus. .53. Crico-thjToideus. 54. Crico-ai'jtenoideus posticus. 55. Crico-arytenoideus lateralis. 56. ThjTo-arytenoideus. 57. Aa-j1enoideus. 53. Hyo-epiglottideus. PALATINE REGION (SIUSCLES OF THE PALATE). 59. Tensor palati. 60. C'ircumflexus palati. MUSCLES OF THE NECK. HLTUERO-CERVICAL REGION (MUSCLES SITUATED ON THE LTPER AND LOWER PARTS OF THE NECK). 61. Khomboideus longus. 62. Levator humeri. LATERAL CERVIC.U, REGION (SIDE OF THE NECK). 63. Splenius. 64. Complexus major. 65. Trachelo-mastoideus. 66. Spinalis colli. su^ERo-CER■\^co-occIP^AL region (muscles situated abo^t; the head). 67. Complexus minor. 68. Rectus capitis posticus, major. 69. Rectus capitis posticus, minor. 70. Obliquus capitis, superior. 71. Obliquus capitis, inferior. INFERIOR CER\1CAL REGION (MUSCLES SITUATED IN THE ANTERIOR PART OF THE NECK). 72. Sterno maxillaris. 73. Sterno-thjTo-hyoideus. 74. Subscapulo-hyoideus. 75. Scalenus. 76. Longus colli. INFERIOR CER■^^CO-OCCIPITAL REGION (MUSCLES BE- NEATH THE BASE ATLAS). 77. Rectus capitis anticus, major. 78. Rectus capitis anticus, minor. 79. Obliquus capitis, anticus. MUSCLES OF THE CHEST. DORSO SCAPULAR REGION (MUSCLES SITUATED ABOUT THE SnOUXDER BLADE). SO. Ti-apezius. 81. Latissimus dorsi. 82. Rhomboideus breris. PECTORAL REGION (MUSCLES SITUATED IN FRONT OF THE ERE.iST BON'S). 83. Pectoralis, transversus. 84. Pectoralis, magnus. 85. Pectoralis parvus. COST.\L REGION (MUSCLES SITUATED EXTERN.iL ANT) INTERN.AL TO THE RIBS). 86. Serratus magnus. 87. Intercostales extcmi. 83. Intercostales interui. STER.\AL REGION (MUSCLES OF THE BREAST BONE). 89. Lateralis sterni. 90. Sterno-costalis, extemi. 91. Stemo-costalis intemi. THE HORSE. 59 DOKSO-COST.VL Ri;GIOX (MUSCLES ON THE SIDES AND ITPCR PAKT OF TDE CHEST). i)2. Superlicialis costarum. 93. TraiisversaKs costarum. 94. Levatores costarum. DOUS.VL REGION' (MUSCLES OF TIIE DACK, .VXTErUOR TO THE LUMBAR ^-ERTEBICE). 95. Longis.simus dorsi. 96. Spinalis dorsi. 97. Semi spinalis dorsi. DLU>HR.\G5LiTIC REGION. 98. Diaphragm or midriif. MUSCLES OF THE ABDOMEN. LLTUBAR REGION (MUSCLES OF THE LOINS). 99. Semi spinalis limiborum. 100. Intertransvcrsales lumbonmi. 101. Sacro lumbalis. 102. Psoas Magnus. 103. Iliacus. 104. Psoas par\Tis. ABDOMLN.\L REGION (MUSCLES OF THE ABDOMEN). 105. Obliquus externus abdominis. lOG. Obliquus internus abdominis. 107. Transversalis abdominis. 108. Rectus abdominis. AN.\L REGION (MUSCLES OF THE ANUS). 109. Retractor aui. 110. Spliincler ani. GEXIT.IL REGION (MUSCLES OF THE M,\I,E ORGANS OF GENERATION). 111. Cremaster. 112. Erector penis. 1 13. Triangulai-is penis. 114. Accelerator urinEe. The muscles in the genital regions of the female are named : Erector Clitoridis, Sphincter Vagina;. COCCTGEAL REGION (MUSCLES OF THE T.\IL). 115. Erector coccygis. 116. Depressor coccygis. 117. Curvator coccygis. 118. Compressor coccygis. MUSCLES OF THE FORE EXTREMITIES. EXTEKN.VL SCAPU'LAR REGION (MUSCLES ON THE OUT- SIDE OF THE SHOULDER BLADE). 119. Antea-spinatus. 120. Porte a-spinatus. INTERNAL SCAPULAR REGION (MUSCLE ON THE LVSIDE OF SHOULDER BLADE). 121. Subscapularis. POSTERIOR SCAPUXAR REGION (MUSCLES BEHIND THE SHOULDER BL.\DE). 122. Teres major. 123. Teres minor. ANTERIOR HU-JIEEAL REGION (MUSCLES IN FRONT OF THE OS HUMERI). 124. Coraco-humcralis. 125. Flexor brachii. 126. IlumeraKs externus. POSTICRIOR UUMER.iL REGION (MUSCLES BEHIND THE 127. ,. . 128. lis 129. £ § g 130. w"' OS HUMERI). Ca|)ut magnum. Caput medium. Caput parvum. Anconeus. MUSCLES OF THE ARM AND FORE LEG. .AXTERIOR BRACHIO CRURAL REGION (MUSCLES IN FRONT OF THE ARM). 131. Extensor metacai-j)i magnus. 132. Extensor pedis. 133. Extensor suflraginis. 134. Extensor metacarpi obliquus. SUPERFICIAL POSTERIOR BR.\CniO CRURAL REGION (MUSCLES ON THE EXTERN.AL SIDE OF THE ARM). 135. Flexor metacarpi externus. 136. Flexor metacarpi medius. 137. Flexor metacarpi internus. 138. Flexor accessorius sublimis. DEEP POSTERIOR BRACHIO CRITI-AL REGION. (THESE MUSCLES ARE SITUATED BENE.VTU THE FORMER.) 139. Flexor pedis perforatus. 140. Flexor pedis perforans. 141. Flexor pedis accessorius profundus. 142. Lumbrici, anterior. 1424. Lumbrici, posterior. MUSCLES OF THE IIIXD EXTREMITIES. GLUTKAL REGION (MUSCLES OF THE SUPERIOR P.ART OF THE QU.VRTER). 143. Gluteus externus. 144. Gluteus maximus. 145. Gluteus minimus. PELM-TROCHANTERLVN REGION (MUSCLES SITUATED AT THE UTPER PART OF THE THIGH BONE). 146. Pj-rilbrmis. 147. Obtm'ator externus. 148. Obtm-ator internus. 149. 150. Gemini. ANTERIOR rLIO-FEMOR.U, REGION (MUSCLES SITU.WED AT THE FORE PART OF THE HAUNCH). 151. Tensor vaginEe. 152. Rectus. 153. Triceps vasti. 154. Rectus parvus. INTERN.AL ILIO FEMORAL REGION (MUSCLES SITUATED AT THE INN-ER PART OF THE HAUNCH). 155. Sartorius. 60 ANATOMY AND PHYSIOLOGY OP 156. Gracilis. 157. Pectineus. 158. » o C Adductor brevis. 159. .§ 3 < Adductor longus. 160. file Adductor magnus. POSTEEIOR ILIO FEMOEAL REGION (MUSCXES ON IHE OUTER ANT3 POSTERIOR FART OF THE HAUNCH). 161. Biceps abductor. 162. Abductor tibialis. MUSCLES OF THE THIGH A^^D LEG. ANTERIOR AND FEMERO-CRURAE REGION (MUSCLES IN FRONT OF THE TIBIA). 163. Extensor pedis. 164. Peroneus. 165. Flexor metatarsi. SUPERFICIAL POSTERIOR FEMORO-CKUEAL REGION (MUSCLES IN THE REGION OF THE HOCK). 166. Gastrocnemius extemus. 167. Gastrocnemius internus. 168. Plantaris. DEEP POSTERIOR FEMORO-CRUHAL REGION (MUSCLES •n-HICn ARE FOUND BENEATH THE FORMER). 169. Popliteus. 170. Flexor pedis. 171. Flexor pedis accessorius. The muscles of the internal ear are named : Laxator tympani, 2 Mcmbrana " 2 Tensor " 2 Stapedius, 2 Total, 8 RECAPITULATION. "We shall now recapitulate, as regards what has preceded, in reference to the num- ber of muscles ; for there exist various opinions regarding the same. It may be proper for us to bear in mind, however, that VETERINARY SCIENCE, here, is yet in its in- fancy ; and it is well known to some prac- titioners, that there are several muscles which remain to be named by some future compiler of veterinary literature. But for all practical purposes we know enough of the anatomy of the horse. The industrious individual, however, who not only desires to make himself conversant with what is al- ready known, but aims to improve in the future, will not rest satisfied with the pro- ductions of his predecessors. To such an one we bow with due deference, and encourage him to proceed in the work of progression. There is a fine field for exploration, and a discerning public are ready and willing to crown the industrious laborer with the laurel of merit. In the preceding table, the number of muscles, including those marked 49 1-2 and 142 1-2, appears to be 173; among these are ten single ones, wliich are thus expressed: Whole mmiber, .... 173 Deduct single ones, .... 10 Pairs, . Multiply by 163 2 Single muscles, 326 Add muscles of the internal ear, four pairs, 8 Single muscles, as above added, 334 10 It appears, therefore, that there are in the system of the horse three hundred and forty- four muscles. It should be borne in mind, that in the preceding classification all are considered as muscles. Among them are found tendons, which are component parts, or rather ap- pendages, to the same. Mr. Percivall says there are, in the horse, 151 pairs, and 10 single muscles ; add the four pairs of the in- ternal ear, which he has omitted in the cal- culation, and we get 155 pairs. On page 72, " Hippapathology," the number of muscles is 312 Add muscles of the ear, omitted, . . 8 The author's estimate, Diiference, 320 344 24 Probably the above author considers the " 24 " as tendons. ^ •& ^ EXPLANATION OF FIGURE VII. NO. 1. — OSSEOUS STRUCTURE. 35. Radius. g. Trapezium. 36. Lower row of the carpal bones. 37. Metacarpus magnus. 38. Sessamoids. 39. Os suf&aginis. 40. Os corona. 41. Os pedis. NO. 2. — MUSCULAR STRUCTURE. INTERNAL VIEW OF THE NEAK-FOKE LEG. o". Pectoralis transversalis. 2 . Flexor metacarpi medius. r". " " iiitcrnus. s". Extensor metacarpi magnus. t" " " obliquus. m". iC. Flexors pedis — perforatus et perforans. v". Suspensory ligament. x". Extensor pedis. z. 8. Bifurcation of the suspensory hgament. NO. 3. The description of No. 1 answers also for No. 3. The letter /. is intended to point out the location of the ulnar, into which is inserted the triceps. g. Region of the carpus. NO. 4. n". Triceps extensor brachii. o". PectoraHs transversahs. o'. P". Flexor metacarpi externus. q". Flexor metacarpi medius. «". Fleshy beUy of the perforatus et perforans. a;". Extensor pedis. g". Extensor sufiraginis. z". u". v". Flexors tendons. k. Hoof. THE HORSE. «1 3 "^ ." CJ ;:3 I is >^ O ' rt S3^ 32 A r^«— * ^ ■So s ° s go a-g o-g a>i o o o =* ■^-' o -^ c H^&H H HE "§-"1 "^^ •- s ^ o S o Td2 ' bD -= J5 2 "S ,r --T " S-2 g 5dS c "3 o &, ^ ^ "^ 5 o C c; fj S S ^; M o o a M C -^ — ^ ^ .2 o bc if 5 s 5 43 -^ :Z! 2 ~ cr ^ CO -C T^ ^ Q,^^ ^ -t-^ =^3 ^ ^ > O IP'S « => o .i bD E n ^ Ch ^ -B -2 -3 o o o o 5§ o aj ?i a; 2. o " o O >- r- •^ "^ S g _m O — ' tw o ,o -o " , :g ° :S ^ -g « ' o J!; o cr-S C3 .iS ■-2 .2 5 -^ p „ O 0) o =" "^ O ^ r^ a " o ~ 'o ^ 5:^ +j 1^ +^ 'a a.-9 afipfi C3 bn m CJ -a D 3 be C g J3 O '-' o t( _^ T-l K o CI J5 (U a. •^ -s tt H H C rrj c 03 "ca •s o -2 >.■-? O Q O O t,_i ono 'S & m cc d -1 is C - ^ J5 ,Q x: [o J3 ■ ^H -+-i "-1 -t-^ n hShh P-( Ph Si a ^ <-< J J IC CO I>QD G5 O .- -£.2 o P _H n !? ;:! ro tn o O O pj ?3 a-i "o "o P P gJ O 3 CT C S-TS TJ ;p ;3 57^ 13 X! ^ •«ti lO to O 00 Pi 62 ANATOMY AND PHYSIOLOGY' OP V c-o-s X o s c ^ O S s « .- s +: iS " c to c c £: 5=3 rt = ,, t^ = c ttJ ^ ~ r i cS fl ~ ° v s ° « s <1 H c^ -^ ■:=13 P =. (u -r ^ i- ^ 7 "" to i 1- C c3 .-f j3 — « T? r! r^ A ^ -; C3 C s r3 ^ is (H -3 a) -■ O C OJ 22 -c o js 2 : _ ■ - S -^ = ~ rt C ? c: • : § > S J-l ^ ' S "S "3) a; .^ cj : 2 ^ c :5 =« s . ,0 CS C3 >-^ r aJ rt £ ^ aj 1 +^ « tx? g u: i o 2 "" o-"^ c, C- aj rt g - c O K -d ' •-■ to H be--; ■£ f; cJ 2 "O S S ^ ^ 5 4i. die — ■ a c p • ; ig o 0) ~ i: J5 o -c -t- o • S3 i^ s '-( X 0) o n) c ^ Jl^ jp C) tn _o S J2 -3 >.o O aj jp 5 O a. P o aJ D S a) _e a) *^ CI 5<« > a5 O "m ■« S 0) (11 ^-tJ UJ c d) o Si a. -*-* P 1 " 1^ fo ^^^ 9-0 3 S Ph s o ^ 13 I bD O THE H0B9E. 63 O — ' •3 ^ r3 c - ->^ -1^ o o -^ o o H H H E-i i ^o -fS 4) .s 0) TS t« m be • s s -3 5 c o^ c • •+e aj TD s =3 ."S fi-be C3 C O P o S . ■2^ S-3 ^2 ^ c 5 oralis ig the '. pperj c iress a baclcw be- ^ C 0) g " y -2 '■i 0) c.~ c r- ^ ?r P -^ c/: — 2 J= 'O 'd •:3 .£■-£ o S Ji o a a -*^ ci ^ C: •-! 5.0 -g "1 £ -^ CD g -K . OJ ^ ^ ci S be c3 cS wj C "2 O n tilage rior a heir lie n o .2 > C3 G. a 3 O O . ^-P .5 ;:; o a ni Si-^ o — "d "^ "> •5 bb^ IS OJ El S -d^ 0) O O g ti. g_, O S C3 := c ••^-•^ o ■ -^ S -d-s ?i S !- OJ a C3 S i3 ^ J3 CO = ^ C 2t:oc.|-| ° e*^ £ S S ^ B ^- « -g S " t3 O O J^ ci -S S S '-' tn S fe OJ m -= ^ s C " ■^ ^ bCrt 3 O o "' 3 -O c3 matic ridge, rou le of the lower j parts of 1he ?• '3 fe o .a trj t(_ o o ■2- c'bc -d rt 3 E beo ,a ^oj gnSPS 'S 2 b-= tc^ ) — c ^ o ^ m S S :S 5 5 o "^ J3 9. S S 0.ii o -r -= ci ^ C o «a o' H Eh V. S rSt^ -a ,1, o •:i ,T! ^-^ c 5 Q, ■M o U Ph s S b 0) ClJ _rt o c +^ y .:_: ci fi lyj ^ e w Ph (54 ANATOMY AND PHYSIOLOGY OF P-. to j3 ^ a rt P te ,0 cS t2 '3 'p '^ s o — ' H ^ o-^ H ^5 gs 41! 03 ■3 >^ c3 -a '.Br'S ^^^ o g S § o J -^ S? ^_§ -t^ Qj a; >- "'^ ^-a£^ o to o C ^ a o U u <1) . ^ « o' o ^ ^ ' O ^ (U ■B'-S rt « 03 O t^ to to >-' 0) c ^ o ,> ^ S 5 (u - g^' o " "" ' -as. s a tongue to draw outh. .S 6i3 d C3 the , and le m C3 C3 S f? ^ 0) ~ c is t; o -;3 ^ g ra ^ i> a 2 Tl C aj d P bn n nd ! ■*^ ^ t4H rt -c C3 02 0-0 o -^ 3 ,/ ^ '-' to >-. bn 2 O ,^ "2 5 S c -? ~ => Eh •T3 i 11 5^ o u. ^ .=* "S ^ cr~" ~ ^ Cd "*"^ r^ to :2 ^ O s Qj rt Ch J2 ,1:3 O OJ c2 .a ^ -M § g-.a CD .2 -^ (U * >„ :£ .2 '^ :g -n .> r5 '^ £-a ss O C TS i2 ^H trr tC CI, c G *aj ; -G T) O cs o rt •5 (D «J TS C3 5 ^ "S . a a Tl 'to U 0) trrg •*^ X fe^ >i a, ^ J P 60 °, O W _>> 6 ffi CS &> i w THE HORSE. 65 • oj to o -d !3 J- to ij, .-1 :S > -2 Q S -^ 3 ?'^ be >-. O J3 ci "^M OJ j bo, ^ I Z Z ^^bh^ ^^M^^'-^c ^^ Is Zs ^s '-^+- ^^ Q ^ O >y a «^ '!? > -^^ Xtu rSn-, Hi Iris|.:ii|3^^ri=ii|i3||ii"=s Is III •rt ^ 2 -I "S^ S ^ -^'^2 5 "c^ .3 5^--:3 rt-M 2 s^bS q 2-1'^ 2-1 •d a~ Cr-^ja a « a~ S-S"^ c3i„*^ o 2 isi+--^^i-w a o-h e^ EcSocsWO-OoaoOGoSOo P--^ &,o+3:t3-Pr|C3Co^+^ ■aa+-'Cr^-ai-'i»«-»^cS--i "'<-i oc.ra-<- ecSoc3M-lo-ooao"'='oS°o ^^-^ ^^^ o „ _ Eh feHHH Eh EhEh H 2 2 ^ go cj ^ OJ a. Oh be 3 a 0) o ^ h "S 3 3 ^ 2 ^ g tc t« QJ -r^ -5 fe bfl ai ^ .jr ° :S ^ -J^ ^ 2 a 2"^ ab a'*^ aQ)^^--2.2a&':3o -g T^ o S,-2 0.0 g^:^ ^ c^ o n,i3 tH S W kS P , P P <1 P 1^ 'tJC hH -c; QJ cS to a R § 6.2 -5 ^ ■c " •c «« 43 u H 66 ANATOMY AND PHYSIOLOGY OF i2 CL. S S -c ^2 J m a) (U ^ tH O 13 ^ O rt M £ o I- -a ~ ■43 ^ ^ c 1:^ 5-- -a E-^is tc, £ S u o ij S > O ni 3 W p CD 6/) « S (D r; ^ c3 m o o a, 2 to i> ~ ' — I .43 0) o > O o "^•^ §^5 §"rt -« ^ E? c 5 J^-ls rt g g< ! J3 — • ■— CO ■ u :S S O -. ^ o -5 m g g -H" D cj O O -P OJ .S^ ^%-.^ o aj cs •- <^ ^ "^ .- ? ■^ ^ P. ^ 'S. „ o m ii >- « o -►^ to a, .S^r^l T3 O +3 ^ O O C _g t*- v^ J3 C -= C o r2 6C S.S O) O 0) 2 £.2 3 -^ •^ C3 6C ^Jlj rt ^ *3 (D CJ u^ ^ ^ lif^ ~'a ^ o Is o 1 rt S 2 ody rt of one. o "S ^ rt -" O > "' -^t:-^ o > S ° g 2 '^ =" o o >" ^ "lit ^ "^ c ^ <=> tn ^ '^ cu o S :S S S <« .2 c O -J^.5Prt o "S o -- K rt +3 "i -T^ s.S 5 S 2^ ^.rt e^ I , ^ o r rt OJ -iri 1;=; 5^ g ° §^ C C rt to O D CL, -3 ^ oj i! o Q HJ 4^ rt C H rt £ rt.S /- o- ^ :r' %:; rt be O -G .rt ' Eh O 0) !« o o ^ ;i b o rt O rt 'So;^ -e "o £'3 g •^ .-- ■*^ — -2 ^ o O-rt--^ ^ ^:^ & S ''B ^ ."S 'S rt O +3 o ^ _S g^ r^ ^ .O g D .c.ia r= 2 B D 'n O "v. O O o. a.2^ •^i ^it o d ^5 S2 0) O y 1) ion the u side of lind the &0 rt 5 3 s ^ c-g o .2 .-ts 5 "C '> "a, ."t; .O ^H -rt -^ 52 "3 P^ P^ THE HORSE. 69 ^ S H Eh H QC Eh Eh II lllis iiii bill's 1^ |ili&i« ^^ !•§» Eh H HH^HEhH 5:S'rt"^SS 'laojtc^^'^bn, o"7'-:=5'S-3 So'S-'-orh'nU'SScs-.m OD"*^ o :S H,^ " ^ 2 ^ ^ s «5_ s IP'S g s g ^„^ 4 ^ .s a g.£ ..£ s'tS : tc s ^ .-2 ■?= ~ ° v (1) fH 0) S- 3 it o a a a 1 O to ^e o m O two last; a The right cr all the lum eral tendin -fJ tc to s ^ -Q o ?2'2q2cijj>beooc3^ aa -^aaja^-wjDS 1 tj o a I a =" cs m ^ ^ -w ^ -7^ -a o :g to to j3 2 o O -+— H H (D a S' i-lT^ be^:?^ "- S o 2 S o 1^ ^^.2 <"' , ^ ^ -S,^ .S^ o o S "S o "o a 2 o o-^ u c ja ^■:| o g S o pq bc^ S,:; o S-a'* ■p 2 o <^ S . — ' .a S ,S 0^ pq fi S c3 be a figs "El ■•5e o <^ EXPLANATION 01^ FIGURE VIII. NO. 1. — OSSEOUS STRUCTURE. 22. Femur. 23. Patella. 24. Fibiila. 25. Os calcis. 26. Astragalus. 27. Inferior row of the tarsal bones. 28. Metatarsus magnus. 29. Sessamoids. 30. Os suffi-aginis. 31. Os corona. 32. Os pedis.' The above explanations will serve to illustrate No. 3. * * are the matatarsi pai'num. NO. 2. INSIDE VIEW OF THE OFF-HIND LEO. g. Rectus. p'. Vastus internus. 2*. X. X. Extensor pedis. q. Flexor metatarsi. r'. v\ Gastrocnemius externus et internus. t. Peroneus. u. The insertion of the gastrocnemi. v'. V. Tendon of the flexor metatarsi. z'. Suspensory ligaments. if. The hoof. 6. 5. The saphena vein. K. Abductor femoris. 8. Bifurcation of the suspensory hgament. it'. (Beneath the jjostern) Perforatus et perforans. NO. 4. K. J. Biceps, showing the manner in wliich it bifurcates. r'. Gastrocnemius internus. t'. Peroneus. v'. Flexor pedis accessorius. 5. " " externus. u'. Insertion of the gastrocnemi. y'. Peroneus. u. u'. v'. Flexors of the foot. «f. The hoof. THE HORSE. 71 rB ^2S| -t3 be •^ OT prox k^erse the bac^ 'P. J3 m •^^ c=l O o cj r; n ap trans nd a- draw s 2 M X ■-' Gh c3 ni . O o u. 9 -0 borum ing th cesses time t o o a, Cj o 1) o > O i O •" ? 5 « S c^ §- S ■« - -5 « 3 i '^ u rt a; ^ S „ ° Th CI, Q,j3 J3 O X 2 H H CS 1^ C3 ^ -fc o o a o ^ s s o c ^1 b !0 o -3 JS o a ,J3 O C- O ' C ii G -^ 1 ^ ^ I^ I ^i" -o ^ g^' ^ o ti •=; ■jj ■*^ s „r p^^ £^ o — "E OJ ttac lese the 3 -^ o g CS*- c-o f- &, H h H mi cS •+f P OJ O -y J 1^ a; -S fe o ° S g rt to „.^ 2 o d .a ^ o o S s- -t- rt . . ji O rl ^. inn thei and I ve limb antei &.-0 B o O s o 1l the ear ody orsa he I •och 2 '5 "rt tc o o O S ^ T) -.- -P o L'o n Xl +^ +3 Eh H H ^•s.e n> a 0) .5 3:2 bc-a a a &H a ■ ■;< H O -►- " g .2 " - '"^ ■-" " in » o -a ,~ a^ !a= J ^^ "S .a >> c » o a m -a ja a2 vm jj c •3 ii ^■^'a-S ■^ ci cTr^ a X J= ''c a ^ . o a a .S -a a as o ^ 5 S o a a C a a « r' o P^ ss 72 ANATOMY AND PHYSIOLOGY OF WJ -a c 1 c o o &1 f^!> . pom o ' '3 ^ c-^ o c o m e -1 CI- CJ S CJ *' tc +^ O J3 OJ "^ m P c3 , _L CO ?::: s-i ,j^ , 3 S §'2 ^■5 s a CS ??nPi2 cS -S Ei) 9 cs to "O O cd O^ ?n ^3, 043 c3 3 ^2p8o •^3 0:3 C3 alba. the inn the false i the lumbf o 3 O re linea the cart ue ribs a QJ 3 C a •^ o-P a. H H r5 OJ " -3 13 O 5 S H g S 2 c "3 "^ cs i 5 c : ? OJ ■ '^ s. 43 O . dj flj ct o- o s ^ ^ i Q CS >^ 1) Eu--<'^ Ct4H "3 li g J2 QJ ill c. ill P <; =s a O „ '- ' 3 c -s !^7 o 2 o 2 .^, ^ -fs .3 ^ J3 ? 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T. _o E 3 u o c c ii ■l-l c o 1 o "3 ci o o -a e" 1 1 _p |> CI. c 3 e" £ o ci c c p -i^ P c 5 £ c: c tc O p -1 o ^ ^ "5 5 .2 +^ •J go T -,0 >" C £ " S S a p "\ CS M p %-l -g o -; -^ £ s o ■*-' ^ ^ ci ^ 'p "^ ''" ^ ^ 3 "^ ^ ^ %. — ^ . 5 = '2 -S o -c :i s o '♦^ J5 ^ :S :~3 "o o i O 'c c J3 o £"0 +2 *-5 £b o £ 3 .3 -^ s t* -*^ H ^ E-1 n H Eh H 0!g tp O m o o 42 -r-t TS !E o _j_-, p ^"i ■s n o o _>, ^ C3 10" — 0. o 3 £1. "3 1 E Cli || [£ g — ^ ^ "3 £ - " "^ -jl '> "5 C -E [tp P ^ 3 g v^ 3 'a 'o c "3 ^1 3 3 c E £■ c t- ^ C P o ^ — £ J § < n_ cc _fi H^ < is o S "3 3 3 1 J d a p 3 s ft 5 ^ c ° O ■C 00 C5 »-( 6 •^ •^ «* Tj« ■^ 'J* ^ LO a l-H tH T-l ""• 1-1 T-( iH 1-1 iH ■^ .s THE HORSE. 77 2 i ■; W) = — — ^ = '^ rt -2 § ? ji , P. It^^l 5-:£:i :-§ >^^tc:: ^ .^ ^^ ^•5 3 &^^ -5 -^-o 1c ii^ -51 -g is 'S'S-JS g o^ St^t-g S,, „ og ^ -^ oJ: 2S-2.H— -K oo^ .- o o S = ^ " S C -v^ebn "^rttoSS-— i _^ t£j3 zi •- hVti »j ~ =— i C |^2§ -g'-g- £ i~ i^ S^, . ^ g g|^.5Pfc£2 H &>s-s S_g3"§5H_ H H J i "^ Eh Et ^r-5-« § 2 ^ Si~ « «§ i^ £.5 S.= g :g § g 2|g o^ .1 111113 I. -"^^^I II • :if i'^ b^-£ i^i:~'^rt t;^ ^j3-gg cjt^ -gcs ^^u^ ^|1 '.Isi| 2 §1 §^ [Ml -nt 1^ -^^ Sis |g^|:s S| |S -S|5.r |o 5 2 ^-3 ^ ^ g.s-3 QJ m H ^ HH 02 H H CC o a ^1 'c3 0) 0) o o ■a H ">> T3 e to •S;3 p I Vi -»^ OJ pl — C3 bD a 'a b*D a3 S o '2 a. to" bC S ^ .W5 CJ QJ ll QJ "3 QJ •i. o o 1 o s 5 _2 T3 ^ Vi o T3 to O c o o3 S > p ^5 2 OJ n QJ g O "*" o ^ ^ -3 s a ° i 5 c3 .a -^ c o O +2 1 i -*^ 3 ■3 be J3 c ° 2 |3 >>2^ ■3 a 'a 8 2 a, § '2-S o s be J3 a- a to c3 a, a _o OJ M 11 OJ J3 OJ -a t4H Ci 5 OJ C.£ CJ -75 -a a '^ o O -1^ J3 to -+^ ° +- -^ -f^ >-< H H C c 1—1 H H H H Is 13 t4-l C3 _o 1 "ij 1 -^XJ .s ^ &■ 'T3-d Oh ^ •i^ o rt E 13 J s 3 bJD tJ _QJ < G o 1 to . abD ~ 2 a ID II C3 £•0 QJ 73 rg 4J 1 S QJ to 1^ C &H -^ a _^ +3 ^ -^^ &.«~ *' D. O j^ CI -! PLl X y.. M ^ P^ rt s QJ OJ rf U3 CO t^ CD OJ 1-H 6 to O CO CO CO CO i> C» 2 tH rH 1— 1 tH l-H l-H t-( tH ON DISSECTION. No man can ever expect to become a practical anatomist or pathologist, unless he practise dissection. It is the only possi- ble way by which he can familiarize him- self with the healthy structmral organiza- tion of the horse. Having made himself acquainted with the healthy aspect of the various parts, their uses, etc., he next is able to judge of the various grades of textural change which occur and exist, between the part that has been studied under its healthy aspect, and that which has now departed from its healthy con- dition. Thus, in the prosecution of the study of anatomy, the student finally be- comes a pathologist ; and, although he may be a beginner, he places himself in a posi- tion only a few removes from the old and experienced practitioner, and can venture to " measm-e a lance " with the renoioned knights of the healing art. DISSfiCTING INSTRUMENTS. The dissector should supply himself mth a beak-pointed scalpel (which is one of Ger- man origin), for superficial dissection, and a myology knife, strong and rounded at its point. For the dissection of blood-vessels and nerves, a more delicate and pointed scalpel is needed. The forceps should be strong, and armed at the points with teeth ; two pair of scissors are needed, one pointed and the other blunt; a saw and blunt chisel, for opening the cranium. A blow-pipe, curved needles, and a few extra scalpels, are all that the student requires. SUBJECTS SUITABLE FOR DISSECTION. For demonstration of the muscular sys- tem, a well-proportioned and fully-developed subject should, if possible, be selected, and one that has died suddenly, or been killed in consequence of some accident, is to be prefeiTed. For making wet and dry prepa- rations, lean, emaciated subjects should be selected. The lymphatic system is best shown on animals of a flabby and cedema- tous organization. Young animals are the best subjects for dissection, in view of de- monstrating the circulatory and nervous systems. RULES IN REFERENCE TO DISSECTION OF THE MUSCLES. As there are abundance of subjects to be had in the United States, and it being in- convenient for one individual to dissect a whole subject, he had better divide it into six parts, viz. : 1st, The head and neck. 2nd and 3rd, The anterior extremities, wliich include the thorax, its contents, and the diaphragm. 4th and 5th, The poste- rior extremities, to which belong the pelvic and abdominal viscera. 6th, Those viscera which cannot be advantageously divided, as the heart, stomach, bladder, organs of generation, &c. Should the dissector de- cide to commence on the whole subject, he first removes the sldn, in order to expose the panniculous carnosus ; this will require some care, as some of the fibres of this subcutaneous muscle are intimately con- nected with the former. There are various ways of removing the skin : the author pre- fers to commence on the back, and dissect off towards the feet. Supposing the sub- ject to lie on the off-side, we commence an incision at the anterior part of the nasal re- gion, and continue the same upward until we arrive at the occiput ; we then in- cline the scalpel from the superior part of the neck, in order to avoid the mane, and (79) 80 ANATOMY AXD PHYSIOLOGY OF continue the incision along the lateral part of +he dorso lumbar spines until the coccyx is reached; the overlapping portion can then be dissected, and turned over to the off-side, so as to expose the tendinous in- sertions of the panniculus into the ligamen- tum nuchas, etc., etc. The panniculus being exposed, it may be divided into three parts, viz.: 1st, The Cervical portion, \\\\ic\i com- prises the head, neck, shoulders, and fore- arms. 2nd, The Thoracic portion, 3rd, The Abdominal portion. Having traced the attachments of the panniculus, the muscles then engage om- attention ; they being composed of nearly parallel fibres, the manner of displaying them is indicated. The cellular tissue should if possible be detached with the skin and panniculus ; without this precau- tion the surface of some of the coarser muscles would have a mangled appearance. The knife should always follow the direc- tion of the muscular fibres, and the part on which a muscle is to be dissected should be placed, if possible, in such a situation as to produce a forcible extension of that muscle ; thus, in tracing the origin and insertion of a muscle, the dissector becomes acquainted with its use. After exposing the external layer of mus- cles they may be detached from their in- sertion, or divided in their centre ; if di- vided, we thus preserve the two points of origin and insertion. The deeper seated mus- cles may be demonstrated in the same way. In the dissection of muscles the scalpel should be used in a free and prompt man- ner; the strokes should be long and bold, using the little finger to steady the move- ment of the hand. In making autopsies and in examining the viscera, the subject is generally placed on his back. ANATOMICAL PREPARATIONS. It is highly important that every student should be acquainted with the methods of maldng wet and dry preparations, and of injecting the blood-vessels; for specimens of this land are the best means of familiar- izing us with the structures of quadrupeds, •and such, when properly prepared, possess a real and practical value. INJECTING INSTRUMENTS. Pole describes three kinds of instruments used in maldng injected preparations. The first consists of a brass sjrringe, made of various sizes ; the nozzle is adapted to pipes into which the syi-inge is to be in- serted ; a short pipe, with stop-cock, also accompanies the syringe, which is to be applied betu'een the syrmge and either of the pipes. The second is a similar instrument, only much smaller ; its pipe is very minute, and its piston is furnished with a ring, so that the thumb may be used to throw its con- tents into a vessel. The third instrument is generally used for injecting the glands and lymphatics w^ith quicksilver. It consists of a glass tube, terminating mth a steel end, and having an extremely fine steel pipe, which screws on to the latter. The syringe used by the author of this work is one manufactured in England (and can be found in some of our agricultural stores), for the purpose of syr- inging plants ; it has the most accurate bore and finely-adjusted piston of any in- strument now in use, and being of medium size it can be used for either large or minute injections. Some alterations, However, have to be made in the nozzle and pipes fitted accordingly. DIRECTIONS FOR USING THE SYRINGE. In using the syringe, a certain amount of tact or experience is necessary, and the be- ginner must not feel disappointed should he fail in a first or second attempt ; for some little oversight might frustrate the whole process. Everything should be in readiness, such as ligatures, forceps, scissors, sponge, hot and cold water, etc. The pipes should be inserted into the ves- sels, and confined there by strong ligatures; and, before the syringe is inti-oduced, its noz- zle must be turned upwards, and the piston pressed until all the air and froth are ejected ; then introduce the nozzle into the stop-pipe ■-< ¥ j4^ EXPLANATION OF FIGURE IX. OSSEOUS srm 1. Frontal bones. 2. Parietal. 3. Occipital. 4. Temporal. 5. Nasal. 6. Lachrjinal. 7. Malar. 8. Superior ma.\illarius. 9. Anterior " 10. Inferior " 11. Cervical vertebrse. 16. The true ribs. 18. The sternum. 33. The scapula. 34. The humerus. c. The incisors. d. Dorsal spines. MUSCUL.YR STRUCTURE. ANTERIOR VIEW. a". Trapezius. c". Scalenus. e". PectoraKs par\'us. f". Antea spinatus. g". Postea spinatus. li". Teres major. I". A portion of the triceps extensor brachii. S. The fasehial covering of the splenius. K. The masseter. g. Levator labii superioris. f. Nasalis longus. e. Orbicularis oris. c. Dilator nai-ium lateralis. d. Dilator narium anterior. m. I. Attolentes et abducens aurem. b. Levator palpebral. a. Orbicularis palpebrarum. o". Pectoralis transversaKs. u. Levator humeri. V. Sterno niaxillaris. X. Subscapulo hyoideus. 2. Maxillary vein . 3. Jugular vein. THE HORSE. 81 and press the piston steadily until a sensible resistance is felt. If much force be used, rupture of a vessel may take place. After. a prudent force has been applied for some time, the syringe may be withdrawn, pre- viously securing the stop-cock. A steady and uniform pressure on the piston will be more likely to secure uniform injection than force, or sudden jerks. Should the first in- jection fail to fiU the vessels, it must be immediately followed by a second. When injecting through a very small pipe, the in- jector must be patient, and steadily con- tinue the pressure on the piston. When using loarm injections, the syringe must be Icept ivarm by immersing it in hot water, and the part to be injected must also be kept at the same temperature, by the same means. DIFFERENT KINDS OF INJEC- TIONS. There are six lands of injections now in use, viz., the cold, coarse, and fine injection, the minute, the mercKrial, and, finally, the plaster of Paris injection. The five first are most employed ; the plaster of Paris is objectionable because it is easily fractured. FORMULA FOR COARSE WARM INJECTIONS. Red. — Beeswax, sixteen ounces; white resin, eight ounces ; turpentine varnish, sLx ounces ; vermilion, three ounces. First liquify the wax, resin, and turpen- tine varnish, in an earthen pot, over a slow fire, or in a water bath ; then add the Ver- million, previously reducing it to a fine powder, so that the coloring ingredients may be intimately and smoothly blended, then add the same to the above ingredients, and, when they have accrued due heat, the injection is fit for immediate use. Yellow Injection. — Take beeswax, eight ounces ; resin, four ounces ; turpentine var- nish, three ounces ; yellow ochre, one ounce and a quarter. White Injection. — Clarified beeswax, eight ounces ; resin, four ounces ; turpen- tine varnish, three ounces ; flake white, two ounces and a quarter. 11 Pale Blue Injection. — Take the preced- ing ingredients, and add to them a small portion of indigo. Black Injection. — Beeswax, resin, and turpentine varnish in the above proportions ; and add lamp-black ad libitum. The same rules are to be observed in pre- paring all the injections. FORMULA FOR FINE INJECTIONS. Red. — Brown and white spirit varnishes, of each four ounces ; turpentine varnish, one ounce ; vermilion one ounce. Yelloiv. — Brown and white spirit varnishes, of each four ounces ; turpentine varnish, one ounce ; king's yellow, one ounce and a half. To make a white injection, add to the last formula two ounces of flake-white instead of king^s yelloiv. Blue. — Brown and white spirit varnishes, of each four ounces ; turpentine varnish, one ounce; Prussian blue, one ounce and a half. This may be made black by adding ivory black instead of Prussian blue. FORMULA FOR MINUTE INJECTIONS. The liquifying principle in minute injec- tions is " size," which is made in the fol- lowing mamier : Take fine transparent glue, one pound, break it into pieces ; put it into an earthen pot, and pour on it three pints of cold water ; let it stand twenty-four hours, stirring it occasionally with a stick ; then set it over a slow fire until it is perfectly dissolved ; skim ofl" aU the scum from the surface, and strain the remainder through flannel ; it will then be fit for the coloring ingredients. Minute Red Injection. — Size, one pint ; vermilion, three ounces and a half. Yellow. — Size, one pint ; king's yellow, two ounces and a half White. — Size, half a pint ; flake white, one ounce and three quarters. Blue. — Size, half a pint ; fine blue smalt, six ounces. PLASTER INJECTION. Before mixing the plaster of Paris, the pipes must be secured to the mouths of 82 ANATOMY AND PHYSIOLOGY OF the vessels at which the injection is to enter. Plaster of Paris (to which some of the preceding dry coloring materials, suitable to the fancy, can be added) must be put in a mortar and rubbed with a pestle in order to pulverize it completely ; water is then to be added until the mixture is of the consistence of cream ; the syringe being in readiness, it is to be filled and im- mediately injected into the vessels. Li the author's opinion, this injection is only suit- able for injecting first-class vessels, for it coagulates or "sete" so quickly that it cannot be used as a minute injection. It is said that a small quantity of olive oil, in- corporated with the liquid plaster, retards its coagulation ; yet if too much were added it would spoil the preparation. The moment the parts are injected the syringe should be washed out in cold water, and when the injection "sets" in the veins, the pipes must be removed and likewise cleansed. FORMULA FOR COLD INJECTIONS. Dr. Parsons recommends, for coarse cold injections, the following formula : Take coloring matter and grind it in boiled lin- seed oil, on a painter's marble, until it has acquired the consistence of common white lead, as sold at the stores. After being finely legivated, a little lime-water, in pro- portion of two table-spoonsful to a pint, is to be incorporated by stuTing. At the mo- ment of filling the syringe with the injec- tion, there should be added to it about one- thud of its measure of Venice turpentine, which should be stirred in briskly and used immediately, as it very soon hardens. For a temporary cold coarse injection, white lead ground in oU answers every purpose ; it requires no addition of lime- water, because the lead is generally adul- terated with carbonate of lime, wliich hardens the mixture, and it can be colored to suit the taste, or the vessels can be colored with a pencil brush, before varnish- ing. For filling the arteries, to dry and pre- serve, red lead is the best and cheapest material ; vennilion, however, resembles more the color of arterial blood. Whatever part we expect to inject with warm injections, must be immersed in water very hot, — not hot enough, however, to crisp the vessels. Attention to this matter is higlily important, in view of successful in- jection. THE COURSE OF INJECTIONS. Injections must follow the course of the circulation ; the arteries, however, having no valves, are easily injected in any dii-ec- tion ; but the veins are furnished with nu- merous valves, which prevent the flow of injection from the heart. Sometimes it is necessary to break these valves by means of a small whalebone probang. In the region of valves are often found coagula- tions, wliich must be washed out before the injection can be introduced ; and tliis, also, must be performed in the direction of the circulation. Small pipes are to be intro- duced, and warm water must be thrown in, which can be made to escape tlirough an incision made with a lancet in a region ap- proaching the right amicle. The incision can afterwards be closed by suture, or otherwise. Many of the veins of the horse, however, are destitute of valves, and therefore admit of injection in a direction contrary to their circulation. To inject a portion of the animal, — the knee or hock, for example, — it is necessary to secure all the branches of the vessels that have been divided where it is separated from the body. The part is then to be in- jected in the same manner as if we were injecting the whole body. QUICKSILVER INJECTION AND PREPARATIONS. The fluid specific gravity and beautiful metallic lustre of quicksilver render it val- uable for displaying minute vessels. Dr. Parsons remarks that the principal objec- tion to its general use is the continuance of its fluidity, which renders dissection, after injection, almost impracticable. Yet there are some very fine specimens of quicksilver injections of glands and deep-seated lym- THE HORSE. phatics, in the WaiTen museum of this city, tiiat cannot be surpassed by any other kind of injection. ' The same authority re- marks, that the specific gravity of quick- silver, when supported in a column, is such as to exert strong pressvn-e upon a blood- vessel or lymphatic that receives it, and therefore in some cases a syringe is unne- cessary. It is to be borne in mind that the force of the injection depends upon the perpendicular height of the column, and not on its diameter, and the former may be such as to burst the vessel. The injections should always be conducted in a shallow dish or tray, so that the quicksilver may not be wasted. When injecting the lym- phatics, it is necessary to be provided with small lancets, straight, cm"ved, and deli- cately-pointed fine needles, which must be armed with waxed threads. For common blood-vessel preparations, glass tubes of the shape of a straight blow-pipe are needed. INJECTING THE LYMPHATICS WITH MERCURY OR QUICKSILVER.* In injecting the lymphatics, our success depends, perhaps, more on the body we choose, than on any other circumstance : bodies slightly anasarcous, if they be ema- ciated, are the best. From the valvular structure of the lymphatics, it is necessary to inject from the extremities towards the trunk. It is almost impossible for one person to succeed in injecting the lymphatics without assistance ; there are so many things requi- site, besides merely holding the tube in the vessel, that an assistant is indispensable. It is very necessary, before beginning, to see that the injector has within his reach sharp-pointed scissors, knives, forceps, lan- cets, pokers for tubes, needles, and waxed threads, so arranged that they can be used instantly, for it will often happen that it will be impossible for either the assistant or the operator to take his eye for a mo- ment ofl' the vessel, without losing it. * Sir Charles Bell. When injecting the superficial lympha- tics, we first cut off a portion of skin, so as to expose the loose cellular texture ; having found a lymphatic vessel, it must be seized by the forceps and dissected from the sur- rounding substance. Having hold' of it with the forcep, snip it half across with fine scissors, and into the incision introduce the tube containing the mercury. A poker or director is often necessary for the purpose of creating a vacuum ; a few drops of mer- cury then introduced by the side of the director wiU open the way for more ; the director being withdrawn, the mercury flows into the lymphatics. If the vessel to be injected be a large one, it must be secured by ligature around the pipe. The quicksilver is to be pressed onward, elevating or depressing the pipe so as to regulate the force of the injection. In injecting a gland, we must en- deavor to find the vessel that has the most influence in filfing it. Havmg found it, we secure the other vessels and fill the gland through the former channel. , The vessels or glands injected with mercury should be dried as quicldy as pos- sible, and varnished, or else preserved in spirits of terpentine. METHOD OF INJECTING THE LACTEALS. Take a small portion of the intestine and mesentery, and make an incision in one of the most conspicuous lacteals, as near as possible to its origin in the intes- tine ; then introduce the point of the injec- ting tube, and conduct the operation agree- ably to the preceding rules. When the quicksilver flows out of any of the divided vessels, they must be repaired by an assis- tant ; when as many of the lacteals are fiUed as will receive the quicksilver from this orifice, introduce the pipe into another, and repeat the process as before, and so on, until as many of them are filled as can be ; then inflate the intestine and suspend it in the air to dry, after which it may be preserved by varnishing both inside and out. 84 ANATOMY AND PHYSIOLOGY OP DIBECTIONS FOR INJECTING THE PAROTID GLAND. This should be injected before removal, on account of the numerous vessels by which it is attached to the adjacent parts. Before commencing the operation, the sldn over the region of the gland and duct must be raised, in view of searching for the duct; having found it, an opening must be made into it with the point of a lancet, sufH- ciently large to introduce the point of the steel injecting pipe ; when introduced, con- fine the duct upon it by a ligature with a suigle knot, which shall serve when the pipe is withdi-awn tt) secure the quicksilver in the gland. The gland having been filled, the pipe withdrawn, and the duct secured, we proceed with all possible care to dissect the gland from its situation. Any branches of vessels going off to surrounding parts must be secured by means of a small curved needle, armed with a single ligature, after which they may be divided with safety. The gland being removed, and all extraneous tissue dissected off, it should be placed in water to extract the blood, etc. This wiU require about thirty-six hours ; the water, however, must be frequently changed ; the gland can then be spread on a piece of pasteboard and exposed to dry. It makes the most beautiful preparation, when pre- served in a glass vessel containing pure spirits of turpentine. BREAKING DOWN THE VALVES. Many of the glands, the surface of the liver included, can be injected contrary to the circulation of the lymph. When the quick- silver passes at fu'st freely into the lympha- tics, and suddenly stops, it will be necessary to force it forward by gentle pressure with the edge of a spatula, in the direction in which it seems most likely to run ; by this means the valves are broken down. The valves of the superficial lymphatics of the liver are easily broken down in this way, but the valves in some of the lymphatics are much firmer, consequently not so easily broken down. WET PREPARATIONS. PREPARATIONS BY -DISTENTION. Hollow organs may be distended for preservation with antiseptic liquids, air, wool, hair, cotton, plaster, quicksilver, etc. Wet preparations by distention, with spirits of wine, oil of turpentine, etc. The intention, in distending preparations by spirits, is to give them their natural figure, to exhibit more fuUy the parts of which they are composed, their vascularity, and occasionally some morbid or preter- natural appearance. METHOD OF DISTENDING AND PREPARING THE LUNGS. The lungs taken from a sheep or calf make a very good substitute for those of the horse, which are too bullvy for ordinary use. The pulmonary arteries and veins should first be filled with red-colored injection ; then immerse the lungs in oil of turpentine, con- tained in a vessel, large enough to admit them without compression; then inject into the trachea such a quantity of the above fluid as shall dilate them without danger of rupture. Then secure the trachea by liga- ture. In the same manner we proceed with other parts. K a portion only of an organ or a part of some viscera be required, we first secure the lower orifice by ligature ; inject as above, and then apply a ligature to the upper opening. It can then be suspended in spirits of wine or turpentine. ANTISEPTIC MENSTRUA FOR PRESERVING SPECIMENS. Alcohol — Spirits of Wine. — This is one of the principal fluids now in use for the preservation of specimens. It may be used of various strengths, according to the size and thickness of the specimen to be pre- served. All those that are thick and bulky should be put into pure rectified spirits ; smaller ones may require only one half the quantity of alcohol with water ; and such as are thin THE HORSE. 85 and membranous, can be preserved in com- , mon New England rum. Turpentine. — This also is an excellent antiseptic, and is highly recommended by Parsons and others, for cartilages, fibro-car- tilages, and fibrous membranes. The acids used are, sulphuric, nitric, mu- riatic, acetic and pyroligneous. Dr. Parsons states that Dr. Hayden, surgeon dentist, in Baltimore, has succeeded in preserving anatomical preparations in a superior man- ner, with pyroligneous acid. It should be rectified and diluted ^vith water. Acids, however, cannot be used when the prepara- tion contains bone. METHOD OF PRESERVING THE BRAIN. The following mixture is a very excellent menstruum for preserving the brain and nerves : Take alcohol, eight parts by weight; oxymuriate of mercury, one part. Rub the oxymuriate in a mortar, and gradually add the alcohol. The brain should remain in this mixture for twenty or thirty days, when it may be withdrawn from the liquid, dried, and varnished. METHOD OF MAKING A DRY PREPARATION OF THE AIR-VESSELS OF THE LUNGS. Throw the lungs of a horse into a barrel of water and allow them to macerate for several months, during the sUmmer season ; then, by repeated washing, cleanse the bron- chia, etc., from the parenchyma, dry, and varnish them. METHOD OF MACERATING AND CLEANING BONES. Remove as much of the flesh, ligaments, etc., as can conveniently be done with the knife ; then lay them in clean water, and change the same daily for about a week, or as long as it becomes discolored with blood. They are now to remain without changing, tUl putrefaction has thoroughly destroyed all the remaining flesh and ligaments, which will take from tliree to five months, more or less, according to the season of the year or temperature of the atmosphere. Li the ex- tremities of large cylindrical bones, holes should be bored, about the size of a quill, to give the water access to their cavities and a free exit to medullary substance. As the water evaporates from the vessel, it should be so far renewed as to keep the bones under its surface, or they wiU acquire a disagreeable blackness, and dust should be excluded by keeping the vessel constantly covered. When the white textures are de- stroyed, the bones must be scraped and again laid in water for a few days, and well washed and . scrubbed with a coarse brush ; then immerse them in lime-water, or a solution of pearlash, made with two ounces to the gaUon of water, and after a week they are to be again washed in clean water. They are then to be bleached on the seashore, where they can be daily washed with sea-water.* M. Bogros approves of the above plan of maceration, but at the conclusion of this he directs them to be boUed four hours in a strong solution of carbonate of potass, or in soap suds, adding hot water as fast as it evaporates. They are then to be washed frequently in cold water, and dried each time quickly, and then moistened (not steeped) in weak muriatic acid. The com- mon bleaching liquor in a diluted state will whiten bones, but they should not be im- mersed in it any length of time. When bleached, they may be varnished with the white of an egg.f TO RENDER SOLID BONES FLEXIBLE AND TRANSPARENT. One-half of the inferior jaw bone, or the scapula, are the most suitable bones for the above purpose. Macerate either or both until they are properly cleansed. Then im- merse in a mixture consisting of water, twenty-fiveparts ; muriatic acid, one part. If the bone is kept well covered during a period of about seven months, it will become flexible like cartilage; but as the phosphate of lime in the bone will neutralize some of the acid, a minute quantity may from time to time be added. * Pole ou " Cleansing Bones." t Parsons on "Macei-ated Preparations." 86 ANATOMY AND PHYSIOLOGY OP When the preparation becomes flexible, immerse in warm water ; then give it sev- eral washings in cold water to remove the acid ; dry, and immerse in a glass vessel of oil of turpentine ; it will assume a beautiful transparency, exhibiting the blood-vessels. METHOD OF CLEANING AND SEPARATING THE BONES OF CRANIUM. Take the head of a young colt, remove the skin and muscles, and wash out the brain, previously brealcing it down with a stick or probe ; macerate and cleanse it as before directed ; then fill the cranial cavity with dry corn from the husk, immerse it in water, and the corn as it swells forces open the suturuses, so that they can be readily separated by the hand. Wash, dry, and bleach the bones, and then cover them with colorless varnish. A BRIEF EXPOSITION OF MR. SWAN's NEW METHOD OF MAKING DRIED ANATOMICAL PREPARATIONS.* The new method has been adopted by Usher Parsons, M. D., Professor of Anat- omy and Physiology, from whose work the following selections are made : DIRECTIONS FOR MAKING DfilED PREPARATIONS. The part of a Kmb, chosen for injection, must be as free from fat as possible. A solu- tion of two omices of oxymuriate of mercury in half a pint of rectified spirits of wine, is to be injected into the arteries ; the next day inject as much white spirit varnish, to which one-fifth of white spii-it varnish has been added, and some vermilion; the limb is then to be put into hot water, where it is to remain until properly heated, when the coarse injection is to be thrown into the arteries and veins, if required, bearing in mind the course of the cii-culation ; the valves of the veins can be broken down by a whalebone probe, if necessary. If the vems are to be injected, it is better to wash the blood out of them with water before the solution of oxy- * Professor Chaussier claims to In; the origiual (lis coverer of this method. muriate of mercury is thrown into the arte- ries. After the limb has been injected, it is to be dissected. Every time it is left, and sometimes during dissection, it is advisable to cover those parts which have been ex- posed, with a damp cloth. There are great advantages to be derived from previously injecting the limb in oxymuriate of mercury, for a Limb thus injected undergoes very little change in many days, and, when the dissec- tion is recommenced, the parts will be found in the same state in which they were left, and destitute of any offensive odor. The oxymuriate of mercury is the best agent for arresting the putrefactive process. After the dissection is finished, the limb, or part, must be immersed in a solution of oxymuriate of mercury for a fortnight or more. The solution of oxymuriate of mercmy must be contained in a wooden vessel, as metaUic vessels do not answer. The limb, or part, having been in the solution during the above period, it should be taken out, di-ied, varnished, and, if neces- sary, painted. SOLUTION OF HARDENING THE BRAIN AND OTHER TISSUES. Take of oxymuriate of mercury, one ounce ; muriate of ammonia, thirty-five gi'ains ; pyroligneous acid, one pint. Rub the oxymuriate of mercury and muriate of ammonia together in a mortar, then add half a pint pyrloigneous acid. OXYMURIATE^OF MERCURY IS A VALUABLE ANTISEPTIC. Dr. Parsons relates, that, when a piece of flesh had been immersed in a solution of oxymuriate of mercury untU it was com- pletely changed, and afterwards put into a large vessel containing water for some days, though the greater part of the oxymuriate of mercury was thus washed away, it did not even then appear in the least degree putrid. I procured half of the head and neck of a large horse, which I first injected THE HORSE. 87 with the solution of oxymuriate of mercury, but as the putrefactive process was not thus sufficiently stopped, without dissecting off the sldn I immersed it in the solution of oxymmriate of mercury for several days; and, as no marks of putrefaction remained (the offensive smell being entirely removed), I then put it into a vessel containing a large quantity of water for t^vo or three days more, by which means nearly aU the solution was removed from it. I was thus able to proceed with the dissection during the hot weather, without being in the least incommoded either by the smell or soreness of the hands, and without finding the instru- ments acted upon in any degree, that ren- dered the process at all objectionable. By putting a wet cloth over it when I left it, I was further enabled to make a very minute dissection of the nerves, which I could not otherwise have done, without the use of a large quantity of spirits of wine, and then not with half the convenience and pleasure I have thus experienced. ON VARNISHES AND PAINTS. The following are the recipes for the manufacture of paints and varnishes : WHITE VARNISH. Canada balsam, spirits of turpentine, of each three ounces; mastic varnish, two ounces. Put them into a bottle and shake them together mitil they are properly mixed. MASTIC VARNISH. This may be made by putting four ounces of powdered mastic into one pint of spirit of turpentine, to be kept iia a stoppered bottle. It should be shaken every day untn the greater part of the mastic is dissolved. TURPENTINE VARNISH. Turpentine varnish is made by melting Venice turpentine over a slow fii-e, and add- ing to it as much spirits of turpentine as will reduce it to the consistence of syrup. WHITE PAINT. Three ounces of the best white paint, and one ounce of spirit of turpentine, are to be put into a bottle and shaken together. When it is used with the varnish, a bottle of each should be mixed together. PAINT FOR THE MUSCLES. This is made by grinding on a slab a small quantity of " lake" with white var- nish, to wliich one-foiu"th part of turpen- tine varnish has been added. Dr. Parsons directs that varnish should be laid on with a fine camels'-hair pencil brush, as large as occasion may require. Hollow preparations should have the var- nish poured into them, and, after turning them about in all directions, it is to be drained out as clear as possible. DIGESTIVE SYSTEM. OF THE MOUTH. It may be observed here (as preparatory to the description of this part), that, in quadrupeds in general, the facial angle is one of very considerable obliquity, in con- sequence of the prolongation of that part of the head which coiTcsponds to the face in the human subject; and this develop- ment of feature is in none more strildng than in the horse and dog. Consequently, in these animals, the nose and mouth are cavities of large dimensions. And in the horse, the mouth appears to have been thus prolonged, not only to enable him to col- lect his food with more facility, but also that he might subject greater parcels of it at a time to the action of the gi-inding teeth, whereby the processes of mastication and deglutition are gi-eatly accelerated. " Conformation. — The mouth is con- structed in part of bone, and in part of soft materials. The superior and anterior maxiOary and the palate bones form the roof; the inferior maxilla, the lower part; the incisive teeth, the front ; and the molar teeth, the sides. The lips, cheeks, soft palate, gums, and buccal membrane, constitute its soft parts. The tongue occu- pies its cavity, and the salivary glands are appendages to it. "lips. " General Conformation. — The lips, two in number, superior and inferior, are at- tached to the alveolar projections of the su- perior and inferior maxillfB, by the muscles that move them ; by the cellular tissue en- tering into then- composition; and by the membrane that lines them. Their borders sun-ound and bound the orifice of the mouth, and are united together on either side ; which points of union are denomi- nated their commissures, or the angles or corners of the month. Exteriorly, the lips are creased down the middle by perpen- dicular lines of division ; exhibit little papillary eminences upon their stuface ; and present a softer and shorter coating of hair than what is found in ordinary places, out of which project several long straggling horse-hairs or whiskers. The inferior lip is altogether smaller, and is thinner in sub- stance, than the superior ; and is distin- guished by a remarkable prominence about its centre, from which grows a tuft of long coarse hairs, vulgarly designated as the beard. " Structure. — The lips are both muscu- lar and glandular in their composition. Several small muscles,* arising from the maxillary bones, are inserted into them, and endow them with great self-mobility : one alone, consisting of circular fibres, is inter- woven in their substance without having any other connection ; this is denominated the orbicularis oris, or sphincter labiorum, from its use, which is that of closing the mouth. Tins muscle is an antagonist to all the others ; they raise or depress the lips, or draw them to one side ; but this contracts them, and occasionally projects them in such a manner that the horse can exert with them a prehensile power, which is most remarkably evinced at the time that he is picking up grain from a plain surface ; indeed, the act of nibbling our hands with his lips demonstrates this faculty, and also the force with which he can employ it. The lips are lined by the same membrane that lines other parts of the cavity of the mouth. * Percivall's Hyppopathology. (88; THE HORSE. Beneath it are seated numerous mucous follicles, that elevate it everywhere into lit- tle papilla, which are perforated by the mouths of these follicular glands, as may be readily seen with the naked eye by evert- ing either the superior, or the inferior lip. The skin covering the lips is extremely thin, and possesses considerable vascularity and sensibility. To the tenuity of it, and to the shortness and scantiness of their pilous covering, is to be ascribed the su- perior sensitive faculty of these parts. " CHEEKS. " The cheeks are constituted substantially of the masseter and buccinator muscles, covered by the sldn upon the outside, and the buccal membrane upon the inside. Their internal or membranous surface is studded with scattered mucous follicles, whose excretory orifices may be seen by everting the part. "gums. " The gums consist of dense, compact, prominent, polished masses, of the nature of periosteum, adhering so closely and tenaciously to the teeth and the sides of their sockets, that it renders the one insep- arable from the other, but by extraordinary mechanical force. Lilie other parts of the cavity of the mouth, they receive a cover- ing from the buccal membrane.- " PALATE. " Two distinct parts are included under this head ; the hard and the soft palate. The hard palate is constituted of the pala- tine processes of the superior and anterior maxillary bones ; and of a firm, dense, periosteum-lilce substance, the vaulted, in- ward part of which is elevated into several semicircular ridges, vulgai-ly called the bars. The fibres of tliis substance, which pos- sess great tenacity, are inserted into the pores of the bone in every part, but are most numerous and dense along the pala- tine suture : the interstices are filled up by a dense cellular tissue, through the sub- stance of which are dispersed the ramifica- tions of the palatine vessels and nerves. " The soft palate, sometimes called the velum palaii, is attached to the superior or crescentic border of the hard palate, the border formed by the palatine bones ; from which the velum extends backward and downward as far as the larynx, and there terminates over the epiglottis, in close ap- position with that part, in a loose semi- circular edge. In consequence of the velum palati being long enough to meet the epiglottis, the cavity of the mouth has no communication with that of the nose — these two parts forming a perfect septum between them ; hence it is that a horse cannot respire and vomit by the mouth like a human being, in whom the velum is so short that there is an open space left be- tween it and the epiglottis, through which air or ahment can pass either upward or downward. The soft palate is composed of extensions of membrane from the nose and mouth, between which is interposed a pale, thin layer of muscular fibres. " The velum performs the office of a valve : it prevents the food, in the act of swallowing, from passing into the nose, and it conducts the air from the windpipe into that cavity, without permitting any to escape into the mouth. " OF THE TONGUE. " The tongue, the principal organ con- cerned in taste and deglutition, is lodged in the mouth ; filling the interspace between the branches of the inferior maxiUa. ^^ Duplicifi/. — Like the other organs of sense, it is double ; being composed of two parts, whose union is marked by a longitu- dinal crease along its middle, the divisions having no vascular nor nervous connection nor in fact any intercommunication what- ever ; so that an animal has to aU intents and purposes two tongues, and apparently for the same reason that he has two eyes, two ears, and two nostrils. Anatomy, as far as we can carry our researches, demon- strates this ; perhaps we have no better 90 ANATOMY AND PHYSIOLOGY OP proof of it, however, than what happens in hemiplegia, a disease in wliich only one half of the body is paralytic : under these circumstances, in the human subject, the patient can only see with one eye, use one arm, and taste with but one (and that the correspondent) side of the tongue. " Division. — The tongue, in description, is commonly divided into root, body, and apex : by the attachments of the two former it is held in its situation ; the latter is loose and unconnected. " Attachment. — At its root, it is deeply and firmly inserted by several muscles which arise chiefly from the os hyoides and the inferior maxUla : it is also connected with the pharynx, and with the soft palate. From the sides of the lower jaw, separate layers of the membrane of the mouth are reflected upon its body, forming by their junction a sort of bridle, which is thence extended to the symphysis : to this part, which serves to restrain the organ in its motions, the name of frcenum linguce has been given. " Papillce. — The dorsum or anterior sur- face of this organ has a peculiar covering, which, though it appears to be continued from the buccal membrane, is a different structure altogether, and serves qiute a different piu-pose. The surface of it is roughened, possessing a villous texture, everywhere studded with numerous little conical eminences, called papilla, which are supposed to be formed out of the extremi- ties of the nerves, and to be the especial seat of the sense of taste. These papillas vary in size and figure, and are more abun- dant and larger upon the base and along the sides of the organ. Interspersed with them are a number of mucous follicles, whose apertures may be seen with the naked eye, through which a mucus is discharged upon the papillary surface, keeping it con- tinually moist, and rendering its perception of taste more acute. " Structure. — The tongue is said to pos- sess a covering of common integument ; and certainly its strong, compact tunic has aU the appearances of skin, and presents the common tests of it : the external layer is laminated, is bloodless, is insensible ; the internal or substantial part is tough, fibrous, vascular, and sensitive, in fact, is like cutis ; and the intermediate or connecting material is delicate, soft, and reticular, and forms a bed for the lodgment of the papillae. The substance of the tongue itself consists of an inter-union, or rather an incorporation, of its muscles, the fibres of which intersect one another, and take a variety of direc- tions ; but intermixed with them is a fine adipose tissue, to which is owing the flabby softness of the organ, and the peculiar aspect it exhibits when cut into. " Use. — Though the tongue is empha- tically denominated, from its essential char- acter, the organ of taste, it is not the only part that possesses this faculty ; for the palate, the pharynx, and the oesophagus, it is be- lieved, participate in it. The tongue, in addition to possessing this faculty, disposes of the food during manducation, and, when sufficiently masticated, collects and thrusts it, portion after portion, into the pharynx ; and furthermore, at the time the animal is drinking, it is not only employed as an in- strument of suction, but also as a canal along which the fluid ascends into the pharynx. " Organization. — Every part of this or- gan is plentifuUy supplied with blood. Its arteries are the lingual, branches of large size from the external carotids. The blood- vessels of either side are generally found free from anastomosis with one another ; if either of the arterial trunks is filled with injection, it rarely happens that the opposite half of the organ receives any coloring from it. Its nerves are the ninth pan, which run to the muscles, and a considerable branch from the fifth pair, in whose extreme rami- fications, which are distributed to the papillse, the perception of taste is supposed to be inherent. "OF THE SALIVARY GLANDS. " Number and Names. — The salivary glands, properly so called, are six in number, three upon each side of the head ; the EXPLANATION OF FIGURE X. OSSEOUS STRUCTURE. (SEE PEEOEBIKQ PIAIE.) MUSCULAR STRUCTURE. LATERAL VIE^V. a'\ Trapezius. • Ligamentum colli. 6". Rhomboideus longus. c". Scalenus. /" '. Antea spinatus. 9"- Postea spinatus. h". Teres major. i". Latissimus dorsi. I". m". n". Triceps extensor brachii. S. Splenius. K. Masseter. a. Orbicularis palpebrarum. c. Dilator nai-is lateralis. e. Orbicularis oris. /■ NasaKs longus. 9- Levator labii superiorus. h. Buccinator. i. Zygomaticus. J. Depressor labii inferiorus. m. Attolentes. n. Retrahentes aurem. 0. Abducens vel deprimens aurem. 2- r. Tendon of the splenius and compleius major. t. Obliquus capitis inferiorus. u. Levator humeri. X. Subscapulo hyoideus. VEINS. 1. Temporal vein. 2. Facial " 3. Branch of the jugular. 10. Parotid gland THE HORSE. 91 parotid, the submaxillary, and the sub- lingual. " The parotid, the largest of these glands, so called from being placed near the ear, lies within a hollow space at the upper and back part of the head, bounded by the branch of the lower jaw before, and the petrous portion of the temporal bone behind : it extends as high up as the root of the ear, and as low down as the angle of the jaw, by which latter a smaU portion of it is con- cealed. This gland, like the others of the same class, is enveloped in a case of dense cellidar membrane, and is constituted, in structm-e, of many little lobes or lobuli, con- nected together by processes transmitted into the interior from this cellular covering. Every lobulus is composed of a distinct set of secretory vessels, from which numerous tubuU arise, conjoin, and at length form one main branch ; these branches, which corre- spond in number to the lobuli, unite and re-unite until they end in one common ex- cretory duct. The duct emerges from the inferior part of the gland, runs along the inner part of the angle of the jaw, and crosses over the posterior edge of the bone immediately above or behind the submaxil- lary artery and vein : in the remainder of its course it corresponds to the border of the masseter, and, about opposite to the second anterior molar tooth, pierces ob- liquely the buccinator, and terminates by a tubercular eminence upon the internal slu:- face of the buccal membrane.* " The submaxillary gland, of smaller volume than the parotid, lies in the space between the angles of the jaw, to which, and to the muscles thereabouts, it, is loosely attached by cellular membrane : a portion of it is also generally found proceeding * To expose this duct, at or near its issue from the gland, an incision should be carried along the posterior border of the branch of the lower jaw : first, dividing the skin ; secondly, the panniculus ; thirdly, the cellular tissue immediately covering the duct, which is readily dis- tinguished by its glistening pellucid aspect. By extending the incision around the angle of the jaw, directing it towards the inner edge of the bone, the duct will be found making its first turn : here, however, it is lodged in a hollow, deeply buried in cellular tissue. backward as far as the trachea. Its struc- ture is similar to that of the parotid gland. The submaxillary duct issues near the centre of the gland, creeps along the under and inner border of the tongue, close to the lower edge of the sublingual gland, and terminates by a little mammiform elongation of membrane, vulgarly called the barb (bar- billon) or pap, upon the frffinum linguae, about half an inch above its attachment to the symphysis. Among the other ridiculous and mischievous practices of farriers is that of snipping off these processes. They were seemingly designed as valves, to pre- vent the insinuation of ahmentary matters into the ducts. The coats of this vessel are extremely thin and translucent. " The sublingual gland is still smaller in volume than the submaxillary, though, al- together, one much resembles the other in figure. It lies along the under part of the tongue, covered by the buccal membrane, where, from the lobular unevenness it gives to the surface, its situation is well marked. Its ducts penetrate the membrane by the side of the frsnum lingua. " The use of the salivary glands is to secrete a saline limpid fluid, called saliva ; which is conveyed and poured by their ducts into the mouth during manducation : here it is mLxed with the food, moUifying it, and rendering it more easy of digestion, and at the same time facilitating the passage of the alimentary bolus into the stomach. " OF THE PHARYNX. " The pharynx is a funnel-shaped sac, lodged in the tliroat for the reception of the food. '" Situation. — The pharynx is contiguous to the guttural pouches, superiorly; the larynx, inferiorly ; and the anterior portions of the parotid glands and branches of the jaw, laterally. Posteriorly, it is continuous in substance with the esophagus ; anteriorly, it presents an opening to the mouth. " Attachment. — In front, to the os hyoides and palate bones ; below, to the larynx ; behind, it grows narrow and ends in the esophagus. 92 ANATOMY AND PHYSIOLOGY OF " Structure. — The pharynx is in part muscular, and in part membranous. Of the muscles belonging to it the constric- tors are those that more immediately enter into its composition. They are so dis- posed as to give the membrane forming the sac a complete fleshy covering, which is rendered the more uniform by their proximate fibres being indistinguishably blended: thus the muscles form the most substantial part of the pharynx. The lining membrane, which is of the mu- cous class, is soft and thick in substance, and palely tinged with red in color, and is papillary and in places rugose upon its sur- face ; being perforated by the ducts of numerous follicles which discharge a mucus that preserves glibness and moisture to its interior. The membrane itself is (where it meets them) continuous both with the buc- cal membrane and that which lines the esophagus. " Although the pharynx is designed for the reception of the food, it does not open directly into the mouth ; the two cavities are separated from each other by the soft palate and epiglottis. Except in the act of swallowing and coughing, they have no communication : in the former case, the velum is pressed upward by the food against the posterior openings of the nose ; in the latter, the larynx is depressed by a convulsive action of the muscles in the vicinity. Into the cavity above the velum there are four openings — two of the cham- bers of the nose, one of the larynx, and one of the esophagus : the eustachian tubes do not open into the pharynx ; they end in two large membranous pouches at the upper part of the fauces. The openihg leading into the esophagiis is constantly closed, except when alimentary matters are passing to or from the stomach ; so that air received into the pharynx through the nose can pass nowhere else but into the wind- pipe; but if food be returned from the stomach, it will be regurgitated into the nose ; at least, only that portion of it which enters the pharynx at the moment that the larynx is depressed in the act of vomiting, can be thrown into the mouth, in the same way that air is in the act of coughing. " OF THE ESOPHAGUS. " The esophagus, or gullet, is the tube through which -the food is conducted from the pharynx into the stomach. " Course. — It has its beginning from the pharynx, and is there placed at the upper and back part of the larynx, taking the first part of its course above and behind the trachea, between that tube and the cervical vertebrEB. Having proceeded a short way down, it incHnes to the left, and soon after makes its appearance altogether on the left side of the trachea, and continues so placed during the remainder of its passage down the neck : this explains why we look for the bolus dxiring the act of swallowing on the left, and not on the right side of the ani- mal. In company with the trachea, the esophagus enters the thorax between the first tu^o ribs, at which place, running above that tube, it quits its companion for the superior mediastinum, which cavity it traverses below and a little to the right of the posterior aorta. Immediately beneath the decussation of the crura, the esophagus pierces the substance of the diaphragm, and enters the stomach, at a right angle, about the centre of its upper and anterior part. " Structure. — The esophagus presents, externally, a strong, red, muscular coat ; in- ternally, one remarkable for its whiteness, which in its nature is cuticular. The mus- cular coat is composed of two orders of fibres — • a, longitudinal, forming an out- ward layer ; and a circular, an inward layer : the former ^vill shorten the tube, and perhaps dilate it for the reception of food ; the latter, by successive contractions of the canal, ^vill transmit the food into the stomach. The second, or internal coat, is called the cuticular, from its analogy to the cuticle of the skin. Although it is contin- uous with the membrane of the pharnyx, it is of a totally different composition : it is thinner, but it is much more compact and THE HORSE. 93 sti'onger in its texture, and, I believe, is both insensible and inorganic. It adheres to the muscular covering by a fine cellular tissue, the extensibility of which gives full play to the latter ; and admits, during the empty or collapsed state of the tube, of the former being thrown into many longitudinal pliccE or folds ; as is demonstrated by mak- ing a transverse section of the tube : such appearances result from the contraction of the one coat, and the want of proportion- ate elasticity in the other. Between the two tunics, imbedded amongst the connect- ing cellular tissue, are numerous follicular glands, whose office is to pour forth a mu- cous secretion upon the internal surface of the lining membrane, to render the passage of food along it glib and free from any friction. " N"ASAL FOSS^.* " The nasal fossae are the frw^o chambers or lateral cavities, whose external openings are the nostrils. Their walls or external parietes are almost entirely osseous; and to the OSSEOUS SYSTEM (page 45) the reader must turn for a description of the manner in which the fossae are formed, and of the bones entering into their formation. But, in addition to bone, they are cartilaginous in their constitution. " The cartilages of the nose are five in number: — of which one (the septum nasi) is situated internally; the other four (enter- ing into the composition of the nosti'ils) ex- ternally. " The septum nasi is the vertical carti- laginous partition interposed between the nasal fossas. It exhibits four borders. The inferior one is received into the groove of the vomer ; while the superior presents a lengthened channel between two elevated edges, into which is admitted the internal crest formed by the union of the nasal bones. Its posterior border is affixed to the ethmoidal plate : its anterior serA'es to sus- tain the cartilages forming the nostrils. Both its sides are completely covered by the Schneiderian membrane. * Hippopathology. ^'■Nostrils. — Four in number: two on each side, distinguished by the epithets true and false. " The true nostrils are the large, ovoid, and ever-open orffices so conspicuous ex- ternally. They have for the base of their structure four pieces of fibro-cartUage, which are involved in doublings of the common integument. Each nostril is formed of two flexible ala or wings : a svperior or internal one, and an inferior or external. The former is supported by a broad circular cartilaginous plate ; the latter is crescentic in shape, and forms a flexm-e outward, within which is perceptible the orffice of the lachrymal duct. They are attached to, and supported by, the nasal peak and sep- tum nasi. " The false nostrils are tv\-o little pouches or cavities (ha\Tng the semblance of culs- de-sacs), situated internally, above the trae nostrils, into which an external opening is found within the commissive formed by the union of the two alae. They are formed out of duplicatures of the skin, which is here thiimer, and finer, and softer in its textiure ; and, except at their enti-ance, are without hair upon their surfaces. Their use is not luiown. ^^Schneiderian membrane. — The cavity of the nose is not only divided into the tw'o nasal fossffi, but each fossa is subdivided into the thi'ee meatus (for a description of which, vide page 45). Every part of these cavities and jDassages is covered by the Schneiderian or pituitary membrane. This is a membrane of the mucous class, dis- tinguished for its thickness of substance, for its vasculai-ity, and for its olfactory pa- pilla;. It has two sm-faces: an exposed or secreting one and an unexposed or ad- herent one. The secreting surface is smooth, and is rendered glib and sliiny by the varnish it derives from the mucous secretion emitted by the numerous small rounded pores everyw'here visible in the membrane, but more particularly upon the lower part of the septum, and upon the inferior tin-binated bone. This sm-face exhibits a pale pink blush, the effect of the 94 ANATOMY AND PHYSIOLOGY OP bloodvessels spread over it, which are here so superficial as to owe their principal de- fence to the mucous exudation : hence it is that the complexion of the membrane (varying with the influence of the atmos- phere and other agents) is extremely fugitive and uncertain. The adherent surface of the membrane contracts a close and firm adherence to the parts it covers, through the insinuation of its fibres into them : in- deed, to the bone it appears to supply the place of periosteum ; to the cartilage, of perichondrium. The substance of the membrane exhibits a fibrous structure, in- terwoven with cellular tissue ; and upon that — as a substratum — is spread a glan- dular and vascular apparatus, from wMch issues the mucous secretion ; together with numerous papiUcB, of small size, constituted of the terminations of those nerves from which the membrane derives ordinary sen- sation, as well as those that endow it with the peculiar sense of smelling. The Schneiderian membrane, inferiorly, within the nostrils, is continuous with the dupUca- tures of skin linuig those parts ; superiorly with the membrane lining the pharynx; be- sides which, it is continued into the several sinuses of the head, through the openings leading from them into the nose, and like- wise gives them a complete covering : it is to be obser\'ed, however, that in the sinuses the membrane is thinner, and assumes a paler and more delicate aspect; its natural secretion is also fomid more sparing. The membrane is abundantly supplied with blood-vessels, as well as nerves ; and also possesses its share of absorbent vessels. Its arteries, wliich ramify and anastomose so as to form a spreading network upon the secreting surface, are derived superiorly from the lateral nasal; inferiorly from the facial and palato-maxillary. Its ner\'es are furnished by the fii-st and fifth pairs. " Sinuses. — These cavities are formed in the interior of several of the bones of the cranium and face : in fact, with the excep- tion of the membrane lining them, they are entirely osseous in their composition. This will account for their description having been already given (at page 46), to which we must again refer. " Ducts. — There are tu^o ducts belonging to, or connected with, the nose. One is the ductus ad nasum — a tube partly osseous and partly membranous in its composition, commencing at the inner angle or corner of the eye, within the substance of the lachry- mal bone, running within a' canal continued from this bone through the superior maxil- lary bone, and terminating at the inner and inferior part of the nasal fossa, underneath the duplicatm-e of the inferior ala, upon the surface of the common skin, about one- fourth of an inch fi"ora its junction with the Schneiderian membrane, by an orifice large enough to admit a crow-quill. The other duct is the ductus communis nariiim, which pursues its course along underneath the vomer to the,pharynx ; after arising from two lateral branches springing from oblong apertures in the floor of the nostrUs." INTERNAL PARTS. COMPREHEXDIXG THE CA^TTIES OF THE CEAXIl'M, DE- BIT, NOSE, AXD MOUTH. I. CAVITY OF THE CRANIUM, Constructed for the lodgment of the brain with its appendages, is in form ovoid, flattened inferiorly, broader anteriorly than posteriorly; its antero-posterior or long diameter measming about seven inches ; its transverse or lateral diameter about four inches ; its vertical or perpendicular diameter about three and a half inches. At the same time it is to be observed, that, although the general form of the cavity is the same, its di- mensions may and do vary in dift'erent heads. The eight bones composing the cranium all present internally surfaces more or less con- cave, which, united, form the cavity under consideration ; hence it is that the interior is not regular or uniform, but presents to view diflerent hollows, which are adapted to distinct prominences of the cerebral mass. Division of the interior surface into roof and base of the cranium : The roof is formed by the frontal, parietal, and occipital bones : its superficis is larger THE HORSE. 95 than the extent of the base, and it is with- out any apparently defective places, observ- able in the latter. It presents — 1st. On the mesian line from front to back, the sagit- tal groove, for the longitudinal sinus formed by the frontal and parietal crests, crossed towards the front by the coronal suture, and bounded posteriorly by the parietal protu- berance, to which is attached the tentorium, and behind which is the occipital cupula, for covering the cerebellum. 2nd. On either side, along the same line, the cerebral con- cavities of the frontal bone; the coronal suture, the boundary line between them and the parietal concavities ; the transverse grooves, for the lateral sinuses ; and, sunk within them, the lambdoidal suture. The base is formed by the temporal, sphenoid, ethmoid, and occipital bones. It presents — 1st. On the middle line, from before backwards, the crista galli, and on its sides the ethmoidal fossa and cribriform plates, bounded laterally by the internal orbital plates of the frontal bones, and there pierced by the internal orbital foramina ; the concave surface of the body of the ethmoid bone; the optic hiatus leading to the optic foramina; a transverse suture between the ethmoid and sphenoid bones. Upon the sphenoid bone, the pituitary fossa, bounded laterally by the two optic fossae ; the latter leading to the foramina lacera orbitalia, over which are the spinal foramina ; a trans- verse elevated line denotes the place of junction of the sphenoid with the occipital bone. Belonging to the occipital bone, are the basilar fossce and the occipital hole. 2d. On either side, in the same direction, the inter- nal surface of the icing of the ethmoid bone, rather more convex than concave, for the support of the anterior lobe of the cere- brum ; the concavity of the iving of the sphe- noid bone, for the reception of the middle lobe ; the concavity of the squamous part of the temporal bone, for lodging the posterior lobe : and the sutures bounding these three cerebral surfaces. The foramen lacermn basis cranii, formed between the wing of the sphenoid anteriorly, the basilar process of the occipital bone internally, and the petrotis portion of the temporal bone exter- nally and posteriorly : it is wide and irregu- lar before, narrow behind, and is distin- guished into the spheno-occipital and tempo- ro-occipital hiatus. The petrous portion of the temporal bone, presenting a naiTow triangular surface forwards and upwards, which contributes to the posterior cerebral concavity ; a broad, smooth, but uneven sur- face inwards, against which inclines the cerebellum, and upon which we distinguish — a, the orifice of the meatus auditorius in- ternus ; b, a transverse prominence, and sev- eral cerebral indentations ; c, an irregular convexity downwards, which forms the boundary wall of the labyrinth ; d, 2i fissure separating it from the former. Lastly, the sutures, uniting the petrous to the squamous portion and to the occipital bone. Of the occipital bone a part of the internal surface assisting in the formation of a concavity for the cerebellum, by the convolutions of which it is indented ; the surface even and smooth, and slightly excavated below this, for the support of the medulla oblongata ; stiU lower, the condyloid foramina, through which the ninth pair of nerves pass out. ■THE ORBITS, Two in number, are formed for the lodg- ment, attachment, and protection of the eyes and their appendages. Figure. — Symmetrical. The cavity, which is extended horizontally backward and inward, has, viewed in front, a pyra- midal aspect : the base, represented by the front, has four sides, and four angles ; one only of the sides, however, is sufficient in extent to reach the apex, the others being all more or less imperfect. A line drawn in a horizontal direction through the axis of this figure, inclines more outwards than forwards, more forwards than downwards, intersecting another horizontal line projected directly forward at an angle of about 70°, and one extended laterally, directly outward, (at right angles with the former), at about 20° : the incUnation downward, however, will in course vary with the erect position of the head. 96 ANATOMY AND PHYSIOLOGY OF THE HOESE. Structure. — The orbit is composed of unequal portions coming from four of the bones of the cranium, and from three of those of the face : viz., the frontal, ethmoid, sphenoid, and temporal bones; the malar, lachrymal, and palate bones. Division — Into sides, angles, base, and apex. Sides. — The superior side or roof of the cavity consists only of the frontal arch; which is concave and smooth internally, to make room for the lachrymal gland, and has anterior and posterior borders, sharp and slightly curvated. The inferior side or floor of the orbit is formed by the orbital surfaces of the lachrymal and malar bones, is broader than the roof, though, like it, is deficient as a whole. It comprises the orbital portion of the lachrymal suture : it is terminated in front, hy a smooth, rounded, curvated border; behind, nearly midway between the base and apex, by a shorter and straighter border. The internal or nasal side, the broadest and ordy complete one, is formed principally by the internal orbital process of the frontal bone, into the notch of which is received the QS planum: the ethmoid bone further con- tributes, and also the sphenoid and palate bones, the three constituting that irregular termination of the cavity behind which represents the apex. The frontal orbital plate is smooth and slightly concave, and is united below by a continuation of the trans- verse suture with the lachrymal bone. Its border in front, though slightly cui-vated, is very irregular, having several notches and one or two small foramina in it; it also presents a little tubercle, to which the lach- rymal caruncle is attached. The external or zygomatic side is formed principally by the zygomatic process of the malar bone, that of the temporal conti-ibuting but little : it is concave, and smooth internally, somewhat broader below than upwards ; is intersected obliquely by the zygomatic suture, and has an interior border, smooth and curvated, a posterior one, sharp and straight. Angles. — The supero-internal angles, one before, the other behind, are formed by the beginning of the frontal arch, through which, midway between them, passes the supra- orbital foramen. The infero-internal angle includes the lachrymal fossa. The supero- external angles, one anterior, the other pos- terior, are intersected by the suture uniting the frontal and zygomatic arches. The infero-external angles, particularly the ante- rior, are rounded and smooth. Base. — Of the circmuferent border, the superior and internal parts, about two-fifths of the entire circle, are formed by the os frontis ; the inferior and internal parts, about one-fifth, by the lachrymal bone ; and the remaining two-fifths by the malar and tem- poral bones, in the proportion of three parts of the former to one of the latter. The apex or back of the orbit, formed by the ethmoid, spenoid, and palate bones, is pierced by five foramina: the two round are the internal orbital and optic, which are ranged in a row with two oval and larger in size, the supero-posterior and infero-pos- terior orbital; the one behind is the spinal foramen. in. CAVITIES OF THE NOSE, Comprehending the nasal fossae or cham- bers, and the sinuses. These cavities occupy about two-thirds of the internal space of the superior maxilla, the remaining third belonging to the cranium ; from which they are partitioned by the cranial septum of the frontal bone, in union with the cribriform plates and crest of the ethmoid. The nasal fosste may be said to include about two-thirds of the entire space de- voted to the olfactory cavities. They con- stitute the interior of the proboscis ; have four boundary walls, one above, one below, and two laterally ; are separated from each other by a septum ; but are open both be- fore and behind. The superior wall presents an irregular concave formed by the internal surfaces of the nasal bones, the cells and grooves of the ethmoid, and small portions of the nasal surfaces of the palate bones. The inferior wall is horizontal ; it extends forward beyond the superior, but is con- siderably overreached by that waU poste- THE HORSE. 97 riorly : it is formed by the palatine por- tions of the anterior and superior maxillary, and by the palate bones. The surface is transversely concave, and presents a slight eminence a little behind its middle. Each lateral wall or side presents an irregular concavity, and is formed by the anterior and superior maxillary and the palate bones. To it are attached the supe- rior and inferior turbinated bones, by which the fossa is divided into three separate pas- sages or meatus. The superior meatus, comprised between the nasal and superior turbinated bones, extends from the angle of the lateral nasal opening, passing over the ethmoidal cells, to the cribriform plate, fol- lowing superiorly the declination of the wall. The middle meatus, included between the turbinated bones, leads superiorly into the ethmoidal grooves and cells, and into the sinuses of the head, and ends below, be- neath the termination of the superior. This passage, lilvc the former one, is narrow ; but its greatest diameter is, obliquely, in the perpendicular direction ; whereas the other measures most from side to side. It re- ceives the apertures of the ductus ad nasum, maxillary sinus, ethmoidal grooves, and turbinated cells. The inferior meatus is the most capacious as well as the most direct one: it extends along the inferior wall, from the anterior to the posterior opening of the nose. The septum nasi is the partition separat- ing one fossa from the other. It is formed, posteriorly, by the ethmoidal plate; infe- riorly and posteriorly, by the vomer ; supe- riorly and anteriorly, (and principally) by a broad perpendicular plate of cartilage. The openings of the nose are : the ante- rior, divided by the nasal peak and septum nasi into two, and formed by the superior borders of the anterior maxillary bones: the posterior, divided after the same manner by the vomer and septum, and formed by the nasal surfaces and crescentic borders of the palate bones. The sinuses of the head communicate with, and may be said to constitute part of, the nasal cavities. They are the frontal, nasal, maxillary, sphenoidal, ethmoidal, and palatine. The frontal sinuses, formed within the frontal bones, are situated so that a straight line extended between the supero-internal angles of the orbits passes opposite to about the angular or deepest parts of then- cavities. The sinus (on either side) has a triangular figure. The superior side or roof is flat, and (barring the septa) even upon its surface ; whereas the posterior side is irregular, being convex inwardly, where it is formed by the cranial septum ; concave outwardly, where it is opposed to the part composing the temporal fossa. The infe- rior side slants from behind forward, and from below upward, is irregular on its sur- face, and open or deficient outwardly, where the cavity communicates with the maxiUary sinus. Of the angles, one is directed upward; another downward, ter- minating in the nasal sinus, with which it is conjoined, the two forming one continu- ous cavity ; the third points backward, and is directly opposite to the imaginary trans- verse line above alluded to. The cavity is traversed and divided into several unequal open compartments and recesses by septa; the principal of which is one extended be- tween the superior and inferior sides ; it is partitioned from the opposite sinus by the nasal spine. The sinus is but small in the young compared to its proportionate dimen- sions in the adult subject : it continues to increase afterwards with age, and ultimately extends throughout the whole of the frontal bone. The nasal sinuses, formed by the nasal bones above and the superior turbinated bones behind, are nothing more than the culs-de-sacs or blind terminations of the frontal sinuses. The maxillary sinuses, the largest of these cavities, are spacious but very irregu- larly formed. They are situated below and in front of the frontal. Of this sinus, on either side, the posterior and external walls are formed by the malar and lachrymal bones, whose orbital processes constitute a tliin partition between it and the orbit ; the 98 ANATOMY AND PHYSIOLOGY OF inferior parts consist of the excavations in the superior maxillary bone ; superiorly, the sinus is open, being there continuous with the frontal : the boundary line between these cavities is marked by the suture uniting the lachrymal to the frontal and nasal bones on the outer side, and by the prominent crest formed by the junction of the superior turbinated with the ethmoid bone on the inner ; underneath which pai-t, through a curved (and in the recent subject sort of valvular) fissure, the sinus opens into the middle meatus, between the bases of the tiurbinated bones. The cavity is but small, and still more irregular, in the young subject, in consequence of the intrusion of the yet uncut molar teeth. The frontal sinus, then, terminates in the nasal, but both discharge themselves into the maxillary ; the maxillary has also a blind termination, but empties itself into the pos- terior part of the middle nasal meatus. The sphenoidal sinus is situated within the palatine portion of the body of the sphenoid bone. It has no existence in the young subject, the bone being solid through- out ; but in process of growth a cavernous hollow is formed, which, from the seces- sion and attenuation of the laminae of the bone, continues to enlarge. It communi- cates, by two ovoid openings, with the eth- moidal sinuses. The ethmoidal sinuses are two cavities, separated by the perpendicular plate, situa- ted beneath the ethmoidal cells. They have openings in front, communicating with the lowermost and largest grooves of the same bone, and with the palatine sinuses. The palatine sinuses are formed between the superior maxillary and palate bones ; are situated below and in front of the for- mer ; are separated from each other by the vomer ; and open into the maxillary sinuses : they are irregidar in form and cavernous interiorly. They are not to be found in the young subject. Some might be inclined to treat them as parts of the maxillary sinuses ; they are, however, as perfectly distinct from the latter as the frontal are. IV. THE MOUTH. The mouth is the cavity included be- tween the superior and inferior maxillae, making (in the skeleton) one common va- cuity with the inter-maxillary space. Its antero-posterior dimensions can be but little varied; but its supero-inferior diame- ter will be increased in the ratio of the dis- tance to which the inferior maxiUa recedes from the superior; the cavity during the distraction of the jaws assuming the figure of a misplaced t> , the angle of which is turned baclrward. The mouth is formed — superiorly, by the palatine and superior and anterior max- illary bones ; interiorly, by the inferior maxilla ; laterally, by the molar teeth ; an- teriorly, by the incisive teeth. Behind, through the posterior opening of the nose, it communicates with the nasal fossae. PERITONEUM.* The whole of the viscera contained within the abdomen proper, including the anterior part of the rectum, bladder, and vasa deferentia, are either entirely or par- tially covered by or in contact with perito- neum. This is a serous membrane reflected also over the parietes of the abdomen, so that a parietal and visceral or reflected por- tion require notice. Like other membranes of the same nature, it forms a closed sac, which, however, is not the case in the female, as its cavity communicates with that of the uterus, owing to the open state of the Fallopian tubes at their fimbriated edges. It is loosely connected with the abdominal parietes by subserous cellular tissue, and the same obtains with regard to its connec- tion with the viscera. But we find some parts more adherent than others, such as along the linea alba and cordiform portion of the diaphragm. Also on the organs it is but loosely connected with them at their attached border, where it forms generally a triangular space, occupied simply by vessels, nerves, and cellular tissue, and allowing of * Prize Essav bv Mr. Gamgee. THE HORSE. 99 their distention and alteration in figure. On the other hand, it is more adherent as it extends over the free surface or margin of the various parts it is in contact with. The peritoneum being considered as ex- tending from the .umbilicus over the ab- dominal parietes towards the median line of the diaphragm and spine, is found there to fold on itself, and proceed from tlie latter on to the intestine, forining the mesenters ; and from the former on to the liver and stomach, constituting ligaments. These folds of peritoneum are also seen extending from organs to other parts of the abdominal parietes, and these also constitute ligaments. Then they may be traced from one organ to another, giving rise to the several omenta ; all of which we shall more especially allude to as we speak of the peritoneal coat of each separate viscus. STOMACH. The stomach is the dilated portion of the alimentary canal, intermediate between the oesophagus and small intestine : through the former it receives the ingested aliment, for which it acts as a reservoir during the process of chymification, the active agent in which is the gastric secretion. In the horse, as well as all other soli- pedes, this viscus is exceptional in not being the most capacious dilatation of the alimen- tary canal. M. Colin, in a paper published in the Recueil de Medecine Veterinaire Pratique for Tune, 1849, states that the capacity of the horse's stomach is very variable. He says, that in a very small horse he found it only nine quarts (accord- ing to his evaluation by litre, which may be considered as thirty -four fluid ounces), while in one of colossal dimensions it was as much as 33 3-4 quarts, both having died at the college ( Alfort) infirmary. He gives the average as being from 13 7-20 quarts to 14 3-5. Then, considering the capacity of the stomach in relation with that of the intestines, he found it in a very small horse as one to thirteen, while in other two cases it was as one to ten. He takes the latter as the standard relative capacity between the two. The stomach is situated transversely to the long axis of the body, in the left hypo- chondrium, extending into the epigastrium and during repletion into the right hypocon- driac region. However, its size and situa- tion vary under different cfrcumstances, as to whether it be full or empty, adapting itself generally to its contents. The stomach is fixed on its left side to the diaphgram by the oesophagus, having the spleen attached to it as well. The duodenum then, by means of the lesser or gastro-hepatic omentum, suspends the pyloric end by getting attached to the con- cave surface of the liver. The shape of the stomach might be ex- pressed as bemg that of a tube bent on itself, and dilated along its convex border, so as to form two cul-de-sacB ; i. e., a right and a left one, whilst it has two borders or curvatures, distinguished as a lesser concave and a greater convex one. The stomach has two smooth surfaces, the anterior one being in contact with the liver and dia- phragm, whilst the posterior one corresponds to the convolutions of the small intestines and gastric flexture of the colon. It has two orifices, i. e., a left oesophageal, or com- monly called cardiac, and a right intestinal or pyloric one ; the latter taking its name from the valve by which it is guarded. A circular depression round the stomach, midway between the cardiac and pyloric orifices, most visible when the organ is replete, marks the external division of the stomach into a cardiac and pyloric portion, coiTCsponding with the point where the mucuous membrane varies in character in- ternally. The sacular projection at the cardiac portion takes the name of fundus, owing to its greater magnitude as compared with a smaller cial-de-sac at the pyloric end, the analogue of which in human anatomy is characterized by the appellation of antrum pylori. Having thus briefly described the striking peculiarities of the stomach, I proceed 100 ANATOMY AND PHYSIOLOGY OP with more detail to the consideration of its constituent parts, such as its coats, nerves and vessels. The coats of the horse's stomacli having been generally described as four, it appears needless to altertheir nomenclature, although the one which I shall allude to as third might quite as justly be described as second, or merely spoken of as connecting cellular tissue, without regarding it as a separate coat. The external peritoneal tunic is found proceeding from the diaphragm on to the cardiac portion of the stomach, surrounding the oesophageal opening, where it is tough, and forms the gastro phrenic ligament. Thus we follow it on to the corresponding surface of the viscus, and, firstly, more especially on to the lesser curvature, where it is loosely connected with the other coats, and, the middle portion being more adherent, gives rise to two folds laterally, which seem to stretch from the cardiac to the pyloric orifices, to bind the two together, necessarily leaving a pit or cul-de-sac between them. At the pyloric end the peritoneum comes off from the concave surface of the liver on to the stomach, constituting the gastro-hepatic or lesser omentum, the anterior layer of which comes from the anterior part of the concave surface of the liver, whilst the posterior layer comes from the posterior part of the same, so that the two enclose the vessels going to and from the porta. Having formed a covering to the corres- ponding surface of the stomach, the layers of peritoneum meet at the greater curvature. In following them from this point the des- cription will be facilitated by alluding to the two separately, as they meet to form the gastro-splenic and gastro-colic omenta, as well as the omental sac. In forming the latter, they so blend as merely to con- stitute a fine reticulated vascular layer, in- separable into two, except near the magins of the viscera. Distinguishing the anterior or external layer as A, and the posterior or internal one as B, their arrangement admits of exposition in the following terms : — A passes from the anterior surface of the stomach, forms the loose omentum, and gets on to the transverse colon and spleen. Reaching the latter, it is reflected over its superior surface at the posterior margin of the hilus, so as to contribute to the form- ation of the gastro-splenic omentum, and extends round the free posterior margin of the viscus on to the inferior surface, passing to the right on to the left kidney, and, anteriorly reaching the supero-anterior part of the spleen, is reflected from it so as to continue as the outer layer of the loose omentum. Further to the right, A is trace- able on to the inferior surface of the trans- verse colon, and, extending round the pos- terior part of the latter, is found to ascend up to the spine, and then turn backward and downward to form the mesentery. B, or the internal layer of peritoneum, passes from the posterior surface of the stomach till it reaches the infero-anterior border of the transverse colon, as well as the hilus of the spleen. After covering the an- terior surface of the colon, it ascends up to the pillars of the diaphragm clothing the anterior part of the pancreas, which is thus held between A and B, or layers of the trans- verse meso-colon. A little to the left of this, B passes on to the anterior margin of the hilus of the spleen, forming the inner or pos- terior layer of the gastro-splenic omentum. From this arrangement it results that the peritoneum, in forming the lesser or gastro- hepatic, the greater or gastro colic, and the gastro-splenic omenta, closes in a space termed the omental sac, the interior of which is inaccessible except by an opening at the posterior part of the gastro hepatic omentum, whose free margin at the right side marks the point where it may be pene- trated ; this passage is termed the foramen of Winslow. It is bovmded anteriorly by the lesser omentum, above by the hver, and posteriorly by the transverse colon. Thus, supposing the inner layer of the omental sac to be separable from the outer, and drawn out through the foramen of Winslow, the following parts would be de- prived of peritoneum, i. e., the posterior surface of the stomach, the gastro splenic omentum of its posterior layer ; so that the / EXPLANATION OF FIGURE XI. MUSCXJLAE STEUCTUEE. a". Trapezius. 6". Rhomboideus longus. S. Splenius. c". Scalenus. e". Pectoralis parvus. f". Antea spinatus. ff". Postea .spinatus. /*". Teres major. i". Latissimus dorsi. J". A portion of the serratus magnus. k". " Humero cubital." I", m". n". Triceps extensor brachii : magnum, medium, et parvium. o". Pectoralis transversus. p. Flexors. q". Flexor metacarpi extemus. r". " " internus. s". Extensor metacarpi magnus. x". Extensor pedis. a'. Serratus. c'. Obliquus extemus abdominis. d'. Obliquus internus abdominis. g'. Region of the patella. 7j'. i. Glutei muscles. m'. Tensor vagina. n'. Rectus. o'. Vastus extemus. q'. Flexor metatarsi. r'. Gastrocnemius externus. t'. Flexor pedis accessorius. u. Sterno maxillarius. V. Internal part of the levator humeri. J/'. Peroueus. x'. E.xtensor pedis. 33. Radius. /'. Triceps. VUINS. 3. Jugular vein. 4. Subcutaneous thoracic vein. 6. Saphena vem. 6. Radial vein. D. Serratus magnus muscle. d. Dorsal spines. /. Ubar. e. e. Fibula. la. True ribs. 17. False ribs. 18. Sternum. 19. Ileum. 22. Femur. 23. Patella. 24. Tibia. 34. Os humeri. 35. Radius. OSSEOUS STRUCTURE. THE HOBSE. 101 vessels going to and from the stomach and spleen would remain uncovered, the anterior part of the transverse colon, the anterior surface of the pancreas, and inner or pos- terior layer of the gastro hepatic omentum. Next to be described to the serous coat is the muscular one, which is constitfited of involuntary plain fibres, whose thickness is verj- variable in different subjects, as well as in different parts of the same stomach. The cardiac end is more muscular than the pyloric, except at the right margin of the latter, where it is verj' powerful and thick, as it surrounds the pylorus. The thinnest part of the stomach is unquestionably the convex border of the lesser cul-de-sac. The muscular coat of the stomach is in- tricately arranged, and authorities differ vastly from each other in the description of the several layers constituting it. The number of layers entering into its compo- sition is three : the outer and inner ones are mostly continuations of the inner layers of the oesophagus, while the middle one is pro- per to the stomSch. The outer layer is composed of the longitudinal fibres of the oesophagus: as these reach the cardiac end of the stomach, they form a peculiar turn, whereby the dis- tribution on the surfaces as a flat layer is facilitated. Some of the fibres of this layer dip down to join the deeper ones, while others continue onwards as the longitudinal fibres of the duodenum. As to the fibres which proceed on to the cur\-atiires, they are not so intricate, as they descend directly from the portion of the oesophagus opposite the part they supply, so that the only alter- ation in direction is that of diverging a lit- tle from each other, and pmrsuing the bent course of the corresponding gastric curva- ture. On the lesser one they soon become scanty, and are lost in the circular fibres of the body of the stomach : very few of them are traced on to the pylorous. The fibres proceeding on to the greater curvature are mingled with other considerable bundles taking the same direction, but which are not traceable on to the oesophagus, as they seem to pass round each side of the cardia, and blend with the circular fibres on the lesser curvature. The middle layer consists of annular fibres, which, though scanty as they encircle the extreme left end of the stomach, increase in bulk towards the middle part of the organ, and are especially developed at the lesser curvature. They again decrease over I the antrum pylori, but are ultimately gready j developed for the formation of a powerful sphincter at the pylorus. The internal or oblique fibres of the stomach have somewhat the same arrange- I , ment as the deep layer of fibres of the : oesophagus, although not perfectly identical, as they are arranged like hoops placed one I within the other ; but while in the former j the one set enters the other without inter- section, in the latter there is a partial decus- ] sation by separate bundles. Thus, in real- ity, the oblique fibres of the stomach are ' constituted of two layers, the one proceed- 1 ing fi"om the left end of the stomach on to \ the right, which pass internally to the next I layer : this one proceeds fi'om the right of I the cardia on to the fimdus. Ow"ing to the scantiness of circular fibres at the base of each cul-de-sac, the fibres are here in con- tact with the superficial longitudinal ones. I The oblique fibres are best studied by dis- secting firom within, and, after removing , these, the circular fibres come into ^■iew [with greater ease than by attempting to j expose them firom without. The third coat of the stomach consists merely of the cellular tissue existing be- tween the muscular and mucus coats, as well as connecting the former to the outer serous tunic, in which case it is more abimd- ant and firm nearest the curvatures. There it is situated between the muscular and mu- cus coats : it was named by the ancients, on account of its white aspect, the Tunica Nervosa. It is loose in some parts and firm in others ; not only serving to connect parts together, but also to form a medium in which vessels ramify for the supply of the organ. The internal or mucus coat of the stom- ach differs in the cardiac fi"om the pyloric 102 ANATOMY AND PHYSIOLOGY OP end, as in the former it is but a mere con- tinuation of the unmodified mucus lining of the oesophagus, being characteristic for its scantiness in gland and but limited sup- ply of blood. The most marked feature it possesses is that of being covered by a cuti- cular layer of extreme thickness, easily separable from the basement structure be- neath after slight maceration or boiling. The cardiac portion of the gastric mucus lining is, in a healthy stomach, of a dirty white, bedewed by more or less mucus, and thrown into folds which have a radiated arrangement at the cardiac orifice, whilst at the fundus they are concentricaUy arranged. This portion of the membrane is also fur- nished with papUlae ; and Sprott Boyd, in an Inaugural Essay on the structure of the Mucus Membrane of the Stomach, pub- lished in the Edinbitrgh Medical and Surgi- cal Journal for 1836, describes a very marked peculiarity of an interposed layer between the epithelium and papiJlated surface of the mucus fining. This intermediate layer, he says, has a smooth equal surface, perforated by numerous foramina about the 600th of an inch in diameter, or perhaps a little smaller, the margins of which are sUghtly thickened. He afterw^ards states that he has not been able to trace in the epithefium of any other animal a structure similar to that existing in the horse. These peculiar- ities in the left pouch of the stomach cease abruptly midway the length of the viscus, where the cuticular lining terminates by a serrated edge. The mucus fining of the right end of the stomach is normally of a reddish color, and presents a villous, gfistening aspect, coated thicldy with mucus, and also pos- sessing a high degree of vascularity ; the epithefium is here scanty, but nevertheless tabular. The villous appearance above referred to suggests itself also when the sur- face is examined by the naked eye and by the aid of a lens ; but it is deceptive, as has been already remarked by Sprott Boyd, who correctly refers it to the raised margins of the arolae which stud the surface. This portion of the gastric mucus membrane is also thrown into folds, which become grad- ually more marked towards the pylorus; whereas they are susceptible of obfiteration by distention, there is one circular fold at the pylorus which is permanent, and so dis- posed as to fulfil the office of a valve. The' arteries of the stomach are derived from the coeliac axis, whose three divisions, i. e. gastric, hepatic, and splenic, all contri- bute to supply blood to the viscus ; but the first is specially destined to that office. The gastric artery, being the smallest of the three divisions, takes a course downwards, forwards, and rather to the right, across the pancreas, getting between the layers of the gastro-hepatic omentum. Being then di- rected to the left towards the lesser curva- ture, it divides into an anterior left or smaUer branch, and a posterior right and more capacious as weU as longer one. The an- terior division is destined to supply the anterior surface of the stomach, and more especially the left cul-de-sac, anastomos- ing with branches (sometimes called vasa breva), coming on to the stomach from the splenic. This division of the gastic also anastomoses with oesophageal twigs, which are occasionally of considerable size. The posterior or right division of the gastric artery, destined for the pyloric end of the stomach, anastomoses with some splenic branches, but more especially with the py- loric branches of the hepatic artery. The veins returning the blood from the stomach are the gastric and splenic, which anastomose with the duodenal veins. These all have a few valves, but they may be easily injected from the porta into which they empty, owing to their very free anastomosis. The lymphatics of the stomach are nu- merous, and in some parts very apparent, entering the lymphatic glands situated along the greater cruvature and around the cardia, where they are numerous and large. The stomach is supplied with nerves from both the cerebro spinal and sympathetic or ganlionic system. The pneumogastric or par vagna nerves, arising from the me- duUa oblongata, are the main conductors of nervous influence to and from that vis- THE HORSE. 103 cus. Their arrangement is simple, as, after they have formed various plexuses •within the thorax, in which they mutually inter- change fibres, they reach the diapluagm, and here are arranged as two nervous branches, /. e., a superior and an inferior one. The former is principally destined for the fundus, whilst the latter supplies the pyloric end, and sends branches off to the duodenum, with one or two to the solar plexus. The sympathetic fibres, destined for the stomach, are derived from the solar plexus, descending on to the viscus, in company with the vessels. INTESTINE. This term is applied to that portion of the alimentary canal extending between the pylorus and anus, destined for the temporary retention of the chymous mass, so that its nutrient parts may be absorbed, whilst its more solid, indigestible constituents, are coUeeted for excretion. The intestine in all monogastria, but es- pecially in soUpeda, occupies by far the greatest part of the abdominal cavity. The bonds of attachment to the various parts of the latter are contracted by the intestine, through its peritoneal investment, more especially to the spine, constituting mesen- ters, which I shall especially allude to when describing with more detail each portion of this capacious tube. Not only the attachments, but also the shape of the intestine, vary at different parts of its course, so that it has been deemed necessary to divide it, either arbitrarily or at natural demarcations. Thus we speak of the small and large intestine, the two being separated naturally by a marked change in direction, size, and confirmation. It is also obvious that, as the situation, at- tachment, and shape of each portion of the intestinal canal differ, so must the relations be equally distinct, and further mention of them will therefore be reserved for fuUer exposition elsewhere. SMALL INTESTINE. This, the smallest although longest, is also the first portion of the intestinal tube, extending from the pylorus to its sudden termination into the large intestine. In it the chymified mass is subjected to the modi- fying influence of important secretions, whereby its nutritive parts are fitted for absorption by the vessels, which, for this pur- pose, are arranged in this portion of the in- testinal track. The small intestine has been divided into three parts: this classification is, however, purely conventional. Since it does not recognize anatomical differences for its basis, it might justly be presumed that this dis- tinction of human anatomists exhibited traces of imperfection, even when applied to the frame of man. Such being the case, it is no matter of surprise that, in referring the distinction to the intestinal canal of animals, the incongruities of the system should be still more apparent. Extending from the pylorus, the first por- tion is termed the duodenum, from its being considered as twelve fingers' breadth in length : it is, however, extended round to the left side of the spine, posteriorly to the anterior mesenteric artery. The middle, or floating portion of gut, takes the name of jejunum, and the thii-d,'or csecal portion, is distinctively designated ileum. The duodenum forms a wide curve from the pylorus round to the right, being situated under the concave surface of the liver, pass- ing above the transverse colon, so as to attain the posterior part of the mesentery, and, reaching the left side of the spine, comes in contact with the colon, where it is said to end in the jejunum. The duodenum is fixed by the gastro-hepatic omentum to the concave surface of the liver, the layers of which enclose the biliary and pancreatic ducts, whereby this bond of union is still further strengthened. The peritoneum com- ing from the right and spigelian lobes of the liver, as well as from the right kidney, forms a loose attachment for the duodenum by 104 ANATOMT Am) PHYSIOLOGY OP extending on to the hepatic flexure of the colon, after it has surrounded the first-named gut. The next portion of intestine is at- tached to the spine transversely to the long axis of the body ; winding round the mes- entery to the left of the aorta, it gets at- tached to the gastric flexure of the colon, and here it proceeds, under the name of jejunum, along the free borders of the mesentery. As to the shape of the duodenum, from the pylorus to the right of the porta, we find its dimensions so very great as to have suggested to the ancients the similitude be- tween it and the stomach, of which they regarded it in some degree as an analogue, as testified by the appellation " Ventriculus Succenturiatus," given to it by them. Fur- ther from the pylorus, we find it constricts and assumes a certain caliber, which it maintains till it loses its name for that of jejunum. With reference to the relations of the duodenum, it may be stated that they ad- mit of detail on account of the fixedness of that portion of the gut, an attribute with which it is endowed in contradistinction to the jejunum and ileum. In the first portion of its course, i. e., from the py- lorus to the posterior part of the right lobe of the liver, the duodenum by its upper surface is in contact with the con- cave surface of the latter organ, crossing the vena portse, near which it is pierced by the billiary and pancreatic ducts, which enter it at about five or six inches from the pylorus, forming an acute angle \vith each other. The inferior surface of the duo- denum rests on the transverse colon, and its superior margin is in close contact with the anterior part of the head of the pan- creas. Round to the right, the duodenum is in contact with the hepatic flexure of the colon, right and Spigalian lobes of the liver, as well as the right kidney. To reach the spine it has to cross the direction of the right flexure of the colon, getting behind the mesentery and gastric flexure of the colon, where it is connected with the left kidney. Alluding next to the general anatomical facts as applied to the jejunum, so called on account of its usual vacuity after death, the limit between it and the ileum is de- fined by imagining the small intestine, with the exception of the duodenum, divided into five equal portions, of which the first two take the name of jejunum, whilst the last tliree-fifths receive that of Ueum. The jejunum is suspended superiorly from the spine by an extensive fold of per- itoneum, termed mesentery, wliich serves also as a medium for the passage of the mesenteric arteries, veins and nerves, as well as for chyliferous vessels, to take their course towards the receptaculum chylo, sit- uated to the left of the aorta. The ^vidth of the jejunum is far from being uniform, it being more constricted at some points than at others: its narrowest part is that which is contiguous to the ileum. The ileum is the terminating portion of the small intestine, so called from the tor- tuous course it takes, emptying itself into the large intestine at the junction of the cfficum and colon, by an orifice provided with a valve. The first portion of the ileum is simply attached by mesentery to the spine ; but, in addition to this, in the last part of its course, the gut is connected with the cseum by a fold of peritoneum, which is not large enough to prevent them deviating more than an acute angle from each other. The ileum is, on the whole, the narrow- est portion of the small intestine, but the thickest in its coats. Having now especially to describe the structru-e of the small intestine, it may be taken as a whole, merely alluding to local peculiarities. This portion of the alimentary canal has four coats, to be described in the same order by those of the stomach, i. e. peri- toneal, muscular, cellular, and internal mucus. • THE HOHSE. 105 The first, or the peritoneal, has nothing peculiar, beyond its enclosing a little trian- gular space all along the upper attached border of the gut. The looseness of the peritoneal folds attaching the small intes- tine is very marked; and Colin (Soc. cit.) notes, that the mesentery is proportionately larger in young than in adult quadrupeds, so that the gradual shortening of this ex- plains the spontaneous reduction of exom- phalus or umbilical hernia. The second, or muscular coat, is mostly developed at the commencement of the duodenum and terminating portion of the ileum. It consists of white involuntary fibres, arranged so as to form an outer longitudinal layer, and an inner circular one, both of which completely encircle the gut. The third, or cellular coat, is similar to that of the stomach, in being disposed in two layers, so as to connect the three coats together. It is especially condensed on the inner surface of the muscular coat, so as to take the appearance of a fibrous tunic, at- tached to the mucus lining by loose cel- lular tissue. ' The fourth, or mucus, coat is thin, hav- ing a velvet appearance, due to villi, pecu- liarly small in the intestines of the horse, but remarkably developed in other animals, especially carnivora and fishes. The villi may be seen by a pocket lens, on a well- washed piece of intestinal mucus mem- brane, and between them are seen numer- ous foramina, which are the openings of tubular glands, known as the crypts of Lie- berkuehn. In addition to the tubular glands, by dis- secting, from without, the muscular firom the mucus coat, lining the commencement of the duodenum, we find clusters of vesi- cles, similar to the vesicular structure of the salivary and pancreatic glands. These form distinct layers, provided with ducts, which open on the free surface of the mem- brane ; and Dr. Todd states that Brunner's glands, or, as he calls them, the duodenal, are more developed in the horse than in any other animal he has hitherto examined them in. We have next to treat of the solitary glands — glandulos solitaris — peculiar and rather scanty bodies, visible at various parts of the small intestine. These are vesicu- lar, and without any opening when in the perfect state, surrounded by villous pro- cesses and Lieberkuehnian follicles. Some of the villi also project from the surface of the so-called glands, which are most ap- parent when distended with secretion. About the second half of the jejunum, and along the whole of the ileum, we see longitudinal patches, varying from half an inch to even three inches in length, scat- tered all over, but more especially situated near the superior or attached border of the small intestine, which is contrary to the faulty description of some recent authors. These patches, distinguished as Peyer's glands or patches, also as Agminated glands — Gladdulae agminatas seu aggregate — con- sist of an accumulation of small bodies, each resembling a glandula solitaria in miniature, being also destitute of a natural aperture. Colin (loo. cit.) states that they are first seen at a distance of about six feet and a half from the pylorus, and the least num- ber of them he has ever counted has been 102, whilst the utmost has been 158. The mucus membrane of the small in- testine is thrown into folds, at different parts, which are transverse, and scalloped near the pylorus, whilst in other parts they are mostly longitudinal ; these are all tem- porary folds. There is no such arrange- ment as the valulse conniventes in the small intestines of the horse, though recent wri- ters of great eminence have described them. About five inches from the pylorus, at the superior border of the duodenum, is a semicircular fold, which, if elevated, ad- mits of the finger being tluust behind it into the wide biliary duct. The opening of the pancreatic duct is also visible beneath this fold, but it is not so capacious as the one last mentioned. LARGE INTESTINE. The large intestine constitutes the termi- nating portion of the alimentary canal, 106 ANATOMY AND PHTSIOLOGY OP being remarkably more developed in soli- pedes than in any other of our domestic quadrupeds. It occupies the greater part of the abdomen, and most of it is loose, whilst its shape and other peculiarities vary considerably at different points. It is divided into three parts — cEBCum, colon, and rectum — the precise extent of each being defined by special anatomical characters. The position of the large intestine being constant, it is necessary, for sake of pre- cision, to speak of the whole as to the course it takes in forming the three divis- ions, extending thus between the small intestine and anus. The caecum, or blind pouch, is the first gut, which protrudes in the middle on cut- ting through the abdominal walls at the linea alba. Its bend or blind extremity is projecting into the left hypochondiiac region ; its body crosses obliquely the floor of the abdomen, to reach the right iliac region, where it suddenly bends at an acute angle, being rather constricted, and forms the colon. At this part the latter receives the ileum, and extends up the right side of the abdomen to the diaphragm, where it ti-averses the direction of the spine, resting on the ensiforra cartilage ; turning round the left side, it attains the left iliac fossa posteriorly, where it forms a twist lilie a letter S, from which similitude it has been termed the Sig- moid Flexure of the Colon. The gut, having diminished in size, retiu-ns up the same side of the abdomen to the diaphragm, where it again crosses the spine. Being now on the right side, it continues back to a point beyond the anterior mesenteric ariery, where it turns upward and for- ward, so as to come in front of the artery in question ; then, from right to left, so as to cross the spine for the third time, consti- tuting the fransverse colon, which is more capacious than the part preceding it. The two curves which it forms, one on the right and the other on the left, are respectively called the hepatic and gastric flexures of the colon. The gut so proceeds backward along the left side of the mesentery, being diminished again in size, and constituting the single colon, till we get to the posterior mesentery artery, where, unaltered in other respects, it takes a straight coru"se through the pelvis, out at the anus, and hence the name of Rectum. The CEBum, so called from having only one outlet, being closed at its anterior part, or caecum caput coli, from its being the blind head of the colon, is vulgarly termed the water-bag, owing to the almost invari- able fluidity of its contents. It is situated, as I have before said, ob- liquely along the floor of the abdomen, ex- tending backwards from left to right. It is attached to the spine by a meso- CBBum, which is a fold of peritoneum, com- ing off from the spine on to the superior part of the pouch. There is then the fold already alluded to, which stretches from the ileum on to the caecum, and, through the medium of the mesentery, indirectly con- necting the latter with the spine. The caecum is cone-shaped, having an apex and a broad base. The former gene- rally protrudes the first, when a medium longitudinal incision is made into the abdominal walls, although it is situated above the left portion of the double colon, whilst the liver is directly in contact with the floor of the abdomen. Like the other divisions of the large intestine, the caecum is sacculated. The bands producing this appearance are three in number at the apex ; but between two and three inches from this, one of them bifurcates, so that four bands result, which are continuous on to the colon. The colon arising from the csecum, re- ceives at first the contents of the ileum, being situated along and occupying the greater part of the floor of the abdomen. The colon is generally distinguished as double and single. By double, is meant the flexures of the gxit from its commence- ment to its gastiic curve ; whilst by the single colon, is understood the continuation of the same intestine to the part where the rectum commences. The double colon is attached by the peri- THE H0R3E. 107 toneura coming off on to it from the cas- cum, in the right iliac fossa, and continues from the outer flexm-e on to the inner, so as to Iceep the two in perfect apposition. Thus, if the abdominal parietes are cut through, the whole of the double colon may hang out, with the exception of the transverse portion. The latter is attached to the right kidney, as well as concave surface of the liver, by folds of peritoneum ; to the spine by the transverse meso-colon ; and still more to the left, it is loosely attached by the gastrocolic omentum to the stomach and spleen; besides which it has a peri- toneal attachment to the left kidney. Then the single colon commencing, it is loosely affixed to the spine by an extensive peri- toneal fold, the meso-colon, similar to the mesentery, but smaller and to its left : this fold is continuous posteriorly with the meso- rectum. The relations of the transverse colon are important, no less than interesting, inas- much as it is in close connection with the most important abdominal viscera. On the right, its upper surface is contiguous to the right kidney, as well as to the right and Spigelian lobes of the liver. In the middle, its superior surface is connected princi- pally with the pancreas; and to the left, but stiU superiorly, it approaches the left kidney and spleen. Anteriorly, the stomach also touches it, especially during repletion. The shape of the colon is very variable in different parts of its course. Thus, the first portion of the double colon, from the right iliac fossa till it forms the signoid flexure, is capacious and sacculated; the latter being due to the four bands con- tinuous on to it from the caecum. At the signoid flexure the bands are completely lost, so that the gut is smooth ; but, as we extend up towards the diapliragm, the an- terior band begins, and then the posterior one becomes apparent; so that the .trans- verse and single portions of the colon are puckered by two longitudinal bands. The Rectum, so called from its compar- ative straight course through the pelvic cavity, arises from the single colon, a little anteriorly to the posterior mesenteric artery, and ends at the anus, where its mucus membrane is continuous with the common tegumentary covering. It is attached in its anterior two-thirds by a meso-rectum ; the posterior third is an exception to any other part of the intestinal track, in so far as it is connected to adjacent parts by special faschite, and at its termination by certain muscles hereafter to be dwelt upon. The size of the rectum is much the same as the single colon. It is puckered in its anterior part by two longitudinal bands ; and the sacculi, resulting therefrom, deter- mine the shape of the faecal matters. The rectum is superiorly related to the spine, whilst inferiorly it comes in contact with the bladder, bulbous portions of the vasa deferentia, vesiculae seminales, and prostate. The structure of the large intestine does not vary essentially from that of the small, as it possesses the four coats, i. e. peritoneal, muscular, cellular, and internal mucus. The peritoneal tunic forms an entire cov- ering to the large intestine, with the excep- tion of the superior surface of the trans- verse colon — which is in contact with the pancreas — and the terminating portion of the rectum. The bands by which it unites the intestine to other parts have been akeady described. In addition to the peritoneum formijig an entire covering to the gut, at the attached margin of the flexures of the colon it constitutes folds loaded with fat, varying in width in difierent parts, and clus- tered so as to have deserved the name of appendices epiploicse. The muscular coat of the large intestine is differently developed in various parts. Its fibres are of the plain variety, and ar- ranged in two orders. The outer longitud- inal set is scanty in some parts, but in others forms the longitudinal bands above alluded to. These are shorter than the actual length of the gut itself, so as effectually to pucker it. The number of longitudinal bands vai'ies from one to four in various parts of the gut, and the shape and breadth of the latter is not everywhere the same. The 108 ANATOMY AND PHTSIOLOGT OF longitudinal fibres are abundant in the rec- tum, but they only form bands in the ante- rior two-thirds, as posteriorly to this they uniformly surround the gut. The inner layer of fibres encircles the whole of the gut, being thickest tow-ards the apex of the Cfficum, as well as in the single colon and rectum ; at the end of the latter the inter- nal sphincter-ani is formed by an accumu- lation of the circular fibres. The circular fibres of the colon are engaged in forming the ileo-colic valve, hereafter to be described. The cellular coat of the large intestine resembles that of the smaU, only not so abundant, except at the terminating portion of the rectum, where it is much more de- veloped. The mucus lining of the large intestine is continuous anteriorly with that of the ileum, posteriorly with the common integu- ment. It is thin, more or less coated with mucus, scantier in glands than the one of the small intestine ; but the orifices of the Lieberkuehnian crypts are more apparent, owing to the surface here being destitute of villi. Saccular recesses, more or less capa- cious, exist in the membrane lining the large intestine. The difference in degree of vascularity gives rise to difference in the color of the mucus coat in various portions of the gut : thus, that lining the caecum is generally more deeply colored than that of the colon, whilst the rectal mucus mem- brane is more vascular, and hence redder than the colic or caecal one. At the termination of the ileum is the ileo-colic or ileo-CEecal valve, w-hich is con- stituted of two folds of mucus membrane, almost parallel to each other, and horizontal, leaving between them an eliptical orifice when partially drawn asunder. The folds consist of the circular fibres of the intes- tine, lined on the iimer or ileac side by the villous membrane of the small, wlulst on the csBcal and colic side they are covered by the mucus membrane proper to the large intestine. It is worthy of notice, that though muscular fibres partly enter into the construction of the valve, its efficiency is explicable on purely mechanical grounds, as proved by the fact, that it is competent in the dead body. The anus is the outlet of the intestine, which is perfectly closed, except during the evacuation of fecident matters, and is made perceptible externally by the elevation of the tail, being situated in a space bounded su- periorly by the sacrum and coccyx, laterally by the ischial tuberosities, and interiorly by the urethra in the male and vulva in the female. It is lined within by the mucus mem- brane of the rectum, which is loose and of a marked red color. Its external covering is of common integument, destitute of hairs. Lying between the skin and mucus mem- brane are two circular muscles, whose office is to keep the anus closed and prevent con- stant evacuation of fceces, whilst there are other muscular appendages situated exter- nally to these, destined either to elevate or retract the anus, being evidently antagonis- tic to the sphincters. The internal sphincter-ani is in contact with the attached sm-face of the intestinal mucus membrane, and separated from the integument by the external one. It is con- stituted of the pale circular fibres of the gut, but towards its free edge certain colored fibres are apparent on it. The external sphincter is situated outside the internal one, and within the anal integu- ment: it is circular, and composed of red fibres, attached superiorly under the first coccygeal bone, and interiorly its fibres blend in the male subject in the accelerator urinffi and triangularis penis, and in the female with the constrictor vaginse. The levatores-ani are two pale muscles, attached on each side of the first coccygeal bones, and, spreading downward and for- ward on to the rectum, form an attachment for the internal spliincter, and blending with the longitudinal fibres, so as to increase the thickness of the muscular coat of the rec- tum. The action of these muscles must be that of elevating the anus, and shorten- ing the rectum from before backward. The retractors proper to the anus are one on each side attached to the inner surface of THE HORSE. 109 the articular extremity of the ischium. Ex- tending from before bacl^ward, and rather upward, they blend witli the external spliincter. Their action is obviously that of reti'acting the anal opening. VESSELS, NERVES, AND LYMPHATICS OF THE INTESTINE. The intestinal canal, as a whole, receives arterial blood from the anterior and poste- rior mesenteric arteries, hepatic branch of the coeliac axis, with branches from the in- ternal pudic. The arteries of the small in- testine are derived from the anterior mesen- teric, whose divisions, varying from twenty- four to twenty-eight, proceed to the small intestine, with the exception of four, which miuister to the nutrition and functions of the large intestine. The branches extend- ing from the main trunk, at acute angles, proceed between the layers of the mesen- tery, to \\'ithin one and a halt" or two inches from the gut, where they anastomose, form- ing vascular arches, from which the second- ary branches arise, and, proceeding on to the intestine, ramify on the several coats, espe- cially the mucus one. The anterior division of the anterior mesenteric artery, proceeding to the duodenum, anastomoses with the duo- denal branch of the hepatic artery. The last iliac division inosculates with the cscal and cplic branches of the same trunk. The caecum and colon receive arterial blood solely from the branches derived from the anterior mesenteric, with a slight contri- bution from the posterior mesenteric arteries. The branches of the former originate oppo- site the flexure made by "the cEecum and colon. The csecal clivisions, two in num- ber, proceed downward and forward till they reach the gut. The posterior one passes round the posterior part of the bor- der of the cfEcum, to get on the under sur- face of the latter, extending to the apex, in somewhat a sti-aight course, and ramifying collaterally ; at its termination it forms a vascular network, by anastomosis with the superior csecal artery. The latter one, reach- ing the gut, extends directly forward towards the apex, and comports itself like the former. Thus we see the flexure, formed by the CEecum and colon, is supplied by collateral branches, from the superior and inferior caecal mesenteric divisions, both these anas- tomosing on the corresponding surfaces with the colic arterial trunks. The two branches going to the colon ex- tend, about parallel to each other, down- wards and forwards and to the left, the one gaining the caecal end of the colon, whilst the other proceeds on to the hepatic flexure. Then these may be traced, the one back- ward and the other forward, relatively to the course of the gut, along its superior border, so as to reach the sigmoid flexiure, where they mutually inosculate. From the mesenteric division going to the transverse colon, is a branch proceeding on to the single portion, which anastomoses poste- riorly with the posterior mesenteric. Tills vessel divides first into two branches, i. e. an interior colic and a posterior rectal one. The anterior colic branch is dkected for- ward and downward between the layers of the meso-colon, and divides into four or five branches, which bifurcate and form arches, like the arteries of the small intes- tine, for the supply of the contiguous gut. The arteries of the rectum are sometimes spoken of as hsemorrhoidals, and these are distinguished as anterior, middle, and pos- terior. The anterior liEEmorrhoidals are formed by the liindermost branch of the posterior mesenteric artery, which, passing into the folds of the meso-rectum, supplies consecutive branches to the gut, till, poste- riorly to the peritoneum, where the arteries pierce the muscular coat, and, forming a network of vessels, anastomose with the middle haemorrhoidals, which are the ramifi- cations of the internal pudic* These inos- culate with the posterior heemorrhoidals derived from the same source. The anus is then supplied with blood from the last named branches, as well as from perineal twigs of the external pudic. The veins of the intestine accompany the * This artery sometimes, erroneously, goes by the name of its lerminatiug branch — the artery of the bulb. EXPLANATION OF FIGURE XII. MUSCULAK STRUCTURE. FORWAKD PARTS. •. Ligamentum colli. a" . Trapezius. 6'. Rhomboideus longus. c". Scalenus. g" . Postea spinatus. K. Teres major. i". Latissimus dorsi. v. Scapulo ulnai-is. m' '. n". Triceps extensor brachii. p". , Flexor metacarpi externus. q". . Flexor metacai-pi medius. r. Flexor metacarpi internus. «' '. Extensor metacarpi magnus. S. Splenius. «. Levator humeri. x". Extensor pedis. c". Obliquus externus abdominis. POSTERIOR PARTS. h'. i'. Gluteal muscles. f- Triceps. 7c. Biceps abductor tibialis, posterior. r. Adductor tibialis internus. m. Tensor vagina. n. Region of the vastus internus. r. Gastrocnemius internus. i. S. Gastrocnemius extenius and intemus. X. Extensor metatarsi. y- Peroneus. i'. Flexor pedis accessorius. «'•, g". Coccj-gcal muscles. 4. Subcutaneous thoracic vein. 5. Saphena vein, 6. Radial vein. OSSEOUS STRUCTURE. 10. 20. 21. The pelvis. 11. Cervical vertebroe. IS. Coccygeal bones. 16. The true ribs. 17. The false ribs. 18. Sternum. 22. Femur. 23. Patella. 33. Scapula. 34. Humerus. 35. Radius. d. Dorsal spines. f- Ulnar. THE HORSE. Ill but bonds of connection between it and other parts, such as the gastro-splenic omen- tum, and the attachment to the Iddney and transverse colon heretofore described. This coat is smooth externally, rather closely attached to the fibrous coat internally, but of considerable elasticity, so as to allow the spleen sufficient freedom for distention. The second or fibrous coat, also termed the albugineous or elastic coat, is that closely applied to the parenchyma of the organ. It consists of yellow and white fibres, and in some parts, such as in the trabeculse, Koelliker has found plain muscular fibres, which he says do not exist in the external portion of the fibrous tunic in the horse. The covering not only envelopes the outer surface of the organ, but sends sheaths and processes into its substance. The sheaths are purposed for covering vessels, whilst the processes, termed also trabeculas, divide the substance of the spleen into areolae or interspaces, which contain a red matter, easily washed and pressed out, known as the splenic pulp. The trabeculae also arise as processes from the vascular sheaths, as well as from the external tunic. When the pulp has been thoroughly washed, the outer coat, with the trabeculas and sheaths, have the appearance of a framework or skeleton. The splenic pulp has a medullary aspect, being composed of cells and blood vessels ; and if the organ be cut clean in any direc- tion, we see, besides the cut ends of vessels and trabeculEB, certain pearlish looking bodies, named, from their discoverer, Malpig- hian Corpuscles. If divided, fluid escapes from the cavity which exists in their in- terior. They are perfectly visible to the naked eye, being about one-thirtieth of an inch in diameter ; and, with a pocket glass, "they may be seen attached to the small ar- terial trunks, if the pulp has been previously carefully washed. The spleen derives its arterial blood through the splenic artery, which is the main division of the coeliac axis. Winding be- tween the folds of the gastro-splenic omen- tum, it not only sends numerous branches through the hUum, and on to the surface of the spleen, but also supplies the stomach, largely inosculating with the gastric artery, so that the two might mutually perform each other's office, if the main trunk of either were obstructed. The splenic vein is similarly distributed to the artery, and it empties its blood into the vena portee, just anteriorly to the pos- terior mesenteric vein. The nerves of the spleen are derived from the solar plexus, and with the splenic artery enter the spleen. The lymphatics of the spleen are said by Koelliker to be scanty; but Dr. Sharpey tends rather to the belief that they are abundant. They are arranged superficially and deep, both sets anastomising freely with each other, and, reaching the hilum, they enter various scattered lymphatic glands in the peritoneal folds, and then empty into the receptaculum chyli. LIVER. The liver is the largest gland in the body, and proportionately largest during certain periods of foetal life. It is of a dark reddish brown color, and destined for the office of biliary secretion. It is situated across the long axis of the body, in the right hypochondriac, epigastiic, and partly in the left hypochondriac regions. It is attached to various parts by five ligaments, four of which are peritoneal folds, and one is the remnant cord resulting from the obliteration of the unbilical vein within the abdomen. These attachments will be more fully described with the peri- toneal tunic. The external aspect of the liver is smooth, being convex superiorly and concave inferior- ly, broad posteriorly, and sharp anteriorly. It has a granular appearance, and a very superficial inspection clearly shoAvs that it is composed of lobules, about the size of a pin's head. The hepatic substance is irregularly divided into numerous segments by fissures, which either extend through the gland from side to side, or are mere grooves of more or less depth. The different segments of 112 ANATOMY AND PHYSIOLOGY OF the gland or lobes are three principal ones — right, middle, and left — to which smaller ones are appended. The right lobe is the largest of the three, situated in the right hypochondrium, being thickest posteriorly and sharp anteriorly. The supero-posterior part of the right lobe is marked by a depression, for the adapta- tion of the anterior part of the right kidney. At the superior part of the right lobe is an excavation for the vena cava, which extends from behind forward, and marks off the division between the right and middle lobe. The vena cava is here more or less imbed- ded in the substance of the right lobe, but, generally speaking, it is superficial in the horse, and only an imperfect channel is formed for it. Projecting from the inferior surface and posterior part of the right lobe, is the lobulus spigelii, which is of considerable size, being broad posteriorly, and attached by its superior and left border, so that it projects anteriorly and narrows ; its apex gradually tapers, and has been capriciously designated, by the lovers of a quintuple hepatic arrange- ment, lobulus caudatus. The middle lobe of the liver is the smal- lest of the three ; it is crossed on its inferior surface by the transverse fissure or porta of the liver, at which the vessels and ducts enter into and issue from the gland. The middle lobe in the horse is divided at its anterior part into five or six portions, and Mr. Percivall, in his Anatomy of the Horse, at page 259, has termed it the lobulus scissatus. It is traversed antero-posteriorly by a channel for the remnant of the um- bilical vein, which eventually joins the vena porta. The left lobe is the thinnest of the three, but occupies an intermediate position in length and breadth. It is very thin at its left margin, and gradually thickens pos- teriorly. At its posterior and left side is a depression, in which the CEsophagus rests. Sometimes the left lobe is divided into two at its anterior part ; at others it is single. The superior surface of the liver is convex, and in contact with the pillars and expanded portion of the diaphragm. The right as well as the Spigelian lobes, are in relation posteriorly with the right kidney and right supra-renal capsule, interiorly with the head of the pancreas, duodenum, and transverse colon> The middle lobe is related interiorly to the pancreas, but partially separated from it by the vena porta. It also suspends the duodenum, and its left edge is loose and in close proximity to the flexures of the colon. The left lobe is related posteriorly to the oesophagus, and inferiorly to the left end of the stomach. The pancreas also stretches across its posterior part, partially separating it from the transverse colon. The liver receives an incomplete covering of peritoneum. The latter, reflected from the diaphragm on to the concave sm-face of the middle lobe of the liver, forms a double membranous layer, known, in accordance with its shape, as the falsiform ligament, and holding in its free and concave margin the round ligament, the representative of a fcetal structure, the umbilical vein. Fur- thermore, the liver is provided with a coro- nary ligament, that surrounds the foramen dextrum of the diaphragm, through which the vena cava passes. The lateral ligaments are distinguished as right and left ; they connect each lateral lobe to the diapliragm. The only connections of the liver that remain to be mentioned are the stomach, duodenum, transverse colon, and pancreas to its inferior sruface, and the right kid- ney to the posterior part of the right lobe. Dissecting off the serous tunic, it is found connected with the biliary surface by cel- lular tissue, continuous at the porta with the so-called capsule of Glisson. The latter extends into the liver as a common sheath to blood vessels, nerves, lymphatics, and biliary ducts. To proceed with further description of the liver would be useless, unless first ex- amining the blood vessels and ducts in that part of their course Avhich is external to the organ. The hepatic artery is quite subor- dinate in size, considering the magnitude of the organ and amount of its secretion. It is a branch of the coeliac axis, at first in THE HORSE. 113 contact with the pancreas, and then be- tween the folds of the gastro-hepatic omen- tum, and it reaches the porta on the left side of the portal vein. After giving off pancreatic and duodenal branches, it di- vides into two, a right and a left one. The right, the largest and somewhat the longest, penetrates into the right lobe, giving ofi" collateral branches, first to the middle and then to the right lobe itself. The left is the smallest division, and is distributed to the lobe corresponding to it in position, and also to the middle one. The liver is exceptional for having, be- sides an artery, another afferent vessel — a vein, known as the portal vein, formed by the splenic, which also receives the gastric and mesenteries, meeting each other at the same spot near the posterior part of the pancreas. From its origin, the portal vein takes an oblique course from left to right through the pancreas, and being surrounded by nerves, it reaches the porta of the liver, and here divides into three principal branches, one for each lobe. At the porta we also see the biliary duct coming out, formed by the union of several branches, corresponding in number to the ramification of the blood-vessels. This duct passes through the gastro-hepatic omentum, meeting the pancreatic duct at almost a right angle, and with it opening into the duodenum about five or six inches from the pylorus. Having thus far considered the main vessels, we may examine further the inter- nal structure of the liver. At the porta the branches of the vessels and ducts are associated together, and surrounded by cel- lular tissue, which sheaths grooves or canals, cut in various directions in the substance of the organ. These are the portal canals, and the cellular tissue in question is Glis- son's capsule. The vessels and ducts ramifying on the sheath acquire the name of vaginal branches, and, as they are traced between the lobules, they are termed interlobular. Here the un- assisted eye ceases to take cognizance of their further relation ; but, with careful dis- section, and a common pocket lens, they may be ti'aced to the lobules, which they enter; and the blood of the hepatic artery and portal vein is emptied into a common set of vessels, the hepatic vein. The rela- tion of these vessels in the lobules may be seen on the surface in a good injected speci- men of Uver, where the hepatic veins have been injected one color, and the other ves- sels differently. By this means the centre of the lobule.is colored with the injection thrown into the hepatic veins, and the cir- cumference with that of the portal vein. The hepatic veins issuing from the lo- bules cross the structm'e of the liver in sep- arate grooves, formed by the coalescence of the hepatic particles, so that their base is in contact with the veins, and hence the name of the latter is that of the sub-lobular he- patic veins. These empty into the posterior cava by several orifices, as well as by two larger ones, guarded by semi-lunar valves, situated just at the foramen dextrum of the diapluragm. In addition to the blood-vessels and ducts of the liver, it is supplied with nerves from the solar plexus, which ramify with the ves- sels. The lymphatics of the liver are abundant, and aiTanged, like in other organs, as a su- perficial and deep set, which inosculate freely in the substance of the organ, and, uniting to form several branches, they issue from the porta of the liver, passing through some lymphatic glands situated round the fissure, and from this they advance to the receptaculura chyli. PANCREAS. The pancreas is a compound vesicular or racemose gland, being much of the same nature as the salivary glands. The pancreas occupies the interval be- tween the layers of the transverse meso- colon, along the upper surface of the trans- verse colon. Its attachments are merely cellular, with the exception of the pancreatic duct, which attaches it pretty closely to the duodenum. The pancreas is spoken of as having a 114 ANATOMY ANB PHYSIOLOGY OF body, a head, and a tail. The body of the pancreas is that part stretched across the middle lobe, whUe the head is longitudinally extended, being almost parallel to the vena portEe, and situated below and to the right of that vessel. The head is broad ante- riorly and rather nan'ow posteriorly, and continuous from below upward, and from right to left, then from behind forward, to gain attachment to the body, so as to form a ring for the passage of the vena portsB. The part to the left of this vein is termed the tail of the pancreas. The pancreas is related by its superior surface to the right, left, and Spigelian lobes of the liver, also to the vena cava and aorta, which separate it from the phrenic crura. The posterior part of the head of the pancreas is in relation with the right supra-renal body. The tail of the pancreas is sti-etched transversely to the branches of the cceiac axis, and attached to the left kidney by loose cellular tissue. The in- ferior surface is in contact with the trans- verse colon. On examining carefvilly the sti'ucture of the gland, it is found to consist of clusters of cells, from which ducts arise, and these unite to form a main trunk, that is trace- able back to the tail of the pancreas, in- creasing in size tUl it reaches the anterior extremity of the head, where it pierces the duodenum together with the hepatic duct. Besides these clusters of cells and ducts, the gland contains connecting cellular tissue. • The pancreas is supplied with arterial blood by branches from the three divisions of the cceUc axis, as well as from the an- terior mesenteric. The pancreatic veins empty themselves into the splenic. The nerves are derived from the solar plexus, and the lymphatics of the pancreas, on issuing from the glandular substance, may be traced to the common reservoir of chyle and lymph. GENITO URIXARY APPARATUS. Having already described the intra-abdo- minal portion of the alimentary canal, and its accessories, I proceed to the considera- tion of that portion of the genito-urinary apparatus as contained within the abdomen, in the widest acceptation of the latter term. By this I mean the kidneys, and with them, for anatomical convenience, I classify the siapra-renal capsules, then the ureters, bladder, membranous portion of the urethra, vasa deferentia, vesiculse semi- nales, prostate and Cowper's glands, with which I shall conclude. The kidneys are a pair of glands, whose function it is to secrete urine. They are distinguished as right and left, being both situated in the lumbar region ; but, so far as. concerns their topographical anatomy, notvvithstanding their similarity in position, they need separate notice. The right kidney is more anteriorly situ- ated than the left, coming in contact with the posterior part of the right lobe of the liver, to which it is attached. It is also fixed to the abdominal parietes by peri- toneum, and to the spine by blood-vessels. Its shape is that of a bent ovoid, being more symmetrical than the left. It has two surfaces and two borders. Though differing in these marked general characters, the Iddneys resemble each other in several equally obvious points of their general anatomy. Both kidneys have a peritoneal and an albugineous coat, both have an excretory duct, vessels and nerves, with a structure also equal in the two, con- stituting the bulk of the organ. Exter- nally to the peritoneal tunic is a more or less thick stratum of fat, which is more abundant in old than in young animals, when in a state of obesity. The peritoneal covering of the kidneys is incomplete, especially that of the right one, whose inferior surface and convex border are the only parts coated by it. The left kidney is also covered on its superior surface to a considerable extent, sometimes more and sometimes less. The attachments which each organ contracts through the THE HORSE. 115 medium of this serous investment have al- ready been described. The albugineous tunic is fibrous, and partly sub-serous. It forms a distinct cap- sule, attached to the substance of the organ by fibrous prolongations, which are in some parts aiTanged in pits and depressions, so as to mark out divisions on the surface of the kidney. In addition to this, the albu- gineous coat surrounds the vessels and ureter at the hilus, and enters the substance of the organ. On cutting the kidney horizontally from the convex to the concave border, there are three different parts brought into view, to be taken into consideration. Firstly, a dark conteur, of about half an inch or more in thickness, being generally less at the ex- treme ends of the kidney than at its middle, which completely encircles the central part of the gland, and is termed the cortical structure, fi'om its being most external. This part of the kidney has somewhat a granular aspect, and, when the vessels are full of blood or injection, they appear more or less arborescent, and clustered at innu- merable minute but visible spots, to form the Malpighian tufts. Next to this is a lighter colored material, rather ash-colored, but having a reddish hue, termed the me- dullary substance. This term is not given to it from the fact that it is medullary in consistence, but used in the metaphorical sense of being internally or centrally situ- ated. Approaching still nearer to the concave border of the kidney, is a funnel-shaped cavity, with its apex towards the hilus, and the base bounded by the medullary sub- stance, which is the pelvis. The apex is tubular, and continuous with the ureter, of which the cavity is but an expansion. The walls of the cavity are lined by a mucus membrane, which is loosely applied to the medullary substance, and thrown into folds, taking a radiated direction from the mouth of the ureter. Opposite the apex of the pelvis, the membrane is adher- ent to a prominent border of the mediillary substance, concave from before backward. but convex from above do%\mward, and is pierced by foramina, into which the hning membrane of the pelvis extends, so as to form the uriniferous tubes. On dissecting carefully away the mucus membranes of the pelvis, we reach to the fibrous tunic, which is not continuous on the medullary ridge, but merely attached to its sides, so as to increase the length of the boundaries of the cavity. The ureter arising from this dilatation is continuous outward toward the spine, and then backward, being related superiorly, as it issues from the liillus, with the renal vein ; and then crossing the posterior part of the kidney at its inferior siu-face, it gets between the peritoneum and psoas muscles, and is then traceable back to the bladder, into which it opens. The renal arteries, one for each kidney, arise at almost right angles fi-om the aorta, after the latter has given off the anterior mesenteric. The right one is more ante- riorly situated, and is longer than the left one. After each renal artery has given ofT a branch or more to the supra-renal cap- sule of the same side, it divides, on reach- ing the hilus, into a variable number of branches, usually eight or ten, which pierce the Iddney at different parts of the hilus, whilst a few branches proceed along the surface, supplying the capsule, and then also piercing the organ. The arterial branches entering the kidney have a de- finite arrangement, forming a kind of arch superiorly to the pelvis, from which second- ary divisions emanate and pierce the organ in aU du'ections, so as to reach the cortical substance, abruptly dividing into numer- ous branches, which eventually subdivide to form capillaries. By this it is evident that the cortical substance is more vas- cular than the medullary ; indeed the latter is very scantily supplied with arterial blood. From the arterial terminations the venous origins occur, and these unite to form branches, having a similar arrangement as the arteries ; only as they reach the pelvis, almost opposite the apex, they meet to 116 ANATOMY AND PHYSIOLOGY OF form a wide, capacious trunk, the renal vein. This is supplied \vith valves, not all of which are perfect. At the opening of each renal vein into the cava is a semi- lunar flap, overlapping the posterior part. The nerves of Iddneys are numerous, and derived from the renal plexuses of the sympathetic ; they accompany the vessels with which they penetrate their respective organs. Lymphatics may be seen issuing from the hiUus of the kidney ; they enter some lymphatic glands there situate, and then convey the lymph into the receptaculum chyli. SUPRA RENAL CAPSULES. These bodies, also called capsulse supra- renales, seu atrabilarias, are two in num- ber, and belong to the class vascular glands, whose office is very indefinitely known. They are situated one on each side of the spine, across the direction of the renal vessels. Their attachments are effected by vessels, as well as by the peritoneum, on their inferior surface, connecting them to the corresponding kidney and around to the spine. The shape of the supra-renal bodies is much the same on either side, bemg that of a slightly bent ellipsis. They vary from three to four inches in length, and from one and a half to two inches in breadth. Their concave border corresponds to the renal vessels, as well as to the anterior mesenteric arteries. The convex border is in contact with the inner margin of the Iddney. The anterior extremity of the right one is in connection mth the right hepatic lobe, whilst its inferior surface is in connection with the commencement of the colon. The left supra-renal capsule is related anteriorly to the pancreas, and infe- riorly to the transverse colon. The peritoneal coat of the supra-renal capsules is merely confined to their inferior surface. The proper substance of the organ is enclosed in a fibrous or albugine- ous coat, which forms a distinct covering externally, and becomes continuous as sheaths to vessels internally. On cutting horizontally across a supra- renal capsule, it is found to consist of an outer cortical and an internal medullary substance. The cortical substance is a brownish yellow, due to fat contained in vesicles, which, according to Professor Hein- rich Frey, are smaller toward the surface than more internally. The medullary sub- stance has a greyish aspect, and vessels are apparent in it, as also a yellow tinge, due, according to the above-named author, to similar vesicles, as in the cortical substance, only much scantier in fat. The arteries of the supra-renal capsules are offsets of the renals and anterior mesen- teric, as well as of the aorta, but very variable in number and origin. They are, however, always abundant, and enter the organ principally at its concave border. The veins are larger than the arteries, and pour their contents on the left into the renal vein, and into the vena cava on the right. The nerves of the supra-renal capsules are very abundant, and derived from the renal plexvis. Professor Frey states, that in the horse, gangloin corpuscles constitute one of the structural elements of the nervous tissue in this situation. The ureters, one to each kidney, are con- duits between the kidneys and the bladder, for the passage of vuine. Their cafiber is various, being about one-third of an inch broad, but getting narrower posteriorly. As the ureters issue from the Iddneys, they converge towards the spine; then pro- ceed suddenly backward, till they reach the brim of the pelvis, having thus greatly diverged ; here they converge again, passing downward and backward to reach the sides of the body of the bladder, which they pierce. In their course, the ureters are attached to the Iddney and psoas parvus by loose cellu- lar tissue, and by the peritoneum, which suspends them, by being stretched across THE HORSE. 117 their inferior surface. After the ureters have crossed the spermatic and iliac vessels, they are received witliin a fold of peritoneum, constituting the false ligaments of the bladder. They pierce the muscular coat of the bladder at a distance of about three inches from each other, if the viscus be distended. They pass between the muscular and mucus coats for about an inch, being somewhat diminished in caliber, when they suddenly open into the cavity by an elliptical orifice, so that if the bladder be distended, the sides of the orifice are stretched, and thus closed. The ureters are externally covered by a ceUulo-muscular coat, consistmg of a cellu- lar tissue, with muscular fibres arranged, partly longitudinally and partly circularly, the latter being most internally situated. The ureters are internaUy lined by mucus membrane, continuous anteriorly with the renal pelvis, and posteriorly with the vesical lining. The membrane is loosely attached to the outer coat, and thrown into longitudinal eflaceable folds. BLADDER. The bladder is dilatable musculo-mem- branous viscus, destined for the temporary retention of urine. It is situated during vacuity entirely within the pelvis, but when distended, even moderately, its fundus en- croaches on the proper abdominal space. The bladder is held in situation by the peritoneum coming off fi-om the rectum and sides of the pelvis, so as to form a serous fold, whicli also encloses the vasa deferentia and vesiculaj seminales. Besides this, the bladder is supplied w^ith true ligaments, as well as bounded posteriorly through the in- tervention of the urethra. The shape of the bladder is pyriform, approaching, however, to a sphere when empty or partially distended. It presents for consideration a projecting anterior portion or fundus, a middle part, or body, and a posterior one, or neck. The fundus is globular and regular, having fixed at its anterior part the two obliterated um- bilical arteries, and the remains of the urachus. The body has no precise limits, but may be considered as that portion on which the bulbous portions of the vasa- deferentia rest. It is circular, but if the bladder be much distended, it bends some- what backward and upward. The cervix vesicae is the most consti-icted part of the organ, and marks the limit between the bladder and urethra. The bladder is related by its fundus to the iliac flexures of the colon, interiorly to the pudic and ischial bones, superiorly to the ureters, vasa deferentia, vesiculse semi- nales, and middle part of the rectum. The bladder has three coats. The peri- toneal investment is merely a partial one, as it is reflected from the body on to the sides of the pelvis. It covers the superior sm-face almost completely, but its extent gradually declines laterally and inferiorly. The attachments contracted by the perito- neum are termed false ones. Thus we have the two umbilical arteries, one on each side, enclosed by peritoneum, forming the two lateral false Tigaments. Then the ves- tige of the urachus is similarly enveloped by peritoneum, and constitutes the anterior false ligament. The peritoneum coming oft" from the rectum on to the superior sur- face of the bladder, gives rise to a pouch, termed the recto-vesical pouch, or cul-de- sac, and laterally to the triangular folds limiting the latter, known as the superior false ligaments. Behind the peritoneal re- flection the bladder is attached to the rec- tum and pelvic parietes, by a continuation of the pelvic fascia, which, leaving the inferior surface of the pelvis at the symphi- sis pubis, comes on to the bladder, forming the inferior true ligaments of the latter ; the fascia is then continuous on to the rectum, blending wdth the cellular coat. The pelvic fascia is also traced on to the prostate and sides of the bladder, from the posterior part of the obturator foramen, constituting the lateral true ligaments. Beneath this fibro-serous coat are muscu- lar fibres, arranged in a peculiar manner. There is an outer longitudinal set, traceable 118 ANATOMY AND PHYSIOLOGY OP from the cervix forward toward the body, where the fibres diverge and become oblique, and some even circular ; this layer is prin cipaUy developed posteriorly. The inner or circular layer is not arranged in concentric rings ; but its fibres, beginning at the fund- us, appear to arise from various centres on the surface, and to be taldng a direction more or less curved in different parts, so as to get transversely to the long axis of the viscus, and thus from the inner side have a circular appearance. These fibres are more decidedly cu-cular at the neck, and act some- what lilve a sphincter. Some of the deeper fibres at the neck of the bladder extend for- ward to each orifice of the ureter, marking the limit of the vesical trigon, whose office must be that of approaching the Ups of the elliptical apertitrcs. The mucus coat of the bladder is gener- ally more or less coated with mucus and epithelium, which guard the structure from the corroding effects of the secretion it has to come in contact with. It is thrown into numerous folds, taking various directions, but principally concenti-ical toward the fundus, and longitudinal at the cervix, aU of which are eifaceable by distention of the bladder, and are most prominent when the latter is collapsed. At the upper part of the urethral orifice of the bladder the mucus lining is smooth and free from folds, mark- ing out a triangidar space, bounded ante- riorly by a line drawn between the orifices of the ureters, and laterally by two lines meeting at a spot at the superior part of the vesical orifice. This is termed the vesical trigon. At its apex is a projecting fold of mucus membrane or uvula vesicEe, which seems to moderate the flow of urine into the uretlu-a. The bladder is supplied with blood from the internal pudic, and its veins empty into the internal pudic vein. The nerves of the bladder are derived from the sympathetic, and partly from the two last sacral pau-s which supply the neck. The lymphatics go to glands surrounding the origin of the iliac arteries, termed pelvic lymphatic glands, from which vessels arise, communicating anteriorly with the recepta- culum chyli. URETHRA. This canal in the male subject is not only purposed for the passage of urine, but also transmits the products of the generative or- gans. It extends from the posterior part of the bladder to the glans penis ; but we shall only occupy ourselves with a description of the intra-abdominal or pelvic portion, which terminates at the bulb of the penis or ischial arch. It is continuous anteriorly with the blad- der, attached to the rectum and sides of the pelvis by fascia and loose cellular tissue and muscles. The urethra is cylindrical, of considerable length, and its coats of no mean thickness. The pelvic portion of the urethra is gener- ally about three or four inches long, taking a direction backward and somewhat up- ward. It is related superiorly to the vesiculae seminales, middle lobe of the prostate, and posteriorly it comes in contact with the rec- tum, but separated from it laterally by Cowper's glands. The first or prostatic portion of the ure- thra is purely membranous, strengthened by cellular tissue and a continuation of the fibres of the bladder, the circular ones in particular, which are abundant anteriorly. The posterior two-thirds of the pelvic por- tion of the m-ethra are covered by a thick red muscular layer, which completely encir- cles it, with the exception of that part coming in contact with Cowper's glands. This muscle is continuous behind with the muscular fibres of the penis, which consti- tute the accelerator m-inae. These fibres are externally mixed with longitudinal ones, a portion of which are merely the inner or inferior bundles of the retractor ani, whilst others are derived from the triangularis penis ; both of these muscles tend to fix the urethra. Postero-superiorly the fibres en- circling the urethra are blended with the external anal sphincter. The retractor penis, which gets attached to the sacral bone, is a THE HORSE. 119 white muscle also, affording fixity to the pelvic portion of the m-ethra. Beneath the muscular tunic of the urethra we find a loose cellular tissue, and pos- teriorly also some erectile structure continu- ous on to the penis. On slitting open the pelvic portion of the urethra, to examine its mucus membrane, we find that it is smooth, glistening, and thrown into longitudinal folds. It is antero- superiorly raised by the sub-mucus tissue into a permanent ridge, termed the crest of the uretlu-a or verumontanum. This has a depression about its middle, and on each side are the elliptical orifices of the ejacula- tory ducts, surrounded by the openings of the prostatic ducts. Posteriorly and later- ally are little papillated projections, pierced by ducts emanating from Cowper's glands. These tubular processes are arranged in two parallel lines longitudinally to the course of the uretlu'a. The pelvic portion of the urethra is sup- plied with blood from the internal pudic, and the veins empty into the vessel of the same name. Its nerves are derived from the two last sacral and accompanying sym- pathetic filaments. The lymphatics of the pelvic urethra are similarly disposed to those of the bladder. GENERATIVE ORGANS OF THE MALE. The last division of our subject is that of the abdominal generative organs, only a part of the generative system, and consist- ing in the vasa deferentia, vesiculae semin- ales, prostate and Cowper's glands. VASA DEFERENTIA. There are two vasa deferentia, one from each testicle, for the passage of semen to seminal reservoirs. The vas deferens arises from the posterior part of the epididymis or globus minor, passing through the inguinal canals, and reaching the abdomen ; it is situated in the sub-serous tissue, taldng a course upward, backward, and inward, to reach the brim of the pelvis ; then, crossing the course of the ureters, it gets on to the bladder, where it is dilated, and forms the biilbous por- tion. Its attachments are serous and cellular to the various parts mentioned, whilst its pos- terior part is connected ^vith the urethra. The vas deferens is related, in its course from the inguinal canal, to the bladder ; after it leaves the constituents of the cord, with the parietes of the abdomen ; crossing the mider surface of the iliac vessels, and reacMng the bladder on the inner side of the ureter ; also lying internally to the seminal vesicles, and the terminating portion being covered by the prostate. The structure of the vas deferens is simi- lar throughout, with the exception of the greater thickness of its coats at the bulbous portion, being thinnest where it contributes to form the ejaculatory duct. This tube, of very various length, is con- stituted of an outer cellular investment, not requiring peculiar notice ; of an intermediate contractile and elastic tunic; and, as its name implies, is composed of muscular fibres and elastic tissue, arranged in two layers, i. e., an outer longitudinal and an in- ner circular one, which are easily perceived. The internal or mucus lining is thrown into longitudinal folds, in the narrow part of the duct ; but in the bulbous part it forms permanent rugs, taking various directions, so as to enclose irregular interspaces. The vas deferens is supplied with blood principally from the artery of the cord, al- though the epigastric furnishes a twig to it as well. The bulbous portion is supplied also by vessels of no small calibre from the iliacs. Its nerves are from the sympathetic, as well as from the second and third lumbar. VESICUL^ SEMINALES. The seminal vesicles are one on each side of the bladder, and act as receptacles for the semen. Each seminal vesicle extends from behind forward, upward, and outward, being ex- ternal to the bulbous portion of the vas 120 ANATOMY AND PHYSIOLOGY OP deferens. It is attached by peritoneum coming off from the sides of the pelvis and rectum on to the bladder. The posterior part is fixed by cellular tissue to the pros- tate and neclv of the bladder. The seminal vesicle is pyriform, being about three inches long and about an inch broad at its fundus, but more constricted at its neck. It is connected with the corres- ponding surface of the bladder and rectum, but partially separated from the latter by the prostate. The seminal vesicle has an incomplete investment of peritoneum, covering only the anterior part, wlulst the prostatic portion is covered by an outer cellular coat. Be- neath this is an intermediate tunic, partly elastic and partly contractile. Lavocat describes this muscular coat as easily studied after maceration in dilute nitric acid, when it may be found to consist of an outer longitudinal and inner circular layer, most developed at the fundus, but very thin at the neck. The mucus membrane is plicated, the folds enclosing similar interspaces to those seen in the bulbous portion of the vas deferens. The vessels are supplied by the internal pudic, whilst the nerves are from the lesser splanchnic and two last sacral pairs. EJACULATORY DUCTS. Two in number, each being the common outlet to its corresponding vas-deferens and seminal vesicle, so that their contents may pass into the urethra by an elliptical orifice each side of the depression on the verumontanum. The relations of these ducts are simply to the prostate and urethra. When they reach the latter, they pass between the mus- cular and mucus coat for some little dis- tance, so that at first sight they appear shorter than what they really are. The structure of the ejaculatory ducts consists in an outer cellular and inner mucus lining, both of which are very thin. PROSTATE GLAND. The prostate belongs to the" class of secreting glands. It is situated on the commencement of the m-ethra and termi- nation of the vesiculas-seminales, being su- periorly related to the rectum. Its attach- ments to these parts are merely cellular, although it has some connection with the sides of the pelvis, rectum and bladder, thi'ough the intervention of the pelvic fascia. It is symmetrical in figure, and very vari- able in size, being quite rudimentary in aged geldings. It is of a gray color, knotty to the feel, although spongy in tex- ture. The prostate consists of a middle por- tion or body and two lateral lobes. The former is in contact with the cervix of the bladder and urethra, the latter with the ejaculatory ducts and seminal vesicles. This gland has a posterior convex and an anterior concave margin, whilst it is flattened from above downward, although from its connection with other parts it is rendered more or less convex from side to side. The prostate is composed of an exter- nal fibrous or cellular coat, which forms a complete covering to it. On cutting the gland in any direction, it is observed by the naked eye to have an areolar appear- ance, being a net-work of variously disposed fibres, the larger ones of which are found to be tubular. The prostate opens into the urethra around the orifices of the ejaculatory ducts by numerous apertures. It is supplied with blood from the pudic vessels, and its nerves are derived from the lesser splanchnic and two last sacral pairs. cowper's glands. These also belong to the class of secreting glands, and have sometimes been called the lesser prostates. They are situated ante- riorly to the bulb of the penis on each side of the membranous portion of the urethra. EXPLANATION OP FIGURE XIII. THE SUPERFICLVL LAYER OF 5IUSCLES TAKEN FROM THE BODY OF THE HORSE, SO AS TO EXPOSE THOSE MUSCLES WHICH ARE MORE DEEPLY SEATED. THE HEAD AXD KECE. a. Buccinator. 6. Caninus. c. Retractor labii inferioris. 1, 1. Orbicularis oris. 2, 2, 2. Complexus major. 3, 3. Trachelo mastoideus. 4, 4. Subscapulo hyoideus. 5, Stcrno maxillaris. 6, 6. Stemo thjTo-hyoideus. 7, Jugular vein. 8, Carotid artery, with the eighth paii-, and sjinpathetic nerves. 9, Trachea. 10. Scalenus. THE FORE LIMBS. 1. Scapulo ulnarius. 2. Caput magnum of the triceps extensor brachii. 3. Caput medium of the same muscle. 4. Anconeus. 5. Flexor braeliii. 6. Extensor metacarpi. 7. Extensor pedis. 8. Extensor metacarpi obliquus. 10. Flexor metacarpi extcrnus. 11. Perforans and perforatus. 12. Uhiarius accessorius. 13. Flexor metacarpi intemus. 14. Flexor metacarpi medius. 15. Perforans and perforatus. 16. Extensor metacarpi. B.. B. Radius. THE TRUNK AND BACK. A. Scapula. F. F. Longissimus dorsi. O. Spinalis dorsi. a, a. Intercostals. h, b. Superficialis costarum. e, e. Rectus Abdominis. d, d. TransversaUs abdominis. e, c, e. Obliquus intemus abdominis. /. Hollow in the longissimus dorsi, which part of the gluteus maximus once filled. EXPLANATION OF FIGURE XIII. CONTINUED. HAUNCH AND mXD EXTREMITT. C. Ilium. B. Ischium. E. Tibia. 1. Sacro sciatic ligament. 2. Sjihincter ani. 3. Depressor coccygis. 4. Muscles of the tail. 5. 5. Triceps abductor tibialis. 6. Vastus externus. 7. Rectus. 8. Gastrocnemius muscles. 9. Plantarius. 10. Extensor pedis. 11. Peroneus. 12. Flexor pedis perforans. 13. Insertion of the gracilis. 14. Gastroenemii muscles. 15. Flexor pedis accessorius. 16. Course of the perforans tendon, inside the cs cCch of the hock joint, 17. Insertion qj the gastrocnemius externus into the point of tie Ix&ck. 18. 18. Popliteus muscles. 19. Extensor pedis. THE HORSE. 121 Cowper's glands are covered by the tri- angularis penis of each side : they are about the size of a filbert. Their structure, as apparent to the naked eye, is similar to that of the prostate in every respect, only the excretory ducts are 11 ten or t\velve in number for each gland and linearly disposed on each side of the pelvic urethra. Cowper's glands are supplied with ves- sels and nerves from the same source as the prostate. ORGANS or GENERATION. TESTICLES AND SCROTUM. The preparation of the seminal fluid is the office of two oval glandular bodies, called the testes or testicles ; they are suspended in a portion of the common integument, termed the scrotum, by means of the sper- matic cord and cremaster muscle. The scrotum is composed of the common integument, sub-cellular tissue, and elastis muscle, (the fibres of the latter run in a lon- gitudinal direction, from the cellular sub- stance of the sheath, to the base of the penis), and lastly the tunica vaginalis, which is a prolongation of the peritoneum. The testicle has a peritoneal covering, termed tunica vaginalis testes, and also another distinct tunic termed tunica albu- ginea. The substance of the testicle is ex- tremely vascular, and the vdtimate branches of its spermatic arteries are collected into small bundles of fine convaluted vessels, separated from one another by septula9, or membranous partitions. From these the vasa seminifera, or beginnings of excretory ducts, take their origin, and gradually unite to form a smaller number of canals of larger diameter, but exceedingly tortuous in their course. The testicle is also supplied with nerves and absorbents, secretory and excretory vessels. SPERMATIC CORD.* The spermatic cord, the substance by means of which the testicle is connected with the abdomen, and by means of which it is suspended within its scrotal cavity, is com- posed in the following manner : 1st. It has four coverings ; there is immediately under- neath the skin the faschia superficialis ; next, the cremaster muscle ; thirdly, the tunica vaginalis; and lastly, the tunica vaginalis reflexa. Within the cavity formed by the vaginal tunic, it is that the intestine protrudes in inguinal and scrotal hernia ; the hernial coverings, consequently, exclusive of the sac, will be the faschia and cremaster muscle. 2ndly. The constituent parts of the cord itself, are : a. The arteries, which are two in number ; the artery of the cord, a small branch of the external iliac, which ramifies and expands itself upon the cord ; and the spermatic artery, which, as soon as it reaches the internal ring, enters the inguinal canal, runs down the posterior part of the cord, growing tertuous as it descends, ser- pentines along the superior border of the testes, between it and the epididymis, winds round the anterior end of the gland, and lastly reaches the convex border, where it becomes extremely convoluted, and whereto its branches are principally distributed. In its descent it detaches small unimportant twigs to the adjacent parts ; and, as it ap- proaches the testicle, becomes surrounded by an assemblage of venous vessels, b. The veuis accompany their corresponding arteries, and they indeed may be said to make up the principal built of the cord, for they are not only numerous, but large and flexuous, and, as they approach the testicle, form a sort of plexus, which has got the name of corpus pampiniforme : they return their blood into the posterior vena cava. c. The nerves, Avhich are derived from the hypogastric plexus, also accom- pany the spermatic artery : they are smaD, but sufficiently numerous. Though the testicle does not possess any very great sen- sibility in health, we may vouch for its being acutely sensitive in a state of disease. d. Absorbents exist, both large and numer- ous, in the cord. They are readily found nam ANATOMY ANJ PHYSIOLYGY OF THE HOESE. 123 alongside of the venous trunks; and not infrequently may be filled by introducing mercury into the spermatic artery, e. The VAS DEFERENS, though a constituent of the cord, takes at first a solitary course, remote from the blood vessels. The duct issues from the summit of the head of the epidi- dymis, beginning in a series of convolu- tions gradually unvi'inding as it proceeds; it takes an oblique course nearly as high as the external ring, where it joins the blood vessels, and continues to accompany them posteriorly through the inguinal canal : at the internal ring it leaves them, turns inward and ascends into the pelvis, where we find it creeping along the side of the bladder infolded in peritoneum to get to the cervix, crossing under its course first the umbilical artery and then the ureter; at length it terminates by rather a contracted orifice within the mouth of the duct of the vj^icula seminalis, just behind a little emi- nence in the urethra — tlie capat galinaginis, about an inch posteriorly to the cervix of the bladder. Within the inguinal passage the duct is accompanied by the artery of the vas deferens,^, long slender branch of the epigastric. Its cang-l, flexuous until the duct has joined the cord, but straight in its subsequent course, is not uniform through- out in caliber ; the area of its tortuous part is large, but as it becomes straight it grows contracted: having entered the pelvis, it gradually enlarges again, and acquires un- usual volume in running along the side of the bladder ; and the canal of the enlarged por- tion presents a riticulated structure, which gives its exterior an irregular, tuberculated appearance ; the most contracted part is that in union with the duct of the vesicula seminalis, which is a comparatively small cylindrical conduit. The parietes of the duct are so remarkably thick and firm to the feel, that we distinguish it at once by the fingers from the other parts of the cord : they consist of two tunics ; the ex- ternal one (in which its main thickness con- sists) is white, fibrous, and approaches in appearance to cartilage ; the internal one is thin and fine in texture, muco-membranous in its nature, and here and there incloses a reticulated structure. The different consti- tuent parts of the cord are connected alto- gether by cellular substance, destitute of any fat ; and from the circumstance of the parts in general being more bullvy below the ring, the cord increases in breadth and thickness as it approaches the testicle. THE EPIDIDYMIS. The epididymis is extended along the su- perior border of the testicle, upon which it rests, and to which it is connected by the tunica vaginalis reflexa. Its ends are buDiy in comparison to its middle : that receiving the vasa eft'erentia, the smaller one, is the caput or globus minor; the other, giving rise to the vas deferens, is the globus major, the part farriers call the nut. The interior of this appendage to the testicle exhibits a structure entirely vascular. The vasa ef- ferentia unite and re-unite until they form a single duct, of whose numberless and very remarkable convolutions the globus major is entirely constituted : these tortu- osities (which, when squeezed, freely emit semen) will admit of being unwound for a considerable extent, so as to have the length of the duct calculated with very tolerable exactness from beginning to end, which has been found to amount to several yards. It is small at its formation, but grows imperceptibly larger in making its manifold windings and turnings, until at length it assumes the size of the vas deferens, in which it ends. Its various convolutions are connected together by cellular membrane, and are interspersed with a sparing supply of blood vessels. The course of the semen is this: It is secreted by the capillary coils of the sper- matic artery, from which it is received by the tubili seminiferi: these tubes carry it into the rete, and the rete discharges it through the vasa efFerentia into the epididy- mis, from which it is conducted by the vas deferens into the urethra. Formation and Descent. — It is a singu- lar fact, that the organs whose structures we have been investigating, are originally 124 ANATOMY AND PHYSIOLOGY OF formed in a situation remote from that in which they are destined to carry on their functions; "the colt has no testicles," is the common observation of the unim- formed on these matters ; and we know ourselves that the purse is without them, bu-t we know, in addition, that they exist ready-formed within the abdomen, and that they will descend at a certain period of age into the proper receptacle, the scrotum. During the foetal state we find the testicles more or less developed, tinged with a blush of red, lodged beneath the psoas muscles, in contact with the inferior borders of the kidneys, covered and retained in their situ- ations by peritoneum, and concealed by the intestines around them. Here they receive their arteries from the contiguous trunk — the posterior aorta ; the vasa deferentia run forward to them, and the cremosters liltewise turn forward instead of backward ; there being at this time no such tiring as a spermatic cord. Thus placed, the testicle may be regarded as one of the glands of the abdomen ; indeed it has considerable similarity to the kidney — receiving its ves- sels from the same contiguous source, and sending a long duct backward into the cavity of the pelvis ; nor does there appear any conclusive reason why it should not perform the same office in that situation that it does in the scrotum, and particu- larly since it is known that in birds the tes- ticles remain within the abdomen during life. From the part where the blood ves- sels enter, we find growing a whitish sub- stance, extending backward, diminishing in breadth as it recedes, passing through the ring where the fUus of the cremoster may be traced upon it, and whence it is prolonged into the scrotum, growing nar- rower and narrower until it vanishes ; this substance, regarded by some simply as a ligament, was considered by Mr. Hunter as the gubernaculum or pilot, by means of which the testicle is directed in its passage from the abdomen into the scrotum. Quit- ting the spot where it has been formed and matured, the testicle gradually retrocedes, guided by the gubernaculum, until it ar- rives on the internal ring, which, at this time (like every other part of the parietes) is closed by peritoneum ; this temporary obstruction it overcomes by drawing the membrane down along with it through the ring, and carrying the pouch made thereby down into the scrotum ; the gubernaculum at the time undergoing a complete in- version. This accounts for the production of the tunica vaginalis, and explains how that membrane comes to be doubled or re- flected; the testicle, receiving originally (as an abdominal viscus) one close adherent peritoneal tunic, and acquiring another which forms a loose covering as it passes through the ring, must necessarily have two ; and since both are derived from one and the same membrane, it follows that one must be a continuation of the other. These elongations of membrane, though everywhere in contact, are prevented from adhering together by a continual exhaifi- tion of the natural serous secretion. Any interval that might stibsist between them, in course, communicates with the cavity of the abdomen, through the ring, a part that remains open through life : this, however, is not the case with man — in his body the communication is cut off, after the testicles have descended, by a natural contraction and obliteration both of the ring and the inguinal passage. In many instances, one, in soxue few, both of the testicles, are known to have remained within the belly through life. As we are unacquainted with the immediate cause of their descent, so we are unable to give any rational explana- tion of this phenomenon. I have under- stood, that in many of these cases the glands have been found to be but imper- fectly developed : this, however, is not with- out exception. Period of Descent. — Most animals have their testicles within the scrotum at the period of birth. In the human fcetus they begin to move about the seventh month ; about the eighth they reach the groins ; and before birth they arrive in the scrotum. In the horse, they pass through the ring about the sixth or seventh month before THE HORSE. 125 birth, and are found within the scrotum at the period of parturition. In some cases, one testicle will not make its appearance for some time after the other ; and as the operation for castration is seldom long de- layed, this will account for the rigs (as horses having but one testicle are called) ■with wliich we meet every now and then. Again, instances are not wanting in which one testicle has descended to the ring and there remained through life.* The penis is composed of the two corpora cavermosa : head, or glans penis : corpus musculosum urethra, and the plexus veno- sus. The corporo cavermosa make up the bulk of the organ, they extend from the pelvis to the glans penis ; at the ischial arch they are invested with fibres of the erectors penis, and are strengthened and confined to the pubes by the suspensory ligaments. It is supplied with blood from a branch of the obturaler arter, by means of the inter- nal pvidic artery. Its nerves are termed pudic, so also are the veins. The g-lans is composed of a soft spongy tissue, highly elastic and distensible, and remarkable as the seat of the plexus venosus penis : the latter structure presents itself in the form of a venous conglomeration, and in the erect state of the organ constitutes its chief bulk. URETHRA. The m-ethra is a muco-membranous canal * In a communication I have been favored witli from Mr. Brettargh (whicli I have inserted in the second vol- ume of The Veterinarian), is contained the following information on this subject : " Colts arc foaled with their testicles in the scrotum, which remain there (in ordinary cases ) until the fifth or sixth month, when they are taken up between the internal and external abdominal rings, and there remain until the eleventh, twelfth or thirteenth month, all depeuding upon the degree of keep, as in some that are well fed the testicles can at all times be found in the scrotum. Were the testicles drawn up into the abdo- men, they would be too large to pass through the inter- nal abdominal ring at the time they are wanted to prepare for secretion ; which is occasionally the case, and at once accounts for our meeting with horses that are said to have but one stone. I have seen one instance where both were wanting in the scrotum at four years old." averaging in length, in the unerected state forty -eight inches ; it extends from the ante- rior part of the glans penis to the neck of the bladder ; its use is to afford a passage for the urine and seminal fluid. FEMALE ORGANS OF GENERATION. The vulva or pudendum comprises the prominence and fissure, commencing imme- diately beneath the anus, and extending downwards some four or five inches. The fissiu'e is longest and most conspicuous in breeding mares. The space between the anus and vulva is termed perineum. The prominences on each side of the vulva are called labia frudinde. They owe their bulk principally to muscular and fatty sub- stance, and cellular tissue. The commissures are the parts uniting the labia above and below. The superior or upper commissure is extended to a sharp angle, and joins the perineum; the lower portion is rounded off, and is bounded by a hollow, at the bottom of which is lodged the Clitoris. — This is brought into view im- mediately after staling: it bears a close comparison to the head of the male penis, and, like the latter, is susceptible of sensual enjoyment. To the clitoris belong a pair of muscles named erector clitoris. They take their origin from the perineum. Their ofHce is to erect that body, and proti-ude it into the vagina in the act of coition. T/ie internal parts are the vagina, uterus, FaUopian tubes, fimbriae, and ovaria. The vagina is a musculo-membranous canal, of large dimensions, extending from the vulva to the uterus or womb. It is situated within the pelvis, having the bladder below and the rectum above it, to both of which it has cellular attach- ments, in addition to the reciprocal connec- tion with the peritoneum. To the rectum it is closely and firmly attached by cellular membrane. The figure of the vagina, when it is dis- tended, is that of an oblong cylinder ; but in the collapsed state, its sides are in con- tact, and it will vary its form according to 126 ANATOMY AND PHYSIOLOGY OP the full or empty condition of the bladder. The largest part of the canal is the poste- rior ; there it exceeds the dimensions of the bladder. The length of the canal is about eighteen inches. Its course is horizontal, and rather shows an inclination to the curve of the rectum. The vagina, at its commencement from the vulva, is much thicker in its walls than elsewhere ; in composition it is partly mus- cular and partly membranous. The orifice of it is clothed in that strong, red, circular, fleshy band, which forms the sphincter vagina ; and the adjoining part of the canal is also encircled by some considerable fleshy covering, and thickly coated with muscular fibres. Farther forward than this the vagina is composed of membrane. The Membrane of the Vagina. — The part of which it is constituted is one of the mucus class, and one that possesses consid- erable density, extensibility, and resistance. Its exterior siu-face is rough. Its interior is smooth, and has a pale pinkish cast; unless the mare be under the venereal oes- trum, and then its redness is heightened, and its secretion augmented. In the ordin- ary state, this membrane is tlirown into folds, larger in breeding mares than in others, technically called rugas. Considerably in advance of the clitoris is an opening leading from the lower part of the canal, large enough to admit with ease any one of the fingers : this is the orifice of the meatus vuinarius, or outlet of the blad- der : it is guarded by a doubling of the vaginal membrane, which hangs over it, and Serves the purpose of a valve. The large and conspicuous protuberance at the bottom of the vagina, is the mouth of the uterus. The uterus, or womb, is a hollow mus- culo-membranous organ, united to the ante- rior part of the vagina, and is destined for the reception of the foetus. We distinguish the uterus by the body, horns, neck, and mouth. The body is the oblong or cylin- drical part, growing out of the anterior por- tion of the vagina, in the centre of whicli it is terminated internally by the os-uteri, or mouth of the womb ; it gives origin, in front, to the horns. This part lies wholly within the pelvis, between the bladder and rectum, and is entirely covered by peri- toneum. The cornua, or horns, rise from the body of the uterus, and diverge towards the loins. Their length and size will be much greater in breeding mares than in others. In figure they are cylindrical ; they bend upward in their course, and terminate in round extrem- ities, to ■which are loosely appended the ovaries, or testicles, through the medium of the Fallopian tubes. The cervLx, or neck, of the uterus is the rugose portion, protruded backward into the cavity of the vagina, which has a flower- like appearance, and can only be seen in a virgin uterus in the undistended state ; dur- ing gestation it undergoes a remarkable change. Independently of its union with the vagina, the uterus is confined in its place by two broad portions of peritoneum, which attach it to the sides of the pelvis, named the lateral ligaments of the uterus. During the period of gestation, the uterus experiences considerable extension. The Fallopian tubes are two trumpet-shaped canals, having a remarkable serpentine course ; running within the folds of the ligaraenta lata, from the extremities of the horns to the ovaries. The tube commences by an aperture in the cornu, having an elevated whitish mar- gin, which is scarcely large enough to admit a small silver probe : from this it proceeds forward, folded in peritoneum, and ex- tremely convoluted, until it reaches the ovary, to which it becomes attached ; it then begins to enlarge in its diameter, grows less convoluted, and serpentines along the lower side of the ovary ; it afterwards ends in a fringed doubling of membrane. The internal membrane of the tubes is similar to that of the uterus. The ovari, or female testicles, are two egg-shaped bodies, situated farther forward than the Fallopian tubes, within the cavity THE HORSE. 127 of the abdomen : they receive close cov- erini^s, and are loosely attached to the spuie. These bodies are about the size of wal- nuts. They are not regular oviform ; they have deep fissures on their sides ; they bear a resemblance, at first view, to the testicles and their ducts in the male. Internally, the ovaries are composed of a whitish spongy substance, in which are, in some instances, found little vesicles, con- taining a yellowish glairy fluid, in others one or more dark yeUow or brownish sub- stances, named corpora lutea : the vesicles are the ova, which, from impregnation, re- ceive further development; the corpora lutea denote the parts from which vesicles have burst, and consequently only exist in the ovaries of those mares whose organs have been engaged in the generative pro- cess. Prior to the age of sexual intercourse, these bodies are small and white ; but, as soon as the season of copulation is at hand, they grow large,* redden externally, and present many yellow spots or streaks through their substance. Mammre, though unconnected with the uterus, anomatically speaking, are in func- tion concurring to the same important end. The mammffi, vulgarly called udder, are two flattened oval-shaped bodies, depend- ing, between the thighs, from the posterior and inferior part of the belly. In quadru- peds, with but few exceptions, this is the situation of the mammae. In virgin mares the- udder is so small that there hardly appears to be any. In mares who have had foals, the udder re- mains prominent or pendulous, and has a flabby feel. Toward the latter part of gestation, this part swells, and becomes distinctly visible. Within a few days of foaling, the udder grows tiu-gid with milli ; it does not, how- ever, acquire its full distention until the foal has di-awn it for a few days, from which time it maintains its volume, with little variation, dinging the period of suck- ling. Soon after the foal begins to forsake the teat, the secretion of milk diminishes, and is followed by a contraction of the bag, which goes on gi'adually, untU it has resumed nearly, or quite, its former flat- ness. The interior of the mammae has a light yellowish aspect, and evidently possesses a lobulated structure, which is held together by a fine cellular tissue, interspersed with granules of fat. It is constituted of glan- dular masses, irregular in magnitude and form, and loosely connected one with another, each of which masses is composed of a number of lobules, closely compacted and united together. These insulated lob- ulous portions receive small arteries, from which the milk is secreted. The former, by repeatedly conjoining one with another, become at length several demonstrable canals, radiating from every part, and dilat- ing to hold the millc. When the udder becomes charged with millv, it flows into the teat and distends it. Suction is apparently an operation purely mechanical. The teat is seized and closely compressed by the lips of the foal ; and the imbibing effort which follows has a ten- dency to produce a vacuum, or raise the valve at the upper part of the teat, and the millc passes from the reservoirs into the mouth. PHYSIOLOGICAL CONSIDERATIONS. ON THE REPRODUCTION OP ORGANIZED BEINGS* ^ " If the changes which living beings un- dergo during the period of their existence, and the termination of that existence by the separation of their elements at a period more or less remote from their first combina- tion, be regarded as distinguishing them in a striking and evident manner from the masses of inert matter which surround them, still more is thek difference manifested in the series of processes wliich constitute the function of Reproduction. A very unneces- sary degree of mystery has been spread around the exercise of this function, not only by general inquirers, but by scientific physiologists. It has been regarded as a process never to be comprehended by man, of which the nature and the laws are alilve inscrutable. A fair comparison of it, how- ever, with other functions, will show that it is not in reality less comprehensible or more recondite than any one of them; — that our acquaintance with each depends upon the facility with which it may be siibmitted to investigation; — and that, if properly in- quired into by an extensive survey of the animated world, the real character of the process, its conditions, and its mode of oper- ation, may be understood as completely as those of any other vital phenomenon. " It may be considered as a fundamental truth of Physiological Science, that every living organism has had its origin in a pre- existing organism. The doctrine of ' spon- taneous generation,' or the supposed origina- tion of organized structures de novo out of assemblages of inorganic particles, although at different times sustained with a consider- * Carpenter's Physiology. able show of argument, based on a specious array of facts, cannot now be said to have any claim whatever to be received as even a possible hypothesis ; all the facts on which it claimed to rest having either been them- selves disproved, or having been found satis- factory explicable on the general principle omne vivum ex ovo. Thus, the appearance of Animalcules in infusions of decaying organic matter, the springing-up of Fungi in spots to which it would not have been supposed that their germs could have been conveyed, the occurrence of Entozoa in the bodies of various animals into which it seemed almost beyond possibility that their eggs could have been introduced, with other facts of a like nature, may now be accounted for, without any violation of probability, by our increased knowledge of the mode in which these organisms are pro- pagated. Thus, it is now well ascertained that the germs of Fungi and of many kinds of Animalcules are diffused tlnrough the atmosphere, and are conveyed by its move- ments in every direction ; and that, if to de- composing substances of a kind that would otherwise have been most abundantly peo- pled by these organisms, such air only be allowed to have access as has been deprived of its organic germs by filtration (so to speak) through a red-hot tube or strong sulphuric acid, no living organisms will make their appearance in them ; whilst in a few hours after the exposure of the very same substances to ordinary atmospheric air, it has been found to be crowded with life.* And when it is borne in mind, in the case of the Entozoa, that the members of * Sec the experiments of Schulze, in the "Edinb. New I Phil. Journal," 1837, p. 165. (128) ANATOMY AND PHYSIOLOGY OF THE HORSE. 129 this class are remarkable for the immense number of eggs which most of them pro- duce, for the metamorphoses which many of them are known to undergo, and for the varieties of form under which there is reason to suspect that the same germs may de- velop themselves, it becomes obvious that no adequate proof has yet been afforded that they have been, in any particular case, otherwise than the products of a pre-existing living organism. This, again, is the con- clusion to which all the most general doc- trines of Physiology necessarily conduct us. For it is most certain that we know noth- ing of Vital Force, save as manifested through organized structures ; whilst, on the other hand, the combination of inorganic matter into organized structures is one of the most characteristic operations of vital force ; hence it is scarcely conceivable that any operation of physical forces upon inor- ganic matter should evolve a Living organ- ism. Nor is such a conception more feasi- ble, if it be admitted that vital force stands in such a relation to the physical forces, that we may regard the former as a mani- festation of the latter, when acting tlirough organized structures ; since no vital force can be manifested (according to this view), and no organization can take place, except through a pre-existing organism. " It may be further considered as an es- tablished physiological truth, that, when placed under circumstances favorable to its complete evolution, every germ wiU de- velop itself into the lOieness of its parent; drawing into itself, and appropriating by its own assimilative and formative opera- tions, the nutrient materials supplied to it ; and repeating the entire series of phases through which its parent may have passed, however multiform these may be.* Now the germs of all tribes of plants and animals whatever bear an extremely close relation to each other in their earliest con- dition ; so that there is no appreciable dis- * The apparent exceptions to this rule, which have been brought together under the collective term, " Alternation of Generations," will be presently considered, and will be shown to be only exceptional when misiutei"preted. tinction amongst them, which would enable it to be determined whether a particular molecule is the germ of a Conferva or of an Oak, of a Zoophyte or of a Man. But let each be placed in the conditions it re- quires ; and a gradual evolution of the germ into a complex fabric will take place, the more general characters of the new or- ganism preceding the more special, as already explained. These conditions are not different in Idnd from those which are essential to the process of nutrition in the adult ; for they consist, on the one hand, in a due supply of aliment in the condition in which it can be appropriated; and, on the other hand, in the operation of certain external agencies, especially heat, which seems to supply the force requisite for the developmental process. Now, although we may not be able to discern any such osten- sible differences in the germs of different orders of living beings as can enable us to discriminate them from each other, yet, see- ing so marked a diversity in their operations under circumstances essentially the same, we cannot do otherwise than attiibute to them distinct properties ; and it will be con- venient to adopt the phrase germinal capa- city as a comprehensive expression of that peculiar endowment, in virtue of which each gem. ievelopes itself into a structure of its own specific type, when the requisite forces are brought to bear upon it, and the requisite materials are supplied to it.* Thus, then, every act of development may be considered as due to the force sup- plied by heat or some other physical agency, which, operating through the organic germ, exerts itself as formative power ; whilst the mode in which it takes effect is dependent I * This term is prefeiTed to that of " germ-power " sug- gested by Mr. Paget, because the latter seems to imply that the force of development exists in the germ itself. Now, if this were true, not only must the whole formative power of the adult have been possessed by its first cell- germ, but the whole formative power of all the beings simultaneously belonging to any one race, must have been concentrated in the first cell-germ of their original proge- nitor. This seems a reductio ad absurdum of any such doctrine ; and we are driven back on the assumption (which all observation confirms), that the _/brc« of develop- ment is derived from eocternal agencies. 130 ANATOMY AXD PHYSIOLOGY OP upon the properties or endowments of the substances through which it acts, namely, the germ on the one hand, the alimentan^ materials on the other, — just as an electric current, transmitted through the different ner\-es of sense, produces the sensory im- pressions which are characteristic of each respectively ; or, as the same cvtrrent trans- mitted through one form of inorganic matter produces light and heat, through another, chemical change, or through an- other, magnetism. " In the development of any li\iing being, therefore, from its primordial germ, we have three sets of conditions to study — namely, first, the physical forces which are in opera- tion; second, the properties of the germ, which these forces call into activity; and third, the properties of the alimentary mate- rials which are incorporated in the organism diulng its development. There is evidence that each of these may have a considerable influence on the result ; but in the higher organisms it would seem that the second is more dominant than it is in the lower. For among many of the lower tribes, both of plants and animals, there is reason to be- lieve that the range of departure from the characters of its parent, which the organism may present, is considerably greater than that of the higher ; and that this is chiefly due to the external conditions under wliich it has been developed. The forms of a number of species of the lower Fungi, for example, appear to be in a great part de- pendent on the nature of their aliment ; so among the Entozoa, there seems strong reason to believe that those of the Cystic order are only Cestoidea, that are prevented by the circumstances under which they exist from attaining their full development ; and the production of a fertile 'queen' or of an imperfect 'worker,' among the hive-bees, appears to be entirely determined by the food with which the lar\-a is supplied. No such variations have been observed among the higher classes ; in which it would seem as if the form attained by each germ is more rigidly determined by its own endow- ments ; a modification in the other con- ditions, which in the lower tribes would considerably afl'ect the result, being in them unproductive of any corresponding change. For, if such modification be considerable, the organism is unable to adapt itself to it, and consequently either perishes or is imper- fectly developed ; whilst, if it be less potent, it produces no obvious effect. Thus, a de- ficiency of food in the growing state of the higher animal will necessarily prevent the attainment of the full size ; but it will not exert that influence on the relative develop- ment of different parts that it does among plants, in which it favors the production of flowers and fruit in place of leaves, or that it seems to exercise in several parallel cases among animals. So, again, a deficiency of heat may slightly retard the development of the chick ; but, if the egg be allowed to re- main long without the requisite warmth, the embryo dies, instead of passing into a state of inactivity, like that of reptiles or insects. The extent, indeed, to which these external conditions may affect the develop- ment of the inferior organisms, must not be in the least judged of by that to which their operation is restricted in the higher ; and it is probable that we have yet much to learn on the subject. At present, it may be stated as a problem for determination, whether, from a being of superior organiza- zation, lower forms of living structure, capable of maintaining an independent existence, and of propagating their kind, can ever originate, by an imperfect action of its formative powers. Various morbid growths, such as cancer ceUs, to which the higher organisms are liable, have been looked upon in this light ; these have cer- tainly a powerful Aitality of their owti, which enables them to increase and multi- ply at the expense of the organism which they infest ; and they have also an enargetic reproductive power, by which they can pro- pagate their kind, so as to transmit the disease to other organisms, or to remote parts of the same organism ; but such growths are not independent ; they cannot maintain their own existence, when de- tached from the organism in which they are THE HORSE. 131 developed; and they have not, therefore, the attribute of a separate individuality. Various phenomena hereafter to be detailed, however, respecting the ' gemmiparous ' production of living beings, when taken in connection with that just cited, seem to ren- der it by no means impossible that the in- dividualization may be more complete in other cases, so that independent beings of a lower type may possibly originate in a per- verted condition of the formative operations in the higher. But no satisfactory evidence has ever been atibrded by experience, that such 'equivocal generation' has actually taken place; and its possibility is here alluded to only as a contingency wliich it is right to keep in view. That no higher type has ever originated through an advance in developmental power, may be safely as- serted ; for, although various instances have been brought forward to justify the asser- tion that such is possible, yet these instances entirely fail to establish the analogy that is sought to be drawn from them.* * Tlius, the author of the " Vestiges of the Natural History of Creation " refers to the various modifications wliich have taken place in our cultivated Plants and Domesticated Animals, in proof that such elevation is possible ; quite overlooldng the fact that these external in- fluences merely modify the development, without elevating it, and that these races, if left to themselves, speedily revert to their common specific t^-pe. And he adduces the phenomena of metamorphosis — the transformation of the worm-like larva into an insect, and of a fish-like tadpole into a frog — as giving some analogical sanction to the same doctrine ; totally overlooking the fact, that these transformations are only part of the ordinary develop- mental process, by which the complete fonn of tlie species is evolved, instead of being transitions from the perfected type of one class to the perfected type of one above it. So, again, he quotes the transformation of the worker- grub of the hive-bee into tlie fertile queen, as an example of a similar advance ; \vithout regarding the circumstance that the worker is physicallij higher (according to human ideas, at least) than the queen, whose instincts appear limited to the performance of her sexual functions ; and that the utmost which the fact is capable of proving, is, that the same germ may be developed into two dift'ereut forms, according to the circumstances of its early growth. It must always be borne in mind that the character of a species, to be complete, should include all its forms, per- fect and imperfect, modified and unmodified ; since in this mode alone can that " capacity for variation " be deter- mined, wliich is so remarkable a feature in many cases, and is that which specially distinguishes the races of plants and animals that have been subjected to human influence. " The development power which each germ possesses, under the conditions just now detailed, is manifested, not merely in the first evolution of the germ into its com- plete specific type, but also in the main- tenance of its perfect form, and, within certain limits, by the reproduction of parts that have been desti'oyed by injury or dis- ease. This reproduction, as Mr. Paget has pointed out,* differs from the ordinary pro- cess of nutrition in this, — that ' in grave injuries and diseases, the parts that might serve as models for the new materials to be assimilated to, or as tissue-germs to develop new structures, are lost or spoiled ; and yet the effects of injury and disease are re- covered from, and the right specific form and composition are retained ; ' — and, again, ' that the reproduced parts are formed, not according to any present model, but according to the appropriate specific form, and often with a more strildngly evi- dent design towards that form, as an end or purpose, than we can discern in the nat- ural construction of the body.' In the re- production of the leg of a full-grown Salamander after amputation, which was observed to take place by Spallanzani, it is clear that, whilst the process was from the fu-st of a nature essentially similar to that by which its original development took place, it tended to produce, not the leg of a larva, but that of an advUt animal. Hence it is obvious that, through the whole of life, the formative processes are so directed as to maintain the perfection of the organism, by keeping it up, so far as possible, to the model or archetype that is "proper to the epoch of its life wliich it has attained. The amount of this regenerating power, however, varies greatly in different classes of organized beings, and at different stages of the existence of the same being; and, as Mr. Paget has pointed out,f it seems to In no instance has this variation tended to confuse the limits of well-ascertained species ; it has merely increased our acquaintance with the number of diversified forms into which the same germ may develope itself. * " Lectures on Reproduction and Eepair." t Loe cit. 132 ANATOMY AND PHYSIOLOGY OF bear an inverse ratio to the degree of devel- opment which has previously taken place in each case. Thus, in the Hydra and other Zoophytes, it would appear (as in Plants) to be almost unlimited ; for the de- velopment process in them is checked at such an early period, that both the form of the organism and the structure of its tissues retain the most simple type ; and by the subdivision of one individual, no fewer than fifty were produced by Trembly. Li this, as probably in aU the cases in which new individuals have been obtained by artificial subdivision, there is some natural tendency to their production by the vege- tative process of gemmation ; but this does not always manifest itself. It is a ciuious fact, that the first attempt at regeneration, in some of these cases, is not always com- plete ; but that successive efforts are made, each of which approximates more and more closely to the perfect type. This was well seen in one of Sir J. G. DalyeU's ex- periments ; for he observed that, having cloven the stem of a Tubularia (a Hydroid Zoophyte), after the natural fall of its head, an imperfect head was at first produced, which soon fell off and was succeeded by another more fuUy formed ; this in its turn was succeeded by another ; and so on, until the fifth head was produced, which was as complete as the original. " As a general statement of the amount of this regenerating power, which exists in most of the different classes of animals, has been already given, it is unnecessary here to do more than allude to some of those facts which most strongly bear out the doctrine just laid down. Next to Zoophytes, there are no animals in which the regenerative power is known to be so strong as it is in the lower Articulata (as the Cestoid Entozoa, and the inferior An- nelida), and in the Planaria, which may perhaps be regarded as rather approximat- ing to the Molluscous type ; and here, again, we see that a low grade of general devel- opment is favorable to its exercise, and that the spontaneous multiplication which occa- sionally takes place in these animals by fission or gemmation, is only another form of the same process. In the higher forms of both these sub-kingdoms, as we no longer meet with multiplication by gemma- tion, so do we find that the reparative power is much more limited ; the only manifesta- tion of it among the fully-formed Arrach- nida and Crustacea being the reproduction of limbs, and the power of effecting even this being usuaDy deficient in perfect In- sects. The inquiries of Mr. Newport, however, upon the reproductive powers of Myriapods and Insects, m different stages of their development,* confirm the general principle already stated; for he has ascer- tained that in their larval condition. Insects can usually reproduce limbs orantennEe; and that Myriapods, whose highest development scarcely carries them beyond the larvae of perfect Insects, can regenerate limbs or an- tennae, up to the time of their last moult, when, their normal development being com- pleted, their regenerative power seems en- tirely expended. The Phas7nidce and some other insects of the order Orthoptera retain a similar degree of this power in their perfect state ; but these are remarkable for the similar- ity of their larval and imago states, the latter being attained, as in Arachnida, by a direct course of development, without anjrthing that can be called a ' metamorphosis.' Lit- tle is known of the regenerative power in the higher MoUusca; but it has been affirmed that the head of the Snail may be reproduced after being cut off, provided the cephalic ganglion be not injured, and an adequate amount of heat be suppUed. In Vertebrata, again, it is observable that the greatest reparative power is found among Batrachian Reptiles, whose devel- opment is altogether lower, and whose life is altogether more vegetative, than that of probably any other group in this sub-king- dom. In Fishes, it has been found that portions of the fins which have been lost by disease or accident are the only parts that are reproduced. But in the Sala- mander, entire new legs, with perfect bones, nerves, muscles, etc., are reproduced after * " Fhilodephical Trasaaetioiis," 1814. THE HOBSE. 133 loss or severe injury of the original mem- bers ; and in the Triton a perfect eye has been formed to replace one which had been removed. In the true Lizards, an imper- fect reproduction of the tail takes place, when a part of it has been broken off; but the newly-developed portion contains no perfect vertebrae, its centre being occupied by a cartilaginous column, like that of the lowest Fishes. In the warm-blooded Ver- tebrata generally, as in Man, the power of true reproduction after loss or injury seems limited, as Mr. Paget has pointed out,* to three classes of parts, namely : (1.) ' Those which are formed entkely by nutritive rep- etition, like the blood and epethelia, their germs being continually generated de novo in the ordinary condition of the body ; (2.) Those which are of lowest organization, and (which seems of more importance) of lowest chemical character, as the gelatinous tissues, the areolar and tendinous, and the bones; (3.) Those which are inserted in other tissues, not as essential to their struc- ture, but as accessories, as connecting or incorporating them with the other struc- tures of vegetative or animal life, such as nerve-fibres and blood-vessels. With these exceptions, injuries or losses are capable of no more than repair, in its more limited sense ; i. e., in the place of what is lost, some lowly organized tissue is formed, which fills up the breach, and suffices for the maintenance of a less perfect life.' Yet, restricted as this power is, its opera- tions are frequently most remarkable ; and are in no instance, perhaps, more strUcingly displayed, than in the re-formation of a whole bone, when the original one has been destroyed by disease. The new bony mat- ter is thrown out, sometimes within, and sometimes around, the dead shaft; and when the latter has been removed, the new structure gradually assumes the regular form, and aU the attachments of muscles, ligaments, etc., become as complete as be- fore. A much greater variety and com- plexity of actions are involved in this * " Lectures on Reproduction and Repair." process, than in the reproduction of whole organs in the simpler animals ; though its effects do not appear so strildng. It would seem that in some individuals this regenerating power is retained to a greater degree than it is by the class at large ; * and here again we find, that in the early period of development the power is more strongly exerted than in the adult condition. The most remarkable proof of its persistence even in Man, has been collected by Prof. Simp- son; who has brought together numerous cases in which, after ' spontaneous amputa- tion of the limbs of a foetus in utero,' occur- ring at an early period of gestation, there has obviously been an imperfect effort at the re-formation of the amputated part from the stump.f By the knowledge of these facts and principles, we seem justi- fied in the surmise, that the occtu-rence of supernumerary or multiple parts is not always due (as usually supposed) to the ' fusion ' of two germs, but that it may result from the subdivision of one; * One of the most curious and well-authenticated in- stances of this kind is related by Mr. Wliite, in his work on the " Regeneration of Animal and Vegetable Sub- stances," 1785, p. 16. "Some years ago, I delivered a lady of rank of a fine boy, who had two thumbs upon one hand, or rather, a thumb double from the first joint, the other one less than the other, each part ha\'ing a perfect nail. When he was about three years old, I was desired to take oif the lesser one, which I did ; but to my great astonishment it grew again, and along with it the nail. The family afterwards went to reside in Loudon, where his father showed it to that excellent operator, William Bromfield, Esq., surgeon to the Queen's household ; who said, he supposed Mr. White, being afraid of damaging the joint, had not taken it wholly out, but he would dis- sect it out entirely, and then it would not return. He ac- cordingly executed the plan he had described, with great dexterity, and turned^ the ball fairly out of the socket; notwithstanding this, it grew again, and a fresh nail was formed, and the thumb remained in this state." The Author has been himself assured by a most intelligent Surgeon, that he was cognizant of a case in which the whole of one ramus of the lower jaw» had been lost by disease in a young girl, yet the jaw had been com- pletely regenerated, and teeth were developed and occu- pied their normal situations in it. t These cases were brought by Prof. Simpson before the Physiological Section of the British Association, at its meeting in Edinburgh, August, 18.50. The Author, having had the opportunity of examining Prof. Simpson's preparations, as well as two*Uving examples, is perfectly satisfied as to the fact. 134 ANATOMY AND PHYSIOLOGY OF for, if it be supposed that this subdivi- sion has taken place when the develop- mental process has advanced no further than in a Hydra or a Planaria, it seems by no means impossible that each part might, as in those creatures, advance in its devel- opment up to the attainment of its com- plete form. " There are many tribes, both of Plants and Animals, in which multiplication is effected not only artificially but sponta- neously, by the separation of parts, which, though developed from the same germ in perfect continuity with each other, are capa- ble of maintaining an independent exist- ence, and which, when thus separated, take rank as distinct individuals. This process, which is obviously to be regarded, no less than the preceding, as a peculiar manifes- tation of the ordinary operations of Nu- trition, may take place in either of four different modes — 1. ti the lowest Cellular Plants, and the simplest Protozoa, every component cell of the aggregate mass that springs from a single germ, being capable of existing independently of the rest, may be regarded as a distinct individual ; and thus every act of growth which consists in the multiplication of cells, makes a corre- sponding augmentation in the number of individuals. 2. Li many organisms of a somewhat higher type, in which the fabric of each complete individual is made up of several component parts, we find the new gro\\i:hs to be complete repetitions of that from which they are put forth ; and thus the composite organism presents the sem- blance of a collection of individuals united together, so that nothing ig needed but the severance of the connection, to resolve it into a number of separate individuals, each perfect in itself The most characteristic example of this is presented by the Hydra, which is continually multiplying itself after this fashion ; for the buds or ' geramEe ' which it throws off are not merely struc- turally but functionally complete (being capable of seizing and digesting their own prey), previously to thfeir detachment from the parent. 3. In by far the larger propor- tion of cases, on the other hand, the 'gemma' does not possess the complete structure of the parent, at the time of its detachment, but is endowed with the ca- pacity for developing whatever may be deficient. Thus, the bud of a Phanero- gamic Plant possesses no roots, and its capacity for independent existence depends upon its power of evolving those organs. On the other hand, the 'zoospore' of an Ulva or a Conferva is nothing else than a young cell, from which the entire organism is to be evolved after it has been set free; and, even in the ' bulbels ' of the Mar- chantia, the advance is very little greater. The 'bulbels' of certain Phanerogamic plants, however, bear more resemblance to ordinary buds. 4. In the preceding cases, the organism which is developed by this process resembles that from which it has been put forth ; but there are many cases in which the offset differs in a marked de- gree from the stock, and evolves itself into such a different form that the two would not be supposed to have any mutual rela- tion, if their affinity were not proved by a knowledge of their history. Sometimes we find that the new individual thus bud- ded off is in every respect as complete as that from which it proceeded, though de- veloped upon a different type ; but in other instances it is made up of little else than a generative apparatus, provided with loco- motive instruments to carry it to a distance, its nutritive apparatus being very imperfect. Of the first, we have an example in the development of Medusas from the Hydroid Polypes ; and of the second in the peculiar subdivision of certain Annelida, hereafter to be described. Now it is obvious that, in this process, no agency is brought into play that differs in any essential mode from that which is concerned in the ordinary nu- tritive operation. The multiplication of individitals is performed exactly after the same fashion as the extension of the parent organism ; and the very same parts may be regarded as organs belonging to it, or as new individuals, according to their stage of development, and the relation of depen- THE HORSE. 135 dence which they still hold to it. The es- sence of this operation is the multiplication of cells by continual subdivision. " We have now, on ihe other hand, to in- quire into (he nature of the true Generative process, by which the original germ is en- dowed with its developmental capacity ; and this we shall find to be of a character precisely the opposite of the preceding. For, vmder whatever ch-cumstances the generative process is performed, it appears essentially to consist in the re-union of the contents of ti'JO cells* of which the germ, which is the real commencement of a ' new generation,' is the result. This pro- cess is performed under the three following conditions: 1. AJl the cells of the entire aggregate, produced by the previous subdi- vision, may be capable of thus uniting with each other indiscriminately ; there being no indication of any sexual distinction. This is what we see in the simplest Cellular plants. 2. All the component cells of each organism may, in like manner, pair with other cells, to produce fertile germs ; but there are differences in the shares which they respectively take in the process, which indicate that their endowments are not pre- cisely similar, and that a sexual distinction exists between them, notwithstanding that this is not indicated by any obvious struc- tural character. This condition is seen in the Zygneraa and its allies. 3. The gen- erative power is restrictad to certain cells, which are set apart from the rest of the fabric, and destined to this purpose alone ; and the endowments of the two sets are so far different, that the one furnishes the germ, whilst the other supplies the fertiliz- ing influence ; whence the one set have been appropriately designated ' germ-cells ' and the other ' sperm-cells.' Such is the case in all the higher Plants among which a true generative apparatus has been dis- covered ; and also throughout the Animal kingdom. * In vci-y rare instances, it is the re-union of the two parts of ilie contents of the same cell, Avhich had pre- viously tended to separate from each other, as if in the process of subdivision. " Thus, then, in the entire process in which a new being originates, possessing lUie structure and endowments with its parent, two distinct classes of actions par- tici])ate, — namely, the act of Generation, by which the Germ is produced ; and the act of Development, by which that germ is evolved into the complete organism. The former is an operation altogether sui generis ; the latter is only a peculiar modification of the Nutritive function; yet it may give origin, as we have seen, to new individuals, by the separation (natural or artificial) of the parts which are capable of existing as such. Now, between these two operations there would seem to be a kind of antago- nism. Whilst every act of Development tends to diminish the ' germinal capacity,' the act of Generation renews it ; and thus the tree, which has continued to extend itself by budding until its vital energy is well-nigh spent, may develop flowers and mature seeds from which a vigorous progeny shall spring up. But the multipli- cation of individuals does not directly de- pend upon the act of generation alone ; it may be accomplished by the detachment of gemvice, whose production is a simple act of development ; and the individuals thus produced are sometimes similar, some- times dissimilar, to the beings firom which they sprang. When they are dissimilar, however, the original type is always repro- duced by an intervening act of generation ; and the immediate products of the true gen- erative act always resemble one another. Hence the plu-ase, ' alternation of genera- tions,' can only be legitimately employed when the term generation is used to desig- nate a succession of individuals, by what- ever process they have originated ; an ap- plication of it which cannot but lead to a complete obliteration of the essential dis- tinction which the attempt has been here made to draw between the generative act and the act of gemmation. For when it is said that ' generation a produces genera- tion B, which is dissimilar to itself, whilst generation b produces generation c, which is dissimilar to itself, but which returns to 136 ANATOMY AND PHYSIOLOGY OF the form of generation a,' it is entirely left out of consideration that generation a pro- duces (the so-called) generation b by a process of gemmation ; whilst the process by which generation b produces generation c is one of true generation. So generation c developes d by gemmation, which resem- bles B ; and d, by a true generative act, produces e, which resembles a and c. This distinction, although it may at first sight appear merely verbal, will yet be found of fundamental importance in the appreciation of the true relations of these processes, and of their resulting products. So, in the Author's opinion, the application of the term ' generation ' to the entire product of the development of any germ originating in a generative act, whether that product consist of a single individual, or of a suc- cession, ■wLU be found much more appropri- ate, and more conducive to the end in view, than the indiscriminate application of it to each succession, whether produced by gem- mation or by sexual re-union. It is of great importance to the due comprehension of certain phenomena of Reproduction, which will come under consideration in the Animal kingdom, that the relations of the products of these two processes should be rightly appreciated; and this appreciation of them wiU, it is believed, be Lest gained by a careful inquiry into the phenomena of Reproduction in the Vegetable king- dom." EXAMINATIONS RESUMED. GLANDULAR APPARATUS. Q. Describe the structure of a glandular body. — A. It consists of a collection of tubes, more or less convo- luted, united by cellular substance into masses of a rounded form, constituting a lobule ; each lobule has a separate investment of membrane ; and the whole aggregate of lobules is furnished with a general mem- branous envelope or capsule. Each gland presents a complex arrangement of nimierous arteries, veins, nerves, and Ij-mphatics, and most of them are provided witli an excretoiy duct, which conducts the secretion prepared in the gland. Q. ^^^lat glands are supposed to be destitute of a secretory duct? — A. The pineal gland, thjioid, thy- mus, and renal capsides. Q. What function do most of the glands perform ? — A. Their function is tvvo-fold, namely, the separation of some material fi-om the eii-cidating fluid, wliich would otherwise prove mjurious to the system, and the elaboration of a product destined to renovate the tis- sues. OF THE ABDOMEN. Q. How is the cavity of the abdomen bounded ? — A. Anteriorly, by the diaphragm; posteriorly, by the pelris ; superiorly, by a portion of the vertebra ; inte- riorly and laterally, by abdominal muscles. Q. Into how many regions is the abdomen divided ? — A. Into nine, as follows : right and left hj^pochon- driac ; right and left lumbar ; right and left iliac ; epi- gastric, umbihcal, and hipogastric. PERITONEUM. Q. Why is the peritoneum called " serous mem- brane?" — A. In consequence of the serous or wateiy fluid with which its surface is constantly moistened. Q. What is the structure of serous membranes? — A. The same as that of the areolar tissue, haring a very smooth and glistening inner sm-face, which is covered with a layer of cells ; constituting a distinct tissue, termed epithelium. This is in contact with the primary membrane, thus isolating it fi-om the tissues beneath. Sub-adj acent to this is a layer of condensed areolai' tissue, wliich constitutes the chief tliickness of the serous membrane, and confers upon it its strength and elasticity; this gradually passes into that baser variety, by which the membrane is attached to the part it lines, and which is commonly known as the sub- serous tissue. A fibrous tissue enters into the compo- sition of the membrane itself, and its filaments inter- lace in a beautiful network, which confers upon it equal elasticit)- in every direction. Q. What is the purpose of this membrane ? — A. To facihtate the movements of the contained organs, by forming smooth surfaces which shall fi-eely ghde over each other. Q. What efiect does the gastric fluid have upon the food? — A. It is supposed to have the propertj' of dis- sohing the albuminous and gelatinous constituents of the food. Q. What is the real solvent of the gastric fliud ? — A. Either hydrochleric, acetic, or lactic acid. Q. Is not the solvent action of the gastric fluid aided by some mechanical means ? — A. Yes. By the move- ments of the walls of the stomach, which are produced by the successive contractions and relaxations of their THE HORSE. 137 muscular fibres, the contents of the stomach are thus kept in a state of constant agitation, which is considered favorable to their chemical solution. Q. Docs absorption of nutritious matter take place in the stomach? — A. Yes. A portion of the nutri- tious matter dissolved by the gastric fluid is at once absorbed into the blood-vessels of the stomach, and never passes into the intestinal tube, nor into the special lacteal system of vessels. Q. What term is appHed to the food after its reduc- tion, in the stomach, to a pulpy mass ? — A. Chyme. Q. Gas is frequently evolved in the stomach and intestines dm-ing digestion : how do you account for this ? — A. It is owing to a disturbed or morbid condi- tion of that process, and by no means a necessary at- tendant upon healthy digestion. Q. Does violent exercise immediately after a feed tend to retai-d the formation of chyme ? — A. It does. The circumstances most favorable to perfect digestion are, a short period of rest, followed by gentle exercise. Q. Does any portion of the food ever pass unchanged through the pylorus along with the chyme ? — A. Yes. Whole oats are frequently found in the horse's excre- ment. INTESTIXES. Q. The aliment now being converted into chyme, and having passed the pylorus, what becomes of it ? — A. It enters the duodenum. Q. Having entered the duodenum, with what does the chyme mingle? — A. The biliary and pancreatic secretions. Q. "V^Tiat effect do they have on the gastric secretion and the ch)-me ? — A. The biharj- and pancreatic secre- tions are supposed to contain an excess of alliali ; tliis neutralizes the acid of the gastric juice, so that there is no further solution of albuminous compounds, but the conversion of starch into sugar, which was interrujjted in the stomach, now recommences. Q. What are the uses of the bile ? — A. The chief, uses of the bile appear to be those of a chemical agent promoting the decomposition of the chyme, and also etimulatmg the secretion of mucus, and the peristaltic action of the intestines. Q. What eflect has the pancreatic juice on chyme or the elements of digestion ? — A. It forms an emulsion with oil and fat. Q. The ch)-me, having been acted on by the preced- ing secretions, what name is then given to it ? — A. Chyle. Q. Describe the properties of chyle? — A. If chyle be taken from the thoracic duct of an animal a few homrs after it has taken food, it has very much the appearance of cream, being a thick fluid of an opaque white color, without smell, and having a sKghtly acid taste, accompanied by a perceptible sweetness. It restores the blue color of Htmus, previously reddened by acetic acid, and appears, therefore, to contain a pre- ponderance of alkali. When subjected to microscopic examination, chyle is found to contain a midtitude of 18 globules, of smaller diameter than those of the blood, and corresponding in size and appearance to those of miUv. Li about ten minutes after it is removed from the thoracic duct, it coagulates into a stifl" jelly, which in the com-se of twenty-fom- hours separates into two parts, pro\iding a fii'm and contracted coagulum, sur- roimded by a transparent colorless fluid. Q. What ai'e the principal ingredients of chyle? — A. A large proportion of albumen, a smaller one of fibrin ; a fatty substance or emulsion, which gives to chyle the appearance of milk ; and several salts, such as carbonate of potassa, mmiate of potassa, and pro- phospate of iron. Q. What change does the chyle undergo in its pas- sage along the various vessels ? — A. Its resemblance to blood increases in each of the successive stages of its progress towards the heart and lungs. Q. How are the chemical changes, and the contents of the intestines propelled through the tract of the alimentary canal ? — A. By the peristaltic action of tlie muscular coat of the same. Q. What becomes of the chyle after it has been pre- pared in the duodenum and fii'st intestines ? — A. It is received by absorption into the lacteals, and by them conveyed to the thoracic duct, which transmits it to large veins in the vicinity of the heart. (See distribu- tion of Ipnphatics.) Q. What do you understand by the " absorbent sys- tem?" — A. The absorbent system of vessels consists of two principal divisions, wliich may be compared to two sets of roots proceeding fi'om a common trunk ; one of these commences upon the walls of the intes- tmes, and is termed the " lacteal " system ; whilst the other takes its origin in various parts of the sub- stance of the organism at lai-ge, especially in the skin and subcutaneous textures, and is known as the " lym- phatic " system. Q. Where do the lacteals most numerously abound ? — ^. In the small intestines, below the point at which the Uver and pancreas discharge their secretions. Q. Where do the lacteals commence ? — A. Near the free extremities of the villi of the intestines. Q. In what way do they commence ? — A. It was formerly supposed that they commenced by orifices upon the internal smrface of the intestine ; but Carpen- ter, and other physiologists, contend that the lacteal vessels form loops by anastomosis with each other, so that they have no free extremity. Q. AVhat are the fimctions of the large intestines ? — A. They are engaged in the conveyance and expul- sion of feculent matter, and there are certain changes which take place in their contents, in aid of the object of nutrition, the exact nature of which has never been clearly determined. According to the best authority, it appears that some important changes are effected in that enlarged portion of the canal, termed coecum, and which has, by some, been regarded as a kind of sup- plementary stomach, in which fresh chvine is formed, and fi-esh nutriment extracted from the materials that have passed through the small intestines. The large 138 ANATOMY AND PHYSIOLOGY OF THE HORSE. intestines also extract nutriment from their contents, ■which is proved by the fact that nutritious matter in- jected into them has been known to support life for a certain time. SPLEEN. Q. What is the function of the spleen ? — A. It serves as a kind of diverticulum, to relieve the vessels of the digestive vescera when they are compressed by undue accummulation of the contents of then' canities, or when they are congested by obstruction to the flow of blood, through the hver or heart. It may also be considered as a lymphatic gland, for, in some instances in which animals have been allowed to survive longest after removal of the spleen, the lymphatic glands of the vicinity have been found greatly enlarged and clus- tered together, so as neai-Iy to equal the original spleen in volume ; hence, in such case we infer that its func- tion must be similar to that of the enlarged Ijmphatic Ln-EK. Q. What comprises the principal bulk of the hver — A. It is made up of a vast number of minute lobules of irregular form, but about the average size of a mil- let seed ; and each of them contains the elements of which the entire organ is composed, viz., a plexus of biliary ducts connected with their main trunks, and a mass of biliary cells ; each of which are connected in like manner with the three blood-vessels which unite to the circulation of this organ. Q. What are the vessels of the hver? — A. The hepatic artery, vena portse, and hepatic veins, to which may be added the excretory ducts and absorbents. Q. Of what use is the hepatic artery ? — A. It is the nutrient artery of the Hver. Q. Of what use is the vena portoe ? — A. It acts both as a vein and arteiy : as a vein, it receives the blood firom most of abdominal viscera ; as an artery, it rami- fies through the hver for the secretion of bQe. Q. What is the use of the hepatic veins? ---.4. They return blood to the vena cava. Q. What is the limction of the hver ? — A. It is an organ of excretion, designed to remove from the circu- lating fluid that portion of the products of disintegra- tion, of which the principal component of the urinary is the largest. Q. Into what substance is the greater part of the excrementitious matter converted ? — A. BUine. KIDNEYS. Q. What is the embryotic condition of the kidneys ? A. The kidneys are preceded in the embryo by a sub- stance first noticed by Wolff, and called after him the Wolffian bodies, or false kidneys, which originally ex- tend along the spine fi'om the heart to the end of the intestines ; but they afterwards become shorter, and after a time diminish by absorption, and wholly dis- appear. Q. TVTiat is the function of the kidneys ? — A. Their principal ftmction is to separate from the blood certain matters which would be injm-ious to it if retained. Q. What does the secretory surface of the kidneys consist of? — A. It is composed of epithehal cells which line the tubuh lu-inifera, which draw the pecuhar elements of the urinary excretion from the vascular plexus wliich surrounds the exterior of the tubes, car- rying ofl" the same to their terminations in the ureter. Q. What other arrangement is provided within the kidneys for the elimination of the superfluous fluid of the blood ? — A. K process of transudation takes place by the ftmction of malpighian bodies, whose thin- walled capillaries allow the transudation of water to take place, under a certain pressure, into the tubuli urinifera. SUPRA-RENAL CAPSULES. Q. What is the function of the supra-renal cap- sules ? — A. Their function has liitherto been involved in obscurity, and was supposed to be identical with other glands destitute of ducts or outlets ; but, lately, M. Brown Seguard has demonstrated that they play a very important part in the nervous system of the horse. T.VSA DEFEEENTIA. Q. What is the ftmction of the vas deferens ? — A. It is the excretory duct of the testicle, and conveys the semen to the vesiculse seminales. The author, instead of introducing examinations on the reproductive organs, has thought it best to substi- tute the opinions of that eminent ph}-isologist. Dr. Carpenter; and therefore the reader's attention is now directed to " Physiological considerations on the re- production of organized beings." REMARKS AND EXAMINATIONS ON THE EYE. The parts which compose the eye are divided into external and internal. The external parts are : First, the eyelashes, or cUia, which, in the horse, can scai-cely be reckoned more than one, there being very few hairs in the under eyelid. Secondly, the eyelids, or palpebrse, upper and under : where they join outwardly, it is termed the external canthus, and inwardly toward the nose, the internal canthus : they cover and defend the eyes. The cartilaginous margin or rim of the eyelid, from wliich tlie eye- lashes proceed, is named tarsus. In the tarsus and internal surface of the eyelid there are small glands, which secrete a fluid, to prevent friction of the eye and its lids, and facUitate motion. Tliirdly, the lachiy- mal gland, which is placed on the upper part of the eyelid toward the external canthus ; from this gland the tears are secreted, and conveyed to the inner surface of the vipper eyelid by several minute ducts, or canals, named lacluymal ducts. There is another small body, having a glandular appearance, in the inner corner of the eye ; on each side of which there are small orifices which are called puncta lachrymaha: these are the mouths or openings of two small canals, which, joining together, form a membranous tube ; and this, passing through a smaU open- ing in the bone, extends to the lower part of the nostril, where its termination may be distinctly seen in the horse. As the lachry- mal gland is constantly forming tears, it must be obvious that some contrivance is necessary to convey them off, and prevent them flowing over the cheek : this purpose is answered by the canal just described. When any irritating matter is applied to the eye, the tears are formed too abundantly to be carried off in this way ; they then flow over the cheek. In the human eye, the puncta lachrymalia terminate in a small sac, from which the lachrymal duct proceeds : this is not the case in the horse. Li the inner corner of the horse's eye is placed a body commonly termed the haw, no resem- blance to which is to be found in the hu- man eye. The horse has the power, by means of the muscles of the eye, to biing the haw completely over its siu-face ; it serves, therefore, as a second eyelid, and effectually wipes off any dust, hay, or seeds, or other matter which may have fallen upon the eye. The conjunctive membrane, or tunica conjunctiva, lines the inner surface of the eyelids, and covers the white part of the globe of the eye. This membrane has numerous blood-vessels, which are conspicu- ous when it is inflamed. The bulb or globe of the eye is composed of several coats and humors. The transparent cornea, which, in the horse, forms the front part of the eye, comprehends a larger part of the globe than in the human subject ; on removing this cornea, a fluid, which is named the aqueous humor, escapes, and the iris ap- pears. The iris is a. muscular curtain, having a hole in the centre, which is termed the pupU. This divides the fore part of the eye into two parts, named chambers, wliich are occupied by the aqueous humor. The pupU is of a dark bluish cast ; is of an oval, or rather of an oblong, form. The iris regu- lates the quantity of light that is required to pass through the pupil. For this purpose, it is composed of two sets of muscular fibres : by means of one the pupil is en- larged, and by the other it is diminished. Thus, if the pupil is first examined in the stable, where there is a moderate light, and immediately after in the sunshine, it will be found quite altered; being so small, in a strong light, as to be nearly closed. On re- 140 ANATOMY AND PHYSIOLOGY OP moving the iris, the second humor, or crys- talline lens, appeEirs : this is retained in its situation by a transparent membrane, named its capsule, between which and the lens is a minute quantity of fluid. The third humor of the eye is the A^treous. This humor is not contained in one general sac, but in numer- ous minute and perfectly transparent cells, and resembles pure water : tliis humor serves to produce a small degree of refrac- tion iji the rays of light, and occupies and distends all the posterior part of the globe of the eye. The next coat to the conjmictive is the sclerotica, or white of the eye, a strong, thick membrane, which extends from the transparent cornea to the optic nerve. The next coat to the sclerotic is the choroid. This is a delicate and very vascu- lar membrane. In the human eye it appears of a black color, and it is this which causes the pupU of the human eye to appear black ; but the choroid coat of the horse's eye is variegated in color ; in some parts black, in others blue, and in others green. The next coat is the retina : this is a delicate expan- sion of the optic nerve over the choroid coat, which it accompanies to the margin of the crystalline lens, and there terminates. The use of the retina is to receive certain impressions made by the light reflected from objects, so as to produce in the mind an idea of their figure and color; the optic nerve being the medium of communication between the retina and brain. From the above explanation of the mechanism of the eye, it wiU readily appear that many cir- cumstances may occur to render vision im- perfect, or to destroy it altogether. K the transparent cornea, for example, became white, light could not pass through it, and the aiaimal woidd be blind, however perfect the other parts of the eye might be. The cornea may be either too convex or too flat; in the former case, causing the animal to be near-sighted ; in the latter, producing an in- distinctness of vision with respect to objects that are near. The iris may, in consequence of disease, become fixed, or lose its power of motion ; in which case, the pupU would be always of the same size, and the animal would not have the power of adapting it to the various distances or objects; or, as sometimes happens, the pupil may become quite closed, by which light would be per- fectly excluded from the retina. Supposing the cornea and iris to be healthy, the crys- talline lens, or its capstile, may become opaque, and thereby cause total blindness. But in this part, as in the cornea, we meet with different degrees of opacity : some- times it is very slight, the pupil appearing of a lighter color, and unusually large : in this state, the pupU is said to look duU or muddy, which causes the horse to start ; but when the opacity is complete, it consti- tutes the disease termed cataract. There is another disease, to which the reader's at- tention is called ; it is named gutta serena, or amaurosis. This disease is known by the pupU being unusually large or open, and by its continuing so when the eye is exposed to a sfrong light. EXAMINATIONS OF THE ORGANS OF SIGHT AND THEIR APPENDAGES. Q. Where are the eyes located? — A. Within the orbits. Q. By what foraminse is each orbit perforated? — A. By the optic foramen. Q. From ■whence is the lining membrane of the orbit derived ? — A. From the dura mater and periostemn. Q. Enumerate the appendages of the eye. — A. The eyeHds, eyelashes, muscles of the eyelids, tarsal carti- lages, meibomian glands, timica conjunctiva, membrana nictitans, laclnymal gland, puncta laclir}TnaHa, lachrj'- mal sac, ductus ad nasum, and the muscles of the eye- balL EYELIDS. Q. "What parts do the eyeUds occupy ? — A. The cir- cumference of the orbits and front of the eyeball. Q. What are the eyehds composed of? — ^. In com- position they are cuticular, muscular, cartilaginous, and membranous ; also glandular, vascular, and nervous. Q. What muscle enters into the composition of the eye? — A. The orbicularis palpebrarum. Q. How are the Uds separated ? — A. By a transverse fissure, bounded by the angles or canthi of the eye. Q. What is, attached to the superior or temporal angle ? — A. The tarsal ligament. EXPLANATION OF FIGURE XIV. HINT) EXTEEMITIES. to'. Tensor vagina. k'. Rectus. o'. Vastus externus. q'. Flexor metatarsi. )•'. r. Gastrocnemius internus. s'. " externus. t'. Flexor pedis accessorius. «'. Insertion of the gastrocnemius. »'. Flexor metatarsi. x'. Extensor pedis. ^. y\ Extensors. u. V. Tendo perforans et perforatus. K'. K'. Abductors tibialis. J'. J'. Triceps. v. I'. Adductors. Sf. Hoof. 5, 5. Saphena vein. 8. Bifurcation of the suspensory ligament. 1)'. (Off-hind leg.) Peroneus. z. Suspensory ligament. FORE EXTREMITIES. J". Pectoralis magnus. m". n". Triceps extensor brachii. o". Pectoralis transversah.'!. p". p". Flexor metacai'pi externus. gr". " " mcdius. r". " " internus. s". Extensor metacarpi magnus. u". (At the upper pai-t of the figure.) Levator humeri. u". u". u". u". V. (Beneath the olecranon and carpus.) Flexors perforans and per- foratus. x". Extensor pedis. y". y. Extensor suffraginis. z". Suspensory ligament. i,: The hoof. 4. Subcutaneous thoracic vein. 6. Radial vein. 8. Bifurcation of the suspensory ligament. THE HORSE. 141 Q. AVhat is fixed to the inferior angle? — A. The tendon of the orbicularis. Q. From whence is the loose portion of sldn, enter- ing into the composition of the upper lid, derived ? — A. It is a prolongation of the skin covering the forehead. Q. From whence is that of the lower lid derived? — A. From the mtegimients of the face. Q. How are the internal sm-faces of the lids shaped ? — A. Into concavities which adapt them to the convex- ity of the globe of the eye. Q. By w^hat membrane ai-o the Kds lined? — A. By the conjunetirial. T.UIS.U. C.UlTIL.iGES. Q. "What are the tarsal cartilages ? — A. They enter into the substance of the borders of the Kds, imparting to them both fii-mness and elasticity. Q. Describe the tai-sal cartilages ? — A. The superior cartilage is broader and more convex than the inferior ; they correspond in shape and size to their respective lids ; they are convex outwardly and concave inwardly, and are inserted into the rims of the orbits. Q. What is the texture of the tarsus? — A. Their texture is tibro-cartilaginous. MEIBOMIAN GLANDS. Q. Describe the meibomian glands ? — A. They have the appearance of white follicular bodies, vertically ranged m parallel lines ; they vary both in caUbre and length, and are m the upper rather than the lower hd. Q. What is the function of the meibomian glands ? — A. To secrete a fluid which guai'ds against friction between the eye and its appendages. TUNICA CONJUNCTIVA. Q. What is the situation of the tunica conjunctiva ? — A. It is the lining membrane of the eyehds, mem- brana nictitans, caruncula lachrymalis, puncta lachr)'- maKa, and is reflected to the globe of the eye. Q. Describe the conjunetirial surface ? — A. The adhe- rent one is rough, lax, and flocculent ; the outer surface is smooth, glossy, and humid with secretion. Q. What are the peculiarities in the organization of the conjunctiva ? — A. It is a continuous membrane, yet varies in textm-e, as follows : 1st. That portion which gives a covering to the conjunctiva palpebralis is higlily organized with blood-vessels, and is often tinged a deep red color. 2d. The conjunctiva sclerotica is not 60 highly organized, yet has a few straggling vessels of larger caHbre than those of the former, and its textm-e is more dense. 3d. The conjunctiva cornea? is thin and transparent, more of a homy textui'e, and has no ajjpearance of vascularity. MEMBRANA NICTIT.UCS. Q. TlHiat is the common name for the above mem- brane ? — A. The haw. Q. What is its structure ? — A. Cartilaginous. Q. What is its situation ? — A. It is located behind the inferior canthus, between the eyeball and side of the orbit. Q. What is its figm-e ? — A. It approaches that of an extended triangle, of wliich the short side is turned forwards, and the lengthened angle backwards. Q. Describe the anterior part ? — A. It is thin and elastic, and bounded by a crescentic edge, tei'minating in two salient angles ; it increases in substance, but grows naiTow posteriorly, and there ends in an obtuse conical point, which appears in the adipose tissue at the bottom of the orbit. Q. What is the form of its smfaces ? — A. Inwardly concave ; outwardly convex. Q. What is the body of the nictitating membrane clothed with ? — A. By a portion of conjimctirial mem- brane. Q. What is the function of the membrana nictitans? — P. To protect the eyeball, in the removal of foreign bodies from its surface. L.iCimYMjil. APPARATUS. Q. What parts compose the lachmyTal apparatus? — A. The lachi-ymal gland, caruncula lachrymalis, lachi-y- mal puncta and conduits, lachr)-mal sac, and ductus ad nasum. LACHETJIAL CLASV. Q. Where is the lachrj-mal gland situated? — .4. In a depression, beneath the process of the orbital arch. Q. What are its coverings, and with what is it in contact ? — A. It is covered by the common aponeurotic lining of the orbit ; it is in contact with the levator pal- pebrae, and is enveloped m fat and cellular membrane. Q. What is its form? — A. It is irregular, slightly convex superiorly; inclining to the concave iuferiorUy. It is a conglomerate gland, constituted of many lobules. Q. Have the lobules any further organization ? — A, Yes, they are composed of minute granules. Q. What vessels do the granules receive and what springs from them ? — A. They receive the terminating ramifications of the supplying arteries, and from them spring the radicles of the excretory ducts. Q. What do the radicles terminate in, and where is their outlet? — A. The radicles unite ■nith one another into a set of tubes, which open upon the conjunetirial Kning of the upper h'd m the som-ce of seven risible orifices near its superior angle; this is their outlet. Q. What is the function of the lachrjTnal gland ? — A. To secrete the tears. Q. What becomes of the superfluous teai's ? — A. They either fall over the lower Hds, or pass into the lachrvTnal sac ; from thence, by the ductus, to their out- lets -Nrithin the nostrils, at theii- inferior parts. CUIUNCULA LACHEYM.U,IS. Q. What is the caruncula lachrjinalis ? — A. It is a small eminence, lodged within the inferior canthus, be- tween the eyeball and Hds. Q. What is its use ? — A. It secretes a light yellow unctuous matter, with which the fine hairs on its surface 142 ANATOMY AND PHYSIOLOGY OF being coated it detains any small foreign bodies that may float in the laehrj-mal secretion ; it also directs the latter fluid into the puncta. LACHRYMAL PUNCTA AND CONDUITS. Q. What are the puncta lachrymalia ? — A. Two small orifices situated on the inward margins of the two lids — superior and inferior — near the radix of the caruncle. Q. What do the puncta terminate in? — A. The lacha-jTnal conduits. Q. AVhat is theu' situation ? — A. Within the sub- stance of the eyelids. Q. How are conduits formed ? — A. A minute carti- laginous circle surrounds them, and they are lined by conjunctiml membrane. Q. What do the conduits terminate in? — A. The lachrymal sac. LACHRYMAL SAC. Q. Where is the lachrj-mal sac situated ? — A. AVith- in the depression which leads into the channel of the lachr)-mal bone, behind and below the small eminence upon the orbital ridge of that bone. Q. Describe the sac and its connections ? — A. It is an oblong membranous bag; its front is crossed by fibres of the orbicularis ; it has also a connection with the tendon of that muscle. The posterior part of the sac adheres fii-mly to the lachrjinal bone. It is com- posed of a dense, wliite, fibrous membrane, furnished with a lining from the conjuncti^'ia. Q. By what is this sac perforated? — A. By the lachi-ymal conduits. Q. AVhat does it open into ? — A. Into the ductus ad nasum. Q. What is the function of this sac? — A. It is a reservoir into wliich the tears flow from the lachrymal conduits, and from thence pass into the ductus ad nasum. DUCTUS AD NASUM. Q. AVhat is the ductus ad nasum ? — A. It is a long membranous canal, commencing at the contracted por- tion of the lachrjinal sac, and running ^^ith the groove thi'ough the lachrjTnal bone ; then along a canal in the superior maxillary bone, between it and the anterior tiu'binated bone ; terminating at the inner and inferior part of the nostril. Q. What is the organization of the ductus ? — A. It appears to be a continuation of the membrane com- posing the lachrymal sac, wliich is strengthened by a fibrous sheath ; its internal surface is probably mucous, ■which protects it from the action of the tears, or lachrymal secretion. Q. Describe the course of the tears, or lachi-j-mal secretion? — A. They are secreted by the lachrjinal gland, and are poured by its excretorj' ducts over the surface of the eyeball ; the puncta lachrj-malia absorb them ; they are then conveyed by the lachrj-mal ducts to the lachi'jTnal sac ; and through the ductus ad nasum pass into the nostril. THE EYEBALL AND ITS COATS. Q. What is the form of the globe of the eye ? — A. Nearly of a spherical figure. Q. Of what is the globe of the eye composed? — A. Of membranes, or coats, fiUed with humors or fluids, which preserve its form. Q. How many coats has the eye ? — A. Five : the sclerotic, choroid, retuia, cornea, and iris. Q. Does not the tunica conjunctivia enter into the composition of the membranes of the eye? — A. Yes: it may be considered as common to both. Q. Where does it adhere most closely ? — A. Over the cornea. SCLEROTIC COAT. Q. What is the use of the sclerotica ? — A. It bounds the form of the eye, protects and supports the parts within. Q. What are its perceivable boundaries ? — A. It ex- tends from the optic nerve to the cornea. Q. What is mserted mto it posteriorly? — A. The fleshy part of the retractor muscle. Q. What is inserted into its anterior margins ? — A. The tendons of the foiu: recti. COKNE.V. Q. What is the cornea ? — A. The transparent, ante- rior part of the globe of the eye. Q. How does its form compare with the sclerotica ? — P. It is more convex. Q. What covers its convex surface ? — A. The con- junctiria. Q. What is its sti-ucture ? — A. Laminated. Q. 'WTiat is the iris ? — A . It is a circular membrane, with an irregular central cavity, in the anterior chamber of the eye. Q. What is its central perforation called ? — A. The pupQ. Q. How is the periphery of the pupU bounded ? — A. By several dark, colored, glandular bodies, termed cor- pora nigra. Q. What is the use of the iris ? — A. By contracting it excludes all superfluous rays of light, and by expand- ing admits thr-ough the pupU all that pass thi-ough the cornea. Q. What is the structm-e of the iris? — A. It is a fibrous membrane, dirisable into tivo layers, pro\ided with blood-vessels and nerves. CHOROID COAT. Q. What is the choroid coat? — -4. It is a dark- colored membrane of deUcate texture, located immedi- ately beneath the sclerotica. Q. AVhat are its boundaries? — A. It extends from around the termination of the optic nerve as far for- ward as the edge of the cornea, and ends in the ciliary circle. THE HOESE. 143 Q. How is the choroid coat connected with the sclerotica? — A. By cellular membrane. Q. What is remarkable at its outer edge? — A. It is throMTi into folds, called ciliary ch'cle and processes. Q. What is the color of the choroides ? — A. Extern- ally, its whole surface is black f internally, the anterior parts are black, and the posterior half is of a brilliant variegated green. Q. What is the black part termed ? — A. Pigmentum nigrum. Q. What name is given to the variegated part ? — A. Tapetum lucidiun. Q. What difference do we observe in the pigment of the choroid siu-faces ? — A. The inner layer is thicker and more consistent than that found on the outer surface. RETINA. Q. yVhat is the retina ? — A. It is the third or inner- most tunic of the eye. It cannot, however, be con- sidered as a tunic, for it pervades the interior of the globular expansion without contracting any adhesions until it has reached the corpus ciHare. Q. How is the retina formed ? — A. The optic nerve, hanng reached the inner and inferior part of the globe of the eye, enters the sclerotic and choroid coats, and in its passage thi'ough them its diameter contracts • having arrived at the inner part of the globe, the neiTe forms an eminence, from the circumference ofwliich issues radiating fibres which form the retina. Q. How is the retina sustained in this state of globu- lar expansion? — A. By the humors of the eye, which keep it in contact ■nith the choroides. Q. The retina having radiated on the interior of the globe, where is it inserted ? — A. Into the corpus ciharc. HUMORS OF THE EYE. Q. Of how many humors does the eye consist, and what are their names ? — A. Of three ; they are called aqueous, ciystalline, and v-itreous humors. Q. What parts of the eye do they occupy? — A. They occupy in succession the spaces termed anterior, middle, and posterior chambers of the eye. AQUEOUS HUMOR. Q. What is the use of the aqueous hiunor? — A. It transmits the rays of light, and aids the fi-ee motions of the iiis. Q. What are its boundaries ? — A. It fills the interval between the cornea and crystalline lens. Q. Describe the aqueous humor. — A. It is a bright limpid fluid, and in properties bears some resemblance to the vitreous. Q. What is the composition of both these humors ? — A. They are composed of albumen, gelatine, and miuriate of soda, held in solution by an aqueous men- struum. <^. How is this fluid secreted ? — A. By secretion from the transparent walls of its capsule. CRYST.U.LINE LENS. Q. What is the use of the crystalline lens? — A. It concentrates the rays of Ught, so as to make a distinct image in the posterior chamber. Q. Where is the crj-staUine lens situated ? — A. Be- tween the aqueous and vitreous humors. Q. By what is the crystalline lens enclosed ? — A. By a tunic, called tunica crj'staDina. VITREOUS HUJIOR. Q. ^^Tiat is the vitreous humor, and where is it situ- ated? — A. It is the most bulky himior of the eye; of a jelly-like consistence, yet quite transpai-ent, and occu- pies that portion of the eyeball posterior to the crystal- line lens. Q. What is the use of the vitreous himior? — A. It supports the form of the eye, and maintains the other hum«rs in their proper positions. RESPIRATORY SYSTEM. PHYSIOLOGICAL CONSIDERATIONS. Respiration and Structure of the Lungs. — The organs of respiration are the larynx, the upper opening of which is named glottis, the trachea or windpipe, bronchia, and the lungs. The air is displaced out of the lungs by the action of the muscles of respiration ; and, when these relax, the lungs expand to a certain calibre by their elasticity. Tliis may be exemplified by means of a sponge, which may be compressed into a small bulk by the hand, but, upon opening the same, the sponge returns to its natural size, and all its cavities become filled with air. The purification of the blood in the lungs is of vital importance, and indispensably neces- sary to the due performance of aU the func- tions. When the lungs, and muscles con- nected with them, are in a physiological state, the horse is said to be in good wind — a very desirable state for an animal to be in, whose usefulness depends on his being capable of a long continuance of quick motion. The trachea, or windpipe, after dividing into bronchia, again subdivides into innumerable other branches, the extremities of which compose an infinite quantity of small cells, which, with the ramifications of the veins, arteries, nerves, lymphatics, and the connecting cellular membrane, make up the whole mass or substance of the lungs. The internal surface of the windpipe, bron- chia, and air-cell, is lined with a membrane, which secretes a mucous fluid : when, in con- sequence of an obstructed surface, this fluid becomes abundant, it is expelled by the nostrils. The whole is invested with a thin, transparent membrane, named pleura : the same membrane lines the internal sur- face of the ribs and diapliragm, and, by a duplicature of its folds, forms a separation between the lobes of the lungs. RESPIRATORY SYSTEM. The function of respiration is the conver- sion of venous into arterial blood. This arterialization of the venous blood is a pro- cess highly essential to the well-being of all animals ; more important is it than the assimulation of aliment ; for a horse may live several days without food, yet cannot exist many minutes unless his blood be arterialized. In considering the function of respiration, our attention is first turned to the mechani- cal means by which the air is alternately admitted and discharged from the lungs. The mechanical act of respiration is divisi- ble into two periods, that of inspiration, during which air is drawn into the lungs so as to increase its volume and distend its parenchema and expiration, diuring which process the air which had been so received is expelled. Inspiration is accompanied by enlarge- ment of the capacity of the thorax in its various dimensions. This is effected by the action of different sets of muscles, operated on by the nervous system. The principal muscle of inspnation is the diaphragm. Among the secondary muscles employed in inspiration are those which articulate the ribs, viz., the intercostales. Each rib is capable of a small degree of motion on the extremity by which it is articulated with the vertebrae. This motion is chiefly for- ward and backward ; the intercostal muscles favor this motion, as they are disposed in two layers, each passing obhquely, but with opposite inclinations, from one to the adja- cent rib. There are two ways in which the (144 J ANATOMY AND PHYSIOLOGY OP THE HORSE. 145 chest may be dilated: first, by the dia- phragm ; and secondly, by the intercostales, which elevate the ribs. In natural respira- tion, the horse breathes cliiefly through the aid of the diaplrragm. Should the respira- tion become quickened, the intercostales are employed, and, when the respiration is labo- rious, the axillary muscles of the abdomen, back, and sides, are brought into use. The glottis is opened during inspiration by the muscles of the larynx. The expulsion of the air from the lungs constitutes expiration. This takes place as soon as the air which has been ex- pired has parted with its oxygen, and re- ceived in return a certain quantity of car- bonic acid gas and vapor. In regard to the elasticity of the lungs, it is now demon- strated that they possess no inherent power of elasticity other than that common with all membranous textures. Hence, if an opening be made in the sides of the chest, the lobes on this side collapse inconsequence of the pressure of air from without. We have next to inquire what changes have, in the meanwhile, been effected in the blood by the action of the air to which it has been subjected in the lungs. A visible alteration, in the first place, is produced in its color, which, from being of a dark pm'- ple, nearly approaching to black, when it arrives at the air-cells by the pulmonary arteries, has acquired the bright, intensely scarlet hue of arterial blood, when brought back to the heart by the pulmonary veins. In other respects, however, its sensible qualities do not appear to have undergone any material change. Judging from the changes produced on the air which has been in contact with it, we are warranted in the inference that it has parted with a certain quantity of carbonic acid and of water, and that it has in return acquired a certain proportion of oxygen. Since it has been found that the quantity of oxygen absorbed is greater than that which enters into the composition of the carbonic acid evolved, it is obvious that at least the excess of oxygen is dh-ectly absorbed by the blood ; anu this absorption constitutes, no doubt, an essential part of its arterialization. It has been much disputed whether the combination which seems to be effected be- tween the oxygen of the air and the carbon furnished by the blood, occurs during the act of respiration, and takes place in the air-cells of the lungs, or whether it takes place in the course of circulation. On the first hypothesis, the chemical process would be very analogous to the simple combustion of charcoal, which may be conceived to be contained in the venous blood in a free state, exceedingly divided, and ready to combine with the oxygen of the air, and imparting to that venous blood its charac- teristic dark color ; while arterial blood, from which the carbon had been eliminated, would exhibit the red color natural to blood. On the second hypothesis, we must suppose that the whole of the oxygen, which disap- pears from the air respired, is absorbed by the blood in the pulmonary capillaries, and passes on with it into the systemic circu- lation. The blood becoming venous in the course of the circulation, by the different processes to which it is subjected for sup- plying the organs with the materials re- quired in the exercise of their respective functions, the proportion of carbon which it contains is increased, both by the abstrac- tion of the other elements, and by the addi- tion of nutritive materials prepared by the organs of digestion. The oxygen, which had been absorbed by the blood in the lungs, now combines with the redundant carbon, and forms with it either oxide of carbon, or carbonic acid, which is exhaled during a subsequent exposure to the air in the lungs. Many facts tend strongly to confirm our belief in the latter of these hypotheses. OF THE LARYNX.* The larynx is the organ producing the voice of the animal. Situation. — It is joined to the top of the trachea (or windpipe), and is placed in the throat, between the posterior and broadest * Percivall's Hippopathology. 146 ANATOMY AND PHYSIOLOGY OP parts of the branches of the lower jaw; having the pharynx and uppermost part of the esophagus situated above it ; the supe- rior portions of the sterno-hyoidei and thy- roidei below it ; the tongue, with its muscles, and the os hyoides, in front of it ; and the trachea issuing from below and behind it. Attachment. — The larynx is retained in its place by its connection with the os hyoides and pharynx ; by its muscles ; and by its coalition with the trachea. Conformation. — The larynx has so com- plete a fleshy covering, that it is not until it is divested of its muscles (which have been heretofore described) that it is dis- covered to be composed of five pieces of cartilage, so joined together as to be move- able on one another, and open both supe- riorly and inferiorly, to admit of the passage of air into and out of the trachea. These cartilages have received the names of thyroid, cricoid (two), arytenoid, and epiglottis. The thyroid or shield-like cartilage, by much the largest of the five, forms the superior, anterior, and lateral parts of the larynx. It consists of two broad lateral portions, continuous and prominent at the upper and anterior part of the neck, the promiirence corresponding to which in human anatomy has received the name of pomurti Adami. Below this pomt of union the divisions recede from each other, leaving a triangular space between them, which is occupied by a ligament denominated the Ugamentum crico-thyroideum. The four projecting corners from the posterior parts of the thyroid cartilage are named its cornua: the two superior are joined by cap- sular articulations to the body of the os hyoides ; the two inferior are connected by very short capsular ligaments to the cricoid cartilages ; the union of all wliich parts re- ceives additional strength firom expansions of membrane. At the roots of the superior cornua are two forainina, that give passage to nerves, of considerable importance, to the interior of the larynx. This cartilage not only constitutes by far the most exten- sive part of the larynx, but, as its name in- dicates, incloses and shields from external injury all the others. The cricoid or ring-like cartilage is placed below the thyroid. In front it appears like part of the trachea; but it broadens so much behind, that it overlaps the first ring of the w^indpipe, somewhat after the form of a helmet. Upon its broad or posterior part are four surfaces of articulation : the two upper receive the hinder extremities of the arytenoid cartilages, the two lower are adapted to the inferior cornua of the thyroid cartilage : they are all furnished with cap- sular ligaments and synovial membranes. Furthermore, it is attached by hgaraentous expansions to those parts, and likewise to the first ring of the trachea. The two arytenoid, or eiver-shaped car- tilages, triangular in their figure, lie over the upper and back part of the trachea, leaving an aperture between them leading into that canal, denominated, from its prox- imity to the tongue, the glottis. Their inward parts are everted, and form a tri- angular prominent border, over which is spread the membrane of the glottis : their outward surfaces are marked by concavities in which are lodged the arytenoid muscles. Posteriorly, they repose upon the cricoid cartilage, and are connected with them by capsular articulations : in front, they have a membranous connection with the cartilage next to be noticed. The epiglottis, so named from being raised over the glottis, and occasionedly covering it like the lid of a pot, is well adapted, from its heart-like shape, to the rima glottidis; whose margin is completed by two narrow slips of cartilage proceeding from the base of the lid to the arytenoid. By some, these slips of cartilage have been separately considered: but in my opinion improperly so ; for they are, in reality, nothing more than prolongations or appen- dices of the epiglottis. The surface of this cartilage presented to the interior of the larynx is smooth and concave, and covered by an extension of membrane from the glottis ; that part opposed to the tongue is THE HORSE. 147 unevenly convex, and is tied to that organ, I as well as to the os hyoides, by a doubling of membrane infolding some muscular fibres: to this musculo-membranous ligature, which assists in retaining the cartilage in its elevated position, the name oi frcenum epiglottidis is properly given. The fraenum receives co-operation in this fmiction from strong elastic Ligaments connecting the base of the epiglottis to the thyroid and arytenoid cartilages. If we detach the epiglottis, or raise it forcibly in order to obtain a more complete view of the rima glottidis, the latter will be found to be stretched into an oblong quad- rilateral figure, whose width gradually diminishes from the middle towajrds either extremity, and bears a ratio of about one to six when compared to its length. The sides turned forward are formed by the arytenoid cartilages ; those directed back- ward by two prominent folds of membrane (which envelop the thyro-arytenoid mus- cles), commonly described as the vocalliga- ments, from their being concerned in the formation and intonation of the voice. Immediately over them are slit-like aper- tures, opening into membranous sacs, each large enough to contain a walnut ; these are the ventricles of the larynx, whose use is also connected with the production and modulation of the voice. The membrane lining the cavity of the larynx is one of great susceptibility; on which account it is kept continually moist by a mucus, oozing from numerous lacuna — the excretory orifices of small subjacent follicles whose situation is denoted by the little round eminences upon its surface. This is the common seat of that species of catarrh wliich is accompanied by cough. OF THE TRACHEA. The trachea, or windpipe, is a cartilagin- ous tube extending along the neck, from the larynx to the lungs, for the passage of air. In horses of ordinary size, it is from twenty-five to thirty inches in length. Course. — The trachea commences from the inferior border of the cricoid cartilage, opposite to the body and transverse proces- ses of the atlas ; takes its coui-se along the anterior and inferior part of the neck, inclin- ing to the near side, between the sterno- myloidei muscles (which by their approx- imation conceal the lower portion of it), and enters the chest between the two first ribs; wherein, under the curvature of the posterior aorta, it divides into two parts the bronchial tubes. Structure. — From fifty to sixty annular pieces of cartilage enter into the com- position of the windpipe ; altogether con- stituting a stnicture so remarkable, for the inequality or asperity of its exterior, that the ancients, in order to at once distinguish it from aU other vessels, called it the aspera arteria. No entire or undivided tubular substance could have partaken of the various motions of the head and neck, without having suffered more or less distortion, and consequent deformity and diminution of cafiber, of some part of its canal, which would have been attended with frequent in- terruptions to the free passage of the air, dangerous, and even fatal, to the respiratory functions; w^hereas, constructed as it is, with the aid of its muscular power, no at- titude into which the animal may naturally put himself will impede the freedom of pas- sage tlnough it. The cartilages, or, as they are commonly described, the rings of the windpipe, have all a close resemblance to one another : if there be any disparity be- tween them worthy of notice, it consists in those that form the superior part of the pipe being somewhat larger and broader than those nearest to the bronchial tubes.* A ring is not uniform in its breadth, in conse- quence of having waving or scolloped bor- ders ; the advantage of which is, that a sort of dove-tailed connection is effected which materially contributes to the compactness and strength of the entire structm-e. Its front and sides measure, in the broadest places, half an inch in breadth, and nearly a * Now and then we find, at the upper part of the tube, two or three or more of these rings accreted together ; it gives rise to some prominence thereabouts generally, and may often be detected by taction in the living animal. 148 ANATOMY AND PHYSIOLOGY OF quarter of an inch in thickness — evidently made so substantial to resist external in- jury; whereas its posterior or unexposed parts grow suddenly thin and yielding, and taper to the extremities ; which, instead of meeting and uniting, pass one over the other, and thus form a shield of defence behind, while they admit of a certain dila- tation and conti-action of the internal dimensions of the tube. These attenuated ends are joined together by a ligamentous expansion, mingled •with a quantity of cel- lular membrane. The rings are likewise attached to one another by narrow ligamen- tary bands, sti'ong and elastic ; which, after they have been drawn apart in certain posi- tions of the head and neck, have the power to approximate them ; when the pipe is re- moved from the body, and suspended by the uppermost ring, these ligaments coun- teract the tendency its weight has to sepa- rate the rings, and still maintain them in apposition. The lowermost ten or twelve pieces of cartilage appear on examination but ill to deserve the name of rings ; indeed, they are little more than semi-annular, the deficiences in them behind being made good by intermediate moveable pieces of cartilage. These pieces, whose breadth in- creases as we descend, are let into the vacuities in such manner as to overlap the terminations of the segments, and they are confined and concealed by the same sort of ligamentary and cellular investment as was before noticed. Muscle. — 'Where the outward extremity of the ring suddenly turns inward, and de- generates into a thin flexible flap on either side, a band of muscular fibres is fixed and stretched across the canal, dividing it into two unequal semi-eUiptical passages. The anterior one is the proper air channel ; the posterior or smaller one is filled with a fine reticular membrane, connecting the band to the posterior part of the ring, and preventing i in action, from encroaching upon the main conduit. This self-acting band appears to me to have been added to the tube to enable it to enlarge its caliber — not to diminish it, as a superficial view of these parts might lead one to imagine ; for, in consequence of the passage being natmally elliptical, and the muscle being extended across its long diameter, the contraction of its sides wiU give the tube a circular figure, by increasing the curvature of the ring anteriorly, and thereby, in effect, will expand and not con- tract the cafiber of the canal. I would say, then, that the trachea was made muscular in order that it might have the power of in- creasing its capacity for the passage of air, whenever the lungs were called into extra- ordinary action : in addition to which, I think that this band may, in some degree, counteract any tendency certain positions of the head and neck have to alter its shape and diminish its circumference. This opinion is corroborated by the circumstance, that the muscle grows slender and pale as we approach the lower end of the pipe, where the canal itself is nearly circular, and where it is placed in the least moveable part of the neck.* Membrane. — The trachea is lined by a soft, pale red membrane, which anteriorly has a close adhesion to the rings them- selves, and presents a smooth, polished in- ternal surface ; but which, posteriorly, is loosely attached to the muscular band, and puckered into fourteen or fifteen longitu- dinal plica; or folds, that extend with regu- larity from one end of the tube to the other. These folds were evidently made to allow of the contraction and elongation of this muscular band ; for I cannot myself assign any reason why they should exist in its relaxed state, unless this fulness of membrane be given to admit of enlarge- ment of the caliber of the tube during the contractions of that muscle ; if this be plausible, I may adduce the corrugation of the membrane as another proof that the caliber of the trachea is susceptible of aug- mentation. This membrane is continuous with that which clothes the rima glottidis ; * In this opinion, says Mr. Percirall, I find I am at variance with Girard. The French professor ascribes to it the power of contracting the caliber of the trachea. " Cetto couche, bien evidemment musculeuse, pent retrecir le cal- ibre de la trachee, en rapportant les extremites des seg- mens." — Anat. Vet.,-p.liG et 147, torn. ii. THE HOBSB. 149 but it is paler than it, and not near so sensi- tive. Its arterial ramifications, also less abundant than upon the glottis, exhale a vapor from its surface ; independently of which, it is kept continually lubricated by mucus, furnished from its numerous lacuna, to defend it from anything acriminous that may be contained in the breath. Bronchial Tubes. — The trachea having entered the thorax, bifurcates into the tw^o bronchial tubes. Of them, the right is the more capacious canal, on account of having communication with the larger division of the lungs ; the left the longer one, in conse- quence of having to cross under the pos- terior aorta, in its course to the left division of the lungs. The last cartilage of the main pipe has a spear-like or angular pro- jection extending down between the bron- chial tubes, filling up that space which would otherwise be left open from the di- vergent manner in which they branch off: it is quite loosely attached, in order that the branches may accommodate themselves to the motions of the neighboring parts. The bronchial tubes vary in structure from the trunk that gives origin to them : instead of their rings being formed of entire pieces of cartilage, they are constituted of several separate pieces, making up so many segments of the circle, overlapping one another, and united together and invested by an elastic cellular substance : they also differ in hav- ing no muscular band, another fact connected with the physiology of that part. The bron- cliial tubes, in penetrating the substance of the lungs, subdivide — the right into three principal branches, the left into two ; from which spring innumerable others, that grow smaller and smaller, untU the ramifications become so reduced that they are no longer traceable by the naked eye. In the larger branches, we may dissect out five and even six segments of cartilage, held together by a thin but dense and elastic ceUular sub- stance ; in the smaller divisions, only two are found, and they are diminished in size ; and, in the smallest visible ramifications of all, cartilage is altogether wanting, though, in many places, marks of the rings may be traced upon the continuation of the lining membrane, which in these intimate parts compose the entire parietes of the tube. In the larger branches this membrane (which is continuous throughout the bron- chial system) assumes a plicated disposi- sition — apparently, to admit the more readily of expansion. THYROID GLANDS. Two egg-shaped, apparently glandular bodies, attached just below the larynx to the sides of the trachea, and united in front of that tube by an intervening portion of the same substance, which, by way of dis- tinction, is by some called the isthmus. They are enveloped and attached in their situation by cellular membrane ; are larger and more vascular in the young than in the old subject ; and exhibit a spongy tex- ture, when cut into, which I am at present ignorant of the precise nature of. They are well -supplied with blood-vessels, and have many small nerves going to them. Their physiology still remains obscure. OF THE LUNGS AND PLEURA. The lungs are the essential organs of res- piration : the pleura is but the membrane by which they are invested. PLEURA. The pleiua is a fine, semi-transparent membrane, lining the cavity of the chest, and giving a covering to the lungs. By that portion of it which is called the medi- astinum, the cavity is divided into the right and left sides of the thorax. General Conformation. — If the lungs be exposed, by breaking off one or two of the ribs, we shall perceive that their surface, as well as that of the cavity itself, is every- where smooth, poKshed, and humid. This is owing to the extensive investment of the pleura, the surface of which is now pre- sented ; so that, in reality, without break- ing the surface, nothing but pleura can be touched ; although, from its extreme tenuity and pellucidity, the viscera appear, on a superficial view, to present their own bare 150 ANATOMY AND PHYSIOLOGY OP exterior. Its other side, on the contrary, is rough, having numerous cellular flocculent appendages, by which it is united to the. parts it invests ; and so close and firm are these adhesions, that to cleanly detach it, in the recent subject, is a very difficult and tedious dissection. The pleura is a reflected membrane ; by which is meant, one that not only lines the cavity in which the viscera lie enclosed, but by dupLicature, or what in anatomical lan- guage is called reflection,, gives a partial or complete covering to the contained organs themselves. It is evident, therefore, that such a membrane admits of division into two portions — a lining or parietal, and a reflected portion ; and these, with regard to the pleiua, have, for the sake of more de- finite description, received the names of pleura costalis and pleura pulmonalis. They are both, however, continuous at aU points, are precisely similar in structure and func- tion, and, in fact, are stUl but one and the same pleura. Mediastinum. — There is yet a third por- tion of this membrane to which a distinct appellation has been given, and that is the mediastinum, the membranous partition be- tween the cavities or sides of the thorax ; it differs from both the others in being com- posed of two layers, which are derived from the two pleursB of the opposite sides. If we conceive the pleurae of the two sides of the thorax to be perfect sacs or bags, with flattened sides tmrned inwardly, and closely applied and united together, in such a manner that the double membrane formed by then- union extends through the middle of the chest, from the dorsal vertebra to the sternum, we shall at once have a toler- ably correct idea of the formation as well as situation of the mediastinum. Structure. — The pleura, from the nature of its secretion, is one of those included in the list of serous membranes to which it has been demonstrated also to be similar in its intimate organization. Like them, it presents a shining secreting surface, of a whitish aspect, and considerable transpa- rency, and is composed of httle else than condensed cellular substance, whose texture is penetrated by blood-vessels, absorbents, and nerves : by long maceration in water, indeed, it may be entirely resolved into cel- lular substance. In most parts it is ex- tremely thin, and by no means tough ; but it is not so in all, for that portion which faces the diaphragm is much denser and stronger than the pulmonary or costal di- vision of it. Organization. — The arteries of the pleura, which come from the adjacent parts, are in the natural state exceeding small, ad- mitting only the colorless parts of the blood — a circumstance that accounts for its pel- lucidity ; under inflammation, however, they contain red blood, and such is the ex- planation of that arborescent vascularity upon the sides of the thorax in horses that die of pnuemonia ; than which state noth- ing can better demonstrate the comparative number and distribution of these blood-ves- sels. The majority of them terminate in exhalent orifices, from which is continually poured, upon the contiguous surfaces of the smooth interior of the membrane, a serous fluid, in the form of steam or vapor, which may at any time be rendered visible by opening the chest of an animal recently dead. The absorbents of this membrane are very numerous ; and, thoiigh their ex- treme exility prevents us from demonstrat- ing them in a state of health, yet may they often be seen in considerable numbers in horses that die of dropsy of the chest ; we have also abundant proofs of their exist- ence from various phenomena that occur in the diseases of the part. We know, for instance, that these vessels take up the serous fluid effused in hydrothorax, for they have been found fuU of it after death ; and it is a fact that no longer admits of doubt, that blood, extravasated into the chest, is absorbed by the mouths of these minute vessels. The nerves of the pleura are too small to be traced by dissection ; but, though it is not possessed of much sensibility in a healthy state, we know, at least we presume from analogy, that it is highly sensitive in the diseased; for few diseases are more >'^ EXPLANATION OF FIGURE XV. NO. 1.— FORE EXTREMITIES. LATERAL VIEW OF THE OFF-FOEE LIMB. h. Humero cubital. — Flexor brachii. 11. Tricep.s externus. 0. Pectoralis transversalis. P". Flexor metaearpi externus. s". Extensor metacai'pi magnus. t'. Extensor metaearpi obliquus. «'. Levator humeri. X. X. Extensor pedis. y. y. Extensor suf&'aginis. ^•. The hoof. NO. 2. (VnrW AS ABOVE.) k". Humero cubital, or flexor brachii.- m". n". Two of the triceps extensor brachii. p". Flexor metaearpi externus. *. Extensor " magnus. t. " " obliquus. M. Levator humeri. f'. w. Flexor tendons. x". Extensor pedis. y". y. Extensor sufiragims. 8. Perforatus et perforans. 4. Subcutaneous thoracic vein. NO. 3. (THE SAIIE VIEW.) r. m\ n'\ Triceps extensor brachii. p". . Flexor metaearpi externus. 2- Extensor suffi-agmis. 8. Extensor metaearpi magnus. u. Flexor tendons. x". Extensor pedis. z. Suspensory ligaments. 4. Subcutaneous thoracic vein. NO. 4. OSSEOUS STRUCTURE. 34. Os humeri. /• Os ulnaris. 35. Os Radialis. 9- Trapezium. 36. Ossa carpi. EXPLANATION OF FIGURE XV. CONTINUED. 37. Metacarpus magnum. X. " paiyvm. 38. Ossa sessamoidea. 39. Os suffi-aginis. 40. Os corona. 41. Os pedis. NO. 5. INSIDE VIEW OF THE OFF-FOKE LEG. o". Pectorales. r". Flexor metacarpi intemus. q". Flexor metacarpi medius. s". Extensor metacarpi magnus. f. Extensor metacarpi obliquus. m". v". Flexor tendons. a;.' Extensor pedis. v". Suspensory ligament. 6. Radial vein. 8. Bifiu'cation of the suspensory ligament. z. Splent bone. S,: Inferior border of the hoof. THE HORSE. 151 acutely painful in the human subject than pleurisy, and we have every reason to be- lieve that horses sutTer much from the same malady. Secretion. — It has been observed that the exhalents of the pleura secrete a serous fluid, which is emitted, in the form of an exhalation or vapor, into the cavity of the thorax ; and that it may be rendered visible at any time, if an animal, recently dead, be opened while yet warm ; or if an open- ing be made into the chest of a live animal. In either case, a whitish steam will be per- ceived to issue from the interior of the cavity. This vapor, shortly after death, be- comes condensed and converted into a liquid ; which accounts for the contiguous surfaces of the pleura being moist, and for a collection of more or less fluid, resem- bling water, existing in the most depending parts of the cavity. Li consequence of every part of the membrane being bedewed in this manner, the lung itself may be said to be in an insulated state ; for the pleura costalis does not, philosophically speaking, touch the pleura pulmonalis, nor is the lat- ter in actual contact with the mediastinum : all friction, therefore, in the motions of these parts, is by this interfluent secretion effec- tually prevented. In this, then, consists the chief use of the pleura, viz., to furnish a secretion for the purposes of lubrication and facility of motion, which it further promotes by its extreme glibness of surface. It is said also to answer the purpose of ligaments to the contained organs, thereby confining and sti-engthening them. The use of the mediastinum is to divide the chest into two compartments. The lungs (by butchers called the lights) are two spongy bodies formed for the pur- pose of respiration. Situation and Relation. — They are con- tained in the lateral regions or sides of the thoracic cavity ; separated from each other by the mediastinum and heart, which occupy the middle region. Prior to any opening being made into the thorax, the lungs con- tinue to liU up every vacuity : no sooner, however, is a perforation made into the thoracic cavity than they shrink in volume, and become in appearance too small for the spaces they occupy. This arises from their being during life — or rather during the unopened state of the thorax — in a con- stant state of inflation with atmospheric air, which preserves them expanded ; and they suffer coUapse of substance the instant air is admitted, in consequence of the pres- sure of the atmosphere upon them, from which they were protected before by the parietes of the thorax. Division. — The lungs are two in num- ber, the right and the left lung ; parti- tioned from each other by the mediasti- num. A further division of these organs has been made into lobes. That on the right side, the larger of the two, consists of three lobes ; the left, only of two. These lobes, which are nothing more than partial divisions of the lung by fissiores of variable extent through its substance, serve to adapt them more accurately to the thoracic cavi- ties, and, at the same time, render them fitter for the purposes of expansion and contraction. Volume. — The lungs of the horse, when inflated, are of great bulk ; * and the right is the larger of the two : in consequence of the heart being inclined to the left side, less space is given for the left lung. Attachment. ■ — The lungs are attached, superiorly, to the spine (which attachment is sometimes called their roots) by blood- vessels, the divisions of the trachea, and the mediastinal portions of the pleura : every- where else, in a healthy subject, they are free and unconnected. Figure. — In form, the lungs of the horse are very like those of the human subject; and the latter have been compared to the foot of an ox, to which the injected lung of the foetus bears indeed much resemblance ; for, though the two lungs are not symmetri- cal, yet, both together, they put on this shape, which is the counterpart of that of * I consider, in comparison with the body, that they exceed in magnitude those of the human subject. 152 ANATOMY AND PHYSIOLOGT OP the cavity they occupy. With regard to their general figure, however, the lungs may be said to be conical : being broad and con- cave posteriorly, where they are opposed to the convex surface of the diaphragm ; nar- row and somewhat pointed anteriorly, where they are received into the blind pouches of the pleura, in the space between the two first ribs. Color. — In color, these organs vary some- what, depending upon the age of the ani- mal, and upon the quantity and distribution of the blood they contain. In the young subject, they are of a lighter and more uni- form shade than in the adult. In perfect health they assume a pink hue ; which, as age advances, becomes mottled with purple and grayish patches. Sometimes, in the dead subject, they are found of the color of the darkest venous blood, which arises fi-om an inordinate congestion of that fluid wthin the pulmonary veins. Structure. — The lungs are composed of the branches of arteries and veins, and of the ramifications of the trachea ; all which vessels are connected together by an abun- dant intervening ceUulai substance, known by the name of parenchyma. Beneath the cm-ve made within the chest by the poste- rior aorta, the trachea divides into the two bronchial tubes, of which the right is the larger, but the shorter ; the left the longer, in consequence of having to pass under the aorta in order to reach the left lung. Having entered the substance of the lung, the right tube divides into four others ; the left only into three ; which difference arises from the right lung possessing an additional lobe. These branches may be traced for a consid- erable extent within the parenchyma, giv- ing off in their passage numerous other smaller tubes of similar structure ; but, as we prosecute our dissection of them, we shall find that, in growing smaller, they par- take less and less of the nature of cartilage, and that the extreme ramifications are not only entirely membranous in their composi- tion, but of so fine a texture as to be per- fectly transparent. It wiU be remembered here, that, in speaking of the trachea, a; membranous lining to it was described of the mucous kind, which, it was observed, thence passed into the bronchial vessels : now, it is of the continuation of this mem- brane in an attenuated state that the minute air-tubes appear entirely to consist ; at the extremity of every one of which the mem- brane is prolonged into a kind of blind bag, or cul-de-sac, to which the name of air-cell has been given. From the arborescent ramification and pecuhar mode of termination of the bron- chical tubes, some anatomists have com- pared them, and the cells at their extremities, to a bunch of grapes — supposing the stalks to represent the ramifications of the former, and the grapes connected with them the air-cells; others have described them as having resemblance to a honeycomb : and so far as the knife, with the aid of glasses, can develope their intimate structure, the first is an apt comparison, insomuch as it relates to the disposition of their cells ; the last, insomuch as it conveys an idea of their ready inter-communication. For, though they do not communicate but tlurough the ramifications of the' bron- chial tubes, this is a medium of inter- course at once so general and free, that numbers of them are inflated at the same time by impelling air into any one of the larger branches. With the parenchymatous substance, however, they have no commu- nication whatever.* The blood-vessels that enter into the com- position of the lungs are denominated the pulmonary. The pulmonary artery, having taken its origin from the right ventricle of the heart, winds upward to the root of the left lung, and there divides into the right and left pulmonary arteries, which divisions * If the substance of tlie lungs be lacerated or rent asunder, the sui'face will be found to present a tabulated aspect. Introduce a blow-pipe into one of these lobuli, and all the other lobules — the entire lung — may be in- flated from this one ; showing the free communication ex- isting between them. The same may be effected by in- jecting quicksilver. You may do the same with the inter- stitial substance ; but in this case you do not fill the lobules. In fine, the lungs with tlTeir cells resemble a sponge ; only that the connecting tissue has no communication with the sponge. THE HORSE. 153 enter their correspondent lungs. The rami- fications of these vessels (which differ from other arteries in having no anastomotic communications one with another) accom- pany those of the bronchial tubes, and, like them, divide and subdivide, grow smaller and augment in number, as they approach the air-cells ; upon the internal * surfaces of which they become capillary, and assume a texture of correspondent thinness and pellucidity with the cells them- selves. Through these minute vessels every particle of blood is impelled every time it is circulated over the system, as was stated when on the blood : a remarkable change of color is thereby effected in it, and we have now an opportunity of seeing in what manner this fluid is exposed to the influence of atmospheric air for the purpose. It is evident that no immediate contact can hap- pen between the air and the blood, for the thin, transparent side of the vessel, if not that of the air-cell likewise, must ever be interposed ; so that, whatever this influence be, it must take effect through one or other or both of these membranes. We might conceive, indeed, that such minute vessels could not transmit through them such a body of fluid as the blood ; but, when we look at the volume of the lungs, and con- sider the incalculable number of air-ceUs they must contain, the globular surface of every one of which is furnished with an ex- pansion of pulmonary vessels, we shall feel more surprise and admiration at the extreme division and diffusion of this fluid in order to receive the necessary change, than that such a prodigious number of capillaries should be equal, in their united caliber, to the- pulmonary artery itself. From the extremities of the arteries, upon the surface of the air-cells, arise the pul- monary veins. These, by repeated union with one another, form themselves, first, into visible branches, which subsequently become branches of larger size, until at length they end in eight pulmonary venous trunks, which proceed to, and by four openings ter- * Some say, "upon the external surfaces." minate in, the left auricle of the heart. The ramifications of these veins, unliive the generality of others, are not more numerous than those of their coiTespondent arteries : and the reason for this is obvious ; for, here, one set of vessels are not more subject to compression than the other, nor does the heart (which is so proximate to them) re- quire any such aid as an additional number of veins affords to carry on the circulation. The pulmonary veins have only to convey the blood back to the heart, after it has received its due change within the capil- laries upon the air-ceUs. Organization. — Besides the pulmonary blood-vessels, there are two others, named bronchial arteries. They come off, by one trunk, from the posterior aorta, and each of them enters a division of the lungs, in the substance of which it branches forth, and takes the course of the bronchial tubes. These tubes they supply, as weU as the coats of the pulmonary vessels, and the paren- chyma of the lungs, with blood: in fact, they may be regarded as the nuti-ient ves- sels of these organs. It has been, ho^vever, and stiU remains, a subject of dispute, whether these vessels do exclusively nourish the substance of the lungs or not ; some say that they do ; whUe others assert that they are assisted in this function by the ptd- monary artery, with some of the branches of which they anastomose. The latter opinion certainly does not appear to be sup- ported by facts of much weight; on the contrary, the blood which the pulmonary arteries contain is dark-colored, and unfit for the nutriment of any organ ; and as for anastomosis, we have no demonstrative proof of its existence. The bronchial veins end in one trunk, which returns the blood into the vena azygos. The nerves of the lungs are derived prin- cipaUy from a large plexus within the chest, constituted of the par vagum and sympa- thetic. They enter the pulmonary structure in company with the bronchial tubes and blood-vessels, and continue their course with them, to be dispersed upon the bron- chial membrane and parietes of the air-cells. 154 ANATOMY AND PHYSIOLOGY OF THE HORSE. The absorbents of the lungs are large and numerous, particularly the deep-seated : and of the superficial, we may often succeed in injecting considerable numbers, by intro- ducing a quicksilver-pipe under the pleura pulmonalis. They aU pass through the absorbent glands situated around the roots of the broncliial tubes. Parenchyma. — The connecting medium of the various constituent parts of these organs, or, as it is termed, their parenchyma, appears to consist of little else than cellular tissue, without any intermLxture of adipose matter : it admits of the free diffusion of any fluid that may be extravasated into it — of air that may have escaped from the air-cells, or of serous fluid poured out when the lungs become anasarcous ; but, as was observed before, there is no intercommuni- cation between it and the cells or vessels, as long as the organs preserve their integrity of structure. Specific Gravity. — The lungs, when healthy, are exceeding light in comparison to their volume ; so that, if they be immersed in water, unlike most other parts, they will float upon the surface, — a fact familiar to every one who has seen the liver and lights of an animal thrown into a pail of water to be washed : indeed, the name of lights itself seem to have been given to them from this very property. If the festal lungs, how- ever, be so treated, they will instantly sink to the bottom of the vessel : and this ex- jjrrimental result at once shows why those ol an animal that has once breathed should swim ; for, in the one instance they contain air, in the other they are wholly free from it. They are not to be regarded as respi- ratory organs in the foetus. It is evident, therefore, that the lungs owe their property of lightness to the air they contain ; and, as a further proof of it, if that air be by any means absorbed or pressed from them, and their bulk diminished by collapse of the air-cells, like other viscera, they will prove heavier than an equal volume of water: hence it is that the lungs of a horse that has died of hydrothorax, even though they be sound, are of a greater specific gravity than those of one in health. It occasionally happens, however, that these viscera evince, in this particular, the properties of airless lung, while their natural volume and general appearance remain the same : there must be present interstitial deposition. BRONCHIAL GLANDS. Small, oval-shaped, glandular-looldng bodies, situated about the roots of the lungs, adhering more particularly to the bottom of the trachea and the bronchial tubes. They exhibit a dii'ty French gray hue, inter- spersed with dark blueish spots, and are about the volume (though this varies much) of a tick-bean. For a long time the nature of these bodies remained obscure : of late, sldlful injections have clearly shown them to be absorbent glands. They possess their capsules, and, when cut open, exhibit a cel- lular structure. They contain a dark fluid, which will soil anything it touches ; whose principal ingredient chemists have found to be carbon. CIRCULATORY SYSTEM. PRELIMINARY REMARKS (oN*" THE BLOOD, ETC.). The appearance of blood is familiar to most persons. It contains the elements for building up and nourishing the whole ani- mal structure. On examining blood with a microscope, it is found full of little red glo- bules, which vary in their size and shape in different animals, and are more numerous in warm than in cold-blooded animals; probably this arises from the fact that the latter absorb less oxygen. If the blood of one animal be transfused into another, it will frequently cause death. When blood stands for a time after being di'awn, it separates into two parts. One is called serum, and resembles the white of an egg ; the other is the clot or crassamentum, and forms the red coagulum, or jelly-like substance : this is accompanied by whitish, tough threads, called fibrine. When blood has been drawn from a horse, and it as- sumes a cupped or hollow form, if serum, or buffy coat, remain on its surface, it de- notes an impoverished state ; but if the whole, when coagulated, be of one uniform mass, it indicates a healthy state of this fluid. The blood of a young horse gen- erally coagulates into a firm mass, while that of an old or debilitated one is gen- erally less dense, and more easily divided or broken down. The power that propels the blood into the different ramifications of the animal, is a mechanico-vital power, and is accomplished through the medium of the heart and lungs ; the latter is a powerful muscular organ contained in the chest. From certain parts of it arteries arise ; in others the veins terminate ; and it is princi- pally by its alternate contractions and ex- pansions, aided as already stated, that the circulation of the blood is carried on. The heart is invested with a membranous sac, called pericardium, which adheres to the tendinous centre of the diaphragm, and to the great vessels at the base of the heart. The heart is lubricated by a serous fluid within the pericardium, which guards against friction. In dropsical affections, the quantity of this fluid is considerably in- creased, and constitutes a disease called hy- drothorax. The heart is divided into four cavities, viz., t\vo auricles, named from their resemblance to an ear, and two ventri- cles, forming the body. The left ventricle is smaller than the right ; but its sides are much thicker and sti-onger : it is from this part that the grand trunk of the arteries proceeds, called the great aorta. The right cavity, or ventricle, is the receptacle for the blood that is brought back by the veins after going the rounds of the cu-culation ; which, like an inverted tree, become larger and less numerous as they approach the heart, where they terminate in the right auricle. The auricle on the left side of the heart receives the blood that has been disti-ibuted through the lungs for purifica- cation. Where the veins terminate in auri- cles, there are valves placed. The coronary vein, which enters the right auricle, has its mouth protected by a valve called semi- lunar, or half-moon shape, which opens only toward the heart, and prevents the blood taking a reti-ograde course. The different tubes coming from and entering into the heart are also provided with valves to pre- vent the blood from returning. For exam- ple, the blood proceeds out of the heart, along the aorta ; the valve opens forward or upward, the blood also moves lapward, and pushes the valve asunder, and passes through ; the pressure from above effec- tually closes the passage. The valves of (155) 156 ANATOMY AND PHYSIOLOGY OP the heart are composed of elastic cartilage, which enables them to work with ease. In some diseases, however, they become ossi- fied. This, of course, is fatal. The heart and its appendages are also subject to other diseases, called dilatation, softening, hard- ening, etc. Now, the blood, having been brought from all parts of the system by the veins, enters into the vena cava ascending and descending portion, which empty them- selves into the right auricle ; and this, when distended with blood, contracts, and forces its contents into the right ventricle, wliich, contracting in its turn, propels the blood into the pulmonary arteries, whose numerous ramifications bring it in con- tact with the air-ceUs of the lungs. It then assumes a crimson color, and is then adapted to build up and supply the waste. Having passed through the vessels of the lungs, it continues on, and passes into the left auricle: tliis also contracts, and forces the blood through a valve into the left ventricle. This ventricle then con- tracts in its turn, and the blood passes through another valve into the great aorta, from which it is distributed into the whole arterial structure : after going the rounds of the circulation, it is again returned to the heart by the veins. EXAMINATIOXS OX THE NATUEE AND PROPERTIES OF BLOOD. Q. What are the properties of blood? — A. In health, it is a smooth homogenous fluid, of unctuous adhesive consistence, of a slightly saline taste, and of a specific gravity somewhat exceeding that of water. It exhales a vapor which has a peculiar odor ; this, how- ever, differs in various animals. Q, Does the blood always preserve the same den- sity P * — A. No. Its density is hable to great variations, under the states of rest, labor, disease, and health. Q. What do you understand by the " crassamentum " of the blood? — A. It is supposed to consist chiefly of fibrin. Q. How is it colored ? — A. It owes its peculiar color to what is termed the red globules, which are entangled in it during its coagulation. Q. How can this be demonstrated ? — A. By long continued ablution in water, the red particles are hber- ated ; and we have remaining a white, soKd, and elas- tic substance, which has all the properties of fibrine, and is almost exactly similar to the basis of muscle. Q. By what name was fibrine formerly known ? — A. Coagulable Ij-mph. Q. What is the form of the red globules of the blood? — A. The Abbe de la Torre, who examined them under microscopes of considerable j)Ower, states that they obtained the appearance of flattened annular bodies, with a depression, sometimes perforation, in the *Dr. B. Babington is of opinion that the blood, whilst circulating in the vessels, consists of two parts only — a fluid which he calls liquor sang^Hi'nys, and red globules; and he is induced to believe, from his experiments, that fibrin and serum do not exist as such in the circu- lating fluid, but that the hquor sanguinis, when removed from the vessels, and no longer subjected to the laws of life, has then, and not before, the property of separating into fibrin and serum, hhd. Chi- rvrg. Transact, vol. xvi. pt. 2. Lond. 1S31, and art. DIood (morbid conditions of the), in Cyclop, of Anat and Physiol. Lond. 1836. centre, but they differ in size and shape in various animals. Q. By what means is the blood colored? — A. By means of n-on and oxygen. Q. Describe the properties of the serum ? — A. It is the yeUow fluid pait that is left after the separation of the crassamentum ; it is of a saline taste, and homoge- nous, adhesive consistence. Q. What effect has a temperature of 160° on it? — A. The whole is converted into a firm white mass, per- fectly analagous to the white of an egg wliich has been hardened by boihng. Q. Can any liquor be extracted fi'om the serum afler ha'\ing been coagulated by heat? — A. Yes. If the coagulum be cut into slices, and subjected to gentle pressure, an opaque liquor drains from it, which is called the serosity. PEEICAEDIUSI. Q. By what is the heart surroimded? — A. The pericardium. Q. What is the structure of this? — A. It is a fibro- serous membranous bag, composed of two coats ; one fibrous, the other serous ; these are united by cellular tissue. Q. What are its connections? — A. It is attached to the sternum, pleura, diaphragm, and to the roots of the large blood-vessels at the base of the heart. Q. AVhat is the function of the serous sm-face of the pericardium? — A. To secrete the liquor pericai-dii. Q. What is the use of this liquor ? — A. It serves to protect its own sm-face, and that of the heart, from friction. Q. What office does the pericai-dium perform? — A. It sustains the heart in its proper situation. THE HOESE. 157 Q. What is the form of the heart? — A. Its form is conoid, yet somewhat flattened on the anterior surface and rounded on the other. Q. Where is the heart situated? — A. Within the thorax, in the region of the fourth, fifth, and sLxth dorsal vertebra;; bounded on tlie sides by the lungs and walls of the thorax ; posteriorly, by the dia- phragm; Inferiorly and anteriorly, by the sternum. Q. How is the body of heart di\ided ? — A. Into a base and apes. Q. Wliat are the divisions internally? — A. It is di\ided into foiu' cavities, \t2 : two auricles, or anterior cavities ; two ventricles, or posterior eanties. Q. Wliat commimications exist between the cavities of the heart ? — A. Between the two auricles there is no commxuiication, nor between the two ventricles ; but the right aiu-icle opens into the right ventricle, and a similar opening exists between the left auricle and ventricle. Q. How do veterinarians describe the relative situa- tion of the cavities of the heart ? — A, The amides are described as anterior and posterior, because the right auricle forms the upper and fore part, and the left is in a posterior dii-ection ; the ventricles being located under then- respective auricles ; thus we have the ante- rior and posterior ventricles. Q. How is the exterior sm'face of the heart pro- tected ? — By a dupHcature of the pericardium. Q. What is the function of the am-icles? — A. To receive the blood from the various vessels and transmit it to the ventricles. Q. What is the function of the ventricles? — A. One propels the blood to the lungs, for pm-ification ; the other distributes it through the arterial ramifi- cations. Q. Name the venous vessels which terminate in the right auricle. — A. Three venous vessels terminate in it, riz : the vena cava, anterior and posterior, and the coronary vein; the vena azygos forms a junction with the anterior cava, just as the latter pierces the walls of the auricles. Q. How are the auricles dirided ? — A. By the sep- tum am-icularum. Q. Describe the internal mechanism of the right ventricle? — A. It has within it numerous fleshy pil- lars, longitudinally distributed; also, three fleshy prominences, termed carnea columnse, from which sev- eral tendinous cords proceed to the edges of those membranous and fibrous productions ; these close the auriculo-ventricular opening; the apparatus alto- gether forms vahida tricupsis. Other cords, similar to the cordis tending, pass between the outer wall and the septum. Q. Where is the origin of the right pulmonary ar- tery ? — A. It emerges from the upper and back part of the ventricle. Q. How is the mouth of this artery protected ? — A. By three semilimar valves, wliich present Uttle pouches within its cavity ; these valves consist of doublings of the Hning membrane of the -parts. Q. Describe the left ventricle? — A. Its cavity is smaller than that of the right, and its wall is tliicker. Its muscuU pectinati appear mostly upon the septum, within the apex and mider the valves ; it has two, in- stead of thi-ee, carna; columnse ; they are more bulky, and project more into the carity than those of the right. Q. From whence does the aorta arise ? — A. From the upper and fore part of the left ventricle. Q. What is remarkable about the mouth of the aorta ? — A. It has three semilunar valves, similar to those at the origm of the pulmonary artery. Q. By what are the ventricles cUrided ? — A. They are divided by a fleshy partition called septum ventric- uloi-um. Q. How is the circidation of the blood cfiected ? — A. By the alternate contraction of the auricles and ventricles, called the dyastole and systole of the heart. Q. By what vessels is the heart itself supplied with blood ? — A. By the coronary arteries. ARTERIAL SYSTEM. DISTRIBUTION OF ARTERIES. The blood is propelled by the heart through the great aorta, which rises out of the base of the left ventricle, in the space between the left auricle and the pulmonary artery. The branches furnished by the main trunk are the coronary arteries. The right coronary artery emerges from between the pulmonary and right auricle, winds round the fissure separating that cavity from the right ventricle, and turns down under the termination of the vena cava; and distri- butes ramifications in its course, which penetrate the substance of the parietes, and end in spiral branches. The left coro- nary artery, in passing out between the pulmonary artery and left auricle, sends off a large branch, which encircles the other auricle ; it then takes its course down- ward, and ends in spiral ramifications. ANTERIOR AORTA. This is a shorter division of the main trunk. The course of this vessel is under the windpipe ; it gives origin to those large arteries which are distributed over the breast, head, neck, brain, and anterior ex- tremities. It divides, at a short distance from the heart, into the right and left ar- teria innominata ; the right is considerably longer than the left, and measures nearly as much again in circumference ; it forms the ti-unk from which the two carotid arteries spring; the left terminates in the following vessels : * 1. The dorsal artery. 2. Posterior cervi- cal. 3. Vertebral. 4. Internal pectoral. 5. External pectoral. 6. Inferior cervical. * The vertebral artery, forming the basilar, gives off tlie posterior cerebellal, anterior cerebellal, posterior cere- bral, and the circular arteriosus. 7. Axillary. Each of these arteries ramify and anastomose with others, and are dis- tributed to muscular and adipose substance. From the axillary artery spring all the ar- teries of the fore extremity. This vessel can only be seen by detaching the shoulder from the body. It arises within the chest, from the arteria innominata ; gains exit by making a sudden turn around the first rib, rather below its middle, crossing the lower border of the scalenus in the turn; it is first directed outward in this flexure, and then backward, and at length reaches the inner part of the head of the humerus, where it makes another turn backward, and afterwards takes the name of the brachial artery. Its branches are — 1. The external thoracic. 2. The internal thoracic, which runs to the point of the shoulder, and gives its branches to the levator humeri and shoulder joint. 3. The dorsalis scapulae ascends, in a flexuous manner, to the shoulder joint, crossing the insertion of the subscapularis. It runs for a short distance along the ribs. 4. The subscapularis, a large artery, which also arises from the upper part of the trunk, but near to its ter- mination. It passes along the ribs, screened from view by the edges of the subscapu- laris and teres major, to both of which muscles it detaches several small branches, and ends near the lower angle of the bone ; it also gives off several branches to the triceps and panniculus. 5. The humeral. The humeral artery descends from the inner and back part of the head of the os humeri, in an obUque direction on the body of the bone, where it divides into the ulnar, spiral, and radial arteries. On its inner side, it has the spiral and ulnar ner^'es ; in front, the radial nerve; and behind, the (158) ANATOMY AND PHYSIOL YGY OF THE HORSE, 159 humeral veins; and it is covered inter- nally by the large pectoral muscle, to which it sends some small branches. But its prin- cipal branches are — 1. One near its origin, which crosses the bone to get to the flexor brachii, and sends twigs to the shoulder- joint. 2. A posterior branch, arising a lit- tle lower down, which enters the muscle called triceps. 3. Near its termination, another branch to the flexor brachii. Where the artery divides, it is covered by the hu- meral plexus of veins, and by the absorbent glands of the arm. The ulnar artery consists of a common root, from which spring three or four ves- sels of considerable size, running in waving lines upon the inner side of the lower end of the humerus. The upper one is directed to the ulnar, splitting before it reaches the bone, and sending one branch upward upon the elbow, and another downward to the heads of the flexors ; to which muscles the other branches of tliis vessel are dis- tributed. The spiral artery, the outermost division, turns round the os humeri, passing under the flexor brachii, and sending a recurrent branch to it, to arrive at the front of the radius, where it splits into several branches, of which — 1. Some run into the elbow joint. 2. Others, larger and more numer- ous, penetrate the heads of the extensors. 3. Two long, slender ones descend upon the radius, and give branches, in their course, to the extensor muscles as low as the knee, and there end in ramifications about and into the joint, joining with others coming from the radial. The radial artery, the principal division humeral, continues its descent along the radius, about the middle of the arm ; the nerve accompanies it first on its outer side, and subsequently behind it. A short way above the knee, it splits into the metacarpal arteries. The small metacarpal artery descends, within a cellular sheath, along the inner and back part of the knee. It continues its descent along the metacarpal vein (which runs to its inner side), tUl it gets below the knee, and then transmits its divisions down the front of the suspensary ligament ; be- tween it and the canon bone, it sends off" branches over the front of the knee, the canon, and suspensaiy ligament. The large metacarpal artery, a continua- tion of the radial trunk, continues its course down the leg, by the side of the tendo per- foratus, passing under the posterior annular ligament, approaches the fetlock just above the joint, and then splits into three vessels ; from the middle division three recuiTcnt ar- teries are given out ; the side divisions be- come the plantar arteries. From the arch below come off" tw^o other branches, which descend into the joint. The plantar arte- ries, external and internal, in the fore ex- tremity, result from the fork of the meta-. carpal ; in the hind, from that of the metatarsal. (Their general distribution is the same, both in the hind and fore feet.) They descend the fetlock upon the sides of the sessamoids, in company with the veins which run in front of them, and with the plantar nerves which proceed behind them ; the artery then passes down to, and into, the substance of what is caUed the " fatty frog ; " it next passes the inner and upper extremity of the coffin bone, and afterwards to the foramen of the posterior concavity of the bone. The branches of the plantar artery are many and important. After detaching some small ramifications inwardly to the fetlock, posteriorly to the ffexor tendons, and anteriorly to the ex- tensor tendon, it then sends off" — 1. The perpendicular artery. 2. The transverse artery. 3. The artery of the firog. 4. The lateral laminal artery. 5. The circulus ar- teriosus. From the latter arise two prin- cipal sets of vessels — 1. The anterior laminated arteries. 2. The inferior com- municating arteries, "thirteen, and some- times fourteen, in number." 3. The circum- flex artery. Then, again, from this vessel spring the solar arteries, which may be so named from their radiated arrangement. These latter are destined for the supply of the sole, upon which they run in radii at equal distances, whose common centre is the 160 ANATOMY AND PHYSIOLOGY OF toe of the frog, where they end in commu- nications with the arteries of that body. THE CAROTID ARTERY. The right arteria innominata, having de- tached seven important branches, which vary but little in their mode of origin, general course, and distribution, from the several arteries into which the left division resolves itself, become the common carotid — a large vessel emerging through the up- per opening of the chest ; it divides, as it quits the chest, into two branches, called the right and left carotids. These arteries ascend, and having reached the top of the larynx, the carotid of either side branches into three divisions — the external and inter- nal carotids, and the ramus anastomoticus -. here, though the trunk itself becomes deeply lodged in soft parts, its situation is well in- dicated by the larynx, with which it is in contact. This vessel detaches — 1. Several unimportant muscular branches in its pro- gress up the neck. 2. The thyi-oideal artery, which furnishes the laryngeal, a small artery that perforates the ligament uniting the cartilages of the throat. Tlie external carotid artery is the large division, which may be regarded as the con- tinuation of the carotid itself. This artery is imbedded in glandular substance, sur- rounded by venous and nervous trunks, and protected by bony prominences and muscles. The first branch of the external carotid is the submaxillary artery ; it comes off behind the horn of the ox hyoides, just as the carotid makes its second curve, and ranks next in size to the trunk itself. After reach- ing the lower jaw (about one-thu-d of its length downwards), it arrives upon the face; here it becomes subcutaneous, ending in an equal division, called the facial and inferior labial arteries. Its branches are, the ascend- ing laryngeal, pharyngeal : smaller branches go to parotid gland, and a large branch, called the lingiial. The latter detaches a few twigs into the submaxillary space ; it then branches into two arteries, the ranine and the sublingual. The ranine, apparently a continuation of the lingual, passes along the under part of the tongue, and transmits branches to the interior, and continues of large size even to the tip of the organ, wherever its extreme ramifications are ex- pended. The sublingual artery winds along the under and outer border of the tongue, preserving a more superficial course than the former. It supplies the sublingual gland, and distributes branches over the membrane of the tongue. The submental artery leaves the submaxillary, follows the course of the branch of the jaw, and de- taches twigs to muscles ; it then transmits its ramifications into the gums internally. The anterior masseter branches pass on the external side of the jaw. The inferior labial artery courses the side of the jaw, invested in the cellular and fleshy substance belonging to the buccinator. It gives off slender ramifications to the in- vesting cellular substance, also the buc- cinator arteries ; the buccal twdgs bifurcate, sending their divisions respectively to the upper and under Ups ; these form the supe- rior and inferior coronary arteries of the lips. The facial artery ascends upon the side of the face, crosses the buccinator, then, having run as high as the bony ridge from whence the masseter arises, it detaches a large branch, and then expands upon the upper and fore part of the face; its termi- nating ramifications are in the cellular sub- stance and skin covering the fore part of the face. • The posterior auricular gives branches to the parotid gland, and to the different mus- cles of the ears. The teviporal artery, i\ie anterior auricular, and the internal maxillary, may be con- sidered as the terminating branches of the external carotid. The internal maxillary gives off deep temporal branches, long slender twigs, to the soft palate, to the ear, and to the articulation of the jaw; the facial artery also gives off the inferior maxillary, the supra-orbitar, the ocular, the infra- orbitar, and the palate maxiUary. The second and smallest division of the carotid is the EXPLANATION OF FIGURE XYI. XO. 1. FROXT AND SIDE MEW OF THE NEAR-HIND LEG. D LEG. K. Abductors. J". Triceps. m\ Tensor vagina. m'. Rectus. o'. Vastus externus. )•'. Gastrocnemius externus. V. s'. Peroneus. u\ Gastrocnemius externus. «'. u'. Flexor pedis. X. Extensor pedis. y. y. Extensors. «. V. Flexor tendons. §-. The hoof. 8. Bifurcation of suspensory ligament. No. 4 is nearly the same as No. 3, and therefore ueeds no further description. z. Suspensory ligament. EXPLANATION OP FIGUKE XVI. CONTINUED. NO. 5. f. Abductor tibialis. q. Glans penis. r. Gastrocnemius extemus. t'. Flexor pedis accessorius. «'. Lisertion of the gastrocnemius. x'. X. q. Extensor pedis. 8. Bifurcation of the suspensory ligament ^-. The hoof. 9. Flexor metatarsi. 5. The saphena vein. z. Suspensory ligament. THE HOKSE. 161 RAMUS ANASTOMOTICUS. It leaves " the trunk of the carotid, joins the vertebral, and from it arises the occipital artery, which gives off twigs to be dispersed upon the dm-a mater, temporal muscle, and muscles of the occiput. INTERNAL CAROTID. This vessel, whose calibre is not more than half that of the external carotid, ascends to the base of the skuU : at its en- trance into the skull, a vessel comes off named the arteria communicans : after having given off tliis vessel, the internal carotid pierces the dura mater, takes its course up near the optic nerve, and branches into four divisions, wliich supply the cere- brum with blood. The remaining vessels of the brain are derived from the vertebral artery, which gives off posterior arteries to the dura mater, and ramifications to the medulla oblongata. The basilar artery sends off branches to the cerebellum. THE POSTERIOR AORTA. Considerably longer and larger than the anterior is the main trunk, from which are derived the artery of the abdomen, pelvis, and posterior extremities, in addition to the posterior intercostals, and some few of the thoracic arteries. It commences opposite the fourth dorsal vertebrae : from its origin it courses first upward, and then backward, having the pulmonary artery on its left, the termination of the windpipe on its right, then takes a course along the spine, inclin- ing to the left side. From the inferior part of the curvatme of the aorta arise the right and left bronchial arteries: these vessels penetrate the lungs in company with the bronchia, to the branches of which they cKng in the course of their ramifications within the substance of the lungs. The esophageal also spring from the con- cavity of the arch near to the former, and proceed backward to the esophagus, where it divides into an inferior and superior artery. The intercostal, the remainuig branches, come off in pairs from the sides of the ves- sel, to supply aU those intercostal spaces posteriorly to the last. These arteries ma along the lower borders of the ribs, and end about the inferior parts of the chest and abdomen. They furnish, near their origin, small branches, which enter the vertebral canal. Having detached these small ves- sels, the posterior aorta continues its pas- sage into the abdomen. In maldng its exit from the chest, it gives off the phrenic or diaplnragmatic arteries. Witliin the abdomen, the aorta continues to be firmly fixed to the spine, by its several cellular attachments, as far as the lumbar vertebra, under the body of which it branches into four large arterial trunks. Prior to this division, the abdominal aorta gives off the cceliac artery, which is nothing more than the common root of the splenic, gastric, and hepatic — arteries that in some instances have separate origins. The splenic artery, after passing between the stomach and spleen, ends in the left gastric artery. In its course it gives off several branches to the pancreas, caUed pan- creatic arteries. The gastric artery, the smallest of the cceliac divisions, ruiis forward to the small curvature of the stomach, between the layers of the omentum, branching, before it reaches this organ, into two vessels, called inferior and superior gastric, which finally ramify upon the upper and under surface of the stomach. The hepatic artery, the largest of the cceliac division, proceeds before the pancreas to the right side of the cavity, and passes over the pyloric end of the stomach, and gives off smaU -branches to the pancreas. Near the pylorus, it sends a branch to the duodenum, which, as soon as it reaches the intestine, divides : one division — ■ the duo- denal — retrogrades along the gut, and ends in anastomosis, with branches coming from the anterior mesenteric ; the other — the right gastric — crosses the gut, proceeds to the great curvature of the stomach, where it inosculates with the left gastric. The hepatic artery itself is continued forward to the porta of the fiver, where it divides 162 ANATOMY AND PHYSIOLOGY OF into the right and left hepatic ; the right, — the larger and shorter one, — after giving off a considerable branch to the portio media, turns back to reach the right lobe ; the left, after giving off a branch or two to the mid- dle portion, penetrates the left lobe. The anterior, or great mesenteric, is the next vessel to the cceliac, and arises from the under part of the posterior aorta. From its origin, it passes downward within the layers of the mesentery, detaching some small twigs to the pancreas ; it then sepa- rates into larger vessels (commonly from eight to twelve in number), from which are derived a branch that runs to the duode- num ; several other branches encircle and ramify on and around the intestines. The renal or emulgent arteries leave the aorta at right angles just below the preced- ing vessel ; they each pass into the respect- ive kidneys, and therein divide into branches that penetrate the glandular substance. The spermatic arteries, right and left, originate from the under part of the aorta ; they pass out of the abdomen, at the ab- dominal ring, to the testicles. In the female, they pass to the ovaries, fallopian tubes, and horns of the uterus. The posterior aorta also gives off the small mesenteric, and five or six pairs of lumbar arteries. Under the last lumbar vertebrsE, the aorta gives off two pairs of arterial trunks, called the external and inter- nal iliacs. The internal iliacs give off a branch called the artery of the bulb, and afterwards branches into three divisions — the obtura- tor, gluteal, and lateral sacral arteries. The artery of the bulb passes to the bulb of the penis, where it terminates. In the female, this artery sends its terminating branches to the vagina. It gives off the fcetal umbilical artery. In leaving the pel- vis, the prostatic artery, which detaches twigs to the vesiculEB seminales, also distri- butes its ultimate ramifications to the pros- tate gland. It also gives off divers branches, anal and perineal, to the posterior portion of the rectum, anus, and parts comprising the perineum. The obturator artery is the lowest of the divisions of the internal iUac. Its branches are the arteria innominati, and ramifications to the obturator muscles and ligaments. Its divisions are the ischiatic, which distributes its branches to the triceps ; next, the pubic : the internal pubic artery gives tw^o sets of branches, which pass to the penis. The gluteal artery is destined principally to supply the gluteal muscles. Tlie lateral sacral artery, having reached the coccyx, divides into two branches. It furnishes the sacro-spinal branches, five or six in number, and the perineal artery. It soon divides into several ramifications, of which many run into the gluteal mus- cles ; others descend on the back of the thigh, and others are distributed to the anal muscles, and to the skin and ceUular sub- stance of the perineum. The lateral sacral also furnishes the lateral coccygeal, and the inferior coccygeal. TJie external iliac artery, right and left, results from a branch of the posterior aorta, which takes place under the body of the last of the lumbar vertebrae, and passes into muscles, forming the inside of the thighs. The vessel gives off the circumflex artery of the ileum, the artery of the cord, and the arteria jDrofunda : the latter, having reached the posterior quarters, it sends its ramifications into the biceps. Before this vessel dips into the substance of the thigh, it gives rise to a large branch called the epigastic artery. The epigastric artery, in passing the mar- gin of the internal ring, forms a branch which divides into several small arteries ; of these a twig runs to the groin, and ramifies among the adipose membrane and absorbent glands ; then, next, a slender branch to the cremaster, and subcutaneous twig to the thigh, and, lastly, the external pudic artery. The fenwral artery. — Regarding the pro- funda femoris as a limb of the external iliac, we descend to the femoral artery, the subsequent continuation of the same trunk. This artery proceeds in an oblique direction down the haunch, preserving nearly the line of its middle ; opposite to the head of the THE HORSE. 163 tibirf, it branches into the anterior and pos-' terior tibial arteries ; the anterior tibial gives ofl' the inguinal artery, also tlnree or four branches to thft sartorius, and one to the side and front of the stifle. Its posterior branches are a large artery to the gracUis (wliich detaches twigs to the long and short heads of the triceps), also one to the biceps. At the back of the stifle come off the pop- liteal branches, four or five in number, taking opposite directions, which are des- tined for the supply of the joint ; one runs down upon the posterior tibial muscles; another — the recurrent branches — climbs the back of the os femoris, and anastomoses with the descending ramifications of the profunda femoris. The tibial arteries are a continuation of the femoral trunk, which branch off into tibial arteries at the head of tibia. The posterior tibial artery, the smaller of the two, passes along the posterior deep region of the thigh, to the hock, where it ends in bifurcation. Its branches are, one that runs into the flexor pedis ; another to the upper and back part of the tibia ; and small twigs to both the flexgrs. There are several terminating branches, some ramifying sub- cutaneously, others continuing down the leg internally over the tendon of the flexor pedis, and ending at the lower part of the canon in divers small ramifications. The anterior tibial artery, after leaving the trunk, passes down the fore part of the thigh to the hock and metatarsal bone, where it becomes the me+atarsal artery. The metatarsal artery pursues its course downwards to about two-tliirds the length of the leg ; it then gains the posterior part of the latter ; a Httle above the fetlock, it divides into three vessels : one forms an arc, (as in the fore extremity), from which come off" the recurrents, and they anastomose with the posterior tibial artery ; the lateral divisions become the plantar arteries. 164 ANATOMY AND PHYSIOLOGY OF REMARKS ON THE DISTRIBUTION OF ARTERIES. Aorta Anterior Aorta The preceding is a brief sketch of the arterial structure, and the professional man will perceive that we have not named the whole of the arteries ; therejpre, in order to supply this deficiency', the author here introduces a table of the arteries, constructed by Mr.*Percivall. TABLE OF THE ARTERIES. Anterior Aorta. Posterior Aorta. Right Arteria Lmominata. Left Arteria Innominata. ''Dorsal. Posterior Cervical. Vertebral, forming the Basilar. Internal Pectoral. External Pectoral. Inferior Cervical. _ Axillary. ('External Thoracic. Internal Thoracic. Axillary < Dorsalis Scapulse. Subscapular. 1^ Humeral, f Ulnar. Spiral Radial. . . i Small Metacarpal. Larare ditto. Left Arteria Innominata ('Posterior Cerebellal. J Anterior Cerebellal. ] Posterior Cerebral, l^^ Circular Arteriosvis. Humeral Large Metacarpal | External Plantar. ) Internal ditto. f Perpendicular. Transverse. Plantar < Artery of the Frog. Lateral Laminal. (^Circular Arteriosus. Ant. Laminal. Inf. Communicatuig. Circumflex. . . • ^ Solar. The Right Arteria Innominata sends off' branches correspondent to those on the left side ; and, in addition, the Right Carotid J External Carotid. Left ditto. > Ramus Anastomoticus. ) Internal Carotid. ' Ascending Pharyngeal. Common Carotid External Carotid < Submaxillary. Parotideal. Int. Pterygoid. Post. Masseter. Post. Auricular. Temporal. Ant. Auricular. Int. Maxillary. Pterygoid. Lingual. . . . Inferior Labial. Facial. \ Ranine. I Sublingual. ( Buccinator. I Angular Oral. ^ Masseter. J Buccal. I Sup. Labial. I False Nasal. 'Deep Temporal. Palatine. Inf. Maxillary. Supra-Orbitar. Ocular. Infra- Orbitar. ^Palato Maxillary. THE HOESE. 165 Dm-a Matral. Ramus Anastomoticus [• Occipital. ^ Temporal. Nuchal. {Ai-teria Communicans. Anterior Dura Matral. Anterior Cerebral. . . <( Ophthalmic IVIiddle Cerebral. Lateral Cerebral. PosTERiou Aorta Thoracic Division. Abdominal ditto. Bronchial Thoracic Division < Right Broncliial. Left ditto. Superior Esophageal. Inferior ditto. Abdominal Division Esophageal. Intercostals. Phrenic. S Pancreatic Branches. Splenic Branches. Left Gastric, j Superior Gastric. I Inferior ditto: ' Pancreatic Branches. Duodenal. Right Gastric. Right and Left Hepatic. Duodenal Branches. SmaU Mesenteric. Coecal Branches.' Anterior Colic Branches. Right Renal. ) External Branches. Left ditto. \ Capsular Renal; Spermatic <( Right and Left, „ , . n/r i ■ i Posterior Colic. Posterior Mesenteric j -p , . ^ Lumbars — five or six pairs. Bifurcation of the Posterior Aorta into External and Internal Iliac Arteries. (Umbilical. Vesical Branches. Prostatic. Anal and Perineal Branches. Arteria Innominata. Foraminal Branches. Ischiatic. r Splenic CffiUe <^ Gastric Hepatic Anterior Mesentric Renal Internal Iliac. Obdurator Pubic Int. Pubic Branches to the Crus Penis. Ditto Corpus Cavernosuni. Ditto Dorsum Penis. Ditto Glans Penisi Cutaneous Branches. The Middle Sacral issues at the Bifurcation of the Trunk. f Circumflex of the Heum. r^ . , T1- Artery of the Cord. External Hiac -^ ArWa Pmfnndn J Arteria Profunda. -{ Epigastric. Femoral. Branch to the Groin. Branch to the Ring. External Pudic" 166 ANATOMY AND PHYSIOLOGY OF Femoral - Inguinal. Muscular Branches. Stifle Branches. Muscular Branches. Popliteal, -j Recurrent. Anterior Tibial Posterior Tibia ' Recurrent Articular. Muscular Branches. Cutaneous Branches. Metatarsal Branclies. Metatarsal Ai-tery. . Muscular Branches. Medullary. Tarsal. Internal Metatarsal. -{ RecuiTcnt. Recurrent. External Plantar. Internal Plantar. DISTRIBUTION OF VEINS IN THE HORSE. The two main venous trunlvs, the vence cava, anterior and posterior, correspond to the anterior and posterior aortas. THE ANTERIOR VENA CAVA Forms the main trunk of the veins, re- turning the blood from the head, neck, chest, and fore extremities. It is principaDy formed by the concurrent union of the jugular and axillary veins, and is situated at its forma- tion in the space between the two first ribs, about midway between the sternum and vertebras ; it also receives the pectoral, ver- tebral, dorso-cervical, and inferior cervical veins, and the vena azygos. THE JUGULAR VEIN. It passes behind the condyle of the lower jaw, under the parotid gland, and joins the external carotid artery, and continues its course down the neck with the latter. It now receives the auricular veins, anterior and posterior, and also internal. The next is the temporal, the third is the internal maxillary ; the latter in its course receives the blood of many small veins, — the palato- maxillary, infra and supra orbitar, ocular, inferior maxillary, and deep temporal ; the fourth branch, received by the jugular vein, is the parotideal, and the last branches from the masseter muscles. THE OCCIPITAL VEIN Descends from the head, along with the occipital artery. It brings blood from the occipital sinuses, receives veins from the posterior lobes of the cerebrum and cere- bellum ; also from the dura mater. The submaxillary vein is a large branch of the jugular. It is formed upon the side of the face by the concurrence of the facial, labial, and varicose veins. It joins the trunk by the side of the trachea, just below the parotid gland. In its course it receives a number of veins ; the principal are — the submental, sublingual, lingual, pharyngeal, and superior laryngeal veins. The facial vein results from an expansion of small veins upon the side of the face, one of which is the varicose from the masseter. The labial vein is formed by the union of a plexus of venous branches, coming prin- cipally from the angle of the mouth, joined by others both from the upper and lower Ups. The varicose vein is buried in the masseter. The jugular trunk having received the submaxillary, proceeds down the neck, and terminates in the anterior vena cava, withiu the space between the two first ribs. Near the junction of the submaxillary the jugu- lar receives the small thyroideal, cutaneous, muscular, and tracheal veins. Near its termination it receives a branch of the superficial brachial, and plait or plat vein. The vertebral vein runs the same course as the artery, through the foramina, in the transverse processes of the cervical verte- brse, with the exception of the last. This THE HORSE. 167 vein has communications with the occipital sinus and posterior cerebral veins, medulla oblongata, and spinal marrow ; it also re- ceives vessels from the deep-seated mus- cles in the vicinity, and ends in the an- terior vena cava, just behind the first rib. The axillary vein returns the blood dis- tributed by the axillary artery to the va- rious parts of the fore extremity ; there is a superficial and deep-seated set; the former run under the skin, the latter among the muscles. The plantar veins are an intri- cate network of small veins, and cover the foot with a venous netting. The veins of the sole pour their blood into the veins of the lamina ; the latter increase in size towards the coronet, and gradually unravel themselves, so as to collect in a great many branches ; these run upward, through the substance of the coronary ligament, and form the superficial coronary vein ; from them other branches proceed and join the deep coronary, and afterwards unite in a single vein opposite the pastern joint. The veins of the frog, oiiex ramifying in the form of networlc over that body, ascend into the heel, growing larger as they leave the foot ; they make a single branch at the pastern joint, then unite wdth the vein com- ing from the laminae, thereby forming the plantar vein. The plantar vein ascends, unites with other vessels, and becomes metacarpal. The vietacarpal veins, two in number, result from the union of the plantar ; these veins pursue their course up the leg, one on either side, to the back of the knee, where they end in anastomosis. The internal metacarpal vein preserves the line of the splint bone. These vessels receive in thefr course cutaneous veins from the front of the canon, and one or two descending veins from the back of the leg; it after- wards forms the deep-seated veins of the arm. The superficial brachial vein ascends along the inner side of the radius to the elbow-joint ; here it crosses over to the front of the biceps and pursues its ascent upon that muscle toward the point of the shoul- der, and then passes inward to the jugular vein. In its com'se to the latter, it receives numerous cutaneous and muscular branches, communicates with the humeral vein, and anastomoses with other veins of the arm. The radial veins, two in number, arise from the junction of the metacarpal veins above the loiee ; they take the course of the radial artery, and receive anastomosing vessels as they ascend from the ulnar and superficial veins. The ulnar veins (witii one exception) end in the common trunk of the humeral vein. The humeral vein accompanies the ar- tery ; it receives small veins from the mus- cles. Tlie axillarij vein is the continuation of the humeral, augmented by the accession of the triceps vein. Its branches are, the subscapular vein, and dorsalis scapular; the latter terminates about midway be- tween the chest and shoulder. The re- maining branches of this vein are the humeral thoracic, and the external thoracic ; it also receives other small veins, which contribute more or less to its volume. The pectoral vein runs the course of the pectoral artery. It originates in branches from the abdominal parietes, continues to receive accessory vessels in its course, and ascends along the inner and lower border of the first rib. The dorso-cervical vein consists of two divisions, ramifying mth the dorsal and posterior cervical arteries ; it receives the anterior intercostal vein. The inferior cervical vein rmis down the lower part of the neck in company with the artery ; the principal branches are muscular, though some come from the skin and ab- sorbent glands in the vicinity. Tlie vena a-ygos ends just as the trunk opens into the auricle ; it returns the blood from the lower intercostal veins. THE POSTERIOR VENA CAVA. This is the corresponding venous trunk to the posterior aorta, retiu-ning the blood from the parietes of the abdomen and pel- 168 ANATOMY AND PHYSIOLOGY OP vis, the urinary and genital organs, and the posterior extremities. It takes its course imder the bodies of the lumbar vertebrae, runs along the great fissure of the liver, perforates the cordiform tendon, and pur- sues its way directly across the cavity of the chest to the lower part of the right auri- cle : in its passage it is joined by the lum- bar spermatic, renal, hepatic, and diaplnagm- atic veins. The common iliac veins are formed under the sacro-iliac, symphysis, by the union of the external and internal iliacs ; they re- ceive a vein from the psoas and Uiacus, cir- cumflex vein of the ileum, middle sacral, and azygos. The ischiatic vein, situated upon the side of the pelvic cavity, midway between the external iliac and lateral sacral veins ; ex- ternal and internal branches unite to form it.. The internal comprise veins coming from the bladder, anus, perineum, and, in the male, from the bulb and prostate: in the female, fr'om the vulva and body of the vagina. The external come principally from the gluteal and obturator muscles. The lateral sacral vein comes from the tail, formed by coccygeal veins ; it runs forward to the sacrum, and receives in its course the permeal and sacro-spinal branches. The external iliac vein takes the same course as the artery ; as it departs from the belly, this vessel receives The inguinal vein (coming fi-ora the groin), also a superficial or cutaneous abdominal vein, known as the milk vein in cattle. The femoral vein is the continuation of the iliac trunk below the brim of the pelvis ; and is the main channel into which the deep-seated veins of the hind extremity poiu their blood. We commence the de- scription, as in the fore extremities, at the leg. The large metatarsal vein ascends the canon by the side of the flexor tendons, and passes over the front and inner part of the hock ; it sends out branches, from which result the Ajiterior tibial veins, which run between the tibia and fibula to the back and lower part of the os femoris, and then are joined by the posterior tibial vein, and all three tmite to form the femoral. 2%e posterior tibial vein is a continua- tion of the smedl metatarsal vein, and cor- responds in size to the small metacarpal. It runs in company with the posterior tib- ial artery, receiving various muscular branches in its course, also the medullary vein of the tibia. The femoral vein results from the t\\'o last-named vessels ; runs behind the femoral artery, and ends in the external iliac vein. It receives muscular veins, as well as veins from the stifle joint, and the medullary vein of the OS femoris ; also, about two-thirds of its length upwards, it is joined by the saph- ena vein. The vena saphena major results from the large metatarsal vein ; at the hock it anas- tomoses with the anterior tibial vein; it also receives cutaneous and muscular branches in its course. The vena saphena minor springs from the small metatarsal vein ; it runs up the back of the hock, over the root of the os calcis, and ultimately reaches the femoral vein. The vena porta circulates the blood through the liver, and is principally formed by the union of the splenic and mesenteric veins. THE HOUSE. 16J> bo ( ^^^:n f^ != 2 £ »: ." 5 o o .2 i- S S S !S n^ ^ •! ^ 5 ?1 S ^-S t5 .2 a e ^ -g •X3 g 5 5 d o o H r^S t; -Q :^ G 13 c3 -3 be ;a.<0 o S c g ? 3 „ w w »w C3 r- 3 .~ ^ ^ ►::;Q fe i-^ > co CO J Ph m o <^ o .5 .2 '3 lo 170 A^■ATOMY AND PHYSIOLOGY OF g o "-B o o S D O 5 ri o o OOOPh Op5 S§ = =3 1;; P_ §g ■^ c c; C rt I " 5 S .2 •" 'rt H Ph K p3 O EXPLANATION OP PIGURE XVII. OSSEOUS STRUCTUKE. 1. Frontal bones. 3. Occipital. 4. Temporal. 8. Superior maxillaris. 10. Inferior " 11. Cervical vertebrse. 14. The sacrum. 16. The false ribs. 18. The sternum. 19. The ileimi. 20. The ischium. 21. Pubis. 22. Femur. 23. Patella. 24. Tibia. c. Fibula. 25. Os calcis. 26. Tarsal bones. 27. The mferior tarsal bones. 28. Metatarsus magnum. 29. Sessamoides. 30. Os suf&aginis. 31. Os corona. 32. Os pedis. 33. Scapula. 34. Os humeri. 35. Radius. f. Os uhiaris. 36. The carpal bones. 37. Metacarpus magnum. 38. Scssamoida. 39. Os corona. 40. Os sufh-aginis. 41. Os pedis. d. Dorsal spines. MUSCULAR STRUCTURE. a. Orbicularis palpebrarum, c. Dilator naris lateralis. e. Orbicularis oris. f. Nasalis longus. li. Buccmator. j. Depressor labii. k. Masseter. 10. Region of the parotid gland. o. Adducens Tel deprimens aurem. EXPLANATION OF FIGUKE XVII. CONTINUED. THE NECK. 6". Cen-ical ligament. Ligamentum colli, c". Trachelo subscapularis, scalenus. s. Splenius. r. t. Tendon of the splenius and scalenus. u. Levator humeri. 6UPERI0E PART OF THE SHOCLDEK AND BACK. i". Latissimus dorsi. m". Trapezius. SHOULDER AND FORE EXTREMITIES. g. Ji". Spinatus muscles. I", in', n'. Triceps extensor brachii. s". x". Extensors metacarpi. p". q". Flexors externus et Intemus. It. u. u. u. Flexors perforans et perforatus. 8. The pastern. S,-. &{. Hoofs. ABDOMINAL REGION. c". Abdominis transversalis externus. 4. Subcutaneous thoracic vein. POSTERIOR E-KTREMITIES. «'. Erector coccygis. g". Compressor coccygis. i'. Glutei. J'. Triceps. h'. Biceps abductor. v. Abductor tibialis intemus. r'. r. s'. Gastrocnemi. t!. Peroncus. g'. Extensor metatar-si internus. w. Insertion of the gastrocnemius. V. Flexor perforans et perforatus. a;, y. Fleshy belly of the extensors. 8. The pastern. if. «,'. Hoofs. THE HORSE. 171 THE BRAIN AND ITS APPEN- DAGES. The cerebrum, cerebellum, and medulla oblongata, and medulla spinalis are invested with three membranes : the dura mater, pia mater, and tunica arachnoides. Of these the exterior is the dura mater ; which, though called a membrane, is of a dense, tough, and inelastic texture. It is so firmly adherent, by means of numerous little prominences, to the sutures of the cranium, that it is diSicult to separate them ; this membrane is to the internal cranium what the pericraninm is externally. The inner surface of the dura mater is lubricated by a fluid furnished by its own blood-ves- sels. The pia mater is that membrane which closely envelopes the substance of the brain, and dips down between its convolu- tions, and adheres to its surface by num- berless minute blood-vessels. It differs in its appearance and texture from the dura mater; presenting a smooth surface exte- riorly, but a rough and villous one next to the brain, and being composed of a beauti- ful network of blood-vessels united together by a delicate cellular tissue. The third membrane has been compared to a spider's web, in allusion to which the name of mcmbrana arachnoides has been given to it. The arteries which supply the brain with blood are the two vertebrals, besides two other branches called the internal carotids. Its blood is returned from the sinuses of the dura mater by the vertebral and jugular veins. It is on the supply of the vertebral ar- teries, however, that the brain mainly de- pends for its supply, for, if ligatures are placed on these arteries, the animal dies; whereas, both the carotids may be tied without occasioning any apparent ill effects. If our memory serves us. Dr. J. C. Warren, of this city, has performed the latter operation on the human subject with suc- cess. In raising the bony covering of the brain, we meet with two processes, called the falx, or longitudinal process, and the ten- torium, or transverse process. The former resembles the blade of a scythe, hence its name. These processes are formed from duplicatures of the dura mater; the first descends for a short distance between the lobes of the cerebrum. It takes its rise from the crista galii, and terminates on the OS occipitis. The tentorium is extended from the inner plate of the os occipitis along the sides of the cranium to its base, whence it may be traced to the sphenoid bone, and is lost in the common covering of the dura mater. It is composed of two laminae : one is continuous with the falx : the other forms that portion of the membrane which covers the cerebellum. The tentorium is equally divided by the fabc into two lateral por- tions. T/ie sinuses. — The superior, or longitu- dinal sinus, riins within the duplication of the faLx, along its superior border. The tivo lateral sinuses are formed within the duplicature of that part of the tento- rium which is attached to the temporal and occipital bones ; one extending to the right, and the other to the left. They receive veins both from the cerebrum and cerebel- lum. The cavernous sinuses, so named from the cavernous appearance of their interior ; they receive some important nervous trunks in their passage from the brain, and for lodging the terminations of the internal carotid arteries. They commonly commu- nicate with the sub-occipital sizius ; these are also of membranous formation, and are found upon the cuneiform process of the os occipitis, running longitudinally to the fora- men magnum. They receive veins from the cerebellum and posterior parts of the cerebrum. Cerebrum. — The largest portion of the cerebral mass, and that which presents it- self to our view in raising the skidl, is the cerebrum. It is equally divided by a longi- tudinal fissure along its middle, into which the fabc cerebri descends ; and its divisions, 172 ANATOMY AND PHYSIOLOGY OF which are symmeti'ical, both internally and externally, are denominated hemispheres. Cerebellum. — The cerebellum is at once distinguished from the cerebrum by its being only one-sixth the size of the latter. Its figure is irregular : it has two oval ends, and its lateral dimensions exceed its longi- tudinal. It is divided into three oblong lobes — a middle and two lateral. Medulla oblongata, the smallest division of the cerebral mass. It rests on the cunei- form process of the occipital bone, and is continued upward and backward to the foramen magnum. The upper surface of the medulla oblong- ata forms, with the tuber annulare, the floor of the fourth ventricle. Pituatary gland, a red body, of an egg form, seated upon the sella tursica, within a fold of the dura mater. It has a mem- branous capsule, surrounded by cellular ad- hesions, by which it is firmly retained in its place. Medulla spinalis. — The spinal marrow is that extended portion of brain-like sub- stance which is continued from the poste- rior part of the medulla oblongata through the entire length of the spinal canal. It is inclosed in the same membranes that en- velop the brain ; but, in addition to them, the superior ligament of the spine serves as a covering and defence to it below. To this, and to the periosteum lining the canal, its proper theca is loosely attached by cellular, adipose, and gelatinous matter. Its dura mater is derived from that which covers the brain: in being continued through the foramen magnum, the membrane is contracted into a cylindrical sheath which loosely encases the marrow, and is generally described under the denomination of theca vertebralis. The arachnoid membrane and pia mater have the same relation to the marrow that the same membranes have to the brain, of which they may be considered prolongations. ORIGIN AND DISTRIBUTION OF THE NERVES. The nerves, being symmetrical in number and distribution on either side of the body. take their origin in pairs, and these pairs are numbered, and so distinguished from one another, according to the order in which they arise. There are forty-six pairs of nerves ; ten, coming from the brain, are distinguished as the cerebral nerves ; thirty- six, from the spinal marrow, denominated the spinal nerves. CEREBRAL NERVES. First pair, or olfactory nerves, arise from the corpora strata, along the posterior bor- ders of which bodies the medullary bands or roots of them may be traced as high up as the middle lobes of the cerebrum. These are the largest of the cerebral nerves, are bulbous at their origin, pulpy in texture, and exhibit, when cut into, comparatively to their size, large cavities, which are walled in by a layer of medullary matter, enclosed wthin a thinner one of cortical substance. Second pair, or optic nerves, arise from the thalamia nervorum. They leave the cranium through the optic foramen, and pass to enter the globe of the eye, within the interior of which it expands, and forms the retina. In its whole course, it is en- closed within a sheath prolonged from the dura mater. Third pair, or mofores ocidorum, take their origin by several filaments, from the inward parts of the crura cerebri. The trunk of the nerve first runs obliquely out- ward, across the back of the crus, then turns downward and enters the cavernous sinus, on through the foramen lacerum-or- bitale. In entering the cavity, the nerve divides into two branches. The smaller is generally received by the levater oculi. The larger branch subdivides into several others ; the longest of these runs round the eyeball, and penetrates the oblique muscle. Two or three others run to the abductor and depressor muscles. Fourth pair of pathetic. — These take a filamentous origin, and pass the border of the tentorium, entering the cavernous sinus, from thence to the orbit. Its destination is the superior oblique muscle of the eye. Fifth pair, or par trigemini. — These are THE HOESE. 173 the largest nerves of the brain. They take thek origin by filaments from the crura cere- belli, and pierce the dura mater. Each nerve appears to form a ganglion; from this ganglion we say that three nerves de- part. One is called the ophthalmic; the second, the anterior maxillary ; the third is the posterior maxillary nerve. The oph- thalmic nerve is the smallest of the three divisions ; as it emerges from the orbit, it divides into three branches, caUed the lachrymal, the super-orbitar, and the lateral nasal branch. Tlie second division, or anterior maxillary nerve, leaves the cranium thi'ough the hole called foramen rotundum, of the sphenoid bone, and takes its passage through the inferior orbital canal, whence it emerges, covered by the levator labii superioris, upon the face : here it splits into several large branches, denominated the facial nerves. But prior to its entering this canal it detaches several important branches to the eyelid, lachrymal duct ;. also several long filaments, which descend on the tube- rosity of the anterior maxiUa, penetrate the bone, and furnish twigs to the antrum, and the two superior molar teeth. The largest branch is the spheno-palatine, or lateral nasal nerve, to which the foramen spheno- palatine gives passage into the nose, wherein it divides into two sets of fila- ments. One of these is spread over the lateral parietes of the nasal cavity; the other ramifies over the sinuses, and sends a filament to the lower border of the sep- tum. A branch also goes to the velum palati, and another branch accompanies the palatine blood-vessels, and ranaifies over the soft palate. The facial branches of this division ter- minate on the front and sides of the face, and receive communicating filaments from the anterior facial branch of the portio dura, and with them form a plexus. The third division, the posterior maxillary nerve, gives off" a branch which runs up in front of the parotid gland, and joins the portio dura ; also branches called the buc- cal nerve, pterygoideus, and gustatory. The latter descends by the side of the tongue, penetrates that organ about its middle, and vanishes in its tip. It also sends ramifica- tions to the roots of the incisive teeth, and to the under lip. Sixth pair, or ahducentes, arise by means of filaments from the medulla oblongata ; this nerve gives ofl^ two or three filaments to the retractor oculi ; but its principal des- tination is to the abductor, along the fasci- cula of which, its ramifications are equally distributed. Seventh pair, or auditory nerves. — This pair includes two separate nerves on either side ; one, from its remarkable softness, is denominated the portio moUis; the other, in conti-adistinction, the portio dura. The portio mollis enters the organs of hearing, and is distributed to the labyrinth. The portio dura arises from the medulla oblongata, and passes to the internal part of the ear, the tympanum, and eustachian tube. It is also distributed to the temples, eyelids, nose, lips, cheek, and neck. Eighth pair, or par vagum. — At its commencement it consists of two separate portions ; the first called the glosso-pharyn- geal nerve, and the second the true par va- gum. They arise from the corpora olivaria, and make their exit through the base of the cranium. The glosso-pharyngeus gives off branches, which join the portio dura, to the constrictors of the pharynx, and form branches which ramify in the base of the tongue. The proper par vagum, having disunited from the glosso-pharyngeal nerve, proceeds downwards to join the carotid artery, and takes its com'se along the neck to the chest. Its filaments are — 1. To the cervical gan- glion. 2. The pharyngeal branch, whose filaments pass to the esophagus and larynx. 3. Two slender branches to the carotid ar- tery, which form a plexus. 4. The laryngeal branch. At the back part of the neck the pai- vagum inclines upwards, and is found above the carotid artery ; it then passes between the two first ribs into the chest. Having entered the thoracic cavity, it runs 174 ANATOMY AND PHTSIOLOGT OF witliiii the superior mediastinum ; the right nerve adheres to the trachea, crosses above the root of the right lung, alongside of the esophagus, and gains the under side of that tube before it leaves the chest. On the left side the nerve accompanies the anterior aorta, and crosses the root of the posterior aorta, and also reaches the esophagus. Its branches mthin the chest are fUaments to the tracheal and cardiac plexuses ; also, a branch called the recurrent nerve ; branches to the pulmonary plexus, and also t^vo cords that branch out and penetrate the walls of the amides. The recmTent nerve of the left side originates from the par vagum, by the side of the anterior aorta, and coils round the root of the posterior aorta. The recun-ent nerve, so denomuiated from its retrogi'ade course, passes upwardly and outwardly, and is fomid between the caro- tid artery and the trachea ; having reached the top of the latter, it spreads into fine ter- mmating branches, several of which run to the muscles of the laiyixs and thyroid carti- lage, and end in ramifications upon the membrane of the glottis. Its branches are filaments to the pulmonary plexus, cardiac plexus, posterior cervical ganglion, and branches to the esophagus and ti-achea. The par vagum runs to the stomach. The left nerve sends filaments to the heart, and others along the small cm'vature, which communicates with the ramifications of the right nerve ; the other crosses to* the left side, and joins the great semilunar ganglion. The right nerve, as soon as it reaches the heart, divides into numerous branches, which join the left, and spread their ramifi- cations upon the under part of the heart ; some run to the pylorus, and others join the hepatic plexus. Accessory nerves to the eighth. — These nerves are considered as accessory to the eighth, in consequence of their being found in close connection in issuing from the cranium; it originates in the vertebral canal, by the union of several fUaments. In its course into the cranium it receives many other fine threads, and in that cavity joins the par vagum. Beneath the atlas, the accessory nerve divides ; the front division runs downward, and penetrates the beUy, transmitting side twags in its course. The posterior division turns round the transverse process of the atlas to the scapula, near which it is lost in muscular substance. The branches of the accessory pass to the par vagum, anterior cervical ganglion, and communicate with the sub-occipital nerve. Ninth pair, or linguales, arise behind the eighth pair, from the corpora olivaria ; it is found in company with the par vagum, near the coronoid process. The nerve passes down the lower jaw, between the muscles forming the root of the tongue, and ends in the tip of the latter. It sends branches to the lingual muscles and to the hyo-glossus longus. Tenth pair, or sub-occipital nerves. — They arise from the meduUa oblongata, and be- ginning of the spinal maiTOw; they pass out through a hole in the fore part of the body of the atlas. It then branches into a superior and inferior division. The superior is distributed to the extensor muscles of the head and neck. The inferior branch goes to the trachea, lymphatic glands, and mus- cles of the neck. CERVICAL NERVES. These consist of seven pairs, originating from the cervical portion of the spinal mar- row. Each nerve, as soon as it issues from the spinal canal, forms two nervous fila- ments, one superior, the other inferior. The first cervical nerve makes its exit be- tween the first and second cervical vertebree. It sends branches to diiferent muscles, and communicates with the Second cervical nerve, which makes its appearance between the second and third vertebrae. Its superior filament sends branches to the muscles of the neck, and levator humeri, communicates with the ac- cessory ner\'e, and Third cervical. — This also sends branches and twigs to the different muscles of the neck, and communicates with the fourth. The fourth, fifth, sixth, and seventh pairs THE HORSE. 175 pass from the spine, between their respec- tive vertebrEB, and send branches to the phrenic nerve, and ramifications to the mus- cles, sympathetic nerve, and unite with the dorsal. THE DIAPHRAGMATIC OR PHRENIC NERVE. This is formed by branches from several of the cervical nerves. It takes its course down along the inferior border of the scale- nus muscle. It terminates by numerous ramifications on the tendinous parts of the diaphragm. DORSAL NERVES. These consist of eighteen pairs. They pass from the vertebral canal in the same manner as the cervical, having superior and inferior branches. The inferior branches foUow the course of the intercostal blood- vessels, and are called intercostal nerves. The superior branches are distributed to the back and loins. LUMBAR NERVES, Consist of five pairs (corresponding to the number of the lumbar vertebras). The first nerve ends in ramifications near the stifle, and gives off" branches to the last dorsEd nerve, to the sympathetic, and to the second lumbar nerve. The second nerve has communication with the first nerve, and sympathetic ; also the crural. It sends one division to the fore part of the haunch, where it becomes sub- cutaneous, and ramifies over the stifle. The other division crosses the ilio-lumbar artery, just below its origin, and takes nearly a similar course to the inward part of the haunch, and then ramifies upon the skin; in its way it detaches a considerable branch, called the spermaticus externus, which passes through the abdominal ring, and sends twigs, in the male, to the scrotum and tes- ticle ; in the female, filaments go fi-om it to the uterus, udder, and external labia. The third nerve contributes to form the crural and obturator. It sends small branches to the sympathetic, psoas, and obturator nerves. Tlie fourth nerve sends a branch to the sympathetic, conti-ibutes to the production of the crural ; and also sends a branch to the obtm-ator. The fifth nerve communicates with the sympathetic, craral, and sciatic plexus. SACRAL NERVES, Consist of five pairs; a superior and ir- ferior fascicula. The superior make thej exit through holes upon the upper part o the sacrum, and are there bmied rmder a thick mass of muscle, and become cutane- ous upon the outer part of the haunch. The inferior fascicida. — The first nerve largely contributes to the origin of the sciatic plexus, and sends a branch to the gluteal ner\'e ; also to the sympathetic and second lumbar nerves. The second nerve communicates Avith the tliird and sympa- thetic, and sends branches to the surround- ing muscles and sciatic plexus. The third and fourth have similar connections. The fifth passes into the coccygeal muscles. COCCYGEAL NERVES. These issue from the spine, in the same manner as the last described. They com- municate with one another, ai-e distributed to muscles in the vicinity, and end in fila- mentous ramifications at the end of the tail. NERVES OF THE FORE EXTREMITY. The fore extremity receives its nerves from the axillary or humeral plexus, and this plexus is formed by the union of por- tions of the sixth and seventh cervical nerves, and a division of the first dorsal ner\'e. The external thoracic nerves, sLx or seven in number, arise from the humeral plexus, and are distributed to the pectoral, triceps, and other muscles ; they finally ramify into the skin. Tlie scapular nerves are called anterior, posterior, and sub-scapular. The former sends its ultimate filaments to the triceps. Tlie posterior scapular nerve sends branches to the sub-scapularis, triceps, teres 176 ANATOMY AND PHrSIOLOGV OP minor, and shoulder joint, and ends in the insertion of the levator humeri. The subscapular nerves run upward be- tween the shoulder and chest, and enter the subscapularis. The spiral or external ciBaneous nerve is furnished by the axillary plexus ; arises be- hind the humeral artery, and passes between the OS hiimeri and the head of the triceps, througji the extensors, to the external flexors of the canon. It gives off several branches to the triceps, ramifies on the fore and out- ward part of the knee, and sends branches to the heads of the extensor muscles. The radial nerve descends with the humeral artery to the inward side of the elbow joint, and runs along the back part of the radius to the knee; passing under the annular ligament, it descends to the leg, and takes the name of the internal metacar- pal nerve. It gives off numerous twigs to the muscles, and finally becomes subcutane- ous. The ulnar nerve origmates from the humeral plexus. It passes down the radius, under the annular ligament, to the tendo perforans, and there becomes the external metacarpal nerve. It gives off internal cutaneous and subcutaneous branches, rami- fies into cellular substance, penetrates the heads of the flexors, and finally disperses its ramifications in front of the leg. The metacarpal nerves continue down the leg, over the fetlock joint, where they become the plantar nerves ; these pursue their course behind their corresponding blood-vessels to the back part of the foot, which they penetrate to the inner side of the lateral cartilages. The plantar nerve detaches a branch from the fetlock to the lateral cartilage ; another passes to the fatty frog. The final branch enters a hole in the back and lower part of the coffin bone, in company with the plan- tar artery, and there divides and distributes its ultimate branches around the edges of the sole. NERVES OF THE HIND EXTREMITY. The crural nerve is derived partly from the second, third, fourth, and fifth lumbar nerves. It makes its appearance under the transverse process of the loins, and proceeds in a line with the external iliac artery. It gives off filaments to the psoas magnus, niacus, rectus, and vastus internus muscles. It also gives off cutaneous filaments ; one runs to the stifle, and ends in ramifications upon the fore part of the thigh. The other continues down the leg, and can be traced as low as the fetlock. The obturator nerve, contiibuted to by thnd and fourth lumbar nerves, sweeps round the brim of the pelvis, and detaches tsvigs to the obturator muscles. Its ultimate filaments are expended on the triceps and gi'acilis. The gluteal nerve, after leaving the cavity of the pelvis, accompanies the gluteal artery, and passes into the substance of the gluteal muscles. The sciatic nerve derives its origin from the sacral and last of the lumbar nerves ; after leaving the cavity of the pelvis, passes between the hip joint and the tuberosity of the ischium, and plunges into the substance of the haunch. Here it divides into branches called the popliteal nerves. At the hock its principal branch separates into the external and internal metatarsal nerves ; the former runs over the flexor pedis to the os calcis. Their subsequent course and ultimate dis- tribution are the same as those of the plan- tar nerves of the fore extremity. The second popliteal nerve passes between the bellies of the gastrocnemii, above the first, detach- ing twigs to them in its passage, and then spreads into many branches, which pene- trate the heads of the flexor muscles of the foot, and send filaments into the stifle joint. SYMPATHETIC NERVE. Tills nerve derives its name from the uni- versal influence which it has on the nervous system. It communicates with the head, neck, chest, pelvis, and abdomen, by its fre- quent intercourse and connection with their respective nerves. It is supposed by some writers to be a nervous system of itself It has, at different distances, a great number THE HORSE. 177 of gangliform tubercles, from which ramifi- cations proceed forward, as well as filaments backward, to the ganglia of the nerves of the medulla spinalis. It is considered gen- erally as beginning from a branch of the fifth and sixth pair, given off at the base of the cranium. The ganglionic structures and the different plexuses are named from their form, location, and distribution ; hence we have the cervical ganglion, semilunar, sacral, etc. From the semilunar ganglion nervous filaments shoot in various direc- tions, which, from their being compared to the rays of the sun, are denominated the solar plexus. From the divergent filaments of the latter, the several smaller plexuses of the abdomen may be said to derive their formation, taking names according to the viscera they are particularly designed to furnish with nerves ; hence we have the splenic plexus, that sends filaments to the spleen, the hepatic plexus, mesenteric, aortic, hypogastric, and renal plexuses. The sym- pathetic nerve in the abdomen travels over the sides of the bodies of the lumbar verte- brae, below the articulations of the ribs, and pursues its course into the pelvis. Here, also, it forms ganglia, which correspond in number to those of the lumbar nerves : and from every ganglion come off two filaments: one which runs to the corresponding lumbar nerve ; the other crosses the aorta, and, by joining the aortic plexus, communicates with nerves coming from the sympathetic of the other side. From the loins, the sympathetic descends into the pelvis, and takes its course along the side of the sacrum, and forms five gan- glia, corresponding to the sacral nerves ; it finally terminates by forming a union with its fellow. EXAJnNATIONS ON NEUROLOGY. Examinations on Neurology, which will include the names of parts not alluded to in the preceding summary of the nervous system. yiERTES. Q. What are nerves ? — A. Long, firm, and wliite chords, which ramify- after the manner of blood-vessels, and are distributed to all parts of the horse's body. Q. Where do they arise? — A. From the brain, medulla oblongata, and medulla spinalis. Q. What communications have the different nerves with each other? — A. They anastomose: forming sometimes a plexus; at others, a knot or ganglion, from wliich other branches arise. Q. What is the structure of nerves? — A. They consist of fascicuh, or bundles, of distinct longitudinal fibres, closely connected together by cellular substance. Q. What are the coverings of nerves ? — A. Contin- uations of those which envelop the brain and spinal marrow, termed nemilcma. Q. What is the structure of ganglions ? — A. They are formed by a close intermLxtiu-e of filaments. BRAIN AND ITS MEMBRANES. Q. Where is the brain situated? — A. It occupies tlie cranial canity. Q. How is the brain divided? — ,-1. Into cerebrum, cerebellum, and medulla oblongata. Q. By what membranes is the brain enveloped ? — A. By three membranes, or meninges : 1st, The dura mater ; 2d, Pia mater ; 3d, tunica arachnoides. DUIl.4 MATER. Q. What is the situation of the dura mater? — A. It is the external covering of the brain. Q. How does it differ from the other coverings of the brain? — A. It is more dense, tough, and inelastic. Q. How is it retained mthin the cranium ? — A. It is fu-mly adherent to the interior of the cranium, more particularly to the depressions between the teeth of the cranial sutures. Q. How does the internal differ from the external surface ? — A. It has a smooth, pohshed, and lubricated surface. Q. Is the dura mater supphed with nerves? — A. Being composed of tendinous fibre, it is supposed to be destitute of nerves. Q. How are the processes of the dura mater formed ? — A. By duphcatures. Q. What are the use of the processes? — A. They steady and protect the various dinsions of the brain. Q. By what names are the processes known ? — A. The longitudinal process is called falx cerebri, and the transverse ditto is called tentorium. Q. What is the situation of the falx cerebri ? — A. It forms a partition under the anterior and superior parts of the cranial cavity extending from the crista galU to the occiput, and ends in contmuity with the tentorium. Q. What is the situation of the tentorium cerebelU ? — A. It is extended, after the manner of an arch, from the cerebral plate of the occipitis along the sides of 178 ANATOMY AND PHYSIOLOGY OP the craniiun to its base ; whence, greatly diminished in breadth it continues onward to the os sphenoides. SDTDSES OF THE DUKA MATER. Q. What are the names of the principal sinuses of the dura mater? — A. The superior or longitudinal sinus; two lateral, cavernous, and sub-occipital sinuses. PIA MATER. Q. What is the situation of the pia mater ? — A. It surrounds and closely invests the convolutions of the brain, and passes into the ventricles, furnishing them with an internal membrane. Q. What is the structure of the pia mater? — A. It presents a smooth exterior surface ; next the brain it is rough and villous, and is composed of a network of blood-vessels, which are united together by a delicate cellular tissue. Being highly vasciJar, it is supposed that the blood-vessels of the brain ramiiy- in it before entering the latter. AR.\CirXOID MEMBRANE. Q. Where is the tunica arachnoidea situated ? — A. It is a delicate and transparent membrane, spread uniformly over the surface of the brain. CEREBRUM. Q. Where is the cerebrum situated? — A. It occu- pies the superior part of the cranium. Q. What is its form, and how is it dinded ? — A. It is oval, convex above and concave below, and is divided by a longitudinal fissure along its middle, into which the falx cerebri descends. Its divisions are denom- inated hemispheres. Q. What is the appearance of the sm-face of the cerebrum? — A. It is covered with eminences called convolutions. Q. Of what is the substance of the brain supposed to consist ? — A. Of two kinds of matter ; the external is called cortical or cineritious, and the internal is termed medullary. Q. What is the color of the cortical? — A. Red- dish-ash. Q. What is the color of the medullary portion? — A. Of a milk-wliite hue. CORPUS CALLO.SUM. Q. What is the situation of the corpus callosum ? — A. It is an oblong wliite body, located at the bottom of the fissure which divides the two hemispheres of the brain. ' Q. What does the corpus callosum join on each side? — A. Its edges blend mth the medullary sub- stance of the two hemispheres of the cerebrum. Q. What name is given to the medullary substance of both hemispheres, together with the corpus callosum, when the usual anatomical section is made? — A. By cutting off the hemispheres of tlie cerebrum nearly even with the corpus callosum, there is seen a large oval mass of medullary substance, called the centrum ovale. LATERAL \'ENTRICLES. Q. What are the lateral ventricles ? — A. Two ca^-i- ties situated beneath the corpus callosum and medullary arches of the cerebrum. Q. What di^■ides the lateral ventricles from each other ? — A. The septum lucidum. Q. Name the parts which are generally considered as the contents of the lateral ventricles. — A. They are the corpora striata, the hippocampi, plexus choroides, fornix, and the thalami nervorum opticorum. Q. What is the situation and form of the corpora striata? — A. They are found on the lower and back parts of the ventricles, projecting into the centre of the cavities, where they expand as they approach the septum ; grow narrower and recede from each other above ; below, they extend to the anterior cornua. HIPPOCAMPI. Q. What is the situation of the hippocampi? — .4. They occupy the superior spaces of the ventricles in contact with the septum. Q. From whence do they originate ? — A. From the centres of the hemispheres. Q. AVhat is their structure? — A. They consist of alternate lamina; of medullary and cortical matter. PLEXUS CHOROIDES. Q. What is the situation of the plexus choroides ? — A. They are situated in the chaunel between the corpus striatum and hippocampus. Q. Describe the appearance of the same? — A. It is a soft vascular substance, consisting of a plexus of minute blood-vessels; it makes its appearance from behind the fornix, and ends abruptly in a round bulbous mass. FORNIX. Q. Describe the fornix audits situation? — A. The fornix is that part wliich receives the posterior border of the septum lucidum. It is extended after the manner of an arch, between the corpora stratia below and the heads of the liippocampi above, where it forms a junction with the corpus callosum. Q. Describe the processes or crura of the formx ? — A. The two inl'erior crura spring from the corpus albicantium, at the base of the brain, and finally unite ; thus united, they appear inthin the ventricles and con- stitute the body of the fornix. The superior crm-a proceed from the upper end of the fornix, and descend into the superior cornua of the lateral ventricles, and end in sharp, pointed extremities. TH.U..«IIA. Q. What is the situation of the thalamia nervorum opticorum ? — A. They form the upper and back parts of the lateral ventricles. Q. Describe the thalamia. — A. They have a wliite THE HORSE. 179 appearance, conoid in form, narrow and approximated inferiorly ; broad superiorly ; they finally contract into medullary bands, the tractus optici, which turn round the crura cerebri to the base of the brain. Q. How are the thalami distinguished from the corpora striata ? — A. They are more dense and firmer in composition. T-HNU. Q. "Whut is the situation of the taenia? — A. They are located in the groove between the thalamus and corpus striatimi, partly covered by the plexus choroides. COMMISUEES. Q. Name the commisures of the brain. — A. 1st, commissm-a mollis ; 2d, commisura inferior cerebri ; 3d, commism-a superior cerebri. Q. How is the commisura mollis formed? — A. By contiguous parts of the thalami, which are united by cortical matter. Q. How is the commisiu-a inferior cerebri formed ? — By a connection between the hemispheres of the brain. Q. Where is the superior commissure located ? — A. Above the commissura mollis ; it has the appearance of a short medullar)- chord. FORAMEN. Q. What is the foramen? — A. It is a triangular depression under the arch of the fornix, into which the lateral ventricles open. VEXTEICLES. lEemai'ls. — Having put the usual question regarding the lateral ventricles, which may be numbered 1 and 2, we now come to the tliird ventricle, which is not so well marked as in the human subject.] TnlRD \-EXTRICLE. Q. How is the third ventricle formed ? — A. By a mere fissure existing between the thalami. [Remarks. — The fourth ventricle, being located in the cerebellum, will be considered under this head.] INTTINDIBtJLUM. Q. Where is the infundibulum located ? — A. At the inferior part of the third ventricle. PDTEAL GLANT). Q. Where is the pineal gland located? — A. Be- tween the summits of the thalami, over the third ventricle, and above and before the superior com- missure. Q. Describe the pineal gland. — A. It is a small conoid body, of grajish color, marked by a slight depression along its centre. Q. What are its attachments ? — A. It is attached by means of the pia mater to the thalami and tuber- cula quadragemina. Q. What is the internal structure of the pineal gland ? — A. It consists of cortical and granular matter. NATES AND TESTES. Q. Where are the nates and testes situated ? — A. Above the third ventricle, behind the pineal gland, and immediately over and within the third and fourth ven- tricles. Q. How do the nates differ from the testes? — A. The former are larger than the latter, and are separated by a groove from the testes, and by a deep perpendicular fissure from each other. Q. What is their form ? — A. Semi-oval. Q. What is their composition? — A. They are com- posed of cineritous and meduUai-y matter. CEKEBELLtni. Q. What is the situation of the cerebellum ? — .4. In the inferior and posterior parts of the cranium. Q. How does the cerebellum compare in size vvith the cerebrum? — A. The former is only about one- sixth the volume of the latter. Q. Describe the appearance of the cerebellum ? — A. Its surface is lobular and convoluted ; its form is irregular, ha\ing two oval ends placed transversely, united in the centre by a broad vermiform belt; its lateral dimensions exceed its longitudinal. Q. How is the cerebellum di\ided? — A. Into three lobes, a central and two lateral. Q. How does the composition of the cerebellum difier from that of the cerebrum ? — A. In the former the cortical substance exceeds the medullar)-, and, instead of forming the bulk of the outer parts, as is the case in the cerebrum, it pervades the inner. FOURTH \-ENTRICLE. Q. What is the situation of the fom-th ventricle ? — A. It is situated between the cerebellum, tuber annu- lare, and medulla oblongata. Q. Where is the choroid plexus of the cerebellmn situated ? — A. Within and across the posterior part of the fourth ventricle, between the cerebellum and medulla oblongata. Q. How is the choroid plexus of the cerebellum dis- tributed ? — A. It is ilistributed into three divisions : one lies in the middle of the calamus ; the two latter are found within fissui'es in the cerebellum, occupjing the spaces between it and the tuber annulare. BASE OF THE BRAIN. Q. How is the base or posterior part of the cerebrum di\ided? — A. It is divided into six lobes. Q. Describe their divisions. — A. There are two anterior or inferior, resting upon the wings of the ethmoid bone ; two middle, upon those of the sphenoid j and two superior or posterior, lodged in the fossa of the squamous portions of the temporal bones. Q. What name is given to two broad, smooth promi- nences which appear over the middle lobes at the base of the brain ? — A. These are the corpora striata. Q. "What nen-es originate from this vicinity? — A. The olfactorv nenes. 180 ANATOMY AND PHYSIOLOGY OF THE HORSE. Q. What lobes rest on the wings of the sphenoid bone? — A. The crura cerebri. Q. From whence do they arise ? — A. From the in- ferior and middle lobes of the cerebrum, and are con- tinueSKnto an ovoid protuberance above them, named tuber annulare. Q. What is observable between the crura cerebri ? — A. A small hemispherical medullary eminence, called corpus albicantium. Q. Where are the tractus optioi situated ? — A. They wind obliquely downward around the crura. Q. Where do they proceed from ? — A. From the terminations of the thalami. Q. What is the situation of the crura cerebelli ? — A. They are located higher up and in a more outward direction than the crura cerebri. Q. Describe the crura cerebelU. — A. They are two cylindroid, medullary chords, which join the lateral lobes of the cerebellum to the tuber annulare. Q. What does"" the tuber annulare rest upon ? — A. On the cuneiform process of the posterior occipital bone. Q. Where are the foruminss caeca situated ? — A. Above and below the tuber. Q. What is their appearance? — A. They are described as little, round depressions, or blind holes. EXPLANATION OF FIGURE XYJU. [from BI^rNE's ■* OCTLIXES.'*] A. The skull, face, and upper jaw, in one piece. B. Lower jaw. a. Incisor teeth. 6. Tushes. c. Molares, or grinders. d. Peak formed by the extremities of the nasal bones. e. Zygomatic spine, to the bottom of which the masseter takes its origin. /. Orbit. g. Ca\-itv above the orbital arch. A. Pole.' i. Zygomatic arch. j,j. Styloid processes for the attachment of the muscles. it. Joint formed by the upper and lower jaws. I. Meatus auditorius, or opening to the internal ear. C. C. Marks the extent of the cervical vertebrse. D. Dentata. m. Atlas. n. AVing of the atlas. 0, Large superior spine of the dentata. p. Body of the dentata. q. Liferior spine of the dentata. s, s, s, s, s. Superior spines of the five remaining cer\-ical vertebrse. r, r, r, r, r. Oblique processes of the five last cervical vertebrte. u, ti, II, u, u. Transverse processes of the same bones. t, t, t, f, t. Inferior spines of the five last cervical vetebrae. THE THORAX. V, V. Cariniform process of the sternum. w, w, 10, w, w, w, w. Cost« or true ribs. y< y< 11' y> t/' V' y> V' H' y- ^^^ ^^ distinguished fi-om the costce. X, X, X, X, X, X, X, X, X. Cartilages by means of which the ribs are attached to the sternum. z, z, z, z, z, z, z. Heads of the ribs. 1, 1, 1, 1, 1. Superior spines of the first five dorsal vertebras, the fifth being generally the longest spine in the body. 2, 2, 2, 2, 2, 2, 2, 2. Superior spines from the sixth to the thirteenth, towards which they slope downward; the thirteenth is generally the most upright spine in the dorsal region. 3, 3, 3, 3, 3. Last five of the superior of the back spines, which have an inclination forward. THE LOWS, OR LtJlIBAR REGION. 4, 4, 4, 4, 4, 4. Superior spines of the lumbar region, thicker than the dorsal spines, and having a decided in- cluiation forward. 5, 5, 5, 5. Projecting transverse processes of the loins. THE SACRITM. 6, 6, 6, 6, 6. Superior spines of the sacrum leaning decidedly backward, thus leaving a large space between the points of the last lumbar and the first sacral spLue, at which place occm-s the great hinge of the back. 8, 8, 8, 8. Bodies of the sacral vertebrse. THE T.UL. 7, 7, 7, 7, 7, 7, 7, 7, 7. Coccygeal bones. THE PELVIS. E. Ossa innominata, consisting of three bones upon each side. a. Ilium. 6. Pubis. c. Ischium : the three bones unite at the cavity which receives the head of the thigh bone. 9, 9. The inferior spines of the ilium. 10, Superior spine, which partly covers the first sacral spine. c. r. Ischiatic spines. EXPLANATION OF FIGURE XVIII. OONTINUED. THE THIGH AND STIFIJ: JOINT. F, F. Femurs. d. Round head of the boue. e. Short neck of tlie femur. P. Great trochanter. g. Small external trochanter. h. Small internal trochanter. i, i. Sulcus whence the gastrocnemii muscles originate. J, J. Posterior condyles of the femur. h, k. Anterior trochlea over which the patella glides. 0, 0. Patellas : the interarticular cai'tilages of the stifle joint, as well as the cartilages tipping the dorsal lumbar sacral spines, and the su])erior margin of the blade bone or scapula, ai-e necessarily omitted in this delineation, which is admirably drawn from a macerated skeleton. THE TIBIA AND FIBULA, OK LEG BONES, AND THE HOCK JOINT. H, H. Tibias. 1, I. Heads of the bones. m, 711. Fibulas. n, n. Inferior head of the tibia. I, I. Hock joint. o, 0. Asti'agalus. p, ]}■ C'alcis forming the point of the hock. THE POSTEKIOR SHANK BONES. K, K. Canons, metatarsals, or shank bones. L, L. Splint bones. THE BONES OF THE PASTERNS, AND FEET, OF THE POSTERIOR LIMBS. M, M. Sessamoids. It, N. Large pastern bone. O, O. Smaller pastern bone. p, p. Pedal bones. BONES OF THE ANTERIOR EXTREMITY. Q. Scapula or blade bone. a. Suijcrior margin whence the cartilage has been removed- b. Spine of the scapula. c. Anterior fossa of the scapula. d. Posterior fossa. e. Shallow cup which receives the head of the humerus : the cartilage, which is situated around the margin of this cup, and which serves to deepen it, has been destroyed by maceration. f. Tuberosity termuiating the spine of the scapula, whence the flexor brachii originates. It. Humerus or arm bone. g. Head of the bone. h. Smooth cartilaginous and synovial pulley over which the tendon of the flexor brachii plays. i. F.xternal trochanter of the himierus. j. Inferior head of the humerus. Ic. Pit into which the ulna is received. S, S. Ulna, the top of wliich is termed the olecranon. T, T. Radius. I. Head of the bone. m. Inferior head of the bone. TJ, V. Carpus or knee joint, consisting of two rows of bones. n. Trapezium, which gives security to the great flexors, and attachment to several of the lessor flexors of the fore leg. V, V. Canon or shanlc bone. o, 0. Head of the bone receiving the lower row of the bones of the knee. W, W. Splmt bones. p. Inferior head of the canon bone. X, X. Sessamoid bones. Y, Y. Large pastern bone. Z, Z. Small pastern bones. P. Pedal or coffin bone. DISTRIBUTION OF THE LYMPHATICS. Mr. Percivall remarks, in his lectures, that " no English veterinarian has, up to the present day (1820), been at the pains to demonstrate, practically, the particular distribution of the absorbing vessels of the horse. Professor Girard, whose ' Traite d' Anatomie Veterinaire ' does no less credit to the talent and industry of its author than honor to the veterinary school over which he presides, has presented us with an arti- cle on the ramification of the lymphatics, wliich I shall translate. " THE THORACIC DUCT. " The largest, longest, and most remark- able of the lymphatic vessels, in which terminate the majority of the lymphatics of the body, is situated within the thorax, on the right side of the dorsal vertebrae, be- tween the aorta and vena azygos; It re- ceives the lymphatics from the posterior extremity, pelvis, parietes, and viscera of the abdomen, head, neck, withers, and left anterior extremity. " It takes its origin under the loins, in a dilation or sinus situated at the root of the great mesenteric artery, and is named the receptaculum chyli : it directs its course forward, enters the thoracic cavity 'by the aortic perforation through the diaphragm, extends along the bodies of the dorsal ver- tebrae, until it arrives opposite the base of the heart, where it curves downward to cross over to the left side in its way to the anterior opening of the thorax ; as it leaves the spine for this purpose, it runs over the trachea and esophagus ; having reached the left side, it stretches forward to the beginning of the anterior vena cava, and terminates in the base of the left axillary vein. Not unfrequently, it ends in the right axillary; in some instances, even in the be- ginning of the anterior cava. At its termi- nation, it dilates and forms a sinus, whose mouth opens into the vein, is guarded by a broad valve, so disposed as to prevent any reflux of blood into the duct* It has also a ligamentous band around it, at this part, wliich confines it to the vein receiving its contents. "the RECEPTACULUM CHYLI.f " This reservoir forms the point of general confluence of all the lymphatics of the pos- terior limbs and abdomen, and from which originates the thoracic duct. It is main- tained by the aorta on one side, the vena cava posterior on the other, and is formed by the union of five or six large lymphatics, of which two or tlu-ee come from the en- trance of the pelvis, two or three others from the iiicoeiiterj, a single one from the environs of the stomach and liver." The Professor here makes a classification of the lymphatics of the body. LYMPHATICS DISCHARGING THEIR CONTENTS INTO THE ABDOMINAL PORTION OF THE THORACIC DUCT. " 1. Lymphatics of the Posterior Extremi- ties. — These are distinguishable into the superficial and deep-seated. The first origi- nate from the skin and subcutaneous cel- lular tissue. They form divers ramifica- tions, which accompany the superficial veins; of which the most remarkable at- tend the vena saphena major, firequently anastomosing with one another, and form- ing an anastomotic network. All these lymphatics run to the subcutaneous ingui- * Notwithstanding this valve, blood often gains admis- sion into the canal ; this is observable in all cases of vio- lent death, or in which struggles and convulsions attend expiration. t Pei'civall's Lectures. (181) 182 ANATOMY AND PHYSIOLOGY OF nal glands, which are lodged upon the superior and anterior part of the thigh. " The deep-seated lymphatics take their rise from the foot, ascend along with the plantar veins, continue upward among the muscles, in company with the deep-seated veins, corresponding in their principal di- visions to those vessels, and proceed to the inguinal glands. "All the lymphatics of the posterior limbs assemble at these glands, and here form a plexus, from which several large branches depart and traverse the iliac glands, clinging to the sides of the iliac vessels, and discharge their contents into the pelvic branch, contributing to the recep- taculum chyli. " 2. Lymphatics of the Pelvis. — The vessels coming from this cavity run in part to the inguinal glands, and in part to the internal pelvic glands. The su- perficial lymphatics about the pubes and the outlet of the pelvis run and join those of the extremities ; those of the perineum and anus enter the cavity, and are accom- panied by those coming from the croup and tail, both proceeding to the glands ^vithin the interior of the pelvis. All the deep- seated lymphatics accompany the veins, make for the pelvic glands, form union with the others, and run and empty themselves into the main pelvic branch, wherem their lymph mixes with that coming from the in- guinal glands. " The lymphatics of the urinary and genital organs, included in the pelvic cavity, also traverse the glands lodged therein, and unite with those of the parietes of the pel- vis. Those of the scrotum enter the in- guinal glands, as also do those belonging to the sheath and penis. The ramifications derived from the testicle and spermatic cord take the course of the veins, and pene- trate one or two of the lumbar glands lodged at the entrance of the pelvis. The lymphatics of the mammae, which are also divisible into superficial and deep-seated, run to the inguinal glands, and anastomose with the superficial set belonging to the in- ferior parietes of the abdomen ; but, before they reach these last glands, they pervade those of the mammae. " 3. Lymphatics of the Parietes of the Ab- domen. — These vessels, in general but little developed, for the most part run to the in- guinal glands. The superficial set of the lower parietes accompany the cutaneous inguinal vein, anastomose with the lympha- tics of the scrotum and mammas, and tra- verse the glands in the groin : some of them direct their course forward, along with the cutaneous external thoracic veins of the thorax, unite with the superficial lymphatics of that part, and proceed to the axillary glands. The deep-seated vessels of the belly run in company with the epigastric vein, and go to the inguinal glands, or else they accompany the pectoral vein, and per- vade the glands in front of the thorax. " The superficial or subcutaneous l)mi- phatics of the loins join either those of the croup or those of the flanks : the deep- seated, which spring from the peritoneum, muscles, or spinal canal, perforate one of the lumbar glands, and pass onward to ter- minate in the main pelvic branch. " 4. Absorbents of the Mesentery. — The mesenteric branches, ordinarily two or three in number, the most considerable of which is constantly united to the great mesenteric artery, receive aU the vessels continued from the mesenteric glands, as well as those coming from the mesentery and intestines. " The mesenteric absorbents, extremely numerous, are sustained between the layers of the mesentery, where they form a vascu- lar network ; many of them issue from the exhalent surface of the mesentery and in- testinal tube ; others take their rise from the interior of the intestines, from which they imbibe chyle. All these vessels con- verge towards the lymphatic reservoir, clinging in their passage around the mesen- teric veins ; some, however, taking a solitary course at a greater or less distance from any blood-vessel. Having arrived at the root of the mesentery, they pass through one or two, sometimes three, of the mesenteric glands, and afterwards join the principal THE HORSE. 183 lumbar lymphatics. The absorbents of the colon and ccEcum caput coli run to the glands set at intervals along the intestinal tube, whence they proceed to the recepta- culum chyli. " 5. Lymphatics of the Liver, Stomach, Spleen, and Omentum. — The hepatic trunk comprises the lymphatics issuing from the above viscera. This branch of the recepta- culum chyli not uncommonly consists of two divisions, and receives in addition to the above-mentioned vessels many ramifications from the crura of the diaplnragm. " The lymphatics of the pancreas, lilie the above, also run with the divisions of its veins, and join either those of the liver or those of the spleen : some proceed di- rectly to the common hepatic trunk. " II. RAMIFICATIONS TERMINATING IN THE THORACIC PORTION OF THE MAIN COMMON DUCT. "1. Lymphatics of the Parietes of the Thorax. — The superficial absorbents of the chest take their ri^ either from the sur- face of the skin or else from the subcuta- neous muscles ; they form several large branches which accompany the thoracic cutaneous vein, unite with the superficial lymphatics coming from the anterior parie- tes of the abdomen, and proceed to the axillary glands. " The deep-seated set take divers direc- tions, and pass through the different sets of glands. The pectoral, which anastomose with ramifications from the abdomen, fol- low the pectoral vein, and reach one or two glands at the entrance of the chest. The intercostal spring from the pleura and in- tercostal muscles, accompany the intercos- tal veins, pervade the internal dorsal glands, and terminate by several branches in the thoracic duct. " The lymphatics of the fleshy part of the diaphragm unite, some with the poste- rior intercostal, others with pectoral ; those coming from the crura run to the dorsal glands, where they anastomose with the intercostal : those from the cordiform ten- don anastomose with the deep hepatic, run forward between the layers of mediasti- num, nearly to the heart, and enter the car- diac glands. " 2. Lymphatics of the Thoracic Viscera. — The absorbents of the difl'erent organs contained within the thorax traverse one or several of the bronchial or cardiac glands, and afterwards form divers branches, which end in the thoracic duct. The pulmonary lymphatics, very numerous, are distin- guished into superficial and deep-seated. The first take their rise from the surface of the lungs, creep along under their envelop- ing membrane, and make for one or more of the bronchial glands. The deep set, which originate from the air-cells and from the parenchymatous tissue, follow the di- visions of the pulmonary veins, run to the roots of the bronchia ; there miite with the superficial, and perforate one or two of the bronchial glands. " The cardiac lymphatics derive their origin either from the surfaces (both exte- rior and interior) of the heart, or from the muscular substance of the organ ; they mount upon the curvature of the posterior aorta, and disappear in the cardiac glands. " The lymphatics of the superior part of the mediastinum, and of the esophagus, join, some the intercostal, and others the bronchial ; those coming fi-om the anterior part of this membranous partition, from the thymus, trachea, and esophagus, unite, either with the pectoral, or close with the cardiac and anterior intercostal. " 3. Lymphatics of the Head. — The lymphatics of the head form two planes, a superficial and a deep one. The super- ficial pursue the course of the cutaneous veins, and run in part to the sublingual and utteral glands. The deep vessels, which come from the nostrils, fauces, palate, etc., also run to the gutteral and sublingual, in which they unite with the superficial. From these two groups of glands, through which pass the lymphatics of the head, de- part several large branches, two or three of which descend upon the anterior face of the trachea; others follow the course of the deep-seated and cutaneous veins, unite with 184 ANATOMT AND PHYSIOLOGY OF THE HORSE. those of the neck, and descend to the front of the chest. Almost all these vessels ter- minate in the thoracic duct; some few alone, on the right side, ending in the right axillary trunk. "4. Lymphatics of the Left Fore Extrem- ity. — The lymphatics of this member present the same disposition as those of the poste- rior limbs, and are divided into superficial and deep-seated. The former, consisting of diverse ramifications, accompany the superficial veins ; the more considerable of them forming a plexus, which accompanies the cutaneous (superficial brachial) vein of the limb. The deep vessels originate from the foot, muscles, and bones, pursue the divisions of the deep veins, and plunge into the axillary glands, wherein they mite with the superficial, and whence they ex- tend to the thoracic duct. " The Right terminating Trunk of the Lymphatics. — This very short lymphatic canal is obliquely situated at the entrance of the thorax, upon the transverse process of the last vertebrse of the neck, extending in a direction from above downward, and from without inward, and terminating most com- monly in the right axillary vein ; though, in some instances, it joins the thoracic duct. This trunk is formed by the lymphatics coming from the right axillary glands, and some from the right lung, and right side of the neck and trachea." (See Appendix.) EXAMINATIONS ON THE PHYSIOLOGY OF THE LYMPHATICS. Q. What is the character of the fluid found in the lymphatics ? — A. It resembles dilute, liquor sanguinis, or the Uquid portion of the blood in which the cor- puscles float. Q. What finally becomes of the lymphatic fluid ? — A. It was formerly supposed that the lymphatic fluid was eliminated from the system; but Carpenter and other physiologists now contend that this is not the case ; that the same is poured into the common recep- ticle with the nutrient materials newly imbibed from the food, whence both ai-e propelled together into the general current of the circulation ; and thus, instead of being eliminated, the lymphatic fluid is employed in the formation of new tissues. Q. From whence is the Ij-mphatic fluid derived? — A. 1st, from the residual fluid, which, ha^'^ng escaped from the blood-vessels into the tissues, has furnished the latter with the materials of their nutrition, and is now to be returned to the current of the circulation. 2d, from the particles of the solid frame-work which have lost their vital powers, and* are therefore unfit to be retained as components of the h\ing system; they therefore reenter the circulation, to be again submitted to the assimulating process, so that nothing shall be lost. Q. By what process do fluids enter the cutaneous IjTnphatics ? — A. By a process of imbibation. Q. What fluid is more readily absorbed than some others? — A. Milk. Q. What authority have you for this ? — A. Schoeger, in the coui-se of his experiments, found that the lympha- tics of a limb, long immersed in milk, became tinged with it, while none of it could be detected in blood drawn from the veins. ^kMrg 0f f d^rmarg Cei:|mtrii&s. (185) A GLOSSARY OF VETERINARY TECHNICALITIES. A. Abdomen. — The posterior part of the body of the horse. Ahdominalis. — Pertaining to the abdomen. Abdominal Regions. — The divisions of the exterior of the abdomen. Abductor. — Muscles are named abductors which draw parts from the axis of the body, or given centres. Abnormal. — Unnatural, irregular. Accelerator. — A muscle of the penis. Acetabulum. — A name given to the cavity in which the head of the thigh bone articulates. Achillis Tendo. — The tendon of the muscle inserted into the hock. Acuminated. — Pointed, like a needle. Adductor. — Muscles which draw parts toward the axis of the body. Adipose. — Fatty matter. Adventitious. — Accidental. Afferent. — A term used to designate the structures which convey fluids to different parts. Ala:. — Wings. Albumen. — An element which constitutes the chief part of the white of an egg. Alimentary Canal. — The passage which commences in the oesophagus and ends in the anus. AU-eolus. — The bony sockets of the teeth. Anal. — Relating to the anus. Anatomy. — To cut, with a view of displaying the struc- ture, relations, and uses of parts. Animus. — The principle of vitality. Annular. — A ring-like ligament, found at the posterior part of the knee of the horse. Antagonist. — A term applied to counteracting muscles or tendons. Anterior. — A term applied to what may be situated before another part of the same kind. Anti. — A prefix, signifying against. Antilabium. — Against the lips. Antrum. — Cavity in bones. Anus. — The posterior extremity of the rectum. Aorta. — The largest artery of the body. Aortic. — Pertaining to the aorta. Apex. — The pointed end of an organ. Aponeurosis. — A tendinous expansion of fibre. Arachnoid. — A membrane of the brain. Arch of the Colon. — Transverse portion of that intestine. Areola. — The spaces between fibres composing an organ. Arterial. — A property belonging to arteries. Arterialization. — The change which occurs in venous blood when brought in contact with air in the lungs. Artery. — The name of blood-vessels which distribute arterial blood. Articular. — Belonging, or relating, to joints. Articulation. — {From articidus.) A joint. Asperity. — A roughness. Astragalus. — The bone beneath the os calcis. Atlas. — The anterior bone of the neck. Attollens. — A name given to muscles which lift, or raise, the parts. Auditory. — Muscles and parts connected with the ear. Auricular. — Relating to the ear. Auricles. — The anterior cavities of the heart. Axilla. — The part between the superior region of the arm and the chest. B. Biceps. — (From, bis — twice, and caput — ahead; two heads.) The term is applied to muscles, having two distinct heads, or origins. Bifurcate. — (Bifurcas; from bis, twice, and furca, a fork). A blood-vessel or muscle is said to bifurcate when it divides into two branches. Bilary. — Relating to the bile. Brachial. — Of, or belonging, to the arm. Bronchia. — Bifurcations of the windpipe. Bronchial. — Relating to the bronchia. Buccal. — (From bucca, the cheek. ) Belonging to the cheek. Buccinator. — A muscle of the cheek. Bulb. — A dilated portion of the tube at the base of the penis. Bursce. — Sacs, or bags. Bursce Mucosa. — Sacs found in the region of joints. Bursal. — Relating to bursse. C(Ecum. — The blind gut. Ccecal. — Pertaining to the cajcum. Calcis Os. — The prominent bone of the hock. Cancelli. — Cellular structure of bones. Canine Teeth. — The eye-teeth, cuspidati. Canthus. — The angle of the eye. Capillary. — Hair-like vessels which are found between the arterial and venous vessels. Capside. — A membranous sac. Capsular. — A term applied to ligaments which surround articulations. Caput. — The head. Cardia. — The heart. Cardiac. — Pertaining to the heart. Carotid. — The name of the principal arteries of the neck. Carpus. — The bones of the knee. Caruncle. — A small fleshy excrescence. Carunculce Lacrymalis. — Small fleshy bodies found in the angle of the eye. (187) 188 GLOSSARY OP VETERINARY TECHNICALITIES, Caarfa. — The taU. Cava. — The largest vein in the body of the horse. Cavity. — A hollow part; the abdominal cavity, for example. Cellular. — Composed of cells. Centrum Ovale. — The appearance of the brain, when a horizontal section is made on a level with the corpus cal- losum. Centrum Tendinosum. — Tendinous centre of the dia- phragm. Cephalic. — Pertaining to the head. Cerebellum. — Inferior lobe of the brain. Cerebrum. — Superior lobe of the brain. Cerebral. — Relating to the brain. Cerebrospinal. — Pertaining to both the brain and spinal cord. Cervical. — Pertaining to the neck. Cervix. — The neck or contracted portion of an organ. ChorcB Tendince. — Part of the internal structure of the heart. Choroid. — The inner tunic of the eye. Chyle. — A fluid found in the thoracic duct and lacteals. Chyme. — A name given to the food after it has passed the pylorus. Cilia. — The eyelids, hair of the same, etc. Cineritious. — A term applied to that part of the brain which is of an ash color. Circuhis. — A ring. Clitoris. — A part of the pudendum of the mare corres- ponding to the glans penis of the horse. Coccyx. — The bones of the tail. Cochlea. — The spiral cavity of the ear. Cascum. — (Sometimes spelt ca;cura.) The blind gut. Cosliac. — Prolongation of the solar plexus, an artery and vein of the abdomen. Colon. — The largest and most dilated portion of the intestines. Cobimnce Camas. — A muscular arrangement within the ca-rity of the heart. Commisure. — A suture, junction, or joint. Complexus. — To embrace or smround. Concha. — External cavity of the ear. Conduit. — A canal. Condyle. — An irregular process or enlargement. Condyloid. — A tubercle, wart-like. Conglobate. — Ball-shape. Conglomerate. — An assemblage of glands. Conjunctivia. — External coat of the eyeball, and inter- nal lining of the eyelids. Conoid. — Conc-likc. Constrictor. — Muscles that are bound together are thus named. Tlie office is to close an outlet. Continuity. — Idcntitj- of parts, having direct connection. Convolute. — Rolled up. Coracoid. — Like a crow's beak ; a process of the scapula. Cornea. — Anterior coat of the eye. Cornu. — A horn. Corona. — A crown, the superior pastem is thus named : OS coronse. Coronal Suture. — The uniting medium between the frontal and parietal bones. Coronary. — Arteries and veins, proper to the heart, are thus named. Coronoid. — Processes of bones are thus named when they form an eminence. Corpora. — A term applied to numerous prominences in the brain and elsewhere. Corpus. — A body . Corpora Striata. — Striped eminences in the brain. Corpuscle. — A minute body. Corrugator. — A muscle which wrinkles the surrounding parts. Cortical. — Resembling bark. Costa. — A rib. Costal. — Pertaining to the region of the ribs. Costalis Pleura. — That portion of the plem-a which lines the interior of the chest. Cotyloid. — Cup-shaped. Cranium. — The skull. Crassamentum. — The clot, or red globules, of the blood. Cremaster. — A muscle of the testicle. Crest of the Ileum. — The anterior, superior parts of the pelvis. Cricoid. — Ring-like. Crista. — A crest. Crucial. — In the form of a cross. Crural. — Belonging to the thigh. Crystalloid. — Resembling a crystal. Cuboides. — One of the bones of the knee, which resem- bles a cube, or die. Cuneiforme. — A bone of the knee, in form resembling a wedge. Cuspidata. — The tushes of the horse are thus named . Cutaneous. — Belonging to the skin. Cuticle. — The scarf skin. Cyst. — A bladder or sac. D. Dartos. — A name given to the muscle which corragates the scrotum. Deferens. — The excretory canal of the testes. Dentatus. — A tooth-liko process on the second cervical vertebra. Denies Incisors. — The twelve front teeth of the horse. Denies Molares. — The twenty-four grinders. Depressor. — A muscle is so named when it depresses the part on which it acts. Diaphragm. — The muscle which separates the thorax from the abdomen. Diastole. — Periodic dilation of the heart. Dilator. — A name given to muscles which dilate certain parts. Diploe. — The cellular structure, which separates bony tablets. Diverticulum. — A blind tube, diverging from the course of a larger one. Dorsal. — Pertaining to the back. Ducts. — Orifices of various canals. Ductus. — A canal for conveying fluids. Duplicate. — Doubled. DupKcalure. — Reflection of a membrane upon itself. Dura Mater. — The outermost tunic of the brain. E. Efferent. — Vessels are thus named wliich convey fluids from glands. Elevator. — A muscle is so called when it lifts or elevates the parts to which it is attached. GLOSSARY OF VETERINARY TECHNICALITIES. 189 Encephalon. — The brain. Ensiform. — Sword-like. Epididymis. — An appendage to the testicle. Epigastrium. — Region of the stomach. Epiglottis. — Cartilage at the root of the tongue. Epiphysis. — A union of bones by means of cartilage. Epithelium . — A transparent membrane covering rarious internal parts. Erector. — A name given to certain muscles, which raise or erect the parts. Eroded. — Rough and jagged. Esophagus. — The gullet. Ethmoid. — Sieve-like. Ecito-Motary. — The true spinal nerves. Excretory. — Vessels and ducts are thus named which discharge fluids. Expiration. — The act of expelling air from the lungs. Extensor. — To stretch out; a name given to several muscles and tendons. Extremity. — The enJ. F. Facial. — Belonging to the face. Falciform. — Scythe-shaped. Falx. — The scythe process of the dura mater. Fascia. — The tendinous expansion of muscles. Fascicular. — Fibres arranged in bundles. Fauces. — Posterior part of the mouth. Femoral. — Of, or belonging to, the thigh. Fenestra. — Part of the internal ear. Fibre. — A thread or filament. Fibrous. — Composed of fibres. Fibula. — A small bone attached to the posterior part of the tibia of the horse. Filament. — A minute fibre. Filiform. — Thread-like. Fimbria. — A fringe. Fissure. — A crack or groove. Flavus. — Yellow. Flexor. — A name given to numerous muscles and ten- dons which bend the limbs. Foliatus. — Leaf-form. Follicle. — A minute sac or bag. Foramen. — An opening. Fornex. — Arch or vault ; one of the structures of the brain. Fossa. — A shallow cavity or depression. Frcenum. — A ligament which restrains motion. Frontal. — Belonging to the anterior part of the cranium. Function. — Any action by which vital phenomena are produced. Fundus. — The base or bottom. Funis. — The umbilical cord. G. Ganglion. — A knot or enlargement in the course of a nerve. Gastric. — Pertaining to the stomach. Gastric Juice. — A secretion, peculiar to the walls of the siomach. Gastrocnemii. — The tendinous portion of muscles in- serted into the OS calcis, or point of the hock, are thus named . Gemini. — Twins : two organs precisely alike are thus named. Gestation. — Pregnancy. Gland. — An organ of secretion. Glandula. — A small gland. Glandular. — Resembling a gland. Glenoid. — The name of articulating cavities. Glissons Capsule. — The fibrous envelope of the liver. G/o6aosei origin of the optic nerves. Thalamus. — A bed or origin of certain parts. Theca. — A sheath. Thoracic. — Belonging to the thorax or chest. Thoracic Duct. — The ti-unk of the absorbents. Tliorax. — The chest. Thyro. — Names compounded with this word belong to muscles which ai-o attached to the thyroid cartilage. Thyroid. — Resembling a shield. Tibia. — The bone beneath the femur. Tibial. — Belonging to the tibia. Tinea. — The name of a fish ; the tench. Tissue. — An organized structure. Trachea. — The windpipe. Tracheal. — Pertaining to the windpipe. Trachelo. — Names compounded with this word belong to muscles located in the region of the neck. Transversalis. — Having a transverse direction. Transversus. — Placed across. Trapezium.— A four-sided-figure, bone of the horse's knee. Trapezoides. — A bone which in figure somewhat re- sembles the preceding ; it also enters into the composition of the horse's knee. Trapezius. -'Eora square; a muscle placed over the region of the withers. Triangularis. — Triangular. Triceps. — Three-headed. Tricuspid. — Having three points ; a name applied to a valve in the right rentricle. Tri^rf. — Three-cleft. Trigastnc. — Having three bellies. Trisplanchic Nerve. — The great sympathetic or gan- glionic nerve. TrocAan^er. —Eminences or tuberosities on the bones. Tuba. — A tube. Tuber. — A solid roundish substance. Tuberosity. — Protuberance or projection. Tubular. — Tube-like. Tunic. — A membranous covering. Turbinated.— Shajped like a sugar-loaf. Turgid. — Swollen. U. Ulna. — Bone of the fore extremity, termed point of the elbow. Ulnar. — Pertaining to the ulna. Umbilicus. — The navel. Uncifiirm. — Shaped like a hook. Ureter. — A tubular connection between the kidneys and bladder. Urinal. — Pertaining to the urine. Uterine. — Relating to the womb. Uterus. — The womb. Uvula. — A pendulous body, posterior to the soft palate. GLOSSARY OP VETERINARY TECHNICALITIES. 193 Vagina. — A sheath; the cavity between the pudenda and womb. Vaginal. — Pertaining to the vagina. Valvular. — Valve-like. Vas. — A vessel. Vas Deferens. — Excretory duct of the testicle. Vasa. — The plural of vas ; vessels. Vascular. — Highly organized with blood-vessels. Vascular System. — The heart and its vessels. Vastus. — Relates to size ; large, thick and fleshy mus- cles of the thigh. Vena. — A vein. Vena Cava. — The great vein. Vena Porta. — The largest vein of the liver. Venter. — The belly. Ventricles. — A term applied to the cavities of the brain and heart. Vermiform. — Shaped like a worm. Verlebrce. — Bones of the spinal column; Vesical. — Foi-med like a bladder ; pertaining to the bladder. Vesicles Graafian. — Small bladders or cysts found in the ovaria (female testes). Via. — Way or passage. Villous. — Velvet-like, applied to the villous coat of a horse's stomach. Viscera. — Internal organs. Visceral. — Eclating to a viscus. Viscus. — An organ within the body. Vital. — Life-like. Vitreous. — Glassy ; transparent. Vivisection. — Surgical operations on living subjects. Vivus. — Living; life-like. Vulva. — The pudendum. Z. Zoology. — The science of animals. Zootherapeutics. — Relates to the curative action of med- icines. Zootomy. — Comparative anatomy. Zygoma. — An arch or yoke. Zygomatic. — Belonging to the zygoma. EXPLANATION OF FIGURE XIX. OSSEOUS STRUCTURE. 1. Frontalis. ^ Parietalis. 3. Occipital. 5.' Nasal. 6. ^ Lachmyral. 8. Superior maxillaris. 5. Anterior " 10. Inferior or lower jaw. 11. Cervical vertebrae. 16. True ribs. 17. False ribs. 18. Sternum. 19; Ileum. 22. Femur. 23'. Patella. 24. Tibia- 25. Os calcis. 26. Astragalus. 21, Tarsal bones. 28. Metatarsus magnum. 29. Sessamoids. 30. Os sufEraginis. 31. Os coroiia. 32. Os pecb's. S3. Scapula. 34i Os humeri. 35. Radius. 36. Carpus. 37. Metacarpus magntun. 39. Os suffraginis. 40. Os corona. 41. Os pedis. MUSCULAR STRUCTURE. FOEWAED PARTS. — THE HEAD. a. Orbicularis palpebrarimi. 6. Levator palpebrte. c. Bilalor nans lateralis. d. " " anterior. e. Orbicularis oris. f. Nasalis longus. » g. Levator labii superiorus. r. Buccinator. J. Retractor labii inferiorus. Jc. ^lasscter. m. Attolentes et abduoens aurem. 2. Facial vein. THE KECK. c". Trachelo subscapularis. — StSenus. 6. Rliomboideus longus. EXPLANATION OF FIGURE XIX. CONTINDF.D. f. Splenius. o. Abducens vel depiimens aiirem. r. t. Tendon of the sijlenius and complexus major. V. Sterno maxillaris. X. Subscapido hyoideus. THE SHOULDER, ANTERIOR MUSCLES, AND FORE EXTREMITIES. a. Trapezius. 6'. Teres. e". Pectoralis parvus. f. Antea spinatus. g". Postea spinatus. I", i: Triceps extensor brachii. o". Pectoralis transversalis. >■". Flexor metacarpi internus. s". s". Extensor metacai'pi magnus. t. t. Extensor metacarpi obliquus. u. Tendons perforatus and jjerforans. u. (At the humeral region.) Levator humeri. x". x". Extensor tendons. 8. The hoof. THE ABDOMEN .VND POSTERIOR PARTS. — ABDOMINAL REGION, AND OF HIE COSTA. a". Serratus lumborum. o". Obliquus externus abdomini.s — (beneath the dotted line). D. Serratus magnus. POSTERIOR PARTS. g". Ligameuts of the patella. /t. d. e. Glutei. k. Extensor metatarsi internus. m. Tensor vagina. m". Rectus. o". Vastus externus. u. Gastrocnemius internus. V. V. Flexor pedis. It. Flexors ])erforatus and perforans. x". x". Fleshy belly of the extensor. X. X. Extensor tendons. 8. The hoof. ^dmnarj C0nc0liJ5itaI C^art. (195) A VETERINARY TOXICOLOGICAL CHART, CONTAINING THOSE AGENTS WHICH ARE KNOWN TO CAUSE DEATH IN THE HORSE; WITH THE SYMPTOMS, ANTIDOTES, ACTION ON THE TISSUES, AND TESTS. BY W. J. T. MORTON, Lecturer on Veterinary, Materia Medico, etc. "Poisons are substances which are capable of altering or destroying, in a majority of cases, some or all of the functions necessary to the support of the vital principle." — Fmdere. They are derived both from the organic and inorganic kingdoms ; and their action is either local or remote. Local action is referrable to, 1st, Chemical Decomposi- tion; 2d, Irritation and Inflammation; 3d, Nervous Impression. Remote action is effected by, 1st, Absorption ; 2d, Sympathy. Animal Poisons rank first in potency; next to these, the Mineral ; and lastly, the Vegetable. Aerial poisons are, perhaps, the most insidious. The manner in which poisons are introduced into the System varies. The Alimen- tary Tube, the Skin, the Circulation, and the Lungs, are the media. 1st, They may be taken into the Stomach inadvertently with the food, or they may be maliciously or acci- dentally administered. They may also be thrown up as Enemata. 2d, They may be placed underneath the Skin ; or injected into the Circulation ; or they may be absorbed from Wounds. 3d, K gaseous, they may be inhaled, and enter the blood during its transit through the Lungs. They are generally arranged according to the effects which they produce upon the Animal Economy. The great end of Toxicological Science is to counteract their influence, which may be accomplished by chemically decomposing them, by their expulsion from the System, and by restoring the Function of the Organ of which they have caused derangement. As comparatively large quantities of the Poisons are required to destroy Life in the Horse, the niceties of chemical manipulation in the application of Tests are uncalled for. It wUl generally be sufficient to collect some of the contents of the Stomach and Litestines, add distilled Water to them, filter and to the Solution apply the Test or Re-agent. Sometimes they require the influence of heat ; and, when the contents are not attainable, portions of the Alimentary Tube which have been most acted upon by the Agent are to be boiled in distilled Water, and similarly treated. • L — IRRITANT POISONS. These produce their action upon some part of the Alimentary Canal, particularly the Stom- ach and Intestines; and by absorption they are often carried to other Organs. The principal Symptoms are those of Irritation and Inflammation. AGENTS. ACIDUM SULPHURICUM. Sulphuric Acid. ACIDUM NITRICUM. Nitric Acid. ACIDUM HYDROCHLORICUM. Hydrochloric Acid. Symptoms. — The liquid mineral acids axe the most powerful of aU local irritants. Indications of their action are uneasiness, frequent pawing and shifting of the posi- I tion, increased secretion of saliva, which is sometimes viscid and fetid, the mouth in- flamed, difficulty in swallowing from corro- sion of the lining of the esophagus, acute gastric irritation extending to the intestines, and giving rise to symptoms resembling a (1971 198 TOXICOLOGICAL CHAET. most violent attack of colic ; pain on pres- sure being applied over the abdomen ; fre- quent attempts to dung and stale ; and, after the fsBces have been voided, a discharge of mucus streaked with blood takes place : tenesmus, pulse quick and feeble, prostra- tion of strength, profuse perspiration, cold- ness of the body, and death, after the ani- mal has endured excruciating agonies. In one case related to me, nitric acid was poured into the ear, and death took place from inflammation extending to the mem- branes of the brain. Treatment. — As the general symptoms of poisoning by the liquid mineral acids do not materially differ, neither wiU the general treatment. This will consist, 1st, In dilut- ing the agent by throwing into the stom- ach large quantities of water by means of Read's pump. 2d, In neutralizing it, by suspending in the water chalk, magnesia, or soap ; or, in the absence of these, the plaster from the walls. 3d, In allaying the su- pervening inflammation by means of blood- • letting, should the urgency of the symp- toms demand it ; and also by the adminis- tration of opium, and a free use of demul- cents. The subsequent nervous debility and prostration of strength are to be com- batted by the milder vegetable tonics, and a gradual return to liberal diet. Morbid Appearances. — The mouth, pha- rynx, and esophagvTS, present traces of the action of the peculiar acid. The stomach is distended with gas, and occasionally lined with its disorganized tissue, which is eroded in patches, and so deeply ulcerated as to form perforations. Intense inflam- mation often exists in this viscus, which extends throughout the whole of the intes- tinal tube, involving its peritoneal tunic; this last circumstance has been thought to be distinctive between poisoning by acids and metallic compounds ; this cannot, how- ever, be relied upon. The blood in the larger vessels sometimes forms a firm clot. These appearances will not be so marked when an acid has been given in small doses for some time, or if much diluted we may then expect to find the coats of the stom- ach and intestines thickened and contracted, the result of chronic inflammation, with here and there eroded spots, but not of any depth. Tests. — General. — Sour taste — neu- tralization by the alkalies — effervescing with the carbonates — reddening of litmus paper. Particular. — Sulphuric Acid. — The parts with which it comes in contact are first whitened, and then changed to a brownish color. By macerating them or the con- tents of the stomach in distilled water, fil- tering, and adding a solution of the nitrate of barytes, an insoluble precipitate, the sul- phate of barytes, is obtained. Nitric Acid. — The tissues changed of a yellow color, which is heightened by am- monia. The filtered solution boiled on copper filings in a test tube emits orange- colored fumes of nitrous acid. Potassa be- ing added to it, by evaporation a salt is ob- tained, which deflagrates ; or a piece of bibulous paper may be satmrated with the solution, dried, and inflamed. Hydrochloric Acid. — Tissues blanched. Its fumes are rendered more manifest by a rod dipped in ammonia being held in them. This test, however, we are rarely able to avail ourselves of. On the addition of nitrate of silver to the solution, it gives a white precipitate, the chloride of silver. AGENT. ACIDUM OXALICUM. Oxalic Acid. Symptoms. — Instances are recorded of horses having been poisoned by this acid, but whether maliciously given, or adminis- tered by mistake for the sulphate of mag- nesia, I cannot say. The symptoms atten- dant on its action, when a concentrated solution is given, will not be dissimilar to those produced by the mineral acids. When dUuted, however, it is said to cause death by palsying the heart and nervous system, or by inducing tetanus or narcotism ; but I TOXICOLOGICAL CHART. 199 am not aware that such action has been ob- served in the horse. Treatment. — Avoid large quantities of water, as it favors the absorption of the acid. Throw into the stomach a mixture of chalk, or of magnesia and water, partic- ularly the former ; or lime from the walls may be used ; either of wliich will form an insoluble salt. The alkalies are inadmissi- ble, because they form soluble salts. De- mulcents to be freely employed, and the remaining irritation to be allayed by opium- Morbid Appearances. — None recorded in the horse. In other animals the stom- ach has been found to contain black extra- vasated blood, its inner coat being of a cherry-red color; in some places the siu:- face is brittle, and the subjacent stratum gelatinized. The intestines are usually in- flamed throughout. When its influence has been through the medium of the blood on remote parts, the heart has been found to have lost its contractility, and to contain arterial blood. Tests. — Acid reaction on litmus paper. A concentrated solution with ammonia forms a salt whose crystals radiate, the ox- alate of ammonia. Hydrochlorate of Lime throws dowia a white precipitate which is soluble in nitric acid, the oxalate of lime. Sulphate of Copper yields a blue or green- ish-white precipitate, the oxalate of copper. Nitrate of Silver causes a dense white precipitate ; also an oxalate which, when dried and heated, fulminates. AGENT. ACIDUM ARSENIOSUM. Arsenious Acid, White Arsenic. Symptoms. — Intense pain, resembling acute enteritis; belly tympanitic, with a rumbling noise in the intestines ; the dejec- tions oflensive, and mixed with mucus ; pulse quick and feeble, becoming scarcely perceptible at the jaw ; respiration labori- ous; surface of the body covered with an extremely cold, clammy sweat ; extremities cold; efforts to vomit; countenance anx- ious, and indicative of great torture ; mu- cous tissues injected ; mouth hot ; increased secretion of saliva, which is singularly fetid ; delirium from pain which has become con- tinuous; exhaustion; death. The action of tliis poison is not merely as a local irri- tant, it being often conveyed to remote parts through the medium of the circula- tion, thus causing death. Even as an external applicant it has been known to pro- duce much general derangement of the sys- tem, independent of its influence as an escharotic, which is powerful. On this account, when the methods usually resorted to have failed to demonstrate its existence in the contents of the stomach and intes- tines, Orfila has succeeded in detecting it in the organic tissues, particularly the liver. Treatment. — A free use of diluents, or of lime water; avoid blood-letting, as this promotes the absorption of the poison ; give large doses of the hydrated peroxide of iron precipitated by ammonia from a solution of the sulphate of iron, so as to form an insoluble arsenic of iron, which may be expelled by the action of active piu-gatives. The subsequent inflammation is to be combatted by the ordinary antiphlo- gistic remedies ; while the debility which supervenes, and wliich is often great, is best counteracted by the vegetable tonics and judicious dieting. Morbid Appearances. — The stomach and intestines, especially the latter, highly in- flamed and ulcerated in patches. The caecum and colon present the most marked action, the villous coat being black from an effusion of altered blood, and the peritoneal tunic involved. Congestion of blood in the lungs, liver, and kidneys; redness of the lining membrane of the windpipe, extend- ing to the air-passages generally ; conjunc- tival membrane highly injected, and the blood in a fluid state throughout the body. Ecchymosis in the heart. Tests. — Ist, by Reduction. — The sus- pected powder, being dried, is to be mixed with twice its weight of newly-burnt and pulverized charcoal, and introduced into a 200 TOXICOLOGICAL CHART. test-tube : the heat of a spirit lamp is now to be applied ; first to the upper part of the mixture, and afterguards steadUy to the bot- tom of the tube, when, if arsenious acid is present, the metal arsenicum will be sub limed, and, encoating the tube, form a ring of a polished-steel lustre, the inner surface of which is crystalline. The little watery vapor, which wiU be condensed within the tube before the metallic crust begins to ap- pear, is to be removed by a roll of bibulous paper. 2d, by Liquid Re-agents. — The contents of the stomach, or such parts of that viscus as have been acted upon, being boiled in distilled water, the solution is to be filtered. The ammoniacal sulphate of copper added to this gives an apple-green precipitate, the ar- senite of copper. The ammoniacal nitrate of silver, a lemon-yellow precipitate, changing to a dark brown on exposure to light, the arsenite of silver. Sulphuretted Hydrogen, — generated by the action of dilute suphu- ric acid on suphm-et of iron, in a flask, hav- ing an emerging tube bent at a double right angle, — passed up through the solution for ten or fifteen minutes, gives a sulphur yel- low precipitate, the sulphuret of arsenicum. Water impregnated with this gas affords the like compound. The solution for tliis test must be perfectly neutral. This pre- cipitate may be aftenvards subjected to re- duction. 3d, by Nascent Hydrogen. — This is ef- fected in Marsh's tube. The fluid con- tents of the stomach, or the filtered solution before spoken of, being introduced into it, zinc and sulphuric acid are added, and the arseniuretted hydrogen as it escapes from the jet inflamed, when luater and metallic arsenic will be condensed upon the glass disc held above it. The former will be dis- sipated by the heat, «,nd around the latter a ring of arsenious acid may be seen. In the absence of a Marsh's tube, a common two-ounce wide-mouthed vial, with a cork perforated by a piece of glass tube or evea tobacco-pipe, may with care be made to an- swer all the purpose. AGENT. HYDRARGYRI BICHLORIDUM. Bichloride of Mercury, Corrosive Suhlimate. Symptoms. — The effects which follow the administration of large doses of this salt, resemble those which supervene when the mineral acids liave been given, except that, generally, super-purgation is present, and the fcecal matter is profuse and highly offensive. Its solubility renders it more energetic than arsenious acid, although it is not so frequently had recourse to for poi- soning. The protochloride of mercury, calomel, when incautiously given, has also caused death, by inducing inflammation of the mu- cous lining of the intestines, accompanied with violent purging and tenesmus. Treatment. — The white of eggs suspend- ed in water, the albumen of which renders the bichloride of mercury insoluble ; or large quantities of wheat-flour, or milk. Iron filings have also been advocated, which, reviving the metallic mercury, may be ex- pelled by piu-gatives ; a free use of dilutents. The treatment of the salivation, which sometimes supervenes, consists in exposure to cool air, the exhibition of saline purga- tives, and nourishing diet. Morbid Appearances. — These would closely resemble the effects produced by the above agent, tho mucous lining of the ali- mentary canal being intensely inflamed throughout, its textmre destroyed, and in parts corroded. The disorganized tissue often contains the poison, which it yields by analysis. Tests. — Lime-ivater, which throws down an orange yellow precipitate, the h yd rated binoxide of mercury. Iodide of Potassium, which gives a beau- tiful scarlet compound, the biniodide of mercury. Protochloride of Tin, which first aflbrds a whitish precipitate, the protochloride of mercury ; and, on adding more of the test a grayish-black powder is formed, which TOXICOLOGICAL CHART. 201 consists of minutely di\ided metallic mer- cury. Sulphurelled Hydrogen, which gives a blackish compound, a sulphuret of mercury. To these may be added the test by re- duction, the reducing agent being the pro- tochloride of tin, assisted by heat. Albumen is not now relied on as a re- agent. AGENT. ANTIMONII POTASSIO-TARTRAS. Potassio- Tartrate of Antimony, Emetic Tartar. Symptoms. — Violent gastric iiTitation; nausea ; efforts to vomit ; profuse perspira- tion; catharsis, accompanied with colicky pains and much flatus ; increased secretion of urine ; the heart's action at. first much quickened, and afterwards scarcely joercep- tible ; labored respu-ation ; injection of the mucous tissues ; extreme distress ; death. Treatment. — • The yellow bark, or any other astringent vegetable that contains tannin, to be administered both in powder and decoction ; a free use of dilutents, olea- ginous purges, and opium, should then be had recourse to for the purpose of aUaying the irritation. Morbid Appearances. — The stomach highly inflamed, and eroded patches on the mucous coat, which are of a deeper color than the surrounding parts ; intestines red- dened, encoated -with slimy mucus, and thickened ; lungs gorged with blood ; and general inflammation of the whole system consequent on its absorption. Tests. — Caustic Potass and Lime-ivater, which precipitates the oxide of antimony. The carbonate of potass acts with still greater delicacy. Hydrochloric and Sulphuric Acids also afford the like precipitate. A strong infu- sion of the gaU-nut gives a dirty, yeUo\\ash white precipitate, the gallate of antimony. Sulphuretted Hydrogen throws down an orange-red precipitate, the red sulphuret of antimony, which is so peculiar as to be al- ways distinguished ; and the reduction of tills precipitate by hydrogen at once dissi- pates aU doubt. AGENT. SALTS OF COPPER. Symptoms. — The salts of copper are rarely employed as poisons to the horse. Large doses of the sulphate improperly given have sometimes caused much intes- tiiral irritation, followed by colicky pains and diarrhoea ; and, in one instance, death from gastro-enteritis. Doubtless inordinate quantities would always destroy life, when symptoms similar to those caused by any other erodent would be manifested, it being a local irritant. The same, perhaps, may be said of the impure acetate of copper. Li the neighborhood of works for smelt- ing of copper, horses are frequently attacked with diseases of the joints, indicated by swelling, bursal distension, exostosis, and, ultimately, anchylosis, arising either from the state of the herbage or the impregnation of the air by the vapors disengaged. Treat merit. — Give a solution of the fer- rocyanide of potassium, or of soap. Albu- men is also an antidote, and metallic iron, which latter precipitates the copper. This is to be expelled by oleaginous purgatives. Tepid water rendered slightly alkaline may also be freely given ; and opium, to allay irritation. A free use of demulcents, as gruel, infusion of linseed, etc., is indicated. For the local affection, punctm-e the bur- sal distention, and, after the evacuation of the cyst, apply an elastic bandage, giving gentle compression. Remove the animal to another locality. Morbid Appearances. — Stomach ulcer- ated where the agent has adliered, and gen- eral inflammation of its mucous lining and that of the intestines, with here and there spots of erosion. Li the instance ad- verted to of the sulphate of copper causing death, there was an engorgement of the blood-vessels of the lungs. Copper is with extreme difficulty detected in any of the secretions. It, however, has been found in the organic tissues, particu- 202 TOXICOLOGICAL CHART. larly the liver after incineration, and also in the blood. Tests. — Water of Ammonia affords an azure-colored precipitate, or a violet-colored solution, the ammoniuret of copper. Ferrocyanide of Potassium causes a brown precipitate, the ferrocyanide of cop- per. Sulphuretted Hydrogen throws down a blackish compound, the sidphurct of copper. A piece of polished iron introduced into the solution is soon covered with metallic copper. AGENT. SALTS OF LEAD. Symptoms. — Of these, like the above, comparatively large quantities are required to cause death. Violent spasms, tremors, obstinate constipation of the bowels, fol- lowed by paralysis, partial or complete, are the usual indications. In the neighborhood of lead works ani- mals are often thus affected, when, in addi- tion to these symptoms, there is a depraved appetite present: the stomach after death being found filled to repletion with strange and incongruous substances. Treatment. — Solutions of the sulphate of magnesia or soda, combined with croton or linseed oil ; afterwards allaying the irri- tation by means of opium. The phosphate of soda has also been ex- tolled as an antidote. The ti-eatment for the paralysis wliicli remains consists in judicious dieting and exercise. Morbid Appearances. — The lining mem- brane of the stomach and intestines is sometimes inflamed, sometimes blanched; the caliber of the latter is diminished, and its coats corrugated ; the muscular tissue throughovit the body has lost its power of contractility ; the buccal membrane is pale, and the blood of a brighter color than nat- ural. Tests. — Chromate of Potass throws down a yellow precipitate, the chromate of lead. Iodide of Potassium likewise gives a yel- low precipitate, the iodide of lead. Sidphuretted Hydrogen causes a black precipitate, the sulphitret of lead. A rod of Zinc introduced into it causes a deposition of metallic lead in a crystalline form. The alkaline carbonates and sulphates, although they give white precipitates with lead, have been objected to as tests. AGENT. POTASS^ NITRAS. Nitrate of Potash, A^itre. Symptoms. — Much uneasiness ; cholicky pains ; pulse feeble, quick, and irregular ; respiration accelerated; mouth hot; mu- cous lining of the eyelids and nostrils highly injected ; increased secretion of vuine ; frequent voiding of feeces. If the qiiantity given is very great, the abdominal pains are -more intense ; the breathing more labored; the pulse quicker; inefTectual ef- forts to stale are made ; the extremities are cold; and the prostration of strength is extreme. If not, after manifesting much uneasiness, the faeces are frequently voided ; diuresis supervenes ; and relief is obtained. Treatment. — A free use of demulcents; oleaginous purgatives ; hot rugs to the ab- domen, and over the loins ; opiate enemata ; if necessary, the abstraction of blood ; with hand-rubbing, and bandages to the extremi- ties. Possibly a mustard cataplasm or a sheepskin over the loins will be of service. Such counter irritants as are likely to be carried to the kidneys are to be avoided. Morbid Appearances. — The villous coat of the stomach highly inflamed and studded with dark spots resembling ecchymosis, varying in size and running into patches ; they are easily scraped off, and contain a deposit of serum. The cuticular coat is also inflamed, but not so highly, and its texture is much weakened ; the small intes- tines are pervaded with an inflammatory TOXICOLOGICAL CHART. 203 blush ; the lungs and heart are congested, and the venous blood is of a brighter color than usual. Constriction and inflamma- tion of the neck of the bladder have also been observed. Tests. — From the fluid contents of the alimentary tube, or from the urine, the salt may be obtained by evaporation and crystal- lization. It is known by deflagi-ating when thrown on a piece of ignited charcoal, of which it animates the combustion ; and by yielding nitric acid when distilled with sul- phuric acid. Heat also disengages oxygen /rom it. Chloride of Platinum added to its solu- tion gives a yellow precipitate. AGENT. CANTHARIS. Tlie Blistering Fly. Symptoms. — Much uneasiness ; intesti- nal irritation ; frequent attempts to stale ; strangury ; bloody urine ; accelerated pulse and respiration ; continual pain, with much coflstitutional disturbance. These symp- toms increase in urgency, until death closes the scene. Treatment. — Expulsion of the agent from out of the alimentary tube by brisk purgatives ; oil is objectionable, on account of the solubity of the active principle of the cantharides in it. A free use of dilu- ents ; opium may be given, and oleaginous and demulcent injections thrown into the bladder. Sheepskins over the loins ; hot rags over the abdomen. Morbid Appearances. — The mucous lin- ing of the alimentary canal throughout in a high state of diffused inflammation ; but the lurinary organs are principally affected : the blood-vessels of the kidneys, bladder, and urethra, are much engorged, and the lining membrane of the latter has in some places a sphaceletic appearance. Tests. — Washing of the contents of the stomach and alimentary tube will develop portions of the beautiful green case-wings of the fly, which appear not quickly to un- dergo decomposition ; Orfila having detected them in a body some months after death. AGENTS. BITE OF THE VIPER, STING OF THE HORNET, WASP, ETC. Symptoms. — Subcutaneous inflamma- tion, indicated by swelling and increased heat, with effusion into the cellular tissue, which sometimes goes on to gangrene; pain ; constitutional excitement ; quickened and irregular pulse ; rigors. The bite of a viper has been known to cause cerebral derangement and death, by its influence on the nervous system. Treatment. — Removal of the stings ; counter-irritants, as liniment of ammonia or turpentine, which, if sufficient, are to be followed by scarification, the use of emol- lients, and the general antiphlogistic reme- dies, such as bleeding, fomentations, and laxatives, with opium to allay the general irritation. The virus of the tooth of the viper may be extracted by cupping, or, which is preferable, let the part be excised, and the nitrate of silver freely appfied afterwards. Morbid Appearances. — But few instances of death are recorded. If it takes place, it is probably the result of sympathetic and general excitement of the whole system ; the usual indications, therefore, of increased nervous and vascular action may be ex- pected to be met with. Tests. — None. 204 TOXICOLOGICAL CHART II. — NARCOTIC POISONS. These produce Stupor, Delirium, and other affections of the Brain and Nervous System, foUoioed by Death. AGENT. OPIUM. Opium. Symptoms. — The horse will bear large doses of this ch'Ug ; the quantity necessary to destroy life is consequently great. Sup- posed instances are recorded of its causing death when given in doses of a few drachms in order to check superpurgation ; also when the annual has been debilitated by disease, when symptoms of enteritis have shown themselves, accompanied with a torpitude of the bowels, and much sufTering previous to death. Much larger quantities have, however, often been given with impunity, and frequently with advantage. Treatment. — Expulsion of the agent from out the alimentary tube by means of oleaginous purgatives, eneiiiata, venesec- tion, and a free use of demulcents, with fomentations to the abdomen, and counter- irritants to the extremities. Should a state of stupor prevail, exercise may be given, and cold water dashed over the head. Morbid Appearances. — The mucous lin- ing of the stomach and intestines inflamed, and easily torn asunder, the inflammation being diffused. This has been thought to be a distinctive between the effects induced by mineral and vegetable poisons ; but it cannot be relied upon, as repeated small doses of an erodent will induce the like appearances, only there will be more thick- ening of the tunics. Tests. — Odor, which is characteristic. To the suspected matter add distilled water acidulated with acetic acid ; agitate for a few minutes, filter and evaporate to the con- sistence of syrup ; boil tliis in alcohol, and again filter and evaporate ; dissolve the re- siduum in distilled water, and add to the solution acetate of lead, which leaves mor- phia in solution : this being heated with sulphuretted hydrogen, any remaining lead will be precipitated. On niti-ic acid being added to the morphia obtained by evapora- tion, it dissolves with cfTervescence, and becomes of an orange-red color. Suspended in water and treated with a drop or two of the permuriate of iron, it is also dissolved, and forms a greenish-blue solution. AGENT. TAXUS BACCATA. The Teio. Symptoms. — Effects variable ; large quan- tities have sometimes failed to cause any injm-y, wliile at others comparatively small quantities have destroyed life. It would appear to be very insidious in its influence, as the animal generally drops down dead without indicating any previous suffering. In some instances slight convulsions have preceded death. The partially dried leaves appear to be more energetic than the green leaves, probably from greater quantities be- ing partaken of. Treatment. — Usually no oppoiiiinity is afforded for the employment of remedies. Should it, however, be the case that the leaves of yew are suspected to have been eaten, I am not aware of any method which could be adopted but that of endeavoring to expel them from the system as quicldy as possible, which may be effected by active purgatives. The after-treatment will de- pend upon the symptoms which may pre- sent themselves. Morbid Appearances. — The alimentary tube distended with fsecal matter in a semi- fluid state, and highly fetid gases. The mucous lining inflamed throughout, particularly of the larger intestines, with here and there patches of extravasated blood. TOXICOLOGICAL CHART. 205 In some few cases scarcely a trace of diseased action in the tissues could be found. Tests. — Portions of the vegetable in the stomach and intestines mixed with the in- gesta. The active principle of the poison is imknown, hence the difficulty in the treat- ment. AGENT. ACIDUM HYDROCYANICUM. Hydrocyanic Acid, Prussic Acid. Symptoms. — Its influence is sudden, and that of a powerful sedative to the system, and, when the quantity is not large, evanes- cent: otherwise it is followed by marked cerebral derangement, manifested by giddi- ness and coma; the breathing becomes laborious ; the nostrils expanded ; the pulse quickened and fluttering ; much debility is present, with loss of power : to these suc- ceed tetanic spasms ; the muscles become rigid ; the jaws locked, and the membrana nictitans is forced over the eye, which is prominent, and has a glassy appearance ; profuse perspiration covers the body, accom- panied by violent convulsions and intense suffering. These effects are succeeded by a remission for a time, during which the animal appears to be in a state of partial insensibility ; but the exacerbations return again and again, and then the paroxysms become less and less powerful, until at length, all action disappearing, the animal is left in a state of exhaustion, the vital powers being much depressed. Li what- ever way the agent is introduced into the system, the effects are similar. The most active form of the poison is that of vapor. When the dose is sufficiently large to cause death, it is unaccompanied with suf- fering. Treatment. — Cold affusions over the body ; the inhalation of dilute ammoniacal and chlorine gases, particularly the latter. The coma may be removed by blood- letting; and diflusible stimulants, such as ammonia, may be administered, combined with tonics to rouse the depressed vital powers. Of course this treatment will only be of service when the drug has been too fre- quently given, or administered in too large quantities. Morbid Appearances. — The inner tunic of the stomach and intestines sUghtly in- flamed; 'the vessels of the lungs gorged with blood; the parenchyma natural; the lymphatics containmg red blood ; the heart inflamed, and spots of ecchymosis on its lining membrane ; the vessels of the brain highly injected, particularly those of the medullary portion, in which organ the odor of the acid is easily recognized, as well as throughout the body, and particularly in the halitus from the blood. The eyes are gfis- tening and prominent. Tests. — Render fluid the contents of the stomach, and distil an eighth part over, when the following tests will be available : The odor, which resembles that of bitter al- monds, and impresses the throat and nos- trils with a peculiar acridity. Sulphate of Copper, the solution being rendered alkaline by potass, throws down a green precipitate, which becomes neai-ly white on adding a little hydrochloric acid, the cyanide of copper. Sulphate of the Protoxide of Iron, simi- larly employed, gives a gi-eenish precipitate, which becomes of a deep blue color on the addition of sulphuric acid, the ferrocyanate of the protoxide of iron. Nitrate of Silver throws down a white precipitate, the cyanide of silver, which is soluble in nitric acid only at its boifing temperature, and which, when dried and heated in a tube, emits cyanogen gas, which burns with a rose-colored flame. AGENT. CARBONIC ACID. Symptoms. — Instances are known of horses having been suffocated during fires, arising from the disengagement of this gas, with, perhaps, some of the compounds of 206 TOXICOLOGICAL CHART. hydrogen. Its sources otherwise are abund- ant. In a state of dilution it causes coma ; when pure, spasm of the glottis, and death by asphyxia. Treatment. — Removal to the ak ; cold affusions over the body ; bloodletting ; dif- fusible stimulants. Morbid Appearances. — Engorgement of the vessels of the lungs with black blood. The v-cssels of the brain and of the heart are in a similar state. The bronchi and trachea filled with frothy mucus. Tests. — The tests for carbonic acid gas are simple enough, but here they are inap- plicable. AGENT. SULPHURETTED HYDROGEN. Symptoms. — This gas, given off fi-om cesspools and other places, has been at times the cause of death. It is rapidly ab- sorbed by the blood, and produces coma and tetanic convulsions. Sometimes death takes place from asphyxia. Treatment. — The same as the above ; to which, perhaps, may be added the inha- lation of dilute chlorine. Morbid Appearances. — The muscles have lost their power of contractility. The blood-vessels are gorged with fluid black blood; the bronchial tubes inflamed, with increased secretion of mucus both in them and the ti'achea ; the odor from the body is highly offensive. Tests. — Carbonate of Lead on a piece of card paper, exposed to an atmosphere impregnated with this gas, is turned black by the formation of the sulphur et of lead; but as the body when undergoing decom- position emits the same kind of gas, this test can only be accepted as a corrobora- tive proof. III. — NARCOTICO-ACRID POISONS. Tliese cause Death, either by irritation or narcotism, and sometimes by both combined. Their influence is first local and then remote, impressing the Nervous System. They are principally derived from the Vegetable Kingdom. AGENT. NUX VOMICA ET STRYCHNIA. Vomic Nut and Strychnia. Symptoms. — The vomic nut induces a quickened and kritable pulse, highly la- bored respiration, snortings, tetanic spasms, loss of muscular power, injection of the mucous tissues, extreme thirst, and death from asphyxia ; previous to which there is intense suflering. The action of its alka- loid, strychnia, is more energetic. It is shown by tremors, followed by a quickness of the pulse and labored respiration, ex- treme irritability, loss of power in the ex- tremities, tetanic convulsions increasing in violence, the legs being thrust from the body, the muscles rigid, opisthotonus, pro- fuse perspiration, insensibility, and the pulse and respiration being scarcely percep- tible ; the paroxysm exists for a few min- utes only, and is followed by a remission of the symptoms, leaving the animal much exhausted and extremely irritable. The exacerbations, however, continue until death takes place from suffocation. Treatment. — From the tenacity with which the powder of the nut adheres to the stomach and intestines, it is with difficulty dislodged. Its removal may be attempted by means of active purgatives, or antidotes may be thrown in ; these consist of chlo- rine and of iodme, which form inert com- pounds with the active principle, strychnia ; but, as the action of the alkaloid is on the spinal marrow and the brain, little good can be hoped to be obtained when a dose suf- ficiently large to destroy life has been given, TOXICOLOGICAL CHART. 207 unless active measures be immediately adopted. If the dose be not sufficiently large for this purpose, there will be a suc- cession of paroxysms, leaving behind them much debility, which is to be counteracted by tonics and diffusible stimulants, with, perhaps, counter-irritants along the course of the spine, lest effusion should take place. Morbid Appearances. — Mucous lining of the alimentary tube inflamed, lungs gorged with blood, and the vascular system through- out the body in a state of congestion. The spinal canal much inflamed. Efl'usion of bloody serum into the theca vertebraUs ; motor division of the spinal cord more in- jected than the other, and the nerves taking their origin from it inflamed. The mem- branes of the brain have been found in- flamed, with effusion on the surface of the cerebellum, and a softening of the whole cortical portion of the brain. Rigidity of the muscles of the body. Rapid decompo- sition, accompanied with much fcetor. Tests. — The powder of the nut has a greenish-gray color, an intensely bitter taste, and the odor of liquorice. Being collected, it is to.be boiled in water acidulated wath sulphuric acid, filtered, and the solution neutralized by carbonate of lime and evap- orated to dryness. The dry mass being acted upon by successive portions of alco- hol, these are to be evaporated to the con- sistence of syrup, when the product will be found to have an intensely bitter taste, and it becomes of a deep orange-red color with nitric acid, which color is destroyed by the protochloride of tin. Sometimes it de- posits crystals of strychnia on standing. These tests will also be available for the alkaloid ; to which may be added its spar- ing solubility in water, the alkaline reaction of its alcoholic solution, and its forming neutral and crystallizable salts with acids. AGENT. SEMEN CROTOXI. Croton Seed. Symptoms. — This purgative, when in- cautiously administered, has produced death by inducing violent inflammation of the intestinal canal, followed by superpurga- tion ; the alvine dejections being profuse, watery, and offensive. Treatment. — A free use of demulcents, with astringents, as catechu, opium, and challi. Bloodletting ; opiate enemas. Hot rugs to the abdomen, counter irritants, etc. Morbid Appearances. — Violent inflam- mation of the intestines, particularly the caecum and colon, involving all the tunics, the mucous lining being easily torn. Fae- ces abundant and semi-fluid. Lungs in a state of congestion. Tests. — None definite. A GENT. DIGITALIS PURPUREA. Fox Glove. Symptoms. — Languor, gastric irritation, coldness of the body and extremities, pale- ness of the mucous tissues, cold and clam- my perspiration, quickened and feeble pulse, death. When it accumulates in the system, af- ter having been repeatedly given in compar- atively small doses, it produces loss of appetite, nausea, languor, a quick and irreg- ular pulse, followed by purgation, and the effects then gradually disappear. Treatment. — Expulsion of the agent by means of a solution of aloes, combined with linseed oil. The free use of demul- cents; diffusible stimulants; counter-irri- tants. Morbid Appearances. — Depending upon the condition and previous state of the animal. If much debilitated, inflammation of the mucous lining of the stomach and alimentary tube may be seen to exist. At other times no trace of its influence on any of the tissues can be detected, and it is then supposed to cause death by exhaustion of the nervous energy. Tests. — None definite. AGENT. VERATRUM ALBUM. White Hellebore. Symptoms. — Efforts to vomit, acceler- 208 TOXICOLOGICAL CHART. ated piilse, untranquil respiration, intestinal in-itation, which, if followed by purging, affords relief; if not, these symptoms be- come more m-gent, the Ijody is covered with perspiration, saliva is secreted in increased quantities, the legs become deathly cold, inflammation of the bowels supervenes, and death. Treatment. — A free use of demulcents. Milk has been strongly advocated ; on what grounds beyond that of its being a bland fluid, I am at 'a loss to conjecture. Olea- ginous purgatives ; counter-irritants. Morbid Appearances. — The villous coat of the stomach will be found inflamed; the intestines also in a high state of inflamma- tion, particularly the csecum and colon ; the heart pale and flabby ; and the lungs con- gested. Tests. — None definite. AGENT. NICOTIANA TABACUM. Tobacco. Symptoms. — Nausea, giddiness, coma, feeble and irritable pulse. Sometimes gen- eral excitement of the system, profuse per- spiration, labored respiration, pulse much quickened, partial insensibility. Treatment. — Expulsion of the agent by purgatives ; diffusible stimulants when coma exists : demulcents. Morbid Appearances. — I am not ac- quainted with an instance of death having taken place, although this agent is fre- quently given as a vermifuge in very large quantities. Tests. — None definite. AGENT. JUNIPERUS SABINA. Savin. Symptovis. — This, like the preceding agent, is given as a vermifuge, and some- times incautiously. Gastric irritation is then evinced, the animal refuses food, and is languid ; this is followed by diuresis, and sometimes by purging ; the pulse becomes irregular and full, and the respiration hur- ried. Treatment. — Expulsion of the agent from out the alimentary canal by oleagi- nous purgatives ; demulcents. Morbid Appearances. — Esophagus and stomach inflamed, particularly the villous portion of the latter viscus, on which patches of extravasated blood are seen to exist ; the small intestines contain much mucus, and are slightly inflamed ; lungs congested ; larynx and trachea of a rusty yello\y color ; glands at the root of the tongue much enlarged. Tests. — The -partially digested vegeta- ble matter found in the alimentary tube, which may be distinguished by its odor. Under the head of Narcotico- Acrid Poisons, perhaps, should be placed the Atropa Belladonna, Deadly Nightshade, which, in large doses, induces singultus, a dilatation of the pupils, feeble and initable pulse, and a relaxed state of the bowels. Also many of the umbelliferous order of plants, as Conium Maculatum, Common Hemlock, the influence of which is probably that of a Narcotic ; Cicuta Virosa, Water Hemlock, which, to some animals proves an energetic Poison ; with a few of the natural family of the Ranunculacae, as the Aconitum Napellus, Monkshood, and Helleboris Niger, Black Hellebore, which cause death by irritation, producing gastro-enteritis, followed by delirium; lilcewise Delphinium Staph vsagria, Stavesacre ; Bryonia Alba, Wild-vine or Bryony, and Felis Foemina, Female Fern; of which latter very large quantities are required to efiect any marked change in the animal system ; and, indeed, it may be said of the Vegetable Poisons generally, that the Horse is enabled to resist the influence TOXICOLOGICAL CHART. 209 of comparatively immense doses of them, which in all probabiiitj- arises from the peculiar structure of his stomach. Wheat and Barley have been designated as poisons to this animal ; and occasionally they have proved to be so, by setting up acute gastritis. A very common sequela of poisoning by Wheat is inflammation of the laminae, the result of metastasis ; and of Barley, a depilation of the skin. We are, however, in want of more correct information than at present we possess, before anything definite can be laid down under this head, as both wheat and barley, given in moderate quantities and with judgment, often prove beneficial. I am induced to pass the agents above enumerated thus cursorily over, my object having been to give a condensed and tabular view of such substances as are known to destroy life in the horse when incautiously or maliciously administered, and to elucidate a Thesis on Poisons which I had the honor to read before the Members of the Veteri- nary Medical Association in 1836 : at the same time, I hope that this attempt may prove of some use to the Student of Veterinary Medicine. 27 k M EXPLANATION OF FIGURE XX. THE HEAD. a. Orbicularis palpebrarum. 6. Levator palpebroe. c. Dilator naiis lateralis. d. Dilator nans anterior. e. e. Orbicularis oris, the circular muscle of the mouth ; the letters are rather too low to indicate the muscle- f. Nasalis longus. g. Levator labii superioris. k. Masseter. m. Attolentes et abducens aurem. THE NECK, c". Trachelo subscapularis. — Scalenus. s. Splenius. . r. t. Tendon of the splenius and complexus major. M. Levator humeri. V. Sterno maxillaris. The jugular vein is here shown between the two preceding muscles. THE SHOULDER AND POKE EXTREMITIES. e". Sterno scapulari. — Pectoralis parvus. /" '. Antea spinatus. 290 APPENDIX TO PAET FIRST. lago-ligamentous substance which unites the pubic bones. Ligaments of the ribs. — Every rib is con- nected to two vertebra by four ligaments, viz., two capsular, internal and external lig- aments. Capsular ligament of the head invests and holds it within the vertebral socket. Two articular cavities are found \\dthin it, one with each vertebrae, which have separate synovial linings. Capsular ligament of the tubercle sur- rounds it at its articulation with the trans- verse process of the vertebra. External and internal ligaments consist of strong fibres, which connect the neck of the rib, above and below, to the spine. Intercostal ligaments are broad fibrous bands which run obliquely across the inter- costal spaces, and hold the ribs and their cartilage firmly together. Sternal ligaments. — These several pieces of the breast bone are united to each other by intervening cartilaginous substance ; in addition to which they are connected by ligamentary bands, both inwardly and out- wardly. The fore part of it is surmounted by a broad portion of cartilage, which runs along its under part. ARTICULATIONS OF THE FORE EXTREMITY. Shoulder joint. — The capsular ligament around this joint is strengthened in many places by additional fibres dispersed upon its exterior. It is fixed to the rough margui of the glenoid cavity, and to the neck of the OS humeri. A synovial membrane lines it, which may be followed upon the carti- laginous surfaces of the bones. Externally, this ligamentous capsule is clothed on every side by muscles, and to them is attributed the main strength of the joint. Elbow joint. — The ligaments of it are two lateral and a capsular. Knee joint. — In the knee there are five distinct articulations ; one between the ra- dius and the three small bones of the upper row ; a second between the small bones, above and below ; a thud betAveen those of the lower row and the metacarpal bones; a fourth between the os trapezium and the OS cuneiforme; and a fifth between the os pisiforme and os trapezoides ; they have all separate capsular ligaments and synovial linings. The ligaments of the knee, and the ten- dons passing over it, are gu-t by broad, glistening, ligamentous bands, which retain the latter in their places, and render the joint stronger and more compact. Between these ligaments, fascia, and the extensor tendons, are some small bursse. External lateral ligament runs from a tubercle upon the radius to the head of the external metacarpal bone. Internal lateral ligament consists of two parts, which proceed from a similar tubercle upon the inside, and from the body of the radius. The longer is fixed to the inner head of the metacarpal bone, and the shorter to the fore part, of the metacarpal. Ligamentum annulare passes from the os trapezium to the ossa scaphoides and cunei- forme : it confines the flexor tendons. Fetlock joint. — Capsular ligament is at- tached to the articulatory sm-faces of these bones; and the synovial membrane, after hav- ing lined it, is reflected upon their cartilages : it ig guarded in front by the extensor tendon. Long lateral ligament is fixed to a projec- tion upon the side of the metacarpal bone, and to the os suffraginis. Short lateral ligament runs underneath the latter. These Ligaments prevent motion sideways. The ligaments of the sessamoid bones are seven, viz.: superior suspensory, the long inferior, the short inferior, the two lat- eral, and the two crucial. Pastern joint is formed by the adaptation of the ossa suffraginis and corona. It has a capsular, and two pairs of lateral ligaments. The capsular ligament is inserted into the smooth cartilaginous ends of these bones : it is blended vnih the extensor tendon in front, and behind with the long inferior lig- aments of the sessamoids. The lateral ligaments are inserted on the sides of the os coronse and suffi-aginis, Cojfm joint is made up of three bones : the os corona, pedis, and naviculare. LIGAMENTAET SYSTEM. 291 Capsular ligament envelopes the articula- tory surfaces, and is inserted beyond their limits ; in front it is united with the exten- sor tendon ; behind, it is sh-engthened by the tendo perforans. Li addition to the capsular, there are three pairs of ligaments. Tlie first pair passes from the superior edges of the os pedis to the lateral parts of the OS corona, and are inserted about its middle. The second pair is sti-etched from the ex- tremities of the OS pedis to the os corona, and are fixed below and behind the first. Third pair arise from the sides of the coronal process, and terminate in the car- tilages. The ligaments of the os naviculare are four, viz. : t\vo single, and one pafr. Superior ligament runs fi-om its upper and posterior part to the tendo-perforans. Inferior is a very broad ligament, arising from the whole of the lower edge of the bone, and thence extending to the os pedis, above the long extensor tendon. Lateral ligaments fix the os naviculare, by its two ends, to the sides of the os corona. ARTICULATIONS OF THE HIND EXTREMITY. The thigh joint is formed by the reception of the head of the os femoris into its socket. Capsular ligament is attached aromid the cervix of the os femoris and the margin of the acetabulum ; it is thickly clothed on every side by muscle, which assists to main- tain its position. The acetabulum is surrounded by the circular ligament, whose border turns in- ward to embrace the cartilaginous head of the OS femoris. The notch in this cavity, to its inward side, is crossed by the transverse ligament, which here makes up for the deficiency in the bone. Ligamentum teres consists of a bundle of ligamentous fibres inclosed in a sheath which proceed from a pit in the inner and upper part of the ball to a similar one in the roof of the socket. Another portion of it leaves the cavity under the transverse ligament, and is implanted in the pubes. The synovial membrane lines the socket, and is reflected over these parts. Stifle joint is composed of the os femoris, the tibia, and patella. Ligamenta patella are composed of four strong cords, which descend over the con- dyles of the OS femoris, and are inserted into the tubercle of the tibia. The external one passes upon the outer and anterior part of the external condyle ; the internal, upon the inward part of the internal con- dyle ; and the middle one, between them. They approach each other in their descent. Concealed by the external one is the fourth ligament of the patella ; it runs to the out- ward part of the tibia. The patella, with its articulatory surface of the condyles in front, forms a joint of its own, perfectly distinct from that between the tibia and os femoris. Its capsular ligament is fixed to its sur- rounding border. Internal lateral ligament descends from the internal condyle to the ixmer and upper part of the tibia. External lateral ligament — stronger than the internal — runs from the external con- dyle to the upper end of the fibula. Crucial ligaments, short and strong, and deeply buried within the joint, run from the space within the condyles to the tibia. The synovial membrane, after having lined the capsule, is reflected upon the car- tilages and ligaments included within it. Hock joint has four lateral ligaments, two on each side, called internal and exter- nal. Capsular ligament includes the lower end of the tibia, and the pully-Hlce part of the asti'agulus ; to both of wliich, and the lat- eral ligaments, and to the os calcis, it is firmly attached. The OS calcis forms a joint with the os cuboides, and the ossa cuneiforme are also a joint, and the middle and small bones make joints with the cuboid above, and the metatarsi below ; hence, there are sLx artic- ulations in addition to what we commonly understand by the hock joint, that between the tibia and astragulua. LEMy'08 K 82 6 »^