^i m OIli^ ®. 1. ItU SIthrara 5Jnrtb (Earoltna ^talp CColUge QK55y v.l NORTH CAROLINA STATE UNIVERSITY LIBRARIES "" lllll'llllllliill S02514809 T ^^2 Pearson ^•1 The Hepaticae of th( British Isles This book may be kept out TWO WEEKS ONLY, and is sul)ject to a fine of FIVE CENTS a day thereafter. It is due on the day indicated below: ^G 1 1 1965 iOM -May-54— Form 8 THE HEPATIC^ OF THE BEITISH ISLES VOL. I.— TEXT THE HEPATIC^ OF THE BEITISH ISLES BEING FIGURES AND DESCRIPTIONS OF ALL KNOWN BRITISH SPECIES WILLIAM HENRY PEARSON CORK. MEM. LINN. SOC. N.S.W., K.S. TASM., VIDENSK. SELSK. CHUISTIAXIA VOL. I.— TEXT LONDON LOVELL REEVE & Co.. Limitki OubliaftevB to tlir il?omr, CToloutal anti Intiiaii vi^oUniiniriits C. HENRIETTA .STREET. COVENT GARDEN 1902 72IS9 PREFACE. The object I have had in view whilst preparing this work lias been to make the study of the British Hepaticae somewhat easier for the growin<^ number of students who are taking an interest in this, hitherto, neglected order. Although much has been written on the structure and arrangement of the genera and species, no attempt has been made, since the appearance of Sir Wm. J. Hooker's "British Jungermanniae " in 1816, to delineate the various organs of each particular species, without which it is most difficult to identify those closely related. A very brief introduction is followed by a full description, with figures of each species, with their organs magnified. I have followed the arrangement proposed by the late Dr. Spruce, to whom I am indebted for the description of most of the genera. I have also made free use of the labours of others which I have or may not have duly acknowledged, but all the species have been critically examined and studied by me, and as the work has been great, I trust criticism may be disarmed on this point. About twenty-five years ago I attended a botanical class taught by tlie late Dr. Carrington, and so made the friendship of one of the most accurate botanists of our time, who then was bringing out his " British Hepaticju," of which only four parts were issued. It was then my ambition to assist him, mid with him I had })reparecl a further part, but it never appeared VI PREFACE. owing to difficulties arising as to its publication and the doctor's failing health. After his lamented death I began to arrange my notes and drawings, and with the kindly encouragement of the late Dr. Spruce and the help of many good friends, I have been able to bring to a conclusion a work which has occupied the scanty leisure of a busy commercial life. My thanks are due to Messrs. James Cash, Richard Crawshaw, George Ely, Symers M. Macvicar, and other friends for help during the progress of the work. No one can be more conscious than I am of its many imper- fections, yet I venture to hope that students will find it of value in the study of these interesting plants. Wm. Hy. Pearson. Park Crescent, Victoria Park, Manchester. CONTENTS PRKFACE . CONTEXTS . INTRODUCTION . GLOSSARY OF TERMS USED CONSPECTUS HEPATICARUM JUNGERMANIACE.E JUBULE.E . JUN(iERMANIE.+: . RADULE^ PORELLE^ . PTILIDEiE . TRIGONANTHE^ SCAPANIOIDE^ EPIGONIANTHEiE FOSSOMBRONIE.« METZCERIEiE MARCHANTIACE.fi RICCIACE^. ANTHOCEROTACE^. BIBLIOGRAPHY INDEX PAGE V vii it 3 11(1 l'4i. 41(j 440 4f.(; 4Sn 4:)s r)(»4 HEPATIC^. INTRODUCTION. Habit. — Frequently growing in matted tufts, sometimes straggling amongst mosses, or creeping over rocks, stones, or trees. (With the description of each species particulars are given of their varied habit.) Flagella are the whip-like processes some genera bear ; they are either furnished with rootlets and minute rudimentary leaves or are destitute of them. Size. — The stems of the British Hepaticse vary from about 2 mm. to 5 or 6 inches in length. The following is an explana- tion of the terms used to indicate their size : " Very long," 3 to 6 inches. " Large," about 3 inches. " Largish," about 2 inches. " Medium," about 1 to l^ inch. "Small," J to 1 inch. " Minute," ^ of an inch to 2 mm. to 3 mm. Colour. — Commonly pale or yellowish to dark green, but some varying from silvery white to black, others red or purple. Stem. — The Hepatica) may be grouped in two divisions, the frondose and the foliose ; but in certain genera both occur, which show that this division is more or less artificial ; yet in the study of our British species it is a useful distinction. The frondose group comprise those genera which have a flattened frond without any distinct stem, stem confused with the lamina, or stem with wings or lamina on either side; they are usually prostrate. D. H. HILL LIBRARY North Carolina State College 2 HEPATIC^.. The foliose species have a distinct stem with leaves; these stems are either cylindrical, frontally or laterally compressed, and various in texture, some being almost ligneous, others loosely cellulose. In position they are either erect, depressed, procumbent, or pendant ; they are either simple or slightly branched, pinnate or bipinnate ; their ramification either lateral or postical ; a cross- section of the stem discloses great variety in the size, arrangement, and texture of the cells, which affords good characters in the determination of species. EooTLETS. — Some species are almost destitute of these; in others they are often very abundant, proceeding from the postical side of the stem, from the base of the stipules, or very rarely from the underside of the postical lobe of some species. They are usually short, simple or rarely forked, unicellular fibres, sparingly dispersed or closely clothing the postical side of the stem. In one genus only (Biccia) they are papillose within ; all the others are smooth, semi-transparent or pellucid, sometimes dirty white, claret or purple coloured. Leaves. — The insertion, position, and shape of the leaves are fully described with each species. The above diagram illustrates the terms used with reference to their position on the stem. Erect, making an angle with the stem of about 10°. INTRODUCTION. 3 Erecto-patent, inakiug an angle witli the stem of about 30°. Patent, making an angle with the stem of about 50'. Patent-divergent, making an angle with the stem of about 70°. Divergent or horizontal, making an angle with the stem of about 90'. The cells of the leaves vary in shape and size, having walls clear and well-defined, with angles thickened or not, most fre- quently with smaller cells at their angles, called trigones. The following table of sizes of cells, suggested by the late Dr. Spruce, has been used in the present work : Comparative Sizes of Leaf-cells of Hepatic.b. Large -10 mm. Largish or rather large . . •()5 mm. Moderate or medium size . 'iy^^t mm. Smallish .... 025 mm. Small -02 mm. Very small or rather minute . -017 mm. Minute . . . . -014 mm. Very minute . -0125 to -01 mm. Texture of Leaves. — In the frondose species the leaves vary from a thin lamina, consisting of a single layer of cells, to thick and fleshy, and numerous cells thick ; in the foliose they are always composed of a single layer, in one or two cases a slight thickening at the base has been observed. Unlike many of the true mosses, they are destitute of a midrib. In one species [Diplophi/Il um albicans) there is a pseudo-nerve, consisting of a line of thickened cells. Stipules, or, as they have been called by various authors, amphigastria, and underleaves, are postical leaves usually very small and rudimentary ; in some genera they are large and equalling the leaves and generally of the same texture, but often absent. Inflorescence. — The reproductive organs of the Hepatic^! are of two kinds, the male (antheridia) and the female (pistillidia) ; their position affords valuable characters in the determination oi' species. The leaves enclosing them are called 4 HEPATIC^. Bracts, which are either slightly or widely different from the leaves, but usually agreeing with them in cell structure. Bracteoles are the floral stipules, and are often a larger modification of the same. Perianth. — This is the term used for the inner involucre, which is surrounded by the female bracts. It consists of one or several layers of cells ; in some genera it is absent, or replaced by the Calyptra, which is usually a delicate sac enclosed by the perianth. Pistillidia, or the female organs, are usually situated at the base and enclosed by the perianth and calyptra; they are oblong in shape and vary considerably in number, one only having been fertilised, developing into the perfect fruit, the others fading away. The Capsule is the seed vessel, enclosing the spores and elaters, supported on a long or short, usually very delicate, hyaline pedicel; it is globose or cylindrical, dividing regularly into 4 valves, either to its base or the lower half, or rupturing irregularly. In the Anthoceros it is pod-like, with a rudimentary columella. Spores or seeds vary in colour, shape, size, and markings. Elaters are spiral threads, enveloped in a hyaline almost invisible sac, and mixed with the spores in the capsule. Antheridia, or male organs, are usually oval bodies, with or without a short stipe, imbedded in the fronds or enclosed in modified leaves called bracts. Gemm^ are detachable cellular bodies found in some species, which develop into perfect plants. Distribution. The Hepaticae are generally distributed throughout the British Isles, from the sand fiats and rocks by the sea-shore to the topmost heights of our mountains, and are found in almost every imaginable locality. They are the more abundant INTJiODUCTION. 5 in shatly woods, l)y waterliills and mountain streams, in hogs or on peaty moors, growing on living and dead trees, on stone walls and rocks or mud-covered walls ; some species are peculiar to limestone, others to subalpine and alpine districts. Full par- ticulars are given with each species, and if the student has any doubt as to the determination of any species its habitat will often afibrd a good clue to its correct name. Census of Distribution. — I have followed the admirable arrangement adopted in the "London Catalogue of British Mosses and Hepaticoe," 18S1. "The numbers 1 to IG and the lettered numerals 17a, 17b, 18a, 18b, and 18c, stand for the twenty-one Watsonian provinces, and show in which of those provinces each species is known to grow. The names of these provinces, with the numbers standing for them in the catalogue, and the counties included in each province, are as follows : 1. Feninmlar (Cornwall, Devon, Somerset). 2. Channel (Wilts, Dorset, Wight, Hants, Sussex). 3. Thames (Oxon, Bucks, Berks, Surrey, Middlesex, Herts, Essex, Kent). 4. Ouso (North'ton, Beds, Hunts, Cambridge, Norfolk, Sutlblk). 5. Severn (Monmouth, Glo'ster, Hereford, Worcester, Warwick, Salop, and Stafford). 6. South Welsh (Glamorgan, Carmarthen, Brecon, liadnor, Cardigan, and Pembroke). 7. North Welsh (Montgom., Merioneth, Anglesea, Carnarvon, Denbigh, Flint). 8. Trent (Lincoln, Leicester, Notts, Derby). 9. Mersey (Cheshire, Lancashire, sine Lake Lane). 10. Hiunher (Yorkshire solus). 11. Tj/ne (Durham, Northumberland). 12. Lake (Lake Lane, Isle of Man, Westmorland, Cumberland). 13. West Lowland (Dumfries, Wigton, Kirkcudbright, Lanark, Ayr, Renfrew). 14. Eaxt Lowland (Berwick, Boxburgh, Selkirk, Peebles, Haddington, Edinburgh, Linlithgow). 6 HEPATIC^. 15. East Highlmid (Perth, Stirling, Forfar, Kincardine, Aber- deen, Banff, Elgin, Easterness). 16. West Hiykland (Argyle, Arran, Islay, Mull, Skye, West- erness). 17a. Ufper North Highland (Sutherland, Caithness). 17b. Lower North Highland (Eoss-shire). 18a. Hebridean (Isles). 18b. Orkney (Isles). 18c. Shetland {lA^^). I. (Ireland) C. (Channel Islands). How TO Examine the Hepatic^. To gain an accurate knowledge of the Hepaticse it is necessary to have both a simple and a compound microscope, the simple one with a good-sized table to deploy and dissect the specimen, and the compound one to study the different organs plainly. Place your specimen in a white saucer, in water, and carefuUy disentangle a single plant from the mass with which it is usually associated, making sure that you have the whole plant before you. Ascertain its inflorescence : with the paroicous species it may be necessary to dissect with a sharp scalpel the bracts immediately below the perianth, and (as the late Dr. Spruce once mentioned to me) do not spare your best specimens to make sure about this character, which is of great value in the deter- mination of species. In taking sections of stems and perianths, dissections of the leaves, stipules, and bracts, and examining spores and elaters, the student will find it convenient to place them on a glass slip, covering them with a cover glass, adding a few drops of water, or glycerine and water, to examine them under the compound microscope, using the 2 or 3 inch for studying the shape of the leaves, stipules, and bracts, the \ ox \ inch for the cell structure, spores, and elaters. GLOSSARY OF TERMS USED. Accrescent, increasing in size. AcROCAurous, fruit terminating the stem or axis. AcROGENOUs, from the growth of the stem taking place at the summit. Acuminate, tapering gradually to a point. AcuTATE, slightly pointed. Acute, terminating at once in a point. Adnate, joined together, adhering to the face of anything. Adventitious, developed in an unusual position. -^ruginose, verdigris-green. Aggregate, crowded. Albescens albicans albidus, whitish. Alternate, one after another, but not opposite. Alveolate, studded with cavities, somewhat resembling the cells in a honeycomb. Amentula, the catkins of (^ inflores- cence. Amplexicaul, clasping the stem. Andrcecium, the male inflorescence. Antheridia, the male reproductive organs. Antical, the upper face of a stem, branch or lobe of a leaf. Apical, belonging to the apex or point. Apiculate, having an apiculus. Apiculus, an abrupt very short point continued from the lamina. Applanate, flattened out or horizon- tally expanded. Apprkssed, applied closely to the stem. AppiiOxiMATE, close together. Archegonia. .S'ee Pistillidia. Arcuate, bent like a bow. Areolation, the network formed by the outline of the cells. Aroma, the flavour peculiar to any plant. Aromatic, having a spicy taste or smell. Articulate, jointed. Ascending, directed upwards. Aspekulous, slightly rough,'?with little points. Assurgent, ascending upward. Attenuate, narrowing gradually. Auricles, earlike appendages. AuRicuLATE, having auricles or small lobes at the base. AuTOicODS, male and female inflores- cence on the same plant. Axil, the angle between the axis and any part produced from it. Axillary, in the basal hollow or axil of a leaf or branch. I Basal, at the base or lowest part. IJiDEXTATE, with two teeth. BiFARious, two ranks or in two opposite rows. Bifid, cleft into two, halfway down. Bifurcate, two-forked. HiLAiiiATE. two-lipped. BiLOiiED, two-lobed. Bipartite, divided nearly to base into two parts. 8 HEPATIC^. BiPiNNATE, doubly pinnate. BiPiNNATiFiD, doiibly pinnatifid. BiSEPTATE, with two partitions. BiSERiATE, arranged in two parallel rows. BiSERKATE, serratures again serrated. Bracteole, the floral stipule. Bracts, the leaves enclosing the repro- ductive organs. Byssaceous, divided into fine threads like wool. Caducous, soon falling off. C/ESious, glaucous-green. C.espitose, forming matted tufts. C^spitulose, in very small tufts. Calcarate, spurred. Calcicolous, growing on limestone. Oalyptra, the sac enclosing the young Calyx, the external whorl of floral leaves. Canaliculate, channelled. Canescent, rather hoary. Capillary, hairlike. Capitate, forming a head. Carinate, keeled. Carnose, fleshy. Catenulate, chainlike. Catkin, a spike of unisexual flowers. Caudex, the axis of a plant. Cauline, belonging to the stem. Channelled, hollowed out like a gutter. Chlorophyl, the green colouring matter of leaves. Cilia, hairlike processes. CiLiATE, fringed with cilia. Circinate, bent circularly. Cladocarpous, the fruit terminating a lateral shoot. Clavate, club-shaped. CoCHLEARiFORM, round and concave like a spoon, Complanate, pressed flat together. Complicate, folded. Compressed, flattened laterally, CoNCOLOROUs, of One uniform colour. Conduplicate, folded face to face. Confervoid, formed of fine threads. Connate, joined together (base to base). Connivent, meeting in one point. Constricted, suddenly narrowed. Convolute, rolled up. Cordate, shaped like the figure of a heart on cards; the point of attachment being at the broader end. Cortical, belonging to the bark, Crenate, having convex teeth. Crenulate, minutely crenate. Cucullate, hooded, the apex curved inward like a slipper. Cuneate, wedge-shaped. Cuspidate, tapering gradually into a rigid point. Cuticle, outer skin of leaf. Cyathiform, cup-shaped. Cylindrical, having a cylindrical figure like a pipe. Decumbent, reclining on the ground and rising again at the apex. Decurrent, applied to leaves when the lamina runs down the stem below the point of attachment. Deflexed, bent downward through the whole length. Dehiscence, mode of bursting. Deltoid, triangular. Dentate, toothed, having sharp teeth with concave edges. Denticulate, minutely toothed. Denudate, bared of leaves. Dependant, hanging down. Depressed, flattened horizontally. DiCHOTOMous, forked divisions in pairs. Dioicous, male and female inflorescence on separate plants. GLOSSARY OF TERMS USED. Discoid, like :i Hat plate. Distant, a{)plied to the leaves when far away from each other. DisTicuous, in two opposite rows. DiVAKicATi:, straggling, spreading widely apart. Discrete, separated and distinct. DiVEKGENT, spreading outward from the centre. EcHiXATE, with rigid bristles. Edentate, without teeth. Elateus, spiral threads with the spores. Elongated, when any part or organ is in any way remarkable for its length, in comparison with its breadth. Emakginate, slightly notched at the summit, as if a piece had been cut out. Emersed, protruding upward. Entii:e, not cloven ; quite entire, not toothed at the margin. Epidermis, the cuticular or outer layer of cells. Erkct, when any part or organ stiinds perpendicularly, or very nearly so, to the surface to which its base is attached. Erose, irregularly notched as if gnawed. Explanate, spread out Hat. Exserted, elevated above the surround- ing parts. Falcate, sickle-shaped. Fastigiate, all branches reaching an equal height. Fertile, bearing fruii. F1LIF0R.M, threadlike. Fimbriate, fringed with proc«'sses. Flaccid, Habby. Flagellikorm, like the thong of a whip. Flagellum, a runner, a small branch. Flavkscent, becoming yellow. Flexuose, bending inward and out- ward. FovEOLATE, imj)ressed with small holes or depressions. Free, not adhering or adnate. FiiONDOSE, foliaceous, or in leafy ex- pansions. Fugacious, dis;ippearing quickly. Galeate, helmet shaped. I Gemm.e, budlike bodies wipable of be- coming plants. Gemmiparous, bearing gemmie. Geniculate, where any part is bent abruptly, so as to form a decided angle. Girrous, very convex or tumid. Glabrous, smooth. Glaucescent, faintly glaucous. Glaucous, covered with bluish-white bloom. Globose, nearly spherical. Granular, rough on the surface. IIaiut, general aspect of the plant.' Habitat, situation where the plant grows. Hamate, hamulose, curved like a hook. Heterophyllous, having leaves of more than one form. Horizontal, standing off at right angles. Hyaline, more or less transparent or translucent. Hyi'ogynous, below the female. Immersed, covered by the surrounding parts. Incised, cut irregularly and sharply. i Included, not extending beyond the surrounding organs. Incrassate, thickened by internal do- posits. 10 EEPATICJE. Incubous, base with the lowest angle on the underside of the stem. Incumbent, lying upon. Incurved, gradually bending from without inwards. Indehiscent, without dehiscence, or regular line of suture. Inflexed, bent inward. Inflorescence, the general arrangement or disposition of the flowers in a plant. Infra-axillary, originating below the axil of a leaf. Innovations, new growths, or supple- mentary extensions of the stem. Insertions, mode of attachment. Interruptedly-pinnate, where the pairs of leaflets in a pinnate leaf are alternately larger and smaller. Involucre, a whorl of bracts free or united, seated below or surrounding the flower or flowers. Involute, rolled inward. Irregular, exhibiting a want of sym- metry. JuLACEOUS, smooth, slender, and cylin- dric. Keel, a central postical ridge like the keel of a boat. Lacerate, irregular, cleft as if torn or lacerated. Lacini^:, small shreds. Laciniate, cut or slashed. Lacuna, hollows. L^TE-viRENS, bright green. LiEViGATE, smooth as if polished. Lamella, small plates. Lamina, the expansion of a leaf exclu- sive of nerve. Lanceolate, narrowly elliptic and taper- ing to each end. Lateral, attached to the side (the sides right and left of the axis). Lenticular, compressed like a double convex lens. Leptodermous, thin-coated. Ligneous, woody. LiGULATE, strap-shaped. Linear, a narrow leaf with parallel Lingulate, tongue-shaped. LoBATE, or Lobed, divided into or bear- ing lobes. Lobe, any division of an organ ; or especially a rounded division or projection. Lobulate, divided into small lobes or lobules. Loricate, equally narrow through- out. Lumen, the internal space or cavity of a cell. Lunulate, crescent-shaped. LuRiDUS, dirty brown. Lutescent, pale yellow. Male, a plant or flower which bears antheridia and no pistillidia. Mammillae, hemispherical with a pro- jecting papilla. Marginal, at the edge. Marginatus, having a border of cells different in form or colour. Membranaceous, or Membranous, thin and semi-transparent. Moniliform, like a necklace of beads. Monoicous, male and female inflores- cence separate but on the same plant. Multifid, cleft into many lobes or segments. Muricate, rough, with sharp promi- nences. I Muriculate, minutely muricate. GLOSSARY OF TERMS USED. 11 Naked, Avhere any part is exposed or uncovered by other surrounding parts or organs. Navicular, boat-sliaped. Nerve, the mid-rib of leaf. NiTiDUS, smooth and polished. Nucleate, with a central substance of greater density. Ob — , in comparison inversely, as OuovATE, inversely ovate. Oblong elliptic, obtuse at each end, with the longitudinal diameter 3-4 times the transverse. Obsolete, scarcely apparent. Obtuse, terminating gradually in a rounded end. Obtusiusculus, rather obtuse. OCHREACEOUS, brownish-yellow. Olivaceous, a dusky-green ; of an olive colour. Opaque, when the surface is dull, or not at all shining. Opposite, when similar organs are arranged in pairs over against each other. Orbicular, circular. Oval, elliptic, and about twice as long as broad. Ovate, elliptic with the lower end broader. Pagina, the expanded surface of the leaf. Pallid, pale. Palmate, 5-lobed from a centre. Papillj:, small rounded prominences. Papillose, covered with papilla;. Paraphyses, succulent pointed threads growing with the reproductive organs. Paroicous, ^ and ? in the same in- florescence, ^ naked in the axils of the lower bracts. Pautitk, divided almost to thf very base. Patent, spreading at an angle of 2(j - \'i . Patentissima, spreading at right angles or nearly so. Patulous, spreading at an angle of 40 -DO . Pectinate, comblike. Pedicel, or Peduncle, the fruit stalk. Pedicellate, having pedicels. Peduncle, or Pedicel, the fruit stalk. Pendulous, hanging down. Perianth, the innermost bracts united. Perigonial Bracts, the leaves sur- rounding or enclosing the pistil- lidia. Perigonium, the involucre of male inflorescence. Perigynium, the involucre of female inflorescence. Persistent, remaining a long time. Piliferous, ending in a line weak point or hair. Pinnate, having branches on two ojtjm- site sides. Pinxatifid, divided half-way to the mid-rib in segments in a feathery manner. PiSTiLLiDiA, the male organs. Plicate, plaited. Plu.mose, feathery. Polymorphous, of many forms. Pore, a small aperture. PosTiCAL, the under or rooting face of a stem or branch, or lobe of leaf. Processes, divisions. Procumbent, spreading on the ground. Proliferous, bearing au excessive de- velopment of parts. PuLViNATE, like a cushion. Punctate, with ojwique dots. Pyriform, pear-shaped. 12 HEPATIC J^. Quadrate, square. QuADRiFiD, cut into four segments. Quadripartite, cloven almost to the very base into fours. Radicles, small rooting fibres. Radiculose, with small rootlets. Recurved, curved back. Reflexed, suddenly bent back. Regular, symmetrical. Repand, slightly sinuoua. Reticulate, having the appearance of network. Retuse, round at the end. Revolute, rolled back. Rhizome, a creeping subterranean stem. RosuLATE, arranged like a rosette. Rotund, round. Rotund ATE, roundish. Roundish, approaching a round form, or merely rounded off at the ex- tremities, RuEESCENT, reddish-brown. Rupturing, irregularly bursting. Scabrous, rough with minute warts. ScABRULOUS, slightly scabrous. ScARiosE, dry, thin, and semi-trans- parent. Secund, turned to one side. Segment, one of the subdivisions of any part or organ. Semiamplexicaul, half clasping the stem. Septate, having partitions. Serrate, with sharp straight-edged teeth pointing forward. Serrulate, with small serrations. Sessile, without evident pedicle. Sinuate, having the margin with alter- nate concavities and convexities. Sinus, the re-entering angle or depres- sion between two projections or prominences. Spicate, where the male flowers are disposed in a spike. Spike, an inflorescence of male flowers on a stem or branch. Spinulose, with minute prickles. Sporangium, the sac holding the spores. Spores, seeds. Sporogonium, the capsule. Squamose, scaly. Squarrose, spreading out at right angles. Stellate, radiating like a star. Stipules, a foliaceous appendage on the postical side of stem or branch. Stipitate, attached to a stipe or foot- stalk. Stoloniferous, producing stolons. Stolons, horizontal or descending shoots from the base of stem with minute Stomata, air-pores in the epidermis of leaves. Striate, marked with striee or slight furrows. Sub — , in composition somewhat, as subacute, rather pointed. Subulate, awl-shaped ; Widely-subu- late, narrowly triangular. SuccuBous, base with the lowest angle on the upper side of the stem, SuLCATE, furrowed, with longitudinal channel. Suture, line of junction of two parts. Synoicous, antheridia and pistillidia in one inflorescence. j I Terete, cylindric and tapering. Transverse, in a plane perpendicular to the axis, or longitudinal direc- tion. Tridentate, having three toothlike divisions. Trifarious, ranged in three rows. Trifid, three cleft midway to the base. GLOSSARY OF TERMS USED. 13 Thioonous, having three faces. Trilobate, three-lobed. Tiiii'AUTiTE, parted nearly to tlie base in three divisions. Triquetuous, triangular. Tristichous, disposed in three rows. Truncate, cut off abruptly. TuBERCULATE, covered with minute knobs. Tubular, hollow and cylindrical. TuuBiNATK, top-shaped. Turgid, slightly swollen. Umbraculiform, umbrella-shaped. Uncinate, hooked, curved back at point. Undulate, with an alternately convex and concave margin. Unequal, the two sides not sym- metrical. Unuuiculate, ending in a point like a claw. Uniseriate, disposed in a single row, Urceolate, pitcher-shaped. Vaguely, without any definite direc- tion. Valves, distinct portions of the capsule which become detached by dehis- cence in a definite manner. Ventricose, unequally swelling on one side, or swelling in the middle. Verrucose, covered with wartlike prominences. Verruculose, when the warts are small and abundant. Vertical, standing quite erect. Vesicular, inflated like bladders. Whitish, not pure white. CONSPECTUS HEPATICARUM [Spruce). Suborder I. JUNaERMANIACEiE. Vegetation foliose, in a very few species frondose. Fruit soli- tary, pedicellate, capsule quadrivalvate, very rarely irregularly rupturing. Elaters mixed with the spores. Androecia spicate, often amentiform ; antheridia enclosed in leafy bracts, in a few frondose species only either scattered or seriate upon the stem and ebractate. Tribe I. JUBULE^. Eamification always lateral, i.e. having distichous, for the most part infra-axillary branches arising from among the lateral leaves only, never from the stipules. Leaves always alternate, incubous, complicato-bilobate, the smaller inferior lobule generally inflated or saccate. Stipules present in nearly all species, radi- culose; in a few species absent, in which case the radicles are in two rows along the stem, in a very few species they are dupli- cate (that is, in two rows). Flowers $ most frequently diandrous, in a few species monandrous. Flowers ? monogynous or 1—4- gynous. Perianth always present, free, more or less frontally compressed, 2-12-angled, constricted at the apex, shortly tubular. Calyptra constantly free both from the perianth and the pistillidia. Capsule on a short pedicel, globose, quadrivalvate for only two- thirds of its length from the apex, the lower third being undivided and forming a neck or apophysis, which is really the dilated apex of the pedicel. Elaters monospiral, truncate, persistent. CONSPECT US HEP. I TK V / A' I'M. 1 5 Tribe 11. JUNGEliMANlEyE. Eamification various, lateral or partly postical (floial branches principally) ; rarely all postical, very rarely antical. Leaves iti- cubous, transverse or succubous, alternate or opposite ; very various in form, subexplanate or concave, entire or very often divided, but very rarely complicato-bilobate. Stipules in some genera constantly present, in others very rare or absent. Flowers ? in some always monandrous, in others oligandrous, in very few polyandrous. Flowers Metzgeria, branches extremely seldom lateral and pinnate, all the floriferous ones (sometimes even those that do not bear flowers) postical, radicles springing from the nerve and the margin, now and then from the whole surface. Reproductive organs of both sexes distichous on the nerve of a shortened branch; pistillidia 2-20 pairs, styleless ; antheridia very fre- quently more numerous, very rarely tetrastichous. Perianth none. Calyptra large, obovate, fleshy, in Aneura very often papillose, in Metzgeria hairy with radicles. Capsule elongate, quadrivalvate to the base, of two series of cells. Apical elaters fixed and interwoven with others that are free, stretched out in four pencils above the open capsules. CONSPECT I ^S HEP A TK 'A R UM. 2 1 Suborder II. MARCHANTIACEyE. Fronds more or less flesliy, prostrate, broadly and indistinctly nerved, venoso-areolate on the upper surface, areolfu as a rule with one pore, with a cavernous stratum below the pores ; dichotomous, sometimes putting out postical f'rondules, villous beneath with very long radicles. Very long hair-like I'asciculated water-carrying ductules fill up the canal beneath the nerve, which afterwards ascend into the capitulum tlirough the little canals in the peduncles. Inflorescence in almost all species dioicous, acrogenous, or epigenous. Androucia either peltate and stipitate or discoid and hall" immersed ; antheridia solitary in the cavities. Female capitula supported on a stout peduncle continuous with the nerve of the frond, from 2 to many-flowered, orbiculate, hemispherical or conical, generally lobate, paleaceous beneath, chambered within the locali looking downwards. Involucre of each loculus one or many-flowered or none. Perianth arcuate, delicate, split into equal segments, rarely absent. Pistillidia from two to twelve set in two rows on a short receptacle. Calyptra thin, often persistent only at the base. Capsule with a short pedicel, globose, of one row of cells, either splitting all round or 4-8-fid from the vertex. Elaters 1-5-spiral, deciduous. Suborder III. EICCIACE^E. Fronds cellulose, fleshy, furnished with a stratum of air- cavities beneath the epidermis, in most instances dichotomous. Fruit valveless, generally immersed in the frond. Involucre most frequently and perianth always absent. Capsule either free or connate and confused with the calyi)tra, globose, bursting irregulcU'ly. Elaters none. Antheridia immersed in the frond. Suborder IV. ANTIIOCEPOTACE/E. Fronds tender, slightly fleshy or thin, becoming flaccid by drying, radiculose beneath but without paleju ; orbicular, lubate at the circumference, nerve very broad, confused with tlie laminx 22 HEPATIC^. Epidermis without pores, sometimes marked with dark gonidia bearing cells. Inflorescence monoicous or dioicous, antical (i.e. epigenous). Antheridia immersed in cavities scattered over the frond, covered, solitary or few, rarely numerous. Involucre shaped like a horn with an oblique scarious mouth. Perianth and calyptra none. Oogonia solitary, naked, nestling in the substance of the nerve. Fruit pedunculate, siliquoid, two-valved, traversed by a columella or dissepiment. Elaters either mono- spiral or containing only a Hexuose (not spiral) fibre. OiiDiK HEPATICiE. Suborder I. JUNGERMANIACE/E. Tribe T. JUBULE^. Genus 1 . FRULLANIA, Raddi. Fndlania, Raddi, Jung. Etr. in Act. Soc. Modena, xviii. p. 20 (1820). Salinatus, Gray and Bennett, Nat, Arr. Br. PI. ii. p. CS? (1H21). Jubula, Duni. Comm. Bot. p. 112 (p.p.) (1H22). Plants usually large, rarely small, depresso-ca}spitose and stratified, often pendulous, somewbat firm, colour rarely green, often reddish brown, sometimes almost black, in a few species rosy, rarely shining. Stems firm, terete, opaque, cells small, pinnaiely branched ; branches exactly axillary, more contiguous to the lobe than to the lobule ; true subiloral innovations wanting. Leaves standing on a very small base, rarely half-embracing the stem, almost exactly transverse complicato-bilobed ; antical lobe larger, obli(|uely ovate or suborbiculate, convex, incubous, often at the antical base semicordate, quite entire, very rarely denticulate near the apex ; lobule cucullate, galeiform or saccate (campanulate or digitiform), rarely evolute ovato-lanceolate, bipartite interior seg- ment, the one nearest the stem small or minute, triangular, subu- late or styliform. Cells usually small, in others minute, in a lew medium, almost subequal, globose-hexagonal ; walls thickened, trigones and angles conspicuous. Stipules constantly present, large, but smaller than the leaves, orbicular, oblong or cuneate, base often cordate or cris])ato-auriculate, rarely subdt>current, apex bifid (in a few species entire). Eootlets where present produced 24 HEPATICjE. from the hilum at the middle base of the stipule, usually short, dark-coloured and stellately spreading. Inflorescence dioicous or monoicous (rarely paroicous). Androecia usually situated on abbreviated branches, globose oblong or cylindrical, bracts crowded 2- (rarely 3-4-) androus. Gynoecia always terminal, acrogynous or cladogynous. Bracts 2-5-pairs, larger than the leaves, lobule larger and evolute, often dentate or laciniate. Perianth almost always projecting, oval or obovate, trigonous, third angle postical or tetragonous, with postical angles two in number, usually smooth, a few species tuberculate, apex constricted, rostellate. Pistillidia 2-4. Calyptra pyriform or obovate, fleshy, below often 6-8 cells thick. Pedicel short, 8 or 9 cells in diameter, 32 cells in circum- ference. Capsule globose, inner layer papillosely spongy, dividing nearly to the base into 4 valves. Elaters 1 rarely 2-4 spiral. Spores moderately large, reddish, tuberculate, verruculose or rough. 1. Frullania Tamarisci (Z.), Dmn. Lichenastrum imbricatum, Tamarisci narhonensis facie, Dill. Hist. muse. p. 499, t. 72, fig. 31 (1741). Jungermania Tamarisci, Linn. Sp. pi. IGOO (1753); Hook. Brit. Jung. n. 6 (1816). Frulkmia Tamarisci, Dum. Recueil Jung. p. 13 (1835). Monoicous, densely or loosely stratified, large, reddish-brown or blackish in colour, rarely dark green. Stems prostrate, imbri- cate, pinnate, branches patent, alternate, unequal in length, beset with still shorter branchlets ; radiculose, rootlets reddish-brown, in dense bunches proceeding from the base of the stipules. Leaves distichous, alternate, closely imbricate, horizontal or patent-diver- gent, unequally bilobed, antical lobe convex, slightly involute, subrotund or ovate-rotund, apex obtuse or acute, postical lobe about 4 times smaller than the antical, much smaller than the stipules, parallel with the stem, cylindrical or obovate, saccate, sometimes evolute, then oblong-ovate, convex on the under surface. Epidermis polished, cells very small to small, roundish-quadrate, walls thick, angles thickened, moniliform cells more or less present which are larger and darker than the others. Stipules FUULLAXIA. 2r. large, about twice as broad as the stem, subquadrate, longer than broad, decurrent, hastate, sagittate or calcarate, apex bidentate, sinus acute, segments acute or obtuse, margin revolute. Flowers y terminal on short branches, proceeding from chief stem or branches. Bracts larger than the leaves, unequally bilobed, antical lobe ovate, lanceolate, acute or acuminate, margin coarsely and irregu- larly serrate, postical lobe smaller lanceolate acuminate, margin irregularly serrate, remarkably revolute. Bracteole oblong, bifid to about the middle, segments lanceolate, subulate, recurved, margin laciniate. Perianth oval or obovate, obtusely tricarinate, rostellate, smooth. Calyptra obovate or pyriform, yellowisii- white, sub-carnose. Pedicel short. Capsule spherical, pale reddish-brown. Spores brownish. Elaters the same. Male plant more slender than the female, amen tula spherical on short lateral branches, perigonial bracts 3 to 4 pairs, turgid, sub-equally bilobed, lobes oval ; perigonial bracteole ovate, bifid, segments and sinus sub-acute, margin plane ; antheridia oval, 2 or 3 in each bract. Dimensions. — Stems 1 to 3 inches long, with leaves 1'25 mm. wide, diameter of stem "2 mm.; leaves, antical lobe ri mm. x '8 mm., postical "3 mm. x -2 mm., antical -8 mm. x "8 mm., postical "2 mm. X.15 mm., antical "G mm. x -5 mm., postical "25 mm. x •1 mm. ; cells '0175 mm., "02 mm. x -015 mm. ; stipules •(> mm. x 35 mm. ; segments -2 mm., 'G mm. x G mm., seg. -Imm., '45 mm. X 3 mm., seg. "2 mm.; bracts, antical lobe 1*25 mm. x "G mm., postical '9 mm. x "25 mm. ; bracteole 1 mm. x 'G mm., seg. '.") mm., ])eiianth '2 mm. x '8 mm. ; perigonial bract "5 mm. x "3 mm., peri- gonial bracteole "4 mm. x "25 mm. ; seg. "lo mm. ; antheridia '175 mm. x 15 mm. Hab. — Spreading in large patches, sometimes of great extent, on trees, stone walls, and rocks, in exposed or shaded places. Common, but often sterile. 1-17. I. Observations. — This, the commonest of the British Fntl/anice, is distinguished from F. ililafata (L.) by its larger size, proportion- ately smaller and more cylindrical postical lobes, margin of 26 HEPATIC^. stipules entire, antical lobes of bracts serrate, and smooth peri- anth. From F. germana Tayl, and F. micropliylla (Grott.). See notes with those species. Description of Plate I. — Fio-. l. Plants natural size. 2. Portion of stem, antical view x 24 (0. and P. Hep. Brit. n. 48). 3. Ditto, postical view x 24 (ditto). 4. Leaf x 24 (ditto). 5, 6. Antical lobes of leaf x 24 (ditto). 7. Postical lobe of leaf X 85 (ditto). 8. Leaf and stipules x 24 (Loch Lomond, Gourlie). 9. Portion of leaf x 290 (C. & P. n. 48). 10-12. Stipules x 24 (ditto). 13. 2nd sub-bracts and sub-bracteole x 24 (Loch Lomond, Gourlie). 14. Sub-bracts x 24 (ditto). 15. Bracts x24 (ditto). 16. Bracteole x 24 (ditto). 17. Perianth x 24 (ditto). 18. Cross-section of perianth x 24 (ditto). 19. Peri- gonial bracts and bracteole x 24 (Croatia, KlingrafT). 2. FruUania microphylla {Gottsche), Pears. Frullania Tamarisci (L.) var, viicrophijlla, Gottsche ex Carrington in Trans. Bot. Soc. Edin. vii. 4.57 (1863). Frullania micfrofhylla (Gottsche) Pears, in " Journ. of Bot." (1894). Exsicc. Gottsche & Rabenh. Hep. Eur. n. 209 k 636 ; Carr. & Pears. Hep. Brit, Fasc. ii. n. 137. Dioicous, shallowly cfespitose, small, reddish-brown to brownish- black. Stems procumbent, dichotomously branched or pinnate, radiculose ; rootlets fasciculate, dirty white, produced from base of stipules. Leaves incubous, closely imbricate, horizontal, unequally bilobed, antical lobe crossing the stem, convex, broadly oval, rotundate, postical lobe about three times smaller, saccate, oval, erect, with apex inclined towards the stem ; epidermis slightly polished ; cells small, 4-, 5-, and 6-angled ; walls thick ; leaves crossed obliquely by a line of slightl}^ larger and darker monili- form cells. Stipules approximate, nearly twice as broad as the stem, oval, ovate or obovate, bifid to about the middle ; sinus sub- acute ; segments obtuse, subacute or acuminate ; margin entire. Bracts about twice as large as the leaves, unequally bilobed; antical lobe oblique, acute ; margin serrate-dentate ; postical about two-thirds smaller, subrhomboid or broadly lanceolate, acute or FRULLANIA. 27 acuminate, irrei^ularly serrate-dentate, sometimes with a laru^e segment on tlie inner mari^in near the base ; bracteolu oval, Ijitid to nearly the middle ; sinus obtuse ; segments lanceolate, acumi- nate ; margin serrate-dentate. Perianth projecting about half beyond the bracts ; antical side carinate ; postical smooth, convex ; epidermis smooth ; cells similar to leaf-cells, only with the dark ones irregularly dispersed ; apex slightly rostellate. Capsule subspherical ; valves with seven trumpet-mouthed elaters attached to each, unispiral. Male stems irregularly bipinnate ; catkins small, on short lateral branches, globose ; perigonial bracts 2-3 pairs, turgid at the base, unequally bilobed ; antical lobe broadly oval ; postical rather smaller, oval ; antheridia with long bearers, subglobose. Dimensions. — Stem ^-f in. long, with leaves 'G mm. wide ; diam. of stem, '075 mm. ; leaves, antical lobe, '35 mm. x •275 mm., postical lobe '175 mm. x "l mm., antical "35 mm. x '25 mm., postical "15 mm. x '1 mm., antical '45 mm. x "3 mm., postical '2 mm. x -15 mm. ; cells "02 mm., '0175 mm., '02 mm. x •0175 mm. ; stipules '2 mm. x ^125 mm ; segments "11, 275 mm. x "175 mm. ; seg., ^15 mm. ; bract, antical lobe "75 mm. x •4mm. ; postical "5 mm. x '25 mm. ; bracteole "4 mm. x "25 mm. ; seg. •225 mm., "275 mm. x "15 mm.; seg. 125 mm.; perianth 1*0 mm. X -5 mm., 1*25 mm. x '55 mm. ; valves of capsule ^3 mm. x •275 mm.; elaters "225 mm. x "02 mm.; diameter of pedicel •3 mm. ; male catkins "4 mm. x "4 mm. ; perigonial bract, antical lobe ^4 mm. x '275 mm. ; postical ^3 mm. x ^175 mm. ; anthe- ridia '125 mm. x 'I mm. Hab. — On smooth face of rocks and on trees in expo.sed or shaded places, mostly near the coast, in shallow patches, closely attached to the rocks or trees. Ixare. 1. The Lizard, W. Cornwall, W. Cfn-noic, July 1883. St. Mary's, Scilly, W. Cuniow, July 1872. 7. Barmouth, ^Merioneth, //'. //. P., June 1878. 10. Ingleton, Yorks, Jf'cbsfer and Pearson, May 1893. South of Ireland, Old Weir Bridge, Killarney, Carriiif/tun, \^i\\. Glengariff, Carriin/ion, 18G1, G. E. Hunt. Obs. — For several years I have considered Fntllania Tamarisci 28 HEPATIC.-E. var. microjjhi/Ila Gotts. to have well-marked specific characters, but whilst the late Drs. Carrington and Spruce did not think fit to raise it to specific rank, I refrained from publishing it as such. Dr. Spruce wrote me some time before he died that he considered it had good claims to specific rank, adding that he had Carrington's and Hunt's specimens from Grlengariff, &c., and Curnow's from Cornwall, and found them essentially different from F. Tamarisci. Herr Stephani, to whom I have submitted descriptions and specimens, writes : '' This is doubtless a good species. The differ- ences between F. fragilifolia Tayl. and it are — F. fragUifolia : F. viicrojjhi/lla : stouter. smaller. postical lobes cucullate. postical lobes longer, saccate. stipules cuneate, segments trun- stipules oblong, segments acu- cate, J- only bilobed. minate, ^ bilobed. bracts $ obtuse, mamillately bracts ? acute, spinosely den- dentate, tate. innermost bracteole large, seg- innermost bracteole small, seg- ments ovate, apex acute, on ments lanceolate, apex subu- one side connate with leaf. late, on both sides free. " There is little difference in the cells, but the dotted line of large brown cells is uninterrupted in F. microphylla, and consists of single scattered cells in F. frag Hi folia y This is certainly the species which it resembles most in size and habit, from which it also differs in the persistent leaves, which have a broader base at their insertion ; the perianth is also smaller and oval. It diff'ers from F. Tamarisci in its much smaller size, leaves more oval, with relatively larger postical lobes, stipules oblong, and not at all calcarate ; from F. dilatata in its smaller size, more oval leaves with dotted moniliform cells, stipules with margin entire, antical lobe of bract serrate, and smooth perianth. Description of Plate II. — Fig 1. Plant natural size. 2. Portion of stem with perianth, antical view x 24. 3. Portion of stem, postical view x 64. 4, 5. Leaves, postical view x 24. FRULLAXIA. 29 ()-8. Ditto, iintii-al view x 04. IK rurtioii of leaf x l".)U. 10, 11. Stipules X G4. \'l. Bract x GL 1 a. Ih'acteole x G4. 13. Cross-section of perianth x L'4. 15. Male catkins x 04. l(i. I Vri gonial bract x 24. 17. Ditto, explanate x 24. 18. Antheridiiun x (\\ (Bannoutli. \V. IF. W) 8. Frullania fragilifolia, Tai/loi-. FritUaiivi fra;/ilifoli(i, Tayl. in Ann. ;in. 172 (IK I;;). Dioicous, creepinr^, shallowly ca^spitose or substratified, small, dark reddish-brown in colour. Stems procumbent, irre(1753); Hook, lirit. Jung. n. T) (1S16). FruUania dilatata, Duni. Kecueil Jung. p. 13 (1835). Dioicous, closely creeping, often forming roundish patches or substratitied, small to largish in size, deep purple colour, often tinged with brown, to olive-green. Stems prostrate, loosely and vaguely pinnate, 8 to 10 cells in diam., cortical cells about 30, walls brown, inner white, equal in size to the cortical ; radiculose, rootlets few, dirty white, })roceeding from the stipules. Leaves imbricate, below smaller and then approximate, incubous, distich- ous, unequally bilobed, antical lobe crossing the stem, horizontal, concave, postical lobe about 3 times smaller, contiguous with the stem, cucullate, saccate, oval-orbicular, sometimes near the bracts widely subulate, plane or recurved ; epidermis dull, not polished : cells smallish, subquadrate or subquadrate-oblong, irregular in outline, lumen brown, walls and trigones hyaline. Stipules much wider than the stem, distant, cuneate or obovate, bitid to |- or .\, sinus acute, segments acute, outer margin slightly notched near their base, margin plane. Stylus small, between the postical lobe and stem, subulate or triangular-subulate. Flowers $ terminal on main stem or branches, no innovant branches immediately below theuj. Bracts larger than the stem leaves, unequally bilobed, antical lobe subrotund, rotundate, margin quite entire, undulate, often rcflexed, postical lobe half the size or less (.bliipioly c 34 HEPATIC^. ovate acute, irregularly dentate, reflexed. Bracteole oblong- cuneate, divided to below the middle, sinus acute, segments lanceolate-acute, outer margin furnished with one or two large teeth ; sub-bract, antical lobe subrotund, postical lobe oblong- acute, with large segment near the base ; sub-bracteole ovate-acute, bifid to near the middle, sinus acute, segments acuminate. Peri- anth projecting two-thirds beyond the bracts, ovate or obcordate, composed of one layer of cells, complanate below, above triangular, antically plane or with a slight elevation, postically carinate, rostellate, beak short, sometimes slightly uncinate ; epidermis tuberculate. Calyptra oblong-obovate, rather fleshy. Capsule spherical, pale yellowish-brown, bursting into 4 oval-acute or obtuse valves. Spores spherical, minutely tuberculate, 2 to 3 times broader than the unispiral, pale golden-brown elaters, also of a darker colour. Male flowers on short lateral branches, catkins roundish or elongate ; perigonial bracts 4-8 pairs, turgid, unequally bilobed, antical lobe oval, postical lobe rather smaller oval-quadrate, perigonial bracteole oval-acute, bifid to about J, sinus acute, seg- ments acute ; antheridia globose. Dimensions. — Stems J to 2 incheslong, with leaves 75 mm. to •1 mm. wide, diam. of stem "05 mm. to "075 mm. ; leaves, antical lobe "9 mm. x '8 mm., postical '4 mm. x -3 mm., antical "6 mm" x •5 mm., postical "3 mm. x "25 mm. ; cells '03 mm. x '015 mm., •03 mm. X '02 mm., "03 mm. x '025 mm. ; stylus '125 mm. x '04 mm. wide at the base ; stipules '225 mm. x -175 mm.; segments •1 mm., -5 mm. x '4 mm., seg. -15 mm., -25 mm. x '17.5 mm., seg. '075 mm.; sub-bract, antical lobe 1-25 mm. x 75 mm., postical '9 mm. x 'd mm. ; sub-bracteole .75 ram. x -5 mm. ; seg- ments -35 mm. ; bract, antical lobe "9 mm. x -5 mm., postical *5 mm. X '35 mm. ; bracteole '1 mm. x '75 mm. ; segments "6 mm. ; perianth 1"4 mm. x 1 mm., I'S mm. x r2 mm. ; capsule '75 mm. X -7 mm. ; valves -6 mm. x '5 mm. ; pedicel '25 mm. diam. ; spores •05 mm. diam. ; elaters "6 mm. x "02 mm. ; perigonial bracts, antical lobe -45 mm. x '3 mm., postical '35 mm. x -25 mm. ; peri- gonial bracteole -25 mm. x -175 mm. ; antheridia '15 mm x -1 mm. JUBULA. 35 Hab. — On trees iiml rocks, often in lar^^^o putelies, closely attached. Common. 1 to Ki. 1. Found on the Continent. Obs. — Distin<(uished Iroui other liritish FrnUania: by its pro- portionately large postical lobes, when fertile by its tuberculate perianth. Description of Plate V. — Fig. I. Plant natural size. 2. Portion of fertile branch, antical view x IG (Dolgelly, W. H. P.). 3. Portion of stem, antical view x 13 (ditto). 4. Leaf, postical view X 31 (Constance, Jack). 5-7. Leaves, postical view x 31 (Dolgelly, W. H. P.). 8. Portion of leaf x 290 (ditto). 9, 10. Styli X 64 (ditto) 11, VI. Stipules x 46 (ditto). 13. Stipules X 31 (Constance, Jack). 14. Bract, postical view x 31 (Dolgelly, W. H. P.). 15. Ditto, explanate x l(j (ditto). 16. Sub-bract x 16 (ditto). 17. Bracteole x 16 (Ditto). 18. Sub-bracteole x 16 (ditto). 19. Perianth X 16 (ditto). 20. Cross-section of perianth X 16. 21. Portion of male stem, postical view x 16 (ditto). 22. Perigonial bracts and bracteole x 1 3 (ditto). Genus 2. JUBULA, Bum. Jnb^il(p, sp. Dum. Comin., p. 112 (lSi'2). Lejeiinife, sp. Corda in Opiz Nat. p. 6'y2 (1S2'.I). Jubuki, Dum. Recueil, p. 12 (1H:>0). Frullani(e, sp. Nees Nat. Eur. Leb. Ill, p. 240 (IS.iH). Plants conspicuous, somewhat large, dark green, prostrate, substratitied, dichotonious, or pinnately branched. Stem 8 cells in diameter, about 32 in circumference. Leaves large, incubous, bipartite ; antical lobe plane, dentato-spinose or ciliate ; lobule minute, saccate, rarely evolute lanceolate acuminate ; cells small or medium, subequal, walls delicate. Antical elobulate leaf axillary at the insertion of every branch, seated partly on the stem and partly on the branch. Stipules half the size of leaf, plane, subrotund, birid, sparsely spinose or quite entire. Inflor- escence monoicous. V innovations two opposite, rarely only one. Bracts 1 pair, une(iually complicate; bracteole bipartite, free. 36 HEPATIC^. Pistillidia delicate, 1 or 2, rarely 3 or 4. Perianth trigonous, obpyramidal, slightly rostellate, smooth. Calyptra lower half broadly tubular, 2 or 3 cells thick, upper half abruptly sub- globose, delicate. Pedicel about twice as long as the perianth, thick. Capsule globose, dividing into 4 valves which are com- posed of two layers. Elaters long. Spores minutely tuberculate. Androecia spicate on branches ; bracts diandrous. 1. Jubula Hutchinsise (Hook.), Bum. Jungermania Uutchinsice, Hooker, Brit. Jung. t. 1 (1816). Jubula Uutchinsice, Dum. Comm. Bot. p. 212 (1822). Frullania ffutchinsice, Nees Nat. Eur. Leb. Ill, p. 240 (1838). Monoicous, in spreading or dense imbricated patches, rarely substratified, medium to large in size, pale to blackish-green in colour. Stems prostrate, laxly and subflabellately branched, sometimes pinnate, sometimes dichotomous, subgeniculate, branches proceeding from below the ? flowers furcately, rarely with uni- lateral innovations, 8x6 cells in diameter, frontally compressed, cortical cells about 30, pale brown, much smaller than the inner which are very large and pellucid ; radiculose, rootlets few, whitish. Leaves incubous, horizontal, subimbricate, antical distichous, bifarious, unequally bilobed, antical lobe plane, broadly obliquely ovate, apiculate-acute or cuspidate, except at the base acutely spinose-dentate, principally at the antical margin, spines 2-12, triangular-acuminate, sometimes longer and ciliaform, lower leaves less, upper leaves more dentate, branch leaves with few, rarely without, antical base dilated but not cordate, covering the stem, sometimes unidentate, postical lobe 4 to 5 cells distant from the stem, subparallel to it, saccate, ovate, 5 or 6 times shorter than the antical lobe or evolute, subulate or lanceolate acuminate ; cells smallish, 5- and 6-angled, near base subelongate, walls thin, no trigones or thickened angles, subpellucid or with numerous chlorophyllose granules which crowd round the cell walls, making them appear thickened ; at the insertion of every branch is present an antical elobulate leaf seated partly on the stem and partly on the branch, broadly subcordate-ovate, acuminate, quite jriU'LA. :n entire or subspinose, no lobule. Stipules suljrotund subplane, acutely decurrent at the base, to about the middle bifid, segments cuspidate-acuminate, connivent, margin with few long spines or entire. Flowers % on terminal branches. Jiracts adnate at their base, longer than the leaves, deeply bilobed, complicate, lobes obliquely lanceolate acuminate spinose or laciniate, postical half the size or smaller. Bracteole oblong, almost bipartite, carinate, lobes unequal, slenderly acuminate spinose. Perianth verruculose, composed of a single layer of cells, about 200 round, near the middle, twice as long as the leaves, projecting about half beyond the bracts, pyriform or narrowly obpyramidal, deeply tricarinate, flattened antically, postically ridged, angles rounded, at the base terete, apex rotundate-truncate, obscurely rostellate. Pistillidia slender, 1 or 2, very rarely 3 or 4. Calyptra below 2 or 3 cells thick, above abruptly subglobose, delicate. Capsule globose, outer la3'er ©f cells dark brown, inner pale yellow, margin of valves when open reflexed, base (apex of pedicel) cruciate, hyaline. Spores brown, elaters pale brown. ]\Iale spikelets on short lateral branches ; perigonial bracts about G pairs, more than twice smaller than the leaves, broadly emarginate-bilobed, turgid below, lobes subacuminate, connivent, lower smaller quite entire, rarely denticulate ; perigonial bracteole ovate-acute, bifid to below the middle, segments subulate, margin entire or unidentate ; antheridia 2, oval. Dimensions. — Stems 1 to 2 inches long, with leaves 2' mm. to 2*5 mm. wide, diam. of stem '1 mm. to 2 mm., leaves, autical lobe r25 mm. x 75 mm., postical •25 mm. x 'IZS mm., antical r5 mm. X -75 mm., postical '15 mm. x -2 mm., antical 1- mm. X -G mm., postical .3 mm. x 2 mm., cells -03 mm., 025 mm., •03 mm. X 02 mm. ; stipules "75 mm. x •G mm. ; segments ^35 mm., G mm. X ^5 mm., seg. "25 mm., "GS mm. x 'G mm., seg. ^25 mm. ; sub-bract, antical lobe 1^25 mm. x 5 mm., postical ^75 mm. x •25 mm. ; l)ract, antical lobe 1*5 mm. x 75 mm., postical 1 • mm. x •35 mm.; bracteole V25 mm. x 75 mm. ; segments '75 mm., perianth rO mm. X '1)5 mm. ; valves of capsule '4 mm. x ^25 mm. ; apex ^^i pedicel 3 mm. diam.; spores •OlS mm.; elaters ^25 mm. x -015 mm. ; 38 HEPATIC^. male spikes 1-25 mm. x "5 inm. ; perigonial bracts, antical lobe •6 mm. X "4 mm., postical lobe '45 mm. x "3 mm. ; perigonial bract- eole '4 mm. x '3 mm., seg. '2 mm. ; antheridia '15 mm. x '125 mm. Hab. — In spreading patches on shaded damp rocks. Rare. 1. Norvah, Cornwall, /. Ralfs. 7. Dolgelly, Merioneth, /. Ralfs, TF. JFilson. Arthog, Merioneth, Dr. Carring ton, John Whitehead^ W. H. P. 10. Near Hebden Bridge, West Yorkshire, James Needham. Ravengill, Pateley Bridge, L. J. Cocks. 12. Isle of Man, G. A. Holt. Lodore, Cumberland, Rev. C. H. Binstead, W. H. P. 16. Dunoon, Gourlie, Lyon. South of Ireland, Miss Hutchins, Dr. Taylor, Dr. Spruce, Dr. Carrinyton, &c. Found in North and South America, Pacific Islands. Obs. — This is a very beautiful and distinct species and not likely to be confounded with any other of the British Hepaticse. By several authorities it has been placed with the Frullanice but Dr. Spruce has kept it distinct, retaining for it Dumortier's genus Juhida; in colour, texture and habit it differs from any of the Frullanice and, as Dr. Spruce points out, "Hep. Am. et An." p. 60, occupies a position intermediate between this genus and LejemiecB, agreeing with the latter in the bracts of the female flower being adnate to subfloral ramuli (or innovations) and other characters. The plant was discovered in the South of Ireland by Miss Hutchins and named after her by Dr. Hooker. Var. inteyrifolia , Nees, is a more slender form with antical lobes less spinose^ sometimes, but rarely, entire, postical lobe e volute ; I do not attach much importance to these characters as plants from different localities vary from type to this form accord- ing to dampness or shade. The description is taken from Dr. Spruce's " Hep. Am. et An." where further valuable notes are given. Description of Plate VI. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 16 (Isle of Man, Gr. A. Holt). 3. Leaf, postical view x 24 (ditto). 4, 5. Leaves, postical view X 24 (Killarney, Dr. Carrington). 6. Portion of leaf x 290 (Isle of Man, G. A. Holt). 7, 8. Stipules x 24 (ditto). 9, 10. LE.IErSKA. '^^ Ditto (Killarncy, Dr. Carnii, p. ;')72 (1H20). Pamhdphinus, Gray ct Bennett, Nat. Arr. Br. PI. 1, p. (!«« (1821). Lejeiinece sect, Lejeimeott/piis Dum. Syll. Jung. p. 32 (18;>1). Plants usually delicate, small, a few very minute, some robust, creeping amongst mosses or other hepatics, stratified or ca3spitose or pendulous, pallid, whitish or yellowish-green, sometimes dark brown. Stems terete, slender, rarely firm, pinnately branched, rarely pluniDeform or dichotomous, all branches contiguous to the outer base of the leaves, i.e. infra-axillary. Leaves oblique or almost longitudinally inserted, alternate, complicato-bilobed, margin entire, serrulate, denticulate, dentate, spinose or ciliate, rarely laciniate, antical lobe larger, broad or narrow, incubous, postical lobe (lobule) usually rhomboidal or ovate, incurved and ventricose rarely plane ; epidermis smooth, papulose muriculate rarely echinate. Cells subequal, small or medium in size, some- times minute, leptodermous ; trigones and angles rarely conspic- uous. Stipules in nearly all s[)ecies present, in a few wanting bifid or rarely entire. Inflorescence dioicous or monoicous (very rarely paroicous). Andra^cia on amentiform branches, rarely on the chief stem ; bracts subequal, diandrous, very rarely triandrous in a few species monandrous. Female flowers monogynous. Sub-floral innovations where present solitary or binate and opposite, in most species adnate to the bracts. Bracts ? 1 pair, rarely several, planilobed. Perianth free from the bracts, oval oblong or pyriform, terete or angular, variously winged, cristate or ciliate at the angles, leptodermous. Calyptra about half the length of the perianth, obovate or pyritbrm, thin. Pedicel cruciate on the section 4 cells across, quasi-articulate whitto, postical view x 64 (Original, herb. Taylor). 4. Leaves, antical view X 85 (Llanberis, E. M. Holmes). 5, G. Ditto, postical view with stipules x G4 (ditto). 7. Portion of leaf x 290 (ditto). 8-10. Stipules X 85 (ditto). 11, 12. Ditto x G4 (Tyn-y-Groes, W. H. P.). 13. Bract X 24 (Borrowdale, Dr. Carrington and AY. H. P.). 14. Bracteole x 24 (ditto). 15. Male catkin x 64 (Llanberis, E. M. Holmes). 3. Lejeunea hamatifolia (JIoo^:), Dinn. Junrjennania hamatifolia, Hook. Brit. Jung. t. r)4 (ISIG). Lejeunea hanuitifoUa, Duni. Comni. p. Ill (isi'i'). Monoicous, densely, intricately but shallowly ca'spitose, small, green in colour. Stems procumbent, very slender but tirm, 44 HEPATICjE. Hexuose, when fertile regularly pinnate, when sterile sparingly branched or almost simple. Leaves approximate, bifarious, alternate, erecto-patent, unequally bilobed, antical lobe convex, ovate or oval-acuminate, apex incurved, margin coarsely and irregularly serrate, postical lobe about half the size of the antical, oval, free margin involute, furnished with a large tooth near the apex, below entire or slightly denticulate (1 or 2 minute teeth). Cells smallish to small, 4-, 5- and 6-angled, walls firm, no thickened angles or trigones. Sti^Dules obtriangular, bifid, sinus broadly sublunulate, segments subulate or triangular, very divergent, 3, rarely 2 cells wide at the base. Female inflorescence on short branches ; bracts smaller than the leaves but somewhat similar, only both the antical and postical lobes more serrate, the postical not being involute. Perianth pyriform, brevi-rostrate, acutely 5 -angled, keels above setose-spinose- winged, mouth when open with few teeth. Androecia terminal on long branches ; perigonial bracts 3 to 4 pairs, erect or erecto-patent, closely imbricate, turgid at the base, antical lobe serrate, postical almost equal in size ; antheridia oval to globose, pale, stipe of equal length, arcuate- curvate ; perigonial bracteoles similar to the stipules, only with segments less divergent. Dimensions. — Stem \ inch long, '05 mm in diameter, with leaves '3 mm. to *4 mm. wide ; leaves, antical lobe '325 mm. x •175 mm., postical '2 mm. x '125 mm., antical '25 mm. x '1 mm., postical '15 mm. x "OTo mm., antical "25 mm. x '125 mm., postical "175 mm. x "1 mm. ; cells "02 mm., "02 mm., x *025 mm., "02 mm. X "03 mm.; stipules '1 mm. x "1 mm.; segments "05 mm., •075 X "1 mm., seg. "05 mm. ; bract, antical lobe "275 mm. x '125 mm., postical '2 mm. x '1 mm.; perianth •G mm. x '4 mm., beak •01 mm. long; perigonial bract, antical lobe '25 mm. x '15 mm., postical '175 mm. x '125 mm. ; perigonial bracteole "1 mm. x '08 mm., seg. "05 mm. ; antheridia '08 mm. x '06 mm. Hab. — On shaded rocks, especially gold-bearing quartz, rarely on trees. Rare. 7. Cwm Bychan, Merionethshire, Rev. Mr. Saltoay. Tyn-y- Groes, Merionethshire, W. H. P., G. A. Holt. 10. Dent, Yorks, G. Stabler. Ingleton, Yorks, G. Webster and IF. H. P. LE.IELWEA. 45 12. Loclure, Cuinlx'i-hind, 6'. SiaUrr, ('. J. Wild, W . II. P. Kuslh- waite, Cumberhiiid, J)r. Carri/if/to/i and //'. //. P. 15. (lien Tilt, Mr. JJi/rnford. I (5. iMuidart, West Inverness, S. M. Mdccicar. I. Killarney, Dr. Tdijlor, Dr. Spruci', //'. Wilson, Dr. Carnni/toN. Brandon, Dr. Moore. Cronia^^lown, Prof. Li/idljcn/. Found on the Continent and in South Africa. Oijs. — Distinguished from all other British Lcjeimcrc bv the long acuminate distantly serrated antical lobe of leaf. 'J'liis hepatic has a most curious distribution ; it is found in lair abundance on the gold-bearing reel's near the celebrated gold mines at Tyn-y-Groes, in North Wales, also on quartz in Borrowdale, at Ingleton in Yorkshire, not on tlie limestone which is so abundant in the last-mentioned place, but on eruptive rocks which arc exposed near the river. In a collection of hepaticse made at Kynsna, a gold-bearing district in South Africa, by Hans Iversen in 1883, specimens of this species were found agreeing in every particular with our native one. Description of Platk IX. — ^Fig. 1. Plants natural size. 2 Portion of stem, antical view x 85 (Africa). ' 3. Portion of male stem, antical view x 85 (Cromaglown, S. O. Lindberg, and the following). 4. Ditto, postical view x 85. 5-7. Leaves x 85. 8-10. Ditto, explanate. 11. Portion of leaf x 290. 12-14. Stipules X 85. 15. Bract, explanate x 85. IG. Perianth x 31 17. Perigonial bracts x 85. 18. Perigonial l)ractcole x 85. 19. Antheridium x 85. 4. Lejeunea serpyllifolia {Dicks.), Lih. Jungenmmia serpyllifolia, Dicks. PI. crypt. Brit. faac. 4 (1801). Lejeunea serpyllifolia, Lib. Ann. gen. sc. phys. 0, p. 374, n. '2, pp. (ISl'd). Dioicous, shallowly or densely csespitose, small, pale or rarely dark green or yellowish colour. Stems laxly creeping, irregularly branched, rarely subpinnate, radiculose, rootlets produced from base of stipules, white, few, short. Leaves incubous, slightly imbricate or approximate, patent-divergent to patent, unequally bilobed, antical lobe convex, obliquely ovato-oval, rotundato- obtuse or obtuse, margin entire, rarely slightly repand, when dry plano-adpressed, postical lobe much smaller than the antical, 46 HEPATIC^. orbicular or ovate, turgid, margin entire, involute; epidermis shining when dry ; cells medium size, 5- and 6-angled, lumen pellucid or chlorophyllose, walls thin, trigones distinct. Stipules distant, larger than the postical lobe of leaf, about twice as wide as the stem, slightly decurrent, oval or subrotund, deeply bifid, sinus broad or narrow, usually obtuse, sometimes acute, segments obtusate or acute. Female flowers on lateral branches. Bracts unequally bilobed, antical lobe obovate, rotundate, or obtuse ; postical lobe much smaller, oval-oblong. Bracteole obovato-oval, bifid to J, sinus and segments obtuse. Perianth projecting about half beyond the bracts, elliptico-clavate, below terete, upper part 5-plicate, angles obtuse, margin smooth, beak short. Calyptra thick, obovate. Capsule ovato-globose. Spores large. Androecia on chief stem or on lateral branches, below the perianth perigonial bracts 2-4 pairs, closely imbricate, unequally bilobed, antical lobe orbicular, postical rather smaller, oval, much swollen ; antheridia 2, oval-globose. Lindberg described two forms as follows : — Var. jplaniuscula. Elongate, about 1 mm. wide, pale or green, pellucid, when dry shining, less branched, intricate and fragile, planiuscula, leaves more or less remote, antical lobe convexulous, slightly decurved, obliquely ovato-oval, rotundato-obtuse, margin entire, postical lobe 5-7 times smaller, stipules subadpressed, 2 to 3 times larger than the postical lobe, convexiuscula, rotundo-oval, sinus more or less broad and obtuse, segments somewhat obtuse, margin entire, perianth elliptico-clavate, below terete, upper part 5-plicate, angles complanate, margin smooth. Var. cavifolia. Smaller and often yellowish-pale or green, very ramose, intricate and fragile, convex, leaves closely imbricate, antical lobe obliquely broadly ovato-oval, very obtuse, apex distinctly narrower but never acute, entire, postical lobe 3-5 times smaller, cells very chlorophyllose and thickened, trigones distinct, stipules subadpressed, as large or larger than the postical lobe, convex, oval-rotund, perianth narrow at the base, oval- pyriform, upper 4th part 5-plicate. Dimensions. — Stems | to f inch long, -085 mm. in diameter, LEJEUNEA. 47 witli leaves 7 mm. to "85 mm. wide; leaves, iinticul lobe, '45 mm. X :) mm., ])Ostical '1 mm. x 'If) mm., antical, "45 mm. x '.\b mm.,postical "1 mm. x I mm. ; cells '03 mm. x 'OW mm., 04 mm. X "0275 mm., -04 mm. x (»;> mm. ; stipules, 25 mm. x "l mm. ; segments, 075 mm., •'! mm. x "15 mm., seij^. "075 mm.; bract, antical lobe 4 mm. x "25 mm., postical, "225 mm. x "125 mm. ; bracteole '4 mm. x 2 mm., segments I mm. ; perianth, "8 mm. X "5 mm. ; perigonial bract, antical lobe, 25 mm. x "2 mm., pos- tical "2 mm. X 15 mm. ; spores, '025 mm. Hab. — On rocks, stones, trees or banks in damp shaded or, more rarely, exposed situations. Moderately common. 1-1 8c. I. Found on the Continent and in North America. Obs. — This, the most generally distributed of British Lejemiece, is abundantly distinct from any of the other British species, with the exception of Lcj. pafcn-t, Lindb., and Lcj. Jlaca (Svv.), which see for distinguishing characters. Description of Plate X. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 31, var. iihnwiscula (Glena, S. 0. Lindberg). 3. Ditto, postical view (ditto). 4. Ditto x 64 var. carifolia (n. 273 G. & R. Hep. Eur.). 5, 0. Leaves x 31 (n. 435 G. k R Hep. Eur.). 7. Portion of leaf x 29U (ditto). 8. Bract X 64 (Ingleton, W. H. P.). 9. Bracteole x 64 (ditto). 10. Perianth x 31 (n. 435 G. k W. Hep. Eur.). 11. Perigonial bract X 64 (ditto). 5. Lejeunea patens, Lindh. Lejexiuea 2)(it€iis, Lindlu-rg Acta Soc, Sci. Fenn. Hep. Hib. x. p. 1^2 (l^?.""). Monoicous, csespitose, small, very pale green or sometimes white. Stems prostrate or sub-erect, irregularly and much branched and interwoven. Leaves more or less closely imbricate, from their base to the height of the postical lobe patent to erect o- ])atent, then acutely patent-divergent, very much decurved and well overlapping the stem, uneipially bilobed, antical lobe remark- ably convex, especially when dry, obliquely ovato-elliptical or very bluntly oval ; postical lobe 2-4 times smaller than the 48 HEPATIC^. antical, oval, very convex, free margin slightly involute or adpressed to the antical lobe. Texture very pellucid, and when dry shining ; cells smallish, 4-, 5-, and 6-angled, lumen but slightly chlorophyllose, walls somewhat thick, trigones present. Stipules smaller than the postical lobes of the leaves, 2 to 3 times as wide as the stem, very convex, subrotund, bifid to nearly the middle, sinus more or less broad and obtuse, segments somewhat obtuse. Female flowers on short lateral branches. Bracts unequally bilobed, antical lobe oval, narrow at the base, apex sub-acute or obtuse, postical lobe 4 times smaller than the antical, oval-lanceo- late ; bracteole oval, bifid to ^, sinus narrow, sub-acute, segments acute. Perianth projecting slightly beyond the bracts, pyriform- clavate, terete below, above 5-plicate, wings prominent, very acute, complanate, upper margin slightly crenulate. Calyptra obovate, large, composed of somewhat large and irregular-shaped cells. Androtcia on short lateral branches ; perigonial bracts few 2-4 pairs, closely imbricate, erect turgid, bifid, antical lobe broadly ovate, postical somewhat similar but rather smaller; antheridia two, oval-globose. Dimensions. — Stem about ^ an inch long, '075 mm. in diameter, with leaves '8 mm. wide ; leaves, antical lobe '5 mm. x •375 mm., postical '2 mm. x '15 mm., antical "45 mm. x "35 mm., postical '225 mm. x '175 mm. ; cells '03 mm. x '025 mm., '025 mm. X *025 mm. ; stipules "19 mm. x '16 mm. ; segments '075 mm., •2 mm. X '185 mm., seg. '08 mm. ; bracts, antical lobe '4 mm x •225 mm., postical '2 mm. x -1 mm. ; bracteole '4 mm. x "225 mm., seg. -11 mm.; perianth -85 mm. x "45 mm.; capsule "275 mm. diam. ; perigonial bracts, antical lobe '3 mm. x '2 mm., postical •275 mm. x -175 mm. ; antheridia "l mm. x "075 mm. Hab. — On rocks, stones, and trees in shaded places. Eare. 7. Arthog, Merionethshire, Dr. Carrington. 10. Saltersgate, Yorks, S. Anderson. 12. Brown Ghyll, G. Stabler and W. H. F. 13. New Gralloway, /. McA?idrew. 16. I. Killarney, W. Wilson. Griena and Tore Cascade, Dr. Carrinr/ton, Prof. lAndherg, G. A. Holt, &c. Conner Hill, Dingle Bay, Prof. Lindberg, and other stations in the South of Ireland. LEJEUNEA. 49 PVnind on the Continent. Norway {Kaalda.s). Uns. — Dr. Spruce wrote me some years ago : " Lindbcrg's Le- jcitnca jxilen-s is an excellent species. There arc fine patches of it in Sir Win. Hooker's herbarium, jj^athered by Wilson at Killarney ; also in Dickson's herbarium ; and it is probably the true type of his L. scrpj/Uifolia. If so we should have to lall back on tlie name cavifulia for what we have hitherto considered nerpyll [folia. I found L. pafcns beiore Lindber^(J3). Lejeunea Moorei, Liridb. Hep. llib. p. tS7 (ls7."i). Monoicous, closely ca'spitose, snuill to medium in size, yelK>wish. rarely very dark green in colour. Stems ])rostrate, sub-simple, vaguely branched or pinnate, but slightly radiculose or rootless. 50 HEPATIC^. Leaves iiicubous, contiguous or subimbricate, to the plica erecto- patent, afterwards patent to patent-divergent, unequally bilobed, antical lobe obliquely oblong or ovato-oblong, rotundate, slightly concave, at the base subdecurrent, complicate, quite entire, plane, postical lobe 3-7 times smaller, subovate or rhomboid-quadrate, saccate, margin indexed, transverse or subascending, rarely obso- lete. Cells smallish, roundish- quadrate, 5- and 6-angled, near base sliglitly elongate, walls and angles thickened, sometimes trigones distinct. Stipules below smaller and distant, above larger and contiguous, about half the length of the antical lobe of the leaf, about 3 times larger than the postical lobe, ovato-oval or orbicular, bifid to the middle, sinus narrow, obtuse or acute, segments subobtuse or acute, sometimes one lobe acute, the other shorter and obtuse, quite entire, rotundate at the base or cordate, rarely decurrent. Female flowers terminal on the branches or ou stem, and subtended by a simple or repeatedly proliferous innova- tion, or sometimes by a pair of opposite ones. Bracts much longer than the leaves, complicato-bilobed, antical lobe oval- lanceolate, obtuse, margin plane, quite entire, postical lobe about half the size or still smaller, elliptical, obtuse. Bracteole oblong, obovate, narrowly lanceolate or cuneato-lanceolate, cloven to about the middle or deeper, sinus very narrow, segments obtuse. Perianth very slightly emersed, green, pyriform, rounded at the apex, beak short, slightly compressed, 5-carinate, keels small, subobtuse. Androecia on short lateral branches or on stem, rotund or oval, perigonial bracts 5-9 pairs, bilobed to the middle, postical lobe rather smaller than the antical, diandrous. Dimensions. — Stems |- to 1 inch long, '1 mm. in diameter, with leaves 1* mm. wide; leaves, antical lobe '55 mm. x "35 mm., postical "15 mm. X '1 mm., antical "5 mm. x '35 mm., postical •125 mm. x "1 mm. ; cells, "025 mm. x '02 mm., '03 mm. x "03 mm., •02 mm. X '02 mm., '03 mm. x "02 mm. ; stipules '3 mm. x '25 mm., segments '15 mm. ; bracts, antical lobe 1* mm. x "5 mm., postical '6 mm. X "25 mm., antical 1* mm. x "6 mm., postical "6 mm. x 3 mm. ; bracteole '8 mm. x '5 mm., seg. '4 mm. Hab. — On rocks and trees in shady situations. Eare. 12 ? LKJKUNEA. 51 South (i'i Ireland, Killanicy, Dr. Cdrr'uiglon. O'Siillivans Cascade, Dr. Moore. Iluiitini,' 'Jowor, Dr. Moore, D. McArdh.-. (jrlciia, Prof. Lhidb('r(j. Kilianioy, 6'. .1. I loll, .]/". B. Slalcr. Soutli America, Rio Janeiro, (llazion, Cuba, Wri(//i(. Obs. — Divstinguished from Lrj. nvrjjijUifolia by the IonL,'er and more narrowly ovate antical lobe, and the smaller postical lolje, smaller cells with thicker walls and anj^les, and lar<^er stipuhs. ''Lcj.xcrpijUi folia is a smaller plant, colour more rarely yellowish, leaves rounder and with a much longer and more turgid postical lobe, sometimes equalling half the antical." (11. Spruce.) Dr. Carrington was the first to publish any notice of this species as native to our Isles, when he described it as var. thymifolia of Lcj. nerpyUi folia in his '* Irish Cryptogams." Proi. Lindberg afterwards ])ublished it as Lcj. Moorci, but later considered it along with Dr. Spruce identical wdtli Lcj. fava (Sw.), which has a very wide geographical distribution. Plants with perfect perianths are extremely rare. Description of Platk XII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 25 (drawn by Tutten West, Moore, " Irish Hepatictu "). 3. Leaf, postical view x (!4 (Croma- glown, S. O. Lindberg). 4. Ditto and stipules x G4 (ditto). 5. Ditto X (54 (Glena,S. 0. Lindberg). 0. Portion of leaf x 21)0 (ditto). 7, 8. Practs x 24 (ditto). 9. Bracteole x 24 (ditto). 7. Lejeunea Holtii, Spruce. Lejeunea Iloltii, Spruce, Journ. of IJutany, Feb. 1887. Monoicous, loosely creeping amongst mosses, largish, yellowish or with a reddish tinge. Stems prostrate, fragile, rarely sub- stratified, subpinnate ; frequently the flower-bearing branclilets, are elegantly pinnulatc. Leaves widely patent, distinctly distichous or bent lightly backwards, dissitous or contiguous, rarely subimbricate, subobli(iue, ovate-oblong or ovate, obtuse, sometimes subacute rarely rotundate, near the base subsinuate- complicate ; lobule very small, less than 1, the size of tlie leaf, subovoid, inflated, with the apex either running gradually into 52 HEPATIC^. the leaf or rather abruptly incurved, prolonged on to the stem, occasionally obsolete, especially where the flowers are wanting; cells medium size, equilatero-hexagonal, except below the middle where they incline to become oblong, rather smooth, walls thin, under ordinary powers (1 inch or f) no trigones are observable, but under the \ inch they are small but distinct ; slightly convex which causes the leaf margin to appear subcrenulate. Stipules delicate, rather less than -J- the size of the leaves, distant, orbicular, subobtusely bifid to the middle, the segments acute or subobtuse. Flowers ? on exceedingly short branchlets, usually with a single pair of leaves, very rarely longer with 4 pairs, usually simple, with no subfloral innovations ; very occasionally there is present a minute microphyllous innovation springing from the base of the flower. Bracts less than ^ the size of the leaves, recurvulo- patent, bilobed to below the middle, lobes slightly complicate lanceolate, acute or obtuse, the postical a little shorter and only half the width ; bracteole lanceolate, narrowly bifid to a J or half of its length, segments acute, connate with the other bract, or free. Perianth distinctly emerging from the bracts for more than double of their length, obpyramidato-pyriforra, depressed at the apex and having a very short slender mucronulate slightly compressed beak ; cells equilatero-hexagonal, pellucid, beautifully reticulated; 5-keeled — the keels having very deep-like wings — occasionally towards the apical margin bordered with a single series of very thin cells. Calyptra half the size, obovate, con- stricted at the base so as to appear shortly stipitate. Capsule globose (not been seen exserted). Androecia very frequently upon short branches ; perigonial bracts 2-5 pairs imbricate, turgidly cymbiform, bilobed, postical lobe rather smaller. Anthe- ridia not known. Dimensions. — Stems f-l|- inch long and -09 mm. in diameter, with leaves Tl mm. wide; leaves, antical lobe '7 mm. x '45 mm., lobule '13 mm., antical lobe "65 mm. x "55 mm., lobule •15 mm. X "15 mm., antical lobe '5 mm. x '.35 mm., lobule '15 mm. X *125 mm. ; cells, near middle, "03 mm., '04 mm., "04 mm. x -03 mm. ; stipules "25 mm. x '25 mm., '3 mm. x '3 mm., LEJEUNEA. •225 mm. x '225 mm.; bract, antical lobe 4 mm. x 1.') hum. to "2 mm., lobule "35 mm. x 'l mm., antical lobe (i inin. X '2 mm., lobule "4 mm. x "1 5 mm. ; bractcole ".'5 mm. x •irjinm., '55 mm. x *275 mm.; periantb '!) mm. x •.").") mm., T mm. X CiS mm. ; pcriL,'onial amentula "85 mm. x '4 mm. Hah. — E.\tremely rare, the only known station bein^^ on shady rocks below Tore W'aterfall, Killarney, near enough to the stream to be moistened by its spray, except when the water is very low. It iij-rows chietly on Tliamninm alopcciirnm, accompanied by Jiadiila Carri/n/tOfii, S(tcco(///na vUiculosa, Mch/jeria conjnyala, and sometimes by Lejciinca diversiloba (6'. A. Hull, June 1885), R. S. Obs. — " Differs from every other British h'jeimea in the female flowers being* borne on exceedingly short branchlets, which normally put forth no subfloral innovation such as con- stantly exists in all our other species. In size it resembles L.Jlava, or luxuriant A. serpi/IJifolia, but usually diff'ers at sight from both by the pale reddish tinge of the foliage. Even where the leaves are of the yellow-green of almost ripe limes, dried speci- mens speedily assume a rufous hue when moistened. Another important and unicpie character is afforded b}' the large pear- shajjcd perianths being so very strongly and sharpl}' keeled that at first sight they seem broadly 5-winged. The female flowers often alternate, or are variously mixed up, with male catkins of about the length of the adjacent leaf, and consi.sting of from two to five pairs of C3'mbiform bracts. Where inflorescences of both sexes are numerous (as they sometimes are), they render the stem or branch elegantly pin nu late. The only species which L. Holt a might l)e conlbunded with is L.Jlara (Sw.), which grows near it and in much greater abundance ; but the former, although nearly equal in size, is iar more delicate and pellucid, and its slight tinge of red is never seen in the yellow or green foliage of L. jlava. The latter, besides the essential difference of the perianths being terminal on branches of various lengths or on the main stem, and invariably putting forth from their base a leafy innovation or even a pair of opposite innovations, each of which may in like manner hear an apical 54 HEPATIC^. flower subtended by a secondary innovation, differs also in the tufted habit, the imbricated leaves (which, although only slightly different in form, have smaller, more chlorophjdlose cells), but, above all, in much smaller obtusely 5-carinate perianths, as com- pared with the large, deeply plicato-carinate perianths of Z. HoltUr Tlie description and observations are taken from Dr. Spruce's communication to the " Journ. of Bot." for Feb. 1887. Descriptfon of Plate XIII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 31. 3. Ditto, postical view X 31. 4. Ditto, X 64. 5. Portion of leaf x 290. 6. Stipule X 64 7. Bract x 31. 8. Bracteole x 31. 9. Perianth x 31. 10. Cross-section of perianth x 31. 11. Perigonial amentula X 31 (Tore Cascade, Gr. A. Holt). 8. Lejeunea ulicina, Tayl. Jungermania ulicina, Tayl. in Trans. Bot. Soc. Edinb. p. 115 (1841). Jungermania minutissima, Hook. Brit. Jung. t. 52, ex parte (nee Smithii) (181 G). Lejeunea ulicina, Tayl. G. L. N. Syn. Hep. p. 387 (18-44-). Lejeunea minutissima, Spruce in Ann. Nat. Hist, et Trans. Bot. Soc. Edinb. (1849) (nee J. minutissima, Sm.) Dioicous, loosely spreading, rarely subcsespitose, often creeping upon mosses, minute, green. Steins straight, not geniculate, vaguely branched, subradiculose. Leaves distant or subcontiguous, erect to erecto-patent (10°-30°), cochleato-concave, rotundo-ovate, at the apex narrower, obtuse or abruptly acute, from the base to J or f (rarely almost to the apex) obtusely complicato-bilobed, postical lobe more or less shorter and narrower, rarely almost equal, turgid at the keel, margin incurved or often plane, apex apiculcito-acute ; cells small, slightly chlorophyllose, subpellucid, smooth or slightly convex, obscurely tuberculose, 4-, 5-, and 6- ant;;ed, walls firm, no trigones or thickened angles. Stipules distant, about 3 times shorter than the leaves, rather broader than the stem, oval, to about ^ or the middle, or rarely below bifid, segments linear-subulate, 2 cells broad below, at the apex 1 cell broad. Female flowers terminal, innovations produced on one or both sides. Bracts large, 2 to 3 times larger than the LEJEUNEA. 65 leaves, subclivergcnt, at an angle of G0°-90°, almost to the middle bifid, complicate, keel narrow, winged, antical lobr somi-ovate acute, postical lobe shorter, rarely ecjualiy as long, semi-lanceolate or obliquely cuneate, margin cellulose-erose, sometimes subden- ticulate. Hracteole a little shorter, erect oblong or oval-lanceolate, to about I acutely bitid, segments acute. Perianth (hardly mature) immersed in the bracts, pyriform-oblong or obovate, apex depressed, terete below, towards the apex obtusely 5-angled, cells slightly convex, papillose or almost smooth. DiMiiNSioNs. — :J— ^ inch long, 02 mm. to "03 mm. in diameter, with leaves "25 mm. to -35 mm. wide, leaves, antical lobe LMnm. x '15 mm., ])ostical 114 mm. xO'Zomm., antical '24 mm x -15 mm., postical -18 mm. x 'l mm. ; cells -025 mm. x '02 mm., "02 mm., 017 mm., '02 mm. x '015 mm. ; stipules '08 mm. x -80 mm., segments 02 mm.; -07 mm. x -05 mm.,seg. 015 mm., *1 mm. x '05 mm. ; lobe oi'bract •4mm. x •2mm.,-5mm. x -Smni-jbracteole 3mm. x -15 mm.; perianth '35-4 mm. x -18 mm. Hab. — On trees, or more rarely creeping uj)on mosses. Eare, extremely so, fertile 2. On beech trees, Southwood, Hants, ]i. S. Mill. 7. Tyn- y-Groes, lllld and lloU. Torrent Walk, Dolgelly, (7. ^lahlrr, W. If. r. 10. Bolton Abbey, Wharfdale, Dr. Spruce. Dent, G. Stabler. Broad wood, Ingleton, G. IFeb.sfer and W. U. P. 12. Levens, Westmorland, G. Stabler. Keswick, C. L>/rlt. 13. Burnfoot Hill, New Galloway, /. Mc Andrew. 15, lO. I. Killarney, Dr. Tatjtor, Dr. Spruce, Dr. Carringtnn, G. J. J/ott and others. Near Lake Brittas, King's Co., B. McArdte. Found on the Continent, France, Mortain, 7\ I/usnot. Obs. — Misled by an original specimen from Sowerby of J//n//- minufis.siwa, Smith, tigured in Eng. Bot., Dr. Spruce was under the im])ression that it was the same as Junr/. idicina of U'aylor, and therefore reduced Jun//. idicina, Tayl. to a synonym of .///////. mifiuti.mnia Sm., and for the fertile Jun//. niinutissinia of Smith coined the name Lejeunea Taylori, Spruce ; man}' years afterwanls he ibund in comparing with the figure in Eng. Bot. of Smith's Jun(/. ininutis.sima that it could never have biMMi drawn frtun that 56 HEPATIC^. specimen, so he reinstated Lejeimea ulicina, Tayl., and made Lej. Taylori a synonym of Lej. minutissima, Sm. Considerable con- fusion as to these two species has consequently arisen, but if we remember that the monoicous form with geniculate stems, no stipules and often found fertile is the Lej. minutissima of Smith, and the dioicous one with somewhat straight stems, stipules present, and flowers extremely rare is Lej. ulicina, Tayl., we shall be right. Shortly before Dr. Spruce died he wrote me : " Lately I came on the specimen given to Mr. Borrer by Sowerby as part of the Jung, minutissima figured in Eng. Bot. You will see it is L. ulicina, and when even Hooker did not discriminate between the two it was impossible Sowerby should." In my plate of Lej. ulicina, fig. 4, is a drawing of this plant ; in Hooker's " Brit. Jung. " the two plants are given as Jung, minutissima, Sm., but as I have never heard of any one collecting Lej. minutissima, Sm., in the Lake District, I should infer that the station given by Hooker, Keswick, refers to Lej. ulicina. Lej. ulicina, Tayl., is quite distinct from any of the otlier British Lejeuneae. Description of Plate XIV. — Fig. L Plants natural size. 2. Portion of stem, antical view x 64 (Ingleton, Webster and Pearson). 3. Ditto, postical view x 85 (ditto). 4. Ditto, postical view X 85 (Herb. Sowerby). 5. Ditto, postical view x 85 (Levens, Stabler). 6. Leaves, antical view x 85 (France, Husnot). 7-9. Ditto, postical view x 85 (Ingleton, Webster and Pearson). 10,11. Portions of leaves x 290 (ditto). 12, 13. Stipules x 85 (ditto). 9. Lejeunea diversiloba, Spnice. Lejeunea diversiloba, Spruce Journ. of Bot. 1876. Lejeimea minutissima, var. major, Carringt. Irish Cryptogams Trans. Edin. Bot. Soc. (1S63). Dioicous, epiphytic on Frullania or other corticolous species, small, pale yellowish-green, sometimes nearly white. Stems slightly branched, branches rigid, subfastigiate, distant, filiform ; radiculose, rootlets very short, fasciculate. Leaves incubous, approximate, erecto-patent, angle about 30°, to below the middle complicato-bilobed (or none), antical lobe slightly concave, LKJEUNEA. 57 obliquely obovate-obloni^, rotundute, rarely obtuse or subacute, quite entire, postical lobe variable in form and size, nearl)' e([unl to the antical lobe or half the size, or very minute or absent alto- gether, sub«|uadrate, or oval, inflated, the upper Ijorder incurved and notched at the outer angle so as to form a rather blunt tooth, at the keel broadly rptundate ; texture somewhat firm ; cells pellucid, from smallish to rather minute, 5- and G-angled, subplane, walls lirm, no trigones or thickened angles. Stipules about .'} times shorter than the antical lobe of leaf, slightly Itroader than the stem, subrotund or oval, to about the middle or below bifid, sinus acute or lunate, segments suberect or incurved, subacute ; sometimes obsolete. Female flowers lateral sessile ; bracts longer than the leaves, suberect, loosely complicate-bilobed to below tlie middle, antical lobe ublicjuely obovato-lanceolate, subacute, obtuse or rotundate, postical lobe almost of equal length or shorter, ligulate or oblong ; bracteole obovato-lanceolate, bilobed to below the middle, segments narrow, obtuse or acute. Perianth immersed in the bracts, subglobose, slightly compressed, highly 5-carinate, very smooth. Androecia on short lateral branches; perigonial bracts '1 pairs, larger than the leaves, very turgid, lobes subequal, almost hemispherical, diandrous. Dimensions. — Stems ^ to ^ sometimes 1 inch long, diameter •05 to "OG mm. with leaves irom 3 mm. to '45 mm. wide; leaves, antical lobe "35 mm. x 2 mm., postical •175 mm. x •! mm., antical •2 mm. X •IS mm., postical '15 mm. x '1 mm., antical "25 mm. x 1 mm., postical '075 mm. x 025 mm., antical '275 mm. x '15 mm., postical '0; cells •02 mm., ^025 mm., "015 mm. ; stipules "1 1 mm. X ^07 mm., segments 075 mm., •125 mm. x '08 mm., seg. •04 mm., ^125 mm. x "1 mm., seg. ^04 mm., •! mm. x "075 mm., seg. •05 mm., OS mm. x 'OG mm., seg. "O."') mm. ; bracts, antical lobe •325 mm. x "15 mm., postical •:• mm. x 075 mm., antical '375 mm. X '15 mm., postical "25 mm. x I mm. ; bracteole ■3mn). x 2 mm., segments "1 75 mm. H.\B. — xSot unl're([uent about Killarney, growing on Fnil/aiiia germana and other prostrate mosses and hepatics, Dr. Tai/hr, l)r. Spruce, Dr. Carrin*/. Clapdale Cave, Craven, Br. Car- rni(/io)}. Bolton Woods, Br. Carriiujlon. Nr. Litton, John Ji'/iitc- head. 12. Heversham Head, G. Stabler. Pasture Beck, Ullswater, //'. //. P. 15. Banks of the River J3urn, A. Croall. Cawdor Wood, Forfar, J. Croall. Glen Tilt, A. Croall. IG. ]\Ioidart, West Inverness, S. M. Macricar, peaty side of ravine, not limestone. 1. Kerry, Br. Taylor. Killarney, Dr. Carrimjton. Found on the Continent and in North America. Obs. — A very distinct species and not likely to be confounded with any other British ones, except the closel}' allied Lcj. lxos.se /liana, Massal., which see. Lej. haniafifolia (Hook.), whicli Hooker considered it to be a variety of, diflers in the antical lobe of leaf beiii^- distantly (U'litate, epidermis smooth, j)resence of stipules, and other characters. Descuiption of Plate XVI. — Fi«;-. 1. Phmts natural size. 2. Portion of stem, antical view x G4. :\, 4. Portions of stems, postical views X (".4. 5-8. Leaves x G4. 9-11. Ditto, cxplanate x G4. 12. Portion of leaf x 290. 13. Papilhe of antical lobe of leaf X 290. 14,15. Styliform appendages x 85. IG. Bract x G4. 17. Perianth X 31. 18. Portion of male branch x (M. 19. Peri- gonial bract x 85. 20. Anthcriiliuiii x '^b. (Teesdale, Dr. Spruce.) 60 HEPATIC^. 11. Lejeunea Rossettiana, Mussed. Lejeunea Rossettiana, Massalongo, Nuovo Giorn. Bot. Ital. vol. xxi. p. 487 (1889). Dioicous, intricately caespitose, minute, yellowish-green. Stems subdicliotomoLis or subpinnate, more or less radiculose, rootlets few, fasciculate, pale. Leaves imbricate, patent, 50°-60°, bilobed, antical lobe somewhat convex, ovate, apices more or less attenuate-acuminate (rarely only acute), usually incurved, upper surface echinate, postical lobe about |- to ^ the size of the antical, subquadrate to roundish-quadrate, tumid at the keel, upper portion plane, margin not incurved, unequally dentate-spinulose, exterior surface echinate like the antical lobe, texture opaque : cells very minute, 4-, 5-, and 6-sided, walls distinct, no trigones or thickened angles ; papillse conical, one to each cell. Stipules absent. Stylus also wanting. Female flowers on short branches, subtended by an innovation. Bracts similar to the leaves, only larger, postical lobe with margin more finely dentate-spinulose. Bracteole wanting. Perianth projecting more than half beyond the bracts, oval to pyriform, upper portion obtusely 5 -angled, epidermis rauricate. Androecia unknown. Dimensions. — Stems about \ of an inch long, "05 mm. diameter, with leaves '85 mm.-l* mm. wide; leaves, antical lobe •45 mm. x -25 mm., postical -225 mm. x "15 mm., antical -45 mm. X "275 mm., postical "225 mm. x '15 mm.; cells -0125 mm.; papillse "0125 mm. long x '01 mm. wide at the base ; bract, antical lobe "5 mm. X -25 mm., postical '25 mm. x '15 mm.; perianth •6 mm. X -45 mm., '55 m.m. x -4 mm. Obs. — Diflers from Lejeunea calcarea, Lib., with which it has been confounded by British botanists in its dioicous inflorescence, more opaque texture, slightly larger size, leaves a little more spreading, echinate postical lobe, with margin more dentate- spinulose, and not incurved, as in L. calcarea (the margin of which cannot be seen without dissection), and by the entire absence of the styliform appendage, usually found between the stem and the leaf of L. calcarea. The above mentioned characters show this IKJEUNEA. 01 plant to be a distinct species, and was discriminated ])y tlic Italian botanist, Prof. Massalongo, from specimens collected in Italy by Dr. Rossetti. Orii^inal specimens from Dr. Taylor of Loj. echinata comprise both this species and Lcj. calcarna, but as Prof Massalonyo was the iirst to recognise its distinctness, his name justly deserves to be preserved. From other British Lcjeunecc it is at once separated by its echinate leaves. Hab. — On limestone rocks, amongst mosses, and often inter- mingled with L. calcarea, Lib. Rare. 8. Millers Dale, Winnats, Ravensdale, Derbyshire, C. A. Jloll. 10. Limestone Cave, Gordale, Yorks, Dr. Carrinf/fon. L Near Dublin, 1830, Br. Tai/Ior. Mucross Desmesnc, Killarney, J)r. Carri)i(/ton^ 18G1. Found on the Continent (Italy, l)r, Jiossetti). Dkscription of Plate XVII. — Fig. 1. Plants natural size. 2. Stem, antical view x 24. 3. Portion of stem, postical view x ()4. 4-7. Leaves x G4. 8. Leaf x 85. 9. Portion of leaf x 21)0. 10. Papilla? X 290. 11. Bract x G4. 12. Perianth x 31. 13. Cross-section of perianth x 31. 14. Apex of perianth x 31 (Winnats, Derbyshire, G. A. Holt). 12. Lejeunea minutissima [Sm.). Jungei-mania mimUissima, Smith, Eng. Bot. t. IG80 (18UC), Hook. Ijrit. Jung. t. 52 (ISIO). Junyefi-inania incoiispicua, Kaddi in Act soc. Modena (iSlS). Lejeunea Tai/loi-i, Spruce, in Trans. Bot. Soc. Edinb. vol. iii. p. 12 (1S|9). Lejeunea inconspicua, de Not. in G. and R. Hen. eur. n. 15, Dum. Hej). Eur. p. 18 (1K74). Lejeiinea minutissima (Sm.), Spruce Journ. of Bot. p. 'M\ (I SSI). Monoicous, closely but shallowy or loosely c.Tspitose, minute, pale green to whitish. Stems filiform, geniculate, llexuose, creeping, irregularly branched. Leaves hilarious, alternate, distant or approximate, erecto-patent to erect, subrotund, almost all the length complicate, very concave or turgid, postical lobe very little 62 HEPATIC^. narrower than the antical ; epidermis subpapillose, texture delicate, cells smallish, 4-, 5-, or 6-angled, walls thin, no trigones or thickened angles. Stipules absent. Female flowers terminal on short lateral branches ; bracts about twice as long as the leaves, broadly oblong, slightly complicate, very shortly bilobed, postical lobe about twice as narrow as the antical. Perianth emersed, turbinate or pyri- fonn, slightly compressed, acutely 5-angled, keel almost smooth, sometimes furnished with a single series of pellucid cells. Calyptra pyriform, texture similar to the leaves. Capsule subspherical, hyaline, cleft halfway to the base. Androecia on short lateral branches, roundish or spicate ; perigonial bracts 3, 4 pairs, cymbi- form, turgid. Antheridia single, oval. Dimensions. — Stems J to J inch long, and "05 mm. in diameter, with leaves '35 mm. to "4 mm. wide ; leaves, antical lobe '225 mm. X -175 mm., postical "2 mm. x 'l mm., antical '225 mm. x •175 mm., postical -2 mm. x '175 mm., antical -25 mm. x 'l mm., postical -225 mm. x -1 mm. ; cells '025 mm. x "02 mm., "02 mm. X '02 mm., '03 mm. x '02 mm., -025 mm. x '025 mm. ; bracts, antical lobe, '35 mm. x '12 mm., postical '3 mm. x '1 mm., antical •275 mm. x -15 mm., postical -225 mm. x 'l mm. ; perianth '5 mm. X -275 mm.; male catkin '4 mm. x -3 mm ; perigonial bract •2 mm. X -15 mm. ; antheridia 'l mm. x '075. IIab. — On stems of trees. Very rare. 1. Trevello Wood, Penzance, Cornwall, IF. Ciirnoio and others. 2. Henfield, Sussex, C. Lijell New Forest, Hampshire, C. Lyell. Langley, Sussex, F. C. Roper. 3. Shiere, Surrey, E. M. Holmes. South of Ireland. Found on the Continent. Obs. — Being destitute of stipules this plant is at once distin- guished from those British species which have them. Lej. calcarea, Libert, and Lej. Rossettiana, Massal. have ecliinate leaves. Lej. microscopica, Tayl. has more spreading, narrower leaves, and is rarely found fertile. See notes under Lej. ulicina, Tayl. Description of Plate XVIIL— Fig. 1. Plants natural size. 2. Portion of stem, antical view x 64 (Langley, Roper). 3. LEJEUNEA. 63 Ditto, postical view x 64 (ditto). 4. Ditto x 64 (Herb. Taylor). 5, 6. Leaves x 64 (New Forest, C. Lyell). 7, 8. Portions of leaf x290 (ditto). 9, 10. Bracts x 64 (ditto). 11, 12. Ditto x 64 (Langley, Roper). 13. Perianth x 64 (New Forest, C. Lyell). 14. Male catkin x 64 (Coimbra, Portugal, Henriques). 15. Antlieridium x 85 (ditto). 13. Lejeunea microscopica, Taylor. Jungermania microscopica, Tayl. in Mackay Fl. hib. 2 p. 59 (1836). Taylor in Hook. Journ. of Bot. 4, p. 97, t. 29. Nees, Nat. eur. Leb. Ill, suppl.p. 5GG (1838). Lejeunea microscopica, Tayl. G. L. N. Syn. Hep. p. 315 (1844). Paroicous, loosely caespitose, minute, very pale green to white. Stems creeping, very slender, Hexuose, pellucid, subramose ; radi- culose, rootlets few, distant, short, single, pellucid. Leaves often unilateral, distant, patent, ovoideo-fusifbrm or ovate-lanceolate, somewhat flattened, slightly unequally bilobed, antical lobe a little larger than the postical, apex subacute, triangular or shortly linguaform ; epidermis papillose, in some specimens almost smooth, texture pellucid ; cells smallish, 4-, 5-, and 6-angled, walls thin, no trigones or thickened angles, near the base in the middle elongate. Stipules absent. Female flowers on short branches, bracts unequally bilobed, antical lobe narrower and longer than the leaves, ovate- lanceolate, postical lobe much smaller, lanceolate acute. Perianth pellucid, obovate or obconical, j^apillose, ecarinate or obtusely angled, apex flat, no beak, mouth contracted. Andrcecia on short lateral branches, perigonial bracts bilobed, more obtuse at the apex than in the leaves. Antheridia oval. Dimensions. — Stems ^ inch long, -03 mm. in diameter, with leaves '35 mm. wide ; leaves, antical lobe "2 mm. x '12 mm., posti- cal '175 mm. x '12 mm., antical '175 mm. x '1 mm., postical •16 mm. X -1 mm. ; cells "03 mm. x "025 mm., -03 mm. x "03 mm., near base of leaf -04 mm. x '02 mm. ; bract, antical lobe '25 mm. X "15 mm., postical "15 mm. x "05 mm. ; perianth "35 mm. x '225 mm. near the apex. Hab. — On Fridlanm, bark of trees or on damp rocks, in shaded or exposed situations. Extremely rare. 64 HEPATIC^. 7. Eocks below Llyn OgweD, Carnarvonshire, Dr. Carrington and W. IT. P. 12. Rostliwaite, Cumberland, Dr. Carrington and W. H. P. Seathwaite, Cumberland, /. R. Bijrom and W. H. P. 16. Moidart, West Inverness, S. M. Macvicar. I. Cromaglown, Dr. Taylor, Dr. Spruce, Dr. Carrington, Dr. Moore, Prof. Lindberg, and others. Glenariff, Co. Antrim, Revs. Lett and Waddell. Obs. — This is one of the smallest and most beautiful of the British Hepaticae ; single plants being scarcely visible to the naked eye. From Lej. minutissima (Sm.), which is also without stipules, it is distinguished by its more distant, spreading, narrower leaves, which are also more papillose and pellucid. Lej. catcarea, Libert, and Lej. Rossettiana, Massal., are larger, coarser species, with more erect leaves, which are distinctly echinate. From all other British LejeunecB it is distinguished by the absence of stipules. Description of Plate XIX. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 85 (Rocks, Ogwen, W. H. P.) 3. Ditto, postical view x 64 (ditto). 4, 5. Leaves, postical view x 85 (ditto). 6. Leaf (ditto) x 64 (Herb. Tayl.). 7, 8. Leaves, antical view x 64 (ditto). 9. Keel of leaf x 290 (Seathwaite, W. H. P.) 10, 11. Portions of leaf x 290 (ditto). 12. Bract X 64 (Glenariff, Lett & Waddell). 13. Perianth x 64 (ditto). 14. Lejeunea calyptrifolia {Hook.), Dum. Jungermania calyptrifolia, Hook. Brit. Jung. t. 43 (1816). Lejeunea calyptrifolia, Dum: Comm. p. Ill (1822). Colura calyptrifolia, Dum. Recueil, p. 12 (1835). Dioicous (?), csespitulose, minute, pale yellowish-green. Stems creeping, slightly and irregularly branched, branches procumbent or suberect ; filiform, flexuose ; radiculose, rootlets extremely minute, pellucid. Leaves closely placed, bifarious, horizontal, patent or erect, largest at the base of the plant, gradually lessening to the apex, unequally bilobed, antical lobe calyptri- form, rostrate corniculate, apex incurved and acute, narrowed at the base, but with a semi-orbicular or broadly oval convolute LEJEUNEA. 65 wing, above whicli the margin is slightly incurved, the fold gradually increasing in size until near the apex, where another erect horseshoe-shaped lobe is observed, which is of an extremely delicate texture and is enclosed in the tubular portion of the antical lobe ; postical lobe about three times smaller than the antical, oval or subquadrate with rounded corners, convolute ; texture delicate, cells smallish, elongate, 4-, 5-, and 6-angled, walls thin, very slightly thickened, trigones absent. Stipules at the base of each leaf, plane, appressed to the stem or a little patent, oblong-cuneate, bipartite or deeply (to two-thirds) bifid, segments patulous, linear-subulate, 3 to 4 cells wide at their base. Female flowers lateral ; bracts 2, 3 pairs, minute, erect and closely ap- pressed to the perianth, subquadrate or oval, apex retuse, emarginate, entire or irregular. Perianth i-elatively large, almost as long as the leaves, projecting three-fourths beyond the bracts, oblong, attenuated at the base, widening towards the apex, obtusely 5-angled, angles furnished with one or two horns ; similar horns are found on other parts of the upper portion of the perianth, apex depressed or flat, mouth closed at first, afterwards slightly opened, contracted, margin irregular. Caiyptra pyriform, com- posed of thicker and smaller cells than the perianth. Capsule oval. Spores minute. Androecia not known. Dimensions. — Stems J inch long; leaves, antical lobe 1"25 mm. X -35 mm., postical "5 mm. x '2 mm., antical 1-5 mm. x -45 mm., postical "5 mm. x '25 mm., upper inner lobe '3 mm. x '15 mm., basal lobe "3 mm. x "225 mm., '3 mm. x "25 mm.: cells •025 mm. x '025 mm., "02 mm. x "035 mm., '02 mm. x '04 mm. ; stipules '25 mm. x "15 mm. at the middle, segments '175 mm. ; bracts "45 mm. x '35 mm., '45 mm. x -3 mm., "45 mm. x 25 mm. ; perianth 1" mm. x -4 mm., -9 mm. x '4 mm. ; caiyptra "6 mm. x "35 mm. ; capsule "375 mm. x "2 mm. ; spores "01 mm. Hab. — In minute, pale yellowish-green tufts, on rocks, trees, or furze bushes. Extremel}^ rare. 1. Trevaylor, Cornwall, /. Ralfs, IF. Curnow.. 7. Tyn-y-Groes, Merionethshire, //o// and Wild. 12. Lodore, Cumberland, G. Ly ell and W. 11. P. Stone- thwaite, Cumberland, -/. U. Byrom and W. H. P. 16. Moidart, E 66 BE PAT ICE. West Inverness, S. M. Macvicar. South of Ireland, Miss Hutchiiis, Br. Taylor, Br. Spruce, IV. Wilson, Br. Carrington, &c. Found on the Continent (France). Obs. — The most curious character about this plant is the inner upper enclosed horseshoe-shaped lobe which Dr. Gottsche was the first to observe, and who seemed to think it had some function which we do not understand, for in an allied Lejeiinea from the Straits of Magellan, the inner structure of which was very similar to our species, he found in the pouch-like lobe the chitinous remains of many small animals which had been unable to find their way out of this trap-like arrangement. A closer investigation of this species will reward the student with particulars which are hardly to be expected in a work of mere descriptive botany. As will be seen from the description and drawings, it is a species which cannot be confounded with any other British one. Description of Plate XX. — Fig. 1. Plants natural size. 2. Plant X 31 (Borrowdale, W. H. P.). 3-7. Leaves x 31 (ditto). 8. Leaf x 31 (Cherbourg, France, Corbiere). 9. Basal lobe of leaf X 64 (ditto). 10. Upper lobe of leaf x 64 (ditto). 11. Portion of leaf x 290 (Borrowdale, W. H. P.). 12-14. Stipules X 64 (ditto). 15, 16. Bracts x 31 (Cherbourg, Corbiere). 17. Bract X 31 (Borrowdale, W. H. P.). 18, 19. Perianths x 31 (Cherbourg, Corbiere). 20. Apex of perianth, showing mouth X 31 (ditto). Tribe 11. JUNGEEMANIE.E. Subtribe I. RADULE^. Genus 4. RADTJLA, Bum. CandoUece, sp. Raddi Jung. Etr. in Mem. Moden. xviii. p. 22 (1820). Martinellii, Gr. & B. Nat. Arr. Br. PI. i. p. 690 (1821). Radulce,STp. Dum. Comm. p. 112 (1822). Radulce, sect. Radulotypus, Dum. Syll. Jung. p. 38 (1831). Radula, Dum. Recueil, p. 14 (1835); Nees, Nat. Eur. Leb. Ill, p. 145 (1838). Stems laxly pinnate or dichotomous ; branches lateral, arising from the outer base of the leaves. Leaves incubous, complicate, RADULA. 67 bilobed, postical lobe smaller than the antical, inflexed, producing rootlets. Stipules absent. ? flowers poly^^onous, fruit terminal on short branches or at the base of the fork. Perianth campanulate, compressed or subterete, truncate, mouth dilated. Bracts entire, 1-pair, somewhat similar to the leaves, deeply bilobed. Calyptra pyriform, persistent, free, opening below the apex. Pedicel stout, composed of 6 to 8 concentric layers of alternate cells. Capsule oval-cylindricai, dividing to the base into 4 valves. Elaters numerous, long, slender, attenuate at both ends, bispiral, decid- uous. Spores large, globose. Androecia spicate ; perigonia bracts 3-15 pairs, small ; antheridia 1 or 2 (rarely 3) in the ven- tricose base of the bracts. 1. Radula voluta, Taylor. Radula voluta, Tayl. MS. G.L.N. Syn. Hep. (1844). Dioicous, shallowly csespitose, medium size, pale yellow to yellowish-brown. Stems procumbent, pinnately branched. Leaves laxly imbricated, incubous, divergent to patent-divergent, un- equally bilobed, antical lobe broader than high, subrotund, obtuse, quite entire, slightly undulate, crossing the stem, decurrent at its postical base, postical lobe about one-third its size, broader than high, subrotund-cordate, plane, undulate or with margin slightly recurved, crossing the stem, auriculate at the base ; texture some- what flaccid ; cells small, 4-, 5-, or 6-angled, walls moderately thick, without trigones. Androecia on short lateral catkin-like branches, perigonial bracts 3, 4 pairs, closely imbricate, erect, turgid, bilobed, lobes oval, postical rather shorter. Female plant not known. Dimensions. — Stems 1 to 1|- inches long with leaves 2' to 3" mm. wide, diameter '2 mm. to '3 mm. ; leaves, antical lobe 1* 6mm. X 1'9 mm., postical lobe "9 mm. x I'nim., antical 1*5 mm. X 1"5 mm., postical "5 mm. x '6 mm., antical Tl mm. x 1"2 mm., postical *5 mm. x '7 mm. ; cells '02 mm. x '02 mm., '02 mm. x •025 mm.; male catkin 3* mm.xl"5 mm., 2*5 mm. x 1'25 mm. ; perigonial bracts, antical lobe V2 mm. x "6 mm., postical 68 HEPATIC.^. 1" mm. X '6 mm., antical 1* mm. x '5 mm., postical *8 mm. x •4 mm. Hab. — On damp or wet rocks, or on earth-encrasting rocks, near streams. Rare. 7. Torrent Walk, Dolgelly, Dr. Carrington and W. H. P. Tyn-y-Groes, Merioneth, JF. 11. P. 12. Lodore Falls, Cumber- land, Dr. Carrington and W. H. P. Rostliwaite, Cumberland, Dr. Carrington and W, H. P. 13. Ness Glen, Dalmellington, Ayrshire, J. McAndreio. I. Dunkerron, Dr. Taylor. Tore Water- fall, Dr. Spruce, Dr. Carrington. Rocks below Eagle's Nest, Cromaglown, G. E. Hunt. Glena, $ Stewart and Holt. Not found on the Continent or in North America, the specimens from Tallulah Falls, Georgia, belonging to a different species. Obs. — Distinguished from other dioicous British Radulce by its pale yellow colour, broad antical lobes, and subrotund-cordate auriculate postical lobes of the leaves, which are more or less undulate and of a flaccid texture. Badula xalajjcnsis, Mont., from Bolivia, according to the opinions of Dr. Spruce and Stephani, who have seen original specimens, is a different species, which I fully agree with, having myself examined them. Description of Plate XXI. — -Fig. 1. Plant natural size. 2. Portion of stem, postical view x 24 (C. & P., n. 44). 3, 4. Leaves, antical lobes x 24 (Tore Waterfall, R. Spruce). 5. (Ditto) x (Herb. Taylor). 6, 7. (Ditto) x 24 (Tore, Dr. Spruce). 8. Portion of leaf x 290 (Glena, Holt and Stewart). 9. Male catkin, postical view x 16 (ditto). 2. Radula Lindbergii,, Gottsche. Radula comiJlanata (L.), var. propagxdifara, Nees,'Nat. Eur. Leb, iii. p. 148 (1838). Radula Lindenhergiana, Gottsche, Hartmann's Handb. Scaud. Fl. 9, Aufl. p. 98 (1864), in error, should be R. Lindhergiana. Radula commutata, Gottsche, Jack, " Flora," n. 23 & 25 (1881), Dioicous, widely and shallowly casspitose, medium to largish in size, from dull greenish-brown to dark sordid brown. Stems prostrate, subpinnate, irregularly or regularly bipinnate, branches RADULA. 69 ascending. Leaves imbricate, patent-divergent, 70°, unequally bilobed, antical lobe plane or slightly convex, crossing the stem, subrotund or broadly obovate, margin quite entire or erose by formation of gemmse, postical lobe 3 to 4 times smaller, appressed to the antical lobe, quadrate, free angle acute, inner margin reaching to the middle of the stem, upper angle rotundate or acute ; cells smallish to medium in size, hexagonal, walls firm, no trigones or thickened angles. Bracts unequally bilobed, antical lobe obovate-elongate, postical lobe about one-third smaller, oblong-quadrate, free angle acute or obtuse. Perianth projecting to about the middle beyond the bracts, obovate, complanate, trun- cate. Capsule brown, dividing into 4 longish oval valves, elaters usually monospiral ; spores roundish. Androecia terminal on the chief stem or on long lateral branches ; perigonial bracts closely imbricate, bilobed, turgid at the base, antical lobe oval, apex rotun- date, postical slightly shorter, oval, apex obtuse ; antheridia spheri- cal. Gemmse often abundant at the margin of the upper leaves. Dimensions. — Stems 1 to 2 inches long, with leaves 2" to 3* mm. wide, diam. of stem "15 mm. ; leaves, antical lobe TS mm. x 1'25 mm., postical lobe 75 mm. x "6 mm., antical 1'25 mm. x 1* mm., postical "6 mm. x '5 mm., antical 1*2 mm. x '8 mm., postical •6 mm. x '4 mm. ; cells '0225 mm. x '0225 mm., '03 mm. x '03 mm., '0225 mm. x '03 mm. ; bracts, antical lobe 1-25 mm. x 75 mm., postical lobe 75 mm. x '5 mm.; perianth 2*25 mm, x 1*1 mm. ; elaters '3 mm. x "006 mm. ; spores "03 mm. Hab. — In spreading flat patches, closely attached to damp rocks in exposed or shaded situations, usually in hilly or subalpine localities. Eare. 2. New Forest, C. Lyell, 1813, March 12, 1814. 7. Clogwyn- du-Arddu, Snowdon, Carnarvonshire, W. H. P. 12. Harter Fell, and Mardale, Westmorland, G. Stabler. Near Keswick, Dr. Carrington and W. H. P. 15. Braemar, A. Croall. Found on the Continent and in Canada (rocks near Belleville), Macoun. Obs. — Distinguished from R. complanata (L.) by its dioicous inflorescence; from R. aquilegia, Tayl., by its colour, and the 70 HEPATICjE. larger, plane postical lobe, which is not turgid at the base ; from B. Carringtoni, Jack, by its greenish sordid brown colour, presence of gemmse, coarser texture, and relatively larger postical lobe. After the examination of numerous specimens of Radula commutata, Gr., I have been unable to find sufficient characters to separate it from R. Lindhergii, G. This species was named by Dr. Grottsche after the late Prof. S. O. Lindberg, of Helsingfors, one of the most distinguished botanists of our time, and whose contributions to the knowledge of the Hepaticse, especially of European species, have been numerous, original, and marked by an accuracy of observation which commands both our respect and admiration. B}^ an error it was first published in Hartmann's " Handbok i Scand. Flora" as Radula Lindawbergiana ; as Lindberg was the discoverer of the species, I have further altered the name from Lindhergiana to Lindhergii. Description of Plate XXII. — Pig. 1. Plant natural size. 2. Portion of stem, antical view x 16 (Clogwyn-du-Arddu, W. H. P.). 3. Ditto, postical view x 16 (Braemar, A. Croall). 4. Ditto X 24 (Hunneberg, Sweden, S. 0. Lindberg). 5. Portion of leaf X 290 (ditto). 6. Perianth with bracts x 16 (ditto). 3. Radula germana, Jack. Radula complanata (L.), var. plumulosa, Nees & var. tenuis, Nees, Nat. Eur. Leb. iii. p. 148 (1838), G.L.N. Syn. Hep. p. 257 (1844). Eadula gerinana, Jack, "Flora," n. 23 & 25, pp. 355, 395-7 (1881). Dioicous, csespitose, small, pale yellowish or bright pea-green in colour, young terminal branches darker green, the lower parts and older stems being of a pale sordid brown colour. Stems procumbent or erect when growing with mosses, imbricate, frontally compressed, narrow, graceful, flexuose ; female plant subpinnate, furcate or dichotomous, barren stems almost simple, with few and very short branches, which are longer near the apex ; radiculose, rootlets few, arising from the underside of the lobule, fasciculate, short, sordid wdiite. Leaves imbricate, below approximate, alternate, une(jually bilobed, antical lobe roundish- BABUL A. 71 ovate or olxjviite, convexulous, entire, or when gemmiparous, erose; postical lobe about one-third the size, rhomboid, subquadrate or quadrate, obtuse or acute at the free angle, tumid at the base, upper portion plane and appressed to the antical lobe ; cells small, hexagonal, lumen usually filled with chlorophyllose granules, walls moderately firm, trigones very minute or not observable. Bracts, antical lobe oblong-obovate, postical oblong- quadrate narrow. Perianth projecting two-thirds beyond the bracts, oblong-obconical, vvith a tapering base, complanate, mouth truncate, quite entire. Pistillidia 8-10. Calyptra with a long neck, delicate, pyriform. Capsule oblong-oval, dividing to the base into 4 oval valves. Spores subspherical, finely granulate. Elaters bispiral, loosely twisted. Male stems irregularly pinnate, catkins lateral, long or short, consisting of from 8-15 pairs of perigonial bracts, which are closely imbricate, erect, tumid, bilobed ; antical lobe oval, postical shorter, roundish-oval ; antheridia oval, large, solitary. Dimensions. — Stems f inch to 1 inch long, with leaves 1' mm.-l*25 mm. wide, diameter '17 mm. x '11 mm., '16 mm. x 'll mm., '15 mm. x '11 mm., leaves, antical lobe 1*2 mm. x "8 mm., 1'2 mm. X 7 mm., I'l mm. x "8 mm., 1' mm. x "7 mm., 1" mm. x •6 mm., postical '7 mm. x "3 mm., '6 mm. x '4 mm,, 6 mm. X '35 mm., '55 mm. x "35 mm., -4 mm. x "3 mm. ; cells "02 mm.; bracts, antical lobe 11 mm. x '6 mm., postical '8 mm. x '4 mm., antical 1* mm. x "5 mm., postical '9 mm. x •4 mm. ; pistil- lidia ■12 mm. X "04 mm. ; perianth 2'2 mm* x 1*1 mm., 2*1 mm. x •9 mm. ; valves of capsule "9 mm. long; spores "044 mm. diam. ; elaters -28 mm. x '006 mm. ; male spikes 1' mm. long, '2 mm.- •4 mm. wide ; perigonial bracts, antical lobe '8 mm. x '4 mm., '7 mm. X '45 mm., '025 mm. x 45 mm., '6 mm. x •45 mm., '6 mm. x •4 mm., postical '6 mm. x '4 mm., '45 mm. x '3 mm. ; antheridia "2 mm. x "15 mm.; gemma) 1 mm. diam. Hab. — In procumbent patches, with stems imbricating, on rocks, or erect when intertwined with mosses (Dicrauum falcatum, &c.) ; in alpine situations. Rare. 15. Pvocks by the Burn, Forfar, A. Croall Loch-na-Gat, Ben 72 HEPATIC J^. Lawers, George E. Davies. Mael Tarmachan near Ben Lawers, C. J. Wild, G. A. Holt. Ben Cruaclian, Loch Awe ; Ben Laoigh, Craig Calleach, 2\ Rogers. Found on the Continent (Germany, Switzerland, Austria). Obs. — Differs from R. comjdanata (L.) in its dioicous inflorescence ; from R. aquilegia, Tayl., in its pale yellowish-green colour, the leaves not being so tumid at the base, and the perianth being longer and narrower; from R. Lindbergii, G. {R. com- mutata, G.) in its paler colour, narrower stems (with leaves), more erect leaves with smaller cells, and lons^er and narrower perianth. Some authorities consider this species to be only a variety of the latter, but the characters indicated induce me to follow Jack in raising it to the rank of a separate species. Description op Plate XXIII. —Fig. 1. Plants natural size. 2. Portion of stem, antical view x 24 (Mael Tarmachan, G. A. Holt). 3. Ditto, ? postical view x 16 (ditto). 4. Leaf x 24 (Germany, Jack). 5. Portion of leaf x 290 (Mael Tarmachan, G. A. Holt). 6. Perianth and bracts x 24 (Germany, Jack). 7. Perigonial bracts x 31 (Ben Cruachan, T. Eogers). 4. Radula Holtii, Spruce. Radula Holtii, Spruce in '' Journ. of Bot." (July 1887). Dioicous, creeping, small, rufulous or olive green. Stems slender, fragile, branches subpinnate or dichotomous. Leaves contiguous or subdissitous, at the base decurrent, complicato- saccate, keel at an angle of 45°, suberect, ascending, convex, abruptly widely patent, antical lobe broadly oblong, rotundate, plane or subconcave, repand, much incurrent, not hiding the stem at the basal angle, postical lobe one-third the length, trapeziform, obtuse or subacute, subplane, inflated at the keel only, running well in upon the stem but not passing over (or across) it ; cells small, 4-, 5- and 6-angled, opaque or sub-pellucid, walls thick, angles but slightly thickened, without trigones. Flowers terminal, innovations unilateral or two opposite. Bracts often only one IIADULA. V3 pair, erect if more, closely imbricate, bilobed to the middle, antical lobe broadly obovate, lobule hardly half the size, subrotund. Perianth projecting very much, slender, subincurved, trumpet shaped, perfectly terete, only at the apex very sliglitly compressed, mouth truncate, obsoletely 4-lobed. Androicia terminal on branches ; perigonial bracts 2, 3 pairs, similar to the leaves only lobule larger, slightly turgid, diandrous. Dimensions.— Stem ^ inch long, 'I mm. diameter, with leaves 1"25 mm. wide ; leaves, antical lobe 'S mm. x "6 mm., "65 mm. x •55 mm., "7 mm. x "G mm., postical lobe or lobule 25 mm. x "3 mm., '2 mm. x "ITS mm., "3 mm. x "275 mm. ; cells '02 mm. x •02 mm,, '03 mm. x '02 mm., '02 mm. x •0175 mm., "0175 mm. x .0175 mm. ; bracts, antical lobe •G mm. x ^4 mm., postical lobe •37 mm. X "35 mm.; perianth, 21 mm. x ^3 mm. (mouth) x ^15 mm. below the middle, 2' mm. x '35 mm. (mouth). Hab. — ^Extremely rare, the only known station being near Tore Waterfall, Killarney, growing with Buniortiera irr'ujua, Juhula Hutcliimice and Lejeunea Mackaii. G. A. Holt, June 1885; D. McArdle, 1897. Obs. — This distinct species cannot be confounded with any other British ones ; from all it is distinguished by its smaller size and slender habit ; R. germana, Jack, which is also dioicous and somewhat slender, has larger and squarer leaf lobules, tlie fold of which is not decurrent nor divergent from the antical lobe, and the perianth being of a different shape and complanate. The most important character of R. Iloltii, however, is the slender, trumpet-shaped, terete perianth, sufficing to distinguish this Radula from every other European species. Justly named by Dr. Spruce after its discoverer, Mr. Gr. A. Holt, of Manchester, who is one of the most accurate students of the British Mosses and Hepaticae. The description and notes are taken from Dr. Spruce's com- munication to the " Journ. of Bot. " for July 1887. Description of Pj>atk XXIV. — Fig. 1. Plants natural size. •2. Portion of stem, antical view, x 24. 3. Ditto, postical view, X 24. 4. Portion of branch, postical view, x 24. 5, 6. Leaves, 74 HEPATICjE. antical view x G4. 7. Ditto, postical view, x 64. 8. Portion of leaf X 290. 9. Perianth x 31. 10. Portion of male stem x 31 (Tore Cascade, G. A. Holt). 5. Radula aquilegia, Taylor. Jungerma7iia aquilegia, Tayl. Trans. Bot. Edin. vol. xi. p. 115-117 (1845). Radula aquilegia, Tayl. G. L. N. Syn. Hep. p. 260 (1844). Jungermania complanata, var. h. minor. Hook. Brit. Jung. t. 81, f. 17 (1816). Dioicous, broadly and shallowy ciespitose, medium to largisli in size, olive- brown in colour. Stems prostrate, irregularly pin- nate or subpinnate, branches often at nearly right angles to the stem; cortical cells 18-20 brown, inner pellucid, 6 cells in diameter. Leaves contiguous or imbricate, distichous, patent- divergent to patent, 60°, unequally bilobed, antical lobe convex, quite entire, obovate or semi-rotund, apex rotundate, crossing the stem, often inflexed, postical lobe about 4 to 5 times smaller, sub- quadrate or oblong-quadrate, remarkably involute, tumid at the base, free upper margin appressed to the antical lobe, apex acute or obtusate, margin nearest the stem covering it to nearly the middle ; cells small, walls thick, angles thickened, sometimes showing trigones. Flowers ? terminal on long lateral branches with one or two innovations produced from below. Bracts about the same size as the largest leaves or slightly larger, unequally bilobed, antical lobe similar in shape to stem leaves, postical lobe about half the size of antical, subquadrate, apex obtuse or rotun- date, plane. Perianth projecting nearly two-thirds beyond the bracts, elongate-obconical, compressed, mouth wide, truncate, quite entire. JMale plants smaller ; perigonial bracts on main stem or on middle or end of lateral branches, closely imbricate, parallel with the stem or erect, smaller than the stem leaves, base rotundate, pendant below its insertion on the stem, unequally bilobed, antical lobe oblong-oval, convex, postical lobe about half the size, oval, much swollen ; antheridia single, large, subspherical. Dimensions. — Stem 1 to 2 inches long, with leaves 1-5 mm. wide ; diameter of stem "l to "15 mm. ; leaves, antical lobe *8 mm, RADULA. 75 X '65 mm., postical lobe '45 mm. x '3 mm., antical lobe 1' mm. x •8 mm., postical 'G mm. x '4 mm., antical '9 mm. x "75 mm., pos- tical '4 mm. X •35 mm., antical "9 mm. x "8 mm., postical '4 mm. X "3 mm. ; cells '02 mm. ; bract, antical lobe 1* mm. x "9 mm., postical '7 mm. x "4 mm. ; perianth 2* mm. x I* mm. wide at the mouth, perigonial bracts, antical lobe '6 mm. x "3 mm., postical •4 mm. X "3 mm., antical "7 mm. x "3 mm., postical '4 mm. x •3 mm. ; antheridia '2 mm. Hab. — On wet rocks, very closely attached. Eare ; fertile plants extremely so. 7. Tyn-y-Groes, Merionethshire, April 1876, IF. H. P.; 1885 G. A. Holt (cum fr.). Cwm Idwal, Carnarvonshire, JF. II. P. Llanberis, Carnarvonshire, /. Cash and W. II. P. 12. Grasmere, Westmorland, Rev. C. II. Binsiead. Lodore, Cumberland, TF. H. P. Stonethwaite, Cumberland, /. R. Bijrom and IF. //. P. 16. Moidart, West Inverness, S. M, Macvicar (cum per). T. Near Bantry, Miss HiitcUns. Killarney, Br. Carrinyton (cum per.). Glena, Prof. Lindhcrg, Knockavohila, Br. Taylor. Killarney, Br. Moore, &c. Blackstair, Slieve Donard, Co. Down, Rev. C. H. Waddell. Obs. — Distinguished from Radida complanata (L.) by its inflorescence and olive -brown colour; from Radida Carri?iytoni, Jack, see notes under that species. It is easily recognised by the swollen base of its leaves, the perigonial bracts being remarkably tumid. Radida physoloba, Mont., from Auckland Island, is a different species. Description of Plate XXV. — Fig. 1. Plant natural size. 2. Portion of stem, antical view x 24 (C. & P. n. 43). 3-7. Leaves x 24 (Ireland, Taylor). 8. Portion of leaf x 290 (ditto). 9. Perianth with bract x 16 (ditto). 10. Portion of male stem, postical view x 24 (C. & P. n. 43). 11, 12. Perigonial bracts X 24 (ditto). 76 HEPATIC^. 6. Radula Carringtonii, Jack. Radula Carringtonii, Jack, in " Flora," p. 385 (1881). Radula aquilegia, Tayl. var. major, Carrington, Trans. Bot. Soc. Edin. vii. p. 455 (1863). Dioicous, closely but shallowly coespitose, medium to largish in size, pale or olive-brown colour. Stems closely imbricate, pros- trate, irregularly bipinnate, moderately firm, cortical cells similar to the inner, about 30, inner 5x8, with brownish walls ; branches alternate, somewhat ascending. Leaves slightly imbricate or approximate, patent-divergent to patent 70°-50°, unequally bilobed, margin quite entire ; antical lobe plane or very slightly convex, rotund or subreniform, crossing the stem; postical lobe ^^ smaller, quadrate or trapeziform, outer angle acute, inner margin reaching to about the middle of the stem, plane, appressed to the antical lobe ; texture thin ; cells smallish, hexagonal, walls firm, without trigones. Flowers ? terminal on short lateral branchlets, produced from the pinnse, always subtended by an innovation. Bracts rather larger than the leaves, unequally bilobed, antical lobe turbinate to oblong-oval, apex rotundate, infiexed, postical lobe, about half the size, somewhat similar in shape, apex rotundate. Archegonia 7, 8. Perianth (young) compresso-campanulate, bilabiate, lips retuse. Male plant smaller, irregularly pinnate, branches alternate, short ; androecia on short catkins, produced from main stem ; perigonial bracts 5-10 pairs, closely imbricate, erect, turgid, bilobed to about J or ^, antical lobe oblong-oval, postical rather shorter, oval ; antheridia small, spherical, bearers long. Dimensions. — Stems 1 to 2 inches long, with leaves 2" mm. to 2*5 mm. wide, diameter of stem -15 mm. to '2 mm., leaves, antical lobe 1*2 mm. x T mm., postical "55 mm. x "5 mm., antical 1"2 mm. X -9 mm., postical '55 mm. x '4 mm., antical 1*5 mm. x 1*1 mm., postical '7 mm. X "5 mm., antical I'l mm. x '8 mm., postical •45 mm. x -35 mm. ; cells "03 mm. x '0225 mm., "0275 mm. x •025 mm., -0275 mm. x '0275 mm., '0225 mm. x '02 mm. ; bracts, antical lobe 1*25 mm. x "9 mm., postical l* mm. x '6 mm. ; male UADULA. 77 ramuli 1^ mm. to 3 mm. long, perigonial bracts, antical lobe '7 mm. X '4 mm., postical '6 mm. x '35 mm. ; antheridia '1 mm. diameter, bearer '15 mm. long. Hab. — In wide closely imbricated patches on rocks or trees, South of Ireland, Tore Woods ; Cromaglown ; Grlena ; Tomes Woods, Killarney, Dr. Carrin(jton, June 1861. Rocks near Tore Cascade, Killarney (male plant), G. E. Hunt, April 1872. O'Sullivan's Cascade ; Grlena ; Cromaglown, Killarney, Prof. S. 0. Lindberg, 1873. Killarney, Br. B. Moore. Damp shady rocks, Tore Cascade, Steivart and Holt (male plants), June 1885. Aunis- caul, Co. Kerry, B. McArdle (cum per.), June 1898. 16. Moi- dart. West Inverness, S. M. Macvicar. Found on the Continent (Norway), B. Kaalaas. Extremely rare. Obs. — This, the handsomest of our native Radida, was deservedly named by Jack in honour of its original discoverer. Dr. Carrington, who collected it in the South of Ireland in 1861, and thought it was a distinct species, although he had not seen either the male plant or perfect perianths, but deferring to the opinion of some of his friends, he published it as var. major of Uadida aquilegia, Taylor ; since then, the male plant along with more perfect female plants have been met with, which clearly show its specific difference from the latter. It is distinguished from R. aquilef/ia by its larger size, rather paler brown colour, which is golden brown by transmitted light, leaves with less convex antical lobes ; postical lobe relatively larger, plane, appressed to the antical lobe, not turgid at the base, free angle more acute, texture thinner and more translucent ; androecia on short catkins proceeding from the main stem. From any of the other British Radulm it is abundantly distinct. Description of Plate XXVI. — Fig. 1. Plant, female, natural size. 2. Ditto, male. 3. Portion of stem, postical view x 24 (Killarney, Dr. D. Moore). 4. Ditto, with male spike x 24 (Tore Waterfall, Holt and Stewart). 5-7. Leaves x 24 (Killarney, Dr. Carrington). 8. Portion of leaf x 290 (Tore Waterfall, G. E. Hunt). 9. Bracts x 24 (Killarney, Dr. Carrington). 10, 11. Perigonial bracts x 24 (Tore Waterfall, Holt and Stewart). 78 HEPATIC^. 7. Radula complanata [L.), Dum. Jungermania complanata, Linn. Sp. PI. 1599 (1753); Hook. Brit. Jung. t. 81 (1816). Radula complanata, Dum. Comm. p. 112 (1822). Paroicous, imbricately, widely, and shallowly csespitose, largish, pale yellowish-green colour. Stems creeping, stratiose, subpinnate, cortical cells about 35, similar in size to inner, only with thicker walls, cells 8 x 10 in diameter; slightly radiculose, rootlets dirty white, produced from postical side of postical lobes of leaves. Leaves imbricate, patent to patent-divergent, 60°, unequally bilobed, quite entire or erose, antical lobe roundish, slightly convex, covering the stem, postical lobe about ^ smaller, quadrate, angle obtuse or subacute, sometimes arcuate ; cells smallish-medium size, hexagonal, walls thin, trigones very small, lumen full of chlorophyll granules. Bracts similar in size to the leaves, antical lobe elongato-obovate, postical lobe about half the size, ovate, apex rotundate. Perianth projecting about half beyond the bracts, obconical, complanate, mouth wide, truncate, quite entire. Capsule dark brown, oval ; spores large, about 3 times the diameter of the elaters, which are bispiral. Perigonial bracts 3-5 pairs, imme- diately below the perianth (or on a lateral branch produced from below it at the apex of which is always an imperfect female flower), closely imbricate, erect or erecto-patent, tumid, unequally bilobed, antical lobe oval, postical about half the size ; antheridia single, globose. Gemmiparous, gemmae discoid, on the margins of leaves, rendering them often irregular and erose. Dimensions. — Stems 1 to 2 inches long, diameter "15 mm., with leaves 225 mm. to 2*5 mm. wide: leaves, antical lobe I'S mm. x 1"1 mm., postical lobe "7 mm. x '5 mm.; cells '035 mm. x "025 mm. ; innermost and perigonial bracts, antical lobe 1'12 mm. x "82 mm. ; postical 7"5 mm. x 5'5 mm. ; perianth 3* mm. x 1*25 mm. j capsule 1"25 mm. x '75 mm. ; spores "03 ram. ; elaters, '2 mm. x 01 mm. ; antheridia 2 '25 mm' x '2 mm. Hab. — On the bark of trees. Common. 1 to 1 7. I. rOllELLA. 79 Found on the Continent and in North America. Obs. — This, the commonest of our British Radidm, is dis- tinguished from all the others by its paroicous inflorescence. The antheridia are normally situated below the young perianth, but with its maturing, they disappear and the bracts assume the form of ordinary leaves ; but a lateral young branch, proceeding from below the perianth, usually bears 3-5 perigonial bracts with an immature perianth at its apex. Description of Plate XXVII. — Fig. 1. Plant natural size. 2. Portion of stem, antical view x 11 (Barmouth, W. H. P.). 3. Ditto X 16 (ditto). 4. Ditto, postical view x 11 (Dove Dale, W. H. P.). 5. Portion of stem with perianth x 21 (Jack del.). 6. Portion of leaf x 290 (Dove Dale, W. H. P.). 7. Perianth with bracts x 11 (ditto). 8. Perigonial bracts x IG (ditto). 9. Young flower, showing ? and 3" x 40 (Gottsche del.). Subtribe IT. PORELLE/E. Genus 5. PORELLA, mil Porella, Dill. Hist. muse. p. 459 (1741) ; Lindb. in Act. soc. sc. fenn. p. 329 (1809). Bellincinia et Antoira, Kaddi Jung. Etr. iu Mem. Moden. xviii. p. 18 et 19 (1820). Cavendishia, Gr. and Benn. Nat. Arr. Brit. PI. i. p. 689 (1821). Madotheca, Dum. Comm. p. 111. (1822). Dioicous. Stems regularly pinnate and feather-like ; branches lateral, axillary. Leaves incubous, often subopposite, complicato- bipartite ; postical lobe much smaller, exterior base often decurrent. Stipules large, basal angles decurrent with a laciniate and crispate wing. Flowers $ lateral, nearly sessile. Bracts 2-pairs, bilobed, dentate or ciliate. Perianth short, ovate, frontally compressed, trigonous on section, mouth plurilaciniate, at first constricted, afterwards bilabiate or campanulate, Calyptra globose, carnose, rupturing below the apex. Pedicel very short, not projecting beyond the perianth. Capsule globose, pale, membranous, divided nearly to the base into 4 valves. Elaters small, short, bi-trispiral, free, attenuate at both ends. Spores rather large, somewhat angular. Androecia shortly spicate; perigonial bracts exactly 8 HEPATIC^E. opposite and connate with the intervening stipules ; closely imbricate, bilobed. Antheridia solitary, large, oval-globose, stipitate, situated in the saccate bases of the bracts. 1. Porella laevigata (Sc/irad.) Lindb. Jungermania IcBvigata, Schrad. Syst. Samm. 11, n. 104, p. 6 (1797); Hook. Brit. Jung. t. 35 (181(5). Madotheca Icevigata, Dum. Comm. p. Ill (1822). Porella Icevigata, Lindb. Muse, scand. p. 3 (1879). Dioicous, loosely csespitose, largish to large, olive-green to yellowish-brown. Stems procumbent, lying over each other in an irregularly imbricated manner, irregularly bipinnate, branches horizontal, irregular in size, number and position, apices somewhat attenuate ; on a cross-section oval, frontally compressed, 24 cells X 32, the cortical and next inner layer of cells being small with very thick brown walls, the third row having brown walls but thinner, and the inner cells becoming gradually larger with thinner and more hyaline walls ; radiculose, rootlets few, very small, proceeding from the lower part of the stem. Leaves closely imbricate, alternate, appressed to stem, incubous, bifarious, unequally bilobed, antical lobe much the largest, horizontal, convex or subplane, obliquely ovate, acute or acurainulate, margin more or less entire, near antical base often furnished with 2 or 3 teeth ; postical lobe erect or erecto-patent, almost parallel with the stem, oblong, plane, appressed to antical lobe, about as long but narrower than the stipules, ciliate-dentate. Texture somewhat thin, epidermis polished, rarely dull ; cells small, roundish, trigones distinct. Stipules appressed to stem, broadest at the base, slightly decurrent, ovate-rectangular, apex subtruncate, margin undulate, ciliate-dentate. Flowers $ produced on short branchlets at right angles to the branches ; sub-bracts somewhat similar to the leaves only larger and ciliate dentate ; sub-bracteole similar to stipules only larger; bracts patent- divergent, much larger than the leaves, antical lobe broadly ovate, rotundate, ciliolate ; postical lobe patent- divergent, broader than the leaf lobules, PURELLA. ^\ ciliolate ; bracteulc broadly ovate, ciliolatc. Perianth cuneate, mouth very wide, irreguhir, ciliohite, cilioUi beiiij^ about 4 cells long-. Pistillidia very large. Dlmensions. — Stems 'I to ;3 inches long, '4 mm. to '5 mm. in diameter, with leaves 3" mm. wide; antical lobe :^"5 mm. x I 75 ram., l2 mm. x I '5 mm., p»>stical M mm. x "4 mm., V^l'^> mm. x "5 mm., Lli mm. x o mm. ; branch leaves, antical lobe TG mm. x 1-25 mm., 175 mm. x I '1^5 mm., postical '7 mm. x "I'l mm. ; cells •02 mm., '0225 mm. x '02 mm., 0225 mm. x -0225 mm. ; stipules, r25 mm. x "75 mm., L* nun. x 'J mm.; sub-bract, antical lobe 2'4 mm. x I'o mm., postical 135 mm. x 7 mm. ; sub-bracteole I "25 mm. X 7 mm.; bract, antical lobe 'o mm. x 175 mm., i)Ostical 175 mm. xrmm. : bracteole 2*25 mm. x 2* mm.; perianths* mm. X 3'5 at the mouth; teeth at mouth of perianth "05 mm. -"075 mm. ; pistillidia "o mm. x 'i uim. Hab. — On trunks ot" trees (especially at the base), stones or rocks. Rare. 1 ? 5. Gt. Do ward Hill, Hereford, B. M. Walkiii'^. 7. Barmouth, ILoU and Wild. Tyn-y-groes, //: 11. P. 10. Nr. Dent, //^. litest. 12. Borrowdale, Cumberland, JJr. Carriuf/lu/t and W. II. P. 13. Frequent in subalpine glens, Dumiries, Cndckskank, Scott., 15. IG. Moidart, West Inverness, S^incris M. Macvicar. I. Found on the Continent. Obs. — Distinguished from other Pordlcc b}' its polished leaves, with antical lobes acute or acun)inate, postical ciliate dentate, margin of bracts finely ciliolate, perianth short, but very wide at the mouth. Plants bearing perianths have, I believe, only been once tbund, by Prof. Henriques in Portugal ; the male plant has never been met with. This plant has the singular character (which was first noticed by Prof. Lindberg), of producing a burning sensation in the mouth when chewed. Description of Pl.\te XXVIII. — Fig. 1. Plant natural size. 1. Portion of branch, antical view x 11 (C. k P. n. 45). 3. Portion of stem, postical view x IG (ditto). 4. Leaf, postical view X 1 I (ditto). 5. Postical lobe or lobule x IG (ditto). 6. Portion of leaf x 2'.)0 (ditto). 7. Stipule x 24 (ditto). F 82 BEPATICJ^.. 8. Sub-bract x 16 (Portugal, hb. S.O.L.). 9. Sub-bracteole x 16 (ditto). 10. Bract x 11 (ditto). 11. Bract and bracteole x 11 (ditto). 12. Perianth x 11 (ditto). 13. Portion of mouth of perianth x 85 (ditto). 2. Porella Thuja {Diclcs.), Moore. Lichemi,struin arhoris vitce facie, foliis rotundioribus, Dill. Hist. muse. p. 502, t. 72, f. 33 (1741). Jungermania Thuja, Dicks. PI. Crypt. Brit. Fasc. 4, p. 19(1801); Tayl. in Trans. Bot. Soc. Edin. xi. p. 116 (1841). Madotheca Thuja, Dum. Comm. p. Ill (1822), Porella Thuja (Dicks.), Moore in Proc. R. I. A. 2nd ser. vol. ii. p. G18 (187G). Dioicous, imbricately csespitose, largish to large, brown or olive-brown, apices of branches often pale yellow. Stem irregularly bipinnate, frontally compressed, cross-section showing about 100 cortical cells very small with thick dark walls, 25 x 30 cells in diameter, inner cells large and hyaline ; more or less robust, firm; radiculose, rootlets fasciculate, coarse, reddish- brown, produced from the base of the stipules. Leaves incubous, appressed, involute when dr\'", closely imbricate, horizontal, unequally bilobed, antical lobe much larger, convex, obliquely rotund-ovate, apex decurved, rotundate-obtuse, margin entire, or rarely with few distant teeth ; postical lobe or lobule hardly decurrent, very slightly attached at its base to the antical lobe, patent, ovate to ovate-oblong, a,pex obtuse or acute, sometimes uncinate, margin recurved, entire or rarely with few small teeth near the base ; a little longer and narrower than the stipules ; texture subopaque, firm but pellucid, epidermis slightly polished, verruculose, cells smallish 4-, 5- and 6-angled, roundish-quadrate, walls thick, trigones distinct. Stipules imbricate, appressed to the stem, decurrent, sub-reniform, sub-cordate or oval, broader than long on stem, longer than broad on branches, margin reflexed, entire or rarely with few teeth near the base. Female flowers on short lateral branchlets, only imperfect ones seen. Bracts larger than the leaves, antical lobe oblong-ovate or obliquely rotund- ovate, apex subacute, margin irregularly dentate, postical lobe r-ORELLA. «.S about 3 times smallor than the antical, oblong' or oblong-ovate, apex obtuse or subacute, margin irregularly- dentate ; bracteole broadly ovate, irregularly dentate. Pistillidia numerous, 10-22 long and narrow. Male stems pinnate, with few short branches androecia numerous on short lateral alternate catkins ; perigonial bracts G-8 pairs, closely imbricate, erect, bilobed, postical lobe small and oblong ; antheridia subspherical to oval, solitary. Dimensions. — Stem 2 to 3 inches long (var. forva 1 inch), diameter "4 to "5 mm., with leaves 2*25 mm. to 3' mm. wide ; leaves, antical lobe 1"8 mm. x 1\S mm., postical 1'2 mm. x '5 mm., antical 1 "5 mm. x 1"2 mm., postical "85 mm. x "5 mm., antical L"8 mm. x L"5 mm., postical 1' mm. x "6 mm., antical 2' mm. x 1*75 mm., postical 1*4 mm. x '75 mm., antical 1' mm. x "8 mm. postical "6 mm. x '4 mm. ; cells "03 mm. x '02 mm., '025 mm. x '025 ram., •025 mm. x -02 mm.; stipules 1' mm. x 1. mm., 1"3 mm. X "D mm., "7 mm. x 7 mm., '7 mm. x "8 mm., 1* mm. x •8 mm. ; bract, antical lobe 17 mm. x 1-2 mm., postical r35 mm. X 'G mm., antical 2*25 mm. x 2 mm., postical 1'5 mm. x L* mm. ; bracteole 1*25 mm. x 1'25 mm., 1*1 mm. x "8 mm. at base ; pistil- lidia "4 mm. X '075 mm. ; perigonial bract, antical lobe 1"5 mm. x 1* mm., postical '8 mm. x i mm. ; perigonial bracteole "75 mm. x '45 mm. ; antheridia 4* mm. x "35 mm. Har. — On rocks, stones or trunks of trees. Rare. 1. Var. forva, Penzance, W. Curnotv. 2. Var. torva, Sussex, 6'. Dcwles. 7. Ravine above Barmouth, Merioneth, Br. Carrhfj- ton. Near Snowdon, Carnarvon, Dr. Carrin/jfo/i. 12.? 16. Braemar, ])r. A. 0. Black. I. Cromaglovvn, Dr. Carri/if//on. By Lough Mangerton, Ifolf and Stewart. Found on the Continent (France, Monta(/nn. Oporto, Portugal, Neioton, S- Var. torva, Italy, de No tar is). North America. Obs. — Remarkable for its somewhat robust habit, dark brown to almost black colour, leaves being very closely imbricate, invo- lute when dry and slightly polished, and rounder than any other of the British species. Distinguished from /-'. j)lati/p/ii/f/a (L.) by its rounder loaves, 84 HEPATIC^^. more quadrate cells with large trigones, bracts very much more dentate or denticulate. From P. lavif/ata (Sclirad.) by its much less polished leaves, its darker brown colour (never greenish), antical lobe being never acute, postical entire and proportionally broader, stipules broader and entire, and by its not having the pungent taste of P. IcBvigata. The North American species P. platyphylloidea (Schweintz.), which has been confounded with it by some authors, is its nearest ally, but is of a lighter yellowish colour, with longer leaves and relatively smaller lobules and stijiules, and with bracts only slightly dentate. Var. torva (de Not.) Lindb. (Madotheca torva, de Not. MSS.) is a much smaller form with tlie stem irregularly branched, and the leaves very closely imbricate and polished. Description of Plate XXIX. — Fig. 1. Plant natural size. 2. Portion of branch, antical view x 11 (Killarney, Dr. Carrington)- 3. Ditto, postical view x 11 (ditto). 4. Leaf x 11 (ditto). 5, 6. Leaves x 16 (ditto). 7. Leaf and stipule x 16 (Mangerton, Holt and Stewart). 8. Leaf x 24 (France, Montague). 9. Lobule of leaf X 16 (Killarney, Dr. Carrington). 10. Portion of leaf x 290 (ditto). 11, 12. Stipules x 16 (ditto). 13. Bract x 24 (France, Montague). 14. Ditto x 11 (Mangerton, Holt and Stewart). 15. Bracteole x 11 (ditto). 16. Ditto x 24 (France, Montague). 17. Perigonial bract x 16 (Oporto, Newton). 18. Ditto, expla- nate x 16. 19. Perigonial bracteole x 16 (ditto). 20. Anthe- ridium x 24 (ditto). 3. Porella platyphylla (Z.), Lindb. Lichenastrwm arboris vitce facie, foliis minus 7-otundis, Dill. Hist. Muse. p. 501, t. 72, f. 32 (1741). Jungermania platyphylla, L. Sp. pi. ed. 2, 2, p. IGOO (1762) ; Hook. Brit. Jung. t. 40 (1816). Madotheca platyphylla, Dum. Couini. p. Ill (1822). Pordla 'platyphylla (//.), Lindb. Muse. Seand. p. 3 (1879). Dioicous, ca3spitose, large, in colour yellowish, brownisli or blackish green. Stems firm, ligneous, opaque, brownish or dull dark green, imbricate, procumbent or suberect, irregularly pinnate, POnELLA. 85 oval on cross-sGction, about 50 cells in circumference, which are small witli firm dark walls, l^tid and 3rd outer layers with dark walls, inner cells lari^er with hyaline walls, '1^) x \(} cells in diam. ; radiculose, rootlets few, proceeding- from the lower portion of the stem. Leaves incubous, closely imbricate, hilarious, distichous, alternate, horizontal, unequally bilobed, antical lobe much the largest, obliquely ovate or broadly ovate, slightly convex, near the base more or less concave, margin curvate and undulate, entire, but usually with a solitary tooth on the upper margin near the middle, a})ex rotundate-obtuse, slightly decurved ; postical lobe or lobule erect or erecto-patent, appressed to antical lobe, slightly decurrent, subobliquely ovate or oval, apex obtuse or subacute, margin recurved, entire, sometimes furnished with one or two small teeth at the base ; texture opaque, epidermis without gloss, cells smallish to medium in size, roundish, walls thin, trigones Very small but disHnct. Stipules approximate, appressed to the stem, decurrent, semi-rotund or oval, margin reflexed, entire. Female flowers produced on very short lateral branchlets at right angles to the branches. Bracts somewhat similar to the leaves but smaller, antical and postical lobes dentate, but the latter larger in proportion ; bracteole semi-obovate, entire or with few small teeth near the base on both sides. Perianth subovate, com- pressed, margin involute, composed of a single layer of cells, mouth truncate, inciso-serrate. Pistillidia oblong, large, 8-10. Calyptra spherical, carnose, somewhat firm, near the base several small cells thick, upper portion composed only of a single layer. Pedicel very short, thick, 15 large cells in diameter. Capsule only just protruding from the perianth, pale yellowish-brown, dividing not quite to the base into 4 valves, which are composed of one layer of cells. Spores smooth, very large, brown. Elaters with about 10 to ]'2 turns of the spii-al, bi-trispiral, much narrower than the spores, and lighter brown. Androecia on very short oval catkin- like lateral branches ; [)erigonial bracts 4, 5 pairs, closely imbricate, ver\' concave, semi-ovate, bilobed, lobes sometimes subequal, antical usually largest, margin slightly involute, entire; perigonial brac- teole ovate, entire or with few minute teeth near the base on both 86 EEPATICJE. sides. Antheridia large, solitary in the axils of the perigonial bracts. Dimensions. — Stems Ij to 3 inches long, '4 mm. diameter, with leaves 3" mm. wide; leaves, antical lobe 2* mm. x 175 mm., postical '8 mm. X '6 mm., antical 2*1 mm. x 7 mm., postical •75 mm. x '5 mm. ; cells "0325 mm. x '0275 mm., "03 mm. x •025 mm., '03 mm. x '0275 mm., ^03 mm. x -03 mm.; ^025 mm. x 025 mm.; stipules 1- mm. x 75 mm., l' mm. x ^85 mm. ; sub-bract, antical lobe 125 mm. x •55 mm., postical "75 mm. x ^35 mm.; bracts, antical lobe 1'6 mm. x 1*3 mm., postical ^75 mm. x '5 mm., antical 16 mm. X 1' mm., postical "8 ram. x ^6 nmi. ; bracteole I'l mm. x 1- mm. ; perianth 3^ mm. x 2*5 mm., teeth at the mouth '1 mm. longx •I mm. broad, at the base "075 mm. x '05 mm., ^05 mm. x •05 mm. ; pistillidia '225 mm. x "06 mm. ; valves of capsule 1*15 mm. X •G mm. ; spores ^055 mm. x '045 mm. ; elaters '2 mm. X '0125 mm. ; male catkin 2^ mm. x 1*25 mm. ; perigonial bract, antical lobe 1^1 mm. x "75 mm., postical '6 mm. x '5 mm. ; peri- gonial bracteole '5 mm. x '4 mm. ; antheridia ^35 mm. x ^3 mm. Hab. — In large patches on stones, rocks, and trees, often common, especially in limestone districts. Fruit somewhat rare- 1 to 3, 5 to 18. I. Found on the Continent and in North America. Obs. — This is the commonest of the British Por^Z/.^-, for distin- guishing characters of which see notes under the other species. It quickly discolours water upon immersion. Description of Plate XXX. — Fig. 1. Plant natural size (Engl. Bot.). 2. Portion of branch, antical view x 11 (C. and P. n. 71). 3. Portion of stem, postical view x 11 (ditto). 4-6. Leaves X 11 (ditto). 7. Lobule of leaf x 16 (ditto). 8. Portion of leaf X 290 (ditto). 9, 10. Stipules x 16 (ditto). 11. Sub-bract x24 (Sweden, S. 0. Lindberg). 12, 13. Bracts x 16 (ditto). 14. Bracteole x 16 (ditto). 15. Perianth x 11 (ditto). 16. Portion of the mouth of perianth x 85 (ditto). 17. Male catkin x 16 (ditto). 18. Perigonial bract X 16 (ditto). 19. Perigonial bract- eole X 24 (ditto). 20. Antheridium x 24 (ditto). PORELLA. 87 4. Porella rivularis {Nees), Lindb. Madotheca rivularis, Nees, Ntit. Eur. Leb. 3, p. lt)(5 (1838). Porella rivularis (Nees), Lindb. Muse. Scand. p. 3 (1S7!)). Dioicous, depresso-csespitose, medium to largish in size, varying in colour from pale to dark olive or brownish-green, when dry often yellowish-green. Stems irregularly pinnate or sub- dichotomous, apices often subfastigiate, somewhat firm, frontally compressed, 1 5 x 20 cells in diameter; cortical cells very small 1-3, outer rows reddish-brown, inner much larger, hyaline. Leaves approximate, slightly appressed, incubous, horizontal or horizontal- patent-divergent, unequally bilobed, antical lobe much the larger, almost plane, obliquely rotund-ovate, apex slightly decurved, margin distantly denticulate or subentire, usually with 1-3 large teeth at the antical base ; postical lobe or lobule erect or erecto- patent, contorted, much decurrent, obliquely ovate or ovate- lanceolate, acute, sometimes uncinate, margin recurved, undulate, repand, subentire or denticulate, often with lew large teeth near the base ; texture somewhat thin but firm, epidermis dull, rarely with a slight polish, cells medium size, roundish, walls thin, trigones very distinct. Stipules appressed to stem or slightly recurved, very decurrent, quadrate- oval, apex rotundate, margin reflexed, undulate, repand, subentire or dentate, especially at the base. Flowers ? produced on short lateral branches, nearer the postical than the antical side of stem ; bracts about 3 times smaller than the leaves, unequally bilobed, antical lobe oval, apex acute, margin entire, postical lobe aljout half the size, quadrate-oval, margin quite entire ; bracteole small ovate-quadrate, retuse, margin quite entire. Perianth broadly oval, complanate, mouth small, sub- entire. Pistillidla about 8. Androscia on short lateral branches, small ; perigonial bracts closely imbricate, 4, 5 pairs, unequally bilobed, complicate-concave, antical lobe oval-oblong, margin entire, postical lobe half the size, oblong-quadrate, margin (|uite entire. Antheridia oval. Dimensions. — Stems U- to 2 inches long, -4 mm. to '5 mm. in diameter, witli leaves 4- mm. wide; leaves, antical lobe 225 mm. 88 HEFATJCJE. X 1*5 mm., 2- mm. x I'G mm., 2'1 mm. x I'G mm., I'S mm. x 1*7 mm., VI mm. x 1*4 mm., 1"75 mm. x 1'5 mm., postical •75 mm. x '4 mm., "7 mm. x '35 mm., "9 mm. x '5 mm. ; cells •035 mm. x -03 mm., -035 mm. x '025 mm., '035 mm. x '035 mm. ; trigones '0017 mm.; stipules "75 mm. x -6 mm., "75 mm. x •75 mm. ; bract, antical lobe -8 mm. x '4 mm., postical '5 mm. x 25 mm. ; bracteole 7 mm. x -45 mm. ; perianth 4*5 mm. x 1-75 mm.; pistillidia '3 mm. x 075 mm.; capsule 1"2 mm. x -9 mm. ; peri- gonial bracts, antical lobe I'l mm. x '6 mm., postical '85 mm. x •4 mm., antical 1- mm. x '55 mm., postical "55 mm. x 3 mm.; perigonial bracteole '5 mm. x 3 mm.; antheridia "14 ram. x "11 mm. Hab. — On stones in rivers, or on rocks and trees by their side. Somewhat rare, and extremely so in fruit. 1? 7? 10. On limestone rocks at edge of stream, Malham, W. JFe^f. On rocks near Collingham, W. Yorks, IF. West. 12. Staveley, Westmorland, G. Sf abler. High Lorton, Cumber- land, Bev. C. H. Bivsfead. Ullswater, Cumberland, E. M. Holmes. 13. Near Kenmure Castle, New Galloway, ./. Mc Andrew. Balmac- lennan, Dumfries, ./. McAndrew. On trees and rocks, Newtondon, Berwickshire, A. Brollierston. I. On trees near River Lagan, Co. Down, Bev. C. H. Waddell. Continent. Obs.— Differs from B. plaiyphylla (L.) in its looser habit, more distant leaves, planer, rather more dentate, smaller, narrower nnd more contorted lobule, thinner texture, larger and distinct trigones, smaller and more irregular stipules, smaller bracts, margin entire, perianth with contracted mouth. Borella pmnafa (D.) is a much more slender species, with more distant leaves, much smaller lobules and stipules, margins of which are all quite entire. Description op Plate XXXI. — Fig. 1. Plant natural size. 2. Portion of stem, antical view X 1 1 (C. and P. n.). 3. Ditto, postical view X 1 1 (ditto). 4,5. Leaves x IG (ditto). G. Lobule X 24 (ditto). 7. Portion of leaf x 290 (ditto). 8. Stipule x 24 (ditto). 9. Bract X 24 (Sweden, S. O. Lindberg). 10. Bracteole POnELLA. 89 X 24 (ditto). 11. Perianth x 11 (ditto). 12. Portion of mouth of perianth x 04 (ditto). 13, 14. Perio'onial bracts x 24 (ditto). 15. Perigonial bracteole x 24 (ditto). J (5. Antheridium x 85 (ditto). 5. Porella pinnata (Z) ///.), Lindh. Jungermania Porella, Dicks. Trans. Linn. Soc. iii. p. 2)V^ (1797), Jungermania distans, Schwein. Muse. Hep. Amer. p. 9, n. "> (1S21). Jimgermania Cordeana, Huhen. Hep. Germ. p. 291 (18P)4). Madotheca Porella, Nees, Nat. Eur. Leb. ?> p. 201 (I808). Porella pinnata (Dill.) Lindb. Hep. Hib. Acta Soc. Sc. fenn. x. p. -lOB (1875). Dioicous, laxly c?espitose, yellowish-brown to dark olive-green in colour. Stems irref^ularly pinnate or subdichotornous, frontally compressed, from 12 x 8 to 20 x 10 cells in diameter, cortical cells very minute with thick walls and reddish-brown 50-80, the 2nd and 3rd outer layers usually tlie same, inner large, walls hyaline. Leaves incubous, approximate or distant, unequally bilobed, antical lobe plane or very slightly convex, ovate-oblong, apices plane or indistinctly decurved, rotundate or obtuse, margin entire, postical lobe or lobule minute, plane or slightly undulate, a little decurrent, erecto-patent, oblong, obtuse or obtusate, margin entire ; texture delicate, thin, cells smallish, 4-, 5- and 6-angled, walls thin, trigones absent or very minute, marginal cells with slightly thicker walls. Stipules appressed to the stem, ovate-rectangular, rotundate-obtuse, plane, margin entire. Flowers ? on long or short lateral branches ; bracts smaller than the stem leaves, unequally bilobed, somewhat carinate, antical lobe oval, postical from ^ to ^ smaller ; bracteole oval, apex rotundate. Perianth (young) compressed, margins decurved, convex antically, broadly ovate, mouth contracted, dentate. Pistillidia 6-8. Cap- sule dividing to the base into 4 valves, bistratose, the outer layer 3, 4 times thicker than the inner. DiMKNSiONS. — Stem 1 to 2 inches long, "15 mm. to '225 mm. in diameter, with leaves 3' to 4" mm. wide; leaves, antical lobe re mm. X r mm., 1-5 mm. x T mm., 2' mm. x 1*5 mm., 1-3 mm. X '8 mm., postical 'G mm. x "2 mm., 'bb mm. x "125 mm., '7 mm. x •3 mm., "8 mm. x '35 mm., "45 mm. x "2 mm., '35 mm. x -175 mm. ; 90 HEPATICJE. cells -02 mm. x '025 mm., '025 mm. x '025 mm., -03 mm. x •02 mm., '04 mm. x "02 mm. ; stipules '8 mm. x '6 mm., '4 mm. x •15 mm., -4 mm. x '25 mm., *35 mm. x '25 mm., 7 mm. x '4 mm. ; perianth 2' mm. x 1'25 mm. Hab. — On damp shaded rocks or stones, very rare. 1. Eiver side, Fingal Bridge, Devonshire, JF. Curnow. 1. South of Ireland, Br. Taylor, Br. Moore, A. Carroll, &c. Lough Mangerton, Stewart and Holt. Found sparingl}^ on the Continent (France), also in North America. Obs. — This is the smallest and most delicate of the British Porellcs, and is nearest related to P. rivularis, which see. The description of the fertile plant is taken from North American specimens by Dr. Gottsche, sterile plants only having been found in Europe. Description of Plate XXXII.^ — Fig. 1. Plant natural size, 2. Portion of stem, antical view x 16 (C. and P. n. 132). 3. Ditto, postical view x 11 (ditto). 4, 5. Lobules x 24 (Mangerton, Holt and Stewart). 6. Portion of stem, witli lobules and stipules x 31 (C. and P. n. 132). 7. Lobule x 64 (ditto). 8. Portion of leaf X 290. 9, 10. Stipules X 24 (Mangerton, Holt and Stewart). 11. Stipule X 31 (ditto). 12. Ditto x 64 (C. and P. n. 132). Genus 6. PLEUROZIA, Bum. Jungermania, Weiss, PI. Crypt. Fl. Gott. p. 123 (1770). Pleurozia, Dum. Recueil, i. p. 15 (1835). Physiotiuin, Nees, Nat. Eur. Leb. 3, pp. 6 and 75 (1838). Involucral bracts two, deeply bilobed, the upper broad and convolute. Perianth much exserted, long, cylindrical, mouth denticulate, decurved and plicate at the apex. Capsule 4-valved, coriaceous. Elaters bispiral, deciduous. Stipules absent. Leaves with postical lobes auriculate, the auricles inflated. PLEUBOZIA. 91 Pleurozia cochleariformis ( Weiss), Dim. Lichenastrum alpmum purpureum foliis cmriiis cochleariformibus, Dill. Hist. muse, p. 479, t. G9, f. 1 (1741). Mnium Jungermania, Linn. Sp. pi. p. 1579 (175;')). Jimgermania cochleariformis, Weiss, PI. Crypt, p. 128 (177U); Sm. Engl. Bot. t. 2500 (1813); Hook. Brit. Jung. t. G8 (181G). Jungermania purpiorea, Scop. Fl. Carn. 2, p. 347 (1772). Pleurozia cochleariformis, Dum. Syll. Jung. Eur. p. 38 (1831). Physiotium cochleariforme, Nees, Nat. Eur. Leb. Ill, p. 79 (1838). Pleurozia purpurea, Lindb. Muse. Scand. p. 3 (1879). Dioicous, loosely or closely cfespitose, large, pale to dark reddish -brown or purple in colour. Stems creeping or ascending, arcuate, simple or with 1-3 long or short branches, frontally compressed, 15 cells in diameter, cortical, with second and third outer layers of cells smaller, reddish-brown, inner whitish, opaque, walls thick. Leaves incubous, closely imbricate, distichous, convex, bilobed, antical lobe ovato-rotundate, emarginate to about a fourth, segments dentate, inliexed, near antical base furnished with 1-3 teeth, postical lobe about \ smaller, attached to the lower portion of the antical lobe, ovate-parabolic, utriculate, base contracted, decurrent, on the inner side deeply depressed and folded longitudinally. At the base of this furrow is a minute slit with a pendant roundish lip, which is slightly concave ; upon this is appressed a very delicate lid of somewhat similar shape but plane, of very delicate cell structure, hanging from the other side of the slit, both these appendages being suspended within the lobule. Texture of leaf firm, epidermis verruculose, cells small to smallish, roundish-hexagonal, trigones very large, often hyaline, lumen deep red or purple, the lip within the lobule is composed of very delicate cells, thin walls, no trigones, about 20 X 30, marginal cells subquadrate, still more delicate, about 50 to GO round. No stipules. Androecia catkin-like, minute, solitary, produced from the axil of the antical lobe, subcylindri- cal ; perigonial bracts G-8, closely imbricate, appressed, concave, ovate or oval, bifid or tridentate ; antheridia oval, solitary. 92 HEFATICJa. Dimensions.— Stems 2 to 4 inches long, '5 mm. x "4 mm. in diam., with leaves 3* mm. wide; leaves, antical lobe 2*5 mm. X ] '5 mm. -1-8 mm.; segments '75 mm., "5 mm., lobule 1"25 mm. -1*5 mm. x '5 mm. -'8 mm., lip of mouth '2 mm. x '2 mm., lid of mouth '25 mm. x "225 mm. ; rells '02 mm. x -0225 mm. "02 mm. X "02 mm., "03 mm. x '02 mm., -03 mm. x "03 mm. ; male spike '75 mm. -1-2 mm. x "4 mm. -'5 mm. ; perigonial bracts •5 mm. X "2 mm., seg. '1 mm., "5 mm. x "5 mm,, seg. 1 mm., •5 mm. X "4 mm., seg. *3 mm., 5 mm. x "5 mm., seg. '2 mm. ; antheridium '2 mm. -'15 mm. x '1 mm. Hab. — In large patches on damp moors in subalpine localities. Rare and local, but where found usually abundant. 13. Black Craig, New Galloway, J. McAndrew. Auchencairn Moss, Dum- fries, C. Scott. 15. Ben Nevis, /. B. Byrom. Ben Voirlich, Gourlie. 16. Glen Finnan, Dr. Carrim/ton. Moidart, S. M. Mac- vicar and W. H. B. Skye, Brof. F. E. Weiss. 17 a. Cape Wrath, Sutherland. I. Mountainous bogs. Near Killarney, Dr. Carritu/- ton. Connor Hill, Brof. S. 0. Lindber//. Mangerton, S. 0. Lind- hercj, Steioart and Holt, male plants. Found on the Continent (Norway, Denmark), Sandwich Islands, Ht). Hoolcer. East Indies, Butam proper, Jowga, 10,000 ft., Hb. Mitten. Obs. — In the lobule of this species there is one of the most remarkable contrivances for the entrapping of small insects it is possible to imagine : on the inner, antical face of the utriculate lobule is a deep longitudinal fold having at the base a slit with two decurrent wings hanging free within, one slightly concave, the other plane — the unwary insect crawling through this slit, the lid opens with the slight pressure and closes after the prisoner has been secured; it is no unfrequent thing to find the chitinous remains of several of these insects within the lobule. It has not yet been clearly shown whether the plant derives nourishment by the assimilation of these animals, but we may rest assured that some object is served by this apparatus, and the plant may not inaptly be called an insectivorous hepatic. The late Dr. Gottsche was, I believe, the first to publish any notes o^ ANTItELlA. 93 this cliciracter ; tterrii Jack and Stepliani have also written on it. This species is a very distinct one, and the only British repre- sentative of the genus. Hooker says, "Weiss, who first adopted the name, describes the species with considerable accuracy," with which opinion I ag^ree. Nees and Jack seem to think that he had a form of Marsupella emart/lnata before him, although Nees has seen speci- mens of the true plant from Weiss's herbarium. Scopoli's Jiiiujeniiania purpurea of 1772 may be the same as our species, but it is very uncertain. It may, however, be said that the description oK J mifjermaiiia purpurea in Lightfoot's "Flora Scotica " (1777) is very inaccurate and vague if applied to this species. Description of Plate XXXIII. — Fig. 1. Stem natural size (Eng. Bot.). 2. Leaf with lobule x 16 (Cromaglown, Stewart and Holt). 3. Ditto x 24 (ditto). 4. Lobule x 24 (ditto). 5. Ditto X ? (drawn by Herr Stephani, "Rev. Bryol," 1886). 6. Lip of inner mouth of lobule x 24 (Cromaglown, S. and H.). 7. Lid of the same x 24 (ditto). 8. Cross-section of lobule near the apex x 24 (ditto). 9. Portion of leaf x 200. 10. Male spike X 24 (ditto). 11-15. Perigonial bracts x 24 (ditto). 16. Antheridium x 85 (ditto). Subtiibe III. PTILIDIE^. Genus 7. ANTHELIA, Dam. Jungermojuia L. Fl. Lapp. (1787). Jimgermanice, sect. Anthelia Dum. 8yll. Jung. p. 63 (1831). Anthelia, Dum. Recueil, p. 18 (1835). Chandonanthus, Mitt, in Hook. f. Handb. New Zeal. Fl. 11, p. 750 (18(17). Plants small or somewhat robust, densely cuespitose, green or olive glaucescent. Stems firm, composed of several layers ot equal, opaque cells ; unequally pinnate ; all branches lateral, at the base sometimes denudate or furnished with minute leaves ; Hagella absent; radicles moderately plentiful in the young state, when 94 HEPATIC j^. mature somewhat rarer. Leaves tristichous transverse, somewhat broad, complicate-carinate, where large confertus equitant, bilobed to the middle or beyond, lobes subacuminate, entire or often (principally the upper) denticulate or spinulose — sometimes the surface spinose-muricate ; cells small pellucid quadrate-hexagonal, at the axis subelongate, walls more or less thickened. Postical leaves (stipules) similar to the lateral and hardly any smaller. Inflorescence dioicous, rarely paroicous, terminal ; ? innovations often present. Bracts several pairs, more or less crowded capitate, distinctly larger than the leaves ; in other characters hardly diflerent, except sometimes trilobed, with the margin coarsely dentate, rarely connate. Perianth free emersed oblong, at the base 2 cells thick, above delicate, frontally subcompressed, antically deeply unisulcate, postical 3 carinate, at the apex 10-8 -plicate ; mouth either wide or subconstricted, denticulate. 1. Anthelia julacea (Z.), Bum.. Lichenastrum alpinum, Bryi julocei argentei facie, Dill. Hist, Muse. t. 73, f. 38 (1741). Jungermania julacea, Linn. Sp. P). 11, p. 1601 (1753); Lightf. Fl. Scot. p. 785 (1777) ; Hook. Brit. Jung. t. 2 (1816). Anthelia julacea, Dum. Recueil, p. 18 (1835); Spruce, On Ceph. p. 81 (1882). Dioicous, densely and intricately cpespitose, flagella wanting, from small to largish in size, brownish or olive-green in colour, glaucescent when dry, often infested with slimy or confervoid matter. Stems erect or procumbent, filiform, firm, composed of equal opaque cells, about 30 cortical, 10 to 12 cells in diam, when dry rigid, unequally pinnate, branches all lateral ; radiculose, in the young state rootlets abundant, when mature somewhat rarer. Leaves tristichous, transversely inserted, appressed, erect, closely imbricate or rarely distant, complicate-carinate, concave, oblong, to about two-thirds bilobed, sinus acute, narrow, segments ovate- lanceolate, acute or subacaminate, entire or rarely erose-denticulate, recurved at both sides ; texture firm, epidermis smooth, cells rather minute to small, subquadrate, 4-, 5-, and 6-angled, near the ANTIIELIA. 95 base, at the middle a little elongate, walls thick, no trigones or thickened angles, composed of two layers of cells at the base, above one layer, upper portion often hyaline. The third postical leaf (stipule) similar to the others, sometimes slightly smaller. Bracts several pairs, more or less crowded, capitate, larger than the leaves, 2 to 3 cells thick near the base, oblong-oval, bilobed to about the middle, sinus narrow, acute, segments ovate-lanceolate, acute or acuminate, margin denticulate, or entire. Bracteole smaller than the bracts, bilobed to about the middle, erose- denticulate. Perianth composed of two layers of cells at the base, one layer and delicate above, near the middle about 100 cells round, free, terminal, projecting about half beyond the bracts, oblong-oval, upper half multi- and deeply plicate, frontally sub- compressed, antically deeply unisulcate, at both sides of the furrow the upper margin carinate, postical 3-(2) carinate, at the apex several small keels added (10-8-plicate), mouth wide or sub- constricted by the folds, laciniate, lacinise denticulate. Calyptra ovoid-globose, 2 to 3 cells thick at the base, pistillidia few (5 or 6) dispersed on the upper portion of the calyptra. Capsule sub- globose, composed of two layers, small, dark brown. Spores pale brown, elaters bispiral about as broad or a little broader than the spores. Androecia at the apex or middle of the stem, on separate ones from the ? although often entangled with them ; perigonial bracts 4-6 pairs, somewhat similar to the leaves only a little broader, sometimes slightly denticulate, closely imbricate, sub- secund, ventricose at the base ; antheridia large, single, oval, bearer short. Dimensions. — Stems from |^ to 2 inches long (rarely more), diameter "2 mm., with leaves "4 mm. wide ; leaves "5 mm. x "2 mm., segments "3 mm., "55 mm. x '3 mm., segments '3 mm., "4 mm. X '25 mm., segments "3 mm.; cells '02 mm., "02 mm., '015 mm. X -01 mm., -025 mm. x '015 mm., -02 mm. x "Ol mm., '02 mm. x •01 mm., -015 mm. x -015 mm. ; sub-bract '7 mm. x •45 mm., segments ^4 mm. ; bracts -9 mm. x '65 mm., segments •G mm., •8 mm. X '5 mm., segments • 5 mm., '75 mm x •65 mm., segments &6 B^PATIC^. •5 mm., 1* mm. X "6 mm., segments "5 mm., l"! mm. x 6 mm., seg- ments '5 mm. ; bracteole '65 mm. x '4 mm., segments "3 mm., "6 mm. X "5 mm., segments '25 mm. ; perianth 2' mm. x 1* mm., 1-75 mm- X 75 mm., segments '25 mm., '4 mm. ; capsule '5 mm. x 'b mm., valves '6 mm. x '3 mm., pedicel 4* mm. x "3 mm. ; spores '015 mm- diameter, elaters *15 mm. x '01 mm. ; perigonial bracts -6 mm. x •4 mm., segments '3 mm. Hab. — Grrowing on moist exposed or shaded rocks in large cushion-like tufts in alpine or subalpine localities. Moderately common, somewhat rare in fruit. 1, 7, 10 to 13, 15, 17. I. Found on the Continent and in Greenland. Obs. — This is one of the commonest hepatics met with, when a certain altitude is attained, often growing in large densely tufted cushions of a silvery lustre ; the stems are so intricately entangled that they are difficult to separate. Distinguished at a glance from Cesia concmnata (Dicks.) and Cesia obtusa (Lindb.) by growing in damper situations and the larger size of tufts ; stems longer, more rigid and filiform ; more closely examined, leaves tristichous, narrower, deepty bilobed, seg- ments acuminate, presence of true and deeply plicate perianth. From Anthelia Jiiratzkana (Limpr.) see following notes. In " Eng. Bot. " 1024 the leaves of this species are figured wrongly as if entire, and in Hooker's " Brit. Jung. " the bracts are described and figured as quadrifid, which is also incorrect. Description of Plate XXXIV. — Fig. 1, Plants natural size. 2. Plant X 11 (Lyn Cwm, W. Wilson). 3-7. Leaves x 24 (Mangerton, Stewart and Holt). 8. Branch leaf x 85 (Meal na Ptargnachan, C. J. Wild). 9. Portion of leaf x 290 (Mangerton, Stewart and Holt). 10. Sub-bract x 24 (Lyn Cwm, Wilson). 11-13. Bracts x 24 (ditto). 14-16. Ditto x 24 (Mangerton, Stewart and Holt). 17, 18. Bracteoles x 24 (ditto). 19. Perianth x 16 (ditto). 20. Portion of the mouth of perianth x 85 (Lyn Cwm, Wilson). 21, 22. Perigonial bracts x 24 (ditto). ANTUELIA. 97 2. Anthelia Juratzkana [Liwpr.), Spruce. Janyermania nivalis, Ssv. in W. M. Ind. iiiu'^c. \>. ."i (Iho;;); Wahlcnb. Fl. Carp. p. :5G:i(lHl !)(?). Jungennauia julacea var. clavuligei'n, Ncos, Nat. Eur. Lcb. 11, p. :507 (1830). Jumjeiinania jvlacea var. t/laucescens, (r.L.N. Syn. Hep. p. 147 (1841), JuTigei'vmnia Jnratzhtna, Linipr, in Kryptogainend. von Schles. 1, p. 289 (187C). Anthelia nivalis ('Sw.), Lindb. Muse. Scand. p. (1H7'.>). Anthelia Juratzkana, S[>ruce, On Ceph. p. H2 (iSSi'). Paroicous, depresso-caespitose, small, pale in colour, apices green or glaucescent, brownish below. Stems creeping, closely entangled, subramose, branches lew ; radiculose, rootlets plentiful, short; on a cross-section 10 cells in diameter; cells hirge, clear, about 40 cortical cells, similar to inner, only slightly darker. Leaves closely imbricate, on sterile shoots distant, tristichous, erect, appressed to stem or secund, oblong-ovate or oblong, con- cave-complicate, bilobed to about {J, sinus acute or slightly rounded, segments nari'owly triangular, or broadly subulate, acute or acuminate, margin entire ; composed of one layer of cells ; texture somewhat lax, cells medium size, quadrate or oblong- quadrate, pellucid, walls thin, no trigones or thickened angles. Stipule (third leaf) similar to the others. Sub-bracts and bracts larger than the leaves, a little broader and swollen at the base, margin denticulate, containing single oval antheridia, rarely the bracts arc trifid. Perianth projecting about ^ beyond the bracts, ovate-oblong or oval, deeply multiplicate, especially at the upper portion, mouth laciniate, lacinia) entire or denticulate ; composed of two layers of cells at the base, above of a single layer. Calyi)tra composed of one layer : pistillidia 3-5. Capsule subspherical ; spores and elaters reddish-brown. Dimensions. — Stems J- to -i- inch long "1 mm. to "3 mm. in diameter, with leaves '5 mm. wide ; leaves "3 mm. x -2 mm., segments "2 mm., ;) mm. x "15 mm., seg. "2 mm., •3.") mm. x 2 mm., see:. '2 mm. ; cells .035 mm. x "02 mm., '04 mm. x '03 mm., •03 mm. x (13 mm. ; sub-bracts '5 mm. x '25 mm., segments •225 mm., 35 mm., '25 mm., seg. "2 mm., "45 mm. x 3 mm., seg. •3 mm. ; bracts 75 mm. x '5 mm., segments "5 mm., D mm. x 98 HEPATICjE. •7 mm., seg. '45 mm., 1. ram. x "6 mm., seg. "4 mm. ; bracteole •7 mm. X '5 mm., segments "55 mm. ; perianth 15 mm. x '7o mm.; lacinise at the mouth '2 mm., "25 mm. ; pedicel "2 mm., "3 mm. diam. ; capsule "45 mm. x '4 mm. ; valves "65 mm. x 'SS mm. ; spores '015 mm. ; elaters *15 mm. x '01 mm. ; antheridia '125 mm. X '1 mm., bearer "1 mm. long. Hab. — On moist rocks in alpine situations. Very rare. 15. Below the summit of Ben Lawers, W. West, August 1880. Ben Laoigh, Peter Eioing, September 1882. Glen Callater, Prof. Balfour, 1841. Found on the Continent. Obs. — Distinguished from A. julacea (L.) by its smaller size, paroicous inflorescence, the antheridia being found at the base of the bracts immediately below the perianth, the somewhat secund leaves, which are much laxer in texture, and the perianth being less cylindrical, with the folds not so deep at the lower portion. I detected this species first as British from specimens collected by Mr. West on Ben Lawers in 1880, who had named it Anthelia julacea. In Dr. Carrington's herbarium, specimens collected in (Hen Callater by Prof Balfour in 1841 and named Anthelia julacea var. clavuligera Nees, belong to this species. Prof. Lindberg refers the Juvg. 7iivalis of Swartz (./. nivalis Sw., W. M. Ind. muse, p. 5. (1803); Whalenb. Fl. Suec, pro parte) to this species, but on this matter Dr. Spruce writes : " However that may be, there is no evidence to show that either Swartz or Wahlenberg discriminated between the two forms, and did not equally include the /. julacea of Lightfoot, as well as ./. Juratzkana, under their name, /. nivalis ; so that to Limpricht belongs the honour of first distinguishing /. Juratzkana by its inflorescence." Description of Plate XXXV. — Fig. 1. Plants natural size. 2. Fertile plant x 11. 3, 4. Leaves x 85. 5. Portion of leaf x290. 6. Sub-bract x 85. 7, 8. Sub-bracts, ^, x 85. 9. Sub-bract x 24. 10. Antlieridium x 85. 11-13. Bracts x 24. 14. Bracteole (?) x 24. 15. Segment of bract x 85. 16 Peri- HERBERT A. 99 antli X 24. 17- Portion of the mouth of the perianth x 85. (All drawn from plants collected by Mr. West, Ben Lawers.). Genus 8. HERBERT A, Gr. 8f B. Herherta, Gr. & B. Nat. Arr. Brit. PI. 1, p. 705 (1821). Schisma, Dum. Comm. p. 114 (1822). Sendtnera, Nees in G.L.N. Syn. Hep. p. 288 (1845). Dioicous ; large ; reddish-brown ; slightly branched ; branches postical ; subfloral innovations sometimes lateral. Leaves incubous or almost transverse, tristichous, hamato-secund, narrow, much longer than broad, sometimes ciliate at the base, otherwise entire, deeply bilobed, lobes narrow, acuminate, vittate. ? flowers terminal. Bracts denticulate, ciliate or spinose, distinctly bisul- cate. Perianth almost hidden by the bracts ; very narrow, ovate- subulate, tricarinate. Calyptra free, small, obovate, deeply trifid, composed of one layer of cells. Capsule globose, 5-8 cells thick ; valves 4 sometimes 2-6. Elaters bispiral, long, persistent. AndrcBcia on short terminal spikes ; perigonial bracts few, some- what similar to the leaves. Antheridia 2 in each bract. 1. Herberta adunca {Dicks.), Gr. ^ B. Jungermania adunca, Dicks. PI. crypt. Brit. fasc. Ill, p. 12 (1793). Jungermania junijierina b. adunca, Hook. Brit. Jung. t. 4 (181G). Herherta adunca, Gr. & B. Nat. Arr. Brit. PI. p. 705 (1821). Schisnut adunca, Dum. Comm. p. IIG (1822). Sendtnera junijoerina var., Nees in G.L.N. Syn. Hep. p. 239 (1845). Sendtnera adunca, Carr. in Trans. Bot. Soc. Edin. 11, p. 454 (1803). Dioicous, densely csespitose, from medium to very large in size, reddish or dark brown in colour. Stems erect, flexuose simple or with one or two short assurgent or pendulous postical branches, flagella frequent, bearing minute leaves, branches and apices of stem very often hamate, firm, opaque, the outer layers of cells (to the 2nd or 3rd) reddish-brown, inner paler, full of dark green colouring matter, cells small, walls ver}^ thick, cortical cells about 50; about 16 in diameter. Leaves incubous or almost transverse, imbricate or approximate, tristichous, falcate-secund, 100 HEPATICJE. vittate, linear-lanceolate, narrow, about 3 times longer than broad, to the middle or below acutely bilobed, lobes straight or divergent, postical usually a little smaller, acuminate, entire, or rarely with one or two small teeth at the base, margin often revolute, leaf and segments channelled ; texture firm, epidermis rough, marginal cells small, quadrate, pachydermous, middle elongate, about twice as long as broad, margin of cells irregular, walls thick, angles thickened, near the base the elongate cells contain very delicate spiral filaments. ? terminal, capitate, bracts several pairs, closely imbricate, attached at their base to the perianth, ovate-lanceolate, margin denticulate, divided to about the middle, sinus acute, segments lanceolate, acuminate. Perianth projecting about a third beyond the bracts, ovate-oblong, strongly plicate, composed of one layer of cells, mouth laciniate, almost to the middle, laciniae 8, entire or sparsely denticulate. Calyptra small, orbicular, uni- cellular, only somewhat thick near the base ; pistillidia about 20. Capsule thick, composed of about 6 layers, inner layer with numerous long, very narrow elaters attached to it. Spores large, pale brown, elaters bispiral, reddish-brown, darker colour and much shorter than those attached to the inner layer of capsule, about half the breadth of the spores or less. Androecia terminal, spicate, perigonial bracts about 12 pairs, closelj^ imbricate, erect, ventricose at the base, oval-lanceolate, margin sparsely denticulate, bilobed to below the middle, segments subulate-acuminate, anthe- ridia solitary, large, oval, bearers long. Dimensions.— Stems 1 to 4 inches long, sometimes shorter and even longer, diam. *2 mm. "3 mm., with leaves 2" mm. wide; leaves 1*25 mm. x "4 mm., segments '75 mm., 2' mm.x "6 mm. seg. 1'2 mm., 2' mm. x 7 mm., seg. 1*25 mm., 1-6 mm. x "7 mm. seg. 1" mm., •75 mm. x '3 mm. seg. '4 mm.; cells, middle of leaf '05 mm. X '02 mm., "04 mm. x "02 mm., '035 mm. x "02 mm., marginal cells -02 mm. x '02 mm., *015 mm. x "015 mm.; bracts 4" mm. X 1*5 mm., segments 2- mm. ; perianth 6" mm. x 1*5 mm. ; calyptra 1'5 mm. x 1" mm. ; pistillidia "3 mm. x '05 mm. ; valve of capsule 2*25 mm. x 1' mm.; spores "03 mm. ; ela.ters free '1 mm. x "01 mm., '15 mm. x "01 mm., attached elaters "3 mm. long; HERBERT A. 101 perigonial bract 2* mm. x 75 mm,, segments 1"25 mm. ; antheridia •275 mm. x "175 ram., bearer -275 mm. x -03 mm. Hab. — Grows in dense tufts on exposed banks or crags in alpine or subalpine situations. Moderately rare, but often grow- ing in great abundance where found. Extremely rare in fruit. 7, 12, 15, IG. L Found on tlie Continent and in North America. The large form H. junipenna (Sw.) in the West Indies, South America, Africa, Madagascar, Java, &c. Obs.— This is a very distinct species and not likely to be con- founded with any other British hepatic. It varies extremely in size from an inch or so to 5 and even 6, and in colour from a reddish to a dark brown. Herberta j unijjerina (Sw.) is usually a more robust plant with larger leaves, broader at the base, often more ventricose, margin at the base distinctly dentate ; Hooker may probably be right in considering //. adimca only a variety of it. I have never seen male or female of //. adunca, so have described and figured those of H. juniperina from Cuban specimens, which I have little doubt will be similar or nearly so to those of H. adunca. llerherta Sauteriana (Hiiben.) {Ilerberta straminea, Dum.) has been reported from Scotland, but all the specimens I have seen under this name are to be referred to H. adimca. Herherta Sauteriana (Hllben. ), which is found on the Continent, has a different habit, leaves close, shorter and broader, coarsely dentate at the base, with segments shorter and broader. Description of Plate XXXVI. — Fig. 1. Plants nat. size. 2. Portion of stem x 24 (Connor Hill, Ireland, Prof Lindberg). 3. Leaf x 24 (III Bell, Troutbeck, Westmorland, C. Stabler). 4. Ditto X 24 (Connor Hill, Prof. Lindberg). 5. Ditto x 24 (Ben Laiogh, Scotland, Gr. A. Holt). 6. Ditto x 24 (Ben Eagh, Bosshire, Jenner & Howie). 7, 8. Leaves x 24 (Clogwyn-d'ur- Arddu, Snowdon, G-. A. Holt). 9. Portion of leaf, middle x 290 (Connor Hill, Prof Lindberg). 10. Ditto, margin x 290 (ditto). 11. Bract X 11 (Cuba, Wright). 12. Periantli x 11 (ditto). 13. Perigonial bract x IG (ditto). 102 HEPATIC^. Genus. 9. MASTIGOPHORA, Nees. Jungermania, Brid. Mss. Web.-F. Inst. muse. hep. prodr. p. 5G, n. 49, (1815); Hook. Brit. Juug. p. 18, n. 64 (1816). Jungermania, sect. 2, Blepharozia, Dum. Syll. Jung. Eur. p. 46, p. p. (1831). Jungermania, sect. 12, Plumulosce, Hiiben. Hep. Germ. p. 265, p. p. (1834). Mastigojjhora, Nees. Nat. Eur. Leb. 1. p. 95, n. 3, et p. 101, n. 10, p. p. (1833); in Lindl. Introd. Nat. Syst. Bot. 2 ed. p. 414 (1835), et Nat. Eur. Leb. 3, p. 89, n. 1 (1838); Mitt, in Hook.-F. Handb. N.-Zeal. Fl. 2, pp. 752 et 754, n. 18-2 (1867). Mastigo2')hora, sect. 2, Cladura, Nees Nat. Eur. Leb. 3, suppl. p. 574 (1838). Blepharozia, Dum. Recueil, 1. p. 16, n. 15, p.p. (1835). Sendtnera, Endl. Gen. pi, 1. suppl. p. 1342, n. 472-16 (1840). Sendtnera, sect. 2, Mastigophora, G. L. N. Syn. Hep. p. 241 (1845). Herberta, Carruth. in. Seem. Journ. Bot. 3, p. 300, p. p. (1865). Branches lateral, furcate or dicliotomous, rarely simple, apices attenuate, decurved, secund. Leaves incubous, bifid to quadrifid, often calcarate at the base. Perianth produced from near the apex of the stem on a short, almost sessile lateral branch, some- times on a longer branch, membranous, tubular, trisulcate, mouth denticulate. Calyptra delicate, membranous. Capsule firm, dividing to its base into 4 valves. Elaters dispiral. Mastigophora Woodsii (Hook.), Nees. Jungermania Woodsii, Hook. Brit. Jung. t. 6(! (1816). BlejyJiarozia Woodsii, Dum. Eecueil, p. 16 (1835). Mastigophm-a Woodsii, Nees Nat. Eur. Leb. Ill, p. 95 (1838). Sendtnera Woodsii, G. L. N. Syn. Hep. p. 241 (1844). Inflorescence (?), loosely csespitose, very large, reddish to purplish-brown in colour, paler below. Stem ligneous, somewhat brittle when dry, reddish-brown, composed of thick-walled cells, cortical, similar in size to the inner, only slightly darker, about 30, inner brownish 10 x 10 ; flexuose, procumbent, bi-tripinnate, branches lateral, branchlets long, attenuate ; rootlets few, pro- ceeding from the attenuate branchlets, hyaline, delicate. Leaves patent-divergent to patent, bifarious, imbricate, very convex, con- duplicate, roundish or broadly subquadrate, unequally bilobed to below the middle, antical smaller, lobes spinulose dentate, sinus MASTIGOPHORA. 103 acute or slightly rounded ; cells smallish, roundish-quadrate margin irregular, walls thick, angles thickened. Stipules very large, considerably broader than the stem, broadly ovate, divided to about the middle, segments broadly subulate, acute, irregularly spinulose, sinus obtuse, at both sides of their base calcarate. Neither male nor female flowers have been observed. Dimensions. — Stems 2 to G inches long, diameter "2 mm., with leaves 1'5 mm. wide ; branches with leaves 75 m.m. wide ; leaves 1"1 mm. X 1"5 mm. ; segments 75 mm., "G mm., '9 mm. x 1*2 mm. seg., "G mm., '5 mm. ; branch leaves 75 mm. x "85 mm., seg. •4 mm. ; cells '03 mm. x 0*2 mm., '025 mm. x -0.25 mm. ; stipules •9 mm. X "9 mm., segments "55 mm., "8 mm. x "G mm., seg. '4 mm., 7 mm. X '6 mm., seg. '45 mm. ; branch stipules '6 mm. x "8 mm., seg. *3 mm. Hab. — Growing in large and rather densely crowded patches in boggy places or damp banks and rocks, in subalpine localities. Extremely rare. Discovered by Mr. Joseph Woods in 1809 on the ascent of Mangerton from Cwm na Cappal, Ireland. Brandon Mountain, Dr. Ta/jlor, W. Wilson. Devil's Punch Bowl, Man- gerton, Br. Taijlor, Steioart and Holt. Conner Hilh Dingle, Dr. D. Moore. Carrantual, Dr. D. Moore. South of Ireland. IG. Moidart, West Inverness, 8. M. Macvicar, 1898. 17. Faroe Isles, Rostrup, 1867. Himalayas (fide Milfeii). Obs. — This fine, beautiful and extremely rare species is remarkably distinct, and not to be confounded with any of our native ones, except perhaps Blcpharozia cUiaris, which might be mistaken for it, but from which it differs in the shape, margin, and structure of the leaves and stipules. Description of Plate XXXVII. — Fig. 1. Plants natural size (Mangerton, Holt). 2. Portion of stem, antical view (after Hooker). 3. Leaf (ditto). 4-6. Leaves x 24 (Mangerton, Holt). 7. Branch leaf x 24 (ditto). S. Portion of leaf x 290 (ditto). 9-11. Stipules X 24 (ditto). 12. Stipule (after Hooker). 13. Branch stipule x 24 (Mangerton, Holt). 104 HEPATICE. Genus 10. BLEPHAKOZIA, Bum. Ju7igerman{a, Linn. Sp. PI. p. 1601 ; Hook. Brit. Jung. (1816). Blejyharozia (Section) Dum. Syll. Jung. p. 46 (1831). Blepharozia, Dum. Recueil Jung. p. 16 (1835). Ptilidium, Nees Nat. Eur. Leberm. 1. p. 1)5 (1833). Leaves incubous, appressed, palmatifid, or complicate-2-lobed, each lobe divided and ciliate. Stipules 4-5-lobed, ciliate. Perianth terminal on short branches. Bracts 2-5-lobed, segments with long cilia. Perianth pyriform, obovate or sub-cylindrical, inflated, smooth, mouth small and plicately contracted, ciliate. Calyptra pyriform, coriaceous. Capsule divided nearly to the base into 4 valves. Elaters bispiral. Antheridia covered by closely imbricated perigonial bracts. 1. Blepharozia ciliaris (Z.), Bmn. Lichenastrum scorjnoides pulchrum villosum, Dill. Hist. Muse. p. 481, t. 69, fig. 3 (1718). Jungermania cilaris, Linn. Sp. pi. p. 1601 (1753); Engl. Bot. t. 2241; Hook. Brit. Jung. n. 65 (1816). Blepharozia ciliaris, Dum. Recueil, p. 16 (1835). Ptilidium ciliare, Hamp. Prod. fl. Here. (1836); Nees Nat. Eur. Leb. Ill, p. 117 (1838). Dioicous, csespitose, large, greenish to purplish-brown. Stems erect or procumbent, bipinnate, slightly frontally compressed, cross-section showing 15 cells x 20, cortical cells about 80, one or two rows with thicker, golden-brown walls, inner cells larger, walls hyaline, thin; branches lateral, short, alternate, ultimate pinnulse very slender, rootlets few, short. Leaves incubous, almost transverse, patent, lower leaves somewhat distant or approximate, upper- and branch-leaves imbricate, bifarious, convex, conduplicate, roundish or subquadrate, quadrifid nearly to the middle, segments lanceolate-acuminate, antical segment largest, on lower leaves entire or furnished with a few long cilia at the base of the leaf, next segment entire also, or with few cilia, the postical segments smaller laciniate- ciliate, upper and branch-leaves with margin and segments ciliate, cilia long. Epi- BLEPEAROZIA. 105 dermis slightly verruculose ; cells medium size, roundish, walls thick, trigones distinct. Stipules almost half the size of the leaves, ohlong, subquadrate, or semirotund, bifid, trifid or quadrifid, ciliate, cilia long. Sub-bracts subrotound, ciliate, quadrifid to the middle, segments lanceolate-acuminate, ciliate. Sub-bracteole oblong-oval, ciliate, bifid to about \, segments lanceo- late, ciliate. Bracts rotund, ciliate, quadrifid, segments unequal, lanceolate-acuminate, ciliate. Perianth terminal on main stem or on short branches, jDrojecting about f beyond the bracts, large, oblong-obovate, composed of a single layer of cells with a few double rows irregularly interposed, about 300 cells round near the middle, slightly folded at the upper portion, mouth contracted, 15-20 cilia of 2-4 cells long. Capsule dark brown, valves dividing to the base, thick. Spores large, pale brown. Elaters very narrow, J less wide than the spores, darker brown in colour. Male stems irregularly bipinnate, slender, androecia usually terminal, on short branches, perigonial bracts much smaller than the leaves, 4-6 pairs, bi-trifid, saccate, antheridia 1 or 2 in each bract, oval, bearers short. Dimensions. — Stems 2 to 3 inches long, *25 mm. in diameter, with leaves about 2* mm. wide; leaves 2' mm. x "2 mm., segments 1* mm., rS X 1*5, segment "75; branch leaves 1*5 mm. x l'25mm., segment -75 mm. ; cells '035 mm. x '04, '04 x "03, "03 ; sub-bract 1"75 mm. xl. '5 mm., segment '1 mm. ; sub-bracteole 1*5 mm. x 1' mm., segment "5 mm.; bract 2'5 mm. x 2'25 mm., segment 1*25 mm.; bracteole 2* mm. x 1"5 mm., segment '75; perianth 5* mm. X 2-25 mm.; spores '035 mm., -03 mm.; elaters 'll mm. x •Ql mm. Hab. — Growing in densely matted tufts of considerable size in bogs or damp j)laces on heaths or moors, especially subalpine. Rare in fruit. 1. 6-1 G. I. Found on the Continent and in North America. Obs. — Following Lindberg, I have assumed that the common large form is the true /. cUiaris of Linnreus, and as such have described it and have separated the small rock and tree form as B. jjulchcrrima (Hoffm.). 106 HEPATIC^. My figure 2, pi. xxxviii. is from the loioer portion of the stem. In Hooker's "Brit. Jung.," pi. 65, f. 3, is a drawing of the ii^)per portion of a stem and branches, with a perianth of the true B. ciliaris. Blejjharozia 2JulGherrima (Hoffm.) differs from this species in its smaller size, more procumbent habit, stems irregularly dichotomously branched, leaves more finely ciliate, never with segments entire on the stem-leaves, perianth cylindrical. Blejiharozia ciliaris is a very beautiful species, and not to be confounded with any other, the nearest approach to it of our British ones being Mast i r/ op ho r a Woodsii (Hook.), which differs in the margin of the leaves being laciniate, not ciliate, in its bifid stipules, which also are only spinose-dentate, not ciliate, and in the very different cell structure. Trichocolea tometitella (Ehrh.) was confounded by some of the older botanists with this species, to which it has a superficial resemblance, but apart from the differences at once revealed by the microscope, it is readily distinguished by its pale green colour, whilst B. ciliaris has a greenish to purplish-brown, or rich yellow. Description of Plate XXXVIII. — Fig. 1. Plant natural size (Cader Idris, W. H. P.), 2. Portion of stem, antical view x 24 (ditto). 3. Stem-leaf X 24 (ditto). 4. Branch-leaf x 24 (ditto). 5. Portion of leaf x 290 (ditto). 6. Stipule x 24 (ditto). 7. Branch stipule x 24 (Ditto). 8. Sub-bract x 16 (Helsingfors, S. 0. Lindberg). 9. Sub-bract x 16 (ditto). 10. Bract x 16 (ditto). 11. Bracteolex 16 (ditto). 12. Perianth x 1 1 (ditto). 2. Blepharozia pulcherrima, [Hoffm.) Lindh. Jungermania pulcherrima, HofFm. Deutschl. Fl. 2, p. 83 (non Linn, f.) (1796). Jungermania Leersii, Eoth. Tent. fl. Germ. 11, p. 402 (1800). Jungermania Hojfmanni, Wallroth, Fl. crypt. Germ. 1, p. 51 (1831). Blepharozia Hoffmanni, Cogn. Hep. in Bull. Soc. Bot. Belg. (1872); Dum. Hep. Eur. p. 54 (1874). Blepharozia jyulcherrima, Lindb. Muse. Scivnd. p. 5 (1879). Dioicous, csespitose, small, tawny or brownish-green in colour. Stems procumbent or rarely suberect, irregularly repeatedly BLEPHABOZIA. 107 dicliotomously ramose, branches ascending, cortical cells about 20, walls golden-brown, inner hyaline 8 x 10, slightly frontally compressed. Leaves crowded, patent, incubous, imbricate, con- duplicate, semirotund, trifid or quadrifid to about a third or more, segments broadly lanceolate subulate, margin and segments with long cilia, cells medium size, roundish, walls thick, angles thickened. Stipules about half the size of the leaves, semirotund, bifid to about a fourth, margin with long cilia. Bracts roundish- oblong-quadrate, trifid to about a third, segments lanceolate subulate, margin and segments with long cilia; bracteole ovate, bifid to about a fifth, margin and segments with long cilia. Perianth terminal on main stem or branches, projecting about two-thirds beyond the bracts, cylindrical, composed of a single layer of cells, about 100 round the middle, slightly obtusely trigonous, mouth contracted, ciliate, cilia 3 cells long, few. Pis- tillidia numerous, linear-lanceolate. Androecia terminal or situ- ated on the middle of the branches, perigonial bracts 4-6, small, closely imbricate, saccate at the base, subrotund, bifid to below the middle, margin and segments ciliate, antheridia oval, large. Dimensions. — Stems ^— | inch long, diam. 'S mm.; leaves 175 mm. X 2" mm., segments '75 mm,, -5 mm., 1"5 x 175, seg- ments 75 mm., '6; cells -03 mm., -03 x '035 ; stipules -85 mm. X 1* mm., segments '4 mm., '2 mm.; bracts 1*25 mm. x "9 mm., segments '6 mm., '5 mm.; bracteole 1.25 mm. x l* mm; perianth 3" mm. X 1* mm., cilia at the mouth -2 mm. long; perigonial bract *5 mm. x "6 mm., segments '3 mm.; antheridia '2 mm. X '15 mm. Hab. — Growing on trees and rocks. Rare. 10. On trees, near Ooneysthorpe. .Z>r. Spruce. 15. Carn Fiaclan, Balmoral, Scotland, G. Stabler. Found on the Continent and in North America. Obs. Distinguished from Blepharozla ciliaris (L.) by its smaller size, more procumbent habit, brownish-green colour (never rose or purple), its irregularly dichotomous spreading branches, not grace- fully plumose and regularly bipinnate, its crowded, much more ciliated stem leaves, segments never entire, and cylindrical perianth, not oblong obovate. 108 HEPATIC^. Description of Plate XXXIX. — Fig. 1-3. Plants natural size. 4, Portion of stem x ?. 5, 6. Leaves and stipules x 24 (Husnot Hep. Gall. n. 45). 7. Leaf x 24 (Greenland, E. Brown). 8. Portion of leaf x 290 (Norway, S. 0. Lindberg). 9, 10. Bracts X 14 (Balmoral, G. Stabler). 11. Bracteole x 24 (ditto). 12. Perianth x 16 (ditto). 13. Cross-section of perianth x 16 (ditto). 14. Cilia from mouth of the perianth x 85 (ditto). 15. Perigonial bract x 24 (Newfoundland, Waghorne). 16. An- theridium x 85 (ditto). Genus 11. TRICHOCOLEA {Diuu,\ Nees. Thricolea, Dum. Comm. Bot. p. 113 (1822). Tricholea, Dum. Syll. Jung. Eur. t. 1. f. 8 (1831). Trichocolea (Dum.), Nees Nat. Eur. Leberm. Ill, p. 98 et 101 (1838). Dioicous. Stems glaucous-green, conspicuous, pinnate com- pound or decompound. Leaves laciniate almost to their base, setaceo-multifid, succubous. Stipules multipartite. Involucre terminal, or from the growth of innovations axillary. Perianth when mature either free, naked and not incrassate, surrounded only at the base or a little beyond by the abortive pistillidia and the narrow ring of involucral scales. Or undergoing more com- plete metamorphosis, in which the cortical layer of the tomentose receptacle invests and becomes blended with it, so as to form a cylindrical coriaceous hirsute involucre bearing at the apex the abortive pistillidia. Calyptra wanting. Capsule oblong, cleft to the base into 4 coriaceous valves. Pedicel bulbous at the base, inserted deep within the receptacle. Elaters free, bispiral. An- theridia large, axillary in the terminal leaves of separate stems. Trichocolea tomentella (^Ehrh.), Bum. Lichenastrum Jilicmum 2ndchru7n villosum, Dill. Hist. muse. t. 73, f. 35 (1741). Jungermania tomentella, Ehrh. Beitr. 2, p. 150 (1785); Hook. Brit. Jung. t. 36 (1816). Tricholea tomentella, Dum. Com. p. 113 (1 bor. p. 28 (1888). Dioicous, cjespitose, medium to largish in size, of a yellowish- brown colour. Stems simple or slightly irrejrularly furcate, flexuose, filiform, rigid and somewhat brittle when dry, erect or procumbent, on a cross-section 15 cells in diameter, cells opaque, cortical cells 40-50, reddish-brown, smaller than the inner ; root- lets few, near the base of the stem. Leaves closely imbricate, bifarious, erect, appressed, semi-amplexicaul, subtransversely inserted, sabquadrate, 1)roader than long, quadripartite, segments lanceolate, acute, furrowed, margin outwardly curved, near the base furnished with long recurved teeth ; texture firm, cells smallish, roundish-quadrate or oblong roundish-quadrate, walls very thick, no trigones. Stipules large, oblong-subquadrate or ovate, bipartite, segments lanceolate, acute or acuminate, furrowed. Bracts larger than the leaves, subquadrate, broader than long, quadridentate to about the middle, segments broadly subulate or lanceolate, acute or acuminate, furrowed, margin of bract and segments laciniate-dentate ; bracteole oblong subquadrate, biden- tate to the middle, segments broadly subulate or lanceolate, acute or acuminate, furrowed, with a smaller outer lobule at both sides at the base, margin laciniate-dentate. Perianth terminal, project- ing about half beyond the bracts, oblong-oval, deeply 9-10-plicate to the base, 2 to 3 cells thick near the base, above one cell layer, mouth delicate, hyaline, laciniate-ciliate, cilia 4-G cells long, 2 cells wide at the base; pistillidia 12-15. Male stems much more slender, with more distant leaves. Dimensions. — Stems 1 to 2 inches long, -125 mm. to "2 mm. in diameter, with leaves *5 mm. to 75 mm. wide; leaves '6 mm. X '7 mm., segments "4 mm. x 2 mm., '5 mm. x 7 mm., seg. 45 mm. X '2 mm., 7 mm. x 1" mm., seg. '45 mm. x '25 mm.; CHANDONANTHUS. 115 cells '025 mm. x -02 mm., '03 mm. x -02 mm., 025 mm. X '025 mm. ; stipules *5 mm. x '4 ram., seg. '35 mm. ; sub- bracteole "7 mm. x '5 mm., seg. "5 mm. ; bract Tl mm. x 75 mm., segments '4 mm., '5 mm., '6 mm.; bracteole I'l mm, x -75 mm., seg. '6 mm.; perianth 1"8 mm. x "9 mm.; cilia at the mouth •175 mm. Hab. — Grrowing in large entangled patches or straggling amongst mosses in alpine situations. Very rare. 15. Balmoral, Geor//e Stabler, July 1884. Var. alpi,ia. 15. Catlow Hills, C. Lt/ell, 1813, Glen Dole, 1855, lib. Carrim/fo)i. Glen Muick, J. Sim.. Slack of Birnie, ./. Sim. Cairngorm, J/ooker. Scotch Alps, Dickson, Donn. Found on the Continent and in North America. Obs. — Very distinct and not to be confounded with any other of our British species. As Dr. Spruce remarks : " The whole habit, the leaves and their cells, &c., recall Chandonanlhus .sqifarrosns and hirfeUii-s." By some authorities it has been referred to Blcp/ia- rostoma, but in that genus the perianth is uniformly trigonous ; by others to A?ithelia, where the leaves are only bifid and along with the stipules, which are similar and give a tristichous insertion to them ; the perianth also is not so multiplicate. Hooker figures the mouth of the perianth without cilia, but in well developed specimens it is beset with them. It is only recently that copious specimens with perianths have been found in Norway. The var. alpina., Hook. [Jmu/ermania jihim, Dum.), is a much smaller form, very slender, filiform, with more distant and less dentate leaves; it might be confounded with Antlielia j/dacea, but is at once distinguished by its quadripartite leaves. The typical form has, so far as I know, only been found in Britain by Mr. George Stabler. I have not seen the male plant nor perfect fruit. Description of Plate XLII. — Fig. 1. Plants natural size. 2. Portion of stem x ?. 3. Leaves x 31 (Balmoral, G. Stabler). 4, 5. Leaves, explanate x 24 (ditto). 6-8. Leaves x 31 (ditto). 9. Leaf and stipule x 24 (ditto) 10. Segment of leaf x 85 116 HEP A TIC J^:. (ditto). 11. Portion of leaf x 290. 12. Stipule x 24 (ditto). 13. Ditto X 31 (ditto). 14. Sub-bracteole x 31 (Norv/ay, A. Blytt). 15. Bract x 31 (ditto). 16. Bracteole x 31 (ditto). 17. Peri- anth X 24 (ditto). 18. Cross-section of perianth x 24 (ditto). 19. Portion of mouth of perianth x 85 (ditto). Subtribe IV. TRIGONAKTHE^. Genus 11. LEPIDOZIA, T)im. Jungermania, Linn. Sp. pi. 1, p. 11.3P> (1753). Pleuroschisma, sect. Lepidozia, Dum. SylJ. Jung. p. 6!) (1831). Mastigo2)Jiora, Nees, Nat. Eur. Leberm., 1, p. 101 (1833), nee. Syn. Hep. Lepidozia, Dum. Recueil, p. 19 (1835); Nees in G.L.N. Syn. Hep. p. 200 (1845). Herpetitmn, sect. 1, Lepidozia, Nees, Nat. Eur. Leberm. Ill, p. 31 (1838). Plants large, rareJy small, pale or yellowish- green, rarely very green, densely osespitose, tufts depressed, rarely erect or pendulous. Stems in many species firm, beautifully feather-like, pinnate or bipinnate; branches lateral, sometimes attenuate and radiculose at the apices, rarely, and principally in small species with postical branches, which are frequently flagelliferous and radiculose. Leaves incubous, small, in some species very minute, sometimes broader than long, very oblique, with antical margin much longer and rotundate, decurvo-concave, palmate or quadrifid almost to the base, rarely 5-6- or only 2-3-fid, segments subulate, entire or furnished at the antical base with a few teeth, in a few species all the margin dentate ; cells usually smallish, quadrato-hexagonal, inferior subelongate, angles slightly thickened, cuticle often rough ; in Micro- Lejndozia very small oblong-quadrate. Stipules similar to the leaves but symmetrical, in Eu-Lepidozia slightly smaller, in Micro-Lepidozia about half the size of the leaves. Inflorescence dioicous, in a few species (chiefly Micro-Lejjidozia) monoicous ; ? on the stem, rarely on the branches, hypogenous. Bracts 3-5 pairs, appressed, concave, innermost frequently about three times (in small-leaved species up to eight times) larger than the leaves, at the apex 2-4-fid, margin denticulate or spinose. Pistillidia 20 or less, the sterile surrounding the base, or a little higher, LEPTDOZIA. 117 of the calyptni. Perianth ahiiost always elongate, ovate-subulate or narrowly fusiform, upper portion obtusely trigonous, in E/i- Lepidozia carnose, mouth entire or denticuhite, Micro- Lepidozia leptodermous, mouth ciliato-laciniate. Calyptra 2-4 times shorter, pyriform or oblong, below somewhat carnose, or in Micro- Lepidozia whole length delicate, afterwards rupturing into two hps. Capsule on a more or less firm shortly exserted pedicel, oblong-cylindrical, dividing to the base into 4 valves, 2-4-strata. Elaters slender, bispiral. Spores minute, smooth or asperulous. Androecia very often constantly on slender postical branches, rarely (and in Micro-Lepidozia principally) terminal on lateral branches ; peri- gonial bracts 5-10 pairs, suborbiculate concave, apex bidentate ; antheridia solitary large pedicellate. 1. Lepidozia cupressina {Sw.\ Junyermmiia ctijjressina, Swartz Prod. Fl. Ind. occid. p. 144 (171)7?). Jumjermaida reptans var. pinnata, Hook. Brit. Jung. n. 75 (18U»). Pleuroschisma reptans war. pi7inata, Dum. Syll. Jung. p. (>'.> (1H;!1). Lejjidozia pinnata, Dum. Kecueil, p. ID (LSoT)). Herpethmi reptans var. pinnatum, Nees Nat. Hist. Eur. Leberui. Ill, p. •>2 (1838). Lepidozia timiidida, Tayl. in G.L.N. 8yn. Hep. p. 20(i (1844). Lepidozia, cupressina var. tumidula, Carr. Trans. Bot. Hoc. Edin. VII., 4i)o, t. xi., f. 7 (18G3). Dioicous, densely csespitose, flagelliferous, medium size, greenish to olive or reddish-brown in colour. Stems creeping or suberect, curled, on a cross-section 16 x 20 cells in diameter, cortical about 50, these and the next inner layer large with thick, dark walls, inner cells smaller with thinner walls, of a light golden brown colour; closely pinnate or bipinnate, branches alternate, unequal, often attenuate, apices fiagelliferous. Leaves incubous, closely imbricate, patent or ])atent-divergent, incurved, very con- cave, cordate-orbiculate or broadly subquadrate-cordate, quadrifid to nearly the ^, segments acute, 8 to 12 cells broad at the base, sometimes even more, postical segments much inflexed ; cells smallish, opaque, subquadrate-hexagonal. \valls thick, angles 118 HEFATIO/E. slightly thickened, no trigones. Stipules large, orbicular or subrotund, or roundish-subquadrate, concave, to nearly the -J quadrifid, segments acutate. Perianth on short postical branches, bracts several pairs, innermost largest, oval-orbicular, apex with 4 or 5 small teeth ; bracteole orbicular, quadri-denticulate ; sub- bracteole ovate, quadri-dentate. Perianth projecting f beyond the bracts, subulate, cylindrical, upper half plicate, mouth con- tracted. Androecia on short postical catkins, often 3 or 4 on a stem ; perigonial bracts closely imbricate, turgid, bitid; perigonial bracteole oval, bidentate, texture as with the perigonial bracts much more delicate than the leaves, cells larger with thinner walls. Antheridia solitar}^ large, roundish-oval, shortly stipitate. Dimensions. — Stems 1 to \\ inch long, diameter "2 mm., with leaves 1' mm. wide; leaves 1* mm. x •S5 mm., segments '4 mm., •9 mm. X "9 mm., seg. '3 mm. ; cells '02 mm. x '025 mm., '02 mm. X '02 mm., '03 mm. x "025 mm., -03 mm. x '03 mm., '035 mm. x '02 mm. ; stipules "55 mm. x "55 mm., segments "2 mm., "5 mm. x •45 mm., seg. '2 mm., "5 mm. x '6 mm., seg. "2 mm., "5 mm. x '5 mm., seg. "2 mm.; bracts 1"1 mm. x '8 mm., teeth "15 mm.; bracteole '8 mm. x "6 mm., teeth "1 mm. ; sub-bracteole "6 mm. x •4 mm., teeth "1 mm. ; perianth 4* mm. x '8 mm. ; perigonial bract "5 mm, x '5 mm., seg. "15 mm. ; perigonial bracteole '25 mm. X '175 mm., seg. '075 mm. ; antheridium "2 mm. x "175 mm. Hab. — Grrowing in large dense patches on exposed or shaded banks, rocks and trees. Rare. 1. Cam Galva, Cornwall, //'. Cnrnoic. 9. Helsby Crag, Cheshire, W. WiUon. Clougha, Lancashire, G. Stabler, July 1881. 10. On gritstone boulders, Idle Woods, Dr. Carrivg- ton, 1857. 12. Borrowdale, Cumberland, Dr. Carriugton and W. 11. P., April 1S84. 16. Moidart, West Inverness, S. M. Macvicar, 1898, W. II. P., 1899. South of Ireland, more or less abundant, Jliss Hut chins, Br. Taylor, Dr. Spruce, and others. Extremely rare on the Continent, Germany, Prof. A. Braiin ; France, near Cherbourg, Mons. Thuret. Found in the West Indies and South America. LEPIDOZIA. I 1 Obs. — Distinguished from L. reptans by its dioicous inflores- cence, its tawny to reddish-brown colour, more robust habit, stems more erect and irregularly pinnate with long flagella, leaves more closely imbricated, cordate-orbiculate (not quadrate), cells across the base of segments more numerous, cells smaller and more opaque with thicker walls. From L. Pear,^o)n, 8p., by its much more robust habit, shape of leaves and stipules aiid androecia on short postical branches. Collected first in the Ih'itish Isles by Miss Hutchins and recoo- nised as distinct from L. repta/is by Hooker, who named it var. pinnata, collected afterwards by Dr. Taylor, who made a new species of it (Z. tumidula, Tayl.). It has been referred to by later botanists as L. cupressina, var. ticinldida, but I am unable to detect any difference between it and the type found in the West Indies and South America. Description or Plate XL I II. — Fig. 1. Plant natural size. 2. Stem, antical view x 11 (Original o^ Lepidozia tumid/da, 'J'avl., Dr. Taylor, Killarney). 3, 4. Leaves x 31 (ditto). 5. Portion of leaf X 290 (ditto). 6-10. Stipules x 31 (ditto). 11, 12. Bracts x 24 (Killarney, S. 0. Lindberg). 13. Bracteole x 24 (ditto). 14. Sub-bracteole x 24 (ditto). 15. Perianth x 11 (ditto). 16. Perigonial bract, with antheridium x 64 (Killarney, G. A. Holt). 17. Perigonial bracteole x 64 (ditto). 2. Lepidozia reptans (//.), Di(m. Jfuscoides ierrestre repens ex obscuro viresceud, foliis superioribus et hoferioribus ad extremitatem dentatis, Mich. Nov. p]. gen. ji. 10, t. 6, f. '2 (1729). Lichenastrum multifidum exigv/um, ad basin florens, p)er siccitatem iinbricatimi, Dill. Hist. Muse. p. 494, t. 71, f. 24 (1741). Jungermania reptans, L., Sp. pi. 1599 (175B). Pleuroschisma reptaiis, Dum. Syll. Jung. Eur. p. 69 (1S81). Mastigophora rejitans, Nees. Nat. Eur. Leb. 1, p. 10 i (1S;;:5). Lepidozia reptans, Dum. Kecueil. p. 19 (1^!35). Herpetium reptans, Nees. Nat. Eur. Leb. Ill, p. ol (18;^S). Monoicous, densely cjespitose, tufts depressed, Hagelliferous, medium size, green, when older pale or yellowish-green, when dry 120 HEPATIC^. cserulescent. Steins creeping, horizontal, irregularly subpinnate or sub-bipinnate, flexuose, on a cross-section 12 to 15 cells in diam., hyaline, cortical larger, about 30 ; branches all lateral, some attenuate ; radiculose, rootlets numerous, near base of the plant, fasciculate, hyaline. Leaves incubous, imbricate or subimbricate on main stem, on branches, and attenuated branches small and distant, patent or patent-divergent, convex, subquadrate or semi- oval, slightly longer than broad, quadri-dentate to about ^, seg- ments acute, 4-7 cells broad at the base, sinus acute, apices incurved, branch leaves tridentate, axillary leaves bidentate ; cells medium size, subquadrate, often broader than long, Lubopaque, walls thick, but somewhat delicate, no trigones or thickened angles. Stipules about twice the width of the stem, very convex, sub- quadrate, broader than long, quadrifid to about a third, on branches trifid, segments acute. Bracts several pairs at the base of the perianth, very delicate texture, subrotund or broadly oval, irregu- larl}'- dentate at the apex ; bracteole very delicate texture, oval or broadly oval, irregularly dentate at the apex. Perianth produced on very short postical branches, projecting f beyond the bracts, almost hyaline, sub-membranaceous, lower half composed of two Iciyers of cells, upper half of a single layer, near the middle about 100 cells round, oblong-subulate, below cylindrical, above some- what plicate, mouth constricted, dentate, teeth about 8. Calyptra large, oval, composed of one layer of cells, Pistillidia few, 4 or 5. Capsule oblong-ovate, deep brown, splitting into 4 equal lanceolate valves. Spores small, verruculose, reddish-brown. Elaters bispiral, not quite as broad as the spores, reddish-brown. Andnpcia postical, amentiform, catkins small, composed of 4-6 pairs of perigonial bracts, which are minute, delicate, complicate, broadly oval when explanate, bidentate to about a fourth ; perigonial bract- eoles oval, bidentate. Dimensions. — Stems 1 to l^ long, "2 mm. to '3 ram. in diam., with leaves 1* mm. wide ; leaves 7 mm. x -6 mm., segments "l 5 mm., •6 mm. X "45 mm. seg. '2 ram. ; cells '025 mm. x '025 mm., '035 mm. x '03 mm., "045 mm. x '035 mm ; stipules "4 mm. x '4 mm., segments "2 mm., 4 mm. x -45 mm. seg. "15 mm.; bract 'TS LF.rinnXIA. 121 mm. X • I '25 mm., sc'Limcnts '."k) mm., '1 mm. ; Wiacteolu 1 '!'.) mm. x •'.) mm.. ."^OLTmcnts "2 mm. ; perianth b' mm. x |- mm., toctli at tin- muutli l mm., OTo mm.; pistillidia 175 mm. x 05 mm.; valve of capsule 15 mm. x 7 mm. ; pedicel :{ mm. diam. ; spores 01 5 mm. diam. ; elaters ;! mm.xOI mm.; perigonial bracts, exj^lanatc ■*VJ5 mm. X "J mm., sej^nnents 075. :> mm. x •'!:) mm. ; sc^Miicnts •075 mm.; antlieridia '125 mm. x "I mm. \\.\\\. — Growing' in dense tul'ts or straL,''nlin;j: amoni^st mosses. Common. I to 17. I. C. Found on the Continent, and in North America. Obs, — This is a common species; lor its distinguishini,^ characters from A. ci/prrs.s'nia and A. Pearsunl, see notes under those species; heim; monoicous it is easil}' recognised hy this character. Dksc'IUI'tion oi Pl.\t1': XL1\^- Fig. 1. Plants nat. size. 2. Portion of stem, antical view x IG. '.\-i\. Leaves Xol. 7. Leaves and stipule X 24. 8. Portion of leaf x 290. \), 10. Stipules x 31. I 1. Bract X 24. 12. Jiracteole x 24. \:\. Perianth x 1 1. 14. Cross- section of the perianth near the middle x I (). 15. Ditto, tlie upper half X IG. 16. Mouth of the perianth, ex|)lanate x 85. 17, 18. Perigonial bracts x S5. !'.». Antheridium x 85 (all Castle Howard Woods, M. K Slater). 3 Lepidozia Pearsoni. Sjjrucc. Lejjidozia I'em'soni, Spruce Jourii. of 15ot. p. ;>l (IHHl). Lepidoiia Wnlfsheryii, Lindb. Soc. F. Fl. Fenn. 1S8L*; l!ol. not. (Issi'). Dioicous, creeping, medium to largish in size, from [)ale to tawny-green in colour. Stems subterete, on a cross-section cells 15x18, cortical 40 to 50, a little larger; flaccid, very slender, elongate, simple or furcate, afterwards la.xly and distantly pinnate : branches short, unequal, simple, very rarely ramulose, some with apices llagelliferous, microphyllous or nidiculose ; rootlets jiroceed- ing more from the stem than I'rom the llagella, or ;dmost absent ; all branches ujore or less lateral, a.xillary, olten with male flowers; rarely a postical stolon is observed and sometimes postical amenti- 122 HEPATIC^. form male flowers. Leaves small, more or less distant, rarely sub- contiguous, obliquely incubouS; patent or erecto-patent, subquadrate, usually more than half or to the middle palmitifid, segments 4, very rarely 5 or 6, subulate, subacuminate incurved, uncinate, at the base 3-5 cells broad, the two middle ones broadest, postical shorter ; cells medium size, subquadrate-hexagonal, a little elong- ate, subpellucid, walls thin, sometimes with the angles slightly thickened, branch leaves smaller 3, 4-fid, upper ones longer and narrower, profoundly bifid. Stipules shorter than the leaves, sub- rotund to subquadrate, divided to about the middle, segments 4 rarely 5 and more rarely 6, broadly subulate, obtuse, incurved. Androecia spicate at the apex of delicate lateral branches, rarely at base, more rarely on short postical branches ; perigonial bracts, 3-10 pairs, slightly smaller than the leaves, imbricate, concave, swollen, bilobed, rarely with an additional antical basal tooth, lobes ovate, acute, incurved ; bracteole subquadrate, longer than broad, bi-trilobed, antheridia solitar}^ large, oval, shortly stipitate. Habitat. Creeping loosely amongst mosses and other hepatics on steep banks or rocks. Rare. 7. Tyn-y-Groes, Merionetlishire, W. H. P., A-pril 1879. 8. Kinder Scout, Derbyshire, ./. Whitehead nnd G. A. Holt, March 1884. 12. Borrowdale, Cumberland, W. H. P., Skelwith Bridge, West- morland, W. H. P. 13. New Grail oway, /. Mc Andrew. 16. Banks of Loch Maree, Argyle, JJr. Carriiifjtou, October, 1889, Moidart, West Inverness, S. M. Macvicrrr, 1898, ir. IT. P., 1899. Norway. Dimensions. — Stems 1 to 2 inches long, 2 mm. in diameter, with leaves '5 mm. wide; leaves '55 mm. x '45 mm. middle seg- ment, '35 mm., "4 mm. x "4 mm. seg., "2 mm., "4 mm. x '3 mm. seg., '2 mm., '5 mm. x "Smm. seg., "25 mm., 4 mm. x "35 mm. seg., "2 mm. : cells '033 mm., '05 mm. x '03 mm., "025 mm. x '035 mm., •04 mm. X -025 mm., '045 mm. x ^03 mm. ; stipules '3 mm. x •4mm., *275 mm. x '3 mm. ; segments "1 mm., '35 mm. x '375 mm. seg., •15 mm., "3 mm. x '35 mm. seg., '15 mm. ; perigonial bracts '4 mm., •475 mm. X '3 mm. seg., '225 mm., "35 mm. x "3 mm. seg., "15 mm., •4 mm. X '4 mm. seg , "15 mm. ; perigonial bracteole "2 mm. x '2 mm. segments, "075 mm. ; antheridia *13 mm. x *1 1 mm. LEPinoZIA, 123 Obs. — " Lepidozia Peari^oni'i^ well distinguished from L. rcplanf^ by the following characters. " In //. Fearso7ii the whole plant is very slender and elongate, and of a pale tawny colour. Stems distantly pinnate, without any rooting flagella from the underside. Leaves small, those of the stem usually distant, or at most contiguous, cloven to heyond the middle into 4, sometimes (though rarely) into 5 or (5, suljulate subacuminate segments. Stipules cloven to the middle into 4, or not unfrequently into 5, more rarely into G, subulate obtuse seg- ments. Inflorescence dioicous. Male spikes terminal (rarely basal) on the lateral branches ; very rarely indeed on a short postical ramulus. "Z. reptcms is of a more robust habit, with shorter, more closely- branched stems ; of a deep green colour when fresh, often turning blueish-white, or glaucous, in drying. Leaves subimbricated, cloven only half-way into never more than 4 segments, which are 4-7 cells wide at the base ; cells broad as long. Stipules with acute segments, which are never more than 4 in number even on the main stem, and are sometimes reduced to 3. The chief dis- tinction, however, is in the monoicous i/i florescence, with the male flowers uniformly in short catkins springing from the underside of the stem (which is their normal position on nearly every other Eii- lejndozla known to us, except L. Pearso)ii)." R. Spruce. . Description of Plate XLV. — Fig. 1. Plants natural size. '1. Portion of stem, antical view x 11. 3-6. Leaves from main stem and branch x 64. 7. Portion of leaf x 290. 8, 9. Stipules x 64. 10. Stipule x 31. 11,12. Perigonial bracts x 64. 13. Perigonial bract x 31. 14. Perigonial bracteole x "60. 15. Antheridium x 85. (All drawn from specimens collected at Tyn- y-Groes, N. Wales, W. H. P.). 124 UEPATIGJ^. 4. Lepidozia setacea [l/'e6.), Miff. Jungemiania setacea, Web. Spic. pi. Goett. p. 143 (1778), Hook. Brit. Jung. t. 8 (1816). Blepharostoma setacea, Dum. Recueil (I8;:!r»). Lepidozia setacea (Web.) Mitt. Proc. Journ. Linn. Soc. v. p. 103 (1841). Dioicous, densely csespitose or creeping, small, pale yellowish- green to dark or olive-brown in colour. Stems very slender, filiform, firm, flexuose, on a cross-section 5 and 6 cells in diameter, large, clear, cortical 10-12, similar in size to the inner ; walls firmer and darker ; irregularly pinnate, or subpinnate, often denu- date below ; branches lateral, unequal, alternate ; flageila few, postical ; radiculose, rootlets few, minute, hyaline. Leaves on main stem contiguous or distant, on branches imbricate ; erect, incurved, in outline subquadrate, tri-, rarely quadri-partite almost to the base, where they are 6 cells broad, segments setaceous, subulate, 3-6 cells long, 2, 3 cells wide at the base, branch leaves often bipartite only ; texture delicate ; cells 4-, 5- and 6-angled, smallish to medium size, lumen clear, walls thin but firm, no trigones or thickened angles. Stipules equal in width to the stem or slightly wider, erect, sometimes appressed to the stem, oval-orbicular in outline, tripartite almost to the base, segments setaceous, subulate, usually 4 cells long, a row of 2 at the base. Perianth postical, arising usually from near the base of the stem from an axil of stipule, on short branches, projecting f beyond the bracts. Innermost bracts much larger than the leaves, oval- orbicular, bidentate, segments ciliate-dentate, acuminate ; bracteole oval, bidentate, segments ciliate-dentate, acuminate. Perianth 4 times longer than broad, oblong, cylindrical below, plicate above, mouth somewhat contracted by the folds, ciliate-dentate, cilia few, 3, 4 cells long, texture delicate, cells laxer than those of the leaves, elongate, almost white in colour, lower half composed of two layers of cells, above of one layer. Calyptra obovate, very delicate ; pistillidia 4-G, linear. Pedicel long, pellucid, delicate. Capsule oval, dark chocolate-brown. Spores dark brown. Elaters bispiral, slightly broader than the spores, similar in colour. Androecia LEPIDOZIA. 125 usually on short lateral branches at the middle or end, amentiform, when terminal often subspherical, rarely on main stem ; perigonial bracts 4, 5 pairs, closely imbricate, of delicate texture, pale colour, bifid, rarely trifid, margin dentate, rarely entire. Antheridia solitary, small, oval, shortly stipitate. Hab. — Growing in dense depressed tufts or straggling amongst Sphagna or other mosses and hepatics on damp shady banks, moors, or marshy ground. Common. 1-3, 5, 7-10, 12-lG, I. Found on the Continent and in North America. Dimensions. — Stems \ to 1 inch long, diameter '075 to -1 mm., with leaves '2 mm. wide; leaves '175 mm. x '15 mm., segments '125 mm., '15 mm., "25 mm. x "2 mm., seg. "2 mm., "15 mm. x •125 mm., seg-. "1 mm., 125 mm. x '1 mm., sesr. '1 mm.: cells •02 mm. x '03 mm., '025 mm. x '035 mm., '04 ram. x 'Go mm., "035 mm. X '02 mm.; stipules "175 mm. x "125 mm., seg. "125 mm.; bracts "45 mm. x "25 mm., seg. "125 mm., "55 mm. x "35 mm., seg. '175 mm., 75 mm. x "35 mm., seg. '15 mm. ; bracteole •4 mm. X "2 mm., seg. "15 mm.; sub-bract '25 mm. x "2 mm.; sub-bracteole '2 mm. x "1 mm. ; perianth 2* mm. x '5 mm., 3 "5 mm. X 75 mm. ; cilia at the mouth '15 mm. ; calyptra '6 mm. x "4 mm. ; pistillidia "2 mm. x "05 mm. ; capsule 7 mm. x "35 mm., valves '8 mm. x '3 mm. ; pedicel "175 mm. diam. ; spores "01 mm. diam.; elaters '175 mm. x "0125 mm.; perigonial bracts "25 mm. X "15 mm., seg. "175 mm., '3 mm. x "2 mm., seg. "2 mm., "325 mm. X "225 mm., seg. "175 mm. ; perigonial bracteole "25 mm. x '1 mm., seg. "2 mm., "1 mm. ; antheridia "08 mm. x "OG mm. Obs. — Distinguished from all other British Lejndoz'ue by its smaller size, deeply divided leaves with subulate segments ; from Blejjharo.sfoma Irkltopliijllum by its smaller size, more robust habit, darker colour, shorter segments of leaves, lateral branches, long- narrow perianth, and dioicous inflorescence. Tlie varieties sertularioldes and Schulfzii {Jiui//. scrtidariuides, Linn, f suppl., p. 449 ; Juftg. Schulfdi, Spreng Pugill., 1, p. 64) with smaller, closer imbricated leaves, are found in drier situations, and pass to the normal form in damper localities. 12G HEPATIC.^:. One form (near Festiniog, Dr. C. and IF. H. I*., and Killarney, H. 0. L.) is larger, more regularly pinnate, of a softer texture with tlie bracts hardly so laciniate as the usual form, but I have observed no other characters to separate it from the type. Dr. Spruce found on Strensal Moor a stout and often fertile form which has leaves mostly 4-cleft, although in laxer forms of the species, leaves with more than 3 divisions are rare. The perfect fruit of this species is very rare. Description of Plate XLVI. — Fig. 1. Plants natural size Portion of stem, antical view x 24 (Killarney, S. 0. Lindberg) 3. Ditto, postical view x 85, showing stipules (ditto). 4, 5 Leaves x 85 (ditto). G, 7. Leaves x 85 (Barton Moss, Lane. W. H. P.). 8. Leaf x 85 (Aust. Hep. Am. Bor. n. 76). 9, 10. Branch leaves x 85 (Killarney, S. 0. L.). 11. Branch leaf x 85 (Barton Moss, W. H. P.). 12. Portion of leaf x 290 (ditto). 13, 14. Bracts x 64 (White Mts., N. America, Gates). 15. Bracteole x 64 (ditto). 16. Sub-bract x 64 (ditto). 17. Sub- brac^eole x 64 (ditto). 18. Perianth x 31 (ditto). 19. Portion of the mouth of the perianth x 85 (Tyn-y-Groes, N. Wales, W. H. P.). 20. Capsule and calyptra x 31 (White Mts., Oakes). 21 Perigonial bract x 85 (Aust. Hep. Am. bor., n. 7Q). 22, 23. Ditto X 64 (Sussex, Gr. Davies). 24. Perigonial bracteole X 64 (ditto). 25. Antheridium x 85 (ditto). Genus 12. BAZZANIA, Gr. ^^ B. Jnngermania, Linn. FJ. Suec. ed. 1, p. .'5;5.") (174;")). Bazzatiia, Gr. & Benn. Nat. Arr. Brit. PI. 1, p. 704 (1S21). Pleuroschisma, sect. Pleuroschismotypus, Duni. Syll. Jung., p. 70 (1S;51). Pleuroschisma, Dum. Recueil, p. 1!) (ISoS). Berpetium, Nees, Nat. Hist. Eur. Leberm., Ill, p. 214 (1838). Masiigobri/um, G.Jj.^. Syn. Hep. p. 214 (1845). Stems firm, almost round, slightly compressed frontally, repeatedly furcate, dichotomous. Branches postical, very short and floriferous, or often elongate, microphyllous and radiculose. Cells of the stem several layers, cortical similar to the inner ones, opaque. Leaves incubous, alternate, in a few species opposite, at the base more or less imbricate, apex more or less distant, decurved, JlJZZ.iXM. 127 sometimes (principally when dry) secund, always oblique, some- times llilcate, about twice as long as broad, at the base semicor- dato-ovate, gradually becoming narrower, the upper half subligu- late, apex plane, almost always truncato-tridentate, rarely 4-dentate or subentire — in a few species equally bidentate, or unequally bilobed ; margin in most species entire, in a few species postical base spinose or dentate. Cells near the base elongate, others small, subquadrate, guttulate, usually smooth, walls and angles distinctly thickened. Stipules everywhere present, about half the size of the leaves, rarely 3-4 times shorter, broader than the stem, appressed, very often subrotundo-quadrate, rarely longer, apex truncate, very often 4-crenate or incised, rarely subentire, margin entire or subdentate ; in a few species cordate at the base, auricles sometimes spinose. Flagella unequally leaved ; leaves minute, tristichous, ovate, concave, apex bidentulate, rarely only apiculate or tridenticulate, producing from near the base long pale radicles. Inflorescence dioicous, both sexes cladogenous, constantly on postical branches. Bracts $ 3- 5-pairs, innermost largest (but often shorter than the leaves), closely imbricate, concave, orbicu- late or ovate, rarely ovato-lanceolate, apex lobulate, laciniate or ciliate, subscariose ; cells rather large, elongate, 4- G-angled. Pistillidia 10-lG. Perianth ovato-subulate or fusiform. 3 to 4 times longer than broad, subcarnose at the base, leptodermous, at first tricarinate at the base, keels broad, quasi G-plicate, on the maturing of the fruit the keels almost obliterated, sometimes sub- terete, with only the apex trigonous constricted, mouth in perfect state with 12-15 long cilia. Ualyptra about half the size of the perianth, pyriform or cylindrical-oblong, at the base 3-strata, surrounded by the sterile pistillidia, above 2-strata. Capsule about half the size of the calyptra, oblong, subcylindrical, about 5-strata, dividing to the base into 4 valves. Elaters slender, sub- obtuse, about half the breadth of the minute spores. Amentula S antical, proceeding from the axil of the stipule, short, incurved : bracts 5, rarely up to 10-pairs, ovate, concave or subcomplicato- convolute, apex truncate, bifid, or bispinose, rarely entire ; antheridia two longistipitate, rarely solitary. 128 HEPATIC^. 1. Bazzania trilobata (Z.), Gr. 8f B. MtisGoides ierrestris repens, ex obscuro virescens, foliis superioribus et inferioribus ad extremiiatem dentatis, Mich. Nov. pi. gen. p. 10, t. 6. f. 2 (1727). Lichenastrmii pinnulis obtuse trijidis nervo geniculato, Dill. Hist, muse, p. 493, 7, 71, f. 22 (1741). Jungermania trilobata, Linn. Sp. pi. 1599 (175?») ; Hook. Brit. Jung. t. 7G (1816). Bazzania trilobata, Gr. it B. Nat. Arr, Brit. PI. p. 704 (1821). Pleuroschisma trilobatum, Dum. Syll. Jung. p. 70 (1831). Jlerpetium ti'ilobatum, Nees, Nat. Eur. Leb. Ill, p. 49 (1838). ^[astigobryum trilobatuvi, Nees in G.L.N. Syn. Hep. p. 230 (1845). Dioicous, densely csespitose, flagelliferous, large, dark to olive green in colour. Stems on a cross-section showing about 20 cells in diameter, cortical about 90, the 2 or 3 outer layers composed of subquadrate cells with firm walls, equal in size to the inner, which are hyaline with thin walls ; simple or once or twice dichoto- mously branched, erect procumbent or creeping horizontally on the ground, llexuose, rather rigid ; flagella abundant, about an inch long, descending from the postical side of the stem, beset with minute leafy scales ; radiculose, rootlets few, fibrous. Leaves more or less imbricate on the antical side of the stem, horizontal, distichous, semi-broadly-ovate, broadly truncate at the apex, tri- dentate, teeth acute, sinuses acute ; texture firm, epidermis slightly polished, cells medium to rather large in size, roundish- quadrate, angles thickened or with large distinct trigones, walls thick. Stipules transversely inserted (not cordate), oblong quad- rate, coarsely and irregularly dentate, cells with thickened angles. Inflorescence cladocarpous, $ on short postical branches, bracts few, much smaller than the leaves, of a delicate texture, cells elongate, walls thin ; orbicular or subrotund, ciliate at the apex. Perianth highly exserted, lanceolate, cylindrical at the base, which is 4 layers of cells thick, near the middle 2, upper portion triplicate and composed of a single layer of cells, about 170 cells round, mouth small, with few teeth 3 to 4 cells long, 2 to 4 cells wide at the base. Calyptra small, oblong-ovate 4 cells thick at the base, 2 above, pistillidia about 8, long and IIAZZANIA. 129 narrow. Pedicel on a cross-section showing 16 large round cortical ceils with firm walls, inner cells 5 in diameter, extremely delicate, hexagonal or subquadrate. Capsule ovate, dark shining brown, dividing nearly to the base into 4 oblong valves, of a thick texture. Spores reddish-brown, verruculose. Elaters bispiral, similar in colour to the spores, not quite as broad. Androecia postical, minute, amentiform ; perigonial bracts very delicate, 4, 5 pairs, closely imbricate, concave-complicate, bifid or with few teeth ; antheridia solitary, small, shortly stipitate. Var. minus Nees. Usually growing on rotting wood, is much smaller than the type, prostrate, dichotomously branched, branches spreading. Dimensions.— Stems usually 3 to 4 inches long, sometimes 1 to 2, rarely 5; *4 mm. in diameter, with leaves 5' mm. wide; leaves 2*5 mm. x 1"75 at the base and '6 mm. at the apex, teeth •2 mm. long, 3* mm. x 2- mm. near the base and '5 mm. near the apex ; cells "04 mm. x 03 mm., "045 mm. x "04 mm., "04 mm. x *04 ram.; stipules 1' mm. x '8 mm., *8 mm. x 7 mm.; bracts 1*5 mm. X 1" mm., 2* mm. x 1*5 mm., 1*75 mm. x 1" mm. ; cilia •75 mm., '5 mm. ; pistillidia "325 mm. x '06 mm. ; perianth 6* mm. X 1*25 mm., teeth at the mouth •125 ram. ; pedicel '5 mm. diam. ; valves of capsule 1*6 mm. x "5 mra. ; spores "0175 ram. diara. ; elaters '3 ram. x 'Ol rara. ; perigonial bracts '5 mm. x •5 mra.; antheridia 1 ram. x '075 mm. Hab. — Growing in dense patches on damp, shady banks or rocks, rarer on rotting wood. Common in subalpine districts, very rare in fruit. 1, 2, 6-8.. 10-16. I. Found on the Continent and in North America. Obs. — A very striking and one of our finest native species. From B. fricrenafa (Wahlenb.) it is distinguished by its much larger size, broader stems, greener colour, less deflexed leaves and its oblong-quadrate stipules, which are coarsely and irregularly dentate. Mr. Greorge Stabler has found on specimens collected on Black Crag, Staveley, Westmoreland, flagella both branched and bearing male catkins. 130 HEPATIC^. Description or Plate XL VII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 11 (C. & P. Hep. Brit. Exsicc. n. 187). 3-5. Leaves x 11 (ditto). 6. Portion of leaf x 290 (ditto). 7, 8. Stipules x 24 (ditto). 9-11. Bracts x 16 (ditto). 12. Perianth x 11 (ditto). 13. Cross-section of the perianth near the base x 16 (ditto). 14. Ditto lower half near the middle X 16 (ditto). 15. Ditto, near the mouth x 16 (ditto). 16. Teeth, mouth of the perianth x 85 (ditto). 17-19. Perigonial bracts x 24 (Killarney, S. 0. Lindberg). 20. Ditto x 64 (ditto). 21. Antheridium x 85 (ditto). 2. Bazzania triangularis (^ScJdeicher), Lindb. Jungermania triangularis, Schleich. PI. Crypt. Helv. ii. n. 61 (1805). Jungermania dejiexa, Mart. Fl. Crypt. Erlang. p. 135, tab. 3, Fig. 8 (1817). Bazzania triangularis (Schleich.), Lindb. Hep. in Hep. 499 (1874). Dioicous, loosely csespitose, or creeping, flagelliferous, small, brownish-green or olive-brown. Stems dichotomously innovantly branched, innovations proceeding from the axil of the stipules, slender, frao:ile, cortical cells about 20, about 8 in diameter, all similar ; flagella abundant, microphyllous ; radiculose, rootlets fasciculate, hyaline. Leaves incubous, horizontal or patent- divergent, distant or approximate, caducous, deflexed, convex or plane, oblong-ovate or ovate, apex entire acute or irregularly bi- tridentate ; cells smallish to medium size, roundish, walls firm, trigones large and distinct. Stipules patulous, orbicular, sub- rotund or obovate, entire, retuse, emarginate, or irregularly crenate. Androecia postical, proceeding from axil of stipule, bud-like, globular. Perigonial bracts 4, 5 pairs, concave-complicate, broadly ovate, bidentate, sinus and segments small, acute; bracteole oval- orbicular, retuse. Antheridia solitary, oval, stipitate. Dimensions. — Stems |- to 1 inch long, -15 mm. to "2 mm. in diameter, with leaves 1" mm. wide; leaves "6 mm. x "4 mm., 7" mm. X '4 mm., "7 mm. x '35 mm., 1'2 mm. x '6 mm., 1*5 mm. X "7 mm.; cells "03 mm.; stipules '4 mm. x "35 mm., '25 mm. X '25 mm., 2 mm. x 2 mm.; perigonial bract '4 mm. i BAZZANIA. 131 X '3 mm. ; sinus '05 mm. ; perigonial bracteole '3 mm. x "25 mm. ; antheridia '115 mm. x '09 mm. Hab. — Grrovving in loose tufts or creeping loosely amongst mosses and other hepatics, on exposed or shaded banks, rocks, or trunks of living trees. 7. Tyn-y-Groes, Merioneth, Dr. Carrington 8f TF. H. P. Cwm Idwal, Carnarvon, W. H. P. 10. Dent, Yorks, George Stabler. 13. Borrowdale, Cumberland, Dr. Carrittgton Sj' IF. H. P. 16. Moidart, West Inverness, ^iS'. WI. Macvicar. I. Killarney, Dr. Carrington and others. Found on the Continent and in North America. Obs. — I am uncertain whether to regard this as the male form of B. tricrenafa, or as a distinct species, or as merely a variety ; the numerous specimens I have been enabled to examine are either sterile or male plants only. It is distinguished from B. tricrenafa by its smaller size, more slender stems, which are usually, but not always, dichotomously innovantly branched, leaves soon falling away, proportionately shorter (except in var. faccida); cells rounder, with large and distinct trigones, stipules patulous, usually sub-entire. In var. faccida the leaves are plane, narrower, often entire and apiculate. From B. Pearsoni (Steph.) it differs in its smaller size, more delicate habit, brownish-green or olive-brown colour, dichotomous ramification, antical base of leaf not large or overlapping, cells with large trigones and broader stipules. Description or Plate XL VIII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 24 (C. k P. Hep. Brit. Ex. 124). 3. Leaf x 24 (ditto). 4, 5. Ditto x 64 (ditto). 6-8. Ditto x 24 (G. & R. Hep. Eur. n. 198). 9. Ditto x 24 (G-. & R. Hep. Eur. n. 401). 10. Portion of leaf x 290 (C. & P. n. 124). 11. Stipule x 24 (G. & R n. 198). 12-14. Ditto x 64 (C. & P. n. 124). 15, 16. Ditto x 85 (ditto). 17. Male amen- tulum X 24 (ditto). 18. Perigonial bract x 64 (ditto). 19, 20. Perigonial bracteoles x 64 (ditto). 21. Antheridium x 85 (ditto). 132 HEPATIC^. 3. Bazzania tricrenata (JVaJilenherg). Jungermania tricrenata, Wahlenb. Fl. Carp. p. 364, n. 1207 (1814). Dioicous, densely or loosely csespitose, flagelliferous, largish to large in size, brownish-green to dark brown in colour. Stems erect or procumbent, slightly branched, branches erect ; cortical cells about 40, similar to the inner cells, only with firmer walls, 15 to 18 cells in diameter; somewhat brittle w^ien dry, flagella abundant, microphyllous, proceeding from the axil of the stipules ; radiculose, rootlets few, fasciculate, hyaline. Leaves approximate or subimbricate, alternate, horizontal, strongly deflexed, when dry involute, semi-cordate-ovate or semi-ovate-oblong, falcate, arcuate at the antical margin, bi-tridentate at the narrow apex ; cells medium size, roundish-quadrate, walls firm, angles thickened or with trigones distinct. Stipules twice as broad as the stem, roundish-quadrate or suborbicular, usually broader than long, margin irregularly crenate or dentate, sometimes emarginate. Inflorescence $ cladocarpous, arising from the axil of stipule ; sub-bracts and sub-bracteole ovate, bifid; innermost bracts ovate, quadrifid. Perianth lanceolate, cylindrical, plicate at the apex, mouth small, denticulate. Pistillidia long, about 7. Capsule not seen. Dimensions. — Stems about 2 inches long, diameter "2 mm., with leaves 1"25 mm. wide; leaves "75 mm. x *45 mm. at apex •175 mm. wide, 1*4 mm. x '9 mm. at apex, "25 mm., 1"3 mm. X '8 mm. at apex, '3 mm., I'l mm. x '7 mm. at apex, "2 mm., •9 mm. X 65 mm. at apex, '25 mm. ; cells '04 mm. x "03 mm., •035 mm. x '03 mm., '03 mm. x '025 mm. ; stipule '4 mm, X '5 mm., ^3 mm. x '5 mm., '35 mm. x *35 mm. ; sub-bracts •55 mm. x "45 mm., '5 mm. x "4 mm., "6 mm. x "5 mm. ; sub- bracteole "4 mm. X "3 mm. ; innermost bracts *5 mm. x "4 mm. segments, '25 mm., "6 mm. x "4 mm. segments '3 mm. ; pistillidia •2 mm. X '05 mm. Hab. — Growing in large tufts on exposed or shaded banks in subalpine localities. 7, 10, 12, 13, 15-17. I. BAZZANIA. 133 Found on the Continent and in North America. Obs. — Differs from B. frilobafa (L.) in its smaller size, brownish colour (not pale or dark green), more deflexed, and when dry, involute leaves, which are shorter and narrower, witli narrower apices, teeth longer and more acute. From B. triangularis (Schleicher) and B. Pearsoni (Steph.) ; see notes under each species. Description of Plate XLIX. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 24 (C. & P. Hep. Brit. Ex. n. 266). 3, 4. Leaves x 31 (ditto). 5, 6. Ditto x 31 (C. & P. n. 122). 7, 8. Ditto x 31 (C. & P. n. 123). 9. Portion of leaf X 290 (Rosthwaite, Cumberland, Carrington & Pearson). 10. Stipule X 31 (C. & P. n. 122). 11, 12. Ditto x 31 (C. & P. n. 123). 13. Stipule x 31 (C. & P. n. 266). 14-16. Sub-bracts X 24 (Killarney, Stewart & Holt). 17. Sub-bracteole x 24 (ditto). 18, 19. Innermost bracts x 24 (ditto). 20, Perianth X ? (from Lindenb. Spec. Hep.). 21. Pistillidium x 85 (Killarney, Stewart & Holt). 4. Bazzania Pearsoni {StcjjJiani). Masligohryum Pearsoni, Stephani, " Hedwigia " (1893). Dioicous, loosely caespitose, flagelliferous, largish, ochraceous. Stems erect or procumbent, simple or with one, rarely two short branches, fiexuose, filiform, uncinate at the apex, on a cross-section cortical cells about 25, with thick walls, inner more delicate, 8-10 cells in diam. ; flagella few, raicrophjdlous ; radiculose, rootlets few, hyaline, fasciculate, delicate. Leaves distant, alternate, much deflexed when dry, somewhat fragile, reniform, with large over- lapping base, apex acute or truncate, entire or irregularly bi- tridentate ; cells small, roundish-quadrate or elongate, guttulate, walls thick, angles very much thickened, no trigones. Stipules patulous, convex, slightly decurrent, obovate, ovate or orbicular, entire or retuse. Inflorescence ? cladocarpous, arising I'rom axil of the stipules. Sub-bracts small, few (4), ovate, bitid ; bracteole obovate, entire. Innermost bracts ovate, bi-tridentate, margin 134 HEPATIC^. entire or denticulate. Pistillidia small, few (4-6). No perfect perianth seen. Dimensions. — Stems 2 inches long, 2 mm. in diameter, with leaves 1'5 mm. wide; leaves 1- mm. x 75 mm. ; cells '03 mm. x •02 mm., '025 mm. x '02 mm., "035 mm. x 02 mm. ; marginal cells "02 mm. x "02 mm., "015 mm. x '02 mm. ; stipules '5 mm. x '4 mm., *4 mm. x '3 mm., '5 mm. x '5 mm., '4 mm. x '3 mm. ; sub-bracts -4 mm. x -3 mm., "3 mm. x '2 mm, ^ sub-bracteole •4 mm. X -3 mm.; innermost bracts "3 mm. x '2 mm,, 'o mm. x •35 mm. ; pistillidia '125 mm. x '04 mm. Hab, — Very rare, the only known stations being on banks and rocks Eagle's Nest, Horses Grlen, Cromaglown, Killarney, Steivart 8f Holt, June 1885. Obs. — Differs from B. fricreuata (Wahlenb.) in its ochraceous colour, the proportionately broader leaves with large overlapping, antical base, smaller cells with thicker walls and much thicker angles. From B. triangularis (Schleich.) by its larger size, ochra- ceous colour and cell structure. Description of Plate L. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 24. 3-5. Leaves x 24. 6. Portion of leaf x 290. 7-13. Stipules x 24. 14-17. Sub- bracts X 24. 18. Sub-bracteole x 24. 19, 20. Innermost bracts x 85. 21. Pistillidium x 85. Genus 13. KANTIA. Gr. 8^- B. Mnium, Dill. Hist. muse. p. 236, n. 5, et p. 237, n. G (1741); Linn. Sp. PI., 1 ed., 2, p. 1114, nn. 16 et 17 (1753). Jungermania, Mich. Nov. pi. gen. p. 8, n. 2, tab. 5, Fig. 14 (1729). Calypogeia, sect. B. Amphif/astriatce, Radd. in Att. soc. Modena, 18, p. 44, tab. 6, Figs. 3 and 4 (1818). Kcmtia, Gr. & Benn. Nat. Arr. Brit. PL 1, p. 706, n. 28 (1821). Citicinnulus, Dum. Comm. bot. p. 113 (1822). Plants rather large, depressed csespitose, pale green or olive, rarely brownish, prostrate or with apices (often gemmiparous), assurgent. Stems moderately firm, simple, rarely furcate, KANT I A. 135 branches postical axillary to the stipules. Leaves rather large, imbricate, incubous, alternate, plane or convexiilous, oblong, some- times twice as long as broad, often subrhomboid, entire, apex rotundate or retuse, rarely acute, bidentate or bilobed ; cells lax, rather large, rarely large or of medium size, quadrate or oblong- hexagonal. Stipules everywhere present, about half the size of the leaves, subrotund, rarely reniform, apex entire, retuse or bifid, margin entire, rarely denticulate or spinulose or on both sides unidentate or lobulate ; often very delicate. Flowers hypogenous, produced from the axils of the stipules, dioicous or monoicous (sometimes paroicous). Bracts ? 2-3 pairs, tristichous, much smaller than the leaves, subrotund, oval, or lanceolate, entire or 2-4-fid, subconnate. Pistillidia about 12. Perianth pendulous, attached by one side of its mouth to the stem, pendant or descending into the earth, clavate or subcylindrical, carnose, pluristratose, hairy. Calyptra membranous, a little shorter than the perianth, attached to the perianth, but free in the upper fourth part. Capsule on a long, firm pedicel, cylindrical, 3-4 times longer than broad, bistratose, inner layer with semi- annular fibres, valves linear, twisted, deeply coloured. Elaters filiform, somewhat short, subobtuse, bispiral. Spores minute, about the same diameter as the elaters, scaberulous. 1. Kantia Trichomanis (Z.). Gr. 8f B. Jungermania terrestris rejiens, foliis ex rotundinate acuminatis, bijidis, apertura 2Jene visihili, Mich. Nov. pi. gen. 8, t. 5, f. 14 (1739). Mniuvi Trichomanis facie, foliis integris, Dill. Hist, muse, p. 23G, t. 31, f. 5 (1741). Mniicm Trichomanis facie, foliis hifidis, Dill. Hist, muse, p. 237, t. -M, f. 6 (1741). Mnium IVichomanis, Linn. Sp. pi., p. 1571) (17");')). Milium flssnm, Linn. Sp. pi., p. 1579 (1753). Janyermania 7'richoma7iis, lYicks. PI. crypt. Biit. fasc. Ill, t. 8, f. 5 (1793); Hook. Brit. Jung. t. 79 (1816). Junyermania fissa, Scop. Fl. Carn. 2, p. 348 (1772). Kantia Trichomanis, Gi\ k B. Nat. Arr. Brit. PI. p. 70(1 (1821). 13G HEPATICjE. Cincinnidus Trichomanis, Diiiu. Comm, Bot., p. 113 (1822). Calypogeia Trichomanis, Corda in Opiz. Nat., p. 653 (1829) ; in Sturm Deutsch. Fl. fasc. 19, p. 38, t. 10 (1835). Heteroicous (monoicous and paroicous), depresso-csespitose, medium to largish in size, pale to dark green in colour. Stems prostrate or apices assurgent, moderately firm, simple or rarely furcate, branches postical, elongate, attenuate, gemmiparous ; radiculose, rootlets many, proceeding in tufts from base of the stipules. Leaves usually small at the base and at apex, largest at the middle of stem, imbricate, incubous, alternate, patent-divergent or horizontal, slightly convex, ovate, entire, apex rotundate or rarely acute, or very slightly bidentate, cells lax, oblong-hexagonal or hexagonal, medium to rather large, walls thin, trigones very small. Stipules nearly half the size of the leaves, about twice as broad as the stem, subrotund, bifid to about a third, refuse or subentire, lobes obtuse, sinus obtuse, margin entire. Bracts very small, 2 or 3 pairs, oblong-quadrate, slightly bifid. Perianth arising from axil of stipule, pendulous, subterraneum, cylindrical or subclavate, carnose, composed of several layers of cells, near base 6 layers, near the mouth surrounded by layers of small and broadly subulate scales, which are often found on other parts of the perianth; radiculose, rootlets long delicate hyaline, interior layer covered with long papillae, composed of single, elongated cells. Calyptra free, 2-4 cells thick near the base, pistillidia long, narrow, about 10. Pedicel thick, about the same diameter as the capsule. Capsule cylindrical, 3 to 4 times longer than broad, composed of two layers, valves spiral, linear. Spores pale brown. Elaters reddish-brown, 20-30 turns of the spiral, bispiral. Androecia bud-like, sometimes catkin-like, postical, situated in the axil of the stipule, near the $, perigonial bracts very small, about 3 to 4 pairs, subrotund, trifid, segments acute, sinus obtuse, antheridia about 10, oval. Gemmae often pedicellate, in spherical heads, yellow-green. Dimensions. — Stems f to 1^ inch long, '3 mm. to '4 mm. in diameter, with leaves 3* mm. broad; leaves 2* mm. x 1*5 mm., sinus "1 mm. ; cells '04 mm. x "03 mm., "07 mm. x '04 mm., KANTIA. 137 •05 mm. X "04 mm. ; trigones "0075 mm. ; stipules 75 mm. x •8 mm,, segments "25 mm., 7 mm. x •9 mm., seg. '2 mm., '55 mm. X "6 mm., seg-. "05 mm., "5 mm. x "G mm. : bract "15 mm. x •! mm. ; perianth 5' mm. x 1* mm. ; scale near the mouth of the perianth •2 mm. X "125 mm., projecting cells, interior of the periantli •075 mm. x 'OS mm., '06 mm. x "035 mm. ; pistillidia '25 mm. x •03 mm. ; pedicel 75 mm. diam. ; capsule 3-25 mm. x 75 mm. ; spores '015 mm. diam. ; elaters '2 mm. x "02 mm. ; perigonial bracts "3 mm. x '25 mm., seg. -1 mm. ; antheridia "125 mm. x •1 mm. Obs. — This is a very distinct species, and cannot well be eon- founded with any other, except the following, wliich see. The attenuate, gemmiparous stems, the rotundate, acute or slightly bidentate leaves, the lateral, pendant perianth and cylin- drical capsule distinguishing it from other species. Kaiitia argufa (Mont.), the other British species of this genus is a much smaller, more delicate plant, with deeply bidentate leaves, and is dioicous. Hab. — Growing in moderate-sized patches in bogs, on damp or wet banks in woods, &c. Common. 1-17. I. Generally distributed on the Continent and in North America. Description of Plate LI.^Fig. 1. Plants nat. size. 2. Stem, antical view x 11 (Loch Bray, Prof. Lindberg). 3. Portion of leaf, showing sinus x 24 (ditto). 4. Portion of leaf x 290 (ditto). 5. Stipule x 24 (Goathland, York., Slater). 6, 7. Stipules X 24 (Loch Bray, Prof. Lindberg). 8, 9. Ditto x 24 (Ottawa, Macoun). 10. Bract x 24 (ditto). 11. Perianth x 11 (Tyn-y-Groes, C J. Wild). 12. Scale, near mouth of perianth x 85 (ditto). 13. Projecting cells from interior of the perianth x 85 (ditto). 14. Capsule x 11 (ditto). 15. Perigonial bract x 85 (Closter, N. America, C. F. Austin). IG. Antheridium x 85 (ditto). 17. Perigonial bracteole (ditto). 138 HEPATIC^. 2. Kantia Sprengelii {Mart.). Junyermania Sjjrengelii, Mart. FI. Crypt. Erlang., p. ];5o, t. ?>, f. 6 (1817). Cincinmdus Sprengelii, Dnm. Syil. Jung., p. 73 (1831). Ccdypoyeia Trichomanis, var. Sprengelii, Nees Eur. Leb., Ill, p. !) (1838). Heteroicous (monoicous and paroicous), cpespitose, medium size, pale to dark-green in colour. Stems prostrate or suberect, simple or furcate, attenuate often at the apex and gemmiparous ; radiculose, rootlets plentiful, proceeding from the base of the stipules. Leaves approximate or imbricate, horizontal or patent- divergent, broadly ovate, apex entire, acute or rotundate or acutely and shallowly bidentate, sinus and segments acute, margin entire ; texture lax ; cells rather large, hexagonal or oblong-hexagonal, walls thin, nucleate granules clinging to the walls making them appear thicker, trigones very minute or absent. Stipules about a fifth to a fourth smaller than the leaves, about twice as broad as the stem, irregular in shape, suboblate to subreniform, bifid to about a third, sinus wide, acute or obtuse, segments spreading acute or obtusate, lobate, outer lobe smaller. Male and female flowers proceeding from axil of stipules, sometimes the two pro- ceeding from the one axil. Perfect fruit and male flowers I have not seen. Dimensions. — Stems from ^ to 1 J inch long, "3 mm. in diameter, with leaves 25 mm. wide; leaves 1-6 ram. x 1*2 mm., sinus •01 mm., 1-4 mm. x I'l mm., 1'3 mm. x 1- mm., sinus '01 mm.; cells "05 mm. x '05 mm., "06 mm. x '05 mm., "05 mm. x "04 mm. ; stipules "25 mm. x "3 mm. segments "1 mm., "4 mm. x "5 mm., seg. "2 mm., '4 x '6 mm. seg. 2', '5 mm. x "6 mm., seg. '2 mm. Hab. — Growing in patches on damp shady banks, in woods, &c. Somewhat rare, probably often mistaken for Kantia Tricho- manis. 1. Penzance, Cornwall, IF. Ciirnow. 4. 7. 10. Castle Howard, Yorks, M. B. Slater. Shipley Glen, JF. West. 16. Found on the Continent and in North America. Obs. — Distinguished from. K. Trichomams (L.) by the laxer texture of the leaves, the thinner cell- walls, stipules irregular in KANTJA. 139 shape, more deeply and acutely divided, segments spreading, often acute and ao^ain lobate, outer lobe smaller. It may be considered by some authorities as only a variety of K. Trichomanis, but as the distinguishing characters are always permanent, I give it as a species. Description of Plate LIT.— Fig. 1. Plants nat. size. 2. Portion of stem, postical view x '24 (Castle Howard, M. B. Slater). 3. Leaf x 24 (Gr. & R. n. 587, as Cali/jjo/jia TricJiomcmis adscendens). 4, 5. Leaves x 24 (Suffolk, Skepper). G. Portion of leaf X 290 (Castle Howard, M. B. Slater). 7. Stipule x 24 (G-. & R. n. 587). 8, 9. Stipules x 85 (ditto). 10. Stipule x 24 (Suffolk, Skepper). 3. Kantia arguta [Mont, et Nees), Lindh. Cal;/pogeia m-yuta, Montagne et Nees in Nees Kat. Eur. Leb. Ill, p. 24 (1S38). Cincinnulus argutus, Dum. Hep. Eur. p. 117 (1874). Kantia aryuta, Lindb. in Not. soc. F. FI. Fenn. 13, p. 363 (1874). Dioicous, laxly and thinly spreading, small, pale green in colour. Stems simple or sparing^ branched, delicate, sometimes elongate, attenuate, depauperate, frequently gemmiparous; branches proceeding from axil of stipules ; radicuiose, rootlets few. Leaves distant or approximate, largest near the middle of stem, smaller below, and often near the apex minute and remote, horizontal to patent-divergent, postical margin usually decurrent, oval or oblong- oval, bidentate to about one-tenth, sinus broad, semilunate, seg- ments a little divergent, acute ; texture fragile, lax, cells rather large, 4-, 5- and 6-sided, walls thick, no trigones. Stipules small, scarcely twice as broad as the stem, irregular in shape, much broader than long, bidentate to below the middle, segments diver- gent, lobate to the middle or below, lobules acute, divergent. I have seen no perfect (^ or ? ; the drawing of the perianth is from a sketch sent me by Mr. Slater, who found a single one on a plant collected by Mr. Curnow near Penzance. Andrcecia on separate plants, bud-like, small, proceeding from axil of stipules. 140 HEPATIC^. Dimensions. — Stem ^^ to f inch long, '2 mm. in diameter, with leaves 1*5 mm. wide; leaves 1* mm. x "6 mm., segments •1 mm. ; cells "07 mm. x '04 mm., "04 mm. x '035 mm., '06 mm. x •04 mm. ; stipules "25 mm. x '35 mm., segments 'l mm., "15 mm. x "25 mm., seg. "075 mm. Hab. — Growing on shady clayey banks, in woods or by road- sides, moderately rare, or overlooked. 1. Near Penzance, IF. Cnmow. 2. Near Ditchling, Sussex, G. Davies. 3. Epping Forest, Essex, ^. M. Holmes. 7. Torrent Walk, Dolgelly, Merionethshire, W. H. P. Near Pandy Mill, Bettws-y-Coed, Carnarvonshire, M. B. Slater. 8. Near Leicester, F. T. Molt. 9. Hazel Grove, Cheshire, Cotterill Clough, Cheshire, G. A. Holt. 10. Park Quarry, Castle Howard, Yorks, G. Stabler, M. B. Slater. Near Mallion Spout, Goathland, M. B. Slater. Scarborough, M. B. Slater. Near Baildon, North- West Yorks, W. West. 12. 14. Helensburgh, Scotland, Br. Graham. T. Kil- larney, D. Moore. Connor Hill, Prof. Lindhercj. Clonallon, Bev. C. H Waddell Found on the Continent (France) and introduced in green- houses in North America. In a collection of Hepaticas made near Natal, South Africa, by Mrs. Ellen Sophie Bertelsen, widow of a Norwegian missionary, specimens of this species were found, agreeing in ever}^ respect with ours. Two other British species were found in the same collection, namely, Lejeimea hamatifolia (Hook), and Cejjhalozia connivens (Dicks.). Obs. — A ver}^ distinct species. Distinguished from K. Tri- ckomanis (L.) and K. Sprengelii (Mart.) by its dioicous inflorescence, much smaller size, leaves more oblong ; wider, semilunate sinus, divergent segments, much laxer and more delicate texture, small stipules which are deeply bidentate, with segments divergent and again lobate, perianth narrower than in K. Trichomanis. In habit it resembles small forms of Lophocolea hidentata (L.), from which it is at once distinguished, even when barren, by its less bidentate eaves and smaller stipules. Description of Plate LIII. — Fig. 1. Plants natural size. CEPHALOZIA. Ml 2. Portion of fertile stem, postical view x 16 (Penzance, Curnow). 3. Portion of stem, antical view x 24 (ditto). 4. Stem, postical view X 16 (Natal, Soutli Africa, Mrs. Bertelsen). 5. Portion of leaf X 290 (Penzance, Curnow). 6. Stipule x 85 (Epping Forest, Holmes). 7, 8. Stipules x 85 (Natal, Mrs. Bertelsen). 9. Stipule X 85 (Penzance, Curnow). Genus 14. CEPHALOZIA, Bum. Lichenastrum, Dill. Cat. pi. Giss. p. 213 (1718), et Hist. muse. p. 181, n. 4 (1741). Jtmgermania, Mich. Nov. pi. gen. p. 9, n. 5, tab. 6, Fig. 17 (1729). „ sect. Cephalozia, Dum. Syll. Jnng. Eur. p. 60 (1831). „ sect. Bicuspides, Nees, Nat. Eur. Leber. 11, p. 211 (1830). Cephalozia, Dum. Eecueil, 1, p. 18, n. 21 (1835). Zoopsis, Hook. f. et Tayl. Crypt. Antarct. p. 55, n. 22 (1845). Trigonanthus, Spruce, Trans. Bot. Soc. Edinb. (1849). Nowellia, Mitt, in C.-Godman Nat. Hist. Azor., p. 321, n. 12 (1870). Phnroschisma Odontoschisma, Dum. Syll. (1831). Odontoschisma, Dum. Kecueil (1835). Sphagnoecetis, Nees in G. L. N. Syn. Hep. (1845). Protliallium slender, linear or almost filiform, consisting of only a single (more rarely in part of a double) series of cells ; either simple or subramose ; often passing at the apex insensibl}^ into the stem and persisting a long time. Plants usually small and tender, in only a few species rather robust ; of almost all shades of green and brown, or whitish and pellucid, sometimes tinged with rose ; growing in depressed matted tufts, or flakes, or creeping over Sphagna and other mosses. Stems usually prostrate or procumbent, leafy throughout, or rhizomatous and leafless at the base — very rarely with the leaves reduced to mere scales — still more rarely frondose ; branches all postical, springing from the underside of the stem, and axillary to the stipules where any exist ; radicles usually copious, pale and slender. Leaves mostly succubous, in a few species transverse, in a very few subincubous ; horizontal or assurgent, never deflexed, roundish, or subquadrate, or cuneate, rarely lanceolate, very seldom plane, usually concave. 142 HEPATIC^. and in most species somewhat complicate and bilobed (but never divided to the veiy base, nor with capillary lobes), in a very few species undivided or variable at the apex ; margins uniformly plane or subincurved — never convex or recurved — mostly quite entire, but in a few species toothed. Eeticulation in the typical species lax and pellucid, in a few species denser and subopaque ; cells often subquadrate; cell-walls mostly thin, rarely conspicuously thickened at the angles ; cuticle smooth or scaberulous. Stipules much smaller than the leaves, and oftener undivided at the apex, but in some species subdentate at the margin ; entirel}^ absent from many species (except in the involucre, where they exist in every Cepltalozid). Inflorescence dioicous or autoicous — very rarely paroicous. Androecia amentiform, occupying the whole, or only a part, of a branch, rarely terminal on the stem. Bracts in many pairs, leafy (even where there are no stem-leaves), bifid, uniformly raonandrous. Gynoecia capitate, usually seated on an abbreviated branch (i.e., cladocarpous), but sometimes terminal on larger branches or on the main stem (acrocarpous). Bracts much larger than the subjacent leaves (where any exist on the same axis) tristichous, i.e., with stipules added, even where absent from the rest of the plant, and in three, or more amplexicaul rows ; all cloven (usually bilobed, sometimes 3-5-lobed), and very often toothed or subspinose ; cells elongate. Pistillidia about 20, shortish and flask-shaped. Perianth free, usually very long and narrow and elongate, reticulate like the bracts, fusiform, trigonous — rarely with the angles varying from 3 to 5 or 6 in the same species, but, whenever reduced to 3, with the third angle always postical, mouth truncate, but usually constricted (from the angles becoming more pronounced and plica3form at the apex), variously toothed, ciliate, laciniate, or entire. Calyptra free (superior), with the sterile pistillidia surrounding its base. Capsule on a long pedicel (which at the calceolate base buries itself deeply in the fertile branch), oblong or sub-cylindrical — usually about twice as long as broad, but in the subgenus Cephalozidla often shorter, oblong- globose- 4- vaived to the base ; capsule-walls of two layers of cells, EUCEPHALOZIA. ] 43 whereof the inner are strengthened by semiannular fibres. Elaters elongate bispiral, about as wide as the diameter of the smooth or scaberulous spores. Propaguhi apical, minute, red or wliitish, polyhedral or amorphous ; rarely present, except in a very few species. Subgenus 1. EUCEPHALOZIA, Spruce. Plants of a moderate size, rarely small or robust, virescent rarely tawny or dull brown, sometimes with a rosy tinge, growing- in broad tufts, or creeping amongst mosses in marshy places. Stems for the most part tender and fragile, rarely somewhat rigid, cortical cells sometimes large and pellucid, simple, very rarely furcate, postical more or less branched, in a few species flagelli- ferous. Leaves obliquely inserted, rarely subtransverse, always broader than the stem, often moderately large (between 0'3 and 1"35 mm. long), more or less oblong, concave or laxly complicate, rarely subplane, bifid, very rarely 3-4-fid, sinus rarely deep, in some subacute, in others lunate; apex of the segments various, but rarely rotundate or cuspidate ; margin entire. Stipules (in a few species normally present) small, entire or bifid. Cells of the leaves moderately constant in size, varying in dift'erent species in diameter between ^}y^ and A- mm., rarely almost large (Jg mm.), very rarely small (^V-^'s" nim.), equilateral-hexa- gonal, or often quadrate-hexagonal or quadrate, in most species subpellucid, in very hw, wall thickened at the angles, cuticle almost smooth ; cells of the bracts and perianth generally larger, rectangular-oblong. Inflorescence dioicous, or monoicous, very rarely paroicous ; ? in some species constantly cladogenous, in some now clado- and now acrogenous, or almost acrogenous, but yet sometimes terminal on the same stem. Bracts 3 pairs, rarely fewer, innermost large bifid, rarely 3-4-fid, entire or very often dentate, spinulose, or incised, free or with the bracteole subcon- formable, of equal length or shorter, connate at the base. Perianth more or less highly emersed, fusiform — sometimes almost linear- trigonous-prismatic, in some species the keels in all stages acute. 144 HEPATIC^. in others (fruit mature) partly obliterated or only at the apex to be seen, mouth constricted denticulate, setulose, ciliate or laciniate ; walls leptodermous, except at the very base, where it changes into the stem or branch, or in a few species, the lower half 2-3 cells thick. Andro3cia spicate or amentiform, in different places, but very rarely hypogenous, $ bracts immediately below the female flowers. 1. Cephalozia catenulata {Huben.) Jungermania catenulata, Hiiben. Hep. Germ. p. 169 (1834). Jungermania reclusa, Tayl. Lond. Journ. Bot. p. 278 (1846). Jungermania hicuspidata, var. ericetorimi, G.L.N. Syn. Hep. p. 139 (1844). Cephalozia serrifiora, Lindb. Medd. Soc. Fennica (1878). Cephalozia catenulata (Hiiben.), Spruce, "On Ceph." p. 33 (1882). Dioicous, usually cladocarpous, densely but thinly caespitose, flagelliferous, small, brown in colour. Stems prostrate, sub- pinnate, cells of the stems about 6 in diameter, cortical 14, inner slightly larger and more pellucid ; branches radiculose ; rootlets whitish. Leaves small, subimbricate, slightly concave, when dry much incurved, catenulate, oval-rotund, slightly decurrent, bifid to about the middle, sinus more or less obtuse, segments patulous or subconnivent, acute, 5 rows of cells, 1, 2, 3, 4, 5 wide; cells small to smallish, subquadrate or rhomboid, leptodermous, chloro- phyllose and subopaque, walls firm, no trigones. Stipules wanting. Branches % postical, short, rarely subelongate. Bracts twice as large as the leaves, oblong, bifid to about the middle, segments subulate-acuminate, margin denticulate or spinulose ; bracteole similar, or entire and broadly subulate, free, margin denticulate or spinulose. Perianth deeply emersed, linear- fusiform, 4 times longer than broad, composed of one layer of cells, the whole length deeply tri-carinate, mouth constricted, setose or ciliolate. Calyptra delicate. Capsule reddish-brown, oval-cylindrical. AndrcBcia terminal on postical branches, jula- ceous ; perigonial bracts somewhat similar to the leaves and of the same size, few pairs; perigonial bracteoles ovate-lanceolate, entire or with a slight notch on one side near the middle ; anther- idia oval. EUCEPHALOZIA. 145 Dimensions. — Stems J to |- inch long, "1 mm. in diameter, with leaves 5 to '6 mm. wide ; leaves 3 mm. x 2 mm., "25 mm. x •17 mm., 2 mm. x 2 mm., segments '1 mm., -225 mm. x "175 mm., seg. '075 mm., '25 mm. x 'IS mm., seg. -1 mm.; cells "025 mm., •025 mm., '0225 mm. x "0225 mm., 02 mm. x "0225 mm., '03 mm. X "02 mm. ; bracts "55 mm. x '4 mm., '6 mm. x •SS mm., segments '3 mm.; bracteole '35 mm. x 175 mm.; perianth 1*3 mm. x "35 mm., 1*4 mm. x 4 mm., teeth at the mouth '15 mm., '1 mm., "075 mm.; perigonial bracteoles '12 mm. x "07 mm.; antheridia "125 X '1 mm. Hab.— On rotting wood, turfy banks and shady rocks. Rare. 2. Tunbridge Wells, R. Spnice. 10. Blaeberry Gill, near Whitby, M. B. Slater. 12. Borrowdale, Cumberland, W. H. P. 13. N. of Black Craig, New Galloway, ./. Mc Andre w ; Col vend, Kirkcudbrightshire,./. Mc Andrew. 15. Banchory, Aberdeenshire, /. Sim. I. Cromaglown and other places in the S.W., Taylor, Spruce, Moore, Carrington and others. Eound on the Continent and in North America. Obs. — Similar in size and habit to Cephalozia hmulcefolia Dum. but more rigid, leaf cells smaller, bracts denticulate or spinulose, perianth composed of a single layer of cells. Ceplialozia pallida Spruce is much paler in colour, more branched, branches subfastigiate, leaves subdecurrent, bifid from \ to \, cells rather larger, bracts entire. Description of Plate LIV. — Fig. 1. Plants natural size. 2. Stem, antical view x 64 (Blaeberry Gill, M. B. Slater). 3. Leaf X 85 (ditto). 4, 5. Leaves x 85 (Canada, Macoun). 6. Portion of leaf X 290 (Blaeberry Gill, M. B. Slater). 7, 8. Bracts x 85 (Canada, Macoun). 9. Bracteole x 85 (ditto). 10. Perianth x 24 (G. & E. n. 594). 11. Portion of the mouth of perianth x 85 (ditto). 12, 13. Perigonial bracteoles x 85 (ditto). 14. Anther- idium X 85 (ditto). 146 HEPATIC^. 2. Cephalozia pallida, Spruce. Cephalozia catenulata var. pallida {Ceph. pallida nobis in hb.), Spruce, On Cephalozia. (1882.) Dioicous, cladocarpous, depresso-csespitose, eflagelliferous, small, pale green to yellowish in colour. Stems much branched, branches subfastigiate ; cortical cells 15, rather larger than the inner ones, 5, 6 cells in diameter. Leaves small, approximate, from patent- divergent to erecto-patent, orbicular to oval-rotund, subdecurrent, from a third to the middle bifid, rarely more, sinus obtuse, seg- ments acute or obtuse, connivent ; cells smallish to medium in size, subquadrate, walls thick, no trigones or thickened angles. Stipules wanting. Bracts larger than the leaves, oval-rotund, bifid to a third, segments acute or obtuse, margin entire, bracteole similar, sub-bracts and sub-bracteole similar but much smaller. Perianth projecting much beyond the bracts, linear-fusiform, composed of a single layer of cells, obtusely trigonous, mouth ciliolate or setulose, ciliola 2 cells long. Calyptra moderately coarse; pistillidia few, 8-10. Capsule oval, dark-brown. Spores pale yellow. Elaters slightly broader and of a darker colour, loosely bispiral. A ndroecia on short postical branches ; perigonial bracts closely imbricate, erect, oval-rotund, concave, bifid to about a third, with usually a third smaller tooth near the middle, seg- ments acute ; antheridia oval. Dimensions. — Stem ^ inch long, "1 mm. in diam., with leaves *75 mm. wide; leaves "35 mm. x "3 mm., segments "2 mm., *4 mm. X "3 mm., seg. '175 mm., '4 mm. x "3 mm., seg. "2 mm. ; cells '03 mm. x -03 mm., '03 mm. x '025 mm., "04 mm. x '03 mm. ; sub-bracts '45 mm. x "45 mm., seg. '15 mm., "5 mm. x "45 mm., seg. '2 mm., '4 mm. x "25 mm., seg. '15 mm. ; sub-bracteole •45 mm. x "3 mm. ; bracts "9 mm. x "7 mm., seg. "3 mm., "8 mm. x •7 mm., seg. 3 mm. ; bracteole '75 mm. x '65 mm., seg. "25 mm. ; perianth 2' mm. x 'Qb mm., 1"75 mm. x "6 mm., teeth at the mouth •05 mm.; pistillidia '125 mm. x "025 mm.; capsule "5 mm. x •3 mm., valve of capsule "85 mm. x '225 mm.; spores "01 mm.; elaters "1 mm. x 0115 mm. ; perigonial bracts '5 mm. x '5 mm.. EUCEPHAWZIA. 147 seg. '2 mm.; perigonial bracteole "25 mm. x "1 mm.; antheridia •15 ram. X 1 mm. Hab. — On turfy banks or moors, or on shady rocks. Rare. 9. Frodsham, Cheshire, G. E. Hunt. 10. Strensall IMoor, G. Stabler ; Stockton Forest, G. Stabler ; Yeadon, Dr. Carrhi/jion. 1. Lachan Bay, Co. Mayo, L. Moore. Loch Bray, Co. Wicklow, B. McArclle. Found on the Continent. Obs. — Distinguished from Ceph. catenulafa (Hi'iben.), which it approaches most in habit, by its paler colour (pale green or yellowish, and not a tawny brown), leaves not so deeply bifid, cells slightly larger and bracts entire. From Cejjh. lunidcefolia Dum. by its smaller leaf-cells, and perianth composed of a single layer of cells only. Description of Plate LV. — Fig. 1. Plants natural size. 2. Portion of stem x 24 (Stockton Forest, G. Stabler). 3-6. Leaves x 64 (ditto). 7. Portion of leaf x 290 (ditto). 8, 9. Bracts x 31 (Loch Bray, D. McArdle). 10. Bracteole x 31 (ditto). 11-13. Sub-bracts x 31 (ditto). 14. Sub-bracteole x 31 (ditto). 15. Perianth x 16 (ditto). 16. Cross-section of perianth x 31 (ditto). 17. Portion of the mouth of perianth x 85 (ditto). 18. Perigonial bract x 64 (Stockton Forest, G. Stabler). 19. Perigonial bracteole x 64 (ditto). 20. Antheridium x 85 (ditto). 3. Cephalozia lunulaefolia, Liim. Jungermania connivens, Mart. Fl. crypt. Erl. p. 109, t. 5, f. -li (1817); Moiig. et Nestl. St. crypt. Vog., 5, n. 432. Jungermania hinuhtfoUa, Dum. Syll. Jung., p. Gl (1831). Cephalozia lumdcefolia, Dum. Recueil p. 18 (183.")). Cephalozia media, Lindb. Medd. af Soc. pro f. fl. Fenn. t!, p. 242 (1881). Cephalozia multijlm'a, Spruce, On Ceph., p. 37 (1882). Dioicous, densely depressed ca^spitose, amongst S/)lia(/)ia laxly creeping, eflagelliferous, small, of a pale green colour. Stem prostrate, subramose, sometimes subpinnate, subcompressed, above almost plane, below convex, cells 6 or 7 in diameter, cortical cells 148 HEPATIC^. 12-14, large, pellucid, interior much smaller, subopaque. Leaves small, subimbricate — in the sterile plant often distant — subascend- ing, rhomboid-rotund, antically decurrent, bifid to about ^, sinus obtuse, rarely lunate, segments connivent, acute or subacuminate ; cells of moderate size, leptodermous, pellucid, slightly chloro- phyllose, quadrate-hexagonal, walls moderately thick, no trigones or thickened angles. Stipules wanting. Inflorescence cladocarpous. ? terminal on very short postical branches. Bracts usually about 3 pairs, tristichous, the innermost considerably larger than the leaves, delicate, oblong-rotund, bifid to the third, rarely trifid or twice bifid, segments acutate, entire ; bracteole somewhat similar, often connate with the bracts. Perianth linear-fusiform, when young triplicate, when mature only towards the apex trigonous, mouth subconstricted, denticulate or setulose (setse 1 or 2 cells long only), carnose, towards the base composed of 3 layers of cells, in the middle 2 and near the apex 1. Calyptra oval-globose, carnose, almost the whole length composed of 3 layers of cells. Capsule oblong- cylindrical, on a somewhat short pedicel. Spores of a beautiful cinnamon colour, smooth. Elaters bispiral, slightly darker in colour than the spores. Androecia on short postical branches, perigonial bracts smaller than the leaves, oval, to about a third bifid, canaliculate-concave ; antheridia solitary, oval, shortty stipitate. Dimensions. — Stems |- to 1 inch long, "l mm. in diam., with leaves •5-'8 mm. wide ; leaves "3 mm. x '25 mm.,- 3 mm. x '3 mm., •3 mm. X 4 mm. sinus -175 mm.; cells '04 mm. x "04 mm., "04 mm. '03 mm., '035 mm. x -04 mm., "0275 mm. x '04 mm., "035 mm. x X -03 mm. ; bracts "7 mm x -6 mm. segments. '3 mm., '8 x "65 mm., seg. '2 mm., '9 mm. x "7 mm., seg. "3 mm., 5 mm. x "4 mm. seg. "2 mm.; bracteole "7 mm. x b mm. seg., 2 mm., '5 mm. x '3 mm., seg., •15 mm. ; perianth 1'25 mm. x '4 mm., 1*25 mm. x -Gmm., 2'1 mm. X '7 mm. ; teeth of the perianth "075 mm. long; pistillidia •2mm. X '05 mm.; capsule '45 mm. x -2 mm. ; spores "0125 mm. ; elaters •15 mm. X '015 mm.; perigonial bracts ^225 mm. x -125 mm., seg. •075 mm. ; antheridia •ll mm. x ^09 mm. Hab. — On shady heathy banks, chiefly in woods and on rotting trunks, m(U*e rarely on sandstone rocks, often fruiting luxuriantly. EUCEPHALOZIA. 149 also on iSpkafjna and other bog mosses where it is usually sterile Common. 2. Tunbridge Wells, Ardingley Rocks, Sussex, G. Davies, E. M. Holmes. 7. Barmouth, Br. Carnufjton ; Tyn-y-Grroes, W. H. F. 8. Kinder Scout, Derbyshire, ./. Whitehead and G. A. Holt. 9. Carrington Moss, Cheshire, G. A. Holt; Alderley, Cheshire, lVilddk\\([ Holt. 10. Castle Howard, Dr. Spruce; Blaeberry Gill, Whitby, M. B. Slater; Esholt, Dr. Carrington, W. West. 12. Foulshaw Moss, Westmorland, G. Stabler ; Borrowdale, Cum- berland, Dr. Carrington 2i\\.^ W. H P. 13. Dumfries, ./. Mc Andrew. 15. Forfar. \7a. Sutherland, Dr. Grerille. I. Found on the Continent and in North America. Obs. — " C. lunidiefolia may be distinguished from C. bicuspidata and comiivens, and from most of their allies, by the dioicous inflorescence ; the small leaves, obtusely cloven to only ^ of their length, and rather more closely reticulate ; the bracts far less deeply cloven, and rarely into more than two segments ; but, above all, by the Jle.shg jjerianth and ealgptra, the perianth being 3 cells thick below and 2 cells thick about the middle, and the calyptra 3 cells thick cdmost up to the very apex : whilst both these organs in C. hictispidata and comiwens consist throughout of but a single layer of cells. Moreover, the perianth is merely denticulate at the mouth, while that of C. conniveiix has the almost unique character, among European Cephalozice, of terminating m long- cilia ; the perianth of C. catenidata being merely ciliolate, or setose at the apex. The purple spores of C. bicuspidata alFord an addi- tional mark of distinction from C. luiiuhefoliar R. S. In 1893 I received the iollowing interesting note from Dr. Spruce : " In writing for a set of my hepaticcTe, C. Delogne (who has charge of the herbarium at the Brussels Bot. Garden) asked for specimens of some of my British Cephcdozia and Lejeunece, which I seiit him. I took the opportunity to ask if any specimens of Dumortier's hepaticse were accessible. It would seem that Dumortier left no herbarium of hepaticse, and there are only a few specimens from him in the national herbarium, where, however, he occasionally worked and annotated some of the specimens. When I was studying the ('ephalozia' I wished very much for a 15U HEPATIC^. sight of some of Dumortier's originals, but in vain. Having convinced myself of the distinctness of C. muWfora (nob.) from C. connivens, I sought for some notice of the former in the works of previous authors, and it seemed to me possibly identical with C. lunidafolia Dum. The name suited well enough — not so well the meagre diagnosis. The first question I asked M. Delogne was about this plant, and he has been good enough to send for my inspection two specimens from the herbarium, the one marked by Dumortier (in the earlier part of his career) Jg. limulata Dum., the other Ceph. hmidafoUa Dum. The latter from the classic exsiccata of Moug. and Nestler is conclusive. I am glad of this, as it gets rid of two names, mulfiflora and media, which were likely to lead to a fight." Description of Plate LVI. — Fig. 1. Plants natural size. 2. Plant X 24 (Moug. & Nestl. n. 432). 3. Portion of stem X 31 (ditto). 4. Ditto x 64 (ditto). 5. Portion of leaf X 290 (ditto). 6-9. Bracts x 24 (Blaeberry Gill, M. B. Slater, herb. Spruce). 10. Bracteole x 24 (ditto). 11. Bracts and bracteole x 31 (Carr. & Pears. Hep. Brit.). 12. Perianth X 31 (Moug. & Nestl. n. 432). 13. Cross-section of perianth, upper half x 24 (Carr. & Pears. Hep. Brit. Exs.). Portion of the mouth of the perianth x 85 (Blaeberry Gill, M. B. Slater). 15. Perigonial bract x 85 (Carr. & Pears. Hep. Brit. Exs.) 16. Antheridium x 85 (ditto). 4. Cephalozia bicuspidata (Z.), Difw. Lichenastrum pin7iulis acutissime bifidis minimum, Dill. Hist, muse, p. 488, t. 70, f. 13 (1741). Jungermania hicusjndata, Linn. Sp. pi. 158'J (1753); Hook. Brit. Jung. n. 11 (1816). Cephalozia bicuspidata, Dum. Recueil, p. 18 (1835). Monoicous, caespitose, flagelliferous, small to medium in size, pale gi'een, dull white or rose coloured. Stem prostrate, or ascending, irregularly ramose, branches few, postical ; flagella radiculose ; rootlets few, more frequent on the fertile branches ; EUCEPHALOZIA . 151 cortical cells about 10, large, pellucid, inner about G in diam., smaller subopaque. Lower leaves smaller and distant, upper larger and subimbricate, diagonal at base or where closer, almost transversely inserted, ovate-orbicular, bilobed to about the middle, concave, sometimes complicate, segments connivent or patent, ovate- lanceolate or subtriangular, postical acute, antical a little narrower, sub-acuminate, sometimes shortly apiculate ; cells rather large, 4-, 5- and G-sided, pellucid, no trigones or thickened angles. Stipules wanting, except rarely on the female flowers. Inflor- escence clado-acrocarpous ; branches ? very short, rarely more or less elongate. Bracts about 3 pairs, innermost almost 3 times longer than the leaves, partly free, to the middle bilobed, segments lanceolate acuminate, entire or towards the base 1-2 spinose, rarely sublaciniate; bracteole similar; sub-bracts smaller, roundish; sub-bracteoles also smaller, lanceolate, spinulose, rarely bifid, sometimes obsolete. Perianth 4 times longer than the leaves, linear, prismatic, or subfusiform, mouth constricted or rarely open, denticulate, setulose or ciliolate ; at first tricarinate at the base, when older subterete, above trigonous, areolation lax, pale green or white, sometimes below of a beautiful purple colour, apex canescent, the whole length (except at the very base, which is 2, 3 cells thick) composed of a single layer of cells, about GO cells round. Calyptra small, delicate. Pistillidia about 12, small. Capsule cylindrical-oblong. Spores purple, Androecia spicate, perigonial bracts situated at the apex or middle of the branches, or occupying the whole length, very rarely immediately below the $ flowers, closely imbricate, ascending, similar to the leaves, but broader at the base, which is swollen, sometimes v/ith a third, small, antical tooth. Antheridia oval, single. Dimensions. — Stems Jto 1 inchlong, 1 mm. in diam., with leaves •5 to 10 mm. wide; leaves "55 mm. x '55 mm., 5 mm. x "4 mm., •7 mm. X 5 mm., segments, "35 mm., 6 mm. x 4 mm., seg. "3 mm., •3 mm. X 25 mm., seg. "15 mm.; cells "05 mm. x 04 mm., '045 mm. X "OSo mm., "04 mm. x 035 mm., "05 mm. x '035 mm., -OG mm. X *04 mm., '05 mm. x -05 mm, ; sub-bract "8 mm. x '8 mm., seg. 5 mm. ; bract 13 mm., 125 mm. x 75 mm., seg. "75 mm,, 15 mm. 152 HEPATIC^. X 1' mm., seg. "75 mm. ; bracteole 1"25 mm. x 1* mm., seg. •6 mm. ; perianth 2* mm., x -75 mm., 2*5 mm. x '85 mm., 35 mm. X r mm., 1'8 mm.-2*2 mm. x "o mm.; teeth at mouth '05 mm., •075 mm.,*l mm. ; pistillidia "15 mm. x "04 mm. ; perigonial bracts "75 mm. x "5 mm., seg. "4 mm., '6 mm. x '6 mm., seg. '25 mm. ; antheridia '125 mm. x "l mm. Hab. — Growing on damp stones, rocks, and earth, or amongst mosses in wet and shady places; rarer on rotting wood. Common. 1 to 17. I. Found on the Continent and in North America. Obs.^ — One of our commonest hepatics, varying extremely in size, colour and habit. Distinguished from C. connivens (D.), which is also monoicous, by the presence of flagella; position, shape and texture of the leaves, and perianth not furnished with long cilia. C. mrvifoUa (Dicks.), which is also monoicous, has no flagella; the leaves are remarkably concave, with long, incurved, hamate, cuspidate segments, and usually of a beautiful purple colour. Description of Plate LVII. — Fig. 1. Plants natural size. 2. Portion of stem x 31 (Pass of Llanberis, G. K. Holt). 3. Ditto X 31 (Penzance, W. Curnow). 4. Leaf x 24 (Kinder Scout, Whitehead & Holt). 5, 6. Leaves x 24 (Tyn-y-Groes,W. H. P.). 7. Portion of leaf x 290 (ditto). 8, 9. Bracts x 24 (Pass of Llanberis, G. A. Holt). 10. Bracteole x 24 (ditto). 11. Sub- bract x 24 (ditto). 12. Bract x 24 (Kinder Scout, Whitehead & Holt). 13. Perianth x 16 (Festiniog, W. H. P.) 14. Cross- section of perianth X 16 (ditto). 15. Portion of the mouth of perianth x 85 (ditto). 16, 17. Perigonial bracts x 24 (ditto). 18. Antheridium x 85 (ditto). EUCEPHALOZIA. 153 5. Cephalozia Lammersiana (^Hiibcn.), ISpnice. Jungermania Lammersiana, Hiibener Hep. Germ. p. 165 (1834). Jungermania bicicsjndata y uliginosa, Nees Eur. Leberm. ii. 253 and (ex parte) yy obliquata, Nees 1, c. 254. Jungermania bicuspidafa, Eng. Bot. t. 2230. Cephalozia Lammersiana (Hiiben.), Spruce, On Ceph., p. 43 (1882). Dioicous, laxly csespitose, flagella none or very rare, medium to largish in size, pale green to pale dull brown in colour. Stems simple or with one or two short postical branches ; slender, delicate; cortical cells large, about 10, inner A, 5 diam, smaller; radiculose, rootlets few, white, delicate. Leaves distant or approxi- mate, alternate, spreading, horizontal or patent-divergent, oval or oblong-oval, bilobed to below the middle, segments connivent or spreading, large, unequal, lanceolate, acuminate ; texture lax, cells medium to rather large in size, quadrate or oblong-quadrate, walls delicate, no thickened angles or trigones. Stipules present on the male plant only, shorter than the leaves, subulate or ovate-lanceo- late, mostly entire, rarely bidentate at the apex. Fertile branches elongate ; bracts oblong-ovate-acute, bilobed to below the middle, segments lanceolate-subulate ; entire or with a small tooth below the middle on one side ; bracteole broadly ovate, coarsely unidentate on both sides ; sub-bracts ovate-lanceolate, only slightly bitid, segments acuminate ; 3rd sub-bract small, ovate, bilobed to about the middle. Perianth much exserted, large, linear-cylindrical, obtusely trigonous, composed of a single layer of cells down to the very base, about 70 cells round near the middle, mouth constricted, denticulate, teeth 1 or 2 cells long. Calyptra oval, delicate. Capsule reddish-brown. Spores reddish-brown ; elaters same colour, bispiral, hardly as broad as the spores. Andrcecia on short or long postical branches; perigonial bracts closely imbri- cate, assurgent, broadly ovate, swollen at the base, bilobed to about the middle, segments acute ; perigonial bracteoles (see stipules). Antheridia single, oval. Dimensions. — Stems 1 to 2 inches long, '125 mm. in diameter, with leaves '75 mm. to 1- mm. wide; leaves "4 mm. x "25 mm., 154 HEPATIC^. segment 2 mm., 45 mm. x '225 mm., seg. "225 mm., -35 mm. x •25 mm., seg. '175 mm.; cells "05 mm. x 02 mm., '04 mm. x 03 mm., "05 mm. x 025 mm.; bracts 1'4 mm. x '6 mm., seg. •75 mm.; bracteole 1*5 mm. x 1' mm., seg. "45 mm., -75 mm.; sub-bracts 1'25 mm. x '45 mm., seg. "25 mm., 3rd sub-bract •5 mm. x '35 mm., seg. '3 mm.; perianth 275 mm. x -5 mm.; teeth at the mouth of perianth -05 mm., -075 mm. ; spores '015 mm. ; elaters -15 mm. x '01 mm. Hab. — Growing in wet places, in bogs, &c., and by the side of rivulets. Moderately common. 1. Marazion Marsh, Penzance, Cornwall, JF. Ciirnow. 8, 9, 10. Eskdale, Yorkshire, Dr. Spruce, M. B. Slater; Maize Beck, Teesdale, Br. Spruce ; Eandy Mere, Egton, Yorks., M. B. Slater. 12. Bogs near Morecambe Bay, G. Stabler ; Grisedale, Cumber- land, G. Stabler. 13. North of Black Crag, New Galloway, /. Mc Andrew ; Barend Moss, Kirkcudbright, /. McAndreio. 16. Moidart, West Inverness, S. M. Macvicar. I. Connor Hill, Ireland, Br. B. Moore. Found on the Continent and in North America. Obs. — Very similar to large bog forms of C. hicusjndata (L.), but distinguished from them by its dioicous inflorescence, the female flowers on long branches, absence of flagella, whitish or dull brown colour (never rose coloured), the large, unequal, acuminate leaf segments, presence of stipules on the male plants, the deeply divided bracts with entire margins. Description of Plate LVIII.— Fig. 1. Plants natural size. 2. Portion of stem, postical view x 24. 3-5. Leaves x 64. 6. Portion of leaf x 290. 7, 8. Bracts x 24. 9. Bracteole x 24. 10, 11. Sub-bracts x 24. 12. Third sub-bract x 24. 13. Peri- anth X 16. 14. Cross-section of perianth x 24. 15. Portion of mouth of perianth x 85. (All Marazion, Cornwall, W. Curnow.) EUCEPHALOZIA. 155 6. Cephalozia hibernica, Spn/ce MS. Cephcdozia hibernica, Spruce MS. Pears, in " Irish Naturalist," Dec. (1894). Dioicous, creeping amongst mosses, with few rooting flagella, smal], of a whitish, crystalline or pale green colour. Stems stout, flexuose, ramose, procumbent or somewhat erect, plano-convex, antical aspect 3-4 cells wide, postical with a band of 4 x 5 much smaller cells; branches postical, irregular, sometimes attenuate and gemmiparous, gemmae capitate ; rootlets hyaline, on the lower part of the stem, or at the apices of the arcuate stolons. Leaves approximate or contiguous, alternate, almost horizontally inserted, decurrent at the base, subalate, plane, so that the stem appears bialate, bifid from a third to the middle of the length, sinus rounded, segments subconnivent or erect, sharply acuminate, apical cells 2-4 in single series, texture delicate, cells large, hyaline, 4-, 5- and 6-sided, surface pitted when dry as in Cejjh. comiivem, walls thick, trigones wanting. Stipules absent. Flowers ? ter- minal on very short postical branches, bracts oblong, divided to about two-thirds into two unequal lanceolate-acuminate segments, sub-bract smaller, divided to about the middle, sometimes with a third small segment on one side, segments acute ; bracteole oblong, bifid to about the middle or entire, unidentate near the middle, segments lanceolate-acuminate ; sub-bracteole oval, divided to about \, segments obtuse, third sub-bracteole small, slightly bifid. Immature perianths only met with, mouth fringed with long cilia, 12 to 15, each composed of 3 or 4 long single cells. Androecia on short postical branches, perigonial bracts 4-5 pairs, closely imbricate, complicate-concave, oval, bilobed to about one- third ; antheridia single, oval. Dimensions. — Stems ^ inch long, with leaves r25 mm. wide; diameter of stem '2 mm. ; leaves '7 mm. x "45 mm., segments •25 mm., -2 mm., '5 mm. x '3 mm., seg. "2 mm., "IS mm., •55 mm. x '35 mm., seg. '2 mm., '175 mm. ; branch leaves "35 mm. X 2 mm. ; seg. -15 mm., "1 mm. ; cells '04 mm. x '05 mm., •05 mm. X -06 mm., '04 mm. x -04 mm., seg. -005 mm., '02 mm.. 156 HEPATIC^. ■05 mm. X "03 mm. ; bracts 1* mm. x "35 mm., seg. '6 mm., 8 mm., 12 mm. X "35 mm., seg. '(3 mm., 9 mm., bracteole '85 mm. x ■4 mm., seg. '6 mm., 1* mm. x '4 mm. ; pistillidia -125 mm. x •04 mm. ; perigonial bract -275 mm. x '2 mm. ; explanate, seg. •075 mm. Hab. — Among Playiotheciiim Borrericumm, Spruce, on banks, Killarney, Dr. David Moore, 1865. Killarney, Mr. Reginald W. ^cnlhj, 1889. Obs. — Specimens were sent by Dr. David Moore to Dr. Carrington in 1865 as Jmig. connkens, who, recognising it as distinct from that species, brought it under the notice of Dr. Gottsche of Altona. He referred it doubtfully to Jung, cra-m folia ^ Lindenb. & Gottsch.. It would probably have remained unrecorded until fertile plants were found, had not a fragment of Moore's specimen been forwarded to Dr. Spruce, who at once referred it to CepJialozia crassifolia {\j. & G.); since then he received further specimens from Killarney, collected in 1889 by Mr. Reginald Scully, and wrote to me some time before his death, that he was inclined to consider the species distinct, and proposed the MS. name of Cephahzia hihernica. I feel doubtful until fertile specimens have been found whether it can be separated from Ceph. crassifolia. In any case it is a distinct addition to our Flora. There is a strong resemblance in habit, texture, and structure of this plant to the genus Zoopsis, its plano-convex stem, with the band of small cells running through it, giving it a vertebrate appearance, its remarkably plane leaves, the cells of which cross the stem unaltered. It is not difficult to trace the transition from the apparent simple ribbon-like frond of Zoopsis argentea (Tayl.) through Z. seffdosa Leitg., with its claw-like leaves, to the more perfect leaf form of Z. Leifgehiaua C. & P. and so to the distinctly foliose Cep)lialozia crassifolia and Ceph. hihernica. It is distinguished from its nearest ally, Ceph. connive^is (Dicks.) by its dioicous inflorescence, the longer segments of its leaves, which are composed of 2-4 single long cells, and other characters. I EUCEPHALOZIA. 157 I am indebted to the late i)r. Carriii<^ton for assistance in the preparation of this description. Description of Plate LIX. — Fig. 1. Plant natural size. 2. Portion of stem, antical view x 24 (Killarney, Dr. Moore). 3. Cross-section of stem x 85 (ditto). 4-G. Leaves x 64 (ditto). 7. Branch leaf x 85 (Killarney, R Scully). 8. Portion of leaf X 290 (Killarney, Dr. Moore). 9. Portion of segment of leaf X 290 (ditto). 10, 11. Bracts x 31 (ditto). 12, 13. Brac- teoles X 31 (ditto). 14. Portion of the mouth of an immature perianth x 31 (ditto). 15. Portion of male stem, antical view X 31 (Killarney, Pt. Scully). 16. Perigonial bract x 85 (ditto). 7. Cephalozia connivens {Dicks.). Junyermania connivens, Dickson, PI. crypt. Brit. Fasc. iv. (1801) ; Hooker Brit. Jung. t. 15 (1816) ; Eng. Bot. t. 2436. Cephalozia connivens (Dicks.), Spruce, On Ceph., p. 46 (1882). Blepharostoma connivens, Dum. Eecueil, p. 18 (1835); Hep. Eur. p. 96 (1874). Monoicous, loosely csespitose or creeping, flagelliferous, small, pale green, pellucid, fragile. Stems subramose, subcompressed, cortical cells about 8, interior moderately large, almost empty; radiculose, rootlets plentiful, long, white. Leaves lower and upper distinctly smaller, middle larger, subimbricate or aj^proxi- mate, almost horizontal, insertion sublongitudinal, antical, very decurrent, obliquely suborbiculate, from J to the middle bifid, sinus obtuse or lunate, segments triangular-acuminate, connivent ; cells moderately large, quadrate-hexagonal or oblong-quadrate, almost empty, walls thick, no trigones or thickened angles. Stipules none. Bracts 3 pairs, the innermost almost twice as long as the leaves, oval or broadly ovate or irregular in outline, deeply 3-5-fid (normally twice bifid), segments lanceolate- subulate, acuminate, margin entire or sub-spinulose ; bracteoles almost free, deeply bifid, exterior margin 1, 2 teeth. Perianth projecting, pyriform-fusiform, terete, towards the apex obtusely trigonous, cells elongate, composed of a single layer, about 60 roundish cells near the middle, these giving the inner and the 158 HEPATIC^. outer " surface a verruculose appearance, mouth sub-constricted, longiciliate, afterwards trilobed ; cilia of the so-called lobe about 4, of about 5 cells long, usually of one series of cells, a few eilioles or teeth interspersed. Calyptra short, delicate. Pistil- lidia few, about 6, long, narrow. Capsule oblong-globose. Androecia conspicuous, occupying the whole branch or often only on the apex or base of it, perigonial bracts 8-10 pairs, a little smaller than the leaves, concave, to the middle bifid, often with a tooth or lobule added on the antical side; perigonial bracteoles more or less absent ; antheridia oval, single. Dimensions. — Stems about an inch long, -2 mm. in diameter, with leaves l" mm. wide; leaves -5 mm. x '5 mm., -4 mm. x •85 mm., segments -2 mm., '45 mm. x -45 mm., seg. '2 mm. ; cells -075 mm., -07 mm. x '05 mm,, "04 mm. x -04 mm., '07 mm. x •04 mm., '05 mm. x •OS mm.; sub-bracts ^75 mm. x '7 mm., seg. •35 mm., -65 mm. x '55 mm., seg. '3 mm., '5 mm. x ^4 mm., seg. '3 mm.; sub-bracteole •SS mm. x '2 mm., seg. '15 mm.; bracts 8 mm., !• mm. x 1' mm., seg. -75 mm., I'l mm. x !• mm., seg. -6 mm. ; bracteole '6 mm. x '5 mm., seg. -4 mm. ; perianth 2^5 mm. — "4 mm. x I'mm., 3. mm. x ^75 mm.; cilia "4 mm., ^5 mm.; pistillidia '25 mm. x -04 mm. ; capsule '5 mm x ^5 mm. ; valve of capsule 1* mm. X -3 mm.; spores 'OlS mm.; elaters '1 mm. x •015 mm. ; perigonial bracts '4 mm. x '35 mm., seg. ^2 mm., •45 mm. x '45 mm., seg. '2 mm. ; antheridia "125 mm. x •I mm. Hab. — Grrowing in loose tufts, thin patches, or creeping among Sphagna on wet moors or peat mosses. Somewhat rare. 1 ? 2. New Forest, Hampshire, C. Lyell. 4. Marshy place in wood near Holt, Norfolk, Hooker. 5 ? 7 ? 9. Chat Moss, Lane, W. H. P. 10. Terrington Carr, Dr. iS/jrac(?; Groathland & Wheeldale, nr. "Whitby, M. B. Slater. 12. 13. Glenlee Glen, New Gralloway, J. Mc Andrew ; Barend Moss, above Eouten Bridge and Crichope Linn, Kirkcudbrightshire, J. Criiickshank. 14. Kelso, A. Brotherston. 15. Strachan, Aberdeen, J. Sim. 16. Moidart, West Inverness, S. M. Macvicar. I. Found on the Continent and in North America, also a variety with flagella in South Africa. ErVEVUMOZlA. l.V» ( )ns.— T)istin}j;ui.sliL'cl Ironi (\ liivuspulnht (L.) |jy tlie Icjugi- tiidinal or lateral insertion of tlie leaves, givin<^ it a flat ribbon- like aj)pc'arance, the very decurrent antical base, connivent segments, the larger cells, absence of flajj^ella, and periantii with long cilia at the mouth. C. lK/iN/<{'/()li(i Duni. is a smaller ])lant with dioicous inflores- cence. Dkscription of Platk LX. — Fig. I. Plant natural si/e. L'. Portion of stem x 24. .'3, 4. Leaves x 64. 5. Portion of leaf X 290. 6,7. Bracts x 24. 8. Bracteole x 24. l)-ll. Sub- bracts X 24. 12. Sub-bracteole x 21-. \'.\. Perianth x 16. 14. Cross-section of perianth x 16. 15. Portion of mouth of perianth x 85. 1 (>, 17. Perigonial bracts x 64. 18. Anlher- idinm x 85 (Terrington Carr, Yorks., Dr. Spruce). S. Cephalozia curvifolia (/>'/r/-.s.), Di/m. Jungennania citrn/olia, Dicks. Tl. ciyi.t. Urit. f-ASc. II, ir>, t. T. (170(>); Hook. Brit. Jiin^. t. 15 et Suppl. t. 1. e.r jxirte (IHIC). Jungermania liaiieri, Mart. Fl. crypt. Erlang. p. 17l', t. •">, f 4C< (1SI7). Cephalozia curvi/olia, Dum. Recueil p. 18 (IS.S.''*). Xouyllia ciirvt/olia. Mitten in Godman's " Natural Hi.story of the Azores ' (1S7(I). Monoicous and dioicous, densely dcpresso-ca?spitose, cflagellife- rous, small, pale green, whitish-red or deep purple colour. Stems slender, pellucid, subterete ; radiculose, rootlets somewhat thick, numerous; simple or slightly branched, branches few, ascending, postical ; very rarely furcate ; cortical cells 8, large, outer walls thick, inner 4 cells in diam., smaller. Leaves loosely imbricate, assurgent-subsecund. succubous, almost transversely inserted, oblique obovate, concave, antical margin almost straight, postical broadly semi-cordate at the base, with intle.xed auricle, with margin appresscd to the leaf, ventricose at the keel, bilobed to below the middle, sinus more or less broadly luniilate; segments incurved-hamate, capillaris-cuspidate, j)oints consisting of 8-10 oblong-(|uadrate uniseriate cells ; cells smallish to medium in size, 160 HEPATIC^. quadrate or obloDg-quadrate, guttulate, verruculose, walls thick, angles thickened. Axillary leaf (when stem furcate) acuminate entire, the other segment abortive. On some elongated stems and branches the leaves are more symmetrical, the inflexed auricle disappearing. Stipules none. Branches $ generally very short, postical. Bracts 3 pairs, tristichous, innermost erect, oblong, complicate-bilobed, segments subovate apiculate, partly free, all the margin, except towards the base, minutely, unequally spinu- lose-denticulate ; bracteole similar ; exterior bracts much smaller ; all the bracts and bracteoles without auricles. Perianth large, often about the middle of a rosy-purple colour, apex white, linear, deeply triquetrous-prismatic, mouth broadly truncate, wide, rarely constricted, setulose, teeth 1-4 minute quadrate cells long ; deli- cate texture, composed of one layer of cells, near the middle about 200 very small, roundish cells around. Pistillidia about 10, large. Calyptra very delicate. Capsule oblong-globose. Androecia terminal or on short postical branches ; perigonial bracts many, closely imbricate, somewhat similar to the leaves but broader ; perigonial bracteoles subulate ; antheridia single, oval. Dimensions. — Stems J to f inch long, diam. -15 mm. ; leaves •65 mm. x '4 mm. ; lobule *3 mm., seg. '35 mm., '6 mm. x "5 mm. lobule, '3 mm., seg. "3 mm., '75 mm. x '5 mm. ; lobule, '2 mm., seg. •4 mm.; cells "045 mm. x '025 mm., '035 mm. x '025 mm., •035 mm. x 03 mm., "03 mm. x "025 mm., "025 mm. x "025 mm. ; bracts 1*25 mm. x '9 mm., seg. "5 mm.; bracteole 1"1 mm. x •7 mm., seg. "4 mm.; sub-bract 1" mm. x '6 mm., seg. '4 mm. sub-bract, "65 mm. x '45 mm., seg. '25 mm. ; sub-bracteole '5 mm. x "3 mm. ; perianth 2' mm. x '6 mm. ; teeth at the mouth •075 mm., 125 mm. ; pistillidia '2 mm. x '04 mm. ; perigonial bracts "6 mm. x '55 mm. ; lobule, '25 mm., seg. '4 mm. ; perigonial bracteole '3 mm. x '05 mm. ; antheridia '1 mm. x "075 mm. Hab. — Growing on rotting trunks of trees, soft sandstone rocks ; and damp heathy moors in subalpine situations. Somewhat rare. 2. Tunbridge Wells, Sussex, Dr. Sprifcc. ^,7. Cvvm Idwal, Carnarvonshire, C. J. JFild and 77^. H. P. 9. Near Todmorden, ErcEl'UALUZIA. Kil John Noire//. 10. Near Whitby, O. S/S'. M. Macoicar. I. About Killarney abundant. Found on the Continent and in North America, also ^Mexico, Azores, and, according to " Syn. Hep.," in South Africa. Obs. — A most beautiful species, usually of a rosy or deep purple colour ; distinguished from all other British Ccp/id/ozuc by the remarkable intle.xed auricle or lobule on the lower side of the leaf. Sometimes the stem is furcate, with a difform (unicural) leaf at the fork; this and the leaf auricle, I suppose, induced Mr. Mitten to make a separate genus of it; but, as Dr. Spruce points out, this rare feature is also seen in other Ccp/ia/ozi(/', and the auricle disappears in some of the ordinary leaves, towards the a})ex of drawn-out stems and branches, and always in the bracts, thus rendering these characters not of sufficient value to separate it from true Cep/ia/ozia. I regret this, as I should have been glad to have followed Mr. Mitten in associating the name of Nowell with this elegant species. John Nowell was a working-man botanist, who had a most intimate knowledge of the Hepaticie, and who was one of the most modest and generous of collectors. My readers will, I trust, pardon me for inserting the following lines by his I'rieud, Mr. Stansfield : — " Nature to Inm Wa.s ever, ever blooming ! Ah, to tell The rapture that, even in winter's depth, To him could yield each tiny, glistening moss, Or lichen grey, clothing the barren rock I AVhat nook of these our vales Ilath he not peeped in— peeped, nay, closely .scanned^ At what clear spring hath he not bent and drank ( Beside what stream, or through what dough, hath he 162 HEPATIC^. Not roved, at his own gentle pace, and there Ta'en note of plant-life in its tiniest forms ? What hill or knoll hath he not clomb, early Or late, or at mid-day, in rain or shine ? Shine sweetly, Sun, upon this honoured grave ! Tread softly ye upon his kindly mould ! And raise a Tablet that the child may ask : ' For whom ? ' and learn that Truth and Gentleness have lived ! " I have remarked upon the colour of the species. Whilst collecting, along with my friend Dr. Carrington, in the woods of Borrowdale, we noticed at some considerable distance a mass of rich reddish-purple which we could not make out. Judge our surprise, when we approached near, to find the huge trunk of a rotting tree covered with this species ! ^We speedily made our- selves " Rich in the wealth, which is collected Among woods and fields," Description of Plate LXI. — Fig. I. Plants natural size. 2. Portion of stem x (Dr. Grottsche, in G. & R. Hep. Eur. n. 232). 3, 4. Leaves x (ditto). 5-8. Leaves x 24 (Naddle Forest, Westmorland, G-. Stabler). 9. Portion of leaf x 290 (ditto). 10. Bracts x 24 (Patterdale, Cumberland, W. H. P.). 11. Bracteole x 24 (ditto). 12. Sub-bract x 24 (ditto). 13. Sub- bracteole x 24 (ditto). 14. Sub-bract x 24 (ditto). 15. Perianth X 24 (ditto). 16. Cross-section of the same x 24 (ditto). 17. Ditto x ?(Dr. G. inG.& E. Hep. Eur. n. 232). 18. Portion of the mouth of the perianth x 85 (Patterdale, AV. H. P.). 19, 20. Perigonial bract x 31 (ditto). 21. Leaf at the base of ■stem X 31 (ditto). 22. Perigonial bracteole x 85 (ditto). 23. Antheridium x 85 (ditto). EUCEPHALOZIA. 163 9, Cephalozia Francisci [Ifoo/i.), Duiit. Jungermania Francisci, Hook. Brit. Jung. t. 49 (1810). Cephalozia Francisci, Dum. llecvieil, p. 18 (1835). Dioicous, densely csespitose, flagelliferous, small, green to reddish-green at the ends of the stems. Stems filiform, yet pro- portionately thick, fleshy, procumbent, subramose, branches ascending, with few leaves near the base ; radiculose, rootlets plentiful at the base, hyaline; cortical cells about 15, inner slightly larger, all opaque ; flagella numerous, leafless or with minute leafy processes. Leaves patent-divergent or patent spreading or subsecund, distant or subimbricate, small, oval- orbiculate or broadly ovate, from ^th to ^rd bidentate, sinus acute or obtusate, segments acute or obtuse, lower one usually connivent; cells smallish, subquadrate, near the middle oblong, marginal cells small, quadrate ; walls thick, no trigones or thickened angles. Stipules large, ovate-lanceolate, obtuse, entire, rarely oblong-quadrate, bidentulate. Inflorescence cladocarpous, ? bracts 3 pairs, innermost much larger than the leaves, ovate- oblong, bifid to about Jrd, segments lanceolate obtuse or acute, recurved, unidentate on the outside ; bracteole free, ovate-oblong, bifid ; sub-bracts oval, shortly bidentate, sinus obtuse, segments acute connivent ; sub-bracteole broadly lanceolate bifid. Perianth twice as long as the bracts, projecting about half beyond them, narrowly ovate-fusiform, tricarinate almost to the base, mouth 6-plicate, entire, at the base 3 cells thick, at the middle 2 cells. Calyptra lower half 2 cells thick. Pistillidia 5-8. Capsule large, dark brown, almost black, oval-cylindrical, spores and elaters deep red colour. Androecia on the middle or the ends of the stem or postical branches ; perigonial bracts 3-7 pairs, closely imbricate, concave- canaliculate, ovate-orbicular, bifid, sinus and segments subacute ; perigonial bracteole linear-lanceolate, acuminate. Antheridia sina^le, laro^e, oval. Sometimes gemmiparous, gemmse large, greenish to reddish- ij^reen, at the ends of the stems. 164 HEPATIC^. Dimensions. — Stems J to J inch long, diam. '15 mm.; leaves •3 mm. X '325 mm., ses^ments "1 ram., '35 mm. x "325 mm., ses^. "075 mm., "35 mm. x '225 mm., seg. '075 mm., '4 mm. x "35 mm., •35 mm. X '35 mm., -25 mm. x 15 mm. ; cells "OG mm. x 02 mm., "035 mm. x "02 mm., '04 mm. x "02 mm., "03 mm. x "02 mm., *025 mm. x "025 mm. ; marginal cells "02 mm. x '02 mm. ; stipules '2 mm. X '1 mm., '2 mm. x "075 mm., -15 mm. x 'l mm., '15 mm. X 08 mm., "2 mm. x "1 mm, ; sub-bract "75 mm. x "5 mm. ; sub- bracteole "5 mm. x '3 mm. ; bract I'l mm. x '7 ram., seg. '4 mm., •0 mm. X -55 mm. ; bracteole "D mm. x "5 mm., seg. "l mm. ; perianth 2" mm. x '6 mm., 2" mm. x '55 mm.; capsule '5 mm. x *4 mm., '65 mm. x '35 mm. ; pedicel '12.5 mm. diam. ; spores "02 mm. diam.; elaters "15 mm. x '02 mm.; perigonial bracts •3 mm. X -25 mm., seg. '075 mm. ; perigonial bracteole '225 mm. X '1 mm. ; antheridia "15 mm. x •[ mm. Hab. — Growing by the sides of ditches in moist turfy soil on moors. Very rare. 2. New Forest, Hants, C. LijpJL 4. Holt & Edgefield Heaths, Norfolk, Francis. 9. Dale Ford, Delamere Forest, Cheshire, W. Wilson. 10. Moors near York, Stockton Forest. Langwith Moor, I)r. Spruce, Strensall Common, Yorks, M. B. Slater §• G. Stabler. 11. Kildale Moor, Cleveland, B. Muchl 13. Eoadside between Rosehall and Brownhall, Dumfries, Cridcksltank. 15. Baker Wells, Strachan, Aberdeen, ./. Sim; Loch-na-gar, Aberdeen,/. Sim. Kinnordy, C. Lyell. I. Bantry, Miss Hut chins. Found on the Continent sparingly. Obs. — Distinguished from the dioicous C. lunultTfolia, Dum. C. catenulata (Hliben.) and C. divaricata (Sm.), in addition to other characters, by the presence of large stipules along the whole length of the stem. C.paitans (N.) is a very different plant with large lax oblong leaves, segments obtuse, large cells. By some strange mistake this species is figured in " Eng. Bot.," t. 2569, as Jung. Francisci. Dr. Spruce remarks that in habit, presence of flagella, concave leaves, small cells, the erect branches with leaves gradually becom- E UCEPHA TJJZL I. 1 05 ing si^ialler at tlie ends and l)caring rosy geninia?, C. Francixri clearly has an affinity with C. [(Jdonfo.) denndahi (Mart.). Description of Plate ]jXII. — Fi0 (1882). Dioicous, loosely ca3spitose, flagelliferous, medium to large in size, light green to pale brown in colour. Stems elongate, laxl}' creeping, subramose, radiculose, rootlets plentiful, white ; 8 cells in diam., cortical cells 14-lG, clilorophyllose, opaque, inner slightly smaller, pellucid ; flagella plentiful, radiculose, leafless or partly leafy. Leaves large, ascending, subsecund, distant, rarely subimbricate, obliquely inserted, suboblique oval-ovate or oblong, sometimes subcuneate at the base, slightly concave, bilobed to a third, rarely almost to the middle, sinus acute or obtusate, narrow, segments unequal, postical larger, lanceolate, apex subcucullate, obtusate, rarely subacute ; cells medium to ratlier large, hexagonal, leptodermous, chlorophyllose, subopaque, basal cells a little larger, marginal ones subquadrate, walls somewhat delicate, angles slightly 166 HEPATIC^. thickened, no trigones. Stipules distant, appressed to the stem, linear, linear-lanceolate, subulate or bifid and irregular segments often unequal, 3-6 cells long, sometimes with 1 or 2 teeth at one or both sides. Inflorescence cladocarpous, ? branches postical, short, about 3 mm. long. Bracts lax, 3 pairs, trisiichous, inner- most about the size of the leaves, erect, ovate-oblong, canaliculate, from a third to nearly the middle bilobed, segments acute or subacuminate, towards the base 1, 2 dentate; cells elongate, sub- pellucid. Bracteole same as the bracts. Sub-bracts much smaller than the bracts, unequally bidentate, or falcate and entire. Perianth projecting 2 to 3 times its length beyond the bracts, linear-fusiform, cylindrical, trigonous only at the apex, mouth truncate-subconstricted, entire, composed of one layer of cells from the middle to apex, lower half 2 cells, at base 2 to 3 cells thick, near the middle about 90 cells round. Calyptra 2 to 3 times shorter, slightly narrower, oblong-pyriform, delicate, at the base 2 to 3 cells thick. Capsule purple, oblong or oblong- cylindrical ; valves linear-lanceolate, composed of two layers of cells. Spores dark reddish-brown, broader than the elaters. Elaters bispiral, same colour as the spores. Amenta i postical, perigonial bracts few (3-6)-pairs, smaller than the leaves, oval or orbicular, concave, shortly bilobed, seg- ments subacute, sometimes with a third, short, antical, incurved segment ; bracteole linear or oval-lanceolate, entire or bifid ; antheridia large, solitary, oval-globose. Dimensions. — Stems from 1 to 3 inches long, '2 mm. to •25 mm. in diam., with leaves 1-25 mm. to 1-75 mm. wide; leaves '9 mm. x '55 mm. segments, "2 mm., '6 mm. x •4mm. seg., •175 mm., 1* mm. x "9 mm. seg., -3 mm., 1. mm. x "6 mm. ; cells •045 mm. x -04 mm., '04 mm. x "03 mm., "035 mm. x -835 mm.; stipules 'b mm. x -08 mm., •35 mm. x 'l mm., ^3 mm. x -1 mm., •275 mm. X •I mm., ^225 mm. x ^075 mm.; sub-bracts ^7 5 mm. X "4 mm. seg., '2 mm., '5 mm. x "3 mm. seg., '2 mm., 'Q mm. X '3 mm. seg., •I mm.; bracts I' mm. x ^6 mm. seg., 'A mm., 1^25 mm. X •e mm. seg., 'o mm., 1*2 mm. x ^5 mm.; bracteole •9 mm. X ^5 mm. ; perianth 3*3 mm. x '8 mm., A' mm. x '7b mm.. EUCEPHALOZIA. 167 4' mm. X '5 mm. ; calyptra 1*6 mm. ; capsule "85 mm. x '5 mm., •8 mm,, X "3-4 mm.; valves 'lb-?>b mm. broad; spores "02 mm.; elaters "2 mm. x '0125 mm.; amenta 2* mm. long; perigonial bracts "5 mm. x '3 mm. sag., "1 mm.; perigonial bracteole "2 mm. X -075 mm. ; antheridia "175 mm. x "15 mm. Hab. — " In tbe wettest part of bogs, creeping upon Sphagna and other mosses, sometimes partly floating." R. S, Somewhat rare. 2. New Forest, Hampshire, C. Lydl, 1813. 7. Glyder Vawr, Carnarvonshire, G. A. Holt. 9. Delamere Forest, Cheshire, W. Wilson. Abbots Moss, Cheshire, G. A. Holt. Carrington, Moss, Cheshire, G. A. Holt. Wybunbury Bog, Cheshire, /. Whitehead. Staley Brushes, Cheshire, /. Whitehead. Barton Moss, Lane, W. H. P. 10. Far Wheeldale and Goathland Moor, Yorks., SauL Anderson. 12. Foulshaw Moss, Westmorland, G. Stabler. 13. Barend Moss, nr. Castle Douglas, Kirkcudbright- shire, /. i)/c^y/r/yYvt\ 16. Moidart, West Inverness, aS'. il/. Macvicar. I. Kylemore, Co, Gal way, ]J. Moore. Found on the Continent and in North America. Obs. — First discovered by Mr. Lyell in the New Forest in 1813, and published and figured in "Eng. Bot." under the false name of Junfjeruianla Francisci, from which it is quite distinct, as will be seen by reference to the description and figures of that species. Nees was the first to recognise it as a new and distinct species, but he appears afterwards to have doubted its claims to specific rank, for he reduced it to a variety of Jungermania injlata in his later works. As Dr. Spruce remarks, how such a con- summate hepaticologist should have confounded them it is diflftcult to conceive, adding : " It may suffice to contrast their chief characters, which are, for C.puitans, the stem rooting by numerous stout flagella; the branches, whether foliferous or floriferous, all postical ; the longer, narrower, and more laxly-reticulate leaves ; the constant presence of stipules ; the cladocarpous inflorescence ; the tristichous female bracts, toothed at the base, the innermost embracing the perianth ; finally, the lineari-fusiform, trigonous, thin perianth. But in Jungermania injtata there are no flagella • 168 HEPATIC^E. the branches arise variously from the mid axil of a leaf, or from its postical angle, and the female flowers are borne on the apex of the stem or of long leafy branches ; there are no stipules at all, except very rarely a small subfloral one ; the bracts are distichous, conformable to the leaves, and usually remote from the perianth (whence the species becomes the type of Dumortier's spurious genus Gymuocoleci) \ and the perianth itself is pyriform, inflated, and obscurely 4-5-plicate, only at the very apex ; it is besides composed of 2 strata of cells up to \ of its height." Description or Plate LXIII. — Fig. 1. Plants % and ^ natural size. 2. Portion of fertile stem with perianth x 11 (Foulshaw Moss, G. Stabler). 3. Portion of male stem x 16 (ditto). 4. Portion of brancli, antical view x 24 (ditto). 5- Portion of leaf x 290 (ditto). 6-8. Stipules x 85 (Delamere Forest, W. Wilson). 9. Stipule x 85 (Wheeldale, Sam. Anderson). 10, 11. Bracts x 24 (Foulshaw Moss, CI. Stabler). 12-14. Sub- bracts X 24 (ditto). 15. Cross-section of perianth, near the base X 16 (ditto). 16. Ditto, near the apex x 16 (ditto). 17. Por- tion of mouth X 85 (ditto). 18. Perigonial bract x 31 (ditto). 19. Perigonial bracteole x 85 (ditto). 20. Antheridium x 85 (ditto). 11. Cephalozia heterostipa, Carr. ct Spruce. Ge'phalozia lieterostipa, CaiT. et Spruce. Spruce, On Ceph., p. 55 (1882). Junyermania inflata, var. heterostipa, Lindb. in Arn. & Lindb. Muse. Asiae bor, p. 47 (1888) ; Kaalaas Leverm. Norge, p. 290 (1893). Dioicous, depresso-ca?spitose, flagelliferous, from a reddish- brown to a dull brown or green, sometimes orange colour, fragile. Stems intricately flexuose, radiculose, rootlets produced the whole length of the stem, white ; simple or twice dichotomously branched, with the young apex almost always bifurcate, sometimes, not always, producing radiculose, stoloniferous, postical branches partly leafy, rarely altogether leafless. Cells of the stem 7 or 8 in diameter, cortical 20-22, subquadrate, subopaque, inner some- what smaller, more pellucid. Leaves diagonally inserted at the EUCEPHALOZIA . 1 G9 base, lower ones distant, patulous, oblong or cuneate -oblong-, to a third subacutely bilobed, segments obtusate or rotundate, often discoloured, upper ones approximate and more or less imbricated, and near the flowers, and where tlie stem bifurcates more deeply coloured, broader, cuneate, assurgent-concave, often to the middle bilobed, very often 3- and sometimes 4-lobed, angular or obsoletely denticulate, lobes alone obtuse, sometimes subacute, subequal, or witli the exterior one, now the antical, now the postical, smaller ; cells small to smallish, 4-6 sided, sul)leptodermous, chlorophyllose, subopaque, lower ones slightly elongate, walls thick, angles not thickened, or only very slightly, no trigones. Axilliar antical leaf (at the fork of the stem) a little smaller than the others, ovate, entire, rarely with a tooth added at the base. Stipules small or minute, sometimes obsolete, rarely altogether wanting, coloured, linear or subulate, entire or bifid, segments erect, narrow, unequal. Sometimes between the normal stipules otlier abnormal ones are found ec[ual in length to the leaves, falcate-lingulate or irregular. Flowers ? terminal on stem, pistillidia 10-16. Bracts 2-3 pairs, laxly imbricate, concave, broader than long, 3-4-lobed, lobes subacute, obtuse, or rotundate ; bracteoles rather smaller than the bracts, oblique, ovate-lanceolate, entire, or deeply bilobed. Perianth projecting much, bej^ond the bracts, green, p3^riform, some- what compressed, very indistinctly trigonous, mouth shortly 6-iobed, lobes 2-4 dentate, whole number of teeth about 18, shortly sub- ulate, unequal; cells quadrate, opaque, pachydermous, composed of one layer of cells, except at its base and a very little higher up, where it is composed of two layers. Dimensions. — Stems |^ to 1 inch long, "2 mm. to "25 mm. in diameter, with leaves 1'5 mm. wide ; leaves, lower "6 mm. x "4 mm., ■6 mm. X '5 mm., segments '2 mm., "65 mm. x "6 mm., seg. '25 mm., upper 75 mm. x "55 mm., seg. -25 mm., 'S mm. x Qtb mm., •9 mm. X '55 mm., seg. "25 mm., 1* mm. x 7i) mm., seg. "4 mm. ; cells '02 mm. x -02 mm., '03 mm. x •025 mm., '0275 mm. x '02 mm., •03 mm. X 03 mm. ; stipules ^2 mm. x "04 mm., '2 mm. x '06 mm., •225 mm. x '05 mm., '15 mm. x ^04 mm., -125 mm. x 075 mm. ; bracts 6 mm. x 8 mm., '75 mm. x 8 mm., '9 mm. x 'll mm. : 170 HEPATIC^. bracteoles 3 mm. x 'ITS mm., '4 mm. x '15 mm., "325 mm. x "1 mm. ; perianth 3' mm. x 1'4 mm., 2*75 mm. x 1*5 mm. ; pistillidia '2 mm. x "05 mm. Hab. — On wet rocks on momitains. 7. Glyders, North Wales, E. M. Holmes, 1876. Extremely rare. Obs. — Distmguished from CcjjJi. fait cats (iSTees) by its smaller size, lurid colour, leaves more frequently trilobed, much smaller cell structure, in being acrocarpous, perianth with mouth laciniate- dentate. From small forms o^ Jiim/. injlata Huds., in having postical, flagelliform branches, distinct polyphyllous involucres, constant presence of bracteole, perianth being visibly (although very obtusely) trigonous upwards and rather wide-mouthed. Description of Plate LXIV. — Fig. 1. Plants natural size. 2. Portion of stem, postical view x 24. 3-7. Leaves x 24. 8. Portion of leaf x 290. 9-13. Stipules x 85. 14. Bract x 24. 15-17. Bracteoles x 85. 18. Perianth x 16 (Glyders, E. M. Holmes). Subgenus 2. OBONTOSCHISMA, Bum, OdontoscJdsma, Dum. Recueil, p. I'J (18oo) (genus). Sphagncecetis, Nees in G.L.N. Syn. Hep. p. 148 (1S45). Jungermania, Dicks, itc. Plants somewhat robust, green or lurid, sometimes rosy, rarely whitish, spreading, in closely intricate la3^ers, or creeping upon mosses, very often densely csespitose. Stems firm, subterete, prostrate or repeatedly arcuate, with the flagelliferous branches rooting; leafy branches few, similar to the stem. Cells of the stem about 8 in diameter, cortical 20-22, about equal in size to the inner ones and of the same colour, sometimes more opaque. Leaves succubous, inserted at the base diagonally or almost longitudinally, broadly ovate or suborbiculate rotundate or retuse, rarely obcordate or emarginate-bidentate, more or less concave, entire ; cells medium size or ver}^ small, equilateral, walls in some ODOXTOSCIimMA . 171 species delicate, in others with the angles thickened, epidermis sometimes scaberulous. Stipules minute and everywhere obvious, or partly obsolete, sometimes none. Inflorescence for the most part dioicous, in a few monoicous ; ? cladogenous. Fertile branches short. Bracts small, tristichous, sometimes bifid, rarel}- 3-4-fid. Perianth larg-e, leptodermous, trigonous-fusiform, very often narrow, mouth ciliate or denticulate, sometimes closed and dividing at a lateral cleft. Calyptra leptodermous. Capsule cylindrical- oblong, in other respects similar to all Cephcdodce. Androecia constantly on whitish small amentula, very rarely more robust and terminal ; perigonial bracts minute ; antheridia situated in their axils. Often gemmiparous, gemmre collected in heads upon the attenuated tips of the branches. 12. Cephalozia Sphagni {JJich.), S/)r//cc. Janyermania Sphagni, Dicks. Crypt. Brit. (1785); Hook. Brit. Jung. t. 33 (^jn'o parte) (181G). Fleuroschisma (Odontoschisma) tSphagni, Dum. Syll. Jung. p. 08 (1831). Odontoschisma Sphagni, Dum. Recueil, p. ID (1835). Sphagncecetis Sphagni, Nees in G.L.N. Syn. Hep. p. 148 (1845). Cejyhalozia {Odontoschisma) Sphagni (Dicks.), Spruce, On Cephalozia, p. GO (1882). Dioicous, loosely ctespitose or creeping sparsely among mosses, flagelliferous, largish in size, green or brownish-green in colour. Stems procumbent, elongate, arcuate, simple or with few postical branches, firm, flexuose, gemmiparous branches wanting, cortical cells 25-30, yellowish-brown, similar in size to the inner ones only with thicker walls, 10-12 cells in diameter, inner cells sub- opaque ; flagella postical, stout, firm, covered with minute root- lets or short conical cells ; radiculose, rootlets numerous, minute, simple hyaline, proceeding from the postical side of the stem. Leaves imbricate or approximate, alternate, bifarious, patent- divergent to patent, subobliquely or almost longitudinally inserted, secund, concave, never repand or undulate, ver}' slightly decurrent antically, ovate-rotund, entire, rotundate, retuse or sometimes 172 HEPATIC^. emarginate, more or less distinctly margined, margin often brownish ; texture firm, epidermis smooth ; cells small to medium size, subopaque, roundish-quadrate or roundish-oblong-quadrate, walls thick, angles large, thickened, trigones rarely present, marginal cells usually quadrate with thicker walls. Stipules none or rarely present at the apex of stem or branches, very minute, ovate, entire or bifid. Inflorescence ? postical ; sub-bracts small, oval, bidentate, concave ; sub-bracteoles oblong, entire ; bracts somewhat larger than the leaves and often very much more pellucid, somewhat recurved, cuneate-oblong, almost to the middle bifid, with 1, 2 or 3 smaller segments, which are apiculate or acute dentate, rarely entire ; bracteole somewhat similar, but smaller. Perianth 3 to 6 times longer than the leaves, subulate-fusiform, cylindrical below, trigonous above, hyaline, near base 3 to 4 cells thick, upper portion composed of one layer of cells, mouth small through the folds, laciniate-ciliate, cilia very narrow, 2 cells long. Calyptra obovate, 1 to 2 cells thick near the base. Capsule spherical, dark brown, valves dividing to the base, thin texture. Spores small, smooth, dark reddish-brown. Elaters bispiral, rather broader than the spores, of the same colour. Andrcecia postical amentiform, catkins minute ; perigonial bracts about 8 pairs, closely imbricate, concave-complicate, oval, bidentate, segments acute; bracteole oblong, entire or slightly emarginate; antheridia single, oval, stipitate. Fruits April, May. Dimensions. — Stems 1 to 3 inches long, "15 mm. diameter; leaves 75 mm. x 'G mm., 7 x "G mm., 7 mm. x "8 mm., 7 mm. X 7 mm., 'GS mm. x "Go mm., 75 mm. x 75 mm. ; cells '035 mm. X '02 mm., '025 mm. x 02 mm., "02 mm. x '02 mm., "03 mm. x '03 mm., -03 mm. x 02 mm. ; sub-bracts '5 mm. x '4 mm. sinus, '1 mm., "5 x '35 mm., "3 mm. x '25 mm. ; sub-bracteoles •3 mm. X "2 mm. ; bracts 1'25 mm. x 75 mm. segments, '4 mm., 'G mm., I'l mm. x Qb mm. segments, '45 mm. ; bracteole 75 mm. x '5 mm. segments, "25 mm. ; perianth 4" mm. x 75 mm., 2"G mm. X "G mm. ; teeth at the mouth '075 mm. ; calyptra 1* mm. x "Smm.; pistillidia '15 mm. x "025 mm. ; capsule "G mm. x "55 mm.; valves ODOXTOSCHISMA. 173 •7 mm. X '3 mm. ; pedicel "15 mm. diam. : spores 008 mm. diam. ; elaters '1 mm. x Ol mm. ; perigonial bract, explanate, '55 mm. X "4 mm. sinus, '15 mm., "55 mm. x "55 mm. sinus, 'i^ mm.; perigonial bracteoles '3 mm. x "lo mm.; antlieridia 08 mm. X -OG mm. Hab. — Growing in dense patches on damp moors or in bogs, often straggling loosely amongst Sphaf/na and other mosses. 1, 2, 4, 7 to 17, I. Found on the Continent and in North America. Obs. — At once distinguished from all other round-leaved species by its postical trigonous perianth and postical amentiform androecia ; when barren, by its arcuate stems and abundant stout flagella. From C. (0.) deniidata (Mart.), see notes under that species. Var. alpina, MS. is a small neat form of a reddish-brown colour with smaller cells, firm walls, with indistinctly thickened angles, perigonial bracts ovate-rotund, with a third, antical, basal tooth, found on the Scotch and Welsh mountains. I have not been able to observe other characters to enable me to separate it from the type. Description of Pj;ATe LXV. — Fig. 1. Plants natural size. 2. Portion of stem x IG. 3-7. Leaves x 24. 8. Portion of leaf X 290. 9 12. Sub-bracts x 24. 13, 14. Sub-bracteoles x 24. 15, IG. Bracts x 24. 17. Bracteole x 24. 18. Perianth x 11. 19. Cross- section of perianth near the base x IG. 20. Ditto near the apex x 16. 21. Portion of the mouth of the perianth X 85. 23-26. Peri'^onial bracts x 24. 27, 28. Perio-onial bracteoles x 24. 29. Antiieridium x 85 (All, Foulshaw Moss, Westmorland, (jfeoro-e Stabler). 174 HEPATIC.^. 13. Cephalozia denudata (Nees), Sjjnice. Jungermania denudata, Nees in Mart. Crypt. Erlang. p. xiv. (1817). Jimgermania scalaris b denudata, Mart. Crypt. Erlang. p. 185, t. G, f. 58, b. (1817). Jungermania S'phagni, Hook, et al. ex p. (181(5). Pletrroschisma denudatum, Dum. Syll. Jung. CD (1831). Odontoschisma denudatum, Dum. Recueil, p. 19 (1835). Sphagnaecetis communis macrior, Nees in G.L.N. Syn. Hep. p. 149 (1844), Sphagncecetis Huehneriana, Rabenh. Handb. ii. 5, p. 338 (1848). Cephalozia deniodala. Spruce, On Cephalozia (1882). Dioicous, intricately csespitose, flagelliferous, small, greenish- to reddish-bi'ovvii in colour. Stems almost leafless at the base, rhizomatous, creeping, subramose, cortical cells about 40, similar to the inner, 10 x 8 cells in diameter; branches subfasciculate, erect, gemmiparous ; radiculose, rootlets numerous on the creeping stems ; flagella postical, few. Leaves imbricate, patent-divergent, never secund, concave, broadly ovate or oval, entire, sometimes retuse or emarginate, slightly decurrent antically, largest near the middle of the stem, becoming gradually smaller above and gemmiparous ; texture firm ; epidermis verruculose ; cells small, roundish-quadrate, walls thick, angles thickened, no trigones. Stipules everywhere present, increasing in size on the gemmi- parous branches and becoming equal in size to the terminal leaves, oblong-roundish-quadrate, broadly oval or lanceolate, entire, acute or subdenticulate. Inflorescence $ postical ; sub-bracts oval or ovate, bidentate to about a sixth ; sub-bracteole oblong, entire ; bracts twice bifid, margin denticulate or rarely entire ; bracteole lanceolate, acute, minutely bidentate. Perianth narrowly fusi- form, cylindrical below, above obtusely trigonous, mouth small, denticulate or shortly fimbriate, at the base 3 cells thick, near the middle 2, upper lialf composed of a single layer of cells. Calyptra very delicate, composed of a single layer of cells. Pistillidia long, narrow, about 12. Androecia postical, amenti- form, arcuate-decurved, perigonial bracts 3-8 pairs, deeply saccate, margin entire, emarginate or incised, segments obtuse or obtusate. ODONTOSCHISMA. 175 Fruits April, May. Dimensions. — |- to 1 inch long, diam. 2 mm.; leaves 'J mm. X "5 mm., 7 mm. x '7 mm., 1' mm. x -9 mm., 'G mm. x '5 mm., '7 mm. X -(3 mm., '8 mm. x 'G mm.; cells '025 mm. x 02 mm., •02 mm. X -0172 mm., '02 mm. x '02 mm. ; stipules '4 mm. x •25 mm., '35 mm. x '25 mm., '5 mm. x 2 mm. ; sub-bracts 'G mm. X -4: mm.; sinus •! mm., "7 mm. x -4 mm., -35 mm. x -2 mm., •4 mm. X •S mm., 'o mm. x ^3 mm. ; sub-bracteole ^3 mm. x ^2 mm. ; bract '7b mm. x •Q mm., segments '3 mm. ; bracteole •!) mm. x •45 mm., sinus "05 mm. ; perianth '2 mm. x -G mm.; pistillidia •2 mm. X ^03 mm. (andnecia '5 mm. — 1^5 mm. long x ^25 mm. thick S.O.L.). Hab. — Growing on rotting trunks of trees or dead vegetable matter, or on crumbling sandstone. Somewhat rare ; very rare in fruit. 2. On crumbling sand rock, Tunbridge Wells, Dr. Spruce; A rdingley Rocks, Sussex, G.Davies. 7. Tyn-y-Groes, Merioneth, irUd §• Holt. 8. Stirrup Wood, Derbyshire, G. J. Holt. 9. Car- rington Moss, Cheshire, G. A. Holt; Clifton Junction, Lane, C. J. Wild. 12, 13. Burmurray, Balmaclellan, New Galloway, J. Mc Andrew. IG. Glen Finnan, Br. Carriiujtou ; Moidart, /S'. M. Macvicar. I. Co. Mayo, D. Moore. Found on the Continent, North America ; also on the Andes of South America, Dr. Spruce. Obs. — Differs from Cephalozia (0.) Sjjhafjni (Dicks.) in its intricately branched leafless caudex, branches usually erect, atten- uate, gemmiparous ; leaves never secund, destitute of any distinct margin, epidermis verruculose, abundant presence of stipules everywhere. Ceph. [0.) Spliacjni (Dicks.) has stems often repeatedly arcu- ately branched, which are equally leaved throughout, often secund, with a distinct margin, stipules absent or few and extremely minute. " Cepli. (0.) deHudaia\'& usually found growing on rotting wood or crumbling sandstone; CcjjIi. {(J.) Sphof/ni on living /iyV/r/y //a and other bo"- mosses." li. S. 176 HEPATICJE. Desceiptiox of Plate LXVI. — Fig. 1. Plants natural size. 2. Portion of stem (after Hooker). 3. Portion of stem, antieal view X 24 (Sweden, S. 0. Linclberg, also the following). 4-9. Leaves x 24. 10. Portion of leaf x 290. 11-15. Stipules x 24. 16-20. Sub-bracts x 24. 21. Sub-bracteole x 24. 22. Bract X 24. 23. Bracteole x 24. 24. Perianth x 24. 25. Cross-section of perianth near the base x 24. 26. Ditto, near the middle. 27. Ditto, near the apex. Subgenus 3. CEPHALOZIELLJ, Spnice. Plants small or minute, often creeping amongst mosses or other hepatics, or densely csespitose, sublurid. Stem for the size of the plant very often firm, generally at the base subrhizomatous, but flagella wanting ; cortical cells 10-20, hardly different from the interior. Lower leaves distant subobliquely inserted (succubous) ; upper more crowded, or almost exactly transverse, the length (•1--15 mm.) rarely exceeding the breadth of the stem, very often cuneate, to the middle or more bifid, carinate, segments subcompli- cate or divergent, entire or in one species subdenticulate, rarely spinulose ; cells small-minute (diam. vg-yVr nim.) subquadrate or guttulate. Stipules very variable, sometimes in one and the same species everywhere present, now altogether obsolete ; always (where present) small, narrow, entire or bifid. Inflorescence for the most part dioicous, in few monoicous ; ? very often terminal on the chief stem, in other species cladogenous, or in various places. Bracts moderately large 3 -pairs (or more) bilobed, tristi- chous, very often with the bracteole highly connate, lobes generally denticulate or spinulose. Perianth for the most part elongate and narrow, leptodermous, acutely prismatic, keels rarely reduced to 3, sometimes 3, 4, 5, and also 6 in the same species, mouth denticu- late or ciliolate, rarely submuticous. Calyptra delicate. Capsule oblong-globose. Androecia in the middle or at the end of the chief stem or elongated branches, bracts not smaller than the leaves, very rarely produced on slender postical branches. CEPHALOZIELLA . 177 14. Cephalozia divaricata (>S////M), Dam. Jungermania divaricata, Sm. Eng. Bot. t. 711) (a. 1X00) et t. 2A^ij'^. Jungermania bijssacea, Roth. Tent. fl. Germ. i. p. 387 (ISOO) ; Hooker. ISrit. Jung. t. 12 (181G), &c. Jungermania con/ervoides, Radcli in Act. soc. Modena, p. 29, t. 4, f. 1. (1818). Jungermania Starkii, Herb. Funck. Nees Nat. Eur. Leb. 11, 225 (18.'5G). Cephalozia divaricata (Sm.) Dum. Recueil (ISoo). Dioicous, densely ctespitose or creeping amongst mosses, flagel- liferous, small, colour very various, namely, green, olive, sometimes reddish, rarely whitish or almost black. Stems prostrate or sub- erect, firm, 6-8 cells in diameter, cortical cells 14-20 scarcely diff'erent from the inner ; simple or slightly branched ; radiculose, rootlets white, Leaves small, distant on sterile stems, on fertile very often subimbricate, accrescent, subsuccubous or almost trans- verse, cuneate or rotund-quadrate, nearly to the middle bifid (rarely a little more), segments complicate or divergent, ovate triangular acute or subacuminate, rarely obtuse, entire, rarely with one or two teeth ; cells from minute to small, subquadrate, leptodermous, pellucid or subopaque, walls firm, no trigones or thickened angles. Stipules lanceolate or ligulate, lower ones minute, entire, upper larger, apex often bifid (rarely trifid), sometimes wanting altogether or only present near the flowers. Flowers % terminal on chief stem or elongated branches (rarely on short ones) ; bracts 3 pairs, tristichous, larger than the leaves, innermost sometimes more or less connate, rotund-quadrate, bilobed, segments acute, denticulate or subspinose ; bracteole adnate to bract or free, ovate-quadrate, bilobed, rather smaller than the bracts. Perianth linear or narrowly fusiform, composed of one layer of cells, often bicolour, towards the base purple, apex white prismatic, witli very rarely only 3 angles, usually 4, 5 or 6 ; mouth subconstricted, denticulate or subentire. Calyptra smaller, delicate. Pistillidia about 12. Capsule oblong-globose. Androecia on proper stems, variously situated on stem or branch, perigonial bracts several pairs, sub- imbricate, equal, incurvo-concave, bilobed, segments somewhat acute ; autheridia single, oval. M 178 HEPATIC^. Fruits March, April. Dimensions. — Stems J to f inch long, '05 to "07 mm. in diameter, with leaves '2 mm. wide; leaves '15 mm. x 'IS mm., seg. -075 mm., -175 ram. x -175 mm., seg. '1 mm., '2 mm. x •15 mm., seg. '075 mm. ; cells '02 mm. x '02 mm., -0225 mm. x •0225 mm., -02 mm. x "03 mm. ; stipules -15 mm. x '075 mm., •125 mm. x '075 mm. ; sub-bract '275 mm. x -3 mm., seg. '1 mm. ; sub-bracteole -175 mm. x -125 mm., seg. "075 mm. ; bracts '35 mm. X '4 mm., seg. •225 mm., ^5 mm. x ^5 mm., seg. •175 mm. ; bracteole '225 mm. x '15 mm., seg. '075 mm. ; perianth '75 mm. x •25 mm., '8 mm. x ^4 mm., 1' mm. x '5 mm. ; pistillidia "15 mm. x *05 mm.; valve of capsule '325 mm. x 125 mm.; antheridia •1 mm. X -075 mm. Hab. — Growing on somewhat damp rocks, stones, earth or rotting wood, exposed or shaded ; on heaths, or creeping upon other hepatics or on mosses. Very common. 1 to 17. I. Europe, Asia and North America. Obs. — A common and variable species, which has stipules always near the bracts, but on other parts of the stems some- times without and sometimes with. The type with stipules has been considered a distinct species [CejjJialozia Stai'lm^ Funck), but agreeing with Dr. Spruce, I find no other characters of sufficient value to make it worthy of specific rank. C. divaricafa is a small species with wiry stems distinguished from its nearest allies Ceph. Jackii^ Limpr. and Ceph. sfellulifera (Tayl.) by its dioicous inflorescence. - Ceph. Jeucantlia, Spruce, which is also dioicous, is of a much paler colour, stems more fragile, leaves more oblong, with segments parallel or connivent, not divergent, stipules always absent. Description of Plate LXVII. — Fig. 1. Plants natural size. 2. Portion of sterile stem x 85 (Sussex, E. N. Bloomfield). 3-5. Leaves x 85 (Warmbrunn, Hb. Nees). 6-8. Ditto x 85 (Glen Finnan, Dr. Carrington). 9, 10. Ditto x 85 (Stockholm, S. 0. Lindberg). 11. Ditto x 85 (Altona, Dr. Gottsche). 12. Portion of leaf x 290 (ditto). 13-15. Stipules x 85 (ditto). CEFHALOZIELLA. 179 IG. Sub-bract x 85 (ditto). 17. Sub-bracteole x 85 (Blanken- berg, Hampe). 18. Bract x 85 (Warmsbrunn, Hb. Nees). 19. Bracteole x 85 (ditto). 20. Perianth x G4 (Altona, Dr. Gottsche). 21. Mouth of perianth x 85 (Alliers, Du Buysson). 22. Antheridium x 85 (Llangollen, W. Wilson). 15. Cephalozia stellulifera {TajjI. MSS.) Jnngermania stellulifera, Tayl. MSS., G.L.N. Syn. Hep. p. 134 (1844), Jnngerrtxania Starkii, var. procerior, Nees G. & R. Hep. Eiir. n. 02.5. Cephalozia divaricata (Sm.), var. stellulifera, Spruce, On Oeph. (1882). Monoicous, acrocarpous or cladocarpous, densely csespitose, eflagelliferous, small, pale green or subolive to reddish-brown. Stems creeping or suberect, simple or slightly branched, flexuose, firm, sometimes thick and fleshy, cortical cells 15-25, similar to the inner, sometimes brown, sometimes hyaline 8-10 in diameter ; radiculose, rootlets few, pellucid. Leaves distant or approximate, near the apex of fertile branches crowded, broader than the stem, patent-divergent, below concave-complicate or spreading, above widely spreading or almost squarrose, oval-quadrate, bifid to nearly the middle, sinus acute or obtusate, segments 5, 6 cells wide at the base, acute or obtusate, divergent or erect ; cells small, 4-, 5- and 6-angled, subquadrate, walls firm, no trigones. Stipules distinct on some stems, lanceolate-acute, sometimes above (sub- bracteoles ?), bifid, sometimes minute or wanting. Flowers $ on apex of main stem or on long or short branches. Bracts much larger than the stem leaves, ovate-quadrate, bifid to about ^, sinus broad, acute or obtusate, segments narrowly triangular, acute, dentate ; bracteole smaller than the bracts, oval or ovate, bifid to about ^, or ovate-lanceolate and entire, margin dentate or denticu- late ; sub-bracts somewhat similar to the leaves but larixer, den- ticulate or quite entire ; sub-bracteole bidentate or entire, margin denticulate or quite entire. Perianth projecting much beyond the bracts, large in proportion to the size of the plant, oblong, or elongate-fusiform, unistratose, trigonous, angles acute, mouth hyaline, scariose and indistinctly crenulate. Pistillidia about 10. 180 HEPATIC^. Calyptra obovate, of a delicate texture. Capsule spherical, dark brown. Spores brown, elaters paler than the spores, Androecia on long or short branches proceeding often from below the peri- anth, catkins long or short, perigonial bracts 4-8 pairs, closely imbricate, roundish-quadrate when explanate, bifid to about \, sin as and segments acute, margin quite entire or dentate, compli- cate-concave, ventricose ; antheridium solitary, oval. Fruits March, April. Dimensions. — Stems \ io \ inch long, "08 mm. in diameter, with leaves '35 mm. wide ; leaves '2 mm. x '16 mm., segments •1 mm., "175 mm. x '125 mm., seg. "075 mm., "175 mm. x '15 mm., seg. -075 mm., '225 mm. x -175 mm., seg. '075 mm. ; upper leaves •3 mm. X -25 mm., seg. "125 mm. ; cells "02 mm., '025 mm., '0175 mm. ; sub-bracteole '2 mm. x '14 mm. ; bracts "425 mm. x •325 mm., seg. '175 mm., '4 mm. x -35 mm., seg. '15 mm.; bracteole '25 mm. x -15 mm., seg. *! mm., '075 mm., '325 mm. x •2 mm. ; perianth -8 mm. x "3 mm., "9 mm. x '35 mm. ; pistillidia '1 mm. X "04 mm. ; spores '01 mm. ; elaters '13 mm. x -01 mm. ; perigonial bracts '21 mm. x •21 mm., seg. ^07 5 mm. ; antheridia •15 mm. X •! mm. Hab. — On hedge banks, rocks and stones, rarely on rotting wood. Rare. 1. Hedge banks near Redruth, Cornwall, 1871, The Lizard, Cornwall 1883, W. Curnow. 5. ? 8. Crich, Derby- shire; W. Wilson, 1833. 12. Westmorland, G. Stabler ; on dead trees, Rigmaden Park, Westmorland, Peter Brecsen. 15. Scottish Highlands, Drummond. Found on the Continent (Schwartzwasser, Sudetia, Hb. Nees, as Jift/f/, Starkii, var. mimiua). Oes. — Differs from C. divaricata (Sm.) in its monoicous inflorescence, leaves often very widely spreading, at the immature flower ends squarrose (hence I suppose Taylor's name), perianth always trigonous (not 4-6-angled), angles acute. In the original specimens from Crich the perigonial bracts are dentate, in those from Cornwall and elsewhere they are quite entire. Description of Plate LXVIII. — Fig. 1. Plants natural size. CEPHALOZIELLA. 1 8 1 2. Portion of fertile plant x 64 (Cornwall, Curnow). 3. Portion of branch, postical view x 31 (ditto). 4-8. Leaves x 85 (ditto). 9. Leaf x 85 (Westmorland, P. Dreesen). 10. Ditto x 85 (Scotland, Drummond), 11-14. Leaves x 64 (Crich, Wilson). 15. Upper leaf x 64 (Cornwall, Curnow). 16. Portion of leaf X 290 (ditto). 17, 18. Stipules x 64 (ditto). 19. Sub- bracteole x 85 (Scotland, Drummond). 20, 21. Bracts x 85 (Crich, Wilson). 22. Bracteole x 85 (Cornwall, Curnow). 23. Perigonial bract x 64 (ditto). 24. Antheridium x 64 (ditto). 16. Cephalozia aeraria, Fearson. Cephalozia m'aria, Pears., Spruce, On Ceph. (1882). Dioicous, cladocarpous, densely depressed csespitose, eflagelli- ferous, minute, tawny or pale brown colour. Stems flexuose, extremely fine, rootlets closely intertwined, moderately strong, sparingly branched, branches subhyaline, foliose; cells of the stem 8 in diameter, cortical ones 10-15, slightly larger than the inner. Leaves dissitous-squarrose, almost patent, minute, sub- cuneate, obscurely carinate, deeply (to fths) bilobed, subentire, sinus acute or obtuse, segments ovate or lanceolate, at the base 2-4 cells broad, sub-acuminate, apical cell single, conical (twice as long as broad), very often incurved, apiculate; cells minute, oblong, pellucid, distinctly guttulate, walls at the angles thickened, epidermis rough. Stipules variable, lower ones often minute or obsolete, upper ones about half the length of the leaves, lanceolate, sometimes connate to the nearest leaf, forming a three-lobed one, the largest rarely bilobed and scarcely smaller than the leaves. Fertile branches very short ; bracts 3 pairs, nearly twice as long as the leaves, in sterile flowers free, a little more than to the middle bilobed, lobes ovate, acuminate. Further organs not yet known. Dimensions. — Stems \ inch long, '05 mm. in diameter, with leaves -25 mm. wide; leaves '125 mm. x '075 mm., segments -06 mm., -15 mm. X -1 mm., seg. "1 mm., 175 mm. x -1 mm., seg. 182 HEPATIC^. •075 mm., •125-'15 mm. x -125 mm. ; cells '015 mm., -015 mm. x •0175 mm., "02 mm. x '0175 mm., '01 mm. x "0125 mm., '0125 mm. X 015 mm. ; apical cell '02 mm. x '01 mm. ; stipules "1 mm. x •02 mm. at base, "1 mm. x '03 mm., '075 mm. x '025 mm. ; bracts "2 mm. x "125 mm., seg. ^075 mm., '1 mm., -25 mm. x •15 mm. ; bracteole '1 mm. x ^06 mm., seg. "04 mm. Hab. — 7. At the mouth and on the roof of an old copper mine between Dolgelly and Tyn-y-Groes, Merionethshire, W.H. Pearson, April 1877. Ehaider Dhu, Tyn-y-Groes, G. A. Holt, July 1882. Mouth of copper mine, near Pistyl Cain, Tyn-y-Groes, C. J. Wild, July 1882. Extremely rare. Obs. — " Cejjli. Macounii, Aust., comes near to it, but is of a green colour, stems slender and much branched, leaves subimbri- cate, sinus sometimes lunate, apices of lobes not unguiculate, cells subquadrate, stipules none. Cejjh. diraricata (Sm.) certainly dis- tinct, flowers acrogenous ; leaves distinctly carinate, rarely more than to the |- divided, exunguiculate, cells quadrate, reticulate (not guttulate). It differs essentially from C. divaricata in being still more minute, in the deeply bifid leaves whose narrow seg- ments end in a clawlike apiculus, the guttulate areolation and especially in the cladogenous inflorescence which brings it near to C. Macoimiir — R. Spruce. Description of Plate LXIX. — Fig. 1. Plants natural size. 2. Portion of stem x 85. 3-14. Leaves x 85. 15. Leaf with connate stipule x 85. 16. Portion of leaf x 290. 17-23. Stipules X 85. 24, 25. Bracts x 85. 26. Bracteole x 85 (Tyn-;y- Groes, W. H. P., April 1877). 1 7. Cephalozia Jackii, Lintprkld MjSS. Cephcdozia Jackii, Limpi\, MSS. Spruce, On Cephalozia, p. 67 (1882). Paroicous, csespitose, eflagelliferous, small, size of Cejj//. divaricata, aero- and cladocarpous, pale green to reddish-green, with the flowering apices often purplish. Stems short, fragile, simple or with few branches, prostrate, radiculose, ends ascending, some branches elongate, with minute leaves. Lower leaves, chiefly CEPHA L OZIEL LA . 183 of the sterile branches, distant, minute, cnneate, upper near the ? flowers imbricate, cuneate-quadrate or subrotund, all subcari- nate, almost or even to the middle acute bilobed, segments ovate- triangular, acute; cells very small to small, quadrate to sub- elongate, 4-, 5- and 6-sided, angles thickened, no trigones. Stipules half the size of the leaves, lower ones linear or lanceolate, upper ones ovate-lanceolate, rarely divided at the apex. Involucre ovate-juliform. Bracts 3-5 pairs, larger than the leaves, oblong- orbiculate, concave, less deeply bilobed, more or less denticulate, enclosing single antheridium, which is oval, the innermost bracts highly connate, forming with the smaller bracteole (with which both bracts are connate) a cuplike receptacle. Perianth twice as long as the bracts, projecting above half its length beyond them, oblong or ovate-oblong, obtusely 4-5-angled (ver}^ rarel}' trigonous), composed of a single layer of cells, about 80 round, near the middle, mouth scariose, truncate, repand, afterwards variously divided. Capsule oblong. Fruits March, April. Dimensions. — Stems J to J inch long, '1 to "15 mm. in diameter, with leaves "5 mm. wide; lower leaves 'l mm. x 'l mm., •15 mm. X '12 mm. ; upper leaves '275 mm. x '225 mm., segments •15 mm., -225 mm. x '175 mm., seg. "1 mm., ^3 mm. x '275 mm., seg. •IS mm. ; cells "02 mm., "015 mm. x •0125 mm., ^0175 mm. X ^0175 mm., ^015 mm. x •OlS mm., '02 mm. x ^0175 mm.; stipules "15 mm. X "075 mm., ^175 mm. x '06 mm.; sub-bracts •3 mm. X "3 mm., seg. "1 mm., •S mm. x ^25 mm., "35 mm. x "325 mm,, seg. '125 mm., '4 mm. x ^4 mm., seg. '175 mm. ; sub- bracteole "225 mm x •IS mm. ; bracts '4 mm. x "3 mm., "55 mm. X "425 mm., seg. '175 mm. ; bracteole ^4 mm. x '175 mm. ; antheridia "15 mm. x •! mm. ; perianth 1"25 mm. x 'b mm., "7 mm. X '25 mm. Hab. — On sandy paths or banks. Very rare. 1. Hayle, W. Cornwall, W. Curiww, April 1842, June 1883. 9. "Four miles from Warrington, Cheshire, towards High Leigh, April 1841, W. Wihon, 2,^ Jang, hpsacea (fruit)." W. Wilson in litt. ad R. Spruce. 12. Whitbarrow, G. Stabler. 184 HEPATICjE. Found on the Continent (Germany). Obs. — Distinguished from Cci^lt. divaricafa (Sm.) by its paroicous inflorescence, Lracts more highly connate, entire brac- teole, not bifid or ever free; remarkable at first sight by its swollen involucre. Description of Plate LXX. — Fig. 1. Plants natural size. 2. Fertile stem with perianth x 24 (nr. Warrington, W. Wilson). 3-5. Leaves x 85 (ditto). 6. Leaf x 64 (Custrin, Germany, Flotow, Hb. Limpricht). 7. Portion of leaf x 290 (nr. War- rington, W. Wilson). 8, 9. Stipules x 85 (ditto). 10. Sub- bract X 64 (Custrin, Flotow). 11. 1st sub-bract x 64 (nr. Warrington, W. Wilson). 12. 2nd sub-bract x 64 (ditto). 13. 3rd sub-bract x 64 (ditto). 14. Sub-bracteole x 64 (Custrin, Flotow). 15. Antheridium x 85 (ditto). 16. Bracts x 84 (nr. Warrington, W. Wilson). 17. Cross-section of perianth near the base x 24 (Cornwall, Curnow). 18. Ditto, near the middle (ditto). 18. Cephalozia elachista (Jac/>:), Lindb. Jungermania elachista, Jack, in G. & R. Hep. Eur. no. 574 (1873) ; Limpr. Ki ypt. Schl. 1. p. 296 (187(5). Cephalozia elachista, Lindb. Hep. in Hib. p. 502 (1874); tSpruce, On Cephalozia, p. 70 (1882). Monoicous, clado- or acrocarpous, laxly csespitose, small, pale green. Stem rooting at the base, rootlets few, distant, very deli- cate, prostrate, subaphyllous, with few branches. Leaves distant, subimbricate only towards the apices of the fertile branches ; oval, deeply acutely bilobed, segments broadly subulate, acuminate, incurved, entire or furnished with one or two teeth ; cells small, subquadrate or elongate, pellucid, sub-leptodermous, walls thin, no thickened angles or trigones. Stipules minute, entire or bifid, acuminate, sometimes wanting. Branches ? short or elongate, often innovantly proliferous ; bracts much larger than the leaves, partly free, deeply bilobed, denticulate or subspinulose ; segments finely acuminate. Bracteole similar. Perianth elongate, trigonous- CEPHALOZIELLA. 185 prismatic, mouth slightly denticulate, teeth few, incurved. Capsule oval. Spores and elaters reddish-brown, elaterebispiral. Amenta ^ , terminal on stem or occupying almost the whole branch ; perigonial bracts narrowly oval, deeply bilobed, denticulate, segments acumi- nate, above secund ; perigonial bracteole oblong, bifid, segments acute ; antheridia small, oval. Fruits March, April. Dimensions. — Stems :f to J inch long, diameter '02 mm., •03 mm. ; leaves '175 mm. x 125 mm., seg. "1 mm., "15 mm. X "125 mm., seg. "11 mm., -135 mm. x -11 mm., seg. 115 mm., •15 mm. X '1 mm., seg. "06 mm.; cells "025 mm. x ^02 mm., •035 mm. x '015 mm., ^025 mm. x "02 mm., '02 mm. x "02 mm., '02 mm. X •0175 mm.; stipules '13 mm. x '05 mm., '1 mm. x •03 mm., seg. "04 ; bracts '35 mm. x ^25 mm., seg. "2 mm. ; perianth l" mm. x '4 mm., I'o mm. x "3 mm., Vb mm. x "4 mm. ; perigonial bracts '2 mm. x "175 mm., seg. '15 mm.; perigonial bracteoles "125 mm. x '075 mm., seg. '05 mm. ; antheridia •I mm. X ^07 5 mm. ; elaters "15 mm. x '01 mm. Hab. — Growing on damp rocks, I., near Lough Bray, S. 0. Lwdher(j, 1873. Brandon Mt., &c., B. Moore. Extremely rare. Found on the Continent ; Baden, Jacl\ Finland, S. 0. Lindbcrij. Norway, Brylui. Obs. — Distinguished from all forms of C. cUcaricafa (Sm.), and from C. Jackii, Limpr. by its monoicous inflorescence ; it is remark- able for its very slender habit, distant, deeply bilobed leaves, which are either entire or slightly denticulate. Description of Plate LXII. — Fig. 1. Plants natural size. 2, 3. Portions of stem x 85. 4-9. Leaves x 85. 10. Portion of leaf X 290. 11, 12. Stipules x 85. 13. Bract x 85 14-16. Segments of bracts x 85. 17. Perianth x 24. 18. Perigonial bract x 85. 19. Perigonial bracteole x "ib. 20. Antheridium x 85. (Gr. & Pi. Hep. Eur. no. 274, Salem, Baden, Germany, Jack.) 186 HEPATICJ^. 19. Cephalozia leucantha, Spruce. Cephcdozia leucantha, Spruce, On Cephalozia, p. 68 (1882). Dioicous, almost always cladocarpous, sometimes acrocarpous, csespitose, eflagelliferous, albescent, small, size of Ceph. divaricafa (Sm.). Stems prostrate, with ^ and ? often intricately entangled, slender, fragile, elongate, flexuose, simple or subramose, radiculose. Leaves small, distant, patulous or assurgent, oblong or quadrate- rotund, bifid to about the middle or more, sinus acute or obtuse, segments often unequal, broadly subulate (3-4 cells broad at the base), acute or subacuminate, parallel or connivent; cells very small, subquadrate, lower ones slightly elongate, walls firm, no trigones or thickened angles. Stipules wanting. Bracts 3 or 4 times larger than the lower leaves, more or less connate, orbiculate, subdenticulate, 2-3-lobed, segments short, acuminulate. Perianth large for the size of the plant, 10 times longer than the lower leaves, ovate-lanceolate-fusiform or sublinear, white, leptoderm.ous, above 3- (rarely 4-) augled, composed of a single layer of very small cells, about 100 round near the middle, mouth minute, often obsoletely setulose. Calyptra delicate. Capsule large, often equal to half the size of the perianth, oblong-cylindrical, badious. Androecia short, rarely occupying the whole of the branch, terminal or medial, julaceous; perigonial bracts closely imbricate, orbiculate, very concave, carinate, to a third or more bi-trilobed, monandrous ; bracteole minute, linear-subulate ; antheridia small, oval. Fruits Spring and Autumn. Dimensions. — Stems |- to J inch long, "075 mm. to "1 mm. in diameter, with leaves "2 mm. wide; leaves '2 mm. x "13 mm., •125 mm. x "15 mm., segments "075 mm., "06 mm., '175 mm. x •175 mm., seg. "075 mm., '15 mm. x -175 mm., seg. '075 mm., •2 mm. X "25 mm., seg. '15 mm., '125 mm., '225 mm. x '2 mm,, seg. '125 mm., 075 mm.; upper leaves '275 mm. x '25 mm., seg. •125 mm., -175 mm.; cells '0167 mm., "015 mm. x '0175 mm., •02 ram. X '02 mm., "015 mm. x "015 mm., '0225 mm. x "02 mm. ; sub-bracts "3 mm. x "25 mm., ses". '15 mm., *4 mm. x '4 mm., ses". CEPHALOZIELLA. 187 •2 mm. ; bracts "75 mm., -4 mm. x "4 mm., seg. '15 mm., -35 mm. X "3 mm., seg. '15 mm. ; bracteole '45 mm. x -25 mm.; perianth 2* mm. X '5 mm., 1*25 mm. x "35 mm.; pistilliclia "075 mm. x "025 mm. ; valve of capsule '5 mm. x '175 mm. ; perigonial bracts •25 mm., '2 mm. x 2 mm., seg. '125 mm. ; antheridia '075 mm. x "06 mm. Hab. — On rotting wood, in very fine fruit. 15. Portach, near Banchory, Aberdeenshire, along with Ccj)]i. catoudata (Hiiben.) and Cepli. ciTvifuIia (Dicks.), /. Sim. Extremely rare. Found on the Continent. Obs. — Cej)h. cateiiuhd(( (Hiiben.) is very different in its larger size and fulvous colour to C. hucantha, leaves subimbricate, broader and more concave, when dry incurved and catenulate, cells larger, but chiefly by the deeply triplicate perianth, with mouth distinctly ciliolate. C. leiiccuitha is also certainly ver}' distinct from CejjJi. lumdafoUa, Dum., which has larger, rhomboid- rotund decurrent leaves, divided only to a third, sinus more or less lunate, cells larger; perianth (in comparison with the leaves) much shorter and like the calyptra fleshy (composed of several layers of cells), &c. &c. "R S." Description of Plate LXXII. — Fig. 1. Plants natural size. 2. Portion of sterile stem x 64. 3-7. Leaves x 85. 8, 9. Tipper stem leaves x 85. 10. Portion of leaf x 290. 11-13. Sub-bracts x 85. 14, 15. Bracts x 85. 16. Bracteole (?) x 85. 17. Pistillidium x 85. 18. Perianth x 31. 19. Cross- section of perianth X 31. 20,21. Perigonial bracts x 85. 22. Antheridium x 85 (Portach, J. Sim). 20. Cephalozia dentata [Baddi), Lindb. Jungermania dentata, Raddi in Mem. della Soc. Ital. di Modena, six. p. 32, tab. 4, Fig. 4 (1818). Anthelia dentata, Dum. Hep. Eur. p. 99 (1874). Cephalozia dentata (Raddi), Lindb. in Journ. Linn. Soc. xiii. (1872); Spruce, On Cephalozia, p. 71 (1882). Monoicous, densely depresso-csespitose, eflagelliferous, minute, pale green w^lien young, when older dark green to brown in colour. 188 HEPATIC^. Stems creeping, simple or with one, rarely two postical branches, proceeding from near the apex, 5 to G cells in diameter, cortical cells about 12, similar to inner, all hyaline; radiculose, rootlets plentiful, long, hyaline. Leaves subtransversely inserted, patent- divergent below 60^-70°, above patent to erecto-patent 50°-30°, approximate, larger, accrescent above, often capitate, below distant and smaller, oval, ovate- quadrate or roundish, concave or slightly complicate, bilobed to the middle or below, segments subincurved, acute, acuminate or mucronate, margin irregularly spinulose- dentate, or subdenticulate ; texture delicate, epidermis smooth, cells minute to small, subquadrate, w^alls thin but firm, no trigones or thickened angles. Stipules everywhere present, minute below, lanceolate or subulate, mucronate, spinulose-dentate or denticulate. Bracts larger than the leaves, bilobed to a third, segments sub- spinose-dentate ; bracteole of equal length, broadly subulate or ovate-lanceolate, entire or bilobed to ^. Perianth projecting f beyond the bracts, delicate, hyaline or with a reddish tinge or brown, subcylindrical, plicate above, mouth slightly or not at all contracted, denticulate ; pistillidia few. Capsule dark brown, nearly smooth, as broad as the dark brown elaters. Androecia produced from the end of the chief stem on one or two very slender branches, catkinlike; perigonial bracts several, larger than the stem leaves, erect, concave, bilobed to about a third ; antheridia large and round. Fruits April. Dimensions, ^Stem \ inch long, "Oo mm. diam., with leaves '3 mm. wide ; stem leaves '25 mm. x "225 mm., segments '125 mm, ; branch leaves '165 mm. x '125 mm., seg. *! mm., cells "0175 mm. ; cells of marginal teeth "03 mm. x "01 mm.; stipules '14 mm. X "05 mm. Hab. — On damp shady earth or rocks. Extremely rare. 1, Curbis Bay, near Penzance, W. Mitten , W. Ciinwiv, G. E. Lewies, Mrs. Mia M. Tindall. 2. Ashdown Forest, Sussex, coll. G. E. Daviesy detected by JF. Mitten. Found on the Continent (Italy, France, Germany). Obs. — Distinguished from Ceph. elacJnsta (Jack.) by its less FHWXOLOnrS. 189 slender habit, shorter stems, with closer upper leaves, which are not so deeply divided, margin spinulose-dentate, stipules larger and spinulose-denticulate. From Prioiiolohiis Turiicr'i (Ilook.) by absence of any lateral branches, presence of stipules and texture of leaves, which is not guttulate. Dr. Spruce conjectured that the inflorescence might be dioicous, but Herr Linipricht has found copiously fertile plants which prove it to be monoicous. All the specimens I have been able to examine were barren ; the description of the lertile plant is taken from Limpricht. Description of Plate LXXIII. — Fig. 1. Plants natural size. 2. Portion of stem, postical view x 85 (Cornwall, Curnow and Davies). 3-5. Branch leaves x 85 (ditto). 6-8. Leaves x 85 (ditto). 9-12. Ditto x G4 (Italy, Carestia). 13, 14. Ditto x G4 (Grascony, Spruce). 15. Portion of leaf x 290 (Cornwall, Curnow and Davies). 16-19. Sti[)ules x 85 (ditto). 20. Stipule X 64 (Italy, Carestia). 21. Ditto x 64 (Grascon}', Spruce). 22. Ditto X 85 (Ashdown Forest, G. E. Davies). Genus 15. PRIONOLOBUS, >S/>/v/rY. Jnngermania, Hooker, Brit. Jung. t. 29 (I SIC). Anihelia, Dum. Recueil, p. 18 (1S;}5). Cephalozia, Lindb. in Journ. Linn. Soc. viii. (187P>). Prionolobns, Spruce, Hep. Am, et And. p. i>OH (188")). Acrocarpous, fertile branches sometimes al)breviate, often elongate. Branches all lateral, very rarel}^ with an occasional postical one. Leaves dentate, complicate, pectinato-distichous. 190 HEPATIC^. Prionolobus Turner! {Hook.), Spruce. Jungermania Tio'neri, Hook. Brit. Jung. t. 29 (1816). Anthelia Turneri, Dum. Recueil, p. 18 (1835). Cephalozia Tarneri, Lindb. in Journ. Linn. Soc. xiii. (1873) ; Spruce, On Cepha- lozia, p. 71 (1882). Prionolobus Tarneri, Spruce, Hep. Am. et And. p. 508 (1885). Monoicous and dioicous, aero- (rarely clado-) carpous, depresso- csespitose, eflagelliferousj small, fragile, pale reddish colour, at the apices greenish. Stems prostrate at the base, subradiculose. assurgent, subramose, branches lateral, rarely postical, pol}^- phyllous, terete. Cells of the stem pluristratose, very delicate, interior opaqne, cortical similar, slightly larger, subpellucid. Leaves pectinate-distichous, close and equitant, to about the middle or more complicate-bilobed, the whole margin irregularly acutely dentate-serrate, often doubly so, lobes ovate or ovate- lanceolate, acute or apiculate, antical lobe erect and subparallel with the stem, postical (a little broader), patent at an angle of about 60°, cells small to minute, quadrate-hexagonal, plane, beautifully guttulate, walls and angles strongly thickened, no trigones, slightly endochromic. Stipules none. Flowers ? terminal, often on elongated branches, innovations wanting. Bracts 1-3 pairs, the innermost almost twice the size of the leaves, connate at the antical base, spinose-dentate, bilobed, lobes subacuminate, acute ; bracteole highly connate with the adjoining bract, ovate, subacuminate, entire or bilobed, spinose-dentate. (Leaves next to the outer bracts a little larger than the others, free at the base, no folioles added.) Perianth projecting much beyond the bracts, delicate, composed of a single layer of cells, about 90 round the middle (except at the base and the angles, where there are some- times two layers) linear (at the base a little narrow), pentagonal - prismatic, keels somewhat high, apex rotuudate, mouth almost closed, obscurely ciliolate, ciliola very narrow, long, hyaline. Cal3q3tra delicate. Capsule oval. Androecia on the same or often on different stems, occupying the middle of the branch ; perigonial bracts assurgent, several pairs, somewhat similar to the leaves, sometimes larger ; antheridia single, oval. PEWNOLOBUS. 191 Fruits ]\Iai'ch, April. Dimensions. — Stems \ to ^ inch long, "OG mm. diam. witli leaves '4 mm. wide; leaves "25 mm. x '2 mm., segments "125 mm. to "15 mm. ; cells 'OlS mm. to '03 mm. ; bracts '45 mm. x 7 mm., seg. '3 mm. ; perianth 1*25 mm. x '275 mm. ; teeth at the mouth •02mm. long; perigonial bracts '25 mm. x '2 mm., seg. "075 mm. to -15 mm. ; antheridia '09 mm. x 07 mm. Hab. — " Sandy or loamy situations, under shade of bushes or on ditch banks, nearly always associated with Africlium undidatum , JJipIojjJij/U/uii albicans, and sometimes with Nardla scalaris and MarsiipeUa Fuiickil. " Very rare. 2. Tilgate and Ashdown Forest, Sussex, E. Jcnner, 1842, G. E. Dacies, 1879. 5. Wolford Wood, Warwick, -/. E. BagiiaU, 1891. 7. Dolgelly, Merioneth, W. H. P., 1875. I. By the side of a mountain rivulet near Bantry, Co. Cork, ^[iss Hutcliins, with female flowers, in March 1811? On a wet sandy bank, Cromaglown, Co. Kerry, bearing autoicous perianths, July 23, 1873, Frof. Lindherg. C. On a gravelly bank near the sea, Guernsey, E. D. Marquand, 1893. Found on the Continent (France), Canary Islands, Northern Africa, and coast counties of California, North America. Obs. — A very beautiful species, remarkable for its guttulate texture and dentate-serrate leaves. Distinguished from CcjjJtalozia deiildia (Ptaddi) by its lateral branches and absence of stipules. Although described by Spruce (whose description I follow) and others as polyoicous (monoicous and dioicous) I have nev^er met with any but dioicous plants, the t ^ind % sometimes growing- entangled together, but on separate plants. Description OF Plate LXXIV. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 85. 3, 4. Leaves x 85. 5,6. Ditto, explanate x 85. 7. Antical lobe of leaf x 85. 8. Pos- tical lobe of leaf x 85. 9. Portion of leaf x 290. 10,11. Bracts x G4. 12. Perianth x 31. 13. Cross-section of perianth x 31. 192 HEPATIC u^. 14. Portion of mouth of perianth x -85. 15, 16. Perigonial bracts x 85. 17. Antheridium x 85 (Killarney, S. 0. Lindberg). Genus IG. PLEUROCLADA, Spricce. Jungermania, Hook. Brit. Jung. t. 72 (181 G). Cephalozia, Dum. Kecueil, p. 18 (1835). Fleuroclada, Spr-uce, On Cephalozia, p. 78 (1882). Differs from its ally Hijf/rohiella by its glaucescent colour stems radiculose, all the length subsequifoliate, neither rhizomatous nor flagelliferous at the base, subpinnate ; all branches lateral, at the base, near the stem-leaf difformed (monolobed) ; leaves very concave (hardly complicate) ; perianth very fleshy, below 8 cells thick, subfloral innovations wanting. 1. Pleuroclada albescens [Hook.), Sjjruce. Jungermania albescens, Hook. Brit. Jung. t. 72 et suppl. t. 4 (181fi). Ceplmlozia albescens, Dum. Recueil, p. 18 (1835). Pleuroclada albescens, Spruce, On Cephalozia, p. 78 (1882). Dioicous, csespitose, tufts depressed, eflagelliferous, small to medium in size, of a whitish or greenish colour, when drj^ with a bluish-white tinge. Stems procumbent, laxly subpinnate, some- times subdichotomous, slightly radiculose, rooting up to apex of stem ; firm oval-terete, composed of 5 concentric subequal layers of opaque cells, cortical about 25, slightly larger than the inner. Leaves somewhat distant or approximate, patulous, almost trans- versely inserted, slightly succubous, orbiculate, very concave, almost hemispherical, to about the third or a little more bilobed, segments ovate, triangular, connivent, acute, sinus narrow, acute or subobtuse ; cells medium in size, quadrate-hexagonal, 4-, 5- and 6-sided, thick but subleptodermous, almost smooth, slight!}^ endo- chromic, subpellucid, walls firm, somewhat thick, angles only slightly thickened, no trigones. Axillary leaf produced partly from the stem and partly from the adnate branch, differs from the other leaves in being broadly ovate, subcordate at base, apex acute, FLEUROCLADA. 193 not bifid. Stipules subcontiguous, appressed, subplane, slightly shorter than the leaves, broadly ovate or ovate-lanceolate, acute or subacuminate, rarely obtuse, on one side above the base deeply uuidentate, sometimes on both sides or wanting altogether. Flowers $ terminal on short or long branches ; at the base, copiously radiculose. Bracts 3 pairs, appressed-convolute, sub- bracts a little larger than the leaves, bracts almost 3 times larger, free at the base or slightly connate, oblong-quadrate, bifid to the third, rarely trifid, segments subacuminate acute, bracteoles smaller, apex entire or bifid, on both sides near the base with 1-3 large teeth. Perianth highly exserted, 7 times longer than the stem leaves, clavate or linear- fusiform, deeply trigonous, mouth constricted, often scariose, afterwards lacerate and erose, texture firm, near the base 5-8 cells thick, near the middle 2-4 cells, at about f high, 2 cells thick, near the apex composed of a single layer only ; cells large, elongate, pellucid. Calyptra pyriform, delicate, only at the base 2 cells thick, where are situated about 8 sterile, somewhat short, lageniform pistillidia, upper portion 1 cell thick. Capsule 4-5 times smaller than the perianth, highly exserted, cylindrical-oblong; valves linear-lanceolate or oval, of a purplish-brown colour like the spores and elaters. Pedicel mode- rately thick, cells large, 4-5 times longer than broad, prismatic- cylindrical, cortical cells 8-9, inner of equal size. Elaters 4 times shorter than the capsule, filiform, obtuse at both ends, bispiral. Spores as broad as the elaters, globose, smooth. Audroecia? Fruits May, June. Dimensions. — Stem ^ to 1 inch long, diameter "2 to '3 mm., with leaves r nmi. wide; leaves explanate, '5 mm. x '5 mm., •65 mm. x '65 mm., 'oS mm. x '05 mm., '3 mm. x 25 mm., segments *! mm., '4 mm. x -35 ram., seg. '15 mm., "4 mm. x •35 mm., seg. 15 mm., ^5 mm x ^45 mm., seg. '2 mm.; cells •025 mm. x "035 mm., ^03 mm. x '03 mm., ^03 mm. x 025 mm., •04 mm. X ^025 mm., -033 mm. ; stipules 'G mm. x "3 mm., "5 mm. X '38 mm., '55 mm. x "4 mm., '3 mm. x '175 mm., 5 mm. x 3 mm.; sub-bracts •G mm. x 'G mm., seg. "2 mm., "75 mm. x '7 mm., 1 • mm. X "7 mm., seg. "3 mm. ; sub-bracteole 75 mm. x 'G mm. ; l)racts N 194 HEPATICjE. 1-3 mm. X '9 mm., 1-mm. x '8 mm., seg. '3 mm. ; bracteole 1* mm. X -55 mm. ; perianth 4' mm. x 'T-'So mm. ; pedicel 10-15-mm. ; capsule -8 mm. x "5 mm., valve 7 mm. x '275 mm. ; spores •0125 mm. diam. ; elaters •2--25 mm. x '0125 mm. Hab. — On damp earth in alpine localities, rare. 15. Clova, R. K. GrcvUle. Ben Mac Dlmi, G. E. Hunt, Thomas Rogers, G. Sf abler. Ben Lawers, G. E. Hirnt. Head of Loch Avon, A. Croall. 16. Ben Nevis, R. K. GrevUle. Found on the Continent and in North America. Obs. — A very distinct species, and only found near the tops of the highest Scotch mountains ; from any of the CepJialoziiB dis- tinguished at once by the lateral branches ; from P. islandica (Nees), see notes under that species. Description of Plate LXXV. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 16 (Clova, K. K. Greville). 3. Ditto, postical view x 24 (ditto). 4, 5. Leaves x 64 (Grimsel, G-. E. Davies). 6. Portion of leaf x 290 (Clova, Greville). 7. Stipule X 64 (Grimsel, Davies). 8. Ditto x 85 (Clova, Greville). 9. 3rd sub-bract x 24 (Lapland, Angstrom). 10. 2nd sub-bract x 24 (ditto). 11. Sub-bracteole x 24 (ditto). 12, 13. Bracts x 24 (ditto). 14. Bracteole x 24 (ditto). 15. Perianth x ? (G. & B. Hep. Eur. n. 468, drawn by Dr. Gottsche). 2. Pleuroclada islandica [Nees). Jungermania islaiidica, Nees Nat. Eur. Leb. 11, p. 29 (1836). Cephalozia islandica, Lindb. in Journ. Linn. Soc. xiii. p. 192 (1873). Pleuroclada alhescens, var. islandica, Spruce, On Cephalozia, p. 79 (1882). Dioicous, csespitose, tufts depressed, eflagelliferous, small, pale green in colour, when dry with a bluish-white tinge. Stems simple or slightly branched, oval-terete, cortical cells about 20, similar to the inner, S cells in diameter ; radiculose, rootlets few, white ; branches lateral. Leaves imbricate, near the apex of stem approximate or somewhat distant below, patent to erecto-patent (40^^), orbicular, complicate-concave, bifid to the middle or below, sinus obtuse, segments broadly lanceolate or triangular, acute, PLEUROCLADA. 195 coniiivent ; texture somewhat thick, cells 4 -, 5-, and G-sided, smallish, walls thick, no trigones or thickened angles. Stipules large, as long as the leaves, ovate-lanceolate, entire. Flowers ? terminal on long branches. Bracts oblong or oblong-ovate, bifid to below the middle, segments lanceolate-acuminate, entire ; bract- eole free or adnate to one of the bracts, much smaller than the bracts, oblong-ovate or ovate, retuse, emarginate or entire ; sub- bracts similar to the innermost, only smaller. Perianth projecting much beyond the bracts, linear or narrowly fusiform, cylindrical below, above deeply 4-5-plicate, folds obtuse, composed of a single layer of cells, near the middle about 80 cells round, mouth con- stricted, irregularly and slightly dentate. Oalyptra delicate, composed of a single layer of cells. Fruit and male stems not yet known. Dimensions. — Stems |^ to 1 inch long, diam. '175 to "1 mm., with leaves '5 to "75 mm. wide ; leaves "3 mm. x '25 mm., segments •125 mm., -275 mm. x '25 mm., seg. "125 mm., -25 mm. x "225 mm., seg. '15 mm., '4 mm. x '3 mm., seg. "225 mm. ; cells "02 mm. x "025 mm., -0225 mm. x '0225 mm., '03 mm. x -025 mm., '02 mm. X "02 mm.; stipules -3 mm. x '125 mm., "35 mnrx"15 mm., '25 mm. x "l mm., "275 mm. x *15 mm. ; sub-bracts '75 mm. x '5 mm., sea:. '4 mm., "9 mm. x '45 mm , seg. *55 mm. ; bracts 1' mm. X '6 mm., seg. '6 mm. ; perianth 2' mm. x '5 mm., teeth at the mouth "04 mm. Hab. — Moist rocks or earth in alpine situations. Extremely rare. 15. Loch-na-gar, Aberdeenshire, /. Sim, April 1878. Found in Iceland, Northern Europe and Greenland. Obs. — Differs from P. albescens (Hook.) in the more slender habit of the plant, closer, more erect leaves, which are usually narrower, with deeper sinus, much less concave, not hemispherical, but simply complicate-concave, stipules narrower with no basal tooth or teeth, bracts longer and narrower, deeply bifid to below the middle, segments lanceolate-acuminate, not to a thirti, with seirments subacuminate-acute, bracteole retuse, emarginate or entire, with no basal teeth, perianth narrower, more plicate near the apex, composed of a single layer of cells, not several. Con- 196 HEFATICjE. sidered by Dr. Spruce as a variety of P. albescens (Hook.), but as it grows under similar conditions and at tbe same altitude, retaining the distinctive characters above mentioned, I look upon it as a distinct species. Description of Plate LXXVI. — -Fig. 1. Plants natural size. 2. Portion of stem, antical view x 31 (Loch-na-gar, J. Sim). 3. Ditto, postical view x 64 (ditto). 4-6. Leaves x 64 (ditto). 7, 8. Ditto X 85 (Lapland, y\ngstrom). 9. Portion of leaf x 290 (Loch-na-gar, J. Sim). 10, 11. Stipules x 85 (Lapland, Anijstrom). 12-15. Ditto x 64 (Loch-na-gar, J. Sim). 16-18. Sub-bracts x 31 (Lapland, Angstrom). 19, 20. Bracts x 31 (ditto). 21. Bracteole x 31 (ditto). 22. Perianth x 24 (ditto). 23. Cross-section of perianth, near the base x 24 (ditto). 24. Ditto, upper half of perianth x 24 (ditto). 25. Portion of mouth of perianth x 85 (ditto). Genus 17. HYGROBIELLA, Spruce. Jungermania, Hook. Brit. Jung. (1816). Gi/mnocolea, Dum. Kecueil (1835). Gephalozia, Lindb. Muse. Scand. (1879). Hygrohiella, Spruce, On Gephalozia (1882). Plants small, csespitose. Stems somewhat thick but fragile, transverse sections about 6 cells in diameter, cortical cells about 14, at the base rhizomatous and flagelliferous ; rootlets very few or absent ; above sparingly branched ; branches lateral and axillary. Subfloral innovations lateral or postical, often repeatedly innovant and floriferous. Lower leaves minute, distant, upper larger and closer, all transverse, complicato-bilobed, lobes equal or antical slightly smaller ; cells large and elongate. Stipules smaller than the leaves, somewhat similar or rarely entire. Inflorescence dioicous. $ terminal on stem or branch. Bracts few pairs, tristichous. Perianth large, subcarnose, fusiform, distinctly obtusely trigonous, mouth small, subentire. Calyptra narrow above. Capsule oblong, bistratose. Elaters bispiral. Spores HYailOBIEL LA. 197 minute. Bracts ^ terminal, few pairs, somewliat similar to the leaves, monandrous. Obs. — The genus lIi/(/rofjlel/a, Spruce, originally contained three species, Jun(/. la,vifoIla, Hook., Juii(/. niyriocarpa, Carr., and Jii)}(/. ncviccnsu, Carr. ; later, Dr. Spruce, not being satisfied with the position of Jiiu(/. mi/riocarpa in the genus, removed it to an unpublished genus proposed by the late Prof. Lindberg, which Dr. Kaalaas has been kind enough to prepare. J//ji(/. jieviceusis having been found in more perfect condition than when Drs. Carrington and Spruce had the opportunity of studying it, proves to be a Mar-sifpcl/a, where it has been placed. Hygrobiella laxifolia (IIoo/,-.), Spr//ce. Jtmyermania laxifolia, Hook. Brit. Jung. t. .")9 (ISKi). •Gijmnocolea laxifolia, Dum. Recueil, p. 17 (18o5). Cephalozia laxifolia, Lindb. Muse. Scand. (1H79). Hi/grohiella laxifolia, Spruce, On Cephalozia, p. 72 (1882). Dioicous, caespitose, flagelliferous, small, pale green to greenish- brown in colour. Stems suberect, almost round, cortical cells 14-20, large, inner 6-10 in diameter, smaller remarkably clear with very dark walls ; often much branched at the base, some branches short, flagelliferous, denudate or microphyllous, others assurgent, fastigiate-corymbose below with few and distant leaves, above (chiefly fertile) leaves closer. Kootlets very few or absent, proceeding from the flagella or stem. Leaves transversely inserted, erect, lower ones minute, ovate-subulate, sometimes entire, upper accrescent, subinibricate, oval and oval-lanceolatf-', complicate- oanaliculate and equitant, from \ to ]^ bifid, segments obtuse or acute, often unequal, sometimes only emarginate, very rarely with a third short segment near the middle. Texture lax ; cells rather large, pellucid, leptodernious, rectangular-hexagonal, two to three times as long as broad, walls thin but firm, no thickened angles or trigones. Stipules similar to the leaves, usually a little smaller, below often entire or emarginate. Flowers % terminal on stem or on long or short branches, often repeatedly innovant. Bracts 198 HEPATIC^. 2-3 pairs, embracing the base of the perianth, or subremote, similar to stem leaves, but often much larger, shortly bifid, unequal, repand, from base to nearly the middle composed of two layers of cells ; pistillidia few, small. Perianth large, lanceolate- fusiform when fruiting often elongate and almost linear, deeply trigonous, from base to near the apex usually composed of two layers of cells (at the keels and lower half often 3), near the middle a cross- section shows a double row of about 150 small cells, smaller perianths with 60-100 cells round, mouth contracted, narrow, with 10-15 short, hyaline, obtusely rounded marginal cells. Calyptra half the size of the perianth, lower half 2 cells tliick, afterwards the apex unequally bivalved, surrounded at the base by the barren pistillidia. Pedicel more than twice the length of the perianth. Capsule oblong, reddish-brown, composed of two layers of cells. Spores globose, smoothish, broader than the elaters, pale reddish-brown. Elaters of a paler colour than the spores, bispiral, short, obtuse, about 12 turns of the spiral. Androecia terminal, short, rarely occupying the whole branch ; perigoniai bracts usually 5-7 pairs, closely imbricate, ovate, bifid, concave, swollen; perigoniai bracteole oblong, entire or bifid; antheridia very large, single, oval, surrounded by leafy processes in the bracts. Fruits May. Dimensions.— Stems J to 1 inch long, -1 mm. diam., with leaves near the apex 1*25 mm. wide; leaves 1- mm. x "55 mm., segments "25 mm., 2 mm., "9 mm. x '45 mm., seg. '3 mm., '25 mm., I'l mm. x '5 mm., seg. '3 mm., '7 mm. x "4 mm., seg. "25 mm., "5 mm. x "3 mm., seg. "275 mm., 35 mm. x "15 mm.; cells "06 mm. x '02 mm., '08 mm. x '02 mm., '05 mm. x '02 mm., •05 mm. X "03 inm. ; stipules "8 mm. x "5 mm., seg. '3 mm., 1'2 mm. X "5 mm., seg. "3 mm., "25 mm., 1"1 mm. x "5 mm., seg. "3 mm., "35 mm. x '12 mm. ; bracts 1'65 mm. x "55 mm., 1'25 mm. x '65 mm., seg. "3 mm. ; bracteole 1*35 mm, x '6 mm., seg. '3 mm. ; perianth 2* mm. x '5 mm., 2*25 mm. x "65 mm. ; pistillidia •13 mm. X "04 mm. ; capsule "6 mm. x '35 mm. ; spores '02 mm. ; elaters •I mm. x "0125 mm. ; perigoniai bracts ^6 mm. x "45 mm.. HYGROBIELLA . 199 seg. "25 mm., 15 mm., G mm. x "5 mm., sei^. 2 mm., 15 mm., •65 mm. x "6 mm., seg-. "3 mm., "25 mm., '4 mm. x 'S mm., seg. •15 mm. ; perigonial bracteole "4 mm. x 25 mm., \'3 mm. x 25 mm.; antheridia 225 mm. x '15 mm. Hab. — Growing on wet rocks by the margin of streams or in moist situations in hilly or subalpine districts. Somewhat rare. 7. Llanberis, Carnarvon, JF. Wilson, IF. H. P. Cwm Idwal, Carnarvon, W. H. P. Glyder Vawr, Carnarvon, TF. 11. P. 10. Goathland Beck, Eskdale, Yorks, Dr. Spruce, M. B. Slater. High Force, Teesdale, Dr. Spruce, M. B. Slater, G. Baker. 12. Langdale, Westmorland, G. Stabler. Borrowdale, Cumberland, TF. H. P. 15. Head of Loch Avon, A. Croall. Clockmaben, near Banchory, Aberdeenshire, T. Sin/. Ben Venue, Dr. Carringtoii. Dalmally, E. George. 16. Glen Finnan, Lr. Carrington. Moidart, West Inverness, S. M. Macvicar. I. Kerry, Dr. Taijlor, Dr. Spruce, Dr. Carrington, and others. Found on the Continent (Norway, Sweden and North Germany), and in Greenland. Obs. — "In specimens from Brandon Mt. (Ireland) and from Teesdale, Eskdale, &c., the branches mostly originate from the leafless lower portion of the stem, and (as there are no radicles to indicate the underside of the stem) it is difficult to ascertain on what face of the stem they are fixed ; but wliere they do spring from a leafy part of it they are lateral, and axillary to the side- leaves. Scotch specimens are more leafy, and the branches are very distinctly lateral. Subfloral innovations are either lateral or postical, and are often repeatedly innovant and floriferous ; as in Hggrobiella myriocarpar Dr. Spruce. A very distinct species, at once separated from any other known British species by the almost equal tristichous leaves (the third postical-stipule) with elongate cells. Not long before he died Dr. Spruce wrote : " Ruminating instead of working, I have turned over some ancient guesses of mine on the classification of Hepatictie. What think you," for a change, "of Isotaclii.s laxifolia (Hook.), Hggrobiella (Spruce), pro parte ^ There are the tristi- chous, nearly transversely inserted leaves ; the elongated cells 200 EEPATICu^. (twice as long as broad) ; the flesh}^ perianth running to a narrow- point often quite closed, &c. The way some genera touch many others, more or less closely, is very remarkable." Description of Plate LXXVII. — Fig. 1. Plants natural size. 2. Fertile stem x 16 (Dalmally, E. George). 3-7. Leaves x 24 (ditto). 8 Portion of leaf x 290 (ditto). 9-11. Stipules x 24 (ditto). 12, 13. Bracts x 24 (ditto). 14. Bracteoie x 24 (ditto). 15. Perianth x 24 (ditto). 16. Cross-section of perianth, near the base x 24 (ditto). 17. Ditto, near the middle X 24 (ditto). 18. Portion of the moutli of perianth x 85 (ditto). 19-22. Perigonial bracts x 24 (Cwm Idwal, W.H.P.). 23. Perigonial bracteoie x 24 (ditto). 24. Antheridium x 85 (ditto). Genus 18. EREMONOTUS, Lindb. §• Kaalaas. JDiplophyllum, Carr. in Carr. et Pears. Hep. Brit. Exsicc. n. 96 (187'.)). Jungermania, Carr. in Trans. Bot. Soc. Edinb. p. 466 (188U). Hygrohiella, Spruce, On Cephalozia, p. 75 (1882). Plants small, with the habit of Cephalozia dicaricata, reddish- or brownish-green, densely and intricately csespitose. Stems very rigid, comparatively thick, fragile, at the base leafless, slightly radiculose, repeatedly ramose, most branches lateral or axillary, below leafless, above leafy, eradiculose, a few (chiefly the lower ones) postical, ieatless, flagelliferous and radiculose. Leaves sub- transverse, lower ones distant, minute, appressed, upper gradually laro-er and closer, ovato-quadrate, complicate, to the middle or more acutely bifid, with acute segments; cells minute, sub- quadrate, walls thickened. Stipules absent. Inflorescence dio- icous. Female flowers terminal on repeatedly innovant proliferous lateral (very rarely postical) branches. Bracts distichous, 2-3 pairs, innermost 3 to 4 times larger than the leaves, ovato-quadrate, from J to ^ bifid, lobes obtuse, rotundate. Perianth free, uni- stratose, relatively large, semiemersed, oblong, Irontally com- pressed, antical deeply unisulcate, postical bisulcate, with a smaller obtuse keel between the furrows, apex broadly rotundate, mouth small, minutely denticulate or setulose. Androecia situated at ERE MO NOT us. 2<)1 tlie end or middle of" branches, amentilbrm ; perigoiiial bracts monandrons, distichous, somewhat simihir to the leaves, only broader and more concave, bitid to about ^, segments obtuse. Obs. — This new genus diH'ers from Cejj/ialozia in its branches being Literal, not postical, and from Hygrohiella, which is its nearest ally, by its frontally compressed, not distinctly trigonous perianth, absence of stipules, bracts of the female flowers being distichous and the very different cell structure ol the leaves. B. Kaalaas in litt. Eremonotus myriocarpus (6V//v.), IJinlb. §■ Kaahia-s. Diplophiillam, myriocarpum, Carr. in Carr. et Pears. Hep. Brit. Exsicc. n. 911 (1870). Jungermania myriocarpa, Carr. in Trans. Bot. Soc. Edinb. p. ^(jO, t. IS, f. -i (18SU). Hyyrohiella mip'iocarpa, Spruce, On Cephalozia, p. 7o (1882). Dioicous, densely csespitose, small, of a reddish-brown colour. ■Stems :f to |- inch long, creeping at the base, rhizoinatous shoots entangled, flexuose, resembling pale brown horsehair, terete, ■ascending, rigid, interrupted, repeatedly innovant, branches springing from one or both sides of the old axis, either ban-en a,nd setaceous or fertile and with rapidly accrescent leaves, some branches (upper) lateral, others (lower) postical flagelliferous and rooting, rootlets sparse ; cells of stem 6 or 7 in diameter, cortical 20, subquadrate, very little larger than the inner ones, at first large and pellucid, afterwards, when dry, opaque. Leaves on the lower portion of stem and branches distichous, approxi- mate, erect, and appressed so close to the stem as to be readily overlooked, ovate, ovate-quadrate, complicate, carinate-concave, when explanate subcuneate, bifid to the middle, segments acute, sinus acute, texture thin, chitinous, polished ; cells minute, punc- tate, subquadrate, leptodermous, subpellucid. No stipules. Flowers $ terminal, often on repeatedly innovant-proliferous branches, innovations lateral, rarely postical, sometimes two oppo.site, elongate. Bracts for the size of the plant large, 2-3 pairs, distichous, closely conduplicate and equitant, keel at an 202 HEPATICjE. angle of 45°, innermost large, 4 times larger than the stem leaves, quadrate-oblong, subcordate at the base, bifid to about the ^th, segments obtusate, rarely subacute, outer ones gradually becoming smaller and changing into stem leaves. Perianth half hidden by the bracts, subcompressed, very obtusely trigonous (third angle postical), antical deeply unisulcate, mouth small, denticulate or setulose, setie 1-4 cells long. Capsule brown, oblong- globose, valves delicate, composed of two layers of cells. Elaters reddish- brown, bispiral. Spores pale brown. Androecia on separate stems growing with the $, simply fur- cate, not repeatedly innovantly dichotomous as in the $ ; perigonial bracts several, terminal or along the stem, slightly imbricate, swollen at the base, bifid to about the -J^rd, segments obtuse, antheridia solitary, oval. Fruits May, June. Dimensions. — Stems about \ inch long, '75 mm. diam. \ leaves '21 mm. x '175 mm., 12 mm. x 08 mm. ; cells "015 ram. X '01 mm., "0175 mm. x "012 mm. ; bracts '475 mm. x "45 mm. ; postical lobe "43 mm. x '25 mm. ; antical lobe '4 mm. x 2 mm. ; valves of capsule "4 mm. x "175 mm.; pedicel "1 mm, diam., at apex -12 mm. diam. ; elaters '1 mm. x "01 mm. ; spores '0175 mm. Hab. — Creeping among spongy peat, along with Jung. Jaxifolia in crevices of moist rocks. Rare. 7. Clogwyn-du-Arddu, Snowdon, Carnarvonshire, August 1881, W. H. P. Pass of Llanberis, Carnarvonshire, May 1883, W. H. P. 10. Brown Ghyll, Laugdale, Westmorland, May 11, 1881, George' Stabler 8r W. H. P. Nan Beild, Westmorland, May 11, 1882, George Stabler. 15. Discovered by Dr. Carri/igton, July 1876, in the crevices of moist rocks, in a stream from Ben Venue, running in the direction of the Trossachs Hotel, Scotland. 16. Moidart, West Inverness, S. M. Macvicar. Norway. Obs. — " In size this resembles CepJialozia dkaricaia, but its true alliance is with Jung, minuta, of which it might be accounted a microscopic form. Owing to the rigid chitinous texture of Jung, ntgriocarpa, the stems and even perianths decay very slowly. EIIEMONOTUS. 203 so that it is not uncommon to find five or six perianths, one above another, representing the growths of so many seasons. This give the plant a very characteristic appearance, when it is remembered the shoots are rarely more than from 2 to 4 lines in length. The same proliferous habit is not uncommon in Ji/m/. uiiiii/fa, and it shares with it other peculiarities, viz., the perianths were invari- ably abortive and the pistillidia barren and undeveloped. The young perianth looks like that of a FndJania ; it is a point where the crests meet. " From J//////, rujida, Lindb., which it also resembles in habit, it may be known by its much smaller size, fasciculate ramification, rapidly accrescent leaves, the lobes of which are equal, and the different form of the perianth ; whereas in Jiiikj. rifjida the shoots are terete from the closely imbricated uniform leaves, the lobes of which are unequal." Dr. Carrington. " This curious little plant differs from /. la.rifo/ia in the total absence of stipules and the dense reticulation ; but in most other respects it is a miniature counterpart of that species ; and they agree so perfectly in habit and in all essential characters that I can hardly doubt they should stand in the same genus. I have cut transverse sections of several perianths of /. uti/riocar/jay and have found them uniformly trigonous, with the third angle at the back, as in J. laxifolm and in all Cephalozice. The furrow along the middle of the upper face of the perianth, with a slight ridge or keel on each side of it, quite corresponds to what is seen in immature, or unfertilised perianths of several of the C('p/ialozi(e." Dr. Spruce. Description of Plate LXXVIII. — Fig. 1. Plants natural size. 2. Plant X 16. 3-4. Portion of barren stems x 64. b,ij. Stem- leaves X 64 (Ben Venue). 7-11. Ditto x 85 (Clogwyn-du- Arddu). 12. Portion of stem and leaf x 200. 13. Apex of leaf X 200. 14. Portion of leaf x 290 (Ben Venue). 15, 16. Bracts x 85 (Clogwyn-du-Arddu). 17. Perianth with bracts x 64. 18. Perianth x 64 (Ben Venue). 19. Spores X 290 (Brown Ghyll). 20. Perigonial bract x 85. 21. An- theridium x 85 (Clogwyn-du-Arddu). 204 HEPATIC^. Genus 19. ADELANTHUS, Mitten. Jungermania, Hook, in Eng. Bot. t. 2567 (1818). Radula, Dum. Syll. Jung. p. 43 (1831). Plagiochila, G.L.N. Syn. Hep. p. 24 (1844). Adekmthtos, Mitt, in Journ. Linn. Soc. p. 2G4 (1864). Sphagncecetis, Hartm. Skand. Fl. ed. 10, 2, p. 144 (1871). Odontoschisma, Lindb. MSS. in Hartm. Skand. Fl. ed. 10, 2, p. 144 (1871). Plants caespitose, pale or dark green. Stems firm, caudex creeping, simple or slightly branched, radiculose, suberect, apex often decurved or subcircinnate, producing from near the base postical tlagella which are minutely leaved and radiculose. Leaves succubous, moderately large, alternate, obliquely rotund or ovate, decurvo-secund, antical margin incurved, subentire, postical sub- plane, very often spinulose-dentate ; cell rather large to minute, guttulate, pachydermous, trigones distinct, marginal cells minute, opaque. Stipules none or very rare and minute. Inflorescence dioicous, cladogenous, postical. Bracts $ tristichous, the inner- most slightly smaller than the leaves, apex bifid or variously incised. Pistillidia 12-15. Perianth pyriform or fusiform, turgid, 3-5 angled, mouth denticulate. Calyptra a little smaller, carnose, below the middle 3-5-stratose, sterile pistillidia surround- ing it. Capsule oval, 4-valved, bistratose. Androecia amentifbrm ; antheridia solitary, very rarely two. Adelanthus decipiens [Hook.), Mitt. Jungermania decipiens, Hook, in Eng. Bot. t. 2567 (1813) ; Brit. Jung. t. 50 (1816). Radida decijnens, Dum. Syil. Jung. p. 43 (1831). Plagiochila decipiens, Dum. E.ecueil, p. 15 (1835). Gymnanthe decijnens, Mitt. Journ. Linn. Soc. 11, p. 166 (1863). Adelanthus decipiens. Mitt, in Journ. Linn. Soc. p. 264 (1864). Odontoschisma decipiens, Lindb. MSS. in Hartm. Skand. Fl. 11, p. 144; Lindb. Soc. Faun, et Fl. Fenn. 13, pp. 357-363 (1874). Dioicous, densely caespitose, flagelliferous, medium size, dark olive green when growing, below and when dry, dark brown in colour. Caudex prostrate, intricately branched below, producing ADELANTHUS. 205 erect or ascending fastigate stems, stems blackish, firm, simple, rarely furcate or subramose, apices decurved ; cortical cells 30-36, lurid, opaque, 10-12 cells in diameter, inner slightly smaller, sub- pellucid ; flagella postical, few, only on the lower part of stem, with very minute, tristichous, distant, subquadrate-rotund, pellucid, rudimentary leaves, bearing fasciculate rootlets, tender, fragile, towards the ends of the fiagella destitute of leaves, but producing numerous rootlets. Leaves incubous, lower smaller, subimbricate, appressed, upper larger, more crowded, decurvo-secund, rarely distichous-patulous, obliquely inserted, antical base long and gradually decurrent, subrhomboid or obliquely orbiculate, entire, apex rotundate retuse, apiculate or spinose, bi-tri-lobate, antical margin incurved, postical subplane or slightly concave. Texture thick, epidermis a little rough ; cells medium to rather large in size, hexagonal, walls firm, angles thickened or trigones distinct, marginal cells distinct, thicker, subquadrate. Stipule on stem none or rudimentary. Inflorescence ? postical, on very short branches produced from near base of stem, rarely from middle. Bracts about 9, in threes, tristichous, outermost minute, scale- like, innermost a little shorter than the leaves, apex more or less deeply bifid, segments patulous, free or subconnate at the base, subentire ; $ branches excavate, cavity pyriform. Perianth twice as long as the stem leaves, projecting far beyond the bracts, narrowly obovate, pyriform, turgid, above obtusely tri- (rarely 4-, 5-) gonous, mouth at first connivent, afterwards subtrifid, sub- denticulate or ciliolate. Calyptra a little smaller than the perianth, pyriform, firm texture, composed of 3 layers of cells, apex afterwards irregularly trifid, at base surrounded by few (about 12?) sterile pistillidia. Andrnecia postical, whitish, slender, julaceous or roundish, concinnate, produced from the whole length of stem, but chiefly from near the base ; perigonial bracts 12 pairs or less, closely imbricate, subcymbiform or cuccu- late, very concave and subsaccate, but scarcely complicate, slightly unequal, very oiten entire, rarely bidentulate, areolation very pellucid; antheridia solitary, shortly stipitate ; perigonial bracteoles everywhere present 2-4 times smaller than the bracts, subrotund ■206 HEPATIC^. or oblong, entire or angular, rarely connate at the base to the nearest upper leaf. Dimensions. — Stems 1 to Ij inch long, '2 mm. in diam., with leaves 2" mm., 2'5 mm. wide; leaves 2' mm. x 2* mm., 2" mm. X 1*75 mm., 1-75 mm. x 1*5 mm., r4 ram. x 1*2 mm.; cells •05 mm. X "05 mm., '045 mm. x "03 mm., "035 mm. x 035 mm. ; bract 1"5 mm. long; perianth 3'4 mm. x 1"5 mm., 2" mm. X '75 mm. ; capsule '75 mm. x '5 mm. ; pedicel "2 mm ; perigonial bracts '5 mm. x "4 mm. ( x "75 mm. explanate) ; perigonial bracteole "25 mm. x "125 mm., '13 mm. x '13 mm. Hab. — Growing on rocks or banks in woods or exposed places, ^ery rare, extremely so in fruit. 7. Tyn-y-Grroes, Merioneth., E. M. Holmes, Dr. Carrington 8f W. H. P., G. A. Holt. 12. Borrowdale, Cumberland, Dr. Carrington ■Sf W. H P. 16. Moidart, West Inverness, S. M. Macvicar, 1898, W. H P. 1899. I. Discovered by Miss Hutcldns near Bantry. Kenmare ^Sound, Glengariff, Dr. Carrington. Rocks about the Upper Lake, Prof. 8. 0. Lindberg. Glenad, Leitrim, Dr. D. Moore. Found on the Continent (Norway), in Cuba, and South America. Obs. — A distinct species and the only British example of the genus. When in fruit the postical $ and $ would readily identify it, but $ has only been sparingly found b}'- Dr. Carrington in Ireland, and no perianths have yet been discovered on British specimens. When barren, distinguished from Plagiocliila punctata Tayl., which it resembles somewhat in size and habit, by its dark lurid green colour, leaves usually unispinose, antical margin incurved, and the large hexagonal cells. The description is taken from Dr. Spruce's valuable notes in "Journ. of Bot." (1876). Description of Plate LXXIX. — Fig. 1. Plants natural size. 2. Stem X 11 (Killarney, Dr. Carrington). 3. Fertile stem x 18 (G. & P. Hep. Eur. n. 474, from Cuba, drawn by Dr. Gottsche). 4-6. Leaves x 16 (Killarney, Dr. Carrington). Leal' x 16 {Canelos, South America, Dr. Spruce). 8. Portion of leaf x 290, SCAPANIA. 20^ (Killarney, Dr. Carrington). 9, 10. l^racts x IS (G. & R. Hep. Eur. n. 474). 11. Sub-bract x 18 (ditto). 13. AndroBcia (Killarne}^, Dr. Carrington, G. & R. Hep. Eur. u. 474, drawn by Dr. Gottsclie). Subtvibe V. SCAPANIOIDE^. Genus 20. SCAPANIA, D/nii. Jungerma7iia, Mich. Nov. pi. gen. p. (! (17:^9); L. Sp. i)l. ed. 1, 2, p. 11. S2 (1758). Martinellia, sect, a Gr. & B. Nat. Ait. Brit. PI. 1. p. G91 (1,S21). Eadula, Dum. Comm. p. 112 (1823). Badula, Dum. sect. 2, Scajmnia, Dum. Syll. Jung. Eur. p. o8 (1831). Scapania, Dum. Recueil, 1, p. 14 (1835). Plants usually robust, some large, otbers rather small, olive- green, reddish-brown or purple, growing in matted tufts or in large masses in streams. Stems firm ; primary shoots creeping, perennial, naked, rhizomatous, entangled ; secondary shoots foliose, ascending, decurved at the apex, simple or dichotomously branched; branches all exactly axillary to the leaves. Leaves disticho- declinate, alternate, conduplicate, unequally bilobed ; postical lobe larefer, convex ; antical lobe incumbent ; maro-ins entire, dentate or ciliate. Stipules absent. Inflorescence dioicous, rarely paroicous, terminal, liracts $ 2, free, resembling the leaves but more equally lobed. Perianth much longer than the bracts, frontally compressed, plane, not winged, mouth wide, truncate, bilabiate, entire or dentate, decurved. Calyptra iree, included. Capsule ovate, of thi(di texture, divided to the base into 4 equal valves. Elaters attached to the centre of the valves, long, bispiral. Androecia terminal or interrupted. Perigonial bracts smaller, ventricose at the base, the lobes shorter and nearly equal. Antheridia (3-12) axillary, oval, seated on a slender stipe. 208 HEPATIC^. 1. Scapania compacta {Hot It), Dum. Jumjermania compacta, Roth, Tent. Fl. Germ. 3, p. 375 (1800). Jungermania resupinata, Web. et Mohr Crypt. Germ. p. 427 (1807) ; Hook. Brit. Jung. n. 23 (1810). Radula resupinata, Dum. Oomm. Bot. p. 112 (1823). Scapania compacta, Dum. Recueil, p. 14 (1835). Plagiochila compacta, M. et N. in Nees Nat. Eur. Leb. Ill, p. 510 (1838). Paroicous, loosely ceespitose, small, yellowish-brown. Steins firm, flexuose, simple or innovantly branched, suberect or prostrate, radiculose up to apex, rootlets dirty-white. Leaves subtrans- versely inserted, approximate, accrescent, shortly and subequally bilobed, antical lobe slightly smaller than the postical, lobes rotund or semi-oval, entire or rarely denticulate, texture firm, epidermis smooth, cells smallish to medium, subguttulate, walls somewhat thick, trigones large. Bracts simihir to the leaves but larger, often denticulate. Perianth large, projecting much beyond the bracts, broadly obovate or turbinate, mouth wide, entire or irregularly denticulate. Antheridia (about 4 to 6 in each bract) oval, situated immediately below the perianth at the base of the bracts, which are slightly swollen. Fruits April, May, June. Dimensions. — Stems \ inch long, -3 mm. to '4 mm. diam., with leaves 2' mm. wide; leaves, upper, antical lobe TS mm. x 1-25 mm., postical 1"75 mm. x 1-25 mm., lower, antical lobe 1- mm. X 75 mm., postical 1-25 mm. x 75 mm. ; cells -03 mm. ; bracts, antical lobe 1"5 mm. x 1' mm., postical 175 mm. x P mm. ; perianth 3* mm. x 2" mm. wide at the mouth, 2-25 mm. x 1-5 mm., antheridium '2 mm. x -175 mm. JJab. — Growing on clayey and sandy banks, and on mud- covered walls. Generally distributed. 1. Common, West Cornwall, W. Curnow. 2. Isle of Wight, G. E. Bavies. 3, 4. About Edgefield and Hempstead Hill, Norfolk, Rev. R. B. Francis. Norfolk and Sufi'oik, Br. W. J. Hooker. 7. Penmaenpool, Merionethshire, E. M. Holmes, Dolgelly, Merionethshire, W. H P. Festiniog, Carnarvonshire, Br. Car- ^OAPAJVJA. 209 rim/fon Sf^ IF. JI. P. 10, 12. On rocks, Witherslack ; in a wood near Staveley Church ; Rayrigg Wood, Windermere ; Oxenden, Great Langdale, Westmorland, G. Stabler. 13. Frequent, Kirk- cudbrightshire and Dumfriesshire, J. McAndreio. 14, 15. Ben Lawers, C. J. Wild. Braemar, JR. K'nhoii. Gateside, Strachan, /. Sim. IG. Frequent on rocks and boulders, Moidart, West Inverness, 8. M. Macvicar. I. Brandon, Co. Kerry, Br. B. Moore. Found on the Continent and Madeira. Obs. — At once distinguished from any of the other known British Scapanice by its paroicous inflorescence, the antheridia being found in the bracts immediately below the perianth, whicli is very rarely the case with any of this genus. Apart from its inflorescence it is a very distinct species, its compact habit, almost equal lobes of leaves and bracts, its habitat, mud-covered walls and clayey banks in the plains, often near the coast, readily enable the student to identify it. Description of Plate LXXX. — Fig. 1. Plants natural size. !2. Portion of fertile stem, antical view x 16 (168 G. & R.). 3. Upper leaf, explanate x 16 (ditto). 4, 5. Leaves x 24 (Isle of Wight, G. E. Davies). 6. Portion of leaf x 21)0 (168 G. & K.). 7. Bract, explanate x 24 (Isle of Wight, G. E. Davies). 8, 9. Bracts x 24 (ditto). 10. Perianth x 24 (Cader Idris, W. H. P.). 11. Portion of mouth of perianth x 85 (ditto). 12. Antheridium x 85 (Isle of Wight, G. E. Davies). 2. Scapania Bartlingii [Hampe), Xce-s. .Tumjermania Bartlingii, Hampe in Nees Nat. Eur. Leb. 11, p. 42.") (188(5). Scapania Bartlingii, Nees in G.L.N. Syn. Hep. p. 64 (1844). Jungerma7via cuspid idiger a, Nees Nat. Eur. Leb. 1, p. 180 (1883). Plagiochila Bartlingii, M. at N. in Nees Nat. Eur. Leb. Ill, p. 520 (1888). .fungermania rupestrit, Schleich. Cat. Exs. (1S21) ^ Scapania C'arestia', De Not. in Mem. Accad. 'I'orin. (IS.'tS) ^ Dioicous, loosely caespitose, small, of a pale green colour. Stems pale brown, somewliat tender, but proportionately thick o 210 HEP AT ICE. for the size of plant, frontally compressed, 20 cells x 15, inner cells distinct, hyaline, two outer rows of cells brown ; short, simple, or with 2-3 innovant branches produced from the axil of old bracts, suberect or creeping ; radiculose, rootlets copious, of a dirty-white colour. Leaves transversely inserted or a little ascending, few, accrescent, bifarious, alternate, lower ones minute, scale-like, middle contiguous, upper closely imbricate, vaginate, unequally bilobed from about a fourth to a third, margin entire, sometimes erose, antical lobe about a third smaller, oblong, oval to subquadrate, apiculate, acute or obtuse, undulate, recurved or incurved, postical lobe oval, obtuse or rotundate ; texture some- what lax, epidermis smooth, cells near middle at the base elongate-quadrate, medium size, near the apex rather minute, subquadrate, cell-walls somewhat thin, angles a little thickened, no trigones. Bracts large, similar to upper leaves, lobes more equal. Perianth projecting about a third to half beyond the bracts, terminal, obovate, compressed, mouth truncate, wide, entire, a little uneven. Male stems with perigonial bracts terminal or at the middle, ventricose, antical lobe more erect, antheridia oval. Gemmiparous, gemmje whitish-green to brown, biseptate, terminal, sometimes forming brownish-black spherical masses. Fruits April, May. Dimensions. — Stems about |- in. long, "3 mm. to "4 mm. diam., with leaves 25 mm. to 275 mm. wide ; leaves, antical lobe 1*2 mm. X -65 mm., postical 1"5 mm. x 75 mm., antical 1*1 mm. x '5 mm., postical 1*5 mm. x 75 mm., antical 1*3 mm. x "75 mm., postical 1*5 mm. X '75 mm. ; upper leaves, antical lobe 175 mm. X 1*25 mm., postical 2* mm. x 1' mm. ; cells, basal '06 mm. X '025 mm., 04 mm. x '02 mm., '04 mm. x '025 mm., near apex of leaf '025 mm. x 015 mm., '02 mm. x '015 mm., "015 mm. x 'OlS mm.; bracts, antical lobe 175 mm. x T mm., postical 2* mm. X r mm.; perianth 2*25 mm. x ri5 mm. at mouth, gemmge "02 mm. x '01 mm. Hab. — Growing on damp, shady rocks by streams ; very rare. 10. "Near the " Strid," Bolton Woods, Yorkshire, Dr. so A PAN! A. 211 Carrli/f//o)i. 1 1 . Egleston, Tees-side, /. G. Baker. Teesdale, r. Mncld. Eare on the Continent. Obs. — Distinguished from small forms of S. feqnilohd by its smooth epidermis, from S. ciirfa by its more equal lobes and distinctly accrescent liabit. Description of Plate LXXXI. — Fig. 1. Stems natural size. 2. Fertile stem x IG (Bolton Woods, Dr. Carrington). :3. Por- tion of sterile stem X IG (ditto). 4. Cross-section of stem x 24 (ditto). 5-7. Leaves flattened out x IG (ditto). 8. Upper leaf X 16 (ditto). 9. Ditto flattened out (ditto). 10. Portion of leaf, middle basilar cells x 290 (ditto). 11. Ditto, near marginal apex (ditto). 12. Bract, flattened out x 16 (ditto). 13. Perianth X 16 (ditto). 14. Portion of the mouth of perianth x 85 (ditto). 15, 16. Perigonial bracts x 24 (Switzerland, Jack). 17, 18. Gemmae x 290 (Bolton Woods, Dr. Carrington). 3. Scapania subalpina {Nccs)^ Duni. Jungermcmia subalpina, Nees in Syn. Hep. Eur. p. 55 (1829). Scapania suhalpina, Dum. Recueil, p. 14 (1885). Dioicous, csespitose, small, pale green to brown in colour. Stems reddish, simple or slightly branched, innovant branches (2, 3 or 4) produced from the postical base of old perianth, erect, rootlets few, hyaline. Leaves patent (50°), imbricate, divided almost to the middle into two subequal lobes, appressed, closel}' incumbent, rotundate or slightly apiculate, antical lobe a little smaller than the postical, subrotund to subquadrate, entire or very slightly denticulate, postical lobe subrotund or broadly rotund, denticulate or rarel}' entire ; texture delicate, lax, epidermis smooth, cells smallish, 4-, 5- and G-sided, walls thin, trigones none. Bracts similar to leaves. Perianth oblong, truncate, mouth entire. Perigonial stems with the c? in the middle or terminal, bracts little different from the leaves. Fruits May, June. H.\B. — On sandy soil or amongst mosses in subalj^ine or alpine districts. Kare. ■212 HEPATICjE. 7. Near Bettws-y-Coed, Carnarvonshire, iT/. ^. /iS7«/pr. 15. Ben Lawers, C. J. Wild, August 1878. Ben Mac Dhui, John WMfehead, July 1876. Ben Nevis, W. JFest, 1880. Glen Tilt, W. JFesf, 1880. Moray, J. Croall, April 1848. Glencoe, .S'. M. Macvicar, 1899. 16. Eesipol, Sunart, S. M. Macvicar 8f W. H. P., 1899. I. Lug- naquilla Mountain, Co. Wicklow, Br. B. Moore. Nephinbeg, Co. Mayo, Br. B. Moore. Found on the Continent. Dimensions. — Stems 1 to \\ in. long, -2 mm. to Smm. diam., with leaves To mm. to 2' mm. wide ; leaves, antical lobe 1"25 mm. X "75 mm., posticallobel*5mm. x 1*25 mm., 1*25 mm. x I'mm. ; cells "025 mm. ; perianth 225 mm. x 1' mm. Obs. — This peculiarly alpine and subalpine species may be distinguished from S. undulata by its neat habit, equal-sized, closely imbricate, appressed, almost entire leaves and the mouth of the perianth being entire. Nearly all the specimens I have examined of this species have the lower portion of the stems thickly matted with earth. Description of Plate LXXXII. — Fio*. 1. Plants natural size. o 2. Portion of stem, antical view x 24 (Glen Tilt, West). 3. Portion of stem, antical view x 24 (Finland, S.O.L.). 4. 5. Leaves x 24 (Sweden, S.O.L.). 6. Postical lobe of leaf X 24 (ditto). 7. Portion of leaf x 290 (ditto). 8. Perianth X 24 (Glen Tilt, West). 9. Perigonial bract x 24 (Sweden, S.O.L.). 4. Scapania aequiloba {Schwae(/.), Bum. Jungermania cequiloba, Schwaeg. Prod. p. 24 (1814). Ju7igermama montana, Mart. Fl. Ci-ypt. Erlang. p. 155, t. 4, f. 31 (1817). Radula cequiloba, Dum. Syll. Jung. Eur. p. 39 (1831). Scajjania cequiloba, Dum. Recueil, p. 14 (1835). •Jungermania tyrolensis, Nees Hep. Eur. 11, p. 440 (1836). Plagiochila tyrolensis, M. et K. Nees Hep. Eur. iv. p. xxxvii. n. 17 (1838). Scapania tyrolensis, G.L.N. Syn. Hep. p. 69 (1844). Martinellia cequiloba, Linclb. Hep. in Hib. p. 521 (1874). Exsicc. G. & R. Hep. Eur. n. 89, 404, 408. Dioicous ; calcicolous, growing in densely matted tufts, medium size, of a dark olive green to dark brown colour. Stems simple SCAPANIA. 213 or slightly branched, black, suberect or erect ; radiculose, rootlets moderately plentiful, hyaline. Leaves horizontally inserted, slightly ascending, closely imbricate, regular, of almost equal size, somewhat pectinate, bilobed barely to the middle, lobes subequal or the antical about a third smaller, convex, ovate or trapezoid, acute or sometimes apiculate, crossing the stem, entire or slightly dentate, postical lobe broadly oval or obovate, acute or sometimes apiculate, subentire, textare firm, epidermis very finely verruculose, cells small, roundish-subquadrate, walls thick, angles thickened, trigones wanting. Bracts similar to upper leaves or slightly larger, lobes subequal, subentire. Perianth projecting considerably beyond the bracts, oblong- obovate, compressed, mouth wide, laciniate, irregularly spinose-dentate. Capsule small, oval ; spores light brown ; elaters short, bispiral, dark brown. Male stems more slender than the others, perigonial bracts terminal or at the middle, little different from ordinary, smaller, ventricose ; antheridia 2, oval, with long bearers, along with several leafy paraphyses. Sometimes gemmiparous, gemmse greenish- white in colour, terminal on the upper leaves of sterile stem. Fruits April, May, June. Dimensions. — Stems 1 to 2 inches long, '3 mm. to "4 mm. diam., with leaves from 2'5 mm. to 35 mm. wide; leaves, antical lobe 1*5 mm. x I'mm., postical 1*5 mm. x 1' mm., antical 1'25 mm. X -75 mm., postical 1"75 mm. x T mm., antical 1"5 mm. x -70 mm., postical 1*75 mm. x 1*25 mm. ; cells 'Old ram., "02 mm., 0275 mm. X '015 mm.; bracts, anticallobe 175 mm. x 1* mm., postical 2- mm. X 1* mm.; perianth 2*75 mm. x I'l mm., lacinias at mouth of perianth '275 mm., cilia ditto '125 mm. ; spores •0125 mm. ; perigonial bracts, antical lobe "75 mm. x "G. mm., postical 1"25 mm. x '7b mm. ; antheridia -15 mm. x •125 mm. Hab.— Growing usually on limestone rocks. Kare. 8. Lover's Leap, near Buxton, C. J. Wild. Miller's Dale, Derbyshire, W.ILP. 10. Winch Bridge, Teesdale, Dr. SprNcc, July 1843. Wharfdale, Dr. Spruce, Dec. 1841. Linton, G. A. Holt. 1(5. On sandy banks, near the sea, Moidart, West Liverness, S. M. Macvicar. Found on the Continent. 214 HEPATIC.^. Obs. — Distinguished from all other Scupanice (except S. aspera, M. & B., which see) by the leaves and perianth being minutely verruculose, in addition to other characters. The best method to observe this feature is to detach a leaf and bend it between the slide and cover glass in water ; with a \ inch, numerous minute warts are observed on the surface of each cell. Its neat habit, regular leaves, not deeply divided, with the dense cells, separate it from allied species. From an exhaustive study of every specimen I have had the opportunity of examining, I am only able to give the few stations above recorded for this species, all the others I have to refer to Scapania aspera, M. et Bern. ; the type seems to be better repre- sented in the male stems, which are more regular and neater in habit. Description oy Plate LXXXIII. — Fig. 1. Plant natural size. 2. Portion of stem, antical view x IG (Moidart, Macvicar). 3. Leaves x 16 (ditto). 4, 5. Leaves x 24 (Winch Bridge, K. Spruce). G. Portion of leaf x 290 (ditto). 7. Bract, explanate x 16 (Austria, Jack). 8. Portion of stem with perianth x 16 (ditto). 9. Portion of mouth of perianth x 85 (ditto). 10. Perigonial bracts x 16 (404 Gr. & R.). 11. An- theridium x 85 (ditto). 5. Scapania aspera, Midler &,' Bertiet. Scapania aspera, M. & B., Henri Bernet, Catalogue Hep. du Sud-Ouest de la Suisse et de la Haute-Savoie (1888). Exsicc. G, & R. Hep. Eur. n. 02 (as Scapania nemorosa), n. ooi (as Scapania nemorosa), n. 602 (as Scajxmia mquiloha forma dentata), Massal. Hep. It.Ven. n. 62 (as Scajxmia cerjuiloba, var. dentata). Dioicous, loosely depresso-ctespitose, of a reddish- or olive-brown colour. Stems tallish, simple or slightly branched, firm, blackish, recurved at the apex, denudate at the base ; radiculose, rootlets few, whitish. Leaves transversely inserted, somewhat smaller and distant below, contiguous or imbricate above, subsecund, unequally bilobed to about the middle, margin ciliate-dentate ; postical lobe more distinctly ciliate, about 25 cilia around the margin ; antical lobe with rather more distant teeth or cilia ; about half the size of SCAPANIA. 215 the postical, convex, oval-triangular, rotundate, appressed to the stem, or upper margin slightly reflexed ; postical lobe oval-oblong, rotundate, reflexed ; texture somewhat firm, epidermis verruculose, several minute papillae on each cell; cells small to rather minute, subquadrate, walls thick, angles thickened, no trigones. Bracts rather larger than the upper leaves, lobes more equal, antical lobe rotundate. Perianth projecting half beyond the bracts, com- pressed, mouth wide, truncate, spinose-ciliate. Male stems more slender, perigonial bracts enclosing leafy paraphyses with the antheridia. Sometimes gemmiparous. Fruits April, May, June. Dimensions. — Stems 1 to 2 inches long, "5 mm. diam., with leaves 5* mm. wide; branches '25 mm. diam., with leaves 275 mm. wide; leaves, antical lobe To mm. x 1* mm. ; postical lobe 2*5 mm. x 1'75 mm., antical TS mm. x T mm., postical 2-5 mm. X 1-5 mm., antical 175 mm. x r25 mm., postical 2-25 mm. x 1-25 mm. ; cells '02 mm., -0175 mm. ; cilia of postical lobe -05 mm.; bracts, antical lobe 2' mm. x 1*5 mm., postical 2*5 mm. x 1-5 mm. ; perianth 3"5 mm. x 2* mm. wide at the mouth, lacinite of the mouth "275 mm., cilia of the same 1 mm. Hab. — Growing on limestone rocks. Somewhat rare. 2. Hill above Studland, Dorset, E. M. Holmes, 26th April 1884. 7. Near Aber, Carnarvonshire, G, E. Hunt, May 1868 (Herb. Spruce). Tower Hill, Abergele, Denbighshire, IF. H. P., August 1892. Llandulas, Denbighshire, IF. H. P., 1899. 9. Locally abundant on limestone rocks in Lane. Silverdale, ./. A. Wheldon, July 1898. Over Kellet, A. Wilson. 10. Bolton Woods, Yorkshire, Dr. Car- rington, July 1877. 12. Yewbarrow, Westmorland, Geor//e Stabler, Nov. 1869. 16. Moidart, West Inverness, S. M. MacDicar, 1899. Eoshven, West Inverness, 8. M. Macoicar, 1899. I. Hill of Howth ; Co. Cavan ; Killarney ; cide B. McArdle. Found on the Continent (Sweden, Norway, Switzerland, Germany, Austria, Italy). Obs. — In habit and general appearance similar to Scapania nemorosa (L.), but in this latter species the leaves, both antical and postical lobes, are usually rounder and margin ciliate or more 216 HEPATIC^. finely dentate, but the most important distinctions of Scapania aspera are the closer texture of the leaves and the epidermis being verruculose ; this latter character separates it from all the other British Scapania, with the exception of Scajjania csquiloba (Schwaeg,). Sccqjania aqiiiloba (Schwaeg.), of which it was considered a variety by such good authorities as Gottsche and Lindberg, is undoubtedly its nearest ally, although strikingly different in habit and smaller in size, and generally of a darker olive-green colour ; the leaves are regularly inserted and almost equal in size the whole length of the stem, and although the lobes are by no means equal, as the name would imply, they are more so than in Scajjaiiia ai<2jera, are more quadrate, with the apex more acute. This character is also observable in the bracts ; in Scapania aspera the perianth is usually wider at the mouth, but the margin is exactly the same. The male plant of Scapania aspera approaches nearer to 5. (squiloba than the barren or fertile one. The founders of the species say, "It has nothing in common with Scapania ceqmloha except in the tuberculose cuticle." This statement will probably be questioned by students who have the opportunity of examining a large series of specimens. Scapania crassiretis, Bryhn (found in Norway), is another allied species, which has not yet been met with in our country. I find the species with which it is most generally mistaken for, even by some expert students, is Scapania resnpiinafa (L.), with which it agrees somewhat in the shape and margin of leaves, but the latter species is usually of a pale olive -brown colour, narrower and more equal leaves, the antical lobe is also more or less reflexed on some portion of the stem, the epidermis is also quite smooth. Description of Plate LXXXIV. — Fig. 1. Plants natural size. 2. Portion of young branch x IG (Yewbarrow, Gr. Stabler). 3. Leaf x 11 (Gf. & E. n. 602). 4. Ditto, explanate (ditto). 5, 6. Leaves x 1 1 (Sweden, Kindberg). 7. Leaf x 11 (G. & K n. 602). 8. Margin of leaf, postical lobe x 85 (ditto). 9. Por- tion of leaf X 290 (ditto). 10. Bract x 11 (Yewbarrow, Gf. SCAPANIA. 217 Stabler). 11. Ditto, explanate (ditto). 12. Perianth x 11 (Sweden, Kindberg). 13. Portion of mouth of perianth x 85 (Yewbarrow, G. Stabler). 6. Scapania resupinata (Z.), Dnm. Lichenastrum auricidatum, jnmudis rotmuUs, afispv/tn, Dill. Hist. Muse. p. 491, t. 71, f. 11) (1741). Jtmgermania resnjmiata, Linn. Sp. pi. 1599, fide Dillenii (1753); Huds. Fl. Ang. p. 512 (1798); With. Arr. Br. pi. 3, ed. Ill, p. 875 (179G); Eng. Bot. t. 2437. Radula dentata, Duin. Syll. Jung. p. 40 (1831). Scapania dentata, Dum. Recueil, p. 14 (183r»). Scapania resujmiata^ Dum. Recueil, p. 14 (1N35). Martinellia gracilis, Lindb. Manip. Muse. Secund. p. 305 (1874). Dioicous, growing in dense tufts of a yellowish-brown or olive colour, pale below, closely entangled with long hyaline root- lets, which are produced from the underside of the stem. Stem ligneous, simple or sparingly branched. Leaves crowded at the apex, equal in size, from about J to the J bifid, closely condupli- cate, antical lobe crossing the stem, appressed, somewhat erecto- patent, convex, yet upper margin frequently reflexed, obliquely reniform-rotund, apex rotundate-obtuse, with distant large teeth, teeth broad at the base, the apical ones generally the largest, postical lobe about twice as large, very convex, upper margin remarkably reflexed and decurrent, obliquely oval-obovate, rotundato-obtuse, dentato-serrate, apical teeth scarcely larger ; epidermis smooth, cells small, quadrato-rotund or quadrate, angles thickened, walls firm. Bracts a little larger than the leaves, antical lobe proportionately larger. Perianth prominent, obconical, piano-compressed, mouth often decurved, truncate, wide, inciso- dentate, usually with about 6 segments, finely dentate, capsule oval, projecting a little. Perigonial spikes usually terminal, bracts closely imbricate, lobes almost equal, ventricose at the base, antheridia oval. Upper leaves often gemmiparous, gemmae oval, usually biseptate. Var. minor. Small, rarely more than ^- in. long, more delicate than the ordinary form, and although it is very constant in size 218 HEPATICJE. and habit, I see no character by which it can be separated from the type. Dimensions. — Stems \ to \\ inch long, '3 mm. diam., with leaves 2' mm. to 3* mm. wide, leaves, antical lobe 7 mm. x "5 mm., *9 mm. X '7 mm., I'l mm. x '8 mm., postical 1' mm. x -7 mm., 1*7 mm. X '9 mm., 1-7 mm. x 1* mm., cells "02 mm., perianth 2*2 mm. long x 1'7 mm. wide at the month, gemmae -03 mm. x •015 mm. Hab. — On shady rocks and walls, more rarely base of trees, generally in subalpine districts. Moderately common. 1. Penzance, W. Curnow. 2. Eridge Rocks, Dr. Spn/ce. 3, 7. Common in Merionethshire and Carnarvonshire, W. H. P. 10. Ilkley, Dr. Carrn)(jton, G. Stabler, G. Webster. Malham Moor, Dr. Carrington. 11, 12. Frequent in Westmorland and Cumber- land, G. StaljJer, Dr. Carrington, W. H. P. 13. Frequent on the hills in Kirkcudbrightshire and Dumfriesshire, /. Mc Andrew. 14, 15, 16. Very common, Moidart, West Inverness, 8. M. Macvicar. I. Very common and widely distributed in Ireland, Dr. D. Moore. Killarney, McArdle §• Lett. Co. Cavan, D. McArdle. Found on the Continent, Teneriffe. Obs. — Distinguished at once from any of the forms of Scapania undidata by its brownish-olive colour, from Seapania nemorosa by the coarser teeth of the leaves, and from Seapania cequiloba and Seapania aqjera by the smooth epidermis. The frequently reflexed antical lobe is also a good character to distinguish it from other species ; this feature is somewhat incon- stant, but is usually found on some portion of the stem. Linnaeus founded his Jtmg. resnpinata on the plant described and figured by Dillenius in his Hist. Muse. p. 491, t. 71, f. 19, which, according to Lindberg, comprises two species, one of which he named Martlnellia graeilis. This, however, so agrees with the description " acutissime crenatos margined " as well as with the figure h, that I follow Dr. Carrington in considering it the true Jung, resupinata. Seapania resnpinata, recorded from several stations in Staffordshire by Mr. J. E. Bagnall (" Journ. of Bot.," March 1896), all refer to Seapania aspera, Mlill. et B. SCAPANIA. 219 Description of Plate LXXXV. — Fig. 1. Plants natural size. 2. Portion of stem x 24 (Scotland, Sim). 3-5. Leaves x 24 (G. & li. n. 225). 6-8. Leaves of var. mijwr x 24 (Ardingley, Davies). *J. Portion of leaf x 290 (New Galloway, J. McAndrew). 10. Perianth x 1 6 (ditto). 11. Portion of the mouth of perianth X 85 (ditto). 12. Gemmae x 290 (Ardingley, Davies). 7. Scapania ornithopodioides {DHL iFifher.), PeavH. Lichenastrum auriculatum, Ornithopodii pinnatis ciliatis, Dill. Hist. Muse. p. 493, n. 21 (1741). Jungermcmia ornithopodioides, Withering, Botanical Arrangement, vol. ii. p. (il),"), n. 14 (177G). Jimgermania planifolia, Hook. Brit. Jung. t. (!7 (1810). Scapania planifolia, (Hook.) Dum. Recueil, p. 14 (18;>.'j). Dioicous (?), forming erect crowded tufts or straggling amongst other mosses and hepatics ; very large ; of a dark brown colour. Stems simple or furcate, ligneous, a cross-section showing it to be oval, the outer layer consisting of one or two rows of dark-red ligneous cells, inner cells white, somewhat homogeneous ; radicu- lose, rootlets short, pale, produced from the postical side of stem. Leaves distichous, imbricate or approximate, divided to the base or almost into two unequal lobes, the antical one about half the size of the postical, erecto-patent to patent (40°), cordate, obtuse or acute, ciliate-dentate, crossing the stem, slightly convex ; postical lobe patent-divergent to horizontal (80°), plane or slightly deliexed, ovate, ciliate-dentate ; at the suture one or two lar^-e cilia are often found, 3 to 4 cells long, 2 cells broad at the base ; epidermis smooth, cells small, roundish, guttulate, cell-walls thick, angles thickened. Specimens were .submitted to bis IrieiKl, the late Dr. Spruce, wbo gave it a MS. name wbicb, bowever, was never publisbecl. In Part I. of Husnot's "Hep, Gall." p. 22, it is publisbed as S. nemorosa, var. infcniicdia, and on Plate 111, Fig. 23, given as /S'. 'inl('nii('(h((. Abbe Lamy, in "Revue Bryol." p. 54 (1876), says be bas observed tins species growing in several ravines about Mont-Dore under different conditions, wbere it retains a perfect conformity of aspect and character, distinguisbing it no less from 8. timbrosa tban from S. iieuiorosa, and be considers it well deserving of specific rank, and approves of the name infer media. Tbe acute antical lobe separates it from >S'. /leii/oro.^a and the rotundate postical lobe from S. imihrom at once. Note on IScapaiiia iiifertiiedia, Husn., by Mr. M. B. Slater : " Scapania intermedia, Husn., approaches in some of its charac- ters to S. piirpurasceni^ (Hook.); it is, bowever, a smaller plant, and resembles more a broad-leaved S. umhrosa ; in cell structure and shape of leaf lobes it much resembles jS. purjourascens. "It is generally of a pale rosy colour, often whitish, and never assumes the fine purple colour of some of tbe forms of S. jj/a-pu- rascen-s. " The latter plant grows very abundantly in the tributarj- streams of the Esk and other moorland rivulets on the North Yorkshire moors, growing on stones by the stream sides, which are often inundated during flood times, and it grog's most luxuriantly on stones where the water is constantly trickling over its roots, assuming fine rosy and purple hues in places exposed to sunlight, becoming greener under the shade of the stream banks or under trees. " S. intermedia always grows on drier rocks out of the stream, often in company with S. urnJjrosa, Diplojjhtjllum albicans, and HarpantJius scutatm. The name is very appropriate to the plant, indicating its intermediate character between S. umhrosa and purpurascens." Description of Plate XCI. — Fig. 1. Plants natural size. 2. Stem, antical view x 16. 3-7. Leaves x 24. 8. Portion 230 HEPATIC^. of the margin of leaf x 85. 9. Portion of leaf x 290. 10. Bract X 16. 11. Perianth and bract x 11. 12. Perigonial bract, explanate x 24 (Littlebeck, Eskdale, M. B. Slater). 13. Scapania irrigua [Nees), Dirin. Jungermania irrigua, Nees, Nat. Eur. Leb. 1, p. 193 (1833). Scapania irrigua, Dum. Recueil, p. 15 (1835). Dioicous ; in spreading tufts or creeping loosely amongst Sphagna, medium size, of a pale green or brownish colour. Stems ascending or prostrate, lax, rooting up to the apex, simple or slightly branched. Leaves horizontally inserted, distant or approximate, lax, divided unequally to about the middle into two lobes, entire, or the upper leaves sometimes distantly denticulate ; antical lobe about half the size of the postical, convex, subquadrate or sub-rotund, acute or sub-mucronate ; postical lobe subquadrate to sub-rotund, acute or sub-mucronate ; texture delicate, cells smiallish to moderate in size, roundish, angles slightly thickened. Perianth obovate to oval, mouth dentate. Perigonial stems with bracts subimbricate, ventricose at the base ; antheridia numerous, sometimes 4 in each bract. The plant is sometimes gemmiparous ; gemmse small, greenish, at the margin of the upper leaves. Fruits April, May. Dimensions. — Stems from ^ to 1 inch long, -3 mm. diam., with leaves 2" mm. to 3" mm. broad; leaves, antical lobe 1* mm. x 1*2 mm., '7 mm. x 1" mm., '6 mm. x "8 mm. ; postical lobe 1'4 mm. X 1*8 mm., r mm. x 1*3 mm. ; cells '025 mm. x '03 mm., -025 mm. X '025 mm. ; perianth 2" ram. long x 1-3 mm. broad, 1'6 mm. X '9 mm. Hab. — Growing in swampy places or on wet banks. Some- what rare. 1. Marazion Marsh; Trungle Moor; Chyanhal ; Tremethick Moor, West Cornwall, W. Curnow (as Scapania ulir/hwsa, Trans. Penz. N. H. Soc. 1881-82). 2. New Forest, C. Li/ell. Guestling near Hastings, Rev. E. N. Bloonipcld. 5. Shirley Heath, SCAPAXIA. 231 Warwickshire, /. J'J. Bcu/ixill. Gospel End ; Newborougli ; Abbot's Bromley ; Arley Wood ; Swynnerton ; Leek ; Cloud, Stafford- shire, J. E. Bcujnall. 7, 8. Kinder Scout, Derbyshire, Holt tb Whitehead. 9. Morley Common ; Walton Swamp, W. Wihon. Oakmere, G. A. Holf. 10. Strensal Common, Yorks, G . S fabler. 12. Foulshavv Moss, /. M. Barnes & G. Stabler. Staveley, West- morland, G. Statj/er. Injebreck, Isle of Man, G. A. Holt. 13. Barend Moss, Castle-Douglas, /. McAmlreic. 15, 16. Moidart, West Inverness, S. M. Macvicar. I. Cromaglown ; Benbulben, Co. Sligo ; Loch Bray, Dr. D. Moore. Found on the Continent. Obs. — This species, which appears to be more peculiar to low- lying localities, is distinguished from 8. ///id/'lata by its subquad- rate, acute leaf-lobes. A small form, growing in drier stations than the normal form, may be confounded with S. ciirfa, but it maybe separated by its stouter habit and the more quadrate shape of the leaf lobes. Description of Plate XCII. — Fig. 1. Plants natural size. 2. Stem X 16 (G. & R. n. 507). 3. Portion of stem x 16 (Thed. Ex. n. 147, as Jun(j. uliginosa). 4. Male stem x 24 (Cheshire, Wilson). 5. Stem x 16 (Essex). 6. Perigonial bract with antheridia x 24 (Foulshaw Moss, Barnes). 7. Leaf x 16 (Husn., Hep. Gall. n. 102). 8. Perianth x 24 (G. & R. n. 319). 9. Mouth of perianth x 31 (Cheshire, Wilson). 10. Portion of leaf x 290 (G. & R. n. 319). 14. Scapania uliginosa [Sirarfz), Dhih. Jungermania uliginosa, Sw, in Lindenb. Syn. Hep. p. .">;) (1820). Radula uliginosa, Dmn. Syll. Jung. p. 40 (1831). Scapania uliginosa, Dum. Recueil, p. 14 (1835); Nees in G. L. N. Syn. Hep. p. 07 (1844). Plagiochila uliginosa, M. &. N. in Nees, Hep. Eur. Ill, p. 522 (1838). Dioicous; in loose tufts or often floating in boggy places, large, brown to reddish olive green in colour, when dry dark 232 HEPATIC^. brown. Stems simple or slightly branched ; leaves regular, sub- transversely inserted, distant to approximate, deeply divided into two lobes, margin entire or rarely minutely and distantly denticulate, aniical lobe about ^rd smaller, reniform, arched, decurrent, incum- bent, convex, postical lobe subrotund, a little reflexed ; cells rather large, sometimes elongate, hexagonal, walls thin. Bracts similar to the leaves. Perianth obovate, mouth wide, entire, rarely minutely denticulate ; spores smooth, pale brown ; elaters reddish- brown. Male stems somewhat similar to others ; perigonial bracts imbricate, ventricose, lobes more equal. Fruits May, June. Dimensions. — Stems 2 to 4 inches long, -2 mm. to -4 mm. diam., with leaves 2-5 mm. to 3-5 mm. broad ; leaves, antical lobe "9 mm. x '9 mm., 7 mm. x 7 mm., "4 mm. x '6 mm., postical 1-9 mm. X 2'2 mm., 1-9 mm. x 175 mm., I'S mm. x 1-25 mm. ; cells '05 mm. x '03 mm., '04 mm. x "035 mm. ; perianth 3*5 mm. X 1-75 mm., 1*5 mm. x 1* mm. ; valves of capsule 1'4 mm. X '65 mm. ; spores "02 mm. ; elaters "08 mm. long x -01 mm. broad. Hab. — Grrows on wet rocks and in swampy places in subalpine and alpine districts. Kare. 7. Clogwyn du Arddu, Snowdon, Carnarvonshire, G. A. Holt, 1883. Nant Francon, Carnarvonshire, Br. Carrington & W. H. P., 1886. 10. Bog, south of Ingleboro', Yorks, Dr. Carrington, 1857. 15. Ben Mac Dhui, A. Croall, W. West, T. Rogers. Ben Lawers, G. A. Holt, IF. West. 16. Loch Rannoch, Br. Carrington. Moidart, West Inverness, S. M. Macvicar. I. Knockavoila, Br. Taylor. Lough Bray, Br. B. Moore. Near the Hunting Tower, Cromaglown, Br. B. Moore. Connor Hill, Co. Kerry, Br. B. Moore. Found on the Continent. Obs. — A few fine teeth are to be observed on the leaves and at the mouth of the perianth of some specimens when highly magnified. The only species with which 8. idiginosa can be con- founded are 8. mididata and S. irrigiia, and from the former it may be distinguished bj' its very reniform antical lobe, which SCAFANIA. 233 retains, all the length of the stem, its relatively small size ; in S. undulata the two lobes usually become more equal near the apex, the cells are usually larger and the leaves more delicate ; in 8. irrifjaa the postical lobe is more or less acute, the antical never reniform, and the lobes are more equal in size. Description of Plate XCIII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 24 (Lapland, S. O. L.). 3. Portion of stem, postical view x 24 (Ben Lawers, West)- 4. Portion of leaf x 290. 5. Perianth x 31 (5iO, G. & K). 15. Scapania rosacea [Corda), Dniii. Jungermania rosacPM, Corda in Sturm. Deutschl. Fl. 11, xxli.-xxlii. p. !)G, f. 29 (1832). Scapania rosacea, Dum. Recueil. p, 14 (183.")). Plagiochila rosacea, M. & N. in Nees, Nat. Eur. Leberm. Ill, p. 525 (1838). Scapania curta, var. rosacea, Carr. Brit. Hep. p. 87 (1874-76). Dioicous ; csespitose ; small ; claret coloured. Stems simple or rarely sparsely innovant, erect or erecto-decumbent, somewhat tender and fleshy; radiculose, rootlets dirty-white. Leaves approximate, patent-divergent, distichous, unequally bilobed, postical lobe obliquely obovate, cultriform, shortly cuspidate or apiculate, plane or slightly concave, antical lobe much smaller, subquadrate, obliquely ovate to cuneiform, acute, ascending ; margin quite entire ; epidermis smooth, texture somewhat lax ; cells small, roundish, walls and angles thickened, no trigones. Bracts rather larger than the leaves, with apices of the lobes more rounded. Perianth projecting about half way beyond the bracts, ovate-oblong, compressed, subplicate, mouth truncate, entire or slightly irregular. Male stems rather smaller ; perigonial bracts on the upper half, approximate or imbricate, erecto-patent, more equally bilobed than the leaves, lobes more rounded, swollen at the base ; antheridia large, roundish-oval. Fruits March, April, May. Dimensions. — Stems J to |- inch long, 3' mm. thick, with leaves 1"5 mm. to 2" mm. broad; leaves, antical lobe 'G mm. long 234 HEPATIC^. by -4 mm, broad, postical '8 mm. x -4 mm., antical '6 mm. x "3 mm., postical '9 mm. x -5 mm. ; cells '02 mm. ; bracts, antical lobe -7 mm. x '5 mm., postical '9 mm. x "6 mm. ; perianth 1*75 mm. long by '75 mm. broad; pistillidia 75 mm. x '3 mm. ; perigonial bract, antical lobe '6 mm. x "4 mm., postical '8 mm. x •5 mm. ; antheridia "2 mm. Hab. — On damp shady rocks. Rare. 9. By the side of a stream, Woodhead, Cheshire, /. Whitehead & W. H. P., July 1880. Alderley Edge, Cheshire, G. A. Holt, March 1883. 12. Kentmere, Westmorland, Rev. H. IF. Lett, August 1899. 13. New Galloway, /. Mc Andrew. 15. Sculty, near Strachan, /. Sim, March 1886. 16. Moidart, West Inver- ness, S. M. Macvicar, 1898. Found on the Continent. Obs. — This is a pretty little species and quite distinct from S. curt a (Mart.). Whether it is the true Jwif/. rosacea of Corda is doubtful, for it does not agree with the figure given in Sturm's Deutschl. Flora, which seems to represent some small form of S. undulata. S. rosacea is at once distinguished from S. curt a by its rather larger size and beautiful rosy or claret-coloured foliage. The stems are somewnat succulent and tender, which character alone separates it from any of the small forms of the other British SeajJanicB. Description of Plate XCIV. — Fig. 1. Plants natural size. 2. Portion of stem x 31 (Woodhead, Cheshire). 3. Leaf x 31 (ditto). 4, 5. Leaves, explanate (ditto). 6. Portion of leaf x 290 (Heisingfors, S. 0. Lindberg). 7. Bract x 31 (Woodhead, Cheshire). 8. Bract, explanate x 31 (ditto). 9. Perianth x 31 (ditto). SOAPANIA. 235 1 G. Scapania curta (^Marf.), Dnm. Jungermania curta, Mart. Fl. Crypt. CJerm. 1, p. 148, t. iv. f. 24 (1817). Jungermania nemorosa,\?iV. denadata, Hook. Brit. Jung. t. xxi, ff. IT-l'.' (181 G)- Radida curta, Dum. Syll. Jung. p. 40 (18:31). Scapania curta, Dum. Recueil, p. 14 (1835). Plagiochda curta, M. et N. in Nees Nat. Eur, Leberm. iii. p. 525 (1838). Dioicous, loosely csespitose, minute, yellowish-green to light- brown. Stems procumbent, simple or subramose, radiculose. Leaves from patent to patent-divergent, 50° to 70°, approximate, subimbricate, accrescent, entire or slightly dentate, to about the middle unequally bilobed, complicate, postical lobe much larger than antical, oblong-oval, acute, antical lobe usually more erect, longer than broad, suboblong-quadrate, acute ; cells small, roundish, angles of cells thickened, trigones distinct, epidermis smooth. Bracts similar to leaves, only larger. Perianth longish obovate, slightly folded above, mouth irregularly dentate. Antheridia terminal on separate stems, perigonial bracts 4-6 pairs, lobes sub- equal, antheridia oval. Fruits April, May. Capsules extremely rare. Dimensions. — Stems |- to f inch long, diam, '2 mm., with leaves 2' mm. broad; leaves, antical lobe *8 mm. broad x "4 mm. high, "9 mm. X "4 mm., "6 mm. x 3 mm., postical lobe I'l mm. broad X "6 mm. high, 1*2 mm. x '6 mm., '9 mm. x "5 mm.; cells 1"50 mm. X 02 mm. ; bract, antical lobe 1" mm. broad x "6 mm- high, postical lobe 12 mm. x '8 mm.; perianth 3*5 mm. long x 1*5 mm. broad. Hab. — Grows on beaten paths and shady banks in woods, &c. Generally distributed. 2. Sussex, ir. MUten. Ardingley Kocks, Sussex, G. E. Dacics. 3. Epping Forest, E. JL Holmes. 5. Gospel End Common ; Seckley Wood, Staffordshire, J. E. BagnalL 7. Cader Idris, Merionethshire, C. J. Ulld (£• G. A. Holt. Tyn-y-Groes, Merionethshire, //'. //. P. Snowdon, Carnarvonshire, G. E. Hunt. 8. Near Woodhead, Derbyshire, G. J. Holt. 9. Staley Brushes, Cheshire, //". Sfajilcj/ <0 C. ./. IJlId. Barton Moss, Lane, Dr 236 HEPATIC^. Carnngton. Easegill, West Lane, /. A. Wheldon. 10. Rigton Hill, Dr. F. A. Lees. Idle Woods, Dr. Carringfon. Yeadon, Dr. Carrington. Shipley Glen, W. West. Near Bingley, W. Wesi. Newsholme Dean, W. West. 12. By the highway side, Staveley, Westmorland, G. Stabler. 14, 15. Trossachs, J. Cruicks/icuik. Near Stirling, Dr. Greville. Ben Mac Dhui, A. C. Black. Forfar Burn, A. Croall. 16. Moidart, West Inverness, very little seen, 8. M. Macvicar. I. Sillaghbraes and Sleemish Mountain, Co. Antrim ; GlenifF, Co. Leitrim ; Benbulben Range, Sligo ; Cromaglown ; near Let- terfrack, Co. Gal way. Dr. D. Moore. Slieve Glah, Co. Cavan, D. McArdle. Found on the Continent and in North America. Obs. — A pretty little species, the smallest of our British Sca- pania ; it can be conlused only with immature forms of other species, and if found with either ^ or $ can be easily identified by the description. Jimgermania Conradi Corda, according to original specimens, is a small imperfect form of this species. Description of Plate XCV. — Fig. 1. Plants natural size. 2. Stem X 24 (Sweden, Lmdb.). 3. Stem x 24 (G. & R 651). 4-8. Leaves x 24 (ditto). 9. Portion of leaf x 290 (Sweden, Lindb.). 10. Bract x 24 (G. & R 651). 11. Perianth x 11 (Sweden, Lindb.). 17. Scapania umbrosa i^Schrad.), Dmit. Jungermania convexa, Scopoli, Flora Carnolica, 2nd ed. p. 349, sp. 1348 (1772) (?). Jungermania iimbrosa, Schrad. Syst. Samml. Krypt. Gew. 2, p. 5 (1707); Hook. Brit. Jung. t. 24 (1810). Radula umbrosa, Dum. Comm. Bot. p. 112 (1823). Scapania umbrosa, Dum. Recueil, p. 14 (1835) ; Howe, Hep. & Antho. California, Mem. Torrey Boc. Club, vol. vii. p. 153 (1890). PlagiocMla umbrosa, Mont, et Nees in Nees Eur. Leberm. iii. p. 525 (1838). Martinellia convexa, Lindb. Musci. Scand. (1879). Scapania convexa (Scop.), Pears. List. Can. Hep. 15 (1890). Dioicous, csespitose, of a yellowish or dark green to br •own 8CAPANIA. ■2:i7 colour. Stems ascending, subramose above, branches subfalcate, radiculose. Leaves imbricate or approximate, patent to patent- divergent (60°), complicate, unequally bilobed to about the middle serrate to tlie upper half, antical lobe |- to ^ smaller, roundisli, ovate, acute, erect, incumbent, postical lobe oval or obovate, acute, convex, subsecund, epidermis smooth, cells guttulate, smallish, roundish or roundish-oblong, cell-walls thick, trigones distinct. Bracts similar, only larger. Perianth oblong, truncate, mouth entire. Capsule oval, dark reddish-brown. Spores pale brown, minute. Klaters sliort, thick, dark reddish-brown. Perigonial stems with leaves closely imbricate, lobes smaller, more equal, slightly ventricose, anther idia oval. Fruits April, May. Dimensions. — Stems |^ to f inch long, diam. '3 mm., with leaves 1*5 mm. broad ; leaves, antical lobe '85 mm. x -45 mm., l-'2 mm. X -575 mm. ; cells "03 mm. x '02 mm. ; walls '005 mm. thick; bracts, antical lobe 1'05 mm. x "5 mm., postical 1*3 mm. X '7 mm.; perianth 1'6 mm. x '65 mm.; pistillidia "2 mm. X '04 mm. ; capsule "7 mm. x "55 mm. ; spores "OlS mm. diam. ; elater "08 mm. x '015 mm. ; antheridia '09 mm. x "06 mm. Hab. — Grows on rocks and rotting wood in shady woods and by the sides of streams. Rare. 1, 2, 5. Ordesley Wood; Mayfield; Dimmings Dale, Staffordshire, ./. U. BagnaU. 7. Dolgelly, Merionethshire, C. J. Wild & IF. H. P. Festiniog, Merionethshire, Dr. Car- ringtoii & W. H. P. 8. Woodhead, Derbyshire, /. Whitehead & W. H. P. 10. Bolton Woods, Dr. Carriiicjton. & G. A. Half, W. H. P. aoathland, M. B. Slater & W. H. P. 12. On decay- ing trees, Naddle Forest, Mardale, Westmorland, G. Stabler. 16. Moidart, West Inverness, common on partly buried rocks and on stumps in ravines and shady woods ; this pretty species is very fine in this district, S. M. Macvicar. I. Nr. Dublin, Dr. Taylor. Loch Bray, Wicklow ; Kylemore, Co. Galway ; Killarney Woods, frequent ; Brandon, Co. Kerry, Dr. D. Moore. Killarney, Dr. Carr'nif/toii. Found on the Continent and in North America. 238 EEPATICjE. Obs. — One of the most distinct species of this genus, smaller than most of them, but more robust than S. curta ; its colour and the acute serrated leaf-lobes distinguish it from them all. The late Prof. Lindberg, in his "Muse. Scand." (1879), stated that the Jim^. umbrosa of Schrader (1797) was without doubt the same as the Jiin(/. convexa of Scopoli (1772), so along with others I adopted the older name. Prof. Howe, in his admirable " Hepaticse and Anthocerotes of California," says, " it is very probable that it may be so, yet, as Scopoli describes the perianth as oval and dentate at the apex, while those of S. umhrosa are oblong and entire at the mouth, we prefer to retain for this species a name concerning the correct application of which we can have no doubt," so I gladly follow my friend Prof. Howe in retaining this very characteristic one. Description or Plate XCVI. — Fig. 1. Plants natural size. 2. Stem X 31 (15, Austin, Amer. Hep.). 3-7. Leaves x 24 (20, C. & P. Hep. Brit.). 8. Portion of leaf x 290 (Woodhead, Whitehead & W. H. P.). 9. Bract x 31 (Husn., Hep. Gall, n. 66). 10. Perianth x 24 (Goathland, W. H. P.). 11. Pis- tillidia x 85 (Husn. Hep. Gall. n. 66). 10. Perigonial bract x 31 (Woodhead, Whitehead «& W. H. P.). 13. Antheridium X 85 (ditto). Genus 21. DIPLOPHYLLUM, Dum. Jungermania, Mich. Nov. PI. Gen. p. 8, tab. 5, fig. 9 (1729) ; Linn. Fl. Suec. 1st ed. p. 335(1745); Hook. Brit. Jung. (1816). Diplophyllum, Dum, Eecueil, 1, p. 15 (1835). Scapania, Mitt, in Hook. Fl. Tasm. 2, p. 233 (1858). Brandies axillary, proceeding from the inner base of the leaves. Leaves conduplicate, unequally bilobed, the antical lobe smallest. Stipules absent. Involucral bracts small, similar to the leaves. Bracteole absent. Perianth free, sessile, erect, terete, subcylin- drical, pluriplicate, mouth denticulate. Capsule coriaceous, 4-valved. DIPLOVHYLLUM. 239 1. Diplophyllum albicans (Z.), Dim. Hepaticoides albescens foliis pinnatis,Ya,i\\. Bot. Par., p. lOO, t. 19, f. 5 (1723). Jungermania repens foliis cordatis carinatis, Mich. Nov. pi. gen. p. 8, t. 15, f. 9 (1729). Lichenasiram auriculatum, pinnulis augustis planis recurvis, Dill. Hist. Muse, p. 492, t. 71, f. 20 (1741). Jungermania albicans, Linn. Sp. pi. p. 1599 (1753), Hook. Brit. Jung. t. 25 (181(5). DiplojyhyUitm albicans, Dum. Recueil, p. 10 (1835). Dioicous ; growing in loose or matted tufts, medium size, of a green, greenish-brown, or often reddish-brown colour, as if singed. Stems simple or subramose, ascending ; radiculose, rootlets short, close, dirty- white, ascending to apex of stem. Leaves bifarious, contiguous, conduplicate, unequally divided to about the J, vittate, margin entire or subdenticulate, often erose ; antical lobe half the size of the postical, incumbent, 30°-40°, lanceolate, acute or rounded ; postical lobe slightly ascending 60^-80^ lanceolate or ovate-lanceolate, acute or rounded, cells subquadrate, small or rather minute, cell-walls thick, no trigones. Bracts similar to leaves, only larger. Perianth terminal, obovate, 5-plicate at the upper portion, mouth contracted, laciniate-dentate, h^^aline. Perigonial bracts in the middle or at the end of the stem, usually 4 to G pairs, often a little smaller, more erect; antheridia 1 or 2 in the swollen base of the antical lobe, oval, very large. Fruits March, April. Dimensions. — Stems from ^ inch to 2 inches long, diam. "2 mm. to '5 mm. ; leaves, antical lobe 1* mm. x '5 mm., postical 1*75 mm. X '75 mm., antical '6 mm. x "25 mm., postical '9 mm. x '4 mm. ; cells '0175 mm., "02 mm. ; bract, antical lobe 1*25 mm. x '5 mm., postical 1"7 mm. x '75 mm. ; perianth 2* mm. x 1' mm. ; perigonial bract, antical lobe '7b mm. x "5 mm. ; postical lobe 1'24 mm. X "5 mm. ; antheridia '3 mm. x "2 mm. Hab. — Grows everywhere, on rocks and banks in shady woods or by roadsides, on the ground or walls; xery common, from the lowlands to considerable heights. 1 to IS. I. C. Found on the Continent, Madeira, North and South America. 240 HEP AT 10 jE. Obs. — This is the commonest species known, being at once recognised bv the presence in the antical and postical lobes of a pseudo-nerve, which is often colourless, and consists of a series of from 4 to 6 elongated cells, a cross-section of the leaf showing the cells to be of equal diameter with others, the outer wall only thickened considerably. Description of Plate XCYII. — Fig. 1. Plant natural size. 2. Stem X 24 (Nova Scotia, Macoun). 3, 4. Leaves x 16 (Killarney, Lindberg). 5. Portion of leaf x 290 (ditto). 6. Cross- section of leaf X 85 (France, Du Buysson). 7. Bract x 24 (ditto). 8. Perianth x 24 (ditto). 9. Portion of mouth of perianth x 85 (C. & P. Hep. Brit. Ex. n. 220). 10. Perigonial stem X 60 (ditto), 11. Perigonial bract x 24 (ditto). 2. Diplophyllum taxifolium [Wahleiib.), Dm in. Jungermania taxifoUa, Wahlenb. Fl. Lapp. p. 383, t. 25, f. A-C. (1812). Jungermania albicans, var. taxi/olia, Nees, Nat. Eur. Leb. 1, p. 228 (1833). Diplophyllum taxifolium, Dum. Recueil, p. IT) (1830). Dioicous ; growing in dense, depressed tufts, small to medium size, of a greenish to reddish-brown colour. Stems simple or ramose, sub-erect to erect, radiculose. Leaves bifarious, a little distant or contiguous, conduplicate, unequally divided to below the middle, margin entire or minutely denticulate, antical lobe about |- to ^ the size of the postical, incumbent, patent to patent-divergent, 40°-60°, oblong-oval to ovate, rotundate ; pos- tical lobe spreading at right angles or slightly ascending, divergent or horizontal, 80°-90°, oblong-oval to lanceolate; cells minute to rather minute, subquadrate, cell-walls thick, no trigones. Bracts similar to the leaves, larger Perianth terminal, obovate, upper portion 5 -plicate, mouth contracted, lacinia;te-denticulate. Pistil- lidia about 20. The male plant I have not seen. Fruits May, June. Dimensions. — Stems from J to |- inch long, diameter "3 mm. ; leaves, antical lobe 1* mm. x -4 mm., postical 1*25 mm. x -5 mm., DIPLOVHYLLUM. IW antical 7 mm. x '.Vd mm., postical 1 ' mm. x '4 mm.; cells 015 mm. bracts, antical lobe I'l mm. x -6 mm., postical r4 mm. x 'G mm. ; perianth 1'75 mm. x '!) mm., 2'25 mm. x V'l'o mm.; pistillidia •2 mm. X '05 mm. Hab. — On rocks in alpine situations. 7. The Grlyders, Carnarvonshire, W. Wilson, 1844. 15. Ben Lawers, Perthshire, Rev. C. IL Butsfead, identified and comm. by G. A. Holt. 1(3. Moidart, West Inverness, S. M. Macuicar. In similar localities on the Continent, in North America and Canada. Obs. — This species has usually been considered a variety of the previous one, but Prof. Lindberg regarded it as distinct, its neater habit, generally smaller and more graceful form, the direction of the postical lobe being more horizontal, the lobes generally more rounded, the absence of the pseudo-nerve — although traces are sometimes observable — sustain this view. Description of Plate XCVIII. — Fig. 1. Plants natural size. 2. Stem x 16 (Thed. 197). 3. Portion of stem x 31 (Norway, Lindberg). 4. Leaf x 24 (ditto). 5. Portion of leaf X 290 (ditto). 6, 7. Bracts x 24 (ditto). 8. Perianth x 24 (ditto). 9. Portion of mouth of perianth x 85 (ditto). 10. Pistillidium x 85 (ditto). 3. Diplophyllum obtusifolium [Hook.), Dion. Jangennania ohtusifolia, Hook. Brit. Jung, t, 2G (181G). Dlplojjhijllum obtusifolium, Dum. Recueil, p. IG (1885). Paroicous or monoicous, densely ctespitose, small, green, greenish-brown or brown in colour. Stems simple or innovantly branched, postical side of stem often of a reddish tinge, sub-erect or ascending ; radiculose up to apex, rootlets thick, dense, long, hyaline. Leaves accrescent, bifarious, imbricate, conduplicate, unequally bifid to about the middle, margin minutely denticulate or entire, antical lobes 2 or 3 times smaller than the postical incumbent, erect or erect-patent (l0°-20"), ovate or oblong, obtuse or acute, postical lobe horizontal or slightly ascending (90-80 ), Q 242 HEPATICJ^. concave, slightly undulate, subaciniciform or oblong, obtuse or sometimes sliglitly apiculate ; cells small, quadrate, guttulate, near base in postical lobe elongate, cell-walls thick, no trigones. Bracts somewhat similar to the leaves, but larger. Perianth projecting from about ^ to ^ beyond the bracts, terminal, obovate or oblong-ovate, upper portion 5 -plicate, mouth contracted, laciniate, irregularly dentate or denticulate. Capsule oval ; spores reddish- brown ; elaters bispiral, same colour. Antheridia situated in the base of the bracts or on branches proceeding from base of perianth, small, one or two in each bract. Fruits April, May. Dimensions. — Stems about \ inch long, diameter '15 mm. to •175 mm. ; upper leaves, antical lobe "6 mm. x '4 mm., postical lobe 1* mm. x "5 mm., antical '5 mm. x -3 mm,, postical "9 mm. X "4 mm. ; lower leaves, antical lobe '4 mm. x '25 mm., postical 7 mm. x -3 mm. ; cells -015 mm. x '02 mm., '025 mm., basal cells •05 mm. X '02 mm., ^04 mm. x ^02 mm., ^03 mm. x '02 mm.; bracts, antical lobe 1' mm. x ^6 mm., postical 1*3 mm. x '7 mm. ; perianth 2^ mm.xl* mm.; capsule '9 mm. x '6 mm.; spores •01 mm. diam. ; elaters "1 mm. x •Ol mm. ; perigonial bracts, antical lobe ^5 mm. x '4 mm., postical ^9 mm. x "5 mm. Hab. — Grows on shady banks and in old quarries, in company with Scapania rosacea and Jung, bicreuata. Rare. 7. Tyn-y-Groes, Merionethshire, G. A. Holt, May 1885. 9. Delamere, Cheshire, Wilson 8f Carrington. 10, 11. Near Heddon on the Wall, Northumberland, Mr. TIiornhilL 15. Gate- side, Strachan, Perthshire, /. Sim, June 1879. 16. Glen Finnan, Dr. Carringtou, July 1876. Moidart, West Inverness, /S'. M. Mac- vicar ^ W. H. 7^., 1899. I. Near Ban try, Co. Cork, Miss Hufcliins. Near Dunkerron, Co. Kerry, Br. Taylor ; Dunscome's Wood, near Cork, W. Wilson. Generally distributed on the Continent. North America. Obs. — Although Dr. Hooker had an impression that the species was dioicous, it is truly paroicous or monoicous, and this character, along with its small size, distinguishes it from any form of Diplo- DIPLOPHYL L mr. 213 pliijllum laxifoUtmi, for which it has been mistaken by Austin and others. Description of Plate XCIX. — Fig. 1. Plants natural size (Eng. Bot. 2511). 2. Portion of stem x 24 (Glen Finnan, Dr. Carrington). 3. Portion of stem x 31 (ditto). 4. Portion of fertile stem x 24 (Delamere, Wilson k Carrington). 5. Upper leaf X 24 (Tyn-y-Groes, Holt). 6. Upper leaf x 24 (Hb. Tayl.). 7, 8. Lower leaves flattened out x 24 (ditto). 9. Portion of leaf X 290 (Delamere, W. & C). 10. Portion of leaf, near base X 290 (ditto). 11, 12. Bracts x 24 (Hb. Tayl.). 13. Perianth X 24 (ditto). 14. Portion of mouth of perianth x 85 (ditto). 15. Spores x 290 (ditto). 16. Perigonial bract x 24 (Tyn-y- Groes, Holt). 4. Diplophyllum Dicksoni [Hook.), Bum. Jmujerman'm ovata, Dicks. PI. Crypt. Brit. iii. p. 11, t. y, f. G (1793);? Lindb. Musci Scand. p. 7 (1879). Jungermania Dicksoni, Hook. Brit. Jung. t. 48 (1816). Diplophyllum Dicksoni, Dum. Recueil, p. IG (18;3o). Dioicous, growing in loose patches or amongst mosses, small, of a pale yellowish-brown colour. Stems simple or rarely branched, innovant branches arising from base of bracts, apparently subpostical ; radiculose, rootlets ascending to apex of stems, hyaline, long, delicate, stem slightly channelled antically, cortical cells about 40, more distinct than the inner, walls flrmer, inner soft, and somewhat indistinct. Leaves transversel}^ inserted or slightly ascending, antical lobe more erect, sometimes erect to erect-patent, equitant, contiguous or imbricate, to below the middle unequally bilobed, complicate, lobes lanceolate, acuminate, antical lobe smaller than the postical, margin entire, upper leaves slightly serrulate ; cells smallish to medium, subquadrate or roundish, filled witli nucleate bodies, cell- walls delicate, hyaline, trigones small. Bracts ovate, bifid to about the middle, segments lanceolate acuminate or obtuse, margin serrate. Perianth pro- jecting about J- to h beyond the bracts, oval fusiform, deeply 244 HEPATIC^E. 8 -plicate to almost the base, composed of a single layer of cells, about 250 round, mouth slightly contracted, laciniate-ciliate. Perigonial bracts usually situated at the middle of the stem, little different from the ordinary ; antheridia 2 at the base, oval. Fruits April, May. Dimensions. — Stems \ io \ inch long, diameter '25 mm. ; upper leaves, antical lobe 1-1 mm. x "4 mm., postical lobe 1*6 mm. X "6 mm. ; lower leaves, antical lobe '8 mm. x '4 mm., postical I'l mm. X "4 mm., antical 1* mm. x -35 mm., postical I'l mm. X "4 mm. ; cells '025 mm., '03 mm. ; bracts 175 mm. x 1*25 mm., 1"6 mm. X 1-25 mm., segments 1*1 mm., '75 mm., '5 mm. ; perianth 2' mm. x 1*1 mm.jlacinise of mouth '5 mm. long; pistil- lidia '15 mm. x '05 mm. ; perigonial bracts, antical lobe '9 mm. X '3 mm., postical 1*1 mm. x -5 mm. Hab. — Grrows ill crevices of rocks and on the earth in shady, usually subalpine situations. Rare. 1. Dartmoor, E. M. Holmes. 7. Cwm Bychan, Merioneth- shire, E. George, E. M. Holmes. Tyn-y-Groes, Merionethshire, C. J. Wild. 9. Kinder Scout, Derbyshire, /. Wkitehead 8^' G. A. Holt. 10, 12. Staveley; Mardale ; Little Langdale, Westmorland, G. Stabler. Isle of Man, G. A. Holt. "Found many years since in Scotland by Mr. Dickson " (Hook. Brit. Jung, 1816). 13. New Gralloway, /. Mc Andrew. 15. Kinnordy, C. Lyell. Strachan, /. Sim. 16. Moidart, West Inverness, 8. M. Macvicar. I. Near Dublin, Br. Taylor. Shauslieve, Mourne Mts., Rev. H W. Lett, 1898. Found on the Continent and in North America. Obs. — Dr. Hooker wrote in his " Brit. Jung," : '"' There are few naturalists to whom Cryptogamic botany is more indebted than to Mr, Dickson. In the genus Jungermania his numerous additions to the list of British species are well known. The present is one of many collected since the publication of the fourth fasciculus of his Flantce OryptogamiccB in the Highland mountains of Scotland, which he kindly communicated to me ; and I have great pleasure in distinguishing it by his name." LorirnCOLEA. 245 Prof. Liiidberg states in "Muse. Scand." p. 7 that Dickson's original specimens ol Juu(j. ovafa PI. Crypt. Brit. Fasci, 3, p. 11, t. 8, fig. 6 (1793) are identical, but evidently, from the above note by Dr. Hooker, neither he nor Dickson considered them the same, and as the description of Dickson is short and unsatisfactory, and the figure poor, I retain Hooker's name. Dijdopliijllum Dicksoni is a very distinct species, no other British one approaching it. Diplop/i/jl/nm arf/enicuii/ (Tayl.), Spruce, a North American species, is its nearest congener, but is distinguislied by its serrate leaves, 12-plicate perianth, &c. Description of Plate C. — Fig. 1. Plants natural size. 2. Portion of stem x 16 (Dartmoor, E. M. Holmes). 3. Portion of fertile stem x 16 (New Galloway, Mc Andrew). 4, 5. Upper leaves x 24 (ditto). 6, 7. Leaves x 24 (Dartmoor, Holmes). 8. Portion of leaf x 290 (New Galloway, McAndrew). 9, 10. Bracts x 16 (ditto). 11. Perianth x 24 (Dartmoor, Holmes). 12. Cross-section of perianth, upper half x 24 (ditto). 13. Portion of mouth of perianth x 31 (ditto). 14. Ditto x 85 (New Galloway, McAndrew). 15. Pistillidia x 85 (ditto) 16, 17. Perigonial bracts x 24 (Dartmoor, Holmes). Subtribe VI. EPIGONEANTHE^. Genus 22. LOPHOCOLEA, Diwf. Jungermania, Mich. Nov. pi. gen. p. H (1729); L. 8p. pi. ed. 1, 2, p. 1132 (1753). Jungermanict} sect. Lophocolea, Dum. Syll. p. 5!) (1831). Lophocolea, Dum. Recueil, p. 17 (1ifidis, Mich. Nov. pi. gen. p. 8, t. .'), f. 12 (1 7211). Lichenastrum pinnulis acutioi'ibus, concavis, Injidis, majits, Dill. Hist. Muse. p. 487, t. 70, f. 11 (1741). Jimyermania bidentata, Jjinn. Sp. pi. p. loOH (1 7r)3) ; Hook. Brit. Jung. t. 30 (1810). Lophocolea bidentata, Dum. Recueil, p. 17 (18;!;")). Lophocolea Hooheriayia, Nees Nat. Eui-. Leb. 11, p. 33G (183(;). Dioicous, loosely and broadly csespitose, medium size, pale green, sometimes almost white in colour. Stems procumbent, flexuose, firm, ramose, branches ascending; radiculose, rootlets small, whitish. Leaves sub-imbricate, approximate or distant, broadly ovate, antical margin decurrent, plane or slightly undu- late, patent-divergent or horizontal, bifarious, bifid to about 1-J, segments lanceolate, subulate, acuminate, unequal, the lower usually smaller than the upper, sinus rounded or acute. Aromatic. Texture thin, epidermis smooth, cells largish, oblong-hexagonal or 4, 5-sided, walls thin — but with the chlorophyl granules adhering to them they often appear thick — trigones minute. Stipules free, large, sometimes reflexed, bifid or quadrifid, usually bifid to below the middle, with a smaller ses^ment on the two outer margins, segments subulate. Bracts erect, larger than the leaves, oval or broadly oval, bifid, trifid or quadrifid to about J, margin here and there dentate or ciliate, segments and sinus acute. Bracteole oblong-oval, bifid, trifid, or rarely quadrifid to about ^-J,, segments and sinus acute, margin slightly dentate or ciliate. Perianth terminal, ovate-oblong, obtusely triangular, mouth widish, tri- lobate, laciniate. Pistillidia numerous (60), often with a purplish tinge. Calyptra thin. Capsule ovate, deep brown. Spores rich fulvous brown, spherical, about twice as broad as the elaters- Elaters bispiral, dark brown. Perigonial bracts 10-12, terminal or at the middle of a branch, closely imbricate, erecto-patent, roundish-ovate, bifid, trifid, or rarely quadrifid to about J-1, segments acute, sinus rounded ; antheridia 1 or 2, oval ; perigonial bracteole quadrifid. Fruits March, April. 248 HEPATIC^. Dimensions. — Stems 1 to \\ inch long, "4 mm. diameter (young stems '25 mm.), with leaves 3' mm. wide; leaves 2"5 mm. X 1-5 mm., segments "45 mm., 2*25 mm. x 1'5 mm., seg. '3 mm., 3' mm. X 1"75 mm., seg. '5 mm. ; cells "08 mm. x '04 mm., "05 mm. X '035 mm., "06 mm. x '04 mm. ; stipules 1* mm. x 6 mm., seg. "6 mm. ; sub-bracteoles 1"6 mm. x "75 mm., seg. "75 mm. ; bract 3 "mm. X 1*6 mm., seg. "3 mm., '4 mm., "5 mm. ; Ijracteole 2*5 mm. X '25 mm., seg. '3 mm.; perianth 2'75 mm. x 1*25 mm.; pistil- lidia "IS mm. x "06 mm. ; perigonial bract 1"25 mm. x 1* mm., seg. •2 mm., '3 mm. ; perigonial bracteole *9 mm. x '6 mm., seg. '6 mm. ; antheridia '15 mm. x -125 mm. Hab. — Grrows in large spreading patches on walls, stones, earth and rotting wood, in tlie plains and subalpine localities, usually in shady situations. Common. 1 to 18. I. Common on the Continent and in North America. Recorded from the West Indies. Obs. — Distinguished from Lo^jJiocoIea cnspidata, Limpr. and LopJt. JieferojjJi'i/lIa (Schrad.) by its dioicous inflorescence. LojjJio- colea latifolia, Nees (Z. Hookenana, Nees) is a variety with darker green leaves of a firmer texture, usually growing in damper situations, but I cannot fix upon any character to separate it from the type. Description of Plate CI. — Fig. 1. Plants natural size. 2. Portion of young stem x 16 (Ashley, W. H. P.). 3-5. Leaves x 16 (ditto). 6. Portion of leaf X 290 (ditto). 7. Stipules x 24 (ditto). 8. Sub-bracteole x 16 (ditto). 9. Bract x 16 (ditto). 10. Bracteole x 16 (ditto). 11. Perianth x 16 (ditto). 12. Por- tion of the mouth of perianth x 16 (ditto). 13. Pistillidium X 85 (ditto). 14. Perigonial bract x 16 (G. & B. Hep. Eur. n. 630). 15. Perigonial bracteole x 16 (ditto). 16. x\ntheridium X 85 (ditto). LOPHOCOLEA. 249 2. Lophocolea cuspidata, JJntpr. Lophocolea hidentata, var. cuspidata, Nees Nat. Eur. Leb. 11, p. 327 (1836). Lophocolea cuspidata, Limpricht in Cohn Krypt. Fl. Schles. p. 303 (187(;). Moiioicous, loosely csespitose, of medium size, pale yellowish- green in colour. Stems branched, branches spreading, lateral sub- postical ; radiculose, rootlets few. Leaves imbricate, spreading, horizontal or patent-divergent, antical margin decurrent, ovate, bifid to about \, segments cuspidate, divergent or connivent, sinus lunate; texture thin, cells medium size, 5- and 6- sided, walls thin, trigones minute. Stipules oblong, bifid to below the middle, with a shorter tooth on one or both of the outer margins, segments subulate. Bracts oblong-subquadrate or semi-oval, bifid or trifid from 1 to the middle, margin plane. Perianth projecting about half beyond the bracts, cylindrical, acutely trigonous, third angle antical, third face postical, mouth dentate. Pistillidia long. Capsule oval, dark brown. Spores reddish -brown, about as broad again as the dark brown elaters. Androecia on long or short branches on the $ plant ; perigonial bracts 5-8 pairs, smaller than the stem leaves, erecto-patent, closely imbricate, ventricose at the base, trifid, the third antical segment smaller incurved, enclosing 1 or 2, rarely 3, almost spherical antheridia. Fruits March, April. Dimensions. — Stems 1 to 2 inches long, diameter "3 mm,, with leaves 3" mm. wide; leaves 1'6 mm. x 11 mm., segments '4 mm. ; cells "04 mm. x '05 mm., "035 mm. x "04 mm., "03 mm. x 03 mm., •04 mm. X '03 mm. ; stipules 1*25 mm. x '75 ram., seg. "5 mm., •6 mm. X '4 mm., seg. "3 mm. ; bracts 2* mm. x "9 mm., seg. 1" mm. and 7 mm., 1*7 mm. x 9 mm,, seg. "Go mm. and '5 mm., 3' mm. X 1*4 mm., seg. "G mm. and "3 mm. ; sub-bracteole 2" mm. X '9 mm.,seg."8 mm. and 7 mm.; bracteole 3' mm. x 1*25 mm., seg. "8 mm. and "7 mm. ; perianth 5* mm. x 125 mm. ; pistillidia "2 mm. X '075 mm,, "225 mm. x '06 mm. ; perigonial bracts 1'2 mm. X '6 mm., seg. •(> mm ; antheridia "11 mm. x "1 mm. 250 HEPATIC a:. Hab. — Growing in spreading patches at the base of trees, on walls or stones in exposed or shady places, perhaps commoner than is thought by being confounded with Loph. hidentata (L.). 5. Hart Hill, Hoare Park, Warwickshire, /. E. Bagnall. 7. Bettws-y-Coed, Carnarvonshire, W. H. P. 8. Stirrup Wood, Marple, Derbyshire, C. J. Wild. 10. Coneysthorpe Bank Wood, Yorks, M. B. Slater. Ash Hag G-ill, Hackness, Yorks, M. B. Slater. 12. Injebrick, Isle of Man, G. A. Holt. Found on the Continent. Obs. — Distinguished from Lophocolea bidentata (L.) by its monoicous inflorescence, narrower leaves, which are more cus- pidate ; from Lophocolea JieterojjJii/lIa (Schrad.), which is also monoicous, by its more spreading habit, its deeply bifid, cuspidate leaves, which are never entire. Description of Plate CII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 16 (Hackness, M. B. Slater). 3. Portion of leaf x 290 (ditto). 4, 5. Stipules magnified (Isle of Man, G. A. Holt). 6, 7. Bracts x 24 (ditto). 8. Bract x 16 (Hackness, M. B. Slater). 9. Sub-bracteole x 16 (ditto). 10. Bracteole x 16 (ditto). 11. Perianth x 11 (ditto). 12. Cross-section of perianth x 11 (ditto). 13. Portion of the mouth of perianth x 16 (ditto). 14. Perigonial bract x 24 (Isle of Man, G. A. Holt). 15. Antheridium x 64 (ditto). 3. Lophocolea heterophylla [Scl/rad.), Dim. Lichenastrum pinnulis ohtusiorihus hijidis 7ninus, Dill. Hist. Muse. p. 488, t. 70, f. 12 (1741). Jungermania heterophylla^ Schrad. Diar. Bot. p. 66 (1801) ; Hook. Brit. Jung. t. 31 (1816). Lophocolea heterophylla^ Dum. Recvieil, p. 17 (1835). Paroicous, csespitose, small, pale green in colour. Stems pro- cumbent, ramose, branches postical, erect ; radiculose, rootlets short, white, few, fasciculate, produced from the base of the stipules. Leaves imbricate, horizontal, slightly decurrent anti- cally, plane or slightly concave, bifarious, spreading or secund. LOriluiJULEA. 251 ovate, (juadrate, entire or obtusely emarginate, rarel}- acutely bifid ; texture somewhat firm, cells medium size, 4-, 5- and G- sided, walls thin, no trigones. Stipules oblong, bifid to below the middle, segments subulate, outer margin unidentate, on young branches bifid only or minutely unidentate on one side. Sub- bracts (perigonial) oblong roundish quadrate, emarginate or entire, lobulate or deeply unidentate at the base, one or two roundish antheridia enclosed in the lobule. Sub-bracteole oblong, bifid or trifid, segments subulate, acuminate. Bracts closely imbricate, oblong quadrate, roundish, or oblong-quadrate, entire or obtusely emarginate, margin entire or furnished with one or two teeth, lobulate or elobulate. Bracteoles oblong bifid to about the middle segments subulate, margin sparsely laciniate-dentate. Perianth terminal on main stem or branches, projecting about half beyond the bracts, ovate, obtusely triangular, mouth wide, trilobate dentate. Calyptra ovate, thin, delicate. Capsule ovate, dark brown. Spores and elaters brown. Fruits March, April, May. Dimensions. — Stems |- to 1 inch long, -25 mm. diam., with leaves 2* mm. wide ; leaves 1" mm. x 'G mm., V'l mm. x 1' mm., 1*25 mm. x '8 mm. ; cells '03 mm. x '05 mm., "03 mm. x "035 mm., •04 mm. X '035 mm. ; stipule '9 mm. x "3 mm., segments "55 mm. ; sub-bract 1*4 mm. x "9 mm.; sub-bracteole \'\ mm. x 6 mm., segments "G mm. ; bracts 1'4 mm. x 1' mm., 1"4 mm. x '8 mm.; bracteole r2 mm. x '7 mm., segments "5 mm. ; perianth 2* mm. X 1* mm. ; antheridia "15 mm. x '14 mm. Hab. — Growing usually at the base of trees or on rotting wood. Common. 1-5, 7-14, IG. I. Found on the Continent and in North America. Obs. — Difi'ers from Lophocolca hhlonlata (L.) by its being paroicous ; from L. ct/spldafa, Limpricht. by its usually smaller size, its variously shaped leaves, entire, emarginate and bifid — hence its name. Growing with it are often very small young stems with leaves deeply bifid, stipules bipartite with a smull tooth on one side. 252 HEPATIC^. resembling small forms of Geoccdyx graveolem, but if their origin, or connection with naature plants be traced, the difficulty is solved. This and the previous species have the strong unpleasant smell, more evident when drying, of L. bidenfata. Description of Plate CIII. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 16 (Dove Dale, AV. H. P.). 3. Portion of young branch x 64 (Abergele, W. H. P.). 4. Leaf X 24 (Castle Mill, Cheshire, G. A. Holt). 5. Ditto x 24 (Bowdon, W. Stanley). 6. Ditto x 24 (Dove Dale, W. H. P.). 7. Portion of leaf x 290 (ditto). 8. Stipule x 24 (Bowdon, W. Stanley). 9. Ditto from young branch x 85 (Castle Mill, G. A. Holt). 10. Sub-bract x 24 (Dove Dale, W. H. P.). 11. Sub-bracteole x 24 (ditto). 12, 13. Bracts x 24 (ditto). 14. Bracteole x 24 (ditto). 15. Perianth x 16 (ditto). 16. Ditto, explanate to show mouth x 16 (ditto). 17. Antheridium X 64 (Bowdon, W. Stanley). 4. Lophocolea spicata, Ta?/L Lophocolea spicata, Taylor in G.L.N. Syn. Hep. 1G7 (1844). Monoicous, closely appressed to rocks, in entangled masses, small, pale green in colour. Stems creeping, irregularly branched, outer layer of cells large, hyaline ; radiculose, rootlets few, small, whitish. Leaves gradually increasing and decreasing in size, horizontal (90°) to patent-divergent (70°), slightly imbricating, antical margin decurrent, ovate, diversiform, bi-, tri or rarely multi- dentate, segments acute, connivent or divergent ; texture delicate ; cells small, subquadrate or oblong-quadrate, 4-, 5- or 6-sided ; walls moderately thick, no trigones. Stipules free, bipartite, segments acuminate, unidentate near the base of the exterior margins. Involucres on short branches ; bracts oblong, upper portion irregularly dentate ; bracteole bipartite, segments acutate, irregu- larly dentate. Perianth projecting beyond the bracts, triquetrous, angles not winged, mouth wide, laciniate-dentate ; pistillidia LOPHOCOLEA. 1h?> about 10. Calyptra, after rupture irregularly lobed. Capsule oval ; spores light brown ; elaters bispiral, hardly as broad as the spores. Male amentula on short branches above or below the ?, 2-5 pairs of perigonial bracts which are closely imbricate, ovate, complicate, swollen at the base ; antheridia single, oval, shortly stipitate. Mixed with the plants are many minute leafy- branches. Fruits April, May. Dimensions. — Stem i to J inch long, T ram. diam., with leaves TS mm. wide; leaves 75 mm. x -5 mm., segments "1 mm., •15 mm., '6 mm. x 4 mm., seg. "1 mm. ; cells '02 mm. ; stipules •2 mm. X "075 mm., seg;. '15 mm., '2 mm. x "1 mm., seii^. "lo mm, ; bracts '8 mm. x '4 mm., seg. "2 mm, ; bracteole "4 mm. x "2 mm., seg. '25 mm. ; perianth "9 mm. x '5 mm, ; pistillidia "125 mm. x •04 mm, ; calyptra "5 mm. x '5 mm. ; capsule -7 mm. x '45 mm. ; pedicel "175 mm. diam.; spores 015 mm.; elaters '15 mm. x •01 mm. ; perigonial bract (not explanate) "45 mm. x ^2 mm., seg. •125 mm. ; antheridia •I mm. x "075 mm. Hab. — On shaded wet rocks ; very rare. 1. St. Just, Cornwall, IF. Ciirnow, 1884. 7. Near Conway, Carnarvonshire, IF. JFilson, 1844. Trefriw, Carnarvonshire, TF. H. Pearson, 1887. IG, Moidart, West Inverness, S. M. Macvicar. I. Cromaglown ; Dunkerron, Dr. Taj/lor. Bantry, Miss Hiitcliins. Killarney, Br. Carringtou, &c, Altadore Glen, Wick- low, J). McArdh'. C. Guernsey, E. D. Marquand. Obs. — Distinguished from the other monoicous species, L. cuspidafa and L. licterophijUa by its habitat, small size and spicate amentula, in addition to other characters. Description of Plate CIV. — Fig. 1. Plants natural size. 2. Plant X 24. Portion of leaf x 290, 4, 5, Stipules x 85. 6. Bract x G4. 7. Bracteole x 85. 8. Sub-bracteole x 85. 9. Upper portion of perianth, explanate x 64. 10. Perigonial bract X 85. (Near Conway, W. Wilson,) 254 HEPATIC^. Genus 23. CLASMATOCOLEA, Spruce. Clasmatocolea, Spi^uce Hep. Am. et And. p. 440 (1885). Plants small, fragile. Primary stem short, sub-erect, densely leaved ; radiculose at the base — sometimes under the terminal flower also ; producing slender, arcuate branches having small and distant leaves, and often rooting at the decurved apex. Leaves alternate, subsecund, ascending, plane or concave, obovate, rotundate, subtruncate or retuse ; branch leaves very often ob- cordato-cuneate. Stipules half the size, heteromorphous, most of them ovato-lanceolate and entire, but others (chiefly the upper ones) bifid. Inflorescence dioicous ; $ bracts larger than the leaves, somewhat similar. Perianths large for the size of the plant, very fragile, obovate or sub-conical, in the upper part obscurely or very slightly trigonous, having a wide 2-4-lobed mouth. Capsule oblongo-globose, in other respects like that of LophocoJea. Clasmatocolea cuneifolia [Hook.), Spruce. Jungermania cuneifolia, Hook. Brit. Jung. t. (i4 (181 G). Mylia cuneifolia, Gr. & B. Nat. Arr. Brit. PI. p. 694 (1821). Leptoscyphus cuneifolia, Mitten in Hook. Journ. Bot. iii. p. 358 (1851). Coleochila cuneifolia, Dum. Hep. Eur. p. 106 (1874). Clasmatocolea cuneifolia, Spruce Hep. Am. et And. p. 440 (1885). Loosely csespitose or creeping, minute, pale olive green to reddish-brown in colour. Stems simple or with one or two very minute shoots, extremely slender, filiform, flexuose, olive-brown, very fragile ; radiculose, rootlets growing in short tufts, white. Leaves approximate or distant, assurgent or erecto-patent, appressed to stem or spreading, slightly decurrent antically, alter- nate, caducous, plane, cuneate with a narrow insertion, entire, truncate or slightly retuse, texture thick, guttulate ; cells minute to small, roundish, lumen clear, containing only few chlorophyl granules, walls thick, reddish-brown or olive colour, angles thickened, no trigones. Stipules distinct, closely appressed to the CLA^SMJ 7V( 'OU'JA . 255 stem, near or connate to the adjacent leaf, broadly subulate, entire or bifid to about J, segments and sinus acute. ? and ^ not seen. Demensions. — Stems \ in. long, with leaves -15 mm. to '2 mm. wide, diam. '05 mm. to '075 mm. ; leaves '2 mm. long x •2 mm. wide at widest part, '\7b mm. x '175 mm., "175 mm. x •125mm., •125 mm. x '1 mm., '1 mm. x '1 mm. ; cells '015 mm. to "02 mm. ; stipules "075 mm. long x '05 mm. broad, "OG mm. x -O:] mm., •04 mm. X 03 mm. Hab. — 1(5. "On birch trees growing on the bark and on FniUania ; on a rock in a ravine creeping among Radiila aqn'degia with Lejcimea ovafa, L. micro-wopica, Plaf/iochila piuicfafa, and P. fri- denticidaia. On the trees it grows in reddish-brown patches of a few inches, or creeping over the Fndlauia.'' Moidart, West Inverness, S. Jl. Macvicnr. I. Growing parasitically on Fndlania Tamarhci near Bantry, Miss Hutchins. Cromaglown, Dr. Taijhr, Dr. Carri/if/ton, Dr. Moore. Killarney, Dr. Carriugton. Connor Hill, Kerry, D. McArdle. Stavenger, Norway, Dr. B. Kaalaas. Extremel}^ rare ; the above are the onl}^ known stations. Obs. — Although no t or % have been observed on this species. Dr. Spruce (Hep. Amaz. et And. p. 440) has no hesitation in referring it to the genus C/asmafocolea founded by him on a species he collected on the Andes. He writes : " These curious little plants come very near Lophocolea, but are well distinguished by the peculiar habit ; the asyurgent leaves, with a plane antical margin — not convexo-deHexed, with the antical margin decurrent and recurved at the base (as in Lophocoh'a); the biform stipules mostly entire, but some bifid. The ])erianth, turgid and indistinctly carinate, is so fragile that the slightest touch breaks off" the short unequal lobes at the wide mouth. I cannot doubt that the Irish Ji()i(/. cuneifolia, Hook. Brit. Jung. t. 64, hitherto known only from sterile specimens, is a true Clasmatocolea. Specimens gathered a few years ago by McArdle are so like the arcuate barren shoots of Cl.frapi/liiiia that, until I compared them closely, I thought them 256 HEPATIC a:. the same species. The Irish plant (like the Andine) has both entire and bifid stipules, and was correctly so described by Nees from original specimens of Miss Hutchins, although Hooker's figure (I.e.) and the Eng. Bot. figure (t. 2700) show only bifid stipules." Mr. Macvicar, in his " Hepaticse of Moidart " (Journ. of Bot. Aug. 1899), says: "Although a small plant, it is not difficult to recognise when growing in patches, there being nothing with its appearance in this district except a small lichen which grows in similar places, and has much the same colour. Single stems creeping among Fndlama can hardly be seen with the naked eye." From flagelliferous shoots of PJagiochila sjnnulosa, Dicks, and JPlagiocMla punctata, Tayl. it maybe distinguished by its assurgent leaves, which are of a different texture, and the presence of stipules along the whole length of the stem. Description of Plate CV. — Fig. 1. Plants natural size. 2. Portion of stem x 85. 3. Ditto, postical view x 85. 4-18. Leaves x 85. 19. Portion of leaf x 290. 20-22. Stipules X 85 (Conner Hill, Kerry, D. McArdle). Genus 24. CHILOSCYPHUS, Bum. Jimgermania, Mich. Nov. pi. gen. p. 8 (1729) ; L. Sp. pi. ed. 1, 2, p. 1131 (1753). Cheilocyphos, Corda in Opiz Nat. p. 631 (1829). Ghiloscyphus, Dum. Syll. Swag. p. 67 (1831). Plants more or less flaccid, pale or dark green. Stems pros- trate, somewhat thick, simple or slightly branched ; radiculose, rootlets proceeding from the base of the stipules. Leaves sub- decurrent, flattened, entire, rotundate, truncate 2 or 3 dentate, rarely bifid. Stipules bifid, laciniate, in some species orbiculate or reniform, margin dentate. Leafy branches postico-lateral, female branches all postical, proceeding from the axils of the stipules, very short. Bracts $ 1-2 pairs, exterior minute, innermost always smaller than the leaves, variously divided. Pistillidia 5-30. Perianth small, obconical or campanulate, only above trigonous, CHILOSGYPHUS. 257 never winged, mouth trilobate, lobes often spinose, postical some- times bifid. Calyptra carnose, below 4-8-stratose, either sub- globose and included in the perianth or clavate and emersed. Capsule on a long pedicel, oblong-globose, about 4-stratose, 4-valved, dividing down to the base. Elaters bispiral, deciduous. Androecia at the middle or apices of postical branches, in the opposite leaved species always on minute-leaved postical amentnla, but in the alternate leaved species at the middle or apices of the stems or branches ; perigonial bracts smaller than the leaves ; antheridia large, situated at the base of the antical lobe. Chiloscyphus polyanthos (Z.), Dmn. Jungermania major, foliis breviorihas et obtusioribus non dentatis, Mich. Nov. pi. gen. p. 8, t. 5, f. 3 (1729). Lichenastrum Trichomanis facie pohjanthemum breve et repens, Dill. Hist. muse. p. 486, t. 70, f. 9 (1741). Jungerviania polyanthos, Linn. Sp. pi. n. 1597 (1753); Hook. Brit. Jung, t G2 (181G). Chiloscyphus polyanthos, Dum. Syll. Jung. Eur., p. G7, t. 1, f. 9 (1831). Cheiloscyphus polyanthos, Corda in Sturm. Deutschl. FL, 19, p. 35, t. 9 (1835). Monoicous, laxly Ciespitose, patches thin, spreading, medium to large in size, green or yellowish-green in colour. Stems creep- ing, lax, flexuose, simple or furcately branched, minute branchlets often produced from the axil of the leaf, antically ; radiculose, rootlets fasciculate, proceeding from the base of the stipules. Leaves alternate, approximate, almost horizontally inserted, antical base decurrent, plane, rotund-quadrate to oblong-quadrate, entire, rotundate or retuse ; texture thin, cells medium size, clear, 5-6- sided, often elongate, walls somewhat thick, no trigones. Stipules small but evident, oblong, bipartite often almost to the base, segments subulate, rarely unidentate on both outer margins. Inflorescence cladocarpous, $ postical on very short branches. Sub-bracts 3, very small, broadly subulate. Bract large, trifid, irregularly dentate. Calyptra exserted, carnose, 4 cells thick near the base, oblong-obovate, mouth irregular, subentire, after protrusion of capsule, whieh is dark brown, oval, composed of R 258 HEPATIC^. 5 laj^ers of cells. Spores pale yellowish-brown, large, almost twice the breadth of the elaters, which are darker brown, long, with about 24 spiral turns. Androecia on short catkins, immediately below the perianth or on main stem ; perigonial bracts but little different from the leaves, except that a small lobe is produced at the antical base, enclosing 1-3 spherical, shortly stipitate antheridia. Fruits April, May, June. Var. pallescens {Jungermania pallescens, Schrad. Syst. Samml. Krypt. Gew. 2, p. 7 (1797); ChUosci/jjhiis jjallescens, Dum. Syll. Jung. p. 67 (1831)) is the small, pale green, neat form, usually fertile, but I can detect no character sufficiently rehable to separate it from the type. On shady rocks. 7. Falls near Barmouth Junction, Merio- nethshire, growing with Juhida Hutchinsice. Var. rivularis, Nees. A large coarse form, of a somewhat thick, greasy texture. 8, 10, 12, 16. Dimensions. — Stems from an inch to several inches long, •2 mm. to "4 mm. in diameter, with leaves 3' mm. to 5* mm. wide; leaves 1*75 mm. x 1'5 mm., 2*5 mm. x 2' mm., cells '025 mm. x •04 mm., '025 mm. x '035 mm., "03 mm. x '03 mm. ; stipules •4 mm. X "175 mm. wide at the base, segments "2 mm., '3 mm., •275 mm. x "125 mm. at the base, segments ^225 mm. ; bract, explanate 1*5 mm. x 175 mm. ; calyptra 3" mm. x 1-5 mm. near apex ; pistillidia ^16 mm. x "06 mm. ; capsule 1*4 mm. x 1*1 mm. ; pedicel "5 mm. diam. ; spores '02 mm. diam. ; elaters '175 mm. x •015 mm. Hab. — Growing in loose or straggling patches in moist and very wet places, on rocks, stones or earth, rarely on rotting wood, often floating in water, where it attains unusual length, stems sometimes 6 to 7 inches long being met with. Common. 1, 3, 5, 7-16, 18c. I. Found generally on the Continent and North America. Obs. — Distinguished from any of the entire leaved JungermanicB by its postical, cladocarpous inflorescence, its large, fleshy, exserted calyptra, the perianth being reduced to a trifid bract; when HARP ANT mm 259 sterile by its horizontally inserted, plane leaves and oblong, deeply bipartite stipules. Description of Plate CVI. — Fig. 1. Plants natural size (Eng. Bot.). 2. Fertile plant magnified (ditto). 3. Portion of stem, antical view x 11 (Bamford, G. A. Holt). 4. Ditto x 11 (Charlesworth, J. Whitehead). 5. Portion of leaf x 290 (ditto). 6. Stipule X 85 (ditto). 7. Ditto x 85 (Bamford, Gr. A. Holt). 8, 9. Sub-bracts magnified (ditto). 10. Bracts x 24 (ditto). 11. Calyptra x 11 (ditto). 12. Pistillidium x 85 (ditto). 13. Spore X 290 (Charlesworth, J. Whitehead). 14. Perigonial bract magnified (after Hooker). Genus 25. HARPANTHUS, Nees. Jmigermania, Web & Mohr. Bot. Taschenb. p. 408 (1807). Harjmnthus, Nees, Nat. Eur. Lab. 2, p. 351 (1836). Pleuraiithe, Tayl. in Hook. Lond. Journ. Bot. 5, p. 282 (1840). Dioicous, plants small, caespitose ; stems decumbent, radicu- lose, stipulate. Leaves succubous, ovate^ emarginate or bilobed, semi- vertically imbricated, sub-decurrent, flattened, secund. Sti- pules numerous, lanceolate, connate with the adjoining leaves, free margin reflexed, unidentate at the base. Fertile shoots very short, postical (issuing from the axils of the stipules), at length sublateral. Bracts small, 1 or 2 pairs with interposed bracteoles. Perianth exserted, fusiform, terete, lower half thickened, mouth contracted, 3-4-fid, segments unequal, entire. Calyptra fleshy, adherent for f its length with the perianth, free only near to the apex. Capsule 4-valved, coriaceous. Elaters bispiral. Male plant more slender; perigonial bracts terminal, more concave, enclosing 1-2 oval antheridia. 1. Harpanthus scutatus [IF. et M.), Spruce. Jungermania scutatus, Web. et Mohr. Bot. Taschenb, p. 408 (1807). Jungermania stipulacea, Hook. Brit. Jung. t. 41 (181G); Sm. Eng. Bot. t. 2538. Lophozia scutata, Dum. Recueil, p. 17 (1835). Harpanthus scutatus, Spruce, Trans. Bot. Soc. Edin. Ill, p. 2G9 (1849). Dioicous, loosely coespitose, of a pale green colour. Stems 260 HEPATIC^. ascending, simple or slightly branched ; radiculose, rootlets plentiful, close, white. Leaves imbricate or approximate, patent- divergent, roundish-ovate or ovate, emarginate to about J-, sinus wide, obtuse, rarely acute, segments acuminate, often somewhat connivent ; cells roundish-quadrate, from smallish to medium size, walls somewhat thick but delicate, trigones distinct. Stipules almost as long as the leaves, somewhat patulous, with an arcuate incurved apex, broadly lanceolate or lanceolate. Female inflores- cence on short postical branches ; third sub-bracts small, bifid or trifid, irregular in shape, oval, subquadrate or subcuneate ; second series of sub-bracts larger, oval ; bracts large, ovate or subquadrate, irregular bifid or trifid to about J, sinus acute or obtusate, seg- ments acute. Bracteole pyriform. Perianth projecting a little beyond the bracts, oblong- oval, acute, obtusely trigonous, composed of a single layer of cells, about 60 near the middle, near the base composed of two layers, mouth constricted, 6-dentate, teeth small. Calyptra mitriform, rostellate, thickened near the base. Pistillidia few, 4, 5, small. Capsule oval, reddish-brown. Spores brown, as broad as the reddish-brown elaters. Androecia on the middle or apex of the main stem or branches, perigonial bracts 3-8 pairs, imbricate, complicate-concave, ventricose, enclosing 1, 2, small, oval antheridia. Fruits April, May. Dimensions. — Stems from |- to 1 inch long, diameter •! mm., with leaves 1* mm. wide; leaves "45 mm. x '4 mm., segments •1 mm., '5 mm. x '4 mm., seg. '1 mm. ; cells '03 mm. x '025 mm., •04 mm. X '03 mm., '03 mm. x "03 mm., '04 mm. x '025 mm.; stipules -3 mm. x 'l mm., "35 mm. x -125 mm. ; sub-bracts '3 mm. X '25 mm., segments "1 mm., '375 mm. x '225 mm., seg. '1 mm. ; bracts '6 mm. x '35 mm., segments '25 mm., -45 mm. x "325 mm., seg. "175 mm. ; perianth 1' mm. x "4 mm. ; pistillidia 'lb mm. x "04 mm.; perigonial bracts '35 mm. x "375 mm.; antheridia •1 mm. X "075 mm. Hab. — Growing on shady banks, rocks and stones, or on rotting wood. Eare. 3. Balcombe, Tunbridge Wells, Surrey, George E. Davies ; BARPANTHUS. 2()I on rocks in Tlung-ersliall Wood, Yieni,Jcnn<'r. 7. Near Llanberis, Cariiarvonsliire, Jf^. IFlhoit ; Tyn-y-Groes, Merionethshire, C. J. Wi/d. 10. Arncliffe Wood, Eskdale, Yorks., Dr. Spruce, M. B. Slater, W. 11. P. ; Bolton Woods, Yorks., Br. Carrinyton. 15. George Don. 16. Moidart, West Inverness, S. M. Macvicar. 1. Bantry, Miss llutchins; Loch Bray, Dr. Taylor; Killarney, Dr. Carrington; O'Sullivan's Cascade, Prof. Lindbercj ; Cromaglown, Stewart 8f Holt. Found on the Continent and in North America. Obs. — When fertile this is at once distiuL,mished from others by its postical inflorescence, with the exception of Harpanthm Flotoioii, which is a larger plant, with more orbiculate leaves, smaller sinus, and cells of a different shape. It differs from any of the forms of Jung, hanlriensis, for which it might be mistaken when barren, in its paler green colour and larger arcuate stipules. Description of Plate CVII.— Fi.g. 1. Plants natural size. 2. Portion of stem, antical view x 31 (Killarney, Dr. Carrington). 3-5. Leaves x 31 (ditto). 6. Leaf x G4 (ditto). 7. Portion of leaf X 290. 8, 9. Stipules x 04 (ditto). 10. Stipule x 85 (ditto). 11, 12. Sub-bracts x 64 (Canada, Macoun). 13, 14. Bracts x 64 (ditto). 15. Bracteole x 64 (ditto). 16. Perianth X 31 (ditto). 17. Portion of the mouth of perianth x 31 (ditto). 18. Pistillidia x 85 (ditto). 19, 20. Perigonial bracts X 64 (ditto). 21. Antheridium x 85 (ditto). 2. Harpanthus Flotowii, Nees. Jiimjerviania Flotoviana, Nees in Diar. Bot. Ratisb. 11, n. 2(i, p. 108 (1833). Jiuujer mania convoluta, Hiiben. Hep. Germ. p. 00, n. 7 (1834). J imyermania Uai'tmanni, Theden. Muse. Suec. Exsicc. v. 1, d. 138, {i/envK/iiieus. Ifarpanthus Flotovianus, Nees, Nat. Eur. Leb. 11, p. 353 (I83racts X 24 (Killarney, Dr. Carrington). 10. Perianth x 24 (ditto). 11. Pistillidium x 85 (ditto). 6. Plagiochila tridenticulata, Taj/l. Jungermania sjnnidosa, var. tridenticulata, Hook. Brit. Jung. p. 9, t. xiv. ft". 9, 10 (1816)?. Radida cornicidata, Dum. Syll. Jung. p. 13 (1831)?. Plagiochila cornicidata, Dvim. Recueil, p. 15 (1835)?. Plagiochila tridenticidafa, Dura. Recueil, p. 15 (1835)?. Plagiochila tridenticulata, Taylor in G.L.IS". Syn. Hep. p. 26 (1844). Dioicous, densely csespitose or rarely creeping, small, of a dark indigo-green colour, when dry almost black. Stems filiform, flexuose, dark brown, erect, ramose, branches lateral ; radiculose, rootlets dull white, ascending almost to apex of the stem. Leaves obliquely inserted, patent to divergent-patent, 60° (fronj 50° to 70°), distant or approximate, alternate, oblong-ovate or cuneate- ovate, from a narrow base, caducous, 2-3-lobulate, bifid or less often trifid to about ^ or ^, sinus rounded or acute, segments acute or apiculate, margin entire ; texture somewhat firm, cells small, roundish, lumen clear, walls thick, angles thickened and pale brown, no trigones. Stipules wanting or rudimentary. Perigonial bracts middle or end of the stem, 3-5 pairs, closely imbricated, more erect, semi-amplexicaul, ventricose at the base, ovate, trifid ; antheridia spherical. Female plants not seen. 1' LAG IOC in LA. 281 DiMKNSioNs. — Stems J to 1 incli lon^-, diam. "L mm., with leaves T mm. wide; leaves '8 mm. x '4 mm., segments '3 mm., "7 mm. X '3 mm., seg. 2 mm., "05 mm. x "4 mm., seg. "3 mm.; cells '03 mm., "02 mm., '0175 mm., -015 mm. ; cell-walls '01 mm. ; perigonial bra(;ts 7 mm. x *55 mm., seg. '2 mm. ; autlieridia •1 mm. H.\B. — Growing on rocks or trees in shady places. Rare. 7. Tyn-y-Groes, Alerionethshire, C. J. Wild, G. A. Holt, Dr. Car- rington, W. 11. P. ; Dolgelly, Merionethshire, JF. Wilson, E. George; Aber, Carnarvonshire, W. Wikon ; Lhinberis, Carnarvonshire, W. II P.- 12. Keswick, C. Lyell, 1812 ; Br. Carriu(jUni $• W. II. P., 1884. 13. New Galloway, /. McAndreio. IG. Moidart, West Inverness, S. M. Macvlcar 8f W. II. P. ; Scotland, W. Wilson. South of Ireland, I)r. Taylor, and others. Bantry, Glengariti'e, Killarney, &c. The Play, tridcniicidata, Tayl., reported as collected in Corn- wall by the late Wra. Curnow, belongs to another species. Obs. — Distinguished from Playiochilu i^anctata, Tayl., by its smaller size, compact habit, darker colour, oblong-ovate, cuneate leaves, which are only bifid or trifid, and which when dry easily fall off, leaving only the naked stem. When dry it turns almost black, and is then at once distin- guished from any of the other PlayiochilcE. It is very doubtful whether Hooker's J/fny. spimdo-sa, var. fridoilicidata, is this species ; the figures given in his " lirit. Jung." certainly do not represent it. Playiocltila Iridenficidata, Dum. Recueil 1835, and Playiocliila corniculata, Dum. Recueil 1835, are both founded on Hooker's Jiniy. .splniflom, var. tridcniicidata, wdiicli probaljly is not the same as Taylor's species. Dr. Spruce remarked in a letter, " the specific name is not a very happy one, for * denticulata' means only rudi- mentarily toothed, whereas the leaves of tliis plant are decidedly 2-3-lobulate." Dkscuiption of Platk CXVIII. — Fig. 1. Plants natural size. 2. Plant X 24(Tyn-y-Groes, North Wales, W. II. P.). 3. Portion of stem X 24 (ditto). 4. Leaf x )i'o (ditto). 5. Portion of 282 HEFATICjS. leaf X 290 (ditto). 6, 7. Perigonial bracts x 24 (ditto). 8. Antheridium x 85 (ditto). 7. Plagiochila exigua, Tai/1. Jungermania exigua, Tayl. in Trans, Bot. Soc. Ed. 1, p. 179 (1843). Plagiochila exigua, Tayl. in Lond. Journ. Bot. v. p. 264, n. 14 (1846). Csespitose or creeping upon other larger species, very small, of a light green to olive-brown colour. Stems ascending or procum- bent, slightly radiculose up to the apex, rootlets dull white, pro- ceeding from the postical side of the stem below the stipules, in small fascicules ; filiform, cortical cells oblong-quadrate, about 10. Leaves obliquely inserted, slightly decurrent antically, patent to patent-divergent, 55°, alternate, remote, but forming small capitula at the apex of the stem, caducous, roundish obovate, obovate, or cuneate, from a narrow base, margin quite entire, bifid from about ^ to J, sinus rounded, segments acute or obtusate ; texture delicate, cells very small to small, subquadrate, lumen clear, walls thick, angles thickened, no trigones. Stipules evident, persistent, subu- late, entire, 5-7 single cells long. Neither male nor female yet seen. Dimensions. — Stems J to |^ inch long, diam. -1 mm., with leaves 75 mm. wide ; leaves "5 mm. x "3 mm., segments "175 mm., •4 mm. X 'S mm., seg. '15 mm., "35 mm. x -25 mm., seg. '1 mm., "275 mm. x '175 mm., seg. "1 ram., '5 mm. x "45 mm., seg. '175 mm., -4 mm. x -3 mm., seg. '15 mm., '35 mm. x "25 mm., seg. '1 mm. ; cells "025 mm. x -0175 mm., '02 mm. x -0175 mm., •0175 mm. x -015 mm. ; stipules -175 mm. x '03 mm. wide at the base. Hab. — Epiphytic on tufts of Frullania, Badula, &c. ; about the bases of moss-covered trees at Cromaglown and Killarney, Br. Taylor. On wet rocks, Knockavoila and Dunkerron, Br. Taylor. Tore Mountain, on Frullania Tamarisci, Br. Taylor. Killarney, Br. Carrington. Glengariffe, Br. Carrinyton. O'Sulli van's Cas- cade and Grlena, Br. Moore. Adrigole, near Glengariffe, Bev. C. H. Binstead, J'LAdlOCHlLA. 283 Extremely rare, no otlier stations known. Obs. — This very rare and minute species, about which many botanists have had their doubts, has generally been referred to the genus Plnf/iochila ; the constant presence of stipules along the length of the whole stem, which remain, whilst the leaves fall oif, and its peculiar habit, lead me to conjecture that it may belong to some other genus, but until more perfect plants are found I place it with the Plagiochila. Shortly before the lamented death of Dr. Spruce, I had some correspondence with him about this plant. He had specimens of what I considered small flagelliferous forms oi Flagiochila punctata under this name, and his opinion was, that my plant (which is an original specimen from Taylor) and the one collected by Dr. Carrington was a new species. As the specimen I have fairly agrees with Taylor's description, I have no desire to further complicate this subject, and I name it P. exigua, Tayl., and refer other small forms which do not agree with Taylor's specimen to either P. punctata or P. spinulosa. Amongst these are to be placed specimens published in C. & P. Hep. Brit. Ex. n. 15, which were doubtfully referred to P. exigua. Dr. Carrington described and figured P. exigua with stipules sometimes acutely bidentate ; in the fine specimens from Dr. Taylor I have only been able to observe long, entire subulate ones. Small, flagelliferous forms of Plag. punctata or Plag. spinulosa are likely to be mistaken for it, but they are at once distinguished by the absence of the long, subulate stipules. The leaves having a very narrow base become easily detached from the stem. A similar feature also has Plag. tridenticulata, Tayl., which is a more robust species, of a darker colour, with longer, sometimes trifid leaves, cells larger, and in this also the numerous long stipules are absent. Description of Plate CXIX. — Fig. 1. Plants natural size. 2. Stem, antical view x 24. 3-9. Leaves x 85. 10. Portion of leaf X 290. 11-13. Stipules x 85. (Original specimens from Dr. Taylor, Cromaglowu, Ireland.) 284 HEPATIC^. Genus 29. LIOCHL-ffiNA, Nees. Jungermania, L. Sp. pi. 1 ed., 2, p. 1597 (1753) ; Hook. Brit. Jung. t. 28 (1816). Liochlcena, Nees in G.L.N. Syn. Hep. p. 150 (1844). Aplozia, Dum. Hep. Eur. p. 58 (1874). Stems prostrate, with rootlets very long, pale or reddish, slightly (laterally) branched, rarely flagelliferous. Leaves ovate- oblong, sublingulate, apex rotundate or retuse, very rarely sub- emarginate, slightly decurved, antical base shortly decurrent, very slightly recurved ; reticulation somewhat lax, subopaque. Stipules absent. Inflorescence dioicous or paroicous. Female flowers terminal. Bracts $ 1-2 pairs, similar to the leaves only broader. Pistillidia 5-12. Perianth at first pyriform, afterwards cylindrical, incurved, eplicate, apex abruptly rotundato-constricted, mouth minutely ciliolate. Capsule highly exserted, dividing down to the base into 4 valves, composed of two layers, inner layer with semi-annular fibres. Elaters attached to the middle of the valves, bispiral. Spores globose, minute, as broad as the elaters. Perigonial bracts similar to the leaves, antical base turgid. Antheridia 2-4. liiochlaena lanceolata (Z.), JVces. Jungermania 2yalust7'is minima rejiensfoliis suhrotundis densissimis Icete virentihus, Mich. Nov. pi. gen., p. 8, tab. 5, fig. 6 (1729). Jungermania lanceolata, Linn. Sp. pi. 1 ed., 1597 (l75o) ; Hook. Brit. Jung. t. 28 (1816). Liochloina lanceolata, Nees in G.L.N. Syn. Hep. p. 150 (1844). Paroicous or monoicous, ^ flowers usually immediately below the ? bracts, sometimes on proper branches which proceed from below the perianth, densely ceespitose, small, pale or dark green to a greenish-brown in colour. Stems dark brown, 30 cortical cells, slightly darker than the inner ones, of equal size, about 12 in diameter, inner very clear, vacuous, walls thin but firm, dark ; procumbent or suberect, simple or slightly ramose, branches lateral, produced at right angles or ascending ; radiculose, rootlets LIOCHLJ^jNA. 285 brownish-white, strong, long, ascending to apex of the stem. Leaves close, horizontal or patent-divergent, obliquely inserted, often projecting anticnlly from the stem, slightly decurrent at the antical and postical base, oblong-oval or oblong-obovate, entire ; texture somewhat thin, epidermis smooth, cells roundish, near the base elongate, lumen pellucid, with few chlorophyllose granules, walls somewhat thin, two bands, trigones distinct. No stipules. Bracts similar to leaves only a little larger and a trifle broader. Bracteole absent. Perianth terminal, erect, somewhat curved, composed of one layer of cells, similar to leaf-cells, but more elongate, clavate-cylindrical, smooth, apex depressed, plane, mouth very small, contracted, conical, cells delicate, hyaline, witli thin walls, elongate-quadrate, about 20 rows, margin crenulate. Pistil- lidia few, about 7, surrounding base of immature capsule. Calyptra delicate, composed of one layer of cells. Capsule oval, spores yellovvisli-brown. Perigonial bracts slightly lobulate, saccate ; antheridia 2-3, spherical, bearers short, composed of 4 rows of cells. Fruits spring. Dimensions. — Stems from ^ io ^ inch long, "4 mm. diam. ; leaves 2- mm. long x 1*2 mm. broad, 1"5 mm. x l* mm., 1-4 mm. X '8 mm. ; cells "05 mm. x "03 mm., '04 mm. x -03 mm., '03 mm. x "03 mm., '02 mm. x "02 mm. ; trigones "01 mm. ; bracts 2" mm. x 1'25 mm., 1'75 mm. x 1*25 mm. ; perianth 3"2 mm. long x 1'5 mm. broad, 2-5 mm. x 1-25 mm. ; pistillidia IQ mm. long x '04 mm. broad ; antheridia '2 mm. diam. Hab. — Grrows in small dense patches in woods or moist shady places, or on banks of streams on claj^ey or gravelly soil, on stones and rocks or at the base of trees. Very rare. 2. Tunbridge Wells, Sussex, IF. Borrer, E. M. Holmes. 10. Cronkley Gill, Eskdale, Yorks, Dr. Spruce; Arncliffe Wood, Eskdale, Yorks, J/. B. Slater. 12. Mill Beck Stock, Rayrigg Wood, Windermere, Westmorland, G. Stabler. Found on the Continent and in North America. Obs. — Hooker figured and described this species from Con- tinental specimens; it was first described from British specimens 286 HEPATIC^. iii"Eng. Bot. Suppl."tab. 2947 (Oct. 1, 1848) by Dr. Spruce, who discovered it in Eskdale, December 1842. Distinguished from Jun^. pumila, With, and Jung. .sjjJiarocarjja, Hook., which are both paroicous, by the insertion and shape of leaves and clavate perianth ; some forms of Jun(/. rijKiria, Tayl. might be taken for lanceolafa, but riparia is always dioicous. Description of Plate CXX. — Fig. 1. Plants natural size. '2. Portion of fertile stem x 11 (C. & P. n. 170). 3, 4. Leaves X 16 (ditto). 5-7. Leaves x 24 (Canada, Macoun). 8. Portion of leaf X 290 (C. & P. n. 170). 9, 10. Bracts x 24 (Canada, Macoun). 11. Cross-section of perianth near the middle x 24 (ditto). 12. Apex of perianth x 16 (ditto). 13. Portion of the mouth of the perianth x 85 (ditto), 14. Pistilliclium x 85 (ditto). 15,16. Perigonial bracts x 16 (ditto). 17. Antheridium X 85 (ditto). aenus 30. JAMESONIELLA, Spruce. Jamesoniella, Spruce, Journ. of Bot. (1876). Large, showy, csespitose, rosy or pale. Stems often suberect, sparingly and fastigiately branched, flagelliferous, circinnate ; branches postico-lateral. Leaves large, alternate, succubous, laterally appressed, decumbent, remarkably decurrent at the base, insertion oblique, very concave, quite entire or very distantly ciliate or denticulate ; cells small or minute, semipellucid, beauti- fully thickened at the angles. Stipules none or very rare and minute, except near the involucre. Inflorescence dioicous ; ? flowers terminal, with innovations below ; bracts and interposed bracteole much laciniated. Perianth more or less projecting beyond the bracts, towards the apex 6-10-plicate, mouth scarcely denticulate. Capsule, &c., not seen. $ cauline, very often terminal, perigonial bracts equalling the leaves, 2-3 androus. JAMESONIELLA. 287 Jamesoniella Carringtoni {Balf\ Spruce. Adelanthus Carringtoni, Balf. MSS. Carr. in Trans. Bot. Soc. Edin. p. 378 (1870). Nardia Carringtoni, Carr. Brit. Hep. p. 27 (1874-0). N'ardia comjjreasa, var. Carringtoni, Lindb. in Hep. Hibern. in Act. S )c. Sc, Fenn. (1875). Jamesoniella Caii'ingioni (Balf.), Spruce, Journ. of Bot. (187G). Dioicous, csespitose, flagelliferous, largish to very large, pale or olive green in colour. Stems procumbent or suberect, simple or innovantly branched, branches postical-lateral, rootlets slender, few, whitish, fasciculate ; firm, flexuose, brownish colour, apex more or less circinnate. Leaves alternate, succubous, laterally appressed, obliquely inserted, secund, very concave, imbricate, regular, lower ones minute, distant, subrotund or cuneate ; obliquely orbicular or reniform, antical base very decurrent, postical semicordate, ampliate, abruptly and very narrowly decurrent, margin entire or furnished with few large teeth, 1-6 ; epidermis polished, texture firm, subpellucid, little altered when dry, cells from very small to medium size, roundish, walls thick, trigones distinct, angles thickened. Stipules absent or rudimentary, towards apex of stem or branches, very minute, sublinear, muchnarrow^er than the stem, appressed ; to about the middle or base unequally or equally divided, segments subulate. Androecia at apex of stem or branch or on lateral branches, spicate, perigonial bracts 6-10 pairs, a little smaller, turgid, antical lobule narrow ; antheridia at base solitary (2-3, Dr. C), spherical, mixed with paraphyses. Dimensions. — Stems 2 to 4 inches long, diam. "2 mm., with leaves 1*5 mm. to 175 mm. wide; leaves 1'5 mm, high from the decurrent base x 1"4 mm. broad, 1*3 ram. x 1*3 mm., 1"2 mm. X 1*4 mm.; teeth "125 mm. long; cells "035 mm., '03 mm., '0225 mm., "02 mm. ; cell- walls '0075 mm. ; trigones "01 mm. ; stipule 2 mm. long x '05 mm. broad at base; antheridia '14 mm. X -12 mm. Hab. — Growing on wet rocks and damp rocky ledges in alpine districts in Scotland. Very rare. 288 HEPATIC^. 15. On rocks above Loch Avon, Dr. GreviUe, 1830; A. Croall 1850; G. Stabler. Ben Lawers, Br. Stirfon, 1866. Ben Voilich, Dr. Stirto7i, 1869. Ben Laoigli, C. J. Wild Sf G. A. Holt, 1880. 16. Loch Maree, C. Howie, 1867. Glen Finnan, Dr. Carrington. Moidart, West Inverness, S. M. Macmcar, 1898 ; 8. M. Macvicar 8r IF. H P., 1899. Faroe Isles. Obs. — This very fine, rare, and beautiful species was first recognised as distinct by Dr. Stirton, who named it in his MSS. Alicularia viridis. Prof. Balfour publislied it as Adelaniluis Carringtoni in the Trans. Bot. Soc. Edin. 1870, and I am glad that the memory of my dear friend sliould be associated with such an unique British species. From Nardia covipressa (Hook.), to which it has even been relegated, as a variety, by Prof. Lindberg, it is at once and readily distinguished by its beautiful cell structure. In N. compressa the cells are more quadrate, with thin walls and small trigones ; the leaves also are nearly plane, appressed to each other, texture more delicate, epidermis not polished, and it has a terminal immersed perianth. Description of Plate CXXI. — Fig. 1. Plants natural size 2. Portion of stem x 16 (C. & P. n. 233, and following figures). 3, 4. Leaves x 24. 5. Leaf with teeth x 24. 6. Leaf x 16. 7. Portion of leaf margin x 85. 8. Portion of leaf x 290. 9. Stipule X 85. 10. Antheridium. Genus 31. JUNGERMANIA, L. L. Sp. pi. 1131 (1753); Dum. Recueil, p. 16 (1835). Plants medium size, rarely very large. Stems simple or furcate, in a few species repeatedly dichotomous, in several vaguely ramose, branches (few) lateral, proceeding from the postical angle of the leaves, true postical branches none or very rare, but rooting fliagella present in some species. Rootlets usually pale, in the prostrate species plentiful, in the caespitoso- JUNGERMANIA. 289 erect species few or almost wanting. Leaves succubous, rarely subtransverse, patent, assurgent or laterally accumbent, mode- rately broad, in several species entire, in others bilobed, in a few 3-5 lobate or dentate ; margin plane or incurved, never recurved, almost always quite entire, rarely denticulate ; cells medium size, rarely small. Stipules usually absent, where present more or less small, entire or bifid, rarely subquadrifid. Inflorescence dioicous or paroicous; female flowers terminal, fertile rarely innovant, sterile frequently present on subfloral innovations. Bracts $, rarely more than 2-pairs, somewhat similar to the leaves or more incised, free from the perianth, sometimes connate with themselves. Pistillidia numerous (8-70). Perianth usually emersed, slightly compressed laterally, rarely all ecarinate, almost always (at least above) 3-10-carinate, smooth, rarely rough, never winged; mouth usually small, quite entire or denticulate, rarel}^ ciliate, sometimes tubular, sometimes at first closed, afterwards rupturing into lobes, very rarely bursting on one side. Calyptra pja'iform or oval, delicate, or only towards the base slightly carnose, where are the free sterile pistillidia. Capsule highly (rarely shortly) exserted, globose or oblong, rarely cylindrical, 2-5 layers of cells, 4-valved, valves rarely bifid. Elaters bispiral. Spores minute, smooth or scaberulous. Subgenus I. APLOZIA, Dim. Jungermcmia, sect. Ajilozia, Dum. Syll. Jung. Eur. p. 47 (1831). Aplozia, Dum. Hep. Eur. p. 55 (1S74); Schiftner, Engl, and Prantl. Pfl. Fam. p. 82 (1893). Dioicous, paroicous or monoicous, caespitose, small to medium in size, green, brown or reddish-brown in colour. Stems creeping or erect, determinate, simple or slightly branched, often with one or two innovations proceeding from below the perianth ; usually radiculose. Leaves succubous, orbicular or ovate, entire or rarely retuse. Stipules absent (except in J. nuf/rnnialis). Bracts simihir to leaves (except in /. anf/zmnnli-s). Perianth terminal, 3-5, very rarely 6 and 7 keeled. 290 HEFATIC^E. 1. Jungermania cordifolia, Hooker. Jungermania cordifolia, Hook. Brit. Jung. t. 32 (1810). Jungermania tersa, Nees Nat. Eur. Leb. 1, p. 329 (1833), fide letter from Dr. Gottsche to Dr. Spruce. A]jilozia cordifolia, Dum. Hep. Eur. p. 50 (1874). Dioicous, laxly casspitose, flag-ella absent, large, dark olive- green in colour, when young pa]e recldish-brown. Stems simple or dicliotomously branched, branches lateral-subpostical {^rising Irum the postical aspect of the leaf axil, often several (3-6) arising from the base of old bracts, suberect or floating, lax, oval on cross- section, 12 cells X 20, cortical cells 40, similar to the inner, hyaline, walls thin, firm, dark-brown ; rootlets absent. Leaves transversely inserted, succubous, decurrent antically, slightly so postically, suberect, subinibricate or approximate, accrescent, sub- opposite near apex, close, lower smaller, alternate, distant, cordate or ovate-oblong, sheathing, amplexicaul ; texture flaccid; cells smallish to medium in size, 4-, 5- and 6-sided, lumen with numerous chlorophyl granules, walls firm, thick, reddish-brown, no trigones or thickened angles. No stipules. Bracts similar to the upper leaves, sub-vertical, ventricose-concave. No bracteole. Perianth almost hidden by the bracts, projecting only about J, fusiform, acute, very slightly plicate near the apex, near the base composed of 3 layers of cells, lower half of 2, upper half of 1 only, interior layer of cells convex ; mouth contracted, very small, laciniate, lacinise entire. Perigonial bracts swollen near the base ; antheridia two in each bract, oval. Dimensions. — Stems from 1 to 4 inches long, "3 mm. to "4 mm. diam. ; leaves 2-25 mm. long x 2-25 mm. broad, 2" mm. x 175 mm., 1-75 mm. x 1'3 mm., 1'6 mm. x 1-3 mm., 1*5 mm. x 1-25 mm.; cells '035 mm. x -025 mm., -025 mm., -02 mm.; perianth 4*5 mm. long x 1-25 mm. broad; young perianth 2'25 mm. x 75 mm. Hab. — Growing in large patches on rocks and earth by the side of mountain rills, often submerged in the water. Somewhat rare. JUNGERMANIA. 291 7. Cwm Idwal, Carnarvonshire, IF. 11. P. 9. Longridge Fell, J. A. Wheldon ; Udale, Jnehh?^ S^' A. Wihon ; EasegiU Kirk, West Lane., A. JFikojL 10. Hovvgill Fells, 7Jr. F. A. Lcch. White- side, Br. F. A. Lce.s. Ingleboro, Joint Nowell. Ingleton, Dr. Carriiigton. Clapliam, W. West. Penyghent, W. West. Middles- more, W. West. Teesdale, Br. Spruce. 12. Plentiful in small streamlets among the mountains in Westmorland, Nan Bield, Mardale ; Hill Bell ; Merke Side ; Middleton Fell; Barbon Fell; Long Sleddale, Westmorland, G. Staljler. 13. Dalveen Scott^ Criffel, Cndckshank. Blackhope Glen, Nichol. 15. Ben Lawers, G. A. Holt. Forfar, C. Li/eU. Mael Tarmachan, C. J. Wild. I. Bather rare in Ireland. Mangerton, in the stream from the Punch Bowl, Br. Taijlor. Coomashana Lake, Br. Car- rington. Brandon, Br. B. Moore. Maghanabo Glen, Co. Kerry, D. McArdle. Cushinden, Co. Antrim, Br. B. Moore. Found on the Continent and in North America. Obs. — This is very distinct, and cannot well be confused with any of the allied species ; small forms of it have been mistaken for Jtfn(/. riparia, Tayl. (and Juntj. tersa, Nees, which, is a mixture of both), but it is distinguished by the absence of rootlets and flagella, the more distinctly cordate-shaped leaves, the transverse insertion of them, and the almost hidden fusiform perianth. When fresh it smells strongly like Baucns Carota. Description of Plate CXXII. — Figs. 1 and 2. Plants natural size. 3. Fertile stem x 16 (C. & P. 27). 4-7. Leaves X 16 (ditto). 8. Leaf x 11 (Penyghent, West.). 9. Portion of leaf X 290 (C. & P. 27). 10. Perianth x 11 (Penyghent, West.). 11. Cross-section of perianth from near the middle x 11 (ditto). 292 HEPATIC^. 2. Jungermania pumila, Withering. Ju7igernmnia jwniila, Wither. Brit. Fl. p. 866 (1776); Hook. Brit. Jung. t. 17 (1816). Aplozia pumila, Dum. Hep. Eur. p. 50 (1874). Paroicous, csespitose, small olive-green colour. Stem simple or branched, creeping or suberect, no flagella ; radiculose, rootlets thick, frequent; branches lateral-subpostical, arising from the postical side of the leaf- axil, or proceeding from the base of perianth. Leaves succubous, upper imbricate, sheathing com- plicate, concave, erecto-patent or patent, lower alternate, spreading, approximate, obliquely inserted, horizontal or patent-divergent, ovate, oval or oblong-oval ; texture delicate, epidermis smooth ; cells 4-, 5- and 6-sided, quadrate or elongate, walls delicate, com- posed of two bands, no trigones or thickened angles. 'No stipules. Bracts similar to the upper leaves, only larger and swollen at the base, where the antheridia are. No bracteole. Perianth pro- jecting about § beyond the bracts, composed of a single layer of cells, about 120 round the middle, fusiform, acute, mouth very small with few delicate teeth or subentire, upper portion antically obtusely 2-plicate with a narrow fold between the folds. Calyptra extremely delicate. Capsule oval, dark brown, spores pale golden- brown, elaters the same, bispiral, 12 turns of the spiral. Peri- gonial bracts immediately below the perianth, 2 or 3 pairs ; anthe- ridia oval, 1 or 2 at the swollen base. Fruits May, June. Dimensions. — Stems J to |- inch long, diam. -1 mm, to 15 mm. ; leaves 1* mm. long x '75 mm. broad, '9 mm. x •? mm., -9 mm. x "6 mm,, '8 mm. x '55 mm. ; cells "05 mm. x 02 mm., "04 mm. x *025 mm., '035 mm. x "03 mm., -03 mm. x '03 mm., '03 mm. x •02 mm. ; bracts 1'5 mm. long x 1' mm. broad, 1'5 mm. x "8 mm., 1*25 mm. x 75 mm. ; perianth 2*25 ram. long x '6 mm. broad, 1"75 mm. X '65 mm. ; pedicel "l mm. thick ; capsule '75 mm. x '5 mm. ; valves '75 mm. x '25 mm. ; spores "025 mm. Hab. — Growing on wet rocks and earth by streams. Some- what rare. JUNGEBMANIA. 293 1. Helston, Reo. C. A. Johns. Riverside, Truro, IF. Curnow. 3. Hungershall Rocks, Kent, Jenner. 6, 7. Tyn-y-Groes, Dr. Carrington 8f JF. H. P. Barmouth, Merionethshire, W. 11. P. Pass of Llanberis, Carnarvonsliire, /. (kish Sf IF. II. P. Cwm Idwal, Carnarvonshire, GrtJ/Uks. 10. Howgill, JF. TFest. Dent, G. Stabler. Clapham, Dr. Car ring ton. Bolton Woods, Dr. Car- rington ^' IF. LI. P. Todmorden, A. Stansjield. Studley, Dr. Hooker. 12. On damp rocks by River Kent, Kentmere ; Black Crag ; by Biver Lune, Sedbergh ; Groom Grhyll ; Barbon Fell, ^estmov\^x\&, G. Stabler. 13. Glenlee Grlen, &c., New Galloway, /. Mc Andrew. Moffat, Dumfries, Nichol. Dalscairth, Dumfries, Crtdchhank. 15. Killin, Perthshire, G. A. Holt. 16. Very common on rocks by the side of streams ; frequent on wet banks, Moidart, West Inverness, S. M. Macvicar. I. On rocks at the side of streams and rivers, not rare. Dr. Taylor. Glen near the Hunting Tower, Cromaglown, Dr. Car- ringfon. Connor Hill, Brandon, Co. Kerry, Dr. D. Moore. Loch Bray, Co. Wicklow, Dr. D. Moore. Grlenad, Co. Leitrim, Dr. D. Moore. Ballyhaise Wood, Co. Cavan, D. McArdle. Found on the Continent and in North America. Obs. — Distinguished from small forms oi Jtingennania riparla, Tayl., with which it is most commonly mistaken, by its paroicous inflorescence, absence of flagella, and its narrower perianth. Description of Plate CXXIII. — Fig. 1. Plants natural size. 2. Fertile stem x 24. 3. Portion of stem x 24. 4-G. Leaves X 24. 7. Portion of leaf x 290. 8. Bract x 24. Perianth with bracts and upper leaves x 24 (Tyn-y-(iroes, North Wales, Dr. Carrington & W. H. P.). 294 HEPATIC^E. 3. Jungermania riparia, Taylor. Jimgermania riparia, Tayl. Ann. and Mag. Nat. ' Hist. August (1843). Spruce in Phyt. March (1843). Jimgermania tristis, Nees Nat. Eur. Leb. 11, p. 461 (1836). Jimgermania j)umila, var. notha. G. et R. n, 396. Aplozia tristis, Dum, Hep. Eur, p. 63 (1874). Aplozia rijmria, Dum, Hep. Eur. p. 65 (1874). Dioicous, csespitose, flagelliferous, small, pale to clarK olive- green colour. Stems creeping or suberect, simple or branched, flagella frequent, postical, branches lateral-subpostical, arising from the postical side of the leaf axil, or frequently from base of perianth ; radiculose, rootlets ascending to apex of stem, whitish, somewhat thick. Leaves imbricate or approximate, alternate^ obliquely inserted, horizontal, spreading, broadly ovate, oval, oblong-ovate or orbicular ; whtre the branches arise the leaves are sometimes (but very rarely) small, and acutely bifid, on sterile stems decreasing in size towards the base and apex ; texture some- what delicate, cells smallish to medium size, 4-, 5- and 6-sided, quadrate or elongate, walls firm, no trigones or thickened angles. Stipules absent. Bracts similar to leaves, usually broadly ovate, spreading. No bracteole. Perianth projecting about f beyond the bracts, fusiform or oblong-ovate, below smooth, slightly frontally compressed, with a somewhat deep, antical furrow, bordered by two distinct folds, near apex with from 5 to 8 folds, mouth contracted, laciniate. Perigonial stems small, perigonial bracts at the middle or terminal, erect, imbricate, complicate, ventricose, smaller but proportionately broader than the stem leaves ; antheridia oval, single. Dimensions, — Sten;is from |^ to 1 inch long, '1 mm,, '15 mm,, •2 mn), diam. ; leaves 1'3 mm. long x 1"2 mm. broad, 1'2 mm, X I'l mm., ri mm. x 7 mm., 9 mm. x 7 mm., "8 mm. x ■7mm. ; cells "04 mm. x "025 mm., '04 mm. x '02 mm,, '03 mm, x "025 mm., •025 mm. X '02 mm,; bracts 1'5 mm, x 1'25 mm., I'i mm. X 1"5 mm.; perianth 2*4 mm, long x '75 mm. broad, "2 mm, x 75 mm., 175 mm, x 75 mm. ; perigonial bract "8 mm. long JUNGEBMANIA. 295 X "7 mm. broad, '8 mm. x "6 mm. ; antlieridia '125 mm. x •1 mm. ' Hab. — Growing in depre.ssed tufts on wet or damp rocks by the side of streams and in shady places. Somewhat rare, more frequent in limestone districts. 1. Near St. Ives, West Cornwall, /¥zY/ey^ §^ C/«5/v/ow. 5. Boulder near Seckley Wood ; Stream near Flash ; Dove Dale, Staffordshire, /. E. Bagnall. 7. Glyder Vawr, Carnarvonshire, W. H. P. Tyn-y- Groes, Merionethshire, Dr. Carnnyioii Sf W. H. P. 8. Miller's Dale, Derbyshire, G. A. Hull. 9. Bamford Wood, Lane, G. A. Holt. Marple, Cheshire, G. A. Holt. 10. Wharfdale, Dr. Sjrruce, Dr. Carrincjton. Horton, /. Whitehead. Saltersgate Beck, M.B. Slater. In^lehovo, IF. West. T>eiii,Dr. Carrui(/toii. Clap- dale, Dr. Cairington. Ingleton, W. West, W. H. P. Gordale, Dr. Carrington. Baildon, W. West. Sheddon Clough, /. Nowell. 12. Leven's Park ; on wet limestone rocks, Brightseer, Kentmere Plantation. Whitbarrow, Westmorland, G. Stabler. Helvellyn, Rev. C. H. Biusteacl. Patterdale, Westmorland, Dr. Carrington i^' W. H. P. Borrowdale, Cumberland, Dr. Carrington Sf W. H. P. 13. The Glen; iTolme Glen, Kirkcudbright, ./. Mc Andrew. 15, 16. Common among wet rocks and sides of streams, ascends to 1700 ft., Moidart, West Inverness, S. M. Macvicar. I. Kerry, Dr. Taylor. Tore Cascade, Killarney, Dr. Carrington. Brandon, Co. Kerry, Dr. D. Moore. Enniscona, Co, Cork, /. Carroll. Loch Bray and near Woodenbridge, Co. Wicklow, Dr. D. Moore. Benbulben, Co. Sligo, Dr. D. Moore. Eathliu Island, S. A. Steivart. Found on the Continent and in North America. Obs. — This is a species which assumes many forms, varying extremely in size and in shape of leaves, the perianth varying also according to its being more or less imperfect. Some of the acute Continental botanists have attempted to separate and describe some of the forms, but, from the study of an extensive collection, I have come to the conclusion that all the differences of size and shape depend upon locality onlj-. It is often confused with Janger- niania jjuiiiilu, from which it is distinguished by being dioicous 296 HEPATIGJS. and the perianth proportionally wider; from small forms of Junger- mania cordifoUa — see notes under the latter species. Description of Plate CXXIV. — Fig. 1. Plants natural size. 2. Stem X 24 (Miller's Dale, W. H. P.). 3. Leaves x 24 (276, G. & Ft.). 4. Leaf (ditto). 5, 6. Leaves x 24 (Miller's Dale, Gr. A. Holt). 7. Portion of leaf x 290 (Miller's Dale, W. H. P.). 8. Bract x 24 (276, G. & P.). 9. Perianth x 24 (Miller's Dale, W. H. P.). 10. Perigonial stem x 24 (ditto). IL Perigonial bract x 24 (Wharfdale, Herb. Slater). 12. Ditto (Canada, Macoun). 13. Antheridia x 85 (ditto). 4. Jungermania sphaerocarpa, Hooker. Jungermania s-phcerocarpa, Hook. Brit. Jung. t. 74 (181G). Jungermania Goulardii, Husn. Hep. Gall. n. 68 et Hepaticol. Gall. p. 20 (1881). Aplozia sjjhKTOcarjxi, Dum. Hep. Eur. p. 61 (1874). Monoicous, rarely paroicous, csespitose, pale to yellowish-green to brown in colour. Stems somewliat fleshy, simple or innovantly branched, innovations arising from base of perianth, erect or ascending ; radiculose, rootlets ascending to apex of stem, long, whitish. Leaves succubous, regular, alternate, approximate, obliquely inserted, slightly decurrent, embracing the stem, secund or spreading, subhorizontal, orbicular or roundish, somewhat slightly retuse, concave, plane or slightly undulate, lower leaves distant and smaller ; texture somewhat fleshy, epidermis smooth ; cells medium size, 5-, 6-sided, walls moderately thin, firm, trigones distinct, marginal cells quadrate, distinct. Stipules none. Bracts similar to the leaves. Bracteole none. Perianth often jDrojecting far beyond the bracts, usuall}^ §, obovate or obovate-oblong, sometimes with a long neck, composed of one layer of cells in the upper portion, in the lower 2 cells thick, 4-angled, sometimes with 5 or 6 angles. Pistillidia 7-10. Capsule spherical, some- times slightly hypophysate. Spores brown. Elaters bispiral. Perigonial bracts terminal on the innovant branches, 3 to 4 pairs, small, oval ; antheridia oval, with long bearers, 1 or 2 at the base of each bract. JUNGERMANIA. 297 Fruits March, April. Dimensions. — Stems |- to 1 inch long, '2 mm. to 3 mm. diam. ; leaves r4 mm. long x r35 mm. broad ; cells 04 mm., •025 mm. x '03 mm., '05 mm. x -025 mm., '02 mm. x '03 mm. ; bracts T mm. x 9 mm., 9 mm. x '9 mm. ; perianth 2*5 mm. long X 1-5 mm. broad, 2' mm. x 125 mm., 2' mm. x 1-mm., 175 mm. X rmm. ; pistillidia -15 mm. long x -05 mm. broad; capsule 75 mm. x '65 mm. ; spores '02 mm. diain. ; perigonial bract •8 mm. long x 7 mm. broad ; antheridia '15 mm. x -125 mm. Hab. — Growing on sandy or clayey damp banks or boggy places. (Generally distributed. 1, 2. Cadnum Bog, New Forest, Hants, C. Lyell 3. North Frith Wood, Kent, Uoiose. Joyden s Wood, near Bexley, Kent, E. M. Holmes. 5. Seckley Wood; Arley Wood; Sherbrook Valley, Staflfordshire, J. E. Bag nail . 7. Near Harlech, Merioneth- shire, Br. Carrington 8f W. H. P. 9. Early Bank Wood, Staley- bridge, Cheshire, 6^. A. Holt. Near Kemple End, J. A. IF/ieldon, Hindburn, A. Wilson. Easegill, West Lane, /. A. Wheldon. 10. Penyghent, W. West. Spofforth, Br. F. A. Lees. Bingley, W. West. Hareley Wood, Todmorden, John NoweU, G. A. Holt. 12, 13. New Galloway, /. McAndreiv. I. On stones by the side of rivulets. Nr. Dublin, Dr. Tuijlor- Kelly's Grlen, Dublin, Br. B. Moore. Lough Bray, Co. Wicklow, Br. B. Moore. Tore Waterfall, Killarney, Br. Carrington. Glenad, Co. Leitrim, Br. B. Moore. Hill of Howth, B. McArdle. Found on the Continent and in North America. Obs. — Distini^uished from all the other British round-leaved Jungermanice by the characters given, from Nardia ^^/^^^'(Schrad.) and Nardia conqjressa (Hook.) by the absence of stipules and bracteole, from Nardia hyalina (Lyell) and Nardia obovata (Nees) by its free bracts and white rootlets, from Jitngermania cord/folia Hook., Jimg. riparia Tayl., Jung, pinnila With, by its inflor- escence and shape of leaves and perianth, from Jungcrmania cremdata, var. graciUiina (Sm.) by its larger size, shape ot perianth, &c. See further notes under var. lurida (Dum.). Description or Plate CXXV.— Fig. 1. Plants natural size. 298 HEFATICE. 2. Fertile stem x 16 (Scotland, Original, Hooker). 3. Ditto (Sweden, Lindberg). 4. Ditto (Yorkshire, Mudd). 5 Leaf x 16 (Sweden, Lindberg). 6. Portion of leaf x 16 (Todmorden, Nowell). 7, 8. Bracts x 24 (Todmorden, Dr. Carrington). 9. Perianth x 16 (Slogdale, Mudd). 10. Cross-section of perianth X 16 (ditto). 11. Ditto (Bay sdale, Mudd). 12. Perigomal bract X 24 (495, Gr. & E.) 13. xintheridium x 85 (ditto). 14. Juwjer- mania Goulardii, Husn. x 16 (68, Husn. Hep. Gall.). 5. Jungermania sphserocarpa, Hook., var. lurida (Diim.). tnia lurida, Dum. Syll. Jung. p. 50 (1831). Jungermania nana, Nees Nat. Eur. Leb. 1, p. 317 (1833). Aplozia lurida, Dura. Hep. Eur. p. 60 (1874). Obs. — Dr. Spruce writes : ''Jungermania sjjharocarpa, Hook, is a species I have gathered abundantly both in England and the Pyrenees, but I have not had occasion to re-examine it for many years. Now, I have gone over all my splicer ocarp a and lurida, and utterly fail to find any specific distinction between them. The former is larger and often lax-leaved ; the latter smaller and with more crowded, more transverse, and proportionately wider leaves ; but there is no essential difference. " The Jungermania Goulardii, Husn. (of which you will have specimens in his Exsiccata) had been previously gathered by myself in exactly the same place — the ascent to the Gorge of Esquierry in the Pyrenees. It is exactl}^ what we Britons should call Jungermania sphcerocarpa, with rather broader leaves than ordinary. " Let us come to the conclusion of the matter, which is, I presume, that /. lurida, or nana, should stand as /. sijlicerocarpa, var. lurida. Other synonyms will doubtless be found in plenty by those who hunt for them." From the examination of numerous specimens from all quarters obtainable, I quite agree with the conclusion arrived at by Dr. Spruce, til at no satisfactory characters can be relied upon to separate J . lurida, Dum. {Jung, nana, Nees) from /. spharocarpa. JUNGERMANIA. 299 What slight differences do exist are owing to either drier localities or higher altitudes. Tj^Dical /. liirida has been named ./. sphcero- carpa by Hooker himself (Scotia, Herb. Hooker). Jimgcrmaiiiasjjhcerocar'pa, var. Inrida, (Dum.), is usually smaller, with less erect, fleshy stems; leaves usually darker in colour, pale brown to reddish-brown, more compressed, crowded, upper ones along with the bracts broader, lower ones on the contrary oval or slightly obovate ; cell structure identical. Jungermariia Goulardii, Husn. (68 Husn. Hep. Gall. Exsicc), is a smaller form, inflorescence like type monoicous, spores same size. Fruits April, May, June. DimensiojNS. — Stems |- to f inch long, '2 mm. to '25 mm. diam. ; leaves I'l mm. long x '9 mm. broad, 1' mm. x -7 mm., "9 mm. X "7 mm. ; branch leaves 'G mm. x "5 mm. ; cells "04 mm. X '03 mm., '03 mm. x '025 mm., '03 mm. ; bracts 1*6 mm. long X I'G mm. broad, 1"2 mm. x 1'6 mm.; perianth 2"25 mm. long xT mm. broad; valves of capsule 1"1 mm. long x "45 mm. broad; spores "02 mm. diam. ; elaters "16 mm. long x '01 mm. broad. Hab. — On wet rocks or in damp places, usually subalpine or alpine. Rare. 7. Pass of Llanberis, W. Wilson, 1830. Snowdon, W. U. P. Glyder Vawr, Carnarvonshire, IV. II. P. Tyn-y-Groes, Merioneth- shire, C. J. Wild. 8. Marple, G. A. Holt. ' Woodhead, /. W/ii/a- head ^' W. H. P. Mellor, G. A. Holt. Kinder Scout, Derby- shire, /. Whitehead 11. Cleveland, W. Mudd. 15. Mael Tar- machan, Perthshire, C. J. Wild. I. Glengarifl", Co. Cork, Miss Hiitchius. Kelly's Glen, Co. Dublin ; near Seven Churches, Co. Wicklovv, Dr. D. Moore. Found on the Continent. Descrh'tion of Plate CXXVL— Fig. 1. Stems natural size. 2. Fertile stem x 16 (Glyder Vawr, W.^H. P.). 3. Ditto, with imperfect perianth x 16 (Mael Tarmachan, C. J. Wild). 4. Ditto {Jnnf/. nana, N., original, ex herb. Flowtow, Nees). 5-8. Leaves x 24 (Mael Tarmachan, Wild). 9. Pranch-leal" x 24 300 HEPATIC^. (Glyder Vawr, W. H. P.). 10. Portion of leaf x 290 (Pyrenees, Dr. Spruce). 11, 12. Bracts x 16 (aiyder Vawr, W. H. P.). 13. Perianth x 24 (Verona, Massalongo). 14. Antlieridium x 85 (Glyder Vawr, W. H. P.). 6. Jungermania crenulata, Smith. Jungermania cremdata, Sm. Eng. Bot. t. 146o (1805); Hook. Brit. Jung. t. 37 (1816). Aplozia crenulata, Dum. Hep. Eui-. p. 57 (1874). Dioicous, loosely caespitose, flagelliferous, small, of a pale green, rosy or reddish colour. Stems reddish-brown, procumbent, cortical cells larger than the inner, whitish, 20 round, inner same colour, 7x8 in diam. ; simple or innovantly branched, innova- tions lateral, arising from axil of leaf, numerous, with small distant leaves ; radiculose, rootlets dull white, plentiful, ascending to apex of stems. Leaves on the upper portion of stem larger and more crowded, imbricate, obliquely inserted, patent, secund or spreading, lower leaves smaller and more distant ; broadly ovate to subrotund; texture somewhat thin, epidermis smooth, cells from smallish to small, roundish-quadrate, 4-, 5- and 6- angled, near the base elongate ('0237 mm.), walls thick, trigones very small or wanting, marginal cells largish ('053 mm.), oblong- quadrate, which give the plant a very distinct appearance, margin quite plane, not crenulate; these large marginal cells are usually wanting in the small leaves of the innovations. Stipules absent. Bracts similar to the leaves, only larger, appressed, sometimes produced from the lower part of the perianth, but very rarely. Bracteole wanting or sometimes rudimentary, then Ungulate. Perianth terminal, projecting about ^ beyond the bracts, com- posed of a single layer of cells, 130 round, larger at the angles, obovate, sHghtly compressed, plicate, 4-angled, angles smooth or irregular, obtuse or acute ; mouth contracted, submucronate, fringed with hyaline, elongate cells, margin crenulate. Calyptra thin, delicate. Pedicel long, pellucid, white, cross-section shows 17 cells round, 6 x 6 in diam. Capsule small, oval, dark reddish- JUNGERMANIA. 301 brown, spores pale reddisli-brown, elaters recldish-brown. Male stems smaller, slender, with several pairs of peri<^onial bracts, small, closely imbricate, semi-amplexicaul, ventricose at base, margin inflexed ; antheridia 1 or 2, oval, pale green, bearers short. Fruits Winter and Spring. Dimensions. — Stems |- to f inch long., diam. '2 mm. ; leaves •7 mm. long x "8 mm. broad, 75 mm. x '75 mm., "G mm. x •6 mm. ; cells "0275 mm., "0225 mm., "02 mm., '03 mm. x 0175 mm. ; marginal cells "06 mm. x -03 mm., '055 mm. x •03 mm., "04 mm. x '05 mm. ; bracts 1*35 mm. long x 1"1 mm. broad, 1*1 mm. x 1*1 mm.; perianth 1"G mm. long x T mm. broad, 1'5 mm. x 85 mm.; spores -015 mm. diam.; elaters •1 mm. long x '01 mm. broad; perigonial bract "5 mm. x "5 mm. ; antheridia "125 mm. x '11 mm. Hab. — Grows in swampy places on moors and in bogs, or in very damp situations by roadsides, &c. Generally distributed. 1. Cornwall, common, W. Curnow. 2. Bogs at Amberley, Sussex, W. Borrer. New Forest, Hants, C. LyelJ. 3. Epping Forest, Essex, E. Forsfer. 4. Holt Heath, Norfolk, Rev. R. B. Francis. Hungershall Eocks, &c., Kent, E. M. Holmes. 5. Can- nock Chase ; Winkshill, near Froghall ; Canwell, Staffordshire, /. E. Bagnall. 7. Roadside, Barmouth, Merionethshire, W. H. P. 8, 9. Longridge Fell, J. A. Wlieldon. Udale, West Lane, Jfheldoti ^' A. Wilson. 10. Ingleton ; Clapham, W. West. Yeadon Moor, Br. Carrington. Todmorden, John Nowell. Rose Wood, Sheffield, A. Carr. 11, 13. Roadsides, New Galloway, /. McAndrew. Glen Mills and Goldielea, Dumfries, Cruichshank. Grey Mare's Tail, Dumfries, Nichol. 15. Kinnordy, Forfar, C. Li/ell. 16. Local; on loamy footpaths, Moidart, West Liver- ness, S. M. Macvicar. I. Not unfrequent, Kelly's Glen, Dublin ; side of river, Seven Churches, Co. Wicklow, Br. B. Moore. Connemara, Dr. B. Moore. Ross Bay, Dr. Carrington. Hill of Howth, B. McArdlc. Slieve Glah, Co. Cavan, D. McArdle. C. Pointe Corbiere, Aurj. Martin. 302 HEPATIC^.. Found on the Continent and in North America. Obs. — Recognised at once, by tlie remarkably large marginal cells, from any of the other round-leaved Jmigermanice. The perianths are generally 4-angled, sometimes more ; Dr. Spruce met with a form in the Pyrenees with 8, some with 7, 6, and 5, but he could find no diflPerence whatever from the ordinary /. crenulaia, and remarks : " So that an occasional duplication of one or more of the perianth-angles would seem a common occur- rence in this section." Description of Plate CXXVII. — Fig. 1. Plants natural size. 2. Fertile stem x 24. 3-6. Leaves x 24. 7. Portion of leaf near margin x 85. 8. Ditto x 290. 9, 10. Bracts x 24. 11. Perianth x 24. 12. Cross-section of perianth, upper half X 24. 13. Portion of mouth of perianth x 85. (All Walton Swamp, Chesliire, Holt & Pearson.) 14. Perigonia] bract X 24 (Baden, Jack). 15. Antheridium x 85 (ditto). 7. Jungermania crenulata, Hook., var. gracillima. {8}ii.\ Jungermania cremdata, var. Hook. Brit. Jung. t. 37 (1816). Jungermania gracillima, Smith, Eng. Bot. n. 2238 (1805). Jungermania Genthiana, Hueben. Hep. Germ. p. 107 (1834). Aplozia gracillima, Dum. Hep. Eur. p. 57 (1874). Aplozia cristulata, Dum. Hep. Eur. p. 57 (1874). Dioicous, ca3spitose, tufts shallow, innovations plentiful, marginal cells similar to the others, on stronger stems the tendency to become larger is observable, bracts similar to the upper leaves, perianth mucronate, margin of mouth slightly crenulate, angles of perianth cristulate, calyptra delicate, claret coloured. Fruits April, May. Hab. — Growing in thin patches on shady sandy or gravelly banks. Generally distributed. 1, 2, 4, 5. Seckley Wood ; Arley Wood; Dimmings Dale, near Dane Bridge, Staffordshire, /. U. Bagnall. 7. Torrent Walk, Dolgelly, Merionethshire, W. II. P. 9. Alderley Edge, Cheshire, C. J. Wild. 10. Castle Howard, M. B. Slater; Teesdale, Dr. JUNQERMANIA. 303 Carrin(jfou. 12. Plentiful in the Lake District ; Gray vigg Forest ; in a quarry near Staveley ; Witberslack ; Whitbarrow ; Barbon Fell ; Long Sledtbile, Westmorland, G. SI abler. 13. Common on damp ground, New GaHoway, -/. Mc Andrew. 15, IG. Frequent on loam and sbale banks, Moidart, West Invei-ness, 8. M. Maccicar. I. Westaston, Co. Wicklow, Br. D. Moore ; Killakeen ; Slieve Glali ; on the shores of Lough Cultra, Co. Cavau, D. McArdle ; Hill of Howth, D. McArdle. Found on the Continent and in North America. Obs. — This form is so nearly related to the type tliat I re<''ard it simply as a variety. Description of Plate CXXVIIL — Fig. 1. Plants natural size. 2. Fertile stem x 31 (Alderley Edge, Cheshire, C. J. Wild). 3. Stem X 31 {Jung. Genthiana, Hub. & Genth. Hep. Germ. ^b. Original). 4-7. Leaves x 24 (n. 248, C. & P.). 8-10. Ditto (Alderley Edge, Wild). 11 Portion of leaf x 290 (n. 248, C. & P.). 12, 13. Bracts x 24 (ditto). 14. Portion of mouth of perianth x 290 (ditto). 8. Jungermania autumnalis, De C. Jimyermania autumnalis, De Candolle FI. Franc, 5, p. 202, sp. Wbib (IS 15); Moug. et Nest. Exsicc. n. 528. Lindb. Muse. Scand. p. 6 (1879). Jungermania subapicalis, Nees, Nat. Eur. Leb. 1, p. 310, n. 33 (1833) ; G. et R. n. 230 (as Jung. Schraderi, Mart.); G. et R. n. 231 (as Jung. Schraderi, Mart.) ; G. et R. n. 592 (as Jung. Schraderi, Mart.) ; Husn. Hep. Gall. Exsicc. n. 67 (as Jung. Schraderi, Mart.); Aust. Hep. Amer. Exsicc. n. 27 (as Jung. Schraderi, Mart.); Underw. Hep. Amer. n. 18 (as Jung. Schradei-i, Mart.). Jungermania Schraderi, Ekart Syn. Jung. Germ. p. 39, t. xl., f. 97, and of most authors (not Martius). Dioicous, csespitose, small, of a yellowish- or reddish-brown colour. Stems procumbent or suberect, simple or slightly branched, branches subpostical ; 40 to 50 cells in circumference, cortical cells slightly darker with the walls rather tirmer, inner 12 to 15 cells in diam., whitish; radiculose, rootlets distant, short, 304 HEPATIC^. dull white. Leaves obliquely inserted, horizontal or slightly ascending, alternate, approximate, plane or slightly concave, broadly ovate, entire or retuse, upper ones subquadrate with rounded angles, sometimes with a small antical tooth ; cells of medium size, roundish, walls thick, of a pale brown colour, trigones distinct, lumen clear or with few chlorophyllose granules, epidermis sraoothish. Stipules subulate. Bracts a little larger than the leaves, broadly ovate to subquadrate, retuse or entire with one or several long segments near to the base on one or both sides. Bracteole free or connate with bract on one side, oblong-quadrate or irregular in shape, laciniate or dentate. Perianth projecting § beyond the bracts, cylindrical or clavate, upper portion plicate, mouth slightly constricted, ciliate, cilia 5-10 cells long, 1 or 2 wide at base. Pistillidia about 30. Capsule oval, reddish-brown. Spores pale to reddish-brown. Elaters dark red, about 20 turns of the spiral. Male stems with 4-6 pairs of perigonial bracts, terminal or situated at the middle of the stem, close, ventricose, with a small incurved segment at the antical base protecting the single, large, shortly stiped, oval antheridium. Fruits April, May. Dimensions. — Stems about J inch long, diam. '15 mm. ; leaves 1*25 mm. x -9 mm., l"! mm. x '8 mm., 1* mm. x 7 mm., •9 mm. X *6 mm., '8 mm. x '7 mm., '75 mm. x '6 mm.; cells •04 mm. X "03 mm., '03 mm., '03 mm. x '02 mm. ; stipules •75 mm. long x "2 mm. broad at the base ; bracts 1*4 mm. x "9 mm., 1-2 mm. X '9 mm., 1" mm. x ^o mm., 11 mm. x '7 mm. ; bracteole 1- mm. X "6 mm. ; perianth 2' mm. x '9 mm., 1'5 mm. x '7 mm. ; cilia of perianth '125 mm., '225 mm. ; pistillidia "2 mm. x '05 mm. ; capsule '25 mm. x -2 mm. ; spores '0175 mm. diam. ; elaters "15 mm. long x '0125 mm. broad ; perigonial bract "8 mm. x "8 mm. explanate ; antheridia '3 mm. x '2 mm. Hab.— Grows in large patches or tufts in somewhat dry situations, on heathy banks or rotting wood in shady places. Bare. 7. Tyn-y-Groes, Merionethshire, Dr. Carrington Sf JF. H. P., JUNGIIBMANIA. 305 April 1879, G. A. Holt, May 1885. 1:^. On a bank by River Lune, near Ingmire Hall, Westmorland, G. Stabler ; Borrowdale, Cumberland, Dr. Carrim/to)i S)' ir. II. P. 13. Troquhain Wood, Balmaclellan ; Blackbank dyke, Glenlee, New Galloway, I. McAndrea'. Found on the Continent and in North America. Obs. — I have taken as type oS^ Jung, autz/nnialis, De C, n. 528 Moug. et Nest. Stirpes Crypt. Vog-Rhen. Exsic, wliich agrees entirely with the description . M' Ardle ; on white limestone near Grlenarm, Co. Antrim, Dr.D. Moore; Killarney on limestone,/);-. Carrington ; Tore Cascade, Killarne}^ and Carrantual, Kerrj',, Dr. B. Moore. Found on the Continent and in North America. Obs. — Under this name I have grouped all the small forms of what were considered Jungermania acuta, Lindenb., which are without stipules ; the form published in Gr. & R. n. 643 as Jung. acuta is the same as a form named Jung, turhinata var. aeutiloba. Spruce, Avhich connects Jung, turhinata with Jung, hantriensis var. Muelleri. Jung, turhinata is a very constant form and may be regarded as the extended type of a group with Jungermania hantriensis, Hook., at the other extreme. It may be distinguished from other dioicous species by the antical segment at the base of the upper leaves and bracts, its delicate cell structure and the turbinate perianth. Description of Plate CXXXII. — Fig. 1. Plants natural size. 2. Portion of stem x 31, antical view (/<'('/;y. //7/^50W2(2;?«. Original. Herb. Taylor). 3. Ditto, postical view (Bedminster, E. H. Read). 4, 5. Leaves x 31 (Cotterill Clough,^ Cheshire, W. Wilson). 6. Portion of leaf x 290 (Hb. Taylor). 7. Bracts x 31 (Bedminster, E. H. Read). 8. Sub-bracts x 31 (ditto). 9. Perianth x 31 (Fib. Taylor). 10. Cross-section of perianth X 31 (ditto). 11, 12. Portions of the mouth of perianth x 31 (ditto). 13. Antheridium x 85 (Monsal, Derbyshire, G. A. Holt). 312 HEPATIC ^E. Subgenus 3. LOP HO ZI A, Bum. •Tumjermania, sect. Lophozia, Dum. Syll. Jung. p. 53 (1831) ; Spruce, Hep. Am. et And. p. 511 (1885). Lopliozia, Dum. Kecueil, p. 17 (1835) ; Scliiflner emend Eng. & Prant. 1 ; Pflanz. 91 und 92 Lief. p. 81: (1893). Leaves suborbicular or subquadrate, bifid, in a few species 3-5 -fid, quite entire or sometimes denticulate. Stipules absent or present, often bipartite. Perianth usually deeply plicate, very rarely cylindrical and smooth. Section 1. BIDENTES, Schiffn. Stem leaves always bifid ; bracts similar or multifid. Stipules absent or present. 12. Jungermania bantriensis, Hooker. Jungermania bantriensis, Hook. Brit. Jung, in note under J. stipiilacea, n. 41 (1816). Jungermania bidentata, var. Hook. Brit. Jung. Suppl. t. Ill (1810). Jtingermania hygro2yhylla, Spruce, Trans. Bot. Soc. Edin. 11 (1840). Jungermania culearis, Wils. MSS. in Spruce, Hep. Pyren. Trans. Bot. Soc. Edin. Ill, p. 20 (1850). Jungermania subcomp-essa, Limpr. Jahr. Schl. Ges. p. 7 (1884). Dioicous, loosely ceespitose, medium size, of a pale, dark green or brown colour. Stems creeping or suberect, simple or innovantly branched; on a cross-section cells alike, walls dark, firm ; radiculose, rootlets long, few, whitish ; firm, flexuose, some- times flaccid. Leaves succubous, alternate, imbricate, secund or spreading, obliquely semi-vertically inserted, horizontal or sub- erect, antical margin decurrent, roundish-quadrate, broadly ovate or orbicular, emarginate-bidentate, from about j^j to 2^ deep, very shallow, broad, rounded, often folded at the base, segments acute or obtuse ; texture somewhat lax ; cells medium size, roundish or roundish-hexagonal, lumen pellucid, with few chlorophyl granules, walls somewhat firm, trigones small but distinct. Stipules lanceo- JUNGERMANIA. 313 late-subul:ite, broadly subulate or ovate-obloug, entire or with a basal tooth on one Side. Bracts broadly ovate or broadly oval, sinus acute, folded at the base, segments acute. Bracteole absent. Perianth projecting about |- beyond the bracts, obovate or ob- conical, smooth or slightly folded near the apex, composed of a single layer of cells, about 100 round the middle, near the base I, 2 or sometimes 3 cells thick, cells quadrate, near the base elongate, with trigones very small but distinct, mouth contracted, laciniate-ciliate, 20 to 25 cilia. Calyptra pale brown, delicate, composed of a single layer of cells above, 2 to 3 near the base. Dimensions. — Stems 1 to 2 inches long, "2 mm. to '3 mni. diam., with extended leaves 2' mm. to 3- mm. broad ; leaves r25 mm. long X 1*6 mm. broad, segments '2 mm., 1"3 mm. x r2 mm., seg. '2 mm., 125 mm. x 1'25 mm., seg. "1 mm.; cells "03 mm., ■03 mm. X -035 mm., '03 mm. x '025 mm. ; stipules '5 mm. long x -15 mm. broad at the base; bracts 2* mm. long x r75 mm. broad, segments 75 and "5 mm. long, 2' ram. x r5 mm., seg. •5 mm..; perianth 2'25 mm. long x 1*25 mm. broad, 2' mm. x 11 mm. Hab. — Growing in loose patches, or straggling amongst mosses on damp rocks or banks in woods. Ptare. 7. Dolgelly, Merionethshire, /. jS«/A' ; near Dolbadarn Castle, Llanberis, Carnarvonshire, JF. II. P. 10. Banks of the Tees, i)r. Spruce; Teesdale, John Noivell ; Bolton Woods, Dr. Carringfon. II, 12. Westmorland, P. Dreeson ; Garburn, Kentmere Park, G. Stabler; Kentmere Waterfall, G. Stabler; Long Sleddale, Rev. a H. Binstead; Stock Ghyll, G. Stabler; Groove Ghyll, Barbon Fell, G. Stabler (These Westmorland stations probably include var. Muelleri). 13. Crummy Park Glen; Ballingear Glen, New Galloway, /. McAndren-. 16. On a wet bank, rare, Moidart, West Inverness, S. M. Macvicar ^' JF. II. P. I. Near Bantry, Miss IhctcJiins ; Glengariff, Dr. Carrington; Brandon, Co. Kerrj^ ; Benbulben, Co. Sligo; Glenifi', Co. Leitrim, Dr. D. Moore. Found on the Continent. Ohs. — This I regard as the most perfectly developed type of a 314 HEPATIC^. series of forms running through Jin^f/. Mneller'i, Noes, Jimg. acuta, Lindenb. to Jung, furbinafa, Eacldi. at the other extreme, which all agree in their dioicous inflorescence, but the variation in size. shape of leaf, stipule and perianth differ to a very great extent ; for further notes see under the varieties following. Distinpruished from var. Muelleri, Nees, bv its larger size, more lax habit, more quadrate leaves, with shallower and broader sinus. From HarjiantliUH Flotowii, Nees, by its smaller stipules and terminal perianth. From Jungermania venfricosa, Dicks., which is also dioicous, by the presence of stipules and absence of bracteole. Description of Plate CXXXIIL— Fiq;. 1. Plants natural size. 2, 3. Portions of stem x 16 (Bantry, Miss Hutchins, Original). 4. Ditto x 24 (Saltzburg, Dr. Sauter). 5. Portion of leaf X 290 (Original). 6. Stipule x 85 (ditto). 7. Bract X 16 (Banks of the Tees, Dr. Spruce). 8. Ditto (Gotland, Lindberg). 9. Perianth x 24 (Original). 10. Ditto x 16 (Saltzburg, Dr. Sauter). 11. Portion of the mouth of the perianth x 85 (Gotland, Lindberg). 13. Jungermania bantriensis, Hook., var. Muelleri [Nees). Jungermania Muelleri, Kees in Lindenb. Syn. Hep. Eur. p. 39 (1829). Lophozia Muelleri, Dum. Eecueil, p. 17 (183.5). Dioicous, caespitose, small, pale green to dark greenish- brown in colour. Stems simple or slightly branched, flexuose, creeping or suberect, innovations arising from base of the perianth ; radiculose, rootlets plentiful, whitish. Leaves succubous, alternate, subimbricate, contiguous or distant, obliquely semi-vertically inserted, horizontal or slightly ascending, slightly decurrent antically, spreading, plane or concave, orbicular, ovate or sub- quadrate, emarginate-bidentate, sinus acute or rounded, narrow or broad, segments acute or obtuse, equal or unequal ; texture somewhat lax, cells medium size, roundish-quadrate, walls mode- JUNGEllMANIA . 315 rately thick, trigones distinct. Stipules small, lanceolate-subulate, acuminate, entire or dentate on one or both sides near the base. Bracts similar to the upper leaves, only larger. Bracteole wanting. Perianth terminal, projecting about ^ beyond the bracts, obovate or subcylindrical, smooth, mouth slightly plicate, contracted, laciniate-ciliate, cilia 2-4-cells long, submucronate. Pistillidia few (5). Perigonial stems smaller, bracts small, imbricate, ventricose, wdli a third antical segment. Dimensions. — Stems ^ to f inch long, •! mm. to "3 mm. diam., with leaves extended 1* mm. to 2" mm. broad ; leaves 1" mm. long X "9 mm. broad, segments '3 mm., 'i) mm. x "85 mm., seg. "2 mm., •8 mm. X "8 mm., seg. '2 mm., '75 mm. x -75 mm., seg. '2 mm., '75 mm. x '65 mm., seg. "2 mm., '5 mm. x "4 mm., seg. '15 mm., "45 mm. x '45 mm., seg. '15 mm. ; cells '033 ('04 mm. x '04 mm., '04 mm. X '03 mm., '03 mm. x "03 mm., '03 mm. x '02 mm.) ; stipules*'9 mm. long x "2 mm. broad at the base, "5 mm. x -15 mm., '4 mm. X '125 mm., 25 mm. x '05 mm., '15 mm. x "05 mm.; bracts 2' mm. long x 1-4 mm. broad, segments "5 mm. and '3 mm. ; perianth 275 mm. long x 1* mm. broad. Hab. — Growing in somew^hat dense, often small patches on damp rocks by streams or shaded places. Somewhat rare. 8. Ash wood Dale, Buxton, Derbyshire, G. A. Holt. 10. Near the Strid, Bolton Woods, Yorks, Dr. CaMngton ^' W. H. P. ; near Linton, Yorks, G. A. Holf, and other stations. 11, 12. Brown Ghyll ; Kentmere ; and other stations in "\^'estmorland, G. Stabler. 15. Near Bumbling Bridge, Dunkeld, Perthshire, Dr. Carrinrjton. IG. JNIoidart, West Inverness, 8. M. Macvicar. Pound on the Continent. Obs. — This is a form connecting Junt/. bantrienfii-'i, Hook., witli Jiaig. iurbinata, Eaddi., and has been variously named; an interesting series of varieties is met with in Europe which has been a source of perplexity to many authorities. I refer to it all the small forms of Jung, broifricnsis, Hook., which have leaves with more acute segments and sinuses, stipules present and with more cylindrical perianths. One of the types of Lindenberg's J/fiif/. acuta (Sillen. Muse. S16 HEPATIC^. Suec. n. 191) is certainly this variety, beino- a larg-e form with stipules. Dr. Spruce observes that Ji(N(/. bantrie/i^is, when young, has perianth pyritbrm or broadly clavate, but var. Jfin'/Zfri, cylindrical. J/i//(/. da/!tne/is^is has often the sinus folded at the base, var. MtieUeri usually plane, but I find the type and var. with and without this feature. Nees describes J>/n(/. Jf/zel/cri with bracts dentate : I should think he had some other species under his notice, for I have seen no approach, in any form, to this character. Forms oiJiing. venfricosa, Dicks., have been confounded with this variety, but are recognised by the postical side of the stem being coloured violet, the absence of stipules, tlie presence of bracteole, which is connate, the bracts and the ovate perianth. Distinguished from Ji(N(j. banfnenms, var. acuta, by the presence of stipules, i-^c. Descrtptiox of Plate CXXXIT. — Fig. 1. Plants natural size. -. Portion of stem, antical ^•iew x 1 6 (as Jung. banfrietisiSt Saltzburg, Sauter, Original, Hb. Flowtow, Nees). 3. Ditto, postical view x 24 (Buxton, Derbyshire,, G. A. Holt). 4-6. Leaves x 24 (Saltzburg, Sauter). 7. Leaf x 31 (Linton. Yorks., G. A. Holt). S, 9. Leaves x 04 (Buxton, G. A. Holt). 10. Portion of leaf x 290 (Craven, Yorks., Dr. Carrington). 11, 12. Stipules x 85 (Linton, Holt). 13-15. Ditto x 85. {Buxton, Holt). 16. Stipule x 24 (191 Silleu. Muse. Suec. n. 191, 2i% Jnnff. acuta). 17, 18. Bracts x 16 (ditto). 19. Perianth X 16 (ditto). 14. Jungermania bantriensis. Hook., var. acuta {^Lindenb.) Jungermania acuta, Lindenb. Syn. Hep. Eur. p. 88 (18:?!1). Lophozia acuta, Diim. Recueil, p. 17 (1885). Dioicous, csespitose, small, pale or dark green to brown in colour. Stems decumbent or suberect, simple or innovantly branched, branches delicate, arising from base of perianth or apex of old stems ; radiculose, rootlets white, copious, ascending to i\pex of stems. Leaves approximate or distant, somewhat secund JUNGEliMANIA. 317 or repand, senii-vertically inserted, sliiilitly ascending-, antical base a little decurrent, ovate, oval or orbicular, bitid to about 1, sinus narrow, obtuse or acute, segments acute or sometimes (jbtuse, margin entire ; texture lax, cells roundisli-(|uadrate, sometimes elongate, walls thick, trigones distinct. No stipules. Sub-bracts similar to leaves, sligbtl}' larger. Bracts larger, oval, ovate or subrotund, bifid to about 1, sinus obtuse or acute, segments obtuse or acute, margin entire or sometimes with a very slight antical indentation. Perianth projecting about rj beyond the bracts, cylindrical or oblong-obovate, smooth, mouth small, slightly dentate. Male stems delicate, bracts more erect, clasping the stem, oval, complicate, saccate at the base, bilid, with usuall}^ an antical seg-ment, antheridia single, oval. Dimensions. — Stems J to J inch long, "1 mm. diani., with leaves "75 mm. to 125 mm. wide ; leaves 3 mm. long x 275 mm. broad, segments 075 mm., 35 mm. x 3 mm., seg. 075 mm., •5 mm. X "45 mm., se"-. '1 mm., "55 mm. x "4 mm., scii'. 1 mm., ■6 mm. X 5 mm., seg. 1 mm., 6 mm. x "6 mm., seg. "l mm.; cells "03 mm., "04 mm. x "03 mm., '05 mm. x '03 mm., -05 mm. x 035 mm.; trigones "01 mm.; sub-bracts "75 mm. x '05 mm., seg. '15 ]nm., '75 mm. x "75 mm., seg. '1 mm. ; l)racts I'l mm. x 11 mm., seg. "1 nun., 125 mm. x T mm., seg. o mm. ; perianth 2' nun. long X 75 mm. broad, "15 mm. x 75 mm.; pistillidia •1 mm. long X •()5 mm. broad; perigonial bract, flattened out ■3 mm. X "3 mm. Hab. — Growing on damp shady limestone rocks, liare. 8. Near Buxton, Derbyshire, /. WliUclicad ; Ashwood Dale, near Buxton, G. J. Jlo/f. 10. Under a bridge (limestone) at Castle Howard, J)r. S/jri'cc 15. Dunkeld, Lr. Cfirri/it/lu/i. Found on tlie Continent. Obs. — This lorm dififers from the type by its smaller size, absence of" stipules ; from var, MuclU'ri by the alisonce of stipules ; ivoiw Jtni(/('rti/n II if/ li/rfjiiiafa, l\addi, b}' the sha] e and texture otits leaves, the sub- bracts and bracts being rarely distinctly antically notched, the i)erianth not turbinate. 318 HEPATIC^. It is another interesting link connecting Jungermania han- trieiisis, Hook., witii Jungerniania furbinata, Eaddi. Description of Plate CXXXY. — Fig. 1. Plants natural size. 2. Portion of fertile stem x 24 (Dunkeld, Dr. Camngton). 3. Portion of stem x 31 (Castle Howard, Dr. Spruce). 4, 5. Leaves x 24 (Dunkeld, Dr. Carrington). 6-10. Lea^^es x 24 (Castle Howard, Dr. Spruce). 11, 12. Ditto x 64 (G. & E. n. 643). 13. Portion of leaf x 290 (Dunkeld, Dr. Carrington). 14, 15. Sub-bracts x 24 (Castle Howard, Dr. Spruce). 16, 17. Bracts x 24 (Dunkeld, Dr. Carrington). 18, 19. Ditto x 24 (Castle Howard, Dr. Spruce). 20. Perianth x 24 (ditto), 21. Perigonial bract x 85 (Dunkeld, Dr. Carrington). 15. Jungermania obtusa, Lindb. Jungermania obtusa, Lindberg Muse. Scand, p. 7 (1879); Kaalaas Leverm. Norge, p. 348 (1893). Dioicous, loosely ciespitose, medium size, pale green to fusce- scent in colour. Stem simple or bifurcate, thick, succulent, fragile, flexuose, below pale brown, above dark or pale green, on the postical side more or less brownish, prostrate, apex slightly ascending, the whole postical side densely radiculose ; rootlets short or long, often in fascicles, near the base purplish-brown, near apex hyaline. Leaves on sterile stems usually decrescent, sometimes of equal size, somewhat remote, upper often closer and contiguous, somewhat concave, obliquely inserted, slightl}' decur- rent antically, rotund to obovato-quadrate, to J or a little more bilobed, sinus semi-lunate, broad, gibbous, obtuse, rarely acute and narrow ; lobes oval, obtuse or rotundate, sometimes somewhat acute, slightly unequal, postical usually larger ; texture tender and soft ; cells smallish to medium in size, somewhat dense, quadrate-rotundate, more or less chlorophyllose but often trans- lucent ; angles slightly thickened ; cuticule verruclose. Stipules usually absent, when present rudimentary, more frequent on the male stems and at tlie apex and fork of bifurcate stems, minute, JUNGERMANIA. 319 subulate-lanceolate. Bracts similar in size to the leaves, or rather smaller, irregularly 2-4-lobed. Bracteole variable, entire or bifid. Perianth projecting, pale green, cylindrical-obovate, upper -J- pluri- plicate, below smooth, composed of one layer of cells, mouth narrow, liyaline, irregularly ciliolate, with 15-20 cilioles, 1 and 2 cells long. Male plant more slender ; andnpcia o to 4 pairs at the middle or apex of stem ; perigonial bracts much smaller than the leaves, concave, at the base saccate, bilobed ; lobes obtuse, connivent, unequal, postical larger; sinus broad, obtuse. Anthe- ridia 1-3 in each bract, oval-globose, yellowish-green, stipitate. Dimensions. — Stems 1 to 2 inches long, with leaves 2" mm. to 3* mm. wide, "5 mm thick, leaves 1*5 mm. long xl "2 5 mm. broad, segments "3 mm., cells '03 mm. x '025 mm., '025 mm., stipules '4 mm. x '15 mm., perianth 3*5 mm. x ■1'25 mm., peri- gonial bract 1' mm. x 1" mm., antheridia "15 mm, x '125 mm, Hat3. — On rockv ledges, in alpine situations. Ben Gaire, 1700 ft., Moidart, West Inverness, Sijmers 31. Maccicar, July 27, 1898. The only known British station. Found in Norway, Switzer- land and Styria. Obs. — There is no other alpine hepatic which this species is likely to be mistaken for, the obtuse lobes of the leaves readily distinguishing it. Drs. Kaalaas and Jorgensen have confirmed the Scotch speci- mens and have also sent me a series of Norwegian specimens with which our species iuUy agrees. The figure given by Dr. Henri Bernet in his " Cat. Hep. Sud- Ouest Suisse " does not well agree with any specimens I have seen. JuHf/. ohfiisa is one of the numerous discoveries made in Scotland by my friend Mr. Macvicar. Description oe Plate CXXXVI. — Fig. 1. Plants natural size. 2. Stem x 11 (Moidart, Macvicar). 3-5. Leaves x IG (ditto). 6. Portion of leaf x 290 (ditto). 7. Stipule x 24 (Norway, Bryhn). 8. Perianth x 16 (ditto). 9. Peri-onial bract X 24 (ditto). 10. Antheridium x ^b (ditto). 320 HEPATIC^. 16. Jungermania socia, Nees. Jungermania socia, Nees, Nat. Eur. Leb, 11, p. 72 (1836). Paroicous, Joosely caespitose, from small to medium size, dark green to j:>ale browu in colour, sometimes with a slight purplish tinge on the upper leaves. Stems simple or innovantly branched, branches slender ; prostrate or suberect, flexuose, thick, somewhat tender; radiculose, rootlets long, whitish. Leaves subinibricate or ajjproximate, accrescent, obliquely semi-verticall}^ inserted, horizontal or slightly ascending, decurrent antically, secund or sjDreading, slightly concave, undulate, orbicular, ovate, broadly ovate, or subquadrate, bifid to about J or ^, segments acute or obtuse, sinus acute or rounded, sometimes trifid, the third seg- ment being basal and smaller ; texture tender, cells smallish to medium size, 4-, 5-, and 6-sided, quadrate or oblong-quadrate, trigones very small but distinct, walls thin, lumen full of chloro- phyl granules which often cling to the walls, giving them a thickened appearance. Stipules generally present on the upper portion of stem, absent below, free or connate with the adjacent sub-bracts, lanceolate, ligulate, subulate, simple or rarely bifid. Sub-bracts {$) orbicular, subquadrate, bifid-trifid, with one or two marginal small teeth, usually two pairs, containing several antheridia, which are large, roundish-oval with rather long stipes. Bracts broadly ovate, irregularly dentate, bracteole oval or ovate, bidentate, connate or free. Perianth terminal, projecting more than J- beyond the bracts, cylindrical, subclavate, upper portion slightly folded, mouth with 5 segments, margin slightly and irregularly bluntly dentate, teeth one cell long, when young with a slight purple tinge. Calyptra mitriform, ovate. Pistillidia linear-oblong, about twelve. Spores pale brown golden colour ; elaters rather short, thick, dark claret colour, very diff'erent in colour from the spores, bispiral, spiral threads thick, closely twisted. Pruits April, May. Dimensions. — Stems from h to 1 1 inch long, with leaves 2- mm, broad, diam. 4 mm., diam. of young stem -25 mm. ; leaves '8 mm. JUNGERMANIA. 321 X 1- mm., segments -3 mm., I'l mm. x •\) mm., seg. '3 mm., •8 mm. X '9 mm., see:- *3 mm., 1" mm. x "8 mm., seg. "4 mm., •8 mm. X "8 mm., seg. "3 mm., '8 mm. x "7 mm., seg. "3 mm., 7 mm. X •() mm., seg. '1 mm. ; cells '03 mm. x '03 mm., -025 mm. X '04 mm., '025 mm. x '03 mm., '02 mm. x '04 mm., "02 mm. x •02 mm.; stipules '45 mm. x "2 mm., '45 mm. x 'IS mm., '225 mm. X -2 mm.; sub-bracts 1*1 mm. x 1'2 mm.; sub-bracteole 9* mm. X "3 mm., '6 mm. x '25 mm.; bract 1' mm. xTG mm.; bracteole "8 mm. x '5 mm. ; perianth 2*5 mm. x "7 mm. ; pistil- lidia "25 mm. x '075 mm. ; spores '02 mm. diam. ; elaters '1 mm. X '01 mm. ; antheridia "175 mm. x "175 mm. PIab. — Growing intermixed with Campijlopm nfrovirens in marshy places on the ascent of Cader Iclris, Merionethshire, to the left of the bridle path to Tow}^!, from Dolgelly, about 1500 ft. IF. IL Pearson, April 1876. Extremely rare. Found on the Continent. Variety. Near Barmouth, Merionethshire. IF. II. Pearson^ June, 1878. Obs. — Distinguished ivom Jiin^ennania capitata, Hook., by its much larger size, more distant leaves, which are more delicate in texture, and by the presence of stipules on the upper portion of the stem, &c. I had some hesitation in adding this species to our list of British Hepaticae, but as I sent good specimens to the late Prof Lindberg, who replied, '' your Jiaig. mjcia, Nees, is correctly named, no doubt left," I think it will stand good. I collected, near Barmouth, another form, where the stipules are found all along the stem. Description of Plate CXXXVII. — Fig. 1. Piants natural size. 2. Portion of young stem x 24 (Helsingfors, Lindberg). 3. Portion of stem x 16 (ditto). 4. Leaves x 24 (ditto). 5. Leaf x 24 (Original, Hb. Nees). 6-11. Leaves x 24 (Cader Idris,W.H.P.). 12. Portion of leaf x 290 (Helsingfors, Lindberg). 13. Sub-bract x 24 (Cader Idris, W.H.P.). 14. Sub-bract with sub-bracteole x 24 (Original, Hb. Nees). 15. Portion ot uracts x 24 (ditto). 16. Sub-bracteole x 64 (Cader X 322 HEPATIC^. Idris, W.H.P.). 17, 18. Ditto X 85 (Original, Hb. Nees). 19. Ditto (Cacler Idris, W.H.P.). 20. Bracteole x 24 (ditto). 21. Perianth x 24 (Original, Hb. Nees). 22. Pistillidia x 85 (ditto). 23. Spores x 290 (Helsingfors, Lindberg). 24. Elaters X 290 (Ditto). 25. Antheridia x 85 (Cader Idris, W.H.P.). 26. Ditto, rather young (ditto). 17. Jungermania capitata, Hook. Jungermania e.vcisa, Dicks. PI. crypt. Brit. fasc. Ill, p. 11 (170,3) ?; Lindb. Muse. Scand. p. 7 (1879). Jungermania capitata. Hook. Brit. Jung. p. 86 (1816). Jungermania excisa, var. crisjyata, Hook. Brit. Jung. t. 9 (1816). Jungermania intermedia, Lindenb. Syn. Hep. Eur. p. 83, p. 93 (1829). Paroicous, csespitose, small, pale or dark green in colour. Stems simple or innovantly branched, frontally compressed, 14 cells X 10, cortical cells darker; prostrate or suberect, somewhat succulent. Leaves above semi-vertically and below sub-horizon- tally inserted, crowded, imbricate,- capitate, crisp and undulate, below smaller and more distant, ovate, orbicular or subquadrate, bifid to about |-, rarely trifid, uppermost leaves and sub -bracts trifid, sometimes quadrifid, segments acute, sinus obtuse, sometimes lunulate ; texture tender, cells from moderate to largish in size (•038 mm.), 4-, 5- and 6-sided, somewhat thin walls, trigones very small. No stipules. Bracts broadly ovate or orbicular, quadrifid, segments irregular. Bracteole free or connate with a bract, obovate or oblong-quadrate, obtuse or apiculate, sub-bracts with a large lateral antical segment to enclose the antheridia. Perianth obovate or oblong-obovate or subcylindrical, upper third portion slightly folded, 4 or 5 furrows, mouth contracted, bluntly dentate, teeth one cell long, usually very short, often tinted slightly purple. Pistillidia about 20. Calyptra delicate. Spores brown. Elaters bispiral, brown. Antheridia inserted immediately below the perianth, 1, 2 or rarely 3 at the base of each bract and sub- bract. This plant is sometimes gemmiparous, geramse diamond- JUNC^EBMAXIA. 323 shaped, pyriform or spherical, pale green or sometimes of a purple colour. Fruits March, April. Dimensions. — Stems \ to h inch long, "2 mm. diam., with leaves i-5 mm. broad; leaves I'l mm. x 1' mm., segments "4 mm. lonc^, 1* mm. x l* mm., sesf. "4 mm., "8 mm. x G mm., seg. '2 mm., •6 mm. X "G mm., seg. '2 mm. ; cells "05 mm. x "04 mm., '05 mm. x "03 mm., '04 mm. x '04 mm., "04 mm. x "03 mm., "04 mm. x •025 mm. ; bracts r2 mm. x i'o mm., segments '2 mm., '4 mm., •5 mm. long, 1"1 mm. x 1"2 mm., seg. -2 mm., '25 mm., "4 mm., 1" mm. X 1*2 mm., seg. "2 mm., 'S mm., "5 mm., 1' mm. x \- mm., seg. *4 mm., 'O mm. x 1" mm., seg. "3 mm., '7 mm. x 7 mm., seg. "2 mm. ; bracteoles "8 mm. x '5 mm. ; sub-bracteole '4 mm. x •4 mm., perianth 2 '5 mm. xl" mm., 2* mm. x "7 mm.; spores •02 mm. diam. ; elaters "1 mm. x -01 mm. ; antheridia "14 mm. x *1 mm. ; stipe OG mm. long x 01 mm. thick. Hab. — Growing in patches on sandy banks or moors. Some- what rare. 1, 2. Guestling near Hastings, Brr. E. K. Blooiufich/ ; Cadnum Bog, New Forest, Hants ; Lyndhurst Eacecourse, C. Li/elL 3. Hungershall Rocks, ./6^;z^/6^/-. 5. Cannock Chase, near Rugeley ; Gospel End Common, Staffordshire, /. E. Ba(jncdL 7. Llanfaelog, Anglesea, W. Wilson. 9. Udale, West Lane, WlteJdoii ^- A. Wilson. 10. Strensall Moor, Br. Spruce; Castle Howard, Dr. Spruce; Grosiwont, M. B. Slater. 12. Witherslack, Westmorland, G. Stabler. 13, 15. Sculty and Gateside, Strachan, Aberdeen- shire, ./. Sim; Kinnordy, C. Lijell. 16. Moidart, West Inverness, S. M. Afacvicar. I. Near Bantr\^, Aliss HutcJdns ; Galtymore Mountain, Co. Tipperary, Br. B. A'loore. Found on the Continent. Obs. — This may be the Jung, excisa of Dickson (PI. crypt. Brit. fasc. 3, 1793), as Lindberg asserts, but his description and figures are vague and incorrect, so I prefer to adopt the very characteristic name of Hooker. Distinguished from Juu//. veniric.osa, Dicks., by its paroicous inliorescence. 324 HEPATIC^. From Ji())(/. xocia, Nees, and Jim^. bicrenata, Lindenb., both paroicous species, see notes under them. Description of Plate CXXXVIII.— Fig 1. Plants natural size. 2. Fertile stem x 16 (Castle Howard, Dr. Spruce). 3-5. Leaves x 24 (Strachan, Sim). 6, 7. Ditto (Kinnordy, C. Lyell). 8. Leaf x 24 (Llanfaelog, North Wales, Wilson). 9. Portion of leaf x 290 (Castle Howard, Dr. Spruce). 10. Bract x 24 (Llanfaelog, Wilson). 11. Ditto (Strachan, Sim). 12. Bract with connate bracteole x 24 (Grosmont, Slater). 13. B^-acteole x 24 (Castle Howard, Dr. Spruce). 14, 15. Sub- bracts X 24 (Strachan, Sim). 16. Ditto (Grosmont, Slater). 17. Ditto (Kinnordy, Lyell). 18. Sub-bracteole x 31 (Castle Howard, Dr. Spruce). 19. Perianth x 24 (Strachan, Sim). 20. Cross-section of perianth x 24 (Castle Howard, Dr. Spruce). 21. Antheridium x 85 (ditto). 18. Jung-ermania bicrenata, Schnidel. Ju/igermania bicrenata, Schmid. Anal. p. 347, t. 64, f. 1 (1747). Lo])hozia bicrenata, Dum. Recueil, p. 17 (1835). Paroicous, csespitose, patches thin ; small, from yellowish to reddish-brown in colour. Aromatic. Stems simple, rarely inno- vantly branched, creeping, procumbent, the perianth and upper portion of stem suberect, stem bearing rarely more than 8 pairs of leaves ; radiculose, rootlets abundant, whitish, by which the stems are firmly attached to the ground. Leaves semi-vertically inserted, ascending, imbricate, concave, not undulate, accrescent, ovate, orbicular, sometimes subquadrate or obovate, bifid from J to ^, segments and sinus acute or obtusiuscuius ; texture thick, guttulate, cells from smallish to medium in size, roundish or roundish-oblong, walls very thick, no trigones. No stipules. Bracts oblong-quadrate, trifid, quadrifid, segments irregularly dentate, rarely entire, from ^ to ^ long ; bracteole subulate, entire or oblong-quadrate and emarginate, connate with one of the bracts, second pair of bracts orbiculate, irregularly dentate, third JUNGEBMANIA. 325 pair broadly ovate, less dentate, but witli a large tooth or segment near the antical base to enclose the antheridia. Perianth terminal^ projecting about ^ beyond the bracts, ovate, oblong-oval, upper half plicate, mouth contracted, hyaline, divided into about 8 segments, which are ciliolate, cilia 20 to 24, from 2 to 4 single cells long, incurved. Pistillidia about 12. Calyptra oval, narrow at the base, delicate texture. Capsule brownish-red, almost spherical. Spores brown. Elaters brown, bispiral. Antheridia situated below the perianth, in one or two pairs of sub-bracts below the innermost pair of bracts, roundish-oval, single, bearers long. Fruits March, April. Dimensions. — Stems about \ inch long, rarely ^ inch, diam. "25 mm., upper portion of stem with leaves 1* mm. wide, lower portion *5 mm.; leaves 1" mm. x '8 mm., sinus '4 mm. deep, •9 mm. X "8 mm., seg. "S mm., "9 mm. x "7 mm., seg. '25 mm., '7 mm. X '7 mm,, seg. "15 mm., '7 mm. x '6 mm., seg. '2 mm. ; cells "04 mm. x "025 mm., -035 mm. x -03 mm., '03 mm. x -025 mm., •03 mm. X -02 mm. ; bracts 1*4 mm. x 1*2 mm., segments '2 mm., •3 mm. and '4 mm., 1"6 mm. x "9 mm., seg. '6 mm. and '4 mm. ; sub-bracts 1"1 mm. x I'l mm., sinus '35 mm. deep, 1* mm. x .8 mm., seg. "25 mm., "9 mm. x '9 mm., seg. "2 mm. , perianth 275 mm. X 1'25 mm., 25 mm. x 11 mm., 2*3 mm. x 1'2 mm., 22 mm. x 1" mm., 1-75 mm. x '9 mm., 1'5 mm. x 7 mm.; cilia at mouth of perianth "15 mm. long x '02 broad, at the base "12 mm. X "02 mm.; pistillidia "15 mm. x "05 mm.; antheridia '16 mm. X '13 mm. ; stipe '1 mm. ; spores '015 mm. diam. ; elaters "1 mm. long X '01 mm. broad. Hab. — Growing on exposed rocks, in woods and on heaths, on sandy soil, soft sandstone or on mud covered walls. Rare. 2. Ashdown Forest, Sussex, G. E. Davies. 5. Sandpit near Aldridge ; Brindley Valley, Staffordshire,/. E. Bar/ nail. 7 . Port- madoc ; Tremadoc, Carnarvonshire, W. Wilson. Barmouth, Merionethshire, /. TFJiiteJiead, C. J. Wild. 8. Swithland Wood, Bloxam, F. T. Mott. Bardon Hill, Leicestershire, Cohnian. 9. Dale Ford Dingle, W. Wilson. Delamere, IV . H. P. Alderley 326 HEPATIC^. Edge, Chesliire, G. A. Holt. Whiteley Dean, near Hollingworth Lake, Lane, G. A. Half. 10. Strensall Common; Grosmont, Yorks., M. B. Slater. 13. S. of L. Dungeon, New Galloway, /. Mc Andrew ; Lochar Moss, Kirkcudbrightsliire, /. Cruichhank. 15. Straciian, Aberdeenshire, J. /S'ii'^w. 16. Moidart, West Liver- ness, on bare peaty banks with Sccqjcmia curta, S. M. Macvicar. I. Temple Michal, Cork, Isaac Carroll; near Letterfrack, and Kylemore, Co. Galway, Dr. D. Moore; Ballykill, Hill of Howth, B. McArdle. Found on the Continent and in North America. Obs. — A very distinct species. From Junff. capiiata Hook., another paroicous species, it is distinguished by its brown colour, the closely and uniformly imbricated, concave leaves, areolation quite different, guttulate, the lumen small, with remarkably thick walls, bracts and sab-bracts more acutely dentate, perianth cilio- late ; in addition to these characters, the plant has a strong aromatic smell when fresh, and even when dry, which becomes more noticeable by moistening it. Description of Plate CXXXIX. — Fig. 1. Plants natural size. 2. Fertile stem x 24 (Sussex, G. Davies). 3. Leaf x 24 (Dale Ford, Wilson). 4-7. Leaves x 24 (Sussex, Davies). 8. Leaf x 24 (G. & K n. 187 ). 9, 10. Leaves x 24 (Nassau, Huebener), 11. Portion of leaf x 290 (Sussex, G. Davies). 12, 13. Bracts x 24 (ditto). 14. Sub-bract, 2nd sub-bract x 24 (Dale Ford, Wilson). 15. Ditto, 3rd sub-bract (ditto, from smaller plant). 16. Ditto, 3rd sub-bract (Nassau, Huebener). 17. Sub-bract and leaf near sub-bract x 24 (Dale Ford, Wilson). 18. Perianth x 24 (Sussex, G. Davies). 19. Cilia x 85 (ditto). 20. Pistillidia with paraphyses x 85 (Barmouth, J. Whitehead). 21. Antheridium x 85 (ditto). 22. Spores x 290. 23. Elaters X 290. JUNGEllMANIA. 327 19. Jung-ermania ventricosa, Dickie. Jungermania minima repens,foUis bifidis, vagina Jiorum ventricosa, Mich. Nov. pi. gen. p. 9, t. 5, f. 15 (172!)). Lichenastrum quod Jtingermania minima repens, foliis bifidis, vagina fiorum ventricosa, Dill. Hist. Muse. p. 489, t. 70, f. 14 (1741). Jufigermania ventricosa, Dicks. Vl. crypt. Brit. fasc. 11, p. 14 (1790); Hook. Brit. Jung. t. 28 (1816). Lojihozia ventricosa, Dum. Recueil, p. 17 (1835). Jtmgermania ventricosa, var porphyroleuca, Limpr. in Cohn, Krypto. Fl. Scliles. 1, p. 280 (1876). Jnnger mania liorphyroleuca, ISTees, Nat. Eur. Leb. 11, p. 78 (1836). Dioicous, csespitose or straggling amongst mosses, small to medium in size, of a light or dark green colour, sometimes with, a reddish tinge. Stems procumbent or suberect, simple or branched, frontally compressed, innovations arising from below the perianth, single and stout, giving the perianth a laterally inserted appearance, or several and then slender, the postical side of the cortical layer usually of a reddish or violet colour ; radi- culose, rootlets abundant, of a whitish colour. Leaves semi- vertically inserted, horizontal or patent-divergent, imbricate or approximate, on fertile stems accrescent, on sterile stems often largest about the middle, concave or reflexed, gemmiparous, obovate, ovate, broadly ovate or subquadrate, obtusely and widely but not deeply (about \) emarginate, rarely and only near the bracts trifid, segments obtuse or acute ; texture somewhat succu- lent, cells from moderate to largish in size, 4-, 5- and 6-sided, roundish-quadrate, lumen usually filled with chlorophyl granules, trigones small but very distinct. No stipules. Bracts broadly ovate, ovate or subquadrate, bifid, trifid or quadrifid ; bracteole simple and Ungulate or oblong-quadrate and emarginate, connate with the bracts or rarely free. Perianth terminal, projecting about \ beyond the bracts, ovate-oblong, obovate, ventricose, when young and barren, smooth, upper portion obtusely folded, with about 5 folds, mouth contracted, margin irregular, dentate, teeth minute. Pistillidia large, long. Capsule dark brown, 328 HEPATIC^. valves splitting down to the base, viewed through transmitted light of a deep dark red colour. Spores reddish-brown, elaters the same, bi-trispiral. Perigonial stems usually slenderer, growing in separate tufts or entangled with the fertile ; bracts clasping the stem, ventricose, usually terminal, two or three pairs, containing in each bract 2 large, oval antheridia. Gemm?e on the ends of many of the upper leaves, yellowish- green, three and four angled. Fruits March, April. Dimensions. — Stems from |- to 1 inch long, diam. '2 to %S mm., with leaves spreading 2* mm. broad; leaves 1*5 mm. long X 1-4 mm. broad, segments '4 mm., 1'5 mm. x 1"1 mm., seg. 4 mm., 1"3 mm. x 1"3 mm., seg-. '4 mm., 1'2 mm. x 1' mm., ses:. 3 mm., I'l ram. x "9 mm., seg. "25 mm., 1"1 mm. x '7 mm., seg. 3 mm. ; cells '06 mm. x '03 mm., "05 mm. x •04 mm., 045 mm. x 04 mm., 04 mm. x "03 mm., '035 mm. x "035 mm.; trigones 0075 mm.; bracts 2" 1 mm. long x 2* mm. broad, seg. "5 mm., 1*7 mm. X 1*3 mm., seg. "5 mm., 1*5 mm. x 1"4 mm., seg. "4 mm., 1-5 mm. X 1'2 mm., seg. "4 mm., 1"4 mm. x 1*2 mm., seg. '5 mm. ; bracteoles -5 mm. long x -175 mm. broad, '325 mm. long x -1 mm. ; pistillidia "3 mm. long x '06 mm. broad, '25 mm. x '05 mm.; perianth 3' mm. long x r3 mm. broad, 1"8 mm. x 1' mm., 17 mm. X 11 mm, 1-3 mm. x '7 mm. ; spores "015 mm. diam. ; elaters 17 mm. long x '015 mm. broad; perigonial bract '9 mm. long X "6 mm. broad; antheridia "2 mm. x '175 mm.; gemmse •02 mm., -015 mm., ^0125 mm. Hab. — Growing in patches on rocks, clayey or sandy ground or straggling amongst mosses on heaths or in bogs, in the plains or subalpine situations, on rotting wood, when it becomes the Jf/,n(/ermajiia poiyln/roleifca of authors. Generally distributed. 1. Penzance, Cornwall, W. Cuntoic. 2. Sussex, G. J^. Dav?e8 ; Tunbridge Wells, Sussex, Dr. Spruce; Hastings, Sussex, Bev. E. N. Bloomfeld. 3. Southborough, Fmvcett ; Roadside near Seal, Sevenoaks, Hon%se ; Ightham, Kent, E. M. Holmes. 4. Tuddenham, West Suffolk, Eafjle. 5. Near Leek, /. E. Bacjiiall ; Dimmings JUNGERMAXIA. 329 Dale, /. E. Bmjnall ; Trentliam, 7?. Garner; Gospel End; Seckley ; Tettensor, &c., StaflPordsliire, /. E. Bapiall. 7. Abergynolvvyn, T. Bo(/crs; Cader Idris, W. 11. P.; Tyn-y-Groes, IF. H. P.; Cvvm Bychan, Merionethshire, E. M. Holmes; Lyn Bodlyn, E. JL Jlohi/e.s; Llanberis, Carnarvonshire, G. E. Huni. 8. Kinder Scout ; Charlesworth Coombs, Derbyshire, G. A. HoU. 9. Green- field, W. West; Bamford Wood, G. A. Holt ; Clifton June, Lane, G. A. Holt; Above Kemple End, West Lane, ./. A. Whet don; Marple, Cheshire, G. A. Holt. 10. Bingley, //'. Wext; Salters- gate Beck, S. Andersoii ; Blaeberry Gill, M. B. Slater ; Bolton Woods, Dr. Carr'ivgton, G. A. Holt. 12. About Staveley ; Saddle Forest ; Foulshaw Moss, among Sphagna ; Witherslack ; Wild- boar Fell, Westmorland, G. Stabler; Borrowdale, Cumberland, Dr. Carrinr/ton S>' W. H P. ; Isle of Man, G. A. Holt. 13. White- hill, C. Scott; Kelton and Lochar Moss, /. Cndchshank. 15. Strachan, Aberdeenshire, T. Sim. 16. Glen Finnan, Dr. Car- riiifjtoii ; Moidart, West Inverness, S. M. Macvicar. I. Banks and rocks in mountain situations, Dublin and Wicklow, Conemore, Dr. Taylor; Antrim; Benbulben, Sligo ; Gal ty more, Tipperary, Dr. D. Moore. Rare at Killarney, Dr. Carrington; Slieve Glah, Co. Cavan, D. McArdle ; Ballykill, Hill of Howth, D. McArdle. Found on the Continent and in North America. Var. Wliitehcadii. 9. Near Woodhead, Cheshire, Johi White- head, 1877. Obs. — This is a very variable species and a great number of varieties have been named by Lindenberg and Nees. With reference to Ju.vgcrmaiiia porphj/roleuca, N., I am con- strained by the examination of an extensive series of specimens to consider it only a variety of .Tfmr/. rentricosa ; the form growing on rotting wood might be accounted distinct enough, if the characters depended upon were only constant, but when the bracts vary from bifid to quadrifid with the bracteoles simple or bifid, connate or free, with segments more or less deep, the examination of a larQ-e series can lead to no other conclusion than that tliev are forms of one species. This is all the more confirmed by the fact 330 HEPATIC^. that in the Herbarium of Nees, are found specimens of both forms and of paroicous species, along with Jiiuf/. alpcstris under the name of Jn/tf/. j-'o^'P^^JP^oleuca. A peculiar type of this species has been collected by the late John Whitehead near Woodhead, Cheshire, which is chiefly remarkable for the extremely large number of pistillidia; I have counted 37, 54 and 57 in three involucres, in this character approaching Jtfn(/. Mildei, G., with which it has other characters somewhat in common — bracts broadly ovate, unequally trifid or quadrifid and perianth plicate. Jun(/. Mildei, G., differs, however, from it, in its yellowish-brown colour not a bluish-green, with the extremities of the leaves tinged violet, the leaves almost vertically inserted and densely imbricated, not semi-vertically and loosely imbricated ; the lobes of the leaves of the young branches are almost equal, not very unequal, and the cells smaller, with large trigones. It has much the habit ol Jung, ccqjitafa, Hook., in its crowded upper leaves, but is dioicous. It affords a striking example of a transition fiom one species to another. To distinguish /. ventricosa from allied dioicous ones is easy. Jim^. incisa has a lighter green colour, leaves almost transversely inserted, upper leaves irregularly toothed. Juu^. bantriensis and its variety Muelleri, have stipules and different bracts and shaped perianth. From Jmi^. aljjestris, see notes under that species. Description of Plate CXL. — Fig. 1. Plants natural size. 2. Fertile plant x 16 (Tunbridge Wells, Dr. Spruce). 3. Male stem X 16 (C. & P. n. 171). 4. Leaf x 24 (Llanberis, W.H.P.). 5. Ditto (Cwm Bychan, E. M. Holmes). 6. Ditto (C. & P. n. 171). 7. Ditto (C. & P. n. 185). 8. Portion of leaf x 290 (Sussex, Gr. Davies). 9, 10. Bracts x 24 (Hastings, Bloomheld). 11. Bract X 16 (Isle of Man, Holt). 12. Young bract x 24 (G. & R. n. 185). 13. Bracteole x 85 (G. & R. 185). 14. Pistillidium x 85 (ditto). 15. Perianth x 24 (Hastings, Bloomfield). 16. Spores and elater x 290 (Sussex, G. Davies). 17. Perigonial bract x 24 (C. & P. n. 171). 18. Antheridium X 85 (ditto). 19. Gemmae x 290 (Bingley, West). Description of Plate CXLI. — Juugermania ventricosa, var. JUNGEBMANIA. 331 porpUyrolenca (N.). — Fig-. 1. Leaves x 17 (After G-ottsche, G. k R n. 288). 2, 3. Sub-bracts x 17 (ditto). 4, 5. Bracts x 17 (ditto). 6, 7. Ditto (Sweden, Lindberg). 8. Perianth x 17 (After Ct. in G. &; R. 288). Jmigprmania ventricosa, var. White- headii {Jmiff. IFIiifeheadii, Pears. MSS). 9. Leaf x 24 (Wood- head, J. AVhitehead). 10, 11. Sub-bracts x 24 (ditto). 12, 13. Bracts X 24 (ditto). 20. Jungermania alpestris, Schleicli. Jungermania alpestris, Schleicher, Exsicc. 2 cent. n. 59 (1804 ?) ; Web. Prod. p. 81 (1815). Jimgermania sudetica, Nees in Hueben. Hep. Germ. p. 142 (1834). Jungermania Goejyertiana, Hueben. Hep. Germ. p. 254 (1834). Cephalozia alpestris, Cogn. Hep. Belg. p. 35 (1872). Dioicous, growing in flat, closely entangled, often spreading tufts, or straggling amongst mosses, small, from green, when young, to reddish-yellow, brown or dark brown in colour. Stems suberect, simple or furcately branched, brown, when old dark brown, serpentine, flexuose, somewhat rigid ; radiculose, rooting up to apex of stem, rootlets close, pale brown, often reddish or reddish-purple. Leaves obliquely semi-vertically inserted, hori- zontal, spreading or secund, sub-complicate and semi-amplexicaul near the apex of fertile stems, antical margin slightly decurrent^ imbricate or approximate, gemmiparous, roundish-ovate, ovate or subquadrate, concave, bidentate, sinus v^ide, very shallow (y\p to \ deep), obtuse, segments unequal, acute or obtuse ; texture firm, epidermis smooth, cells smallish, 4-, 5-, and 6 -sided, roundish- quadrate, trigones very small but distinct, marginal cells slightly thickened and quadrate. No stipules. Bracts larger and broader than the stem leaves, roundish, margin entire or slightly irregular, bifid from -i^ to \, sinus acute or obtuse, not so wide. Bracteole simple. Ungulate or rarely oblong and slightly retuse, connate with the adjoining bract or rarely free. Perianth terminal, pro- jecting about ^ beyond the bracts, oblong or oval, smooth, slightly obtusely folded near the apex, mouth obtusely complicate, margin 332 HEPATIC^. dentate. Pistillidia 5-8. Capsule roundish-oval. Spores small, yellowish-brown, elaters reddish-brown, not much broader than the spores. Male plants usually growing in separate tufts or more rarely mixed with fertile, male stems slender, often furcate* branches short, $ terminal or middle of the stem, 3-6 pairs of bracts, with 2 (rarel}^ 3) antheridia at the base, perigonial bracts clasping the stem, bifid with a narrow sinus, ventricose at the base, antheridia roundish-oval. Growing with the antheridia are often leafy paraphyses. Gemmee often abundant, at the ends of the uppermost leaves, 3-5 angled, brown or purplish-brown. Fruits April, May, June. Dimensions. — Stems from |- to 1 inch long, "2 mm. diam., with leaves 1" mm. to 2" mm. wide ; leaves 1 "3 mm. long x 1.2 mm. broad, segments "25 mm., 1*3 mm. x 1*3 mm., seg. '25 mm., '8 mm. X •/ mm., seg. '1 mm., '7 mm. x '7 mm., seg. '05 mm., '7 mm. x *7 mm., seg. "05 mm., '6 mm. x '6 mm., seg. '05 mm. ; cells "02 mm. x '025 mm., -025 mm. x '025 mm., *02 mm. X "03 mm.; bracts 17 mm. long x I'o mm. broad, I'l mm. X 1* mm. (including bracteole), seg. •2mm.,'3mm.,'8 mm. x -8 mm., seg. '1 mm., '8 mm. x '7 mm., seg. "2 mm., 1. mm. x '8 mm., seg. ^1 mm., '8 mm' x '9 mm. (including bracteole), seg. '1 mm. ; bracteole 1' mm. long x '6 mm. broad, "6 ram. x '6 mm. ; pistillidia '16 mm. long x "06 mm. broad ; perianth 2'5 mm. long x 'I'l mm. broad, 2 '2 mm. x I'l mm., 2'1 mm. x '8 mm., 1'8 mm. x 1. mm. ; perigonial bracts *8 mm. long x '7 mm. broad ; antheridia '16 mm. X '14 mm. ; gemm-se '0125 mm. x "01 mm., '01 mm. x '015 mm. Hab. — Growing in alpine and subalpine situations, on rocks or earth, often in dry and exposed places. Eare. 7. Glyders, Carnarvonshire, W. Wilson. Cwm Idwal, Car- narvonshire, IF. H. P. Cader Idris, Merionethshire, W. H. P. Nr. Dolgelly, Merionethshire, Dr. Carringfon i^' W. H. P. 12. Staveley, Westmorland, G. Stabler. Eydal, Westmorland, G. Stabler. 15. Bei\MiicJ)h\n, JF. JFest, T. Ro(/ers. Ben Lawers, Sadler. Tillylair, /. Sim. 16. Ben Nevis, Z>/'. Carrinf/ton, IF. West, J. Whitehead. Moidart, West Inverness, S. M. Macvicar. JUNGERMANIA. 333 I. Slieve Glah, Co. Cavan, D. McArd/c ; 13enbulben, Co. Sligo, D. McArdh. Found on the Continent and in North America. Obs. — This species is nearly rehited to Jiuiger mania ventricom (Dicks.), and its var. porjjhyroleuca (Nees), on the one hand, and JiuH/ermama baiitrlensi^ var. acuta (Lindenb.), on the other. On examination of numerous specimens from several collectors, and those given in different Fasciculi, I am confirmed in the opinion that some uncertainty exists as to its characters. In my copy of Austin's Hep. Am-Bor., the specimens would refer better to a form of Jungermania ventricosa, which may have been given in mistake, for 1 have good typical specimens from the same station (White Mountains, Oakes). According to Crypt. Bad. n. 96li, J((ngeriuaina aJpe^itris a. latior is a form also qI Jungermania ventricosa, having the violet cortical stem layer, subquadrate leaves with more acute segments, but exactly the same type is given in G. & R. Hep. Eur. n. 304 and n. 264 as Jnngerniania ventricosa. To add to the confusion, I have specimens from Schleicher's Cent. n. 59, which are {ecc parte) the original Jang, alpestris, but my specimens are Jungermania bantriensis var. Aluelleri (Nees), having the subulate stipules on all the stems. In Carr. k Pears. Hep. Brit. Fasci. II. n. 110 is given a small iorm of Jungermania aljjestris as var. geJida (Tayl.), but since it was publislied I have had the opportunity of examining the original specimens of Taylor, and am inclined to look upon the type as quite a distinct species (see Jungermania gelida, Tayl.). From Jungermania ventricosa Dicks, and its var. jjorjjiigrolcuca (Nees), J. a/pest ris maybe distinguished by its often slenderer and more graceful stems, with smaller leaves, which are rounder, not subquadrate, with a wider and shallower sinus, the cortical layer of the stem being destitute of the violet colouring, the bracts only bifid, perianth narrower. Jungermania /jantriensis var. acuta (Lindenb.) is more delicate,^ of a darker green colour, leaves more oval, with narrow sinus. 334 HEPATIC^. larger cells, no bracteole, perigonial bracts with usually an antical basal tooth. Description of Plate CXLII. — Fig. 1. Plants natural size. 2. Fertile stem x 24 (C. k P. n. 109). 3. Male stem x 16 (a. & R. n. 265). 4. Leaf x 24 (Dolgelly, Dr. C. & W. H. P.). 5. Ditto (Staveley, Q. Stabler). 6-9. Ditto (Ben Nevis, Dr. Car- rington). 10. Portion of leaf x 290 (Staveley, G. Stabler). 11. Bract X 24 (Dolgelly, Dr. C. & W. H. P.). 12, 13. Ditto (C. &P. n. 109). 14. Bracteole x 24 (ditto). 15. Ditto x 16 (Dolgelly, Dr. C.&W.H. P.). 16. Pistiilidium x 85 (Ben Nevis, Dr. C). IT. Perianth x 24 (Dolgelly, Dr. G. & W. H. P.). 18. Spores x 290 (ditto). 19. Elaters x 290 (ditto). 20, 21. Perigonial bracts x 24 (C. & P. n. 109). 22. Antheri- diam x 85 (ditto). 23. Gemmas x 290 (G. & Pv. n. 190). 21. Jungermania gelida, Tai/I. Jungermania gelida, Taylor, Journ. of Bot. p. 277, ii. 5 (1845); G. L. N. Syn, Hep. p. 67G (1847) ; Dum. Hep. Eur. p. 79 (1874). Jungermania alpestris, var. gelida, Cooke, Hanclb. Brit. Jung. p. 185 (1894). Dioicous, growing intertwined in tufts of Cesia concimiata, small, reddish-brown at the apices, below, and old stems brown in colour. Stems creeping, or ascending, simple or innovant near apex, flexuose, filiform, graceful, 10 cells in diameter, cortical cells 20-30, somewhat similar to the inner ones ; radiculose, rootlets plentiful, long, whitish. Leaves approximate, erecto-patent, secund, subrotund, bifid to 1 or ^, rarely trifid, segments acute, unequal, incurved, sinus obtuse or acute ; cells small (large for size of plant), roundish ; lumen filled with brownish granules ; walls thick, angles thickened. No stipules. 3 and $ not been seen. Dimensions. — Stems ^ to 1 inch long, diam. "15 mm., with leaves "5 mm, leaves "5 mm. x "4 mm., sinus "l mm., '8 mm. X '6 mm., sinus "2 mm., "7 mm. x '7 mm., sinus '2 mm., '7 mm. X '6 mm., sinus "15 mm., cells "02 mm. Hab. — -Scotch Alps growing intertwined with Cesia concinnaia. JUXGERMANIA. 335 l)riniiiiioii(l. 1 ('). Ben Nevis, Dr. GrcriJ/c. Kosliven, West Inverness, S. M. Macrirar. Obs. — This is a pretty little species, and thougli it may possibly be only a form of Jinir/. aljjestr}>< Sehleich., yet its liabit, small, secuud, acutely bifid leaves witli acute segments and different cell structure lead me to look upon it as distinct. Deschtption of Plate CXLIII. — Fig. 1. Plants natural size. 2. Plant X 24. 3-8. Leaves x 24. 9-14. Ditto x 31. 15. Portion of leaf x 290 (Scotch Alps, Drumniond original, Herb. Tayl.). 22. Jungerniania incisa, Sc/irnd. Janyerinania incisa, Schrader, Syst. Samml. krypt. Gew. 2 p. ."» (17'.i7) ; Hook. Brit. Jung. t. 10(181(5). Lophozia incisa, Dum. Recueil, p. 17 (1835). Dioicous, caespitose, small to medium in size, of a pleasant, bright, light green colour. Stems prostrate or suberect, ver}^ thick for the size of plant, frontally compressed, 9 and 10 cells by 14, of a brown colour beneath, the postical cortical layer of cells lig- neous and indistinctly cellular, the antical cortical cells smaller and delicate, inner hexagonal, thin walls, with numerous dark granules in each cell ; radiculose, rootlets very copious, on all the under side of stem, whitish, thick, long ; simple or furcately branched, branches often arising from base of perianth. Leaves succubous, horizontalh^ inserted, ascending, imbricating or accres- cent, closer and larger near the apex, subcomplicate, obovate, deltoid, broadly ovate, bifid or irregularly trifid to about the middle, margin entire or irregularly dentate or erose, segments acute or somewhat obtuse, often dentate about the middle, sinus rounded or acute ; texture somewhat coarse, epidermis smooth ; cells rather large, 4-, 5-, and G-sided, lumen filled with chlorophyl granules giving the plant a character distinct from its allies, walls moderately firm, trigones small. No stipules. Bracts larger than the leaves, l)ifid with large lateral segments, segments irre- gularly and coarsely dentate. Bracteole none. Perianth project- 336 HEPATIC^. ing about half beyond the bracts, obovate, oval-rotund or oblong-oval, slightly plicate near the mouth, mouth hyaline with 4 or 5 irregular segments, furnished with about 20 long or short cilia, which are 1, 2, or 3 cells long, incurved; their irregularity gives the mouth a lacerated appearance. Calyptra mitriform, cells delicate, yet regular and well defined, not so delicate and indistinct as in some allied species. Capsule oval, dark reddish- brown, valves dividing down to the base. Spores light brown,, elaters purplish-brown, bispiral. Male stems similar to others, sometimes more slender, perigonial bracts accrescent, crowded, forming small capitula, ventricose, bifid with incurved basal lateral segments, dentate ; antheridia spherical, large, one or two in each bract. Fruits March, April. Dimensions. — Stems J- to 1 inch long, "5 mm. diam., leaves 1'3 mm. long x 1* mm. broad, segments '7 mm., 1*2 mm. X 1'2 mm., sees'. "6 mm., 1'2 mm. x '7 mm., seg. '7 mm., I'l mm. X 1'2 mm., seg. '6 mm., '6 mm. x "5 mm,, seg. '4 mm.; cells •05 mm. X *04 mm., "05 mm. x '03 mm., .04 mm. x "04 mm., '03 mm. X "03 mm.; bracts 1'7 mm. long x 1'9 mm. broad, seg- ments "6 mm., '5 mm., 1'5 mm. x 1"8 mm., seg. "8 mm., seg. •4 mm., 1*5 mm. x I'O mm., seg. '6 mm., "3 mm. ; sub-bract 1*3 mm. long x 1*1 mm. broad, segment "5 mm.; perianth 2'7 mm. long x 1*5 mm. broad, 2' mm. x 1"1 mm., 1"8 mm. X 1*3 mm. ; cilia '15 mm. long, "12 mm., "07 mm. ; capsule •25 mm. diam. ; pedicel ^08 ram. diam. ; spores '015 mm. diam. ; elaters •I 2 mm. long x •Ol mm. broad ; perigonial bracts '7 mm. long X 1"1 mm. broad, segments 'A. mm., 1. mm. x.8 mm., seg. •4 mm. ; antheridia 'IQ mm. diam. Hab. — Growing in large patches on heaths, banks, rocks, or amongst mosses on moors ; most frequent in subalpine localities. Generally distributed. 1, 3-5, 7, 8, 10-16. I. Found on the Continent and in North America. Obs. — This is an easily recognised species, and from any of the allied dioicous species may be distinguished by the remarkably JUNGEIIMANIA. 387 frontally flattened upper portion of stem, its pleasant li^lit ^reen colour, the crisped, dentate leaves, and the leaf cells lilled with numerous granules. When dry the specimens do not retain their distinctive colour, but, on the contrary, turn a dingy black. Description of PL.vrE CXLIV. — Fig. 1. Plants natural size. 2. Portion of male stem x K> (Bettws-y-Coed, W. H. P.). 3. Upper portion of fertile stem x IG (France, Du Buysson). 4-6. Leaves x 24 (487 G-. & R). 7. Leaf x 24 (Ben Nevis, West). 8. Ditto x 24 (Da Buysson). 1). Portion of leaf x 200 (487 Gf. &R). 10, IL Bracts x 24 (Du Buysson). 12. Ditto X 16. 13. Sub-bract x 24 (ditto). 14. Perianth x 16 (228 Gr. & E.). 15. Mouth of perianth x 16 (Du Buysson). 16. Cilia from mouth of perianth x 85. 17. S]iores x 290. 18. Elaters x 290. 19, 20. Perigonial bracts x 24. 21. An- theridium x 85 (ditto). 23. Jungermania exsecta, Schmid. Jungermania exsecta, Schmidel, Ic. et anal. p. 241, t. (12, f. 2 (excl. f. fi-uctif. et 10, 20), (1747); Hook. Brit. Jung. t. 14 et suppl. t. 1 (1^/^ No well ; Teesdale, />/-. /§^r«6r. 11, 12. Wliitbarrow ; Heversham Head c. fr. ; Witherslack ; Low Wood, Windermere ; Bow Fell ; Naddle Forest ; Kentmere, Westmorland, G. Stabler. 13. North side of Black Craig, New Gralloway ; (irej Mare's Tail, W.Nichol; Dalscairth & MoH'at, /. C/7^/6'/('.s7/rty//-. 15. Ben Lawers, Perthshire, :Z'. /%6^Ay ; Banks of the Clyde, //'. Zj/o//. 16. Moidart^ West Inverness, S. M. Macvicar. I. Grlenmaluer, Co. Wicklow, Br. JJ. Moore. Found on the Continent and in North America. Obs. — This is the Jaitcjenuaula qiciuquedentata., Web., of Conti- nental authors, but it is absurd to call a plant which has uniformly only 3 teeth " qifi/iqicede/itata " ; certainly ail the authors write of it as having leaves with 3 to 5 and bracts witii 4 or 5 segments, but in ail Continental and British specimens examined I find them uniformly tridentate, without exception. It is also the Jiiiigerinania Lyoni of Taylor, and, since there is a great uncertainty as to what the early authors understood, I use Dr. Taylor's name. Dr. Spruce wrote me : " Juny. qiujiqiiedrnfata Huds. is no doubt what we call J. barbataJ' This, tlie most distinct species of the '' barbata " group, has the lower (antieal) margin of the leaf tridentate, and the upper (postical) margin is rounded and plane, which characteristics dis- tinguish it at once from any of the others. The perianth often appears to be lateral by the production at its base of robust, subpostical branches. Descript-ion of Plate CXLVI. — Fig. 1. Plant natural size. 2. Portion of stem antieal view X IG (Theden. Muse. Suec. n. 144). 3. Ditto X IG (Miller's Dale, AV. H. P.). 4, 5. Leaves X 24 (Bettws-y-Coed, .M. B. Slater). 6. Portion of leaf x 290 (Arthog, W. H. P.). 7. Bract x 24 {Jnn/j. Lijoni, Tayl. Clyde, Hb. Tayl.). 8. Ditto (Arthog, W. H. P.). 9. Bracteole x 24 {J any. lyoni, Tayl. Clyde, Hb. Tayl.). 10. Perianth x 16 (ditto). 11. Portion of the mouth of perianth x 85 (Arthog, W. H. P.) 12. Perigonial bract x 24 (Bettws-y-Coed, M. B. Slater). 13. Antheridium x ^b (ditto). 342 HE PAT 10 M 25. Jungermania gracilis, 8ch/eic/i. Jnngermania gracilis, Schleichex', PI. crypt. Helv. Cent. 3, n. CO (1804). Jungermania hctrbata, var. minor, Hook. Brit. Jung. t. 70, figs. 18-20 (1816). Jungermania harhata, var. attenuata, Mart. Fl. crypt. Erl. p. 177, t. 6, f. 50, c. (1817). Jimgermania attenuata, Lindenb. Hep. Eur. p. 48, n. 44 (1829). DioicoLis, loose]}' csespitose, sniall to medium in size, of a green or brown colour. Stems simple or furcate, or innovantly branclied, suberect, flexuose, branches subj)ostical-lateral, proceed- ing from apex of stem, filiform, attenuate, with small closely appressed leaves ; radiculose, rootlets plentiful, long, whitish. Leaves imbricate or approximate, obliquely inserted, patent or erect- patent, ov^al-quadrate, quadrate, concave, bifid, trifid, or rarely quad- rifid, from about ^ to ^ deep; leaves on the attenuate branches small, closely appressed, erect, imbricate, erose by the development of gemmoe; cells small, 4-, 5-, and 6-sided, lumen filled with chloro- phyllose granules, walls moderately thin, trigones small, distinct. Stipules Avanting or minute, linear, subulate, near the bracts distinct (sub-bracteoles), ovate-lanceolate, bifid to about the middle, margin entire. Bracts larger than the leaves, spreading, quadrate or broadly roundish-quadrate, trifid or quadrifid to about \, segments acute, unequal. Bracteole ovate-lanceolate, bifid to about the middle, connate with an adjoining bract, rarely free. Perianth terminal, projecting about \ beyond the bracts, cylindrical, oval- oblong, oval or ovate, upper half acutely 5-plicate, lower half slightly folded, 2 layer of cells thick at the base, higher one cell thick, mouth contracted, ciliolate. Calyptra 2 cells thick at the base, one higher where it is thin and delicate. Capsule oval, reddish-brown. Spores round, brown. Elaters bispiral. Male stems spicate ; perigonial bracts usually terminal, trans- versely inserted, closely imbricate, 5-6 pairs, ventricose, condupli- cate, bifid with a large tooth on the antical side (trifid), apices incurved ; antheridia one or two in each bract, oval. Gemmae terminal at the ends of the leaves, yellowish-red or green, irregular, oval or angular. JUNOEUMANIA, 343- Fruits April, May. Dimensions. — Stems from h to 1 inch long, rarely more, with leaves from 1* mm. to 1"5 mm. wide, attenuated portion 4 mm. wide, diam. of stem '2 mm. ; leaves "8 mm. long x "8 mm. broad, segments '25 mm., '9 mm. x "9 mm., seg. '3 mm. ; upper leaves •55 mm. x "5 mm., "5 mm. x -4 mm., '45 mm. x -4 mm., -4 mm. X "25 mm. ; cells '02 mm.; l)racts 1' mm, long x "9 mii). broad, !• mm. X 1" mm., seg. "4 mm. ; bracteole "8 mm. long x "5 mm. broad, seu'. "4 mm. ; sub-bracteole "8 mm. x '5 mm., sej^. '4 mm. ; perianth 2" mm. long x "9 mm. broad ; cilia "075 mm. ; peri- gonial bracts '5 mm. long x "7 mm. broad; antheridia '2 mm. X "15 mm. Hab. — Growing in loose tufts or creeping amongst mosses, on heaths or in woods, in the plains or subalpine localities. 1. Near Penzance, U'm. C/wnotf. 3. Abbey Wuod, Kent, K M. Holmes. 4. Holt, Norfolk, Rev. B. B. Francis. 5. Walls, Alton Towers ; Ramshorne ; Dimmings Dale ; Star Wood, Oak- amore, Stafford, ./. E. Bctf/naU. 7. Cwm Bychan, Merioneth, E. J/. Holmes. Barmouth ; Cader Idris, Merioneth, //'. //. B. 8. Charlesworth Coombs, Derbyshire; near Woodhead, Derb}'- shire, G. J. Holt. 9. Alderley Edge, Cheshire, C. J. Wild. Staley Brushes, Cheshire, G. A. Holt. 10. Bolton Woods, 7>r. Carri/i(/ton, Ingleboro' ; Penyghent ; Bingley ; Baildon, Yorks., IJ^. West. 12. Westmorland, G. Stabler; Skiddaw, Bev. C. H. Binstead. 13. Crichope Linn, Kirkcudbrightshire, C. Scott. 15. Kinnordy, C. Li/ell. 16. Moidart, West Inverness, S. M. Macvicar. I. South of Ireland, Miss Hut chins. Near Dublin, Br. Taijlor. Ballykill ; Howth demesne, D. McArdle. 0. Piemont et Grosnez, Aii(/. Martin. Found on the Continent and in North America. Obs. — This is a very polymorphous species, and varies much in size ; it is not likely to be confused with any other species on account of the singular attenuated stems. I should like to have adopted the characteristic name of Martins, Jnn//. hdrtjata var. attennata {Jnnf/. atteiinata (Mart.) Lindenb.), but Sclileicher's nauu' has the priority, and is quoted by Nees, " Eur. Leber.,"' 3U HEPATIC^. vol. 2 p. 164, as a variety. If there be any diflference, Martius's name should be the varietal, but Lindberg, who has seen the original, states they are the same. Description of Plate CXLYII.— Fig. 1. Plants natural size. 2. Stem X 16 (Abbey Wood, Kent, Holmes). 3,4. Leaves x 31 (Sweden, Lindberg). 5. Leaf x 31 (Kent, Holmes). 6-10. Leaves from attenuated stem x 24 (Canada, Macoun). 11, Portion of leaf X 290 (Kent, Holmes). 12. Stipule, free. 13. Sub- bracteole x 24 (Sweden, Lindberg). 14, 15. Bracts x 31 (ditto). 16. Eracteole x 31 (ditto). 17. Perianth x 24 (ditto). 18. Portion of mouth of perianth x 85 (ditto). 19-21 Peri gonial bracts x 31 (ditto). 26. Jungermania barbata, Sclireb. Jitngermania harhata, Schreber, F. Lip. p. 107 (1771); Hook. Brit. Jung. t. 70, ff. 7, 8 (1816). Jtmgermania barhata, var. Schreheri, Nees, Nat. Eur. Leb. 11, p. 189 (1836). Lophozia barbata, Dum. Recueil, p. 17 (1835). Dioicous, loosely csespitose, medium size, green or brow^nish- yellow in colour. Stems prostrate or snberect, thick, brown, simple or slightly branched; radiculose, rootlets close, whitish, copious. Leaves semi-vertically inserted, horizontal, spreading or secund, alternate, slightly imbricate or approximate, plane, sub- quadrate, subcuneate or digitate, antical margin decurrent, divided into 3, 4 rarely 5 subequal, acute or obtuse segments, sinuses rounded or acute; texture somewhat firm, cells small, 5- and 6-sided, walls thin, but with numerous chlorophyl granules attached, apparently thick, trigones very small. Stipules simple or bifid, variable, segments lanceolate or subulate, entire or with few, long teeth. Sub-bracteole oblong-quadrate, bifid, with several other smaller segments. Bracts roundish quadrate, quadri- dentate to ^, with a smaller segment on both sides, segments and sinuses acute. Bracteole oblong-roundish-quadrate, with 4-6 large segments. Perianth terminal, projecting about f beyond the bracts, oblong-oval, oblong-obovate or cylindrical, near apex JUNGEEMANIA. 345 6-plicate, folds short, mouth contracted, ciholate-dentate, 50-GO ciliok. Capsule roundish-oval, elaters and spores brown. Male stems little different from others, perigonial bracts terminal, 4, 5 pairs, closely imbricate, subcomplicate, slightly ventricose, antheridia oval, large paraphyses frequently mixed with the antheridia. Fruits April, May. Dimensions. — Stems from 1 to 1|- inches long, "o mm. to '4 mm. diam., with leaves 2*75 mm. to o" mm. broad; leaves 18 mm. longxl'5 mm. broad, segments '(3 mm., 1'4 mm. x 1*25 mm., seg. "o mm., 1*25 mm. x 1"25 mm., seg. "4 mm. ; cells '03 mm. x •02mm., "025 mm., -02 mm.; trigones '005 mm.; stipules r5 mm. long X '4 mm. broad; sub-bracteole 1*7 mm. longxl* mm. broad; bracts 1*9 mm. long x 1*7 mm. broad, .^eg. 6 mm.; bracteole 1'9 mm. long x I'G mm, broad; perianth 4 '25 mm. long X Ivo mm. broad; cilia at mouth of perianth 1 mm. long; perigonial bracts r25 mm. longxl" mm. broad, seg. '3 mm.; antheridia "25 mm. x "2 mm., "2 mm. x •175 mm. ; paraphyses *25 mm. long x "075 mm. broad. Hab. — Growing in loose tufts or creeping amongst mosses on damp shad}^ rocks, walls or banks. Somewhat rare. 1. Trevaylor Bottom, Cornwall, .7. RaJfo. 7. Tyn-y-Grroes, C. J. Wild §• G. A. Holt; Barmouth, Merioneth, W\ H. P. 8, 9. Longridge Fell, /. J. Wheldon ; Udale, West Lane, Wheldon 8f A. Wihou. 10. Malham, IF. IFesf. 11, 12. Patter- dale, C. Lyell; Black Crag near Staveley ; AVitherslack ; Winder- mere; Kentmere ; Barbon Fell, Westmorland, G. Sf abler. 13. Crichope Linn, C. Scoff; Dalbeattie A¥ood, C. Scoff; Craigs, J. Cniickshaiili ; Moffat, •/. Cndcksl/ank. 14, 15. Ben Lawers, C. J. Wild. 16. Moidart, West Inverness, S. J/. Macvicar. L "General in Ireland," Br. B. Moore; Hill of Howth, B. McArdle; Slieve Glah, Co. Cavan, D. McArdle. Found on the Continent and in North America. Obs. — Easily distinguished from other species of the same group by the plane, digitate leaves, which have neither basal teeth nor cilia. 346 EEPATIG^. Description of Plate CXLVIII. — Fig. 1. Plants natural size. 2. Portion of stem x 16 (142, Thed. Muse. Suec). 3. Leaf x 24 (Vosges, Schimper). 4. Portion of leaf x 290 (142, Thed.). 5. Stipule x 64 (Vosges, Schimper). 6. Bract X 24 (ditto). 7. Bracteole x 24 (ditto). 8. Sub-bracteole x 24 (ditto). 9. Perianth x 11 (Helsingfors, Lindberg). 10. Cross-section of perianth, near the base x 11 (ditto). 11. Ditto, near tlie apex x 11 (ditto). 12. Cilia from mouth of the perianth x 85 (ditto). 13. Perigonial bract x 24 (British Columbia, Macoun). 14. Antheridium x 31 (ditto). 27. Jungermania lycopodioides, JJallroth. Jungermanioj hjcojjodioides, Wallr. Fl. crypt. Germ. Ill, p. 70 (1831). Jungermania harhata, var. lycopodioides, Nees, Nat. Eur. Leb. 11, p. 185 (183G). Lophozia lycopodioides, Cogn. Hep. Belg. p. 31 (1872). Dioicous, csespitose, medium to largish in size, pale or dark green in colour, turning brown when old. Stems simple or fur- cate, somewhat thick, greenish or brown, prostrate, firm ; radicu- lose, rootlets whitish. Leaves semi-vertically inserted, horizontal, imbricate, orbicular or broadly subquadrate, 3, 4, rarely 5 dentate to about \, sinuses acute, segments mucronate, undulate, postical base of leaf ciliate, cilia very long, hyaline ; texture somewhat delicate, cells from small to medium size, roundish-quadrate or oblong, walls somewhat thin, trigones very distinct. Stipules large, bipartite, segments lanceolate, ciliate. Bracts subquadrate, to about \ quadridentate, sinuses acute, segments mucronate, one or two cilia near the base of bract; bracteole narrowly subquadrate or cuneate, quadridentate to about \, sinuses acute, segments mucronate. Perianth terminal, oval or oval-oblong, upper portion 5-plicate, mouth contracted, ciliolate or dentate, cilia distant, unequal. Pistillidia 8-10, cylindrical. Calyptra oval, delicate. Pedicel moderately thick, white. Capsule almost round, dark brown. Spores pale brown. Elaters bispiral, pale brown. Peri- gonial bracts 4-6 pairs, closely imbricate, smaller than stem leaves, broader than long, ventricose ; antheridia numerous, 4 or 5 JUNGEEMANIA. 347 in each bract, oval, beurer long. (Tcmnife sometimes present, brownish-red. Fruits Marcli, April. Dimensions. — Stems 1 to 2 inches long, diam. -4 mm., "5 mm., with leaves 3- mm. broad; leaves 2*5 mm. long x 2-5 mm. broad, segments T mm. long, 2' mm. x 2-25 mm., seg. '75 mm. ; basal cilia l" mm. long; cells '025 mm., '025 mm. x -03 mm., •04 mm. X '02 mm., "03 mm. ; trigones '0075 mm. ; stipule l' mm. high X 1- mm. broad, segments '075 mm. ; ])erianth 3' mm. long X L-25 mm. broad; cilia at the mouth "2 mm., "l mm., "05 mm. ; bracts 2-5 mm. long x 2*5 mm. broad, segments I'mm. ; bracteole 225 mm. long x 1*25 mm. broad, segments 75 mm. ; perigonial bract 1-25 mm. long x 175 mm. broad; antberidia '2 mm. X -275 mm. Hab.— Growing in somewhat large tufts on l)anks or walls, in hilly district-. Hare. 7, 10. 13. Near Hannahstown Bridge, New Gralloway, /. ^[cAiKfn'ir. 15. Strachan, Aberdeenshire, ./. Sim. 16. Very rare, Hoshven Hill, Moidart, West Inverness, at 2500 it. in a tuft of Saxi/rcif/a /ii/pii aides, S. M. Macvicar. Found on the Continent, in North America, and Northern Asia. Obs. — Easily separated from allied species by the large cilia at the postical base of leaf, and the mucronate leaf segments. Distinguislied from the variety F/ocrh'i by its much larger size. Description oe Plate CXLIX. — Fig. 1. Plants natural size. 2. Portion ol' young, slender stem, antical view x IG (Strachan, Sim). 3. Leaf x 1 G (ditto). 4. Cilia from postical base of leaf X 85 (ditto). 5. Portion of leaf x 200 (ditto). G. Stipule X 24 (Canada, Macoun). 7. Bract x IG (Sweden, Lindberg). 8. Bracteole x IG (ditto). 9. Perianth. 10, U. Cilia from mouth of perianth x 85 (Sweden, Lindberg). 12. Perigonial bract X 31 (ditto). 13. Antheridium x 31 (ditto). 348 HEPATIC^. 28.. Jungermania lycopodioides, IFallv., var. Floerkii {Web. et Mohr.). Jungermania FloerJdi, Web. et Mohr, Bot. Taschenb. p. 410 (1S07). Jungermania harhata, var. Floerkii, G. L. N. Syn. Hej). p. 12d (1S44:). Jungermania lycopodioides, var. Floerkii, Lindb. Muse. Scand. p. 7 (1H79). Dioicous, ci^spitose, small to medium in size, pale or dark green to greenish-brown in colour. Stems erect or procumbent, arcuate serpentine, rigid, simple or furcate ; radiciilose, rootlets whitish, few. Leaves sub- vertically inserted, horizontal, imbricate, subrotund, tridentate (very rarely bi-quadridentate) to about ^, sinuses acute or rounded, segments acute, margin of leaf entire with the exception of a small postical basal tooth ; texture some- what firm, cells smallish, roundish-quadrate, walls firm, trigones small, distinct. Stipules simple, bifid or bipartite, with one or two longish cilia or teeth near the base, segments linear-lanceolate, entire. Bracts quadrifid, undulate. Bracteole broadly ovate, bidentate. Perianth terminal, oblong, acutely plicate above. Male plant not seen. Dimensions. — Stems J to 1 J inch long, diam. "3 mm. to '4 mm., with leaves 2' mm. broad; leaves 1'2 mm. long x I'S mm. broad, segments '3 mm., 1* mm. x 1* mm., seg. -3 mm., '9 mm. x l"! mm., seg. "3 mm.; cells -03 mm., '02 mm., '03 mm. x '025 mm.; stipules '7 mm. long x '3 mm, broad at the base, '7 mm. x "25 mm., •35 mm. X *2 mm. Obs. — There is little doubt that this is merely a variety of Jung er ma Ilia lycopodioidei<. Wallr., from which it is distinguished by its smaller size, segments not mucronate, and, instead of being furnished with large basal cilia, a short simple segment only is evident. The stipules are not so ciliate. I have seen neither ^ or $, the description of bracts, bracteole and perianth being taken from Nees "Eur. Leber." A number of forms are described by Nees, the most striking being var. Nauinauniana, N., which is found with us ; except in the largeness of its size, I see nothing to separate it from var. FloerldL JUNGERMANIA. 349 Hab. — Growing on rocks tmd walls, moderately rare, iisuall}^ in subalpine localities. 5. Stream above Flash ; Itoaches, Stafibrdshire, ./, E. Jia//>ialL 7. Cwm Bj'cban, Merioneth, E. M. Holmes ; Tyn-y-Groes, Merioneth ; Barmoutb, Merioneth, /r. //. P. ; Crib Coch, Car- narvonshire, irUd ^- Holt ; Llanberis, Carnarvonshire, W. H. P. 8. Kinder Scout ; Cliarlesworth Coombs ; near Woodhead, Derby- shire, G A. Holt ; BensclifF, Leicestershire, F. T. Molt. 9. Staley Brushes, Cheshire, G. A. Hoil ; LonL2:ridi>e Fell ; Tootle Heii^ht, A^'est Lane, ./. A. Vliddon. 10. Bingley ; Baildon, Yorks., //: West. 12. Isle of Man, G. A. Holl; Staveley; Grayrigg Forest, G. 81 abler ; Kentmere, Westmorland, Rev. C. H. Bin-slead. 13. North side of Black Craig, New Gallowa}-, /. Mc Andrew \ Crichope Linn and Terragles, C. Scott. 15. Ben Lawers, G. A. Holt; Ben Mac Dhui, ./. Whitehead. L Muckish, Co. Donegal, Dr. D. Moore. Found on the Continent and in North America. Description of Plate CL. — Fig. 1, Plants natural size. 2. Portion of stem x IG (Todmorden, C A. Holt), 3-5. Leaves X 24 (Ben Mac Dhui, J. Whitehead). 6. Leaf x 24 (Wood- head, Cheshire ; Whitehead & Pearson). 7, 8. Leaves x 24 (Ben Lawers, Of. A. Holt). 9. Portion of leaf x 290 (Helsing- fors, S. 0. Lindberg). 10, 11. Stipules x 85 (Ben Lawers, G. A. Holt). 12. Stipules x 85 (Isle of Man, G. A. Holt). 13. Ditto x 24 (Ben Mac Dhui; J. Whitehead). 29. Jungermania Kunzeana, Huehener. Jungermania Kumeana, Ilueben. Hep. Germ. p. 115 (18o4). Dioicous, growing in loose patches or amongst mosses, medium in size, of a reddish-brown colour. Stems simple or dichotomously branched, ascending or erect, firm ; radiculose, rootlets long, hyaline. Leaves subhorizontal to patent-divergent, sub-vertical to obliquely inserted, bifarious, subrotund, broadly rotund or sub- quadrate, margin entire, often unidentate at the base, bifid to aljout a third, lobes erect or spreading, ovate or broatlly ovate, 350 HEPATIC^. obtuse, incurved, sinus narrow, rounded, cells small, subquadrate, 4-, 5-, and G-sided, walls thick, angles thickened, no trigones. Stipules usually bipartite almost to the base, rarely simple, seg- ments subulate, lanceolate, divergent, entire, sometimes unidentate at base. Bracts larger than the leaves, subquadrate, quadrifid, some- times unidentate near base, segments widely subulate, acuminate or acute, margin entire ; bracteole free, narrowly subquadrate, dentate- spinose near base, bifid to about the third, segments widely subulate, acuminate, entire, sinus narrow. Perianth terminal, projecting beyond the bracts, oblong-oval or oblong-ovate, terete, upper half plicate, mouth slightly constricted, laciniate, dentate or denticulate. Pistillidia about 12. Male stems more slender, leaves more erect, perigonial bracts often trifid, segments incurved, terminal or in the middle of the stem, antheridia oval. Dimensions. — Stems 1 to 1|- inches long, -3 mm. diam.,175 mm. wide with the leaves ; leaves '9 mm. high x 1* mm. broad, segments •45 mm., '9 mm. x '9 mm., seg. "3 mm,, 75 mm. x '85 mm., seg. "35 mm., "7 mm. x "6 mm., seg. '25 mm. ; cells -0175 mm., "02 mm., •0225 mm. ; stipules "4 mm. long x '275 mm. broad at base, seg- ments "35 mm., "35 mm. x -175 mm., seg. '3 mm., "35 mm. X '125 mm., ses^. '3 mm. ; bracts 1*25 mm. x 1" mm., sesrments •4 mm., '5 mm., *6 mm. ; bracteole 1"25 mm. x 65 mm., seg. •55 mm. ; perianth 2 '5 mm. long x 1-25 mm. broad ; lacinise at mouth of perianth "1 mm. long; pistillidia "2 mm. long x ■05 mm. broad. Hab. — Grows on rocks and heaths in alpine localities. Very rare. Scotch Alps, Bruminoiid (Herb. Taylor). Found in alpine districts on the Continent and in North America. Obs. — This extremely rare alpine British species is abundantly distinct from any other by its colour, shape of leaves, bracts and stipules. Description of Plate CLI. — Fig. 1. Plants natural size. 2. Portion of stem x 16 (Scotland, Druinmond, Hb. Taylor). 3. Portion of male stem x 16 (ditto). 4-6. Leaves x 24 JUNGEBMANIA. 351 (Norway, Kaalaas). 7-9. Ditto x 24 (Lapland, Hb. Lindb.). 10. Portion of leaf x 290 (Scotland, Drummond). 11. Portion of leaf, showing basal tooth x G4 (Norway, Kaalaas). 12,13. Sti- pules X 64 (ditto). 14. Stipule x 85 (Lapland, Hb. Lindb.). 15, IG. Bracts x 24 (Finland, Hb. Lindb.) 17. Bracteole x 24 (ditto). 18. Perianth x IG (Scotland, Drummond). 19. Portion of mouth of perianth x 85 (Finland, Hb. Lindb.). 20. PLstil- lidium X 85 (ditto). 30. Jungermania Helleriana, Nees. Jungermania Helleriana, Nees in Lindenb. Syn. Hep. Eur. p. 64 (1820). Diplophyllum Hellerianum, Dum. Recueil, p. 16 (1835). Jungermania verrucnlosa, var. Helleri, Lindb. Muse. Scand. p. 8 (1870). Cephalozia Ilelleri, Lindb. Medd. af Soc. F. Fenn. 14, j). 65 (1887); Kaalaas Leverme. Norge, p. 101 (189;>). Dioicous, growing in matted tufts, small, of a pale yellow, yellowish-brown or reddish-brown colour. Stems slender, flexuose, simple, or slightly branched, with slender, small-leaved innovations 23roduced from below the perianth ; densely radiculose, rootlets short. Lower leaves small, distant, upper larger and crowded near the apex, bifarious, secund above, transversely inserted, semi- amplexicaul, subquadrate, ovate or oval in outline, divided to about I" into two equal acute, sometimes cuspidate, rarely obtuse segments, sinus acute, narrow; cells small, roundish, roundish- oblong or oblong-quadrate, guttulate, cell-walls thick, angles thickened, no trigones, upper leaves and perianth distinctly verru- cose. Stipules rarely present, subulate. Bracts larger than the leaves, subquadrate, 2-3-lobed, segments acute, spinose-dentate, sinus acute, sub-bracts bifid to about J or -J-, segments acute, margin entire. Bracteole simple, narrowly oblong, subquadrate, laciniate, sparingly dentate, or obovate, subquadrate, bifid, dentate, connate on one side to adjacent bract or free. Perianth projectino- about |- beyond the bracts, ovate or oval, slightly plicate above, mouth a little constricted, laciniate-ciliate or laciniate-dentate. Male stems rather more slender, perigonial bracts middle or end 352 HEPATIC^. of the stem, several pairs, ventricose ; antlieridia single, roundish, bearers of equal length. Gemmiparous ; gemmiparous stems with leaves closely imbri- cate, erect, bifid or sub-entire, erose, gemmas small, quadrate, crimson to purple, terminal in buds. Dimensions. — Stems about ^ inch long, '1 mm. diam., with leaves 'o mm. broad; upper leaves '45 mm. x -35 mm., segments •175 mm., '45 mm. x '3 mm., seg. '15 mm., "375 inm. x '25 mm., seg. '1 mm.; middle leaves '25 mm. x '2 mm,, seg. '125 mm.; lower leaves "2 mm. x '15 mm., seg. '075 mm. ; cells "03 mm. x •02 mm., "025 mm. x "02 mm., '02 mm. x "02 mm., "02 mm. x •0175 mm., '02 mm. x '015 mm., '0175 mm. x "015 mm.; sub- bract '4 mm. x -375 mm., seg. "125 mm. ; bracts '45 mm. x -35 mm., seg. "275 mm. ; bracteole -45 mm. x "3 mm., seg. "275 mm. ; perianth 1" mm. x '5 mm. ; lacinise at mouth of perianth ^075 mm. ; pistillidia -125 mm. x "025 mm. Hab. — Growing on decaying wood, in subalpine districts. Very rare. Discovered in this country by 31 r. George Stabler. 12. Mardale, Westmorland, May 12, 1882, Georr/e Stabler. 15. Balmoral, April 16, 1884, /. Micliie ; on dead trunk oi Pinus sylvestris, Braemar, July, 1884, G. Stabler. Found on the Continent and in North America. Obs. — This very rare and small species is not likely to be confounded with any other; it grows on rotting wood, which is its only habitat. Lindberg reduced it to a variety of hhJiuu/. verrucnlosa(l874:), but as it is a very distinct and full}^ developed species, I do not see any reason why Nees's name Ji/n//. Helleriana (1829) should have been superseded. Later (Medd. Soc. F. Fenn. 14, p. 65 (1887), be removed it to CephaJozia, in which he has been followed b}^ Dr. Kaalaas. Description of Plate CLII. — Fig. 1. Plants natural size. 2. Fertile stem x 31 (Herb. Lindb.). 3, 4. Upper leaves x 64 (ditto). 5-8. Middle and lower leaves x 64 (ditto). 9-11. Upper leaves x 64 (Braemar, G. Stabler). 12. Erose leaf x 64 (ditto). 13. Portion of leaf x 290 (Herb. Lindb.). 14, 15. JUXnEnMAXfA. 353 Stlpiilos free (Canada, Macoiin). HJ, 17. Sub-bracts x 04 (llcrl). Liiidb.). IH. IJracts and connate braeteole x 04 (ditto). \\). liracteole x 1 (ditto). I'O. IVrianth x 31 (ditto). •J I. Portion of nioutli of })erianth x 85 (ditto). l22. Pistil- lidiuni x 85 (ditto). Subgenus 4. SPHEXOLOBTiS, fJudh. Junffetin/inia, sect. c. S phenol oh m^, \j\\\([h. Mu.sr. Sriind. p. '.• (1H7;)). Dioicous, stems simple, furcate or dichotomonsly branclied, leaves transversely inserted, bilobcd ; stij)ule.s absent; bracts ((uadriHd ; })eriantli oval or obovate, upper portion 5-, G-plicate. 31. Jungermania minuta, Cranlz. Jnnget'mania minuta, Crantz ex Dicks. Pi. crypt. Brit. fasc. 1 1 , p. 1:! (17!l(»); Hook. Brit. Jung. t. 44 (IHIC); Sm. Eng. Bot. t. 22.; I. JiuKjermania hiconiis, Web. et Mohr Bot. Taschenb. p. 12.') (1S07). Juivjernuinia Weheri, Mart. Fl. crypt. Erlang. p. l")?, t. .'), f. W.) (1H17). DiplophjiUnm minnttnn, Dum. Recueil, p. K! (IH;*),")). Dioicous, growing in loose or entangled patches, or amongst mosses, small to medium in size, of a brownish or yellowish-green colour. Stems creeping, suberect or erect, simple or dichotomously branched, branches produced from the axils of the leaves, between lateral and postical, more postical, filiform, rigid, fiexuose, some- what brittle when dry, cortical and next inner layer of cells dark brown, inner white, indistinct, small ; radiculose, rootlets few, long, hyaline. Leaves bifarious, distant, contiguous or imbricate, pec- tinate, transverse or patent-divergent (70 ), outline subcjuadrate or ovate when flattened out, subconduplicate, margin <|uite entire, bilobed to about v^, sinus acute or rounded, segments unequal, antical slightly smaller, acute or rarely obtuse, sometimes apicu- lato; texture iirm, e])iilermis slightly verruculose, cells minute to rather minute, subcpuidrate, near base of loaf st)mctiines a little elongate, walls thick, no trigones, guttulate. No stipules. Jiracts larger than the loaves, two, (puKlriful om- bract often with a large 354 HEPATIC JE. tooth near the base, sometimes one bract will be quadrifid and the other only bifid, divided to about the middle, rarely only to about ^ or J, segments lanceolate, acutate or acute, margin quite entire. Perianth projecting beyond the bracts, oval, terete, upper half 5-plicate, composed of a single layer of cells, about 150 cells round, mouth constricted, laciniate-denticulate. Spores pale brown, elaters dark reddish-brown. Male stems similar to others, perigonial bracts middle of stem, concave, ventricose at base, 3 to 4 pairs, antheridia roundish-oval, with long bearers. Gemmiparous, gemmae whitish. Fruits May. Dimensions.— Stems from |- to Ij inch long, '125 mm. diam., with leaves 1' mm. wide ; leaves "8 mm. x '7 mm., segments '4 mm., •75 mm. X '5 mm., seg. '3 mm., "8 mm. x '6 mm., seg. "4 mm. ; cells •015 mm.; bracts 1"1 mm. broad x "95 mm. high, "95 mm. x -95 mm., seg. '5 mm. ; perianth 2'35 mm. x 75 mm. ; spores •01 mm. diam. ; elaters 'I mm. long x '01 mm. broad. Hab. — On rocks and heaths in alpine and subalpine districts. Somewhat rare. 7. Borthwen, Rev. T. Sahraj/ ; Cwm Bychan, Merionethshire, jr. H. p. 8. Kinder Scout, Derbyshire, Wkifehead 8^' Holt, 9. Clougha, West Lane, Wheldou 8f A. Wilson. 10. Near Green- field, //". Wilson; Eibblehead, Br. F. A. Lees; Haworth, Dr. Car- rlnfjion; near Todmorden, /o//y/ iV(:>2d;(:'//. 11,12. Mardale ; Brunt Knot; Kentmere, Westmorland, G. Stabler. 13. Frequent on banks and on the hills, New Galloway, /. McAndreiv. 15. Strachan, Aberdeenshire, /. 8im ; Ben Lawers, C. J. Wild. 16. Common among rocks in peaty soil, rare on the hills, where it was observed at 1800 ft. Moidart, West Inverness, 8. M. Macvicar. I. Mangerton, Dr. Carrington ; Loch Bray, Co. Wicklow, Dr. D. Moore, D. M'Ardle, cum jjer. ; Seven Churches, Co. Wick- low, Dr. D. Moore. Found on the Continent, North America, Mexico, Africa. Obs — This species is distinguished from Jiint/. saxicola by its smaller cells, in addition to the other characters mentioned under JUNGEliMANIA. 355 that species. In its sterile state it may be confounded with some Ibrms of Marsi/pel/a ciitartj'nKila or MarHiipclhi Funckii, but if l)racts be found it is at once identified. Description or Plate CLIII. — Fig. 1. Plants natural size. 2. Portion of stem x 31 (G. & P. G29). 3-0. Leaves, flattened out, X 32 (Loch Bray, M'Ardle). 7. Leaves x 31 (ditto). 8, 9. Leaves, flattened out, x 31 (G. &. P. 029). 10. Leaf x 31 (Loch Bray, M'Ardle). 11. Portion of leaf x 290 (G. & R. G29). 12,13. Bracts x 31 (Loch Bray, M'Ardle). 14. Perianth x 24 (ditto). 15. Cross-section of upper part of perianth x 31 (ditto). 16. Portion of the mouth of perianth x 85 (ditto). 17. Peri- o'onial bract free, after Carrino-ton. 32. Jungermania saxicola, Sc/irnder. Jungermania saxicola, Schrad. Syst. Samml. krypt. Gew. n. '.•7 (1797). Diplo'phyllum saxicolum, Dum. Recueil p. 10 (1835). Dioicous, growing in thick patches or amongst mosses, medium to largish size, of a brown, yellowish, red-brown, or rarely of an olive-green colour. Stems simple or furcate, somewhat robust, firm, branches proceeding from below the leaves, sub-postical, between lateral and postical, but more postical ; about 15 cells in diam. ; cortical cells dark and firm, about 40 to 50 cells round, inner white, somewhat firm and distinct, walls firm ; radiculose, rootlets long, close, hyaline, some stems destitute. Leaves hori- zontally inserted, equitant, imbricate or contiguous, regular, almost of equal size, somewhat oblique, amplexicaul, not decurrent, sub- conduplicate, broadly ovate, bifid to the middle, rarely trifid, lobes unequal, antical a little smaller than the postical, segments sub- acute, obtuse or roundish, margin quite entire ; cells smallish, roundish, roundish-oblong or subquadrate, cell walls thick, angles thickened. No stipules. Bracts larger than the leaves, 2 to 4- lobed to near the middle, segments acute, dentate. Bracteole bifid to below the middle, segments acute, dentate ; sub-bracts broadly ovate or subquadrate, bifid to about the middle, segments roundish, irregularly dentate; sub-bracteole very small, obovate, entire, or refuse. Perianth projecting about half beyond the 356 HEPATIC.^. bracts, oblong-ovate or obovate, cylindrical, upper part plicate, mouth contracted, laciniate-dentate, about 30 fine teeth, pistillidia very numerous. Perigouial stems more slender, growing in separate tufts, perigonial bracts terminal, containing 1 to 3 antheridia. Dimensions. — Stem 1 to 2 inches long, diam. "2 mm., with leaves 1"25 mm. broad; leaves, antical lobe "75 mm. x "55 mm., postical lobe "85 mm. x '55 mm., antical 75 mm. x "5 mm., postical *9 mm. X 7 mm.; cells "025 mm., "03 mm., "04 mm. x "02 mm. ; sub-bract 1"25 mm. x 1*25 mm. (both lobes); sub-bracteole •4 mm. X '35 mm. ; bracts 15 mm. x 1-25 mm., segments "75 mm. ; bracteole 1' mm. x '55 mm., segments 6 mm. ; perianth 2 "2 5 mm. x 11 mm. Hab. — An alpine species, growing on rocks or amongst mosses; very rare. Ehonas Hill, Shetland, Dr. GreviUe. Found on the Continent and in North America. Obs. — This extremely rare British species is not likely to be confounded with any other native hepatic. From Jun^. minuta, Crantz, its nearest congener, it is distinguished by its robustness, the more obtuse lobes of the leaves and its dentate bracts ; from Jung. Kunzeana, Hiib., by the absence of stipules, &c. Description of Plate CLIV. — Fig. 1. Plants natural size. 2. Portion of stem x 24 (Shetland, Greville). 3, 4. Leaves x 24 (Finland, Lindberg). 5, 6. Leaves, flattened out, x 24 (ditto). 7. Portion of leaf x 290 (ditto). 8. Sub-bracts and sub-brac- teole X 24 (ditto). 9, 10. Bracts x 24 (ditto). 11. Brac- teole X 24 (ditto). 12. Perianth x IG (ditto). 13. Cross-sec- tion of perianth X 16 (ditto). 14. Mouth of perianth, flattened out, X 16 (ditto). Subgenus 5. JNASTREPTA, Lindb. Junger mania, sect. Anastrejita, Lindb. Bot. Not. (1889). Dioicous, plants moderate size, csespitose, or creeping singly amongst mosses; stem erect, radiculose, simple or innovantly JUNGERMANIA. 357 branched ; leaves obliquely inserted, enuirginate, margin remark- ably recurved ; stipules absent ; perianth laterally compressed, upper portion plicate, mouth constricted. 33. Jungermania orcadensis, Hook. Jimyermania orcadensis, Hook. Brit. Jung. t. 71 (IMl(l). Mesophylla (?) orcadensis, Dum. 8yll. Jung. p. 80 (IH:;!). Mesophylla orcmlensis, Dum. Hep. Eur. [>. I'M) (1S71). Dioicous, loosely csespitose, largish, brownish-green or dark reddish-brown in colour. Stems simple, or with a few innovant branches arising from the postical side of axil of leaf-subpostical, or truly postical, arising from the postical side of stem ; outer layers of stem dark coloured, inner white ; erect or ascending, flexuose, filiform, reddish-brown in colour; radiculose, rootlets short, dense, ascending to apex of stem, dull white. Leaves obliquely semivertically inserted, closely imbricate, or when young, smaller and approximate, horizontal or slightly ascending, spreading or secund, semi-amplexicaul, antical margin decurrent, broadly ovate to cordate, emarginate, sinus shallow, rounded, segments acute, margin entire, upper (remarkably so) and lower recurved, concave at base ; texture somewhat thin but firm ; epidermis smooth; cells small, roundish to roundish-oblong, guttulate, walls thick, angles thickened, no trigones. Stipules wanting, except where the innovant branches arise, irregular in shape, much broader than long, usually 4-laciniate, the two middle segments often long and acute. Bracts 2, large, recurved above, the outer one embracing the inner, usually trilobate (2-4), sinus somewhat acute, segments more or less acute. Bracteole large, very broad, surrounding the lower half of the perianth, deeply 4-laciniate (often with G lacinice), segments unequal, some triangular, others smaller and more acute, the smallest and most acute at the margins. Perianth projecting somewhat beyond the bracts, slightly arcuate, narrowly oblong (below somewhat narrower than above), distinctly laterally compressed, upper third portion deeply plicate, with 5, sometimes more, rounded fold.*^, 358 HEPATIGjE. which are not all equally deep and perfect ; mouth constricted, ciliate (cilia 3 cells long), between the cilia are short teeth. Andrcecia situated on the middle of stem, 3-5 pairs of perigonial bracts, which are very different from the stem leaves, they are hardly half the size, saccate at the base, obtusely bilobed, lobes broad and obtuse ; antheridia 2, shortly stipitate. Dimensions. — Stems 2 to 3 inches long, diam. "3 mm., with leaves 2* mm. broad; leaves 1*25 mm. long x 1"1 mm. broad, segments '3 mm., 1-2 mm. x '9 mm., seg. '2 mm. ; cells '02 mm., •0225 mm. x -0125 mm. Hab. — Grrowing in loosely matted patches or straggling amongst other mosses and hepatics, in alpine or subalpine locali- ties. Kare. 7. Snowdon, Carnarvonshire, W. Wilson. 12. Ambleside, West- morland, C. Li/cIl ; Naddle Forest, &c., Westmorland, G. Stabler. 15. Clova Mountains, Dr. Greville ; Ben Mac Dhui, W. JFest. 16. Catlow Hills, Kinnordy, C. Lyell; Loch Maree, I)r. Carringto7i ; Glen Finnan, Dr. Carrington; Moidart, West Inverness, S. M. Alac clear. 183. Orkney Isles, Dr. Hooker. I. Brandon Mountain, Dr. Taylor. Found on the Continent. The Jung, oreaclensis recorded from the Himalaya is a different species, insertion of leaves more transverse, different in shape, segments more obtuse, cells larger, &c. Obs. — This is a very distinct species, easily recognised by the remarkably recurved margins of the leaves. No ^ or $ plants have yet been met with in this country. Dr. E. Jorgensen has found them in Norway and I have taken my description and figures of them fi'om his interesting paper in ■'Bergens Museums Aarbog," 1894-95, No. xviii. Description of Plate CLV. — Fig. 1. Plants natural size. 2. Portion of stem, antical view x 16. 3. Portion of stem, postical view x 24. 4. Leaf x 24. 5. Portion of leaf x 290. (All from original specimen, Clova, Herb. Hooker.) 6. Stipule X 30. 7. Bract x 30. 8, 9. Bracteoles x 30. 10. Perianth X 30. (After E. Jorgensen, Berg. Mus. Aar. 1894-5, No. xviii.) JUNGERMANIA. 359 Subgenus G. CEPBALOZIOPSIS, Spruce. Jungermania, subgenus LopJiozia, sect. CejihaloziojJsis, Spruce, Hep. Am. et And. p. 511 (1885). Plants small, leaves minute, somewhat similar to CophaJozia but ramification different and perianth, which when only trii^onous has the third angle antical ; stipules normally absent ; flowers monandrous. 34. Jungermania Pearsoni, Spruce. Jungermauia Pearsoni, Spruce, Journal of Botany, n. s. vol. 10 (1881). Dioicous, depresso-csespitose, small, lurid green to reddish- brown in colour, apex often with a reddish-purple tinge. Stems flexuose, flaccid, when dry somewhat rigid and brittle, once or twice (rarely more) furcate, sometimes swollen at apex, no pos- tical branches ; radiculose, rootlets few. Leaves almost equal the whole length of the stem, distant, rarely subcontiguous, hori- zontal to patent-divergent, broadly cuneate or subquadrate, seg- ments suberect, subparallel with the stem, somewhat plane, ovate, subacuminate, acute or subobtuse, sometimes the antical lobe is furnished with a small tooth at the base, otherwise margin entire ; texture firm, epidermis slightly rough, cells very small, quadrate or a little elongate, opaque or subpellucid, walls thick, no trigones or thickened angles. Stipules subulate-linear, usually more fre- quent near the ap3x, sometimes wanting. Female inflorescence not met with. Androecia on the middle of the stem or branches, perigonial bracts larger than the adjoining leaves, swollen at the base, some- times with an incurved antical tooth at the base, antheridia roundish-oval, solitary, stipe long. Dimensions. — Stems ^ to J^ inch long, with leaves i mm. broad, diam. '05 mm. to '075 mm. ; leaves *35 mm. long x "3 mm. broad, segments '175 mm., '3 mm. x '275 mm., seg. '15 mm., '3 mm. x •225 mm., seg. '125 mm. ; cells "015 mm. x "02 mm., -015 mm., 360 HBFATICJ^. •01 mm.; stipules "175 mm. long x "03 mm. broad at the base, •225 mm. x "02 mm. ; perigonial bract '4 mm. long x '3 mm. broad ; antheridia '125 mm. x '11 mm., ^125 mm. x '1 mm. Hab. — Growing in the interstices, or on the shady side ot rocks, in subalpine localities. Very rare. 7. Glyder Vawr, W. H. P. May 1877 ; on boulders interspersed with the immense broken rocks, at the foot of Twl Dhu, Cwm Idvval, Carnarvonshire, W. H. P. August 1884. 12. Brown Ghyll, Langdale, Westmorland, Gcor(/e Stabler §• TF. H. P. 1881 ; Bleak Eigg, Easdale, Westmorland, Rev. C. H. Binsiead, 1886. 13. North side of Black Crag, New Galloway, /. Mc Andrew, 1884. 16. On a travelled granite boulder in a ravine, Eesipol, Sunart, West Inverness, S. M. Macvicar Sf IF. H. P. 1899. Obs. — " Somewhat similar in appearance to Ceplialozia divari- eafa (Sm.) but larger. It is at once distinguished from that and every other species of Cephalozia by the forked branching, and the entire absence of the postical branches characteristic of CejjJialozia. From MarsKjjella {Sarcoscj/jj///fs) it differs in wanting the rhizoma- tous base, in the ramification, the deeply cloven leaves and their texture, and (so far as I have seen) in the absence of oil-granules from the cells. As we have only the male plant, wliich has monandrous bracts, its place among other bifid-leaved Juvfjer- mani(B is not easy to assign. Tt is not unHke small forms of /. minuta, Schrad., and /. rif/ida, Lindberg ; but its nearest allies are doubtless /. opacida, n. sp., gathered by myself in the Andes, between Eioamba and the Plateau of Chimborazo, on the rocks shattered by the great earthquake of 1796 ; and /. intricata, Lindenb. et G., found by Liebmann in Mexico, near Oaxaca, where it grew closely interwoven with /. eolorafaT — Dr. Spruce. The description is taken from notes by Dr. Spruce in " Journ. of Bot.'Teb. 1881. Description of Plate CLVI. — Fig. 1. Plants natural size. 2. Portion of stem, antical view, x 85 (Glyder Vawr,- May 1877, W.H. P.). 3. Leaf x 85 (Easdale, Eev. C. H. Binstead). 4. Leaf, with stipule x 85 (ditto). 5. Leaf, explanate, and K'- jrX(;Ki:MAXi.\. nr;i stipulo X 85 (ditto), (j-l.'i. Ijcaves, (.'Xijlaniito, x Hij (/ii/Uuiu, ii\n-uce, .Umvu. oi Hot. p. 23'j (1>s7<>); llt'p. Am. ft And. p. 51:5 (Ish:,). Plants large, ruleseent or puri)urescent, assurgent, slightly ramose, subradiculo.>e, rarely Hagelliferous. Leaves large, succu- bous, a.ssurgeiit-secund, ovate, concave, apex bifid (very rarely entire), segments unequal, antical slightly smaller, more or less acute, often incano-cuspidate, (juite entire or denticulate, antical base sometimes laciniate. Cells small, auglrs distinctly thickened, cuticule scaberulous. Stipules absent. InHorescence dioicous. Female flowers terminal, rarely innovant. IJracts $ more deeply divided than the leaves (rarely :> t-liil) denticulate or at both sides of the base sparsely laciniate. Perianth rosy-purple, apex canescent, large, elongate, when young subulate, when mature fusiform or below narrowly cylindrical, Irom almost the base deeply 3-, 5-, or 8-plicate, mouth constricted, fimbriate. Calyptra delicate. Capsule shortly pedicellate, oblong-cylindrical, rarely subMobose, dividinfj to about the base into 4 valves, valves sometimes biiid. Flowers ^ L'-4-androus. 35. Jungermania Doniana, //oo^: Jmujerniduia Donniana, Hook. Hrit. Jung. t. ;!'.• (l^lil). I'lenroschi.tma /huiiiaiia, Duni. Syll. Jung. p. 7- (ls>*I). Juwjermania {AimstrophyUum) /./ojmVimm, Spruce, Journ. of l5ot. p. !';;."• (1>>7(I); Hep. Am. et Ami. p. T.l;! (1X85). JiiiDjcniutnia /kniii, HtMik. Kaalaas Ix'Vi'ini. Noigc, p. .17 H (I s'.i;'.). l>i()icous, c.-i'spitose, medium to laiLTc in size, reddish-brown to dark brown in colour. Stems simi»le or slightly branched. 362 HEPATW/E. branches few, postical-lateral, arising from the postical axil of leaf; a cross-section shows 12 by 16 cells in diam., with 40 to 50 cortical, all somewhat similar, walls firm, dark ; flexuose, erect or procumbent, rigid and brittle when dry, somewhat ligneous, opaque, rootlets few. Leaves closely imbricate or approximate, subhorizontal or slightly ascending, semi-obliquely inserted, bifarious, subsecund, slightly falcate, succubous, concave, canalicu- late, ovate-acute or oblong-ovate-acute, apex bifid to about y\y, sinus acute, segments acute, unequal, margin incurved, quite entire; texture firm, cells guttulate, rather minute, sinuate, roundish-hexagonal, elongate and larger near the middle base, lumen containing a few nucleate bodies, cell walls very thick, pale golden brown, no trigones. Stipules wanting. Bracts usually a little larger than the leaves, broadly ovate, acute, bifid from j^j to \, sinus acute, segments acute and unequal, margin quite entire. No bracteole. Perianth projecting about f beyond the bracts, terminal, reddish-brown, cylindrical or oblong-obovate, terete below, 5-7-plicate above, mouth contracted, laciniate-ciliate, cilia hyaline, delicate, 2-4 cells long. Male plants not seen. Dimensions. — Stems 1 to 3 inches long, diam. '2 mm. to "3 mm., with leaves 3* mm. wide ; leaves 1'4 mm. long x 1* mm. broad, seg- ments "1 mm., 1"3 mm. x 1*2 mm., seg. -125 and "15 mm. ; cells •02 mm. X '01 mm., -01 mm., '03 mm. x '0125 mm. ; bracts 1"4 mm. long X ri mm. broad, segment "2 mm., 1 '3 mm. x 1*2 mm., seg. •15 mm. ; perianth 2 25 mm. long x -75 mm. broad, 2* mm. long x "85 mm. broad; cilia '1 mm. long; pistillidia '2 mm. long x •06 mm. broad. Hab. — Growing in rather small and loosely entangled tufts in alpine situations. Eare. 15. Rocks above Loch Avon, Clova, with perianths, Br. Greville ; Braemar, A. Croall ; Ben Mac Dhui, G. Stabler. 17? Europe, Danubian Provinces, vide Dr. Spruce. Norway, Kaalaas. Obs. — A very distinct species and not likely to be mistaken for any other British one ; the dark colour of the plant with the NARDIA. 363 remarkably incurved margin of the leaves, remove it far from any other of the British species. Description of Plate CLVII. — Fig. 1. Phmts natural size (Eng. Bot.). '1. Portion of fertile stem x IG (Clova, Br. Grreville). o, 4. Leaves x l>4 (Ben Mac Dliui, CI. Stabler.) 5. Portion of leaf x 290 (ditto). G, 7. Bract x 24 (Clova, Dr. Greville). 8. Perianth x 24 (ditto). 9. Cross-section of perianth, lower portion, x 24 (ditto). 10. Ditto, upper portion (ditto). 11, 12. Portions of the mouth of perianth x 85 (ditto). 13. Pistillidium x 85 (ditto). Genus 32. NARDIA, Gr. §• Bcjui. Jimgermania, Schiad. Syst. Samml. crypt. Gew. 2, p. 4 (1 7'.>7) ; Sm. Engl. Bot. 21 (1805); Lyell in Hook. Brit. Jung. t. 03 (1810); Nees Hep. Eur. 1, p. 27.") (1833). N'ardia, Gray & Benn. Nat. Arr. Brit. PI. p. C>[)i (1821). Meso2)hi/lla, Dum. Comm. Bot. p. 112 (1822). Alicularia, Cord, in Opiz Beitr. 1, p. 052 (1820), Southhya, Spruce, Muse. Pyr. in Trans. Bot. Soc. Edinb. 5, 3, p. 197 (1850). Plants medium size, rarely large, csespitose. Stems prostrate or suberect, simple or furcate, branches proceeding from the postical angle of the leaves, rootlets usually numerous, pale or reddish, flagella absent, except in N. comprcssa. Leaves alternate, succubous, or somewhat close and subtransverse, orbicular or reniform, entire or refuse, very rarely bilobed, margin quite entire ; cells medium size, subquadrate, walls rarely thickened, lumen sometimes filled with chlorophyllose granules. Stipules present, subulate or lanceolate ; in the subgenus Euccdij,v absent. Inflorescence dioicous or paroicous; female flowers terminal, often sterile on innovations. Involucre (in subgenus Eaiiardki) urceolate ; bracts 2-5-pairs, opposite, connate with the bracteole. Perianth (in Einiardia) slightly laterally compressed and concrete with the innermost bracts and connective tissue of the receptacle to form the urceolate involucre, apex free, ovoid or conoid, obscurely 3-5- carinate, apex often at first closed, afterwards dividing into 364 HBPATIO^, several lobes. Calyptra free, obovate, delicate, surrounded at its base by 10-12 sterile pistillidia. Capsule shortly pedicellate, oval-globose, coriaceous, dividing to about the base into 4 valves. Elaters moderately long, bispiral. Spores small, asperulous. Androecia subspicate, on the middle or end of stem or branch ; perigonial bracts 6-10 pairs, somewhat similar to the leaves, 2-4-androus. Subgenus 1. EUCALYX, Lindh. Nardia, sect. 1 Eiicalyx, Lindb. in Bot. Not. p. 167 (1872). Stipules absent. Bracts 1-2 pairs, innermost adnate at the base to the perianth. Perianth projecting, moderately firm, acutely 3-8-carinate, mouth constricted, sometimes tubular. 1. Nardia hyalina [Li/ell), Carr. Jungermania hyalina, Lyell in Hook. Brit. Jung, t, G3 (1816). Aplozia hyalina, Dum. Hep. Eur. p. 58 (1874). N'ardia hyalina, Carr. Brit. Hep. p. 35, pi. 11, f. 36 (1874). Nardia [Eucahjx) hyalina, Lindb, Muse. Scand. p. 8 (1879). Dioicous, broadly csespitose, small to medium size, of a light green colour, often witli a rosy tinge. Stems procumbent or suberect, simple or innovantly branched, branches lateral, pro- duced from the axil of the leaves or bracts ; radiculose, rootlets claret or dull purple coloured, very rarely dull white, copious, long, produced from the postical side of stem, base of the leaves and perianth. Leaves obliquely inserted, horizontal, subimbri- cate, contiguous or distant and smaller below, broadly rotund, orbicular or broadly oval, semi-amplexicaul, sometimes subcompli- cate, undulate or plane, margin entire ; texture somewhat tender ; cells largish, roundish, walls thin, trigones distinct. Stipules absent. Bracts larger than the leaves, adnate to the lower portion of perianth, sometimes almost free, erect, enclosing perianth, subrotund, entire. Perianth oval, oblong or ovate, upper portion 4-6-plicate, contracted, mouth small, margin NABBTA. 3G5 composed of elongate, hyaline cells. Pistillidia 15-20. Capsule oval, dark brown. Spores pale brown. Elaters bispiral, brown. Male stems slender, distinct, perigonial bracts smaller, antical side incurved, saccate at the base, $ all the length, antheridia 2-3 in each bract, oval, stipitate. Fruits March, April. Dimensions. — Stems |- to 1 inch long, diam. '25 mm., with leaves 2* mm. to 3* mm. broad; leaves 1"4 mm. long x 1*6 mm. broad, 1*3 mm. x TG mm., 1'4 mm. x 1-2 mm., 12 mm. x 1-3 mm., I'l mm. X -1 mm. ; cells '05 mm.; trigones '01 mm.; bract 1*G mm. high X 2-25 mm. broad ; perianth 2- mm. x 75 mm., 175 mm. X 75 mm. ; pistillidia 'IS mm. long x "03 mm. broad; perigonial bracts 7 mm. x 7 mm. Hab.— Growing in large patches on damp, clayey banks, usually near streams, sometimes on wet rocks. Moderately rare. 1. Lower Ninnis, Madron, Cornwall, E. D. MarquancL 2. New Forest, Hants, C. LijeJL 5. Dove Dale ; Star Wood, Oakamore ; Chartley Moss, Staffordshire, /. E. Bagucdl. 7. Near Snowdon, Carnarvonshire, IV. Wilson ; Dolgelly, Merionethshire, 7K IL F. 8, 9, 10. Eskdale, Br. Spruce; Banks of the Eye, /. G. Baker; Hebden Bridge, G. E. Uuid. Whitby ; Ingleboro, Dr. Carrington. Cleveland, W. Mudd. 12. Stock Ghyll Force, Ambleside, West- morland, C. Lyell ; Naddle Forest; Long Sleddale; Kentmere ; Barrow Field, Helsington, G. Stabler; Scandale, Westmorland, IF. JFesf. 13. Eouten Bridge; Closeburn ; near MofFat Spa Well, /. Cruicl-i^lianh. 15. Dumbarton, A. McKinlcuj ; Glen Esk, Forfar, Br. Greville ; Braemar, A. Croall ; Kinnordy, C. Lyell. 16. Not uncommon on damp loamy banks and at the side of ravines, Moidart, West Inverness, S. M. Macvivar. I. Aooreagh River near Sneem, Co. Kerry, Br. Taylor; Brandon, W. Wihoii ; Luggielaw and Seven Churches, Dr. ]). ^foorc ; Slieve Glah, Co. Cavan, B. M'Ardle; Seefing Mountain, Dublin, Br. B. Moore. Found on the Continent, in the Canary Islands and North America ; Br. Spn/cc also found it in South America. Ol3S. — The only other round-leaved Jinn/crma/iia which has 366 HEPATIGAi. coloured rootlets is Ncirdia [Ei/calj/x) obovata (Nees), from which it is distinguished by its dioicous inflorescence, broader leaves and bracts, and slightly larger cells ; from NarcJla scalarls (Schrad.) by absence of stipules ; from other round-leaved species by its bracts being adnata with the lower half of the perianth. Jungermania crenulafa, Sm., has been placed by Lindberg and others in the section Eiicalyx of Nardia on account of the some- times, although very rarely, sliglit attachment of the bracts to the lower portion of the perianth, this so rarely happens and is so slight that I have placed it next to Jmif/. spJ/aerocarjja, Hook., in the subgenus Aplozhi o^i Jung er mania, which seems to me to be its most natural position. Description of Plate CLVIII. — Fig 1. Plants natural size. 2. Portion of stem, antical view x 16 (Original, Lyell). 3. Portion of fertile stem, antical view, x 24 (Salem, Jack.). 4-7. Leaves x 16 (Original, Lyell). 8. Leaf x 24 (Finland, Lindberg). 9. Portion of leaf x 290 (Original, Lyell). 10. Bract X 16 (ditto). 11. Perianth x 24 (ditto). 12. Pistilhdium x 85 (Finland, Lindberg). 13. Perigonial bracts x 24 (Salem, Jack.). 2. Nardia obovata {Nees), Carr. Jungermania obovata, Nees, Nat. Eur. Leb. 1 p. 332 (1833). Eucahjx obovata, Lindb. Bot. Not. (1872). Southbija obovata, Dum. Hep. Eur. p. 133 (1874). Nardia obovata, Carr. Brit. Hep. p. 32, pi. 11, f. 3.') (1874). Nardia [Eucahjx) obovata, Lindb. Muse. Scand. p. 8 (1879). Paroicous, csespitose, flagelliferous ; flagella small, radiculose ; small, of a dark green colour. Stems procumbent, or suberect, simple or ramose ; branches lateral, subpostical, proceeding from the axil of the leaves or bracts ; radiculose, rootlets purple in colour, short, ascending to apex of stem, often arising from base of leaves and postical side of the perianth. Leaves obliquely inserted, horizontal, slightly decurrent antically, alternate, im- bricate or contiguous, gradually becoming more distant and smaller lower down, in sterile stems they become smaller and XARDTA. 367 more distant near to the apex also, plane or slightly concave, upper leaves on fertile stems subcomplicate, obovate or ovate, margin entire ; texture somewhat flaccid, cells roundish or sub- quadrate, of medium size, chlorophyllose, walls thin, trigones distinct, marginal cells quadrate. Stipules absent. Bracts similar to the leaves only larger, erect, appressed, adnate to the lower half of the perianth. Bracteole absent. Perianth almost hidden by the bracts, oval to obovate, upper portion 5-6-plicate, cells elongate, mouth contorted, contracted, small, laciniate, denticulate. Pistillidia 15-30. Male bracts below the perianth, more complicate, saccate at base, antheridia oval. Fruits March, April. Dimensions. — Stems |- to 1 inch long, diam. '2 mm. to -3 mm. with leaves 2" mm. to 2'5 mm. wide ; leaves 1"3 mm. lono- x T mm. o broad, V2 mm. x '9 mm., I'l mm. x -Qmm. ; cells '025 mm. ,'03 mm., •03 mm. X *04 mm. ; tris^ones, "01 mm. ; bracts 175 mm. Ions: x r25 mm. broad; perianth 2*25 mm. long x 125 mm. broad, 2- mm. X 1' mm. ; spores '02 mm. diam. ; elaters '15 mm, long x '01 mm. broad; perigonial bracts '9 mm. x '7 mm.; antheridia •11 mm. X '1 mm. Hab. — Grows on shaded wet rocks, usually in subalpine districts. Rare. 7. Dolgelly, Merioneth, Dr. Curnngton ^- W. 11. P. ; Snowdon, Carnarvonshire, IF. Wihoii ; Llanberis, Carnarvon- shire, G.E. ILiiit. 9. Bamford Wood; Clifton Junction, Lane, G, A. lIoJl ; Longridge Fell, West Lane, J. A. Wltchhii ; Ease- gill, near Leek, West Lane, A. Wilson. 10. Kildale, N. York., W. Mudd. 12. Kentmere; Little Langdale ; Harter Fell; Mardale; Long Sleddle, Westmorland, C. /^/^(^/c^;-. 13. Opposite Waulk Mill, New Galloway, ./. Mc Andrew. 15. Glen Esk, A. Croall ; Banks of Forfar Burn, A. Croall ; Strachan, Aberdeen- shire, ./. Sim. IG. Common on wet rocks, especially at the sides of streams, ascends to 2500 feet, Moidart, West Inverness, S. M. Macvicar. L KiUarney, IF. Wihon ; Tore WaterfVill, IF. WUson, Br. Car- ritigton ; Ravine near Hunting Tower, Cromaglown, Dr. Carriiifj- 368 HEPATIC.^. ton; Connor Hill, Co. Kerry, Br. D. Moore; Kylemore, Co. Galwa}^ Br. B. Moore; Loch Bray, Co. Wicklow, Dr. D. Moore; Slieve Glali, Co. Cavan, D. MArdle. Found on the Continent, and doubtfully in North America. Obs. — Only to be mistaken for Nardia {Eucahjx) liyalina (Lyell), which see. In addition to other distinguishing characters mentioned there, the rootlets are usually fewer and darker coloured, the bracts adnate higher up the perianth. The specimens published in C. & P. Hep. Brit. Ex. n. 100, as Nardia liyalina belong to this species. Description of Plate CLIX. — Fig. I. Plants natural size. 2. Portion of stem, antical view X 24 (Yosges, Mougeot). 3, Por- tion of fertile stem with perianth x 16 (C. & P. Hep. Brit. Ex. n. 100). 4. Portion of stem, antical view, x 16 (Killarney, Wilson). 5. Portion of stem, postical view, x 16 (G. & E. Hep. Eur. n. 266). 6. Leaf x 24 (Vosges, Mougeot). 7. Leaf x 24 (C. & P. Hep. Brit. Ex. n. 100). 8. Leaf x 24 (Vosges, Mougeot). 9. Portion of leaf x 290 (Vosges, Mougeot). 10, 11. Bracts x 16 (C. & P. n. 100). 12. Perigonial bract X 24 (Vosges, Mougeot). 13. Antheridium x 85 (ditto). Subgenus 2. EUNARBIA, Lindb. Eimardia, sect. 3IesophiyUa (Dum.), Lindb. Muse. Scand. p. 8 (1879). Eunardia, Spruce, Hep. Am. et And. p. 519 (1885). Mesojyhylla (Dum.), Lindb., Kaalaas Lev. Norg. p. 388 (1893). Stipules present, small, subulate. Perianth immersed, deli- cate, obscurely carinate, adnate to the lower half of the bracts. 3. Nardia compressa [Hook.), Gr. §• Benn, Jungermania compressa, Hook. Brit. Jung. t. 58 (181G). Nardia comjyressa, Gr. & B. Nat. Arr. Brit. pi. p. G91 (1821). Mesophylla comp'essa, Dum. Comm. Bot, p. 112 (1822). Alicidaria compressa, G.L.N. Syn. Hep. p. 12 (1844). Dioicous, csespitose, flagelliferous ; flagella leafless, slightly j-adiculose ; from medium to large in size, pale green colour NARDIA. 369 with a purple or claret tinge. Stems simple or branched, innovant, suberect, laterally compressed, cortical cells large, hyaline, 35 to 50, inner very small, densely crowded, 20 to 25 in diam. ; radiculose, rootlets few, white. Leaves closely imbri- cate, erect, horizontally inserted, decurrent, appressed to stem laterally, plane or slightly concave, reniform or subrotund, entire ; texture pellucid, thin, cells medium size, roundish- quadrate, walls thin, firm, reddish, lumen with few oil corpuscles, trigones distinct. Stipules small, triangular, irregular, dentate, rarely subulate. Bracts similar to the leaves, only larger and more distinctly reniform. Bracteole ovate, irregular. Perianth immersed in the bracts, delicate, conoid, lower portion adnate to the innermost bracts, mouth twisted, dentate, teeth small, few. Capsule oval. Sj)ores brown. Elaters bispiral, brown. Andrcecia terminal, 2-3 pairs of bracts, but little different from the stem leaves, slightly ventricose at the base ; antheridia large, oval, 2-3 in each bract. Fruits April, May. Dimensions. — Stems J to 2 inches long, "3 mm. diam., with leaves 2' mm. wide, leaves r4 mm. x TS mm., 1'5 mm. x 2*0, 1-25 mm. x 175 mm. ; cells '05 mm. x 03 mm., "035 mm. x -03, •03 X '04 mm., 03 mm. x '03 mm. ; stipules "6 mm. x -3 mm., •55 X "2 ; perigonial bracts 1'4 mm. x I'S mm., 1^3xl'75 mm.; antheridia '225 mm. x "2 mm. Hab. — On wet rocks and stones by or in alpine or subalpine rivulets. Somewhat rare. 1. Dartmoor, Devonshire, E. M. Holmes. 7. Glyder Vawr. Carnarvonshire, JV. Wilmu, G. A. Holt, W. H. P. Snowdon, Carnarvonshire, Dr. Carnngton ; Pass of Llanberis, Carnarvon- shire, W. H. P. ; Cader Idris, Merionethshire, W. H. P. 9. Staley Brushes, Cheshire, G. A. Holt; Carr Meadows, Derbyshire, G. A. Holt; Kinder Scout, Derbyshire, Whitehead & Holt. 10. Near Todmorden, /. Noicell, G. A. Holt; Ingleboro, Br. Car- rington ; Blaeberry Gill, M. B. Slater ; Falling Foss, Little Beck, North Yorks., M. B. Slater ; Cleveland, W. Mudd. 12. Little Langdale ; Oxendale, Great Langdale, Westmorland, G. Stabler; 370 HEPATICJ^. Borrowdale, Cumberland, W. 11. P. 13. New Galloway, J. McAn- clrew. 15. Succoth Hill, Arroquhar, A. McKinlay ; Ben Sligott, C. Howie; Ben Mac Dliui, G. E. Hunt, J. Wliifehead ; Glen Clunie, Aberdeen, J. Whifehead ; Ben Laoigb, T. Fwfjers. 16. Moidart, West Inverness, 600 feet, 8. M. Macvicar. I. Bantry, Miss Hut chins ; Aooreagb River near Sneem, Br. Taylor ; abundant at Upper Lougb Bray, Br. B. Moore ; Luggie- law and Seven Cburcbes, Dr. D. Moore ; Connernara, near Kyle- more, Gal way, Br. B. Moore; Kelly's Glen, Dublin; Killarney, Dr. Carrington; Co. Wicklow, Prof. S. 0. Lindherg. Found on the Continent, also in Greenland. Obs. — This species, being semi-aquatic, sometimes grows to the length of 4 or 5 inches, and forms in some subalpine rivulets large patches over which the water is constantly flowing. In drier alpine situations a small variety is found, of a deep purple colour, steins J to |- inch long, simple, leaves orbicular, closely imbricate, stipules dentate. From Nardia scalaris [ScJirnd.) it is distinguished by its reniform, closely laterally appressed leaves, which are also thinner and more delicate, and its large cortical stem cells. From JamesonieUa Carrincjtoni {Balf.\ S^jriice^ see notes under that species. Description of Plate CLX. — Fig. 1. Plants natural size. 2. Portion of stem x 16 (Ireland, S.O.L.). 3-5. Leaves x 16 (ditto). 6. Portion of leaf x 290 (ditto). 7, 8. Stipules x 24. (ditto). 9. Stipule X 24 (Cader Idris, W. H. P.). 10. 11. Longitudinal sections of bracts and perianth (Dr. Gottsche, G. R. Hep. Eur. 443). 12, 13. Perigonial bracts x 16 (Ireland, S.O.L.) 14. Antheridium x 85 (ditto). 4. Nardia scalaris [ScJtrad.), Gr. §• B. Jungermania scalaris, Schrad. Syst. Samml. Krypt. Gew. 2, p. 4 (1797). Nardia scalaris, Gr. (Si B. ISTat. Arr. Brit. pi. p. 694 (1821). Mesophylla scalaris, Dum. Comm. Bot. p. 112 (1822). Alicularia scalaris, Corcla in Opiz Nat. p. 653 (1829). Dioicous, loosely or densely ca}spitose, eflagelliferous, small^ NAIWTA. 871 pale, or dark to brownish green, rarely reddish in colour. Stems creeping or suberect, simple or furcate, slightly laterally com- pressed, cortical cells smaller and darker than the inner, which are large and distinct, about 9x9 cells in diam. ; radiculose, rootlets plentiful, dull white. Leaves distichous, imbricate, suc- cubous, erecto-patent or patent, obliquely inserted, appressed or spreading, orbicular or rotund, slightly concave, entire or slightly retuse, lower smaller and distant ; texture somewhat thick ; cells smallish, roundish quadrate, walls moderately firm, trigones large, distinct ; lumen containing several oil corpuscles. Stipules large and distinct, subulate or triangular-subulate, entire. Involucre obovate, laterally compressed, composed of two bracts and brac- teole ; bracts much larger than the stem leaves, orbicular, entire or emarginate, bracteole triangular-subulate, connate at the lower half of the bracts. Perianth immersed in the bracts, the lower half adnate with the innermost bracts, composed of more delicate cells, conoid, mouth contorted. Capsule oval-spherical. Spores brown, elaters bispiral, brown. Androecia terminal or on the middle of stem ; perigonial bracts closely imbricate, smaller than the leaves, ventricose, enclosing 2 roundish oval, small antheridia. Fruits March, April. Dimensions. — Stems |- to 1 inch long, -2 mm. to -25 mm. in diam., with leaves V to 1*5 mm. wide; leaves '6 mm. x '8 mm., •7 mm. X -9 mm., 7 mm. x 7 mm., '6 x 75 mm.; cells '03 mm., -025 mm., '035 mm. "02 mm.; stipules '225 mm. x -125 mm. at the base, '25 mm. x '125 mm., -35 mm. x -075 mm. ; bracts 1-25 mm. x I'l mm. ; bracteole 1- mm. x -6 mm.; pistil- lidia -15 mm. x '075 mm. ; perigonial bracts -G mm. x -G mm., •G mm. X 7 mm., '5 mm. x -G mm.; antheridia 'll mm. x 'l mm., '11 mm. x -08 mm. Hab. — Grows in loose or dense patches on damp banks, moors, rocks, or stones; generally distributed 1 to 18. I. Common on the Continent. Found in North America. Obs. — A common species, easily recognised, even when barren, by the presence in the leaf-cells of several oil corpuscles, from any 372 HEPATIC a:. oi" the other round-leaved JangermanicB, excspt Nardla comprcssa, which has them also. This is a larger species ; stem with large cortical cells, leaves broader and more closely compressed. Nardia hyalina and N. obovata have reddish or purplish root lets, which at once distinguishes them. Description of Plate CLXI. — Fig. 1. Plants natural size. 2. Sterile stem x 24 (Cader Idris, W. H. P.). 3. Portion of fertile stem x 24 (Dalarne, S. 0. L.), 4, 5. Leaves with stipules X 24 (Cader Idris, W. H. P.). 6-10. Leaves x 24 (ditto). 11. Portion of leaf x 290 (ditto). 12, 13. Stipules x 85 (ditto). 14. Stipule X 85 (Sweden, S. 0. L.). 15, 16. Bracts x 24 \Sweden). 17. Bracteole x 24 (ditto). 18. Pistillidium x 85 (ditto). 19, 20. Perigonial bracts x 24 (Sweden, S. 0. L.). 21. Antheridia x 85 (ditto). 5. Nardia silvrettse (Goffsc/te). Syn. Jungermania Silvrettce, Gottsche in G. R. Hep. Eur. Ex. n. 470 (1869). Sarcoscyphus anomalus, Jack in lit. Dr. Gottsche, G. R. Hep. Eur. Ex. n. 470 (1869). Alarsupella (?) silvrettce, Dum. Hep. Eur. p. 128 (1874). Alicularia minor, Limpr. p.p. in Cohn Krypt. Fl. Schles. p. 251-433 (1876). Nardia hanrnatosticta, var. suberecta, Lindb. Muse. Scand. p. 8, n. 153 (1879). Nardia geoscypha, var. suherecta, Mass. Epat. delle Alp. Penn. in Nuov. Giorn. Bot. Ital. vol. xiv. tab. xiii. Exsicc. Theden. Muse. Suec. Ex. n. 136 (as Jung, scalaris); G. R. Hep. Eur. dec. 42 et 44, n. 416 (as Alicidaria scalaris, var. minor); dec. 45 et 47, n. 470 (as Jung. Silvrettce) ; Husn. Hep. Gall. n. 101 (as Alicidaria Paroicous, caespitose, small, light to dark or brownish green in colour. Stems simple or furcate, with often single slender inno- vations arising from below the involucre, suberect, fleshy, some- what fragile, on a transverse section oval, frontally compressed, cortical la3^er of cells smaller, darker, about 30, inner large, hyaline, 12 cells x 9 or 10 ; densely radiculose, rootlets white or purplish. Leaves alternate, subdistichous, obliquely inserted, patent, horizontal or suberect, lower smaller, distant, upper larger NARDIA. 37a accrescent, concave, orbiculate, broadly ovate or ovate, emarginate, rarely entire, sinus shallow, /, to J deep, obtuse or rarely acute, segments obtuse, rarely acute ; texture somewhat thick, cells moderately large, roundish subquadrate, lumen with oil corpuscles as in N. scalaris, trigones distinct. Stipules subulate, entire, or with one, two, or three irregular segments on one or both sides. Involucre oval or clavate, composed of two large bracts with bracteole (irregularly cuneate) behind, which is overlapped by the bracts, connate with them at the base ; bracts 2 to 3 pairs, larger than the leaves, subrotund, emarginate, margin irregular, crisped, ventricose at the base. Perianth immersed, composed of elongate, hyaline cells, mouth irregular, contorted, lower half adnate to the innermost bracts. Calyptra delicate, cells irregular in shape, walls very thin ; at its base are 8-10 barren pistillidia. Capsule brown. Spores smooth, light brown. Elaters darker brown than the spores, bi-trispiral. Antheridia roundish-oval, stipitate, usually two in each bract below the perianth. Fruits March, April. Dimensions.— Stems J to f inch long, diam. '4 x '3 mm., with leaves 2" mm. wide; leaves 2' mm. x I'l mm., segments '8 mm., I'l mm. X -9 mm.,seg. "2 mm., '9 mm x '9 mm., seg. "2 mm., '9 mm. X 7 mm., seg. "2 mm., "8 mm. x '8 mm., seg. "2 mm., '8 mm. X 7 mm., seg. '2 mm., '5 mm. x -4 mm., seg. '15 mm. ; cells •04 mm.; stipules "4 mm. x -1 mm. at base, '35 mm. x •125 mm., "275 mm. x •! mm.; bracts l^l mm. x 1* mm., segments '2 mm., •3 mm.; sub-bracteoles •G mm. x '25 mm. at the base; bracteole •6 mm. X •SS mm. at the base, '4 mm. x -15 mm. ; involucre 2-5 mm. x 1' mm.; cells of perianth •OG mm. x ^02 mm. ; pistilHdia '175 mm. x ^05 mm.; valves of capsule 8 mm. x •G mm.; spores '02 ram. diam.; elaters "15 mm. x •Ol mm. Hab. — Grows on banks and moors. Kare. 9. Gorpley Clough, Todmorden, Lane, G. A. Holt. Alderley Edge, Cheshire, G. A. Holt. 10. Strensall Common, Yorks., G. Stabler. Baildon Moor, Yorks., IF. West. 16. Moidart, West Inverness, S. M. Macvicar. Found on the Continent. 374 HEPATIC^. Obs. — The history of this hepatic is somewhat curious. Nees, in his admirable "Nat. Eur. Leb.," vol. 11, p. 453, speaks of a form of Jimgermania scalaris (var. minor) which ma}^ prove to be a distinct species and to which he would give the name Jim^. hamatosticta. To this Professor Lindberg referred the Jlicidaria geoscyphu8 of de Notaris. Herr Limpricht objects to the name as being inappropriate, for the plant varies very much in colour, according to exposure, shade, &c., and he promotes Nees' varietal name minor to specific rank. To a large, suberect form of this. Professor Lindberg has given the name var. subereda, which he had previously supposed to be the Jmiff. scalaris var. repanda, Hliben., and as such published it as Nardia repanda, but an examination of Hlibener's original speci- mens showing it to be dioicous, the species repanda was dropped. In 1869 Herr Jack forwarded to Dr. Gottsche specimens of this var. subereda under the name Sarcoscijphus aiioiualus, which the Doctor published in G. R. Hep. Eur n. 470 as Jnug. Silvrettce ; as the specific name of Herr Jack is used for another hepatic, I adopt Gottsche's name, which should have priority over the others, if we raise it to the rank of a species. A peculiarity about this species is that the stem is slightly frontally compressed, not laterally as in Nardia com^wessa and partially in N. scalaris. Transverse sections cut from all my specimens give 12 cells x 9 or 10. Distinguished from Nardia scalaris (Schrad.) by its paroicous inflorescence, and the emarginate leaves and bracts. Nardia geoscyplia (de Not.), of which I have not seen British specimens, agrees with this species in its inflorescence, but is much smaller, has a curious creeping habit, is without stipules, and the bracts are not so irregularly margined. Description of Plate CLXII. — Fig. 1. Plants natural size 2, 3. Stems x 16 (Thed. Muse. Suec. Ex. n. 136). 4. Cross- section of stem X 31 (Todmorden, G. A. Holt). 5, 7. Leaves X 24 (Baildon, W. West). 6, 8-10. Ditto x 24 (Todmorden, Holt). 11. Portion of leaf x 290 (ditto). 12, 13, 15, 16. Stipules X 85 (ditto). 14. Ditto x 85 (Baildon, West). 17. MARSUPELLA. 375 Bract X 24 (Todmordeii, Holt). 18 Ditto x 31 (Strensall, G. Stabler). 19. Sub-bracteole x 31 (ditto). 20. Portion of iiiner- luost bract and connate bracteole x 16 (Thed., 136). 21. Brac- teole X 31 (ditto). 22. Pistillidia x 85 (Baildon, West). 23. Antheridium (ditto). 24, 25. Spores and elater x 2'JO (Tod- morden, Holt). Genus 33. Marsupella, Di/iu. Jungermcmia, Ehrh. Beitr. ;5 p. 80 (1788). Nardius, Gr. ct B. Nat. Arr. Brit. PI. 2 p. (;;)4 (1821). Jliii'supella, Dum. Comm. Bot. p. 114 (1822). Spruce " Rev. Bryol." p. 92 (1881). Sarcosci/phas, Cord, in Opiz Beitr. 1 p. (i52 (1829). Nardla, Carr. in Trans. Bot. Soc. Edin. x. p. 309 (1870) ; Brit. Hep. p. 10 (1874). Nardia, sect. c. Marsupella, Lindb. Hep. in Hib. p. 531 (1874). Stem (witli leaves) frontally compressed, usually with rhizomes and flagella proceeding from the base subleafless. Leaves trans- verse, imbricating or spreading, canaliculate-concave, sub- complicate or carinate, always bilobed. Cauline stipules always wanting. Fructification terminal. Bracts few or several pairs, innermost (1-2 pairs) connate ; also at the base with the peri- anth, bracteole normally wanting. Perianth frontally compressed, mouth 5-7 lobed. Calyptra included. Capsule 4-valved. Elaters bispiral. Androecia subspicate ; antheridia in the saccate base of the perigonial bracts. 1. Marsupella emarginata [Ehrlt.), Dion. Junrjermania emao'yinata, Ehrh. Beitr., 3, p. 80 (1787-92). Jungermania macrorhiza, Dicks. Crypt. Brit. f. 2, p. 16, t. 5, f. 10 (1785). Marsuj)ella emarginata, Dum. Comm. Bot. p. 114 (1822). SarcoscT/phus Ehrharti, Corda in Opiz Nat. p. (532 (1828). Dioicous, densely caespitose, flagelliferous, small to large, varying very much in colour, from pale or dark green to brown, reddish-brown, bright red or purple to almost black. Stems slightly or intricately branched, suberect or erect, firm, cortical 376 BEPATICM cells larger and darker than the inner ; radiculose, rootlets long, white or purplish, thick. Leaves spreading, suberect or erect, closely or loosely imbricate, cordate, subrotund or ovate, emarginate to about ^th, sinus acute or obtusate, segments broad, acute or obtuse ; texture firm or lax ; cells smallish, roundish, walls firm, angles thickened, no trigones, marginal cells smaller, quadrate. No stipules. Bracts larger than the leaves, subrotund, emarginate to about ^th, segments and sinus acute or obtuse. Perianth usually smaller than the innermost bracts, adnate to them for about half its height, mouth 4, 5-lobate. Capsule spherical, pedicel short, spores brown, elaters bispiral. Androecia terminal, 2-4 pairs of bracts, which are closely imbricate, ventricose at the base, somewhat similar to stem leaves, enclosing 3, 4, large, oval antheridia, with long stipes (10-24 cells long). Fruits March, April, May. Dimensions. — Stems |- to 3 inches long, diam. -1 mm. to •4 mm., with leaves 1' mm. to I'S mm. wide; leaves 75 mm. x 75 mm., segments 2 mm., 75 mm. x -6 mm., seg. "2 mm., 9 mm. X "8 mm., seg. '2 mm., 1* mm. x l" mm., seg. '2 mm., 175 X 2* mm., seg. '2 mm. ; cells -03 mm., '025 mm., '025 mm. x "02 mm.; bracts 1"5 mm. x 1*5 mm., seg. "3 mm.; perianth 1-5 mm. long; antheridia '225 mm. x '15 mm., -2 mm. x -125 mm.; stipe '15 mm. long. Hab. — Grows on rocks or at the base of stone walls, from the plains to alpine regions, most frequently in or by the side of subalpine rivulets. 1, 3, 7, 10 to 17. I. Found on the Continent and in North America. Var. robusta Glyder Vawr, IF. H. P. Var. saccata ( ?) Killin, Perthshire, G. A. Holt. Obs. — A very variable species in size, colour, and shape of leaves ; several varieties have been named by Nees and other authorities, and some have been esteemed species by Lindberg and others. Two of the most remarkable are var. robusta (Marsupella robust a, Lindb.) and one which I believe is var. saccata of Nees ; the former is distinguished by its large size, firm texture MARSUPELLA. 377 of stem and leaves, subrotund leaves, with shallow sinus and broad rotimdate' segments. Var. mccala N. (?) is a very graceful form, leaves orbicular, sinus shallow, segments broad, rotundate. From the dioicous M. sphacckda, Faiickii, and Ccsia alpuia, see notes under those species for distinguishing characters. Dkscription of Plate CLXIII. — Fig. 1. Plants natural size. 2. Portion of fertile stem x 24 (Mael Tarmachan, C. J. Wild). 3. Leaf x 24 (Langdales, G. Stabler & W. H. P.). 4-6. Leaves x 24 (Snowdon, Nuttall). 7. Leaf x 24 (Festiniog, W. H. P.). 8-10. Leaves x 24 (Arthog, J. Whitehead). 11. Portion of leaf x 2U0 (Mael Tarmachan, C. J. Wild). 12, 13. Bracts x IG (ditto). 14, 15. Linermost bracts with portions of perianth adhering. IG. Antheridium x 85 (Arthog, J. White- head). Description of Plate CXLIV. — Fig. 1, 2. Marsupella emar- (jinata var. robust a. Leaves x 24 (Glyder Vavvr, W. H. P.). 3-5. Ditto X 24 (Finland, S. 0. Lindberg). G. Perigonial bracts with antheridia x 24 (Glyder Vawr, W. J I. P.). Fig. 7. Marsupella emarginata var. saccafa Nees ? . Portion of stem X 16 (Killin, G. A. Holt). 8-10. Leaves x 24 (ditto). 2. Marsupella sphacelata [Gieseke), Bum. Junrjermania sphacelata, Gieseke in Lindeub. Syn. Hep. p. p. 7(> t. i. f. D-12 (1839-41). Marsivpella sphacelata, Dum. Recueil Jung. p. 24 (1835). Sarcoscijphus sphacelatics, Nees Eur. Leberm. i. p. 129 (1833). Xardia sphacelata (Gieseke), Carr. Trans. Bot. Edin. 5, 10, p. 878 (1870). Dioicous, loosely crcspitose, from medium to large in size, pale or dark green, with apices tinged brown, or sometimes black in colour. Stems simple or slightly ramose, flagelliferous, leafless below, erect or procumbent, slender, flexuose, rootlets few. Leaves usually distant, erect-patent, decurrent, amplexicaul at the base, concave, cordate, bilobed to about I, sinus obtusate or acute, 378 HEPATIC^. segments rotundate ; texture lax, delicate, cells smallish, 4-5 and 6-sided ; walls reddish, firm, trigones more or less distinct. No stipules. Bracts erect, somewhat similar to the leaves, only larger and longer. Perianth delicate, small, about f the height of the bracts with which it is adnate, mouth laciniate-dentate. Male stems more slender, antheridia terminal or in middle of the stem; perigonial bracts imbricate, turgid; antheridia 1, '1, or 3 at the base of each bract, oval. Dimensions. — Stems from 1 to 3 inches long, diam. 'IS mm. to '2 mm., with leaves Vb mm. to 2' mm. wide : leaves 1'75 mm. long X 1-25 mm. broad, segments -5 mm., I'S mm. x 1* mm., seg. '3 mm., '9 mm. x 7 mm., seg. '2 mm.; cells "02 mm. x •02nmi.,-02 mm. x '03 mm.; bracts 2' mm. long x 75 mm. broad ; segments '3 mm.; perigonial bracts r25 mm. long x 1* mm. broad, segments '2 mm. ; antheridia -4 mm. x -3 mm., -3 x -25 mm. Hab. — Growing in extensive patches in wet situations, by the margin of pools or streams in alpine situations. 7. Glyder Vawr, G. A. Holt ; Margin of Llyn Bodlyn, Cwm Idwal, W. H. P. 15. Ben MacDhui, G. E. Hunt, JoJm Whitehead; Loch Kandor, G. E. Hunt. 16. Moidart, West Inverness, S. M. Macvicar, 1899. 18c. Uist, ./. Sivi. I. Loch Bray, Lindber^ ; D. McArdle. Found on the Continent and in North America. Obs. — From all forms H). Jangermania Fanckii, Web. et Mohr. Deuts. krypt. p. 422 (1807.) Sarcoscij2)hus Ficnckii, Nees Eur. Leberm. t. i., p. 130 (18;5;>). Marsupella Funckii, Dum. Recueil Jung. p. 24 (1835). Dioicous, densely csespitose, stoloniferous, stolons leafless, nidiculose ; small, lurid or dark brown colour. Stems erect or procumbent, innovantly fasciculately ramose ; radiculose, rootlets few, wanting- on the ascending branches. Leaves imbricate, on young plants or innovations distant, patent to erecto-patent, concave, subrotund, obovate or oval, emarginate from J to J, sinus and segments acute ; texture firm, cells rather minute to small, 4, 5 and 6-sided, walls thick, angles thickened. No stipules. Bracts larger than the leaves, broadly ovate, emarginate from ^ to ^, sinus acute, segments acute or obtusate. Perianth smaller than the innermost bracts to which it is adnate for about half its length, free margin 4, 5 dentate ; pistillidia 5-7. Capsule small, brown ; spores pale brown ; elaters a little darker, bispiral. Androecia terminal on chief stem or branches, 4, 5 pairs of bracts, closely imbricate, tumid ; antheridia solitary, oval, stipitate. Fruits April, May. Dimensions. — Stems |- to f inch long, diam. "1 mm., with leaves '7b mm. wide; leaves '6 mm. x '5 mm., segments '2 mm., •5 mm. X *5 mm., seg. '2 mm., "6 mm. x "6 mm., seg. "2 mm. •5 mm. X "4 mm., seg. '15 mm.; cells "015 mm. x "015 mm. .0225mm. x -0175 mm., '02 mm. x '02 mm.; sub-bract '8 mm. X -8 mm., seg. '2 mm. ; bracts -9 mm. x "8 mm., seg. "25 mm.; perianth '7b mm. high; perigonial bracts "5 mm. x '5 mm., seg. "2 mm. ; antheridia '15 mm. x •12 mm. ; spores "01 mm. ; elaters •15 mm. X '015 mm. Hab. — Grrows usually on subalpine and alpine pathways, or on rocks on which there is a layer of soil, rarely on the bare rock. 2. Tilgate Forest, G. E. Lavics. 7. Bettws-y-Coed, Dr. Carrinyton. Cader Idris, W. 11. P. ; Tyn-y-Groes, G. A. Holt, 380 HEPATIC.]^. W. H. P. 12. Bow Fell, G. Stabler. Borrowdale, IF. H. P. 13. Miljea ; North of Black Craig; Ballingear Woods, New Galloway, /. McAjidreiv. 15. Jjoch-na-Gar, ./. 8i7n ; Ben Challum, Breadalbane, A. 0. Black; Ben Voirlich, Craig-na-Gour, and Ben Lawers, J. McKinlay ; Locli Kandor, Ben MacDhui, G. E. Hunt. 16. Moidart, West Inverness, S. M. Macvicar. I. Black Mountains, Dr. Moore. Found on the Continent. Obs. — Distinguished from M. emarginata (Ehrh.) by its usually darker colour, by its never cordate leaves, with segments more acute. Cesia alpina (Gotts.) has polished leaves of a different texture. Description of Plate CLXVI. — Fig. 1. Plants natural size. 2. Fertile stem x 31 (Llanberis, W. H. P.). 3. Male stem x 31 (ditto). 4-8. Leaves x 24 (Tyn-y-Groes, W. H. P.). 9. Leaf X 31 (Llanberis, W. H. P.). 10. Portion of leaf x 290 (ditto). 11. Sub-bract x 31 (ditto). 12, 13. Bracts with portions of perianth x 31 (ditto). 14-17. Perigonial bracts x 31 (ditto). 18. Antheridium x 85 (ditto). 19. Spores x 290 (ditto). 4. Marsupella olivacea, Spruce. Marsupella olivacea, Spruce in " Rev. Bryol." p. 97 (1881), Synoicous and paroicous, csespitose, small, olive green colour. Stems rigid, rhizomatous, stolons long, creeping, leafless or clothed with minute leaves, radiculose, erect, subsimple, or innovantly elongate — rarely repeatedly ramose, apices of the fertile stems clavate, sterile stems filiform- small-leaved. Leaves small, subimbricate, erect, appressed — upper sometimes subpatent at the apex — at the base decurrent-vaginate, oblong, canaliculate- concave, hardly subcarinate, bilobed from \ to J, sinus obtuse, lobes obtuse, rarely abruptly subacute ; texture thick, opaque ; cells minute, walls thickened, angles thickened, no trigones, cells of the upper portion of the leaf and middle subequal, near the base larger, a little elongate, marginal cells quadrate. Bracts terminal, fusiform, 3-4 pairs, broadly ovate, canaliculate-concave, at the apex very shortly (^V to \) bilobed, lobes incurved, MAllSUPELLA. 381 obtuse, rarely rotundate ; inner bracts twice as large or larger than the leaves, slightly connate at the base, apex sometimes divided to \. In the axils of all the bracts are large, stipitate solitary (rarely two) antheridia, sometimes in the same receptacle with the pistillidia.. Pistillidia about 5, rarely about 10. Peri- anth very variable, sometimes as long as the bracts, oblong, apex rarely closed and symmetrical, very often cucullate, sometimes shorter and open at the side, mouth, then plurilaciniate, concrete from the base to about |- high with the bracts, sometimes only at the base, rarely free. Calyptra smaller, oval-globose, 2 cells thick, at the base and for a little way up adnate with the perianth, to above the middle strewn over with 4-9 sterile pistillidia. Capsule oblong-globose, composed of two layers, inner layer composed of semiannular fibres. Spores brown. Elaters reddish-brown. Fruits March, April, May. Dimensions. — Stems |- to J inch long, '1 mm. diam., with leaves '4 to "5 mm. wide; leaves '5 mm. long x '3 mm. broad, seg. •1 mm., "4 X "3 mm., seg. '075 mm., '4 x '25 mm., seg. "075 mm., '3 X "2 mm., seg. '05 mm., 2 mm. x 15 mm., "Smm. x -3 mm. ; cells, near the middle "01 mm., •0125 mm , "01 mm. x '0125 mm., near the base "0125 mm., -01 mm. x "02 mm., 01 x -0125 mm., -015 mm. X "015 mm. J perigonial bracts "5 mm. x -45 mm.; seg. '1 mm., •525 mm. x ^4 mm., seg. "1 mm. ; bracts '65 mm. x '6 mm. ; anthe- ridia •IS mm. X "1 mm., bearer ^5 mm. long ; pistillidia ^075 mm. x ^035 mm. ; capsule ^4 mm. x "325 mm. ; pedicel "1 mm. diam. ; spores "0175 mm. ; elaters ^075 mm. long x "01 mm. broad. Hab. — Growing on rocks in alpine localities. Very rare. 7. Clogwyn-du-Arddu, Snowdon, Carnarvonshire, IF. H. P. 12. Little Langdale ; Bow Fell, Westmorland, G. Stabler. " This species grows along with JSLarsupcUa Sfahlcri on the large stones and rocks immediately to the left of the track from the top of Rosset Ghyll to Angle Tarn," G. Stabler. IG. Moidart, West Inverness, S. M. Macvicar. Found on the Continent, Riesengebirge, Germany, G. L'lm- prlcht. 382 HEFATIGM. Obs. — This rare species, discovered on Bow Fell by Mr. George Stabler, is distinguished from an}^ of the Ceslm by the presence of a true perianth, from Marsupella usfidafa {Hilben.) by its closer appressed leaves, which are more oval, with shallower sinus and more obtuse segments. For further interesting notes see Dr. Spruce's excellent paper m Revue Bryologique for 1881. Description or Plate CLXVII. — Fig. 1. Plants natural size. 2. Portion of fertile stem x 64 (Bow Fell, G. Stabler). 3, 4. Leaves x 85 (Snowdon, W. H. P). 5, 6. Ditto x 64 (Bow Fell, a Stabler). 7. Upper leaf x S5 (ditto). 8. Portion of leaf near middle x 290 (ditto). 9. Ditto, near the base (ditto). 10. Sub-bract x 85 (ditto). 11, 12. Perigonial bracts X 85 (ditto). 13. Bract and portion of perianth x 85 (ditto). 14. Portion of perianth x 85 (Snowdon, W. H. P). 5. Marsupella ustulata [hilben.)^ Spruce. Jtmgermania ustulata, Hubener. Hep. Germ. p. 132 (1834). Ilarsupella ustulata, Spruce in " Rev. Bryol." p. 100 (1881). Marsupella ustulata (Hiiben.), Spruce in litt. Paroicous, shallowly csespitose, minute, reddish or purplish brown, in shady places green, in exposed alpine situations purplish black in colour. Ehizomes creeping ; radiculose, root- lets frequent, intricately flexuose, stoloniferous. Stems suberect or erect, simple or slightly branched ; fertile clavate, leaves accrescent, subimbricate or imbricate, sterile small, equal, distant or approximate, patent or erecto-patent, broadly oval, ovate or sub- rotund, sometimes subovate, laxly complicate, carinate, from J to ^ bilobed, sinus acute, rarely obtuse, lobes acute, rarel}^ obtuse ; tex- ture firm, cells very small to small, roundish-quadrate, walls firm, angles slightly thickened, sometimes slightly trigonous, basal cells a little larger and elongate. Inflorescence terminal. Bracts $ 2-5 pairs, almost twice as large as the lower or sterile stem leaves, broadly ovate, orbiculate, perigonial bracts subcordate, ventricose and antheriferous at the base, -| to \ bilobed, lobes acute or obtuse, innermost longer and erect, margin reflexed or undulate, more or le^s connate ?it the base, lobes often obtuse, rarely acute. Anthe- MAKSUPELLA. 383 ridiii 1 or 2 in the axil of the bracts, oval. Perianth small, immersed, delicate, composed of larger cells than the leaves, oval-globose or tubular, 4, 5 laciniate, sometimes composed of 2 bilobed leaves rarely with a further segment, adnate from the base to nearly the middle with the innermost bracts, very rarely free to the base. Calyptra oval-globose, surrounded about the base by 6-8 sterile pistillidia. Capsule globose, composed of two layers, inner with semi-annular fibres. Spores small, brown. Elaters as wide as the spores, same colour, bispiral, rarely trispiral. Dimensions. — Stems 2* to 5" mm. long, 075 to '15 mm. diam., with leaves (fertile) "05 mm. wide (sterile) '35 mm ; leaves "3 mm. X "25 mm., seg. '11 mm., •Gmm. x '5 mm., seg. '15 mm., '5 mm. X '4 mn., seg. '1 mm., "25 mm. x '25 mm., '35 mm. x '3 mm., •4 mm. X '33 mm., '55 mm. x '45 mm. ; cells '02 mm. x 0125 mm., '015 mm. X '015 mm., '0175 mm. x '0175 mm., '02 mm. x '015 mm. ; sub-bract '4 mm. x 4*75 mm.; seg. '1 mm; bracts 1' mm. x 1' mm. ; seg. '225 mm., '5 mm. x '5 mm., 'G5 mm. x '5 mm., '85 mm. X I'l mm., I'l mm. x I'l mm.; perianth '7b mm. high; pedicel 2' mm. long; capsule '3 mm.; spores '01 mm. diam.; elaters '1 mm. long x '01 mm. broad. Hab. — Grows in very thin shallow patches on soft sandstone rocks in moist situations, in the plains or in subalpine or alpine localities. Eare. 2. Blackdown, Sussex, Jciiuer, Georfje E. JJacies;. 7. Fairy Glen, Bettwys-y-Coed, Dr. Carriiu/fon ; Snowdon, Carnarvonshire, IF. TL P. ; Tyn-y-Groes, Dr. Carringlon c^ W. II. P. ; Cader Idris, Merionethshire, Wild ci: Holt. 10. Park Quarry, Castle Howard, Yorks., Br. Sjjrnce, M. B. S/a/er, G. Stabler. 12. Harter Pell; Bow Fell; Oxenden, Westmorland, G. Stabler. IG. Moidart, West Inverness, S. M. Macvicar cO JF. II. P. Found on the Continent and in North America. Obs. — Mars/ij)('I/a .sparsifo/ia, Lindb., which is also paroicous, is a much larger and robust species, of a darker colour, with lobes of the leaves obtuse. Marsupclhi olivacea, Spruce, has more oval lcave.«, with shallower sinuses, 384 HEPATIC^. Description of Plate CLXVIII. — Fig. 1. Plants natural size. 2. Fertile stem x 24 (Hercynia, Hampe). 3. Ditto x 64 (Castle Howard, Dr. Spruce). 4. Portion of sterile stem x 64 (ditto). 5. Leaf x 64 (Bettvvs-y-Coed, Dr. Carrington). 6. Ditto X 64 (Hercynia, Hampe). 7. Ditto x 85 (Bettws-y-Coed, Dr. Carrington). 8. Portion of leaf x 290 (ditto). 9. Sub-bract perigonial, x 64 (ditto). 10, 11. Bracts with portion of peri- anth X 24 (Castle Howard, Dr. Spruce). 12. Antheridium x 85 (Hercynia, Hampe). 6. Marsupella sparsifolia {LincJb.), Bum. Sarcoscyphus sparsifolia, Lindb. in Not. pro F. Fl. Fenn. Forh. 1, p. 280, No. 13 (1808). Nardia {Marsuj^ella) sjyarsifolia, Lindb. in Not. pro F. Fl. Fenn. Forh. xiii., p. 270 (1874). Marsupdla sparsifolia, Dum. Hep. Eur. p. 123 (1874); Pears, in " Jovirn. of Bot." p. 225 (1884). Paroicous, csespitose, stoloniferous, stolons without or with a few leaves ; small, sphacelate, olive or blackish-brown colour. Stems creeping, intricately entangled, shoots simple or rarely irregularly furcate, annotinous, often one, two or three inno- vations on a single stem, erect; rootlets short, purple, mostly on the postical side of the creeping stems and stolons, rarer on erect shoots. Leaves 12-20 pairs, distichous or subsecund, alternate, clasping the stem at the decurrent base, ascending, lower leaves approximate or somewhat distant, vaginate, the few leaves nearest apex more closely imbricating, roundish-ovate, cordate-orbiculate, obcordate, subquadrate, some leaves narrow at the base, emarginate, sinus :^ to J deep, obtusely angular, wide, rarely acute, segments broad, obtuse, divergent in the lower leaves, postical lobe larger; epidermis not polished, cells smallish, roundish -quadrate, 4, 5, and 6-sided, walls thick, reddish-brown through transmitted light, trigones indistinct or wanting. Perigonial bracts 2-5 pairs, below the perianth, broadly orbiculate, bifid to about J or ^, sinus and segments obtusate or ^cute. Innermost bracts large, ovate, subquadrate, bifid to ^, MA I? SUP ELLA. 335 sinus and segments acute. Perianth of much more delicate cell- strncture, with distinct trigones, adnate with the lower third of the innermost bracts, divided for lialf its length into 5 irregular, denticulate se^'nients, terminal cells elongate, hyaline. Antheridia oval, stipitate, two in each bract. Pistillidia 8, surrounding base of calyptra. Capsule small, dark brown, almost spherical, walls composed of two layers. Fruits in Summer. D]:\[ EN SIGNS.— Stems ^ to 1 inch long, •!() mm. x -13 mm. diam., with leaves l" mm. wide; cells of stem, cortical layer •02 mm. X '025 mm., interior '015 mm. ; leaves "8 mm. x '8 mm., sinus -25 mm. deep, "85 mm. x "05 mm., sinus '25 mm.; cells •02 mm. X 'Oo mm., -02 mm.; bracts 1-2 mm. x l"! mm., sinus •4 mm.; perigonial bracts '9 mm. x "9 mm., sinus '2 mm.; inner- most bracts I'P) mm. x I'lmm., sinus '5 mm., 1'5 mm. x 1*3 mm., 1-4 mm. x I'l mm.; perianth 1-2 mm. high, segments '4 mm.; pedicel -21 mm. diam. ; capsule '5 mm.; antheridia '02 mm. x •0125 mm. Hab. — Grows on rocks in alpine situations. 15. Loch-na- gar, Aberdeenshire, /. Sj' T. Sim, August 14, 1870. Only known British station. Found on the Continent in Norway, Sweden, Lapland, Austria, and Switzerland; also in North America. Obs.^ — 'This is a very distinct species, altliough it has been confused even by Lindberg himself with M. if.^f/data (Hilben.), who has distributed both species under the name of M. fV. M. Macvicar ^- W. H. P. Obs. — One of the most beautiful and distinct of liepatics, dis- covered by Mr. George Stabler, of Levens, on Bow Fell, in 1875, and named after its discoverer by Dr. Spruce. 388 HEPATIC^. It is to be deeply regretted that, owing to ill-health and the loss of one of his eyes, this most acute and genial of naturalists was compelled to relinquish working among the hepaticie, for there were few better qualified to carry on the work left unfinished by the late Dr. Carrington. It will always be a pleasure to remember that his name is associated with one of the gems of his Lakeland. A remarkably distinct species, distinguished by its rosy purple colour from most others ; the only one with which it can be con- founded being Ereinonofiis myriocarpiiH (Carr.), with which it agrees in habit, and often grows with it. This species never has the purple colour of M. Sfableri, is furnished with numerous leafless stolons, stem leaves are smaller, divided to the middle, bracts quadrate, not broadly ovate, closely complicate, always free, lobes obtusate, entire, perianth rounder, carinate, protruding beyond and quite free from the bracts. M. Sfahh'vi is distinguished from the small purple form of Ceplicdozla (llraricafa (Sm.) without stipules, by its more imbricate, appressed leaves in its sterile state, and when fertile by its very different perianth. The description is taken from Dr. Spruce's contribution to "Eev. Bryol." for 1881. Description of Plate CLXX. — Fig. 1. Plant natural size. 2. Portion of fertile plant x 24 (Oxenden, G. Stabler). 3. Por- tion of stem X 85 (Mardale, G. Stabler). 4-9. Leaves x 85 (Langdale, G. Stabler & W. H. P.). 10. Portion of leaf x 290 (Mardale, G. Stabler). IL Outer bract x 85 (Langdale, G. Stabler & W. H. P.). 12. Bract x 85 (Oxenden, G. Stabler). 13. Ditto (Langdale, G. Stabler & W. H. P.). 14. Perianth x 85 (Bow Fell, G. Stabler). 15. Cross-section of perianth x 85 (Langdale, G. Stabler & W. H. P.). IG. Perigonial bract x 85 (ditto). 17. Antheridium x 85 (ditto). MABSUFELLA. 389 8. Marsupella nevicensis [Carri)/(/(oit), Kaalaa-s. Jumjermania nevicensis, (Jarr. in C. &, P. Hep. iJrit. Exsicc. ii. 85 (1870); Carrington in Trans. Bot. Soc. Edinb. (1880). Sarcosci/phas capillaris, Limpr. in Jaliresb. Schles. Gesellsch. vater]. Cult. 58, p. 182 (1880) {vide Kaalaas). Jli/l/robiella nevicensis (Carr.), Spruce on Cephalozia, p. 77 (1882). Xardia latifolia, Lindb. ap. Soc. F. FJ. Fenn, die 2 Deer. (1882). Marsupella latifolia, Lindb. in Meddel. soc. F. FJ. Fenn. 1:'., p. 288 (1886). Mars'iqjella nevicensis (Carr.), Kaalaas, Leverm. Norge, p. 117 (18'Jo). Dioicous, cnespitose, flagelliferous, small, pale green, here and there with a slight reddish tinge, when dry a dull yellow colour, Stems simple or slightly branched, flexuose, scorpioidal at the apex, naked below, 12 cells in diam., cortical cells '24, rather larger than the inner ; rootlets wanting or extremely sparse, reddish-purple tinge. Leaves distant, alternate, small, scarcely broader than the stem on sterile specimens, vaginate, upper ones erect, roundisli-ovate, lower erecto-patent, ovate, subquadrate, complicate-concave, rounded at the base, or subcordate, apex boat- shaped, J to ^ acutely bidentate, lobes acute, sinus acute ; cells small, 4-, 5-, and 6-sided, walls moderately thick but weak, without trigones in the British imperfect specimens, in perfect ones fully developed trigones distinct. No stipules. Bracts larger than the leaves, suborbicular to subquadrate, bifid to almost the middle, segments acute, sinus acute. Perianth small, delicate, hidden in the bracts, and adnate to about the middle to them, mouth laciniate, sterile pistillidia surrounding base of calyptra. Androecia terminal on main stem, 2, 3 pairs of leaves, imbricate, roundish, bifid to ^, complicate, ventricose, enclosing 1 or 2 large, oval antheridia. DniENSioNs. — Stems h to | inch long, l mm. to '2 mm. diam. leaves "ob mm. x -."jO mm., '375 mm. x '35 mm., "7 mm, x •7 mm., segments "2 mm. ; cells •0125 mm., '015 mm., '017 mm., •02 mm., '02 mm. x '03 mm.; bract r mm x "D mm., seg, ■5 mm.; perigonial bracts "5 mm. x b mm., seg. "15 mm.; antheridia '15 mm. x '125 mm. 390 HEPATIC^. Hab. — Discovered on moist shelving rocks, near the last spring below the summit of Ben Nevis, July 1875, by the late Mr. John Whitehead. This is the only known British station. Found on the Continent. Obs. — With only imperfect sterile British specimens collected, this species has been a puzzle to our leading authorities, but since good fertile ones have been met with on the Continent the difficulty has been solved, and it turns out to be a true Marsupella. No doubt the specimens collected by Mr. Whitehead were growing in a very moist situation, and had differentiated in a remarkable manner from the perfect state. A distinct species, and not likely to be confused with any other when met with. Dp:script]on of Plate CLXXI. — Fig. 1. Plants natural size (Ben Nevis, John Whitehead). 2. Portion of fertile stem x 24 (Norway, S. 0. Lindberg). 3. Portion of $ stem x 31 (ditto). 4. Sterile stem x 31 (Ben Nevis, J. Whitehead). 5-8. Leaves X 64 (ditto). 9-12. Ditto x 24 (Norway, S. O. Lindberg). 13. Portion of stem with leaves x 120 (Ben Nevis, J. White- head). 14. Portion of leaf x 290 (Norway, S. 0. Lindberg). 15. Bract x 24 (ditto). IG. Portion of involucre, showing portion of bract and perianth x 31 (ditto). 17-19. Perigonial bracts x 31 (ditto). 20. Antheridium x 85 (ditto). Genus 34. CESIA, Gr. & B. Jungermania, Lightf. Fl. Scot. 2, p. 786 (1777). Cesius, Gr. & Benn. Nat. Arr. Brit. PI. i., p. 705 (1821). Gijmnomitrium, Corda in Opiz Beitr. 1, p. 651 (1829). Acolea, Dum. Syll. Jung. Eur. p. 76 (1881). Cesia, Lindb. Muse. Scand. p. 9 (1879). Stems fasciculate, ascending or erect, creeping at the base, flagella radiculose ; glaucous green, reddish brown, or almost black. Leaves succubous, distichous, closely imbricated, emar- ginate-bidentate. Stipules wanting. Perianth wanting. Involucre double, inner shorter, composed of two or more involute deeply cleft and dentate free bracts, which enclose the short campanulate CESIA. 391 calyptra. Base of tlie younjs: fruit immersed in the hollow apex of the stem. Capsule globose, 4-valved, wlieii empty reflexed. Elaters bispiral, caducous. AndnL'cia distinct. Antheridia oval, stipitate, seated in the axils of the perigonial bracts. Section 1. IIOMOCRASPIS, Lhulb. S. 0. Lindberg, ]«8G, Schili". Engl. & Prantl, Pilanzenf. Dl k 1)2 Lief. p. 77 (1S;)3). Habit similar to Marst/jjcJla, but diflers usually in the absence of a true perianth. 1. Cesia alpina [Gotlsc/i.), Lliidh. Sarcoscyphus alpinus, Gottsch. in G. & R. Hep. Eur. n. 585 (1872). Cesia alpina, Lindb. in Meddel. soc. F. Fl. Fenn. 13, p. 251 (188G). Dioicous, broadly caBspitose, small, dark olive-green to reddish- brown, through transmitted light of a golden-brown colour. Stems creeping or suberect, simple or innovantly fastigiate, 10-12 cells in diameter; 2, 3 outer layers brownish-yellow, inner hyaline; radiculose, rootlets very short, white, or with a slight purplish tinge. Leaves loosely inserted, alternate or regularly pectinate, patent-divergent to patent (80^ - 40*^), somewhat distant, near apex imbricate, subiunplexicaul, decurrent at the base, ovate, cordate-ovate, cordate-orbicular, emarginate to \ or \, lobes obtuse or obtusate, slightly unequal, sinus obtuse ; texture firm, epidermis polished, cells small, hexagonal, containing 2 or 3 nucleate granules, walls thick, angles thickened, no trigones. No stipules. Bracts ovate-cordate, emarginate from 1 to \ deep, segments and sinus obtusate ; sometimes there is present an ovate lanceolate bracteole connate with the bracts. Perianth delicate, adnate with the bracts at the lower half or free, trilobate, margin irregular. Pistillidia about 10. Spores seen through transmitted light pale brown. Elaters bispiral, short, 4-G turns of the spiral, dark brown. Andrcecia terminal, crowded, 3-4 pairs of bracts, 302 HEPATICM which are broadly ovate, slightly swollen at the base, containing 1-3 oval antheridia. Fruits May, June. Dimensions. — Stems ^ to 1 inch long, '15 mm. diam., with leaves 1* mm. wide; leaves "55 mm. x "?>b mm., segments •125 mm., I'mm. x 75 mm., '8 mm. x -625 mm. ; cells "02 mm. x •025 mm., '02 mm. x '0275 mm.; sub-bracts "85 mm. x 'GS mm., •85 mm. x 75 mm.; bract 1*1 mm. x 7 mm., "8 mm. x 'G mm., segments 'lo mm.; perianth 'S mm. long; pistillidia "1 mm. X '04 mm. ; pedicel ' I ram. x "IG mm. ; capsule "5 mm. x 45 mm. ; spores '0125 mm., '015 mm.; elaters '1 mm. x "01 mm.; peri- p-onial bracts '65 mm. x '55 mm., seo^ments "125 mm. Hab. — Grows in broad patches on slanting rocks, usually unmixed with other species in alpine or subalpine situations. 7. The (ilyders, Carnarvon, ]^. M. Holmes; Pass ot" Llanberis, Carnarvon, April 1876, C. J. Wild t^" W. H. P.; Snowdon, near the Capel Curig Path, about 2500 ft., April 1878, //'. H. P. 12. Bow Fell, Cumberland, June 1881, Gcurfjc Sfah/cr cV //: H. P. (^ and $ fruit). 15. Glen Callater, Braemar, Aberdeen, July 1871, G. E. Hunt; Loch Kandor, Aberdeen, G. iJ. ILiiif; Tillylair, Aberdeen, May 1874, /. Sin/; Loch-na-gar, Aberdeen, October 1876, J. ^- T. Sim: Ben MacDhui, Aberdeen, ;3500 it., August 1880, ir. West; Ben Lawers, Perthshire, A. O. Plack. 16 Moidart, West Inverness, S. M. Macvicar ^- W. II. P. Found on the Continent. Obs. — Easily distinguished from all forms of J/, emaq/i/iafa by its colour, and the polished epidermis and close texture of its leaves ; and from M. FtincMi by the same characters and its obtuse lobes. These are the most likely species which it could be mistaken for, without a careful study of the involucre. Sometimes the perianth is a true one, and adnate to the lower portion of the bracts, but generally it is free and not perfectly tubular. Description of Plate CLXXIL — Fig. 1. Fertile plant, natural size. 2. Male plant, natural size. 3. Portion of stem x 24 (Ben MacDhui, W. West). 4-8. Leaves x 24 (Bow Fell, G. Stabler & W. H. P.). 9-12. Ditto x G4 (ditto). 13. Por- CESIA. 393 tionofleaf x 200 (ditto). 14. Bract x G4 (ditto). 15. Perianth, spread out x 24 (ditto). IG. Pistillidia x 85 (ditto). L7. Spores X 290 (ditto). J 8. Elater x 290 (ditto). 19. Perigonial bracts x 24 (Ben MacDhui, W. West). 20. Antheridium x 85 (ditto). 2. Cesia revoluta {Nec><), Liiulh. Sarcoscyphus revolutus, Kees, Nat. Eur. Leberm. U, p. il'.' (l!S;5(;). Xardia revoluta, Lindb. Revis. crit. Fi. Dan. p. 113 (1871) ; Carr. in Grevillua, 2, p. -SH (1873). Marsupellarevolata, Liudb. in Meddel. Hoc. F. Fl. Fenn. 13, p. 23S (1Sifoiunm among mosses." Differs from C. conc'iiinata, C. ohiiim and C. corallioide-s by its smaller size and darker colour, absence of the silvery glaucous tinge, margin of leaves never scariose. From C. crcin/Iafa, G., which it resembles in size, by being- less intricately entangled, of a darker colour, almost black, not greenish-brown ; margin of leaves entire, not crenulate-dentate. Description of Plate CLXXIV. — Fig. 1. Plants natural size. 2. Stem x 30 (nr. Ben Lawers, Dr. Black). 3. Portion of male stem x 62 (C. & P. n. 76). 4. Leaves x 64 (nr. Ben Lawers, Dr. Black). 5. Leaf x 24 (Ben Lawers, G. Davies). 6. Leaf x 31 (Ben Nevis, W. West). 7. Ditto x Qi (ditto). 8. Portion of leaf, near middle base x 290 (Ben Lawers, G. Davies). 9. Ditto x 200, from near apex of leaf 10, Bract x 31 (Ben Nevis, W. West). 11. Ditto x 64 (C. & P. n. 76). 12. Innermost bract, calyptra and capsule x 64 (ditto). 13-15. CKSIA. 397 rnncrmost bracts x 'M .(I'cii Lawers, (\. Davics). Ki. I*eri- gouial bract witli aiitheridiuiu x G2 (C. & 1*. n. 7(5). Figures 3 and IG drawn by Dr. (iottsche, othex's by Dr. Carrington and W, 11. P. 4. Cesia confer ta [Llm/jr.). S'arcoscj/phvs C07}fertus, Liiupvicht, Jahrb. Hchles. Cios. p. :\]l] (1X80). (Ij/innomitriinn confortum, Limpr. Flora n. 5 (18S1). Jfarsiipella covferia (Limpr.) Spruce, Rev. Bryol ii. (i. !)."i (l.SSI) ; Pears. Journ. of Bot. Sept. (1802). Monoicous, cjospitose oi* straggling among mosses, pale yellowish-green to reddish-brown in colour. Stems simple or branched, prostrate, flexuose, filiform, terete, denudate at the base ; branches short, ascending, about 10 cells in diameter, cortical cells witli slightly firmer walls ; radiculose, rootlets fasciculate, short, ascending to apex of the stem, few, more frequent below, dull white or reddish-purple. Leaves closely imbricate, bifarious, from a vaginate base, appressed, erect, on fertile stems gradually accrescent, oval, bifid to about 1 ; sinus and segments acute; texture firm, epidermis smooth, cells very minute to minute, roundish-quadrate or indistinctly 5-G angled, walls thick, trigones large and distinct. ? terminal, on short branches or on main stem ; bracts larger than the leaves, oval or oval-rotund, the innermost bracts free or sometimes united into a tube, small, tender, margin irregular. Pistillidia 5-10, dispersed over the calyptra. Pedicel long, thick. Capsule dark brown, almost spherical. Spores yellowish-brown, granulate. Elaters o-4- spiral, sometimes furcate. Andrtecia spicate, situated on the main stem or on short branches, perigonial bracts ovate, a little swollen at the base ; antheridia solitary, stipitate. Fruits Summer. Dimensions. — Stems from | to 1 in. lono-, diam. --Jmni., with leaves '5 mm. broad, leaves '65 mm. x T) mm., segments ']'25 mm., "G mm. x '45 mm., seg. '1:25 mm., '5 mm. x "4 mm,, seg. •125 mm., cells '0 1 mm., -OL^nim. x -ol mm., sub-bracts "45 mm. 398 HEPATIC^. X -45 mm., "5 mm. x "45 mm., bracts 75 mm. x "45 mm., segments •1 mm., -65 mm. x '5 mm., seg. -1 mm., "65 mm. x "5 ram., seg. •15 mm., perigonial bract "55 mm. x '425 mm., seg. '1 mm., antheridia -125 mm. x '075 mm. Hab. — 16. Grows in depressed tufts, or straggling among mosses. Ben Nevis, at about 4000 ft., W. West, August 12, 1880. The only known British station. Found on the Continent. Obs. — The British form differs from any of the Continental specimens I have seen in its larger size, but, as it agrees in its monoicous inflorescence, $ and ? on the same plant, but on different branches, and in the shape and texture of the leaves, I have little doubt they are one and the same species. Herr Limpricht, the founder of the species, described it at first as a Sarcoscjjphi(s[Marsiq)elIa), with a distinct perianth, but soon after- wards referred it to the genus, or, as Dr. Spruce maintains, sub- genus Cesia. In carefully examining every stem sent me by Mr. West, I met with two fertile ones which appeared to have true perianths, but the specimens were weathered and too imperfect to rely upon ; others, as figured, well represent the genus Cesia, where there is no true perianth, and the innermost bracts are free. Distinguisbed from other species of the same genus and from any of the Marsiipellce by its monoicous inflorescence. Dr. Spruce found in original specimens from Herr Limpricht a fertile invo- lucre which had a tubular perianth reduced to a short multilobate cup, confirming his contention as to the position of the supposed genus Cesia. Cesia covferta has been referred by some authorities to Cesia varians, Lindb. (Meddel. soc. F. Fl. Fenn. 13, p. 238 (1886), Nardia {Marsupella) varians, Lindb. Muse. Scand. p. 9 (1879), but I have found no variableness in the inflorescence of C. covferfa, nor does it agree in other particulars. When barren it might be overlooked for a small form of Jiivr/. miniita, Crantz, but from which it is very distinct. Description or Plate CLXXV. — Fig. 1. Plants natural size. 2. Portion of fertile stem x 24. 3-6. Leaves x 31. 7-9, CESIA. 399 Ditto X 24. 10. Leaf X G4. 11. Portion of loaf x 290. 12, 13. Sub-bracts x 64. 14, 15. Bracts with inner perichaetium x 64. 16. Bract x 64. 17. Pericfonial bract x 64. 18. Antheridium x 85 (Ben Nevis, W. West). '&' 5. Cesia brevissima [D/nn.). Jungerynania coyiclnnata 11 viinm', Schleich. Cat. exsicc. (1821). Acolea brevissima, Dum. Syll. Jung, p 7G (1831). Gjfmnomitrinm adustum, Nees, Nat. Eur. Leb. 1, p. 120 (1833); SchifFn. in Engl. & Prantl, Pflanzenf. 91 & 92 Lief. p. 77 (1893). Cesia aclusta (Nees), Lindb. in Meddel. soc. F. Fl. Fenn. 13, p. 238 (188(;)et 14, p. (59 (1888); Kaalaas Leverm. Norge, p. 42(5 (1893). Paroicous, shallowly ca^spitose, minute, dark brown colour. Stems intricately entangled, rhizomatous, rhizomes creeping, stems somewhat erect, rigid, ramose ; radiculose, rootlets ^ew, somewhat reddish, fertile stems clavate-fusiform. Leaves closely imbricate and accrescent on the fertile stems, more distant on the sterile, erect or erecto-patent, oblong or oval, bifid to J or ^, segments acute, somewhat unequal, sinus acute, rarely obtusate; texture firm, cells very small, roundish subquadrate, walls firm, trigones wanting, or small and very indistinct. Bracts larger than the leaves, oval or broadly oval, with one rarely two oval antheridia at their base, innermost bracts 2, of a very delicate texture, pale colour, connate with each other, broadly oval or roundish, quite free from the outer bracts, irregularly dentate, sometimes with a small ovate-acuminate bracteole present. Calyptra oval, sterile pistillidia 8-10, dispersed over its convexity. Capsule dark brown, spores small, pale yellowish or reddish- brown, elaters bispiral, as wide as the sjoores, dark reddish-brown. Dimensions. — Stems 2* to 3* mm. long, •] mm, diam. with leaves -3 mm. wide ; leaves "5 mm. long x -3 mm. broad, segments •075 mm., "4 mm. x 25 mm., seg. -05 mm., '4 mm. x -3 ram., seg. •075 mm., ^35 mm. x -2 mm., seg. ^075 mm. ; cells -02 mm. x •015 mm., ^02 mm. x -02 mm., -015 mm. x '015 mm. ; bracts '6 mm. long X -425 mm. broad, segments '1 mm., '5 mm x -4 mm., seg. "075 mm. '55 mm x '5 mm,, seg, "1 mm. ; bracteole -175 mm. x 400 HEPATIC.^. •i mm. ; antlieridia "15 mm. x '1 mm. ; innermost bract '5 mm. long X "35 mm. broad ; pistillidia '1 mm. long x "035 mm. broad ; spores '01 mm. diam. ; elaters •] mm, long x -01 mm. broad. Hab. — Growing on soft sandstone in alpine localities. Ex- tremely rare. 7. Clogwyndu Arddu, Snowdon, Carnarvonshire, August 1Scand. p. 10 (18711). Dioicous, closely caespitose, small, of a silvery green colour. Stems suberect, simple or ramose, cylindrical, about 15 cells in diam., filiform below, towards the apices incrassated and com- pressed, when dry brittle ; radiculose, rootlets ascending to apex. Leaves erect, bifarious, loosely imbricate, somewhat secund, concave, ovate, bidentate to about J,, sinus acute, segments acute, margin quite smooth, often hyaline; texture firm, epidermis smoothish, cells smallish, roundish-quadrate or roundish-oblong, walls distinct, trigones small but distinct. No stipules. Bracts bifid or trifid, somewhat similar to the upper leaves, innermost bracts smaller, two or three, free, irregularly laciniate-dentate, very delicate. Calyptra ovate, extremely delicate, cell structure hardly visible, arranged like a spider's web ; pistillidia dispersed over it. Capsule small, nearly spherical, of a reddish shining brown colour. Spores spherical, deep fulvous brown colour. Elaters similar in colour, bispiral. Male stems somewhat clavate ; perigonial bracts terminal, broadly ovate, swollen at the base, antheridia about 8, 1 or 2 at the base of each bract, almost spherical, bearers long, of 24 cells in two rows. Fruits May, June. 404 HEPA TICJE. Dimensions. — Stems |- to |- inch long, -2 mm. diam., with leaves from '4 mm. to '6 mm. wide, at apex of fertile stem 1* mm. wide; leaves l* ram. long x "75 mm. broad, segments '2 mm., •9 mm. X '7 mm., seg. '15 mm., '8 mm. x "6 mm.,seg. '15 mm. ; cells -025 mm. x '025 mm., '025 mm. x '02 mm., -03 mm. x •02 mm., "03 mm. x '03 mm. ; trigones '005 mm. ; bracts "75 mm. x "65 mm., "6 mm. x '5 mm., "55 mm. x "4 mm. ; perigonial bracts '8 mm. long x '75 mm. broad, '7 mm. x •7 mm. Hab, — Grows in cushion-like patches on exposed rocks in subalpine or alpine localities. Not uncommon at these alti- tudes. 7. Cader Idris, Merionethshire, /. RaJfs ; W. H. P. ; Snowdon, G. E. Hunt ; J. R. Bijrom ; Pass of Llanberis, E. M. Holmes; G. A. Holt; Glyders, Carnarvonshire, W. Wilson. 10. Summit of Ingleborough, W. J. Hooker; Br. Carrington ; Howgill Fells, W. West; Brimham Rocks, R. Teesdale, 1798; Cronkley Scar, Teesdale, /. G. Baker. 12. Helvellyn, John Nowell ; Bow Fell, Westmorland, G. Stabler. 13. Rocks on Kells Hills, /. Mc Andrew ; Hart Fell, Blackhope, Nichol. 14. 15. Mael Tarmachan, C. J. Wild ; Ben Cruachan, T. Rogers. 16. Glen Finnan, JJr. Carrington ; Moidart, West Inverness, S. M. Macvicar, not seen below 1400 It.; frequent from this to the summits. 17. 18. I have not seen any specimens from Ireland. Found on the Continent and in North America. Obs. — Distinguished from Cesia ohtiisa, Lindb. and the rare Cesia corallioides (Nees), in addition to other characters by the acute segments of the leaves. It is frequently mistaken for Anthelia julacea (Lightf ) by inexperienced collectors, from which it is very different. The innermost bracts differ considerably in various indi- viduals ; some stems having two, some three, and others only one, and are more or less laciniate-dentate, but are always free from each other and the outer bracts. Description of Plate CLXXVIII. — Fig. 1. Plants nat. size CESIA. 405 (Sowerby, Eng-. Bot. 2220). 2. Ditto. ,'}. Portion of fertile stem X 24 (Snowdon, W. H. P.). 4. Portion of barren stem x 24 (ditto). 5-7. Leaves x 24 (ditto). 8-10. Ditto (Mael Tarmachan, C. J. Wild). 11. Portion of leaf x 290 (Snowdon, W. H. P.). 12-14. Bracts x 64 (ditto). 15. Perigonial bract X 24 (Mael Tarmachan, C. J. Wild). 16. Ditto (Glen Finnan, Dr. Carrington). 8. Cesia obtusa, Llndh. Cesia ohtusa, Lindb. Muse. Scantl. p. 'J (1S7'J). Gymnomitrmm concinnata7n, var. crenulatum, Limpr. in Cohn Krypt. Schles. 1 , p. 24G (1877)? Dioicous, densely csespitose, small, silvery white, where ex- posed and old it is darker coloured, where shaded it has a greenish tinge. Stems erect when tufts crowded, with few branches assurgent, ascending to the height of the chief stem, when tufts loose, stem prostrate, creeping, fertile stems increasing in size to apex, which is blunt and swollen, barren shoots catenulate, on a transverse section almost round, cortical layer of cells firmer and darker coloured ; rootlets few, hyaline, short, arising from the postical side of stem where the two series of leaves join, ascending to about the middle of the stem, the leaves having to be carefully separated from the stem before the origin of the rootlets can be seen. Leaves closely and regularly imbricate, erect, hilarious, ovate, roundish-ovate or obovate, concave, bidentate to \ or J, sinus acute or slightly rounded, segments broad, obtuse, finely crenulate on both sides, lower margin of leaf quite entire ; texture somewhat fleshy, cells medium size to smallish, roundish- quadrate or roundish- oblong, marginal cells smaller, trigones small but very distinct. No stipules. Bracts larger than the leaves, broadly ovate, bidentate to about \, segments obtuse, finely crenulate. Innermost bracts free, very delicate, ovate, laciniate-dentate or denticulate ; bracteole small, oblong-quadrate, laciniate-dentnte or denticulate. Pistillidia lew (about 8), oblong, dispersed over the calyptra. Male stems slightly clavatej 406 HEPATIC^. antlieridia terminal or rarely situated at the middle of the stem ; perigonial bracts little different from the others, slightly swollen at the base and rather broader ; antlieridia few, oval, the outer layer persistent ; bearers as long or longer than the antheridia (some 12 cells long). Fruits May, June. Dimensions. — Stems i to l inch long, "2, mm. diam., with leaves '2 mm. to "3 mm. wide; leaves -9 mm. long x -7 mm. broad, '85 mm. x 7 mm., '65 mm. x "5 mm., segments "25 mm. high X from 'lo mm. to '4 mm. broad at the base; cells •05 mm. X '025 mm., "04 mm. x '03 mm., "04 mm. x '02 mm., •03 mm. X '02 mm. ; marginal cells '02 mm. x "02 mm. ; trigones "0075 mm., bracts "9 mm. long x "7 mm. broad, segments "2 mm.; pistillidia "15 mm. long x "05 mm. broad ; antheridia '25 mm. x 'IS mm. Hab. — Grows on exposed rocks in subalpine and alpine localities. Not uncommon in North Wales, the Lake District, and Scotland. 7. Cader Idris, Merionethshire, U. M. llohues ; IF. II. P. ; Cvvm Bychan, Merionethshire, G. A. Ilo/f ; Llanberis and Snow- don, Carnarvonshire, E. 31. Holmes, J. 11. Byrom, James Neild. 12. Harter Fell, Mardale, G. Stabler; Hill Bell,/. A. Marluidale (^- G. Stabler; Long Sleddale ; Langdale, Westmorland, G. Stabler. 15. Loch-na-gar, J. ^- T. Sim. 16. Glen Finnan, Dr. Carrin(jton ; Moidart, West Inverness, S. M. Macvicar, rare on low ground, but descends to sea-level ; common on the hills from 1200 ft. to 2000 ft. I. Mwellrea Mountains, Co. Mayo, Dr. B. Moore, 1874; Slieve Donard and other mountains in the North, Bevs. Lett & Waddell. Found on the Continent. Obs. — From Cesia concinnata (Lightf.)it may be distinguished b}" its rounded, obtuse, finely crenulate segments, closel}^ and regularly imbricating leaves, bifariously inserted, never secund (as Lindberg remarks). Cesia ereuiilata (Gott.) is a widely different species, with finer CESIA. 407 stems, arcuately brancliecl, dark brown or almost black in colour, in shaded situations greenish ; in this species the segments are acute, with the hyaline fringe of leaf composed of narrow sharp- pointed, irregularly projecting cells. Cesia coral lloidcH (Nees) has a more acute leaf, without sinus, or if present, very shallow, or very irregular through being weathered, and always has a very delicate diaphanous hyaline border, so delicate that rarely any cellular structure is to be observed. This character, along with the entire absence of any approach to crenulation of the segments, at once separates it from C. ohtuHa. Cesia ofjfiisa (Lindb.) was first recognised as British by Mr. George Stabler and published in " Journal of Botany " for 1880. Description of Pj.ate CLXXIX. — Fig. 1. Plants nat. size. 2. Portion of stem x 64 (Cader Idris, E. M. Holmes). 3-7. Leaves x 81 (Glen Finnan, Dr. Carrington). 8, 9. Ditto x 31 (Scotland, Croall). 10. Lobes of leaf x 85 (Norway, Lind- berg). 11. Portion of leaf x 290 (Grlen Finnan, Dr. Carring- ton). 12, 13. Bracts x 31 (ditto). 14, 15. Portions of the innermost bracts x 04 (Mardale, G. Stabler). 10. Bracteole x G4 (ditto). 9. Cesia crenulata [Gollsc/ic). Gymnomit7'iiim crenulatuvi, (\ottsche in dxvrington'a Iii!-li TTepatica^, Trans. Bot. Soc. Edinb. vii. (18r.:5). Acolea crenulata, J)um. Hep, Eur. p. 125(1874). Dioicous, densely coespitose, tufts very depressed, small, of a dark reddish or greenish brown or almost black colour. Stems with about 20 cortical cells, reddish colour, 10 cells in diam. ; intricately entangled, creeping, prostrate or suberect, arcuaiely branched, filiform, wiry, catenulate, terete, or somewhat com- pressed then lanceolate in outline ; radiculose, rootlets few. ]jeaves closely imbricate, erect, scarcely broader tlnm the stem, oval, ovate or broadly ovate, bidentate to about \, sometimes 408 HEFA TWxE. sub-entire, segmiuts acute, margin scariose, irregularly crenulate- dentate to the base ; texture firm, marginal cells delicate, hyaline, narrow, the next inner row suddenly tliickened and coloured, small to smallish, roundish-quadrate ; walls thick, angles thickened. No stipules. Fertile stems short ; bracts about 3 pairs, larger than the leaves, innermost bracts smaller than the outer, bifid or rarely trifid, fringed with long teeth to the base, free from the outer bracts. Cal3q3tra oval, extremely delicate ; pistillidia dispersed over it. Capsule spherical, pale brown. Spores small ; elaters somewhat short, brown, bispiral. Androecia terminal on short branches ; perigonial bracts not so closely imbricate as the leaves, slightly secund, broader than the stem leaves ; antheridia oval, stipitate. Fruits Mav and June. Dimensions. — Stems i to f inch long, diam. -15 mm. with leaves '3 mm. wide; leaves '35 mm. x '3 mm., seg. *05 mm., •275 mm. x •25mm., seg. "05 mm., "3 mm. x '3 mm., seg. •075 mm.. *25 mm. x '2 mm., seg. '03 mm. ; cells '02 mm., ^02 mm. x •025 mm., "025 mm. ; marginal cells ^04 mm. x ^02 mm., •03 mm. X '015 mm. ; innermost bracts ^25 mm. x "25 mm., seg. "05 mm., "3 mm. x '275 mm., seg. '075 mm. ; pistillidia -15 mm. x •05 mm. ; perigonial bracts ^5 m.m. x ^45 mm., seg. '1 mm., •35 mm. x "4 mm., seg. "075 mm. ; antheridia •IS mm. x •! mm. Hab. — Growing in intricately entangled round cushions or in patches on exposed rucks or stones in alpine or subalpine localities, and also at a lower elevation than cither Cviila conciuuala or Ce^ia obtiiHci. Not unfrequent. 1. Dartmoor, Devon, G. E. Bavies. 7. Cader Idris, Snowdon and other Welsh mountains, W. Wlhon; E. M. Holmes ; IF. H. P. 12. Mardale, Westmorland, G. Stabler; Borrowdale, Cumberland, IF. II. P.; frequent in the Lake District, G. Stabler. 13. Fre- quent on the hills. New Galloway, ./. McAmlrew. ] 6. Glen Finnan, Dr. Carriufjton ; frequent on rocks and boulders on the hills, rarely below 1200 ft. ; noted from 700-2500 ft., and in one locality at sea-level, Moidart, West Inverness, S. M. Macvicar. I. Frequent on the higher mountains, and in some few ami A. 409 instaaces descending to sea-level, Dr. D. Moore. On rocks near the tunnel, Cromaglovvn, Dr. Carriiujlon ; Dankerron and Knock- avoila, Dr.Tai/lor ; Carnintual, Dr. J). Moore; Lugnaquilla, Co. Wicklow, and Graltymore, A. Carroll ; rocks on the Hill of Howth, Br. B. Moore ; Golden Kiver, Co. Louth, llev. 11. W. Lett. Found on the Continent (Norway). Obs. — This is a very distinct species and abundantly different from any other of the genus, Cesia ohtma, Lindb., is far removed from it by its larger size, different colour and shape of leaves, the marginal cells of which are rounder and smaller. Dr. Carrington was the first to distinguish this species, which had been referred by Dr. Taylor to Cesia coraUioideH (Nees). He sent specimens as a new species to Dr. Grottsche, who gave it the characteristic name of crenidata. Although generally distributed in the mountainous parts of the British Isles, it appears to be very rare on the Continent, and not well understood by some authorities, Herr Limpricht, a most acute hepaticologist, and whose devotion to the European mosses has evidently interfered of later years with his study of the Hepaticae refers Cesia oblitsa to C. crenulala, as a variety of C. concuuiaia (lA^\\it),[G^uiiioniitriuiu eonclnnat((ui,Ni\M. crenulattim), (Cohn's Krypt. Schles. 1, p. 24G (1877) ). It generally grows at a lower elevation than any other species of the genus. Mr. Macvicar has found it at sea-level in Scotland, and it is one of the commonest species on rocks and walls about Llanberis in North Wales, and Seathwaite in Borrowdale. Description of Plate CLXXX. — Fig. 1. Plants natural size. 2. Branch x 20 (Dr. C). 3. Fertile branch x 40 (ditto). 4. Apex of barren branch x GO (ditto). 5. Portion of stem x 24 (Pass of Llanberis, W. H. P.). G. Leaf x 85 (ditto). 7. Ditto (Cader Idris, W. H. P.) 8. Leaves x (Dr. C). 1). Portion of the apex of a leaf, showing the crenate margin x oOO (Dr. C). 10. Portion of leaf x 290 (Long Stile, G. Stabler). 1 1. Marginal cells X 290 (ditto). 12, 13. Bracts x 85 (Pass of Llanberis, W. H. P.). 14. Bract enclosing the calyptra and capsule x (Dr. C). 15. Pistillidia surrounded by the innermost bracts x 410 HEPATIC^. (ditto). IG. Pistillidium x 85 (Pass of Llanberis, W. H. P.). 17. Spores and elaters x (Dr. C). 18. Perigonial bract x 85 (Pass of Llanberis, W. H. P.). 19. Antheridium x 85 (ditto). Genus 35. ACROBOLBUS, Nces. Acrobolbus, Nees in G. L. N. Syn. Hep. p. 5 (1844). Gijmnanthe, Tayl. MSS. G. L. N. Syn. Hep. p. 192 (1844). Plants small, semi-parasitic. Stems prostrate, radiculose, simple or furcate, rarely with one or two sublateral branches. Leaves succubous, alternate, subquadrate, bifid, upper sometimes trifid, quite entire, ^or here and there dentate ; cells medium size. Stipules absent. LiHorescence dioicous. Bracts $ 1-2 pairs, large, plurilobate, undulato-crispate. Pistillidia 10-22, Mar- supium terminal, seated at right angles with the stem, short, bulbous, obovato-globose, carnose, mouth without or very rarely with subulate scales. Calyptra adnate up the apex with the marsupium, crowned with the sterile pistillidia. Pedicel calceo- late at the base. Capsule oblong. Antheridia terminal. Acrobolbus Wilsoni (Te/j//.) Nces. Jungermania Wilsoni, Tayl. in schedis. Gymnanthe Wilsoni, Tayl. G. L. N. Syn. Hep. p. 102 (1844). Acrobolbus Wilsoni, Nees, G. L. N. Syn. Hep. p. 5 (1844). Dioicous, csespitose, tufts scattered, small, pea or dark green in colour. Stems creeping, flexuose, colour olive or olive brown near the base, cells 10 x 10, cortical similar to the inner; radi- culose, rootlets abundant, clothing the postical side of the stem, dull white, somewhat coarse. Lmovations postical or sublateral, slender. Fertile stems suddenly accrescent at the apex and bearing much larger erect leaves. Leaves appro:5:imate, semi- vertical, roundish or obovate, divided for one third or even half their length into two acute unequal lobes, the antical somewhat larger, rarely trilobate ; sinus acute ; margin entire or with an ACnOBOLBUS. Ill occasional tooth, insertion oblique, contracted at the base, nearly plane and horizontal at the inCerior portion of tlie stems, but more concave, ascending and connivent near tlie apex ; texture fragile, somewhat thin, but flesh}^ cells medium to rather large in size, roundish-hexagonal or roundish-quadrate, walls thick, angles thickened, trigones hardly perceptable, lumen filled with chlorophyllose granules. Stipules wanting or rarely present as minute scales at the apex of innovations. Fructification terminal, but from the growth of innovations sometimes appearing lateral. Bracts 2, larger than the leaves, broadly ovate or cordate, 2 to 3-lobed, the lobes repand-dentate, so as to have a crisped appear- ance, convolute-concave, base saccate-amplexicaul. Bracteole semi- ovate, acuminate, margin irregularly dentate, connate with one of the bracts, sub-bracts rather smaller, similar in size, but only bilobed, and with margins more or less entire. The bracts are originally free, surrounding the convex apex of the stem, on which 15-20 pistillidia are crowded. But after impregnation the rapid cell-growth of the young germ extends to the ' torus pistillorum,' and the bases of the adjacent leaves, so that the whole are blended together to form an involucre, the lower half of which is ob-conic and entire, projecting below the level of the stem, and forming a kind of bulb, from which numerous rootlets proceed. The mouth of the involucre is surmounted by two connivent bracts and bracteole, which retain their old form. On longitudinal section the structure will be better under- stood, the sub-bracts (which approach the terminal ones in size) obscuring it. The capsule is found occupying a cavit}^ composed in part of the metamorpliosed apex of the stem and bases of the bracts, and in part of the calyptra, which, with the exception of the dome-like apex, is concrete with the outer walls, bearing around it the remains of the abortive pistillidia, which surround the mouth of the involucre like a frimj^e. Perianth wantino;. Calyptra campanulate, of thin hyaline texture, adnate except at the apex with the fleshy involucre. Capsule oval, dark brown, divided into four valves. Pedicel stout, fleshy, white, bulbous at the base, and inserted into the thickened gibbous portion of 412 HEPATWjE, the involucre. Spores reddish-brown, minutely granular. Elaters obscurely bi-spirah Male inflorescence on slender branches, perigonial bracts 2 or 3 in number, terminal, sub-complicate and saccate at the base, enclosing one or more oval, shortly stipitate antheridia. Fruits in Autumn and Winter. Dimensions. — Stems J to |- an inch in length, diam. '2 mm. with leaves explanate 2* mm. wide; leaves 1*6 mm. x 1"5 mm., segments '6 mm., 1'2 mm. x 1"2 mm., seg. '6 mm., lower leaves 1* mm. X 75 mm. ; cells "05 mm. x "035 mm., '04 mm. x '04 mm., •035 mm. x "035 mm.; sub-bracts 2' mm. x 2' mm., seg". '5 mm. 2* mm. x 1'75 mm., seg. '75 mm.; bracts 1*75 mm. x 2' mm., seg. *75 mm., 1'75 mm. x 1'75 mm., seg. '75 mm. Hab. — In scattered tufts, epiphytic on BculuJa or Frullcuiia. Extremely rare. I. Near Bantry, Miss Hutchins, 19th Nov. 1812, young fruit. Banks of a ravine near the Hunting Tower,Cromaglown, Killarney, Nov. 1829, fr., IF. IFilson. Tore Mountain, Killarney, Sept. 1841 ; Glengariff, Sept. 1839, growing on FruUania germa7ia. Dr. Taylor. Obs. — This is a curious and extremely rare species ; the absence of a true perianth separates it from Nardia geoscypliiis (De Not.) and Jung, capitata. Hook , which it resembles somewhat in habit, but both are paroicous. This character serves to distinguish them from it, even when their perianths are not fully developed. Jung, capitata has also larger and more translucent cells with thinner walls. The above description is almost entirely taken from the late Dr. Carrington's uncompleted " British Hepatica?." Descuiption oe Plate CLXXXL— Fig. 1. Plants natural size. 2. Portion of stem, antical view, x 16 (Knock~a-voila, Dr. Taylor). 3. Portion of fertile stem x (Dr. Carrington del.) 4-7. Leaves x 16 (Knock-a-voila, Dr. Taylor). 8. Portion of leaf X 290 (ditto). 9, 10. Sub-bracts x 16 (ditto). 11, 12. Bracts X 16 (ditto). 13. Involucre x (Dr. Carrington, del.) SACCOGYNA. 413 Genus 30. SACCOGYNA, /)>>>,/. Saccoijyna, Dum. Comm. But. (18i';5). Dioicous. IMaiits with the habit of C/iiloxri/p/nix, procumbent, sparingly radiculose ; leaves succubous, horizontal, entire or emarginate ; stipules triangular, adiuite with the adjacent leaves. Perigjnium at first bud-like, postical, proceeding iroin the axil of a stipule at length oblong, fleshy, saccate, naked, smooth or everywhere pilose, mouth circular ; pendulous, affixed laterally by the margin of the apex, sublateral, shortly stipitate, surrounded by the remains of the involucral bracts. Perianth absent. Calyptra connate for most of its length with the outer walls, the dome-like ajjex alone free. Capsule oblong, 4-valved, valves straight, erect. Elaters bispiral, deciduous. Male spikes minute, postical, arising from the axils of the stipules of separate individuals. Saccogyna viticulosa {Linn.) Dum. Junyertnania terrestris viticulis longis, foliis perexigiiis, densissimis, ex roiundi- date acuminatis, Mich. Nov. gen. p. 8, t. 5, f. 4 (1729). Jungermania viticulosa, Linn. Sp. pi. 1.597 (1753) ; Hook. Brit. Jung. t. Go (1816). Saccogyna viticulosa, Dum. Comm. Bot. p. 113 (1822). Dioicous, broadly and shallowly csespitose, medium to laro-ish in size, yellowish green or olive brown in colour. Stems simple or slightly branched, branches postical ; radiculose, rootlets i'ew, fasciculate, wdiite, proceeding from the base of the stipules. Leaves succubous, regular, imbricate, sub-opposite, horizontal or patent-divergent, plane or slightly concave, ovate or ovate- triangular, antical margin decurrent ; texture firm, cells smallish, 4, 5, and G-sided, walls thick, trigones absent or indistinct. Stipules connate with the adjoining leaves or leaf, broadlj' subu- late, acuminate or orbicular-acuminate, irregularly dentate. Periantli produced from the postical side of stem below a stipule, and hanging by a short pedicel proceeding from its upper border, 4 H EPA TIC M. oval or oblong, cylindrical, clothed with long delicate, white rootlets, mouth and upper portion furnished with oval or sub- quadrate, dentate, delicate bracts, base solid. Calyptra oblong- obovate, adnate at the lower portion to the perianth ; pistillidia 7 to 12, very long and narrow. Capsule oblong, reddish brown. Spores reddish brown. Elaters bispiral. Androecia bud-like, ovate, proceeding from the postical side of stem, in the axil of the stipules, composed of 4 or 5 pairs of bracts with bracteoles, bracts crowded,broadly ovate, trifid,ventricose at the base, bracteole broadly subulate, or subulate, bifid, texture much more delicate than either the leaves or stipules, antheridia single, oval, stipes long. Fruits in Spring and eaidy Summer. Dimensions. — Stems 1 to 2 inches long, "o mm. in diam., with, leaves 3" mm. to 4' mm. wide; leaves 2' mm. x 1'5 mm., 1'5 mm. X 1"1 mm.; cells "Oo mm. x "02 mm., '035 x "03, '025 mm. x -025 mm., '03 mm. x "025 mm. ; stipules '7 mm. X '4 mm., 'Gmm. x "Smm., '45 mm. x "2 mm., perianth 2* mm. X -9 mm.; scales on the perianth "7 mm. x -5 mm., •6 mm. x •4 mm., ■4 mm. x '3 mm., "3 mm. x •2 mm.; pistillidia, •3 mm. X -04 mm., perigonial bract, "35 mm. x -3 mm. ; antheridia •2 mm. X '14 mm. ; perigonial bracteoles "25 mm. x '125 mm., segments -15 mm., "3 mm. x '2 mm., seg. '075 mm. Hab. — In various parts of Britain, but more especially in subalpine localities, on damp walls, rocks or stones, often in exposed situations, more rarely in bogs amongst Sjjliafjna. Somewhat local in its distribution, but is often ver}^ abundant where found draping damp walls, which are its favourite habitat to the exclusion of other species. 1. Common in Cornwall, ^V . Ct/rnoir ; Linton, North Devon, T. H. Bifssell. 3. Tunbridge Wells, G. E. Bavic^. 5. Banks of Severn by Seckley Wood, Staffordshire, /. E. Bagnall. 6. Neath, South Wales, Er. Carri7}gion ; Swansea, South Wales, C. W. Slafer S JF. 11. P. 7. Near Bangor, Carnarvonshire, IF. Wihon ; Dolgelly, Jo/iu Wliiidicad ; Maentwrog, W. H. P. ; Cader Idris, J. Balfs ; Barmouth, Dr. Carrivcjion & W. B. P. ; Tyn-y-Groes, Merionethshire, Er. Carriii(jioii tf JF. //. P. 9. Bamford Wood, SACCOGYNA. 415 Lanes., ('. J. Wild. 10. Bracken Gliyll, Dent, with niarsupia and ^ plants, 6'. Shihlcr ; Mai ham Cove, II. 'rcc^dalc; Halifax, /. Ballon, 1775; Dulosg-atc, Todniorden. John Xoircll. 10. Uydal, G, Slaljlcr ; by river Lune, near Orton, Dr. Carrin/jlon ; Jilack Crag, Staveley, G. Slaljler, Stock Ghyll ; Windermere $ and $ ; Langdale; Brown Ghyll, Oxenden, Westmorland, G. Slnhler ; Ulpha, Westmorland, /. Barnes ; Borrovvdale, Cumberland, Dr. Carrlnfjlon ^' W. II. P. 13. Wet rocks, Lochar Moss, James Crnicks/iank ; Brennan Hill, &c., Kirkcudbright, J. Mc Andrew. 15. Kinnordy, C. Li/cll. Ki. Very common on wet banks, iMoidart, West Inverness, S. M. Macvicar. I. Frequent occurrence in many parts of Ireland, but more especially in the south and west ; very fine at Lough Bray, AVicklow, and the woods about Killarney. It also extends to the counties of Antrim and Donegal in the north, and Mayo in the west, Br. D. Moure ; on a damp bank, Howtli demesne, D. McArdle ; on damp ground, Ballyhaise woods, Co. Cavan, D. McArdle. C. Saint-Ouen, /tiff/. Marl in. Found on the Continent, but ver}^ rare. The male flowers and fruit are extremely rare. j\[r. Wilson met with fruit near Bangor, March 1835, C. Lyfdl young fruit at Kinnordy, Prof. TAndberg fruit at Killarney, July 1872, J//-. Ralfs young fruit on Cader Idris, Mr. John Whitehead male flowers and fruit at Dolgelly, Mr. Geortje Slabler young fruit and flowers near Dent and Windermere, and some years ago I collected it in fruit in great abundance on the left-hand side wall, on the road to Harlech, about a mile beyond Maentwrog, North Wales, where it was again found in 1899 with abundant fruit by Mr. D. A.Jones, of Harlech. Obs. — A very distinct species and only type of the genus, not to be confounded with any other British species. Dksciiii'tton of Plate CLXXXII. — Fig. 1. Plants, sterile, ^ and ?, natural size. 2. Portion of stem, antical view x 1 1 (Dolgelly, J. Whitehead), o. Portion of stem, postical view x 24 (Dolgelly,. J. llalfs). 4. Portion of leaf x 290 (Dolgelly, J. 41 G HEPATTC.F.. Wlntelieacl). 5. Stipule x 85 (Dolgelly, J. Ralfs). 6. Perianth X 16 (Dolgelly, J. Wlntelieacl). 7, 8. Scales from near mouth of perianth x 24 (ditto). 9, 10. Ditto from, lower down X 24 (ditto). 11. Pistillidia x 85 (ditto). 12. Perigonial bract X 85 (ditto). 13. Antlieridinm x 85 (ditto). 14, 15. Perigonial bracteoles x 85 (ditto). Subtribe YII, F08S0MBE0NIE^. Genus 37. FOSSOMBRONIA, Racldi. Fossomhronia, Raddi in Act. )Sc. Soc. Moden. (1S20). Dioicous or monoicous, growing in tliin layers, usually densely caespitose, small, pale or yellowish green in colour, inodorous or fetid. Stems short, fragile, simple or furcate, often innovant, no postical branches, antical plane or canaliculate, postical rotundate or narrowly carinate ; creeping ; densely radiculose, rootlets very long, violet-purple, red or rarely white. Leaves large, succubous, obliquely or sublongitudinally inserted, antical margin decurrent, subquadrate, often longer than broad, usually narrow at the base, undulate, rarely plicate, apex broadly lobulate or sinuate, some- times subdentate ; cells moderately large, in some species two and three layers thick. Stipules absent. Female flowers terminal. Bracts similar to the leaves, but rather larger and more incised. Perianth campanulate, pyriform or turbinate, mouth wide, lobate, often longitudinally plicate, upper part tender and composed of a single layer of cells, low^er portion composed of two or three laj^ers. Calyptra free, subglobose or pyriform, much smaller than the perianth, at the base thickened, sometimes with a few sterile pistillidia surrounding it. Pedicel thick, short. Capsule globose, rupturing irregularly into 4 valves, bistratose. Spores large, more or less brownish, tetrahedral, convex face variously armed, lamellate or papillate. Elaters short, obtuse, normally bispiral, sometimes 1 or 3 loosely twisted. Antlieridia 2 or 3, aggregated, exposed on the antical face of stem, orange, shoitly ]:edice]late, afterwards mixing with the pistillidi^,. FOSSOMBRONTA. 417 Obs. — The genus Fossombronia is a very curious one, and naturally comes at the end of the foliose genera ; it has characters in common both with the foliose and frondose ; for instance, a true stem with leaves, but the latter are usually composed at their base of several layers of cells as with most of the frondose. An extremely interesting note on the genus is to be found in the" Species Hepaticarum "of Herr Stephani, whose arrangement of the species I have followed. Six British species have been observed up to now, which are distinguished by their inflorescence, smell, shape of stem and leaves, and more especially by the markings of their spores, full particulars of which are given with each species. FOSSOMBEONIA. (A) Spores vvith lamellae resembling crests, furcate. a. LamelUe laxly disposed. Fossombronia pusilla (L.), Dum. h. Crests crowded and numerous. Fossombronia cristata, Lindb. (B) .Spores vvith lamellfe regulai'ly reticulated. c. Margin appearing as if winged. Fossombronia angulosa (Dicks.), Raddi. d. Lamellae prominent at the margin. Fossombronia Dumortieri (H. et G.), Lindb. (C) Spore.s with lamellae compressed into papilla'. e. Papillae truncate. Fossombronia crfspitiformis, De Not. f. Papilla? long, hispid. Fossombronia Mittenii, Tindall. 2d 418 EEFATlCJi:. (AS pores with lamellae resembling crests, furcate. a. Lamellae laxly disposed, 1. Fossombronia pusilla {Bill., L.), Bum., Lindh. Lichenastrum minimum, capitulis nigris lucidis, Dill, in Ray. Syn. stirp. Brit. iii. ed. p. 110, n. 5 (1724). Jungermania pusilla, L. Sp. pi. i. ed. ii. p. 1136, n. 27 (1753), Maurocenia pusilla, Gr. & Benn. Nat. Avr. Brit. PI, i. p. G87 (1821). Codonia pusilla, Dum. Comm. Bet. p. Ill (1822). Fossombronia pusilla, Dum. Recueil, i. p. 11 (1835). Monoicous, very fetid, caespitose, small, green in colour. Stems procumbent, about three times broader than thick, simple or furcate, antical side plane, postical rounded ; radiculose, root- lets numerous, long, purple-violet. Leaves 3 to 4 pairs, close, obliquely decurrent, horizontal, subquadrate or narrow at the base, undulate, crisp, entire, irregularly emarginate or slightly incised, segments erect, acute or obtuse, those surrounding the perianth (bracts) larger; cells medium size, subquadrate, walls thin, no trigones or thickened angles. Perianth obpyramidal or turbinate ; mouth wide, recurved, irregularly dentate or slightly incised. Calyptra pyritbrm, whitish. Capsule spherical, reddish- brown, walls thin, delicate, bursting irregularly into 4 valves. Spores brown, tetrahedral-globose, with radiating furcate darker coloured lamellse laxly disposed, margin with 16 to 24 blunt teeth, Elaters pale brown, bi-rarely uni- or tri-spiral. Anthe- ridia situated on the antical side of stem, near the base of the leaves, exposed or enclosed by a bract-like process, spherical, with rather a long pedicel. Fruit ripens in Autumn. Dimensions. — Stem about ^ inch long x 2* mm. wide with leaves ; '7 mm. wide, x 2 mm, thick ; leaves 1'5 mm. x 1*5 mm., 1'25 mm. x 75 mm., 1'5 mm. x T mm,, 1' mm. x I' mm., seg- ments "2 mm., '3 mm. ; cells "05 mm. x '04 mm,, '04 mm, X -04 mm., -035 mm. x '03 mm. ; spores '04 mm., 'Oo mm,; elaters '\b mm. x '01 mm. Hab, — On clay or sand, in shady ditches and on damp wood- FOSSOMBRONIA. 419 land paths, by the margins of dried pools, in fallow fields or plou(,^hed lands. Somewhat rare. 1. Sidmouth, South Devon, Mrs. Ella M. Tindall 2. Hay- ward's Heath, Sussex, G. E. Bavies, New Forest, C. Li/ell. 3. Ditching, Surrey, G. E. Davles. Woolwich, Middlesex, Billenius. 10. Barmby Moor, Yorks, R. Spruce. 13. New Gallowa}'', /. Mc Andrew. T. Dingle Bay, Prof. Lindberg S(' Br. B. Moore. Found on the Continent and in North America. Var. oc/tros/jorr/, Lindb. Not. pro. F. et Fl. Fenn. p. 387 (1874). 9. Ashley, Cheshire, G. E. Hi(nt. Winwick, Lancashire, W. Wihon. Eccles, Lancashire, Br. Carrington. Todmorden, Lancashire, J. Nowell. Crumpsall, Lancashire, C. J. Wild. 10. Yeadon, Yorkshire, Z>/*. Crt^v^/zy/oy^ 12. Foulshaw, Westmor- land, G. Stabler. I. Kerry, Br. D. Moore. Obs. — Under this name all the other British species of the genus, with the exception of Fossombronia angidosa (Dicks.), were formerly grouped. It belongs to that section of the arrangement of Herr Stephani, with the laraelLc of the spore furcate, in which only another British species is included, F. cristafa, Lindb., and in addition to other characters, is distinguished from it by the fewer number of crests, which are more laxly disposed. The var. oclirospora, so far as I am able to understand it, differs but slightly from the type. Description of Plate CLXXXIIL — Fig. 1. Plants natural size (Hooker). 2. Fertile plant magnified (ditto). 3. Cross- section of stem X 24 (Sidmouth, Tindall). 4-7. Leaves x 11 (Hay ward's Heath, G. E. Davies). 8. Leaf x IG (ditto). 9. Portion of leaf x 290 (ditto). 10. Perianth magnified (Hooker). 11. Spore x 290 (Sussex, Davies). 12. Ditto x 400 (S. 0. Lindberg). 13. Ditto (Corbiere). 14. Elater magnified (Hooker). 15. Portion of stem showing antheridia, magnified (Hooker). IG. Ditto. 17. Antheridium magnified (ditto). 420 HEPATIC^. h. Crests crowded and numerous. 2. Fossombronia cristata Lindb. Fossomhronia cristata, Lindb. apud Soc. pro F. et Fl. Fenn. (1873), Not. pro F. et Fl. Fenn. 382 (1874). Monoicous, inodorous or very slightly fetid, csespitose or sparsely gregarious, small, green or yellowish-green in colour. Stems pro- cumbent, apex suberect ; radiculose, rootlets reddish-purple, numerous. Leaves crisp, subquadrate, obtusely or acutely lobed ; texture delicate, cells medium size, quadrate, walls thin, no trigones or thickened angles. Perianth obpyramidal, mouth wide, irregularly dentate or incised. Capsule globose, dark brown. Spores dark brown, tetrahedral-globose, densely cristate, about 10 crests across, sinuous or sub-parallel, 28-36 minute spines projecting around the margin. Elaters pale brown, short, thick, bi- rarely uni- or tri-spiral. Fruit ripens in Autumn. Dimensions. — Stems about \ inch long, '3 mm. thick ; leaves 1* mm. X 1- ram., segments '2 mm., '3 mm., 7 mm. x •8 mm. ; seg. '1 mm., "2 mm. ; '7b mm. x "75 mm. seg. "2 mm., •3 mm.; '9 mm. x '8 mm., seg. "3 mm., '4 mm. ; cells -05 mm. x •04 mm., -04 mm. x '03 mm. ; bract 1-25 mm. x 1-25 mm., seg. '4 mm. ; calyptra "7 mm. x '5 mm. ; capsule "4 mm. ; spores •04 mm., •OS mm. ; elaters "13 mm. x •Ol mm. Hab. — On bare damp soil on footpaths, sides of ditches, margins of ponds, and fallow fields. Eare. 2. Aston and New Forest, Hampshire, C. Lyell. Ashdown Forest, G. E. Davie-s, November 1884. 5. Staffordshire, Rev. H. P. Reader. 10. Side of pool, near R. Foss, Yorkshire, W. Ingham. 16. Moidart, West Inverness, 8. M. Macvicar. I. On whitish clay, shore of Loughbrickland, Co. Down, Rev. H. IF. Lett, October 1890. Found on the Continent and in North America. Himalaya, Gamble vide Stephani. Obs. — Distinguished from Fossombronia jy?^5z7/, 1(5 (1832), Jungermania Ralfsii, Wils. Eng. Bot. Suppl. t. 2874 (1843). BiplolcEna Lyellii, var. lamellata, Nees, Nat. Eur. Leberm. Ill, pp. 345, 852 (1838). Petalophyllum Ralfsii (Wils.), Gott. in Lehm. Pug. pi. nov. et min. cogn. viii. p. 29 (1844), G. L. N. Syn. Hep. p. 472 (1846). Vodonia Ralfsii, Dum. Hep. Eur. p. 16 (1874). Petalophyllum lamellatmn, Lindb. in Meddel. Soc. F. Fl. Fenn. 1, p. 91 (1874). Fossomhronia corbukfformis, Trabut, Atlas Fl. Alg. p. 7 (1886). Dioicous, csespitose, closely attached to the ground, small, light green in colour. Fronds procumbent, furcate, broadly obovate or reniform, flattish, beset on the antical side with lamellar processes disposed in radii towards the extremity of the frond, which is somewhat emarginate ; texture rather flaccid, cells largish to large, walls thin, with numerous chlorophyl granules attached to their sides, making them to appear some- what thick, midrib prominent, postically thick, often lengthened out at the base of the frond and denuded so as to resemble a stipe, covered beneath with numerous rootlets. Perianth antical, proceeding from the middle of the frond, tubular, wide at the mouth, which is dentate. Calyptra smaller. Pedicel thick and somewhat persistent. Capsule oblong-spherical, slightly apophy- sate, coriaceous, splitting to the base into two valves which are usually bidentate. Spores brown, reticulate, walls raised, giving them a bordered appearance not unlike F. aiigdosa, margin hyaline, areola? 5, 6 across, 20 to 25 cells round. Elaters brown, narrow, bi-tri-spiral. Male fronds broader, antheridia numerous, oval, enclosed in the laminse, more frequently near the midrib or enclosed in scale-like processes on the mid-rib. Dimensions. — Fronds about \ inch long x \ inch broad, stems 2* mm. broad x 1* mm. thick; cells of frond '06 mm.; perianth 4*5 mm. x 2*25 mm. ; capsule 2' mm. x 1"5 mm.; spores •07 mm. ; antheridia '275 mm. x -175 mm. Hab. — Damp sandy flats on the coast. Very rare. 1. Near Penzance, /. Balfs. Formerly on the roadside PETAL OPHYLL UM. ' ^ between Loggan's Mill and Treve, Pbillack, and between Hayle Causeway and St. Erth, W. Curium; Gwitbian, near Penzance, W. Curnow. 7. Aberfiraw, Anglesey, /. Ualfs, W. Wilso7i, W. H. P., 1900. 9. Southport, Lancasbire, Dr. Carrington , a J. Wild, W. IL P. 10. Coatbam Marsb, Yorks, IF. Iii(jhaw, 1901. I. Malabide, Br. D. Moore. Algiers, Trabut. Obs. — Mr. W. Curnow, in bis " Hepaticoe of West Cornwall," says : " One of tbe most interesting additions was a new species, PefalopJii/Ihou llalfsii, first discovered by Mr. Kalfs in an immature state on tbe sand-flats of Aberffraw, in Anglesey, and in tbe spring following I gatbered it witb perfect fruit at Hayle, growing in company witb Moerchia kibernica. I regret to say tbat tbis locality for it bas since been destroyed by drainage, &c. In consequence of tbis find an animated contest ensued between Dr. Taylor and Mr. Wilson and Mr. Ralfs, eacb of our opponents considering it a form of one or otber of tbe known species of Fossombronia. Mr. Ealfs, bowever, witb bis usual clearness of perception, and a praisewortby pertinacity in maintaining wbat be believed to be tbe trutb, ultimately won bis case, and estab- lislied tbe riglit of tbe plant to tbe rank of a new species, wbere- upon Mr. Wilson courteously appended to it tbe name of tbe discoverer.'^ It is a singular species, easily recognised by tbe lamellate fronds. Description of Plate CXC- — Fig. 1. Fertile Irond, nnt. size. 2. Male frond, sligbtly magnified. 3. Frond witb cross- section of periantb (after Wilson, Eng. Bot.). 4-(j. Cross-sections of stem X 11 (Soutbport, C. J. Wild). 7. Portion of frond x 290 (ditto). 8. Periantb x 11 (AberfFraw, Wilson). 9. Spore x 290 (Penzance, J. Ealfs). U». Antheridium x 24 (Soutbport, Wild). 432 EEFATICjE. Genus 40. PALLAVICINIA, Gr. 8f Benn. Pallavicinia, Gr. t Benn. Nat. Arr. Brit. PI. (1821). Dilmia, Dum. Comm. Bot. (1822). Bhjttia, Endl. (1840), G. L. N. Syn. Hep. (184G). Fronds prostrate or suberect, green, radiculose, strongly costate^ simple or with a few postical branches, rarely furcate at the apex. Inflorescence dioicous, antical. Bracts $, 2-3 series, variously divided, more or less connate. Perianth large, tubular, fleshy, mouth ciliolate. Calyptra included, equal or shorter than the perianth, irregularly torn at the apex, somewhat delicate, below the middle only somewhat fleshy and surrounded by the (about 20) sterile pistillidia. Capsule on a long pedicel, cylindrical, composed of one layer of cells, dividing to its base into 4 valves, often incompletely. Elaters slender, bispiral, deciduous. Spores minute. Androecia uniseriate, situated on the margin of the midrib, at the base of small incised or dentate peri gonial bracts. Antheridia solitary. 1. Pallavicinia Lyellii [Hook.), Gr. §' Benn, Jungermania Lyellii, Hook. Brit. Jung. t. 77 (1810). Palhvicinia Lyellii, Gr. & Benn. Nat. Arr. Brit. PL p. 775 (1821). Dilcena Lyellii, Dum. Comm. Bot. p. 114 (1822). Blyttia Lyellii, Endl. Gen. PI. (1840). Steetzia Lyellii, Lelim. PI. Preiss., TI. p. 129 (1846). Dioicous, in spreading patches, often stratified, medium size, pale to yellowish green in colour. Fronds horizontal, simple or with one or two lateral branches proceeding from the postical side of the frond, oblong, nearly linear or obovate-ligulate, strongly costate, costa plano-convex, postically rotundate, composed of several layers of cells, with a strand of narrow, elongated, darker coloured ligneous cells along the centre, sometimes two strands are observed, apex of frond emarginate, very rarely furcate, gradually or abruptly narrowing at the base, margin undulate, entire or subserrate; radiculose, rootlets few, proceeding chiefly PALLAVICIXFA. 433 from the costa, simple, pellucid; texture delicate, cells of the pagina large, oblong-hexagonal, walls thick, no trigones, lejito- dermous, pellucid. Flowers $ antical, often two, proceeding from the costa, about the middle of the frond. Bracts veiy short, connate, palmatiiid-laciniate. Pistillidia 12-20, long and narrow. Perianth erect, linear-cylindrical, fleshy, below 5 cells thick, mouth sub-constricted, lobate-ciliolate. Calyptra a little shorter than the perianth, above delicate, near to the base 2, 3 cells thick, apex irregularly ruptured. Pedicel elongate, delicate, white. Capsule sub-cylindrical, 5 times longer than broad, 4-valved, valves often cohering at the apex, delicate, walls composed oi" one layer of linear cells. Spores reddish-brown, roundish-angular, finely verruculose. Elaters very long and narrow, pale Iji-own. Male plant more delicate. Perigonial bracts antical, situated along the middle of the frond, imbricate, scale-like, laciniate- ciliate. Antheridia large, solitary, sub-globose, shortly slij)itate. Dimensions. — Fronds 1 to 2 inches long, 2* mm. wide, costa •85 mm. x '35 mm. thick ; cells "OS mm. x '05 mm., '00 mm. x •04 mm., •05 mm., '04 mm. ; bracts i V5 mm. high ; perianth 10^ mm. X 1'3 mm.; pistillidia "6 mm. long; pedicel 3*5 mm. long; capsule 4' mm. x '8 mm., 3* mm. x "75 mm. ; spores "04 mm. diam. ; elaters '5 mm. x "02 mm. ; antheridia 3 mm. x •25 mm, Hab. — Boggy spots amongst Spliafjna, and on wet dripping rocks. Very rare. 2. New Forest, Hampshire, C. LyeJI ; nr. Tunbridge, Sussex. Foster. 7. On wet rocks near Cader Idris, Merionethshire, ./. Ba/f^^ 11, 12. Foulshaw Moss, Westmorland, G. Slahlcr. 13. Lochar Moss, Dumfriesshire, James Cruickshank. I. South of Ireland, Bantry, Miss Hufchins ; Lough Biay, Dr. Tai/Jor ; Maghanabo Glen, near Fermoyle ; Castlegregory and by the lakes between Maghanabo Glen and Connor Hill, Co. Kerry, Dr. D. Moore. Found on the Continent and in North and South America and other parts of the world, Obs. — Distinguished when sterile from any British liondose 2 i: 434 HEPATIC^. species by the presence of a strand of strong, ligneous, narrow, elongated cells running along the centre of the costa; when fertile by the short, laciniate bracts, surrounding the erect linear- cylindrical perianth, which encloses a relatively large calyptra. Description of Plate CXCI.— Fig. 1 and 2. Plants natural size. 3. Portion of fertile plant x (Hooker). 4. Cross-section of frond x 24 (Herb. Austin). 5, 6. Ditto x 16 (ditto). 7. Portion of frond x 290 (ditto). 8. Female flower (bracts, perianth, and calyptra) x . 9. Section of female flower, showing bract, perianth, pistillidia, and germen x. 10. Bract x. 11. Capsule and portion of pedicel x . 12. Portion of male stem, showing perigonial bracts and ligneous strand x. 13. Perigonial bract and antheridium x. 14. Antheridium x (Hooker). 2. Pallavicinia hibernica (Hook.), Gr. 8f Benn. Jiongermania hibernica, Hook. Brit. Jung. t. 78, & suppL t. 4 (1816). Pcdlavicinia hibernica, Gr. & Benn. Nat. Arr. Brit. PI. p. 684 (1821). Dilaena hibernica, Dum. Comm. bot. p. 114 (1822). Biplolaena hibernica, Dum. Syll. Jung. p. 83, t. 2, £. 21 (1831). Diplolaena Lyellii, var. hibernica, Nees Nat. Eur. Leberm. Ill, p. 343 (1833). Diplomitrio7i hibernicum, Oorda in 8turm Deutschl. Fl. fasc. 22, p. 87, t. 23 (1835). Moerchia hibernica, Gottsche in G. &. R. Hep. Eur. Exsicc. n. 295 (1866). Delin. Hook. Brit. Jung. tab. 78 et Suppl. tab. 4 ; Ekart Syn. Jung. Germ. tab. X. fig. 84, et tab. xiii. fig. 116a ; Husn. Hep. Gall. pi. xi. fig. 124. Dioicous, loosely C2esj)itose, medium to largish in size, odorous, pale green colour. Fronds procumbent, loosely imbricate, once or twice dichotomously furcate, oblong, margin entire, undulate ; texture somewhat delicate, cells medium to rather large in size, quadrate or oblong-quadrate, walls thick, no trigones, midrib thick, broader than thick, 20 cells broad x 16 cells thick, gradually diminishing in thickness into the lamina, usually with no ligneous strands, lamina on each side of the midrib S to 10 cells wide composed of 2 or 3 layers of cells ; radiculose, rootlets hyaline, rarely reddish. Female flowers several, produced anti- cally from the mid-rib ; bracts two, surrounding base of the rALLAVICINIA. 435 perianth to about one-fourth its hei<,^ht, suborbicular, laciniate- (lentate. Perianth larij^e, oblong-ovate, slightly [)licate, unistratose near the middle, about 130 cells round, near base two or three cells thick, mouth a little constricted, irregularly and sparsely dentate. Calyptra half the size. Pistillidia long, about (>. Capsule dark reddish brown, oblong-ovate, dividing to the base into 2 or 3 bidentate or entire valves ; spores dark reddish brown, with numerous irregular ridges, giving them a minutely verruculose marginal appearance ; elaters very narrow, pale brown. Andrcccia produced in scale-like bracts antically from the mid-rib, perigonial bracts broadly oval, subconcave, apex irregularly dentate ; anthe- ridia large, oval spherical. Fruits about the middle of April. Dimensions. — Fronds J to 2 inches long, 3' to 4" mm. broad, mid-rib "G mm. broad x "4 mm. thick; cells "04 mm. x "05 mm., •03 mm. X "04 mm., -04 mm. x -04 ram.; bract 1' mm. high x •75 mm. broad; pistillidia "3 mm. x '075 mm. ; perianth 4* mm. x •2 mm. ; pedicel '175 mm. diam. ; valve of capsule 2" mm. x 1'25 mm. ; spores '035 mm. ; elaters "4 mm. x "Ol ; perigonial bracts '75 mm. x *6 mm., '05 mm. x *5 mm. ; antheridia "275 mm. x '2 mm. Hab. — On elevated marshy situations amongst /Sy^/zr/y/zr/. Ver}' rare. 15. Catlaw Hills, Kinnordy, Forfar, C. Li/cIl, 1822. I. Amongst Sphagnum cusjndatum and Jaiigermania emarf/ii/a/n on the shores of Lough Bray, a very elevated situation near Dul^lin, Dr. Tajjlor. J). McArdle, July, 1S78. ( )bs. — The description is that of the original Jfuif/. Jiibcniica published by Hooker in his "British Jung."; two other forras have been grouped with it which probably deserve to rank as distinct species. The type is distinguished from them by its much larger size, long, slender, slightly undulate fronds in addition to other characters. Distinp^uished from Pallavicinin Li/el/ii (Hooker) by the absence of a nerve in the midrib, by its strong smell in addition to other characters. Var. irH.suiiia/ia, Gottsche [Pa/lacicuiia M'lhoiiiana (Ci.), 436 HEPATIC^.. Pears. MS.). Jnut/. Jnbeniica, Wilson in Eng. Bot. 2, tab. 2750, excl. lower half of plate (1834). Delin. Eng. Bot. 2. tab. 2750, pp. Husn. Hep. Gall. pi. XT. fig. 124, b, c, d. Exsicc. G. etE. n. 163 ; C. and P. n. 143. Closely cajspitose, small. Fronds more upright, crisp, much folded ; mid-rib thicker, bracts more laciniate than in either of the two other forms. Dimensions. — Capsule 2*5 mm, x 1* mm., spores "05 mm., '04 mm., elaters '35 mm. x '01 mm., '2 mm. x "Ol mm. Hab. — In marshy sandy stations, usually on the coast. Rare. 1. Phillack; Hayle Sands, Cornwall, W. Curnow, 1842. 7. AberlFraw, Anglesey, IF. Wilson, fruit April 1829, September 1830. W. H. P., 1900. 9. Southport, Lane, Dr. Carrington, 1863, 1865; C. J. IFihl, 1882; W. H. P. Coatham Marshes, Yorks., Wm. Ijigliam, 1899. 15. Drumlie Airy, a waterfall on the Noran, Forfar, C. Lyell. Found on the Continent (Finland, Grermany, France, Switzer- land, and Italy) ; also in North America. Var. lejJtodesma (P allavicinia leptodesma, Tansley in Utf.). Fronds |- to 1 inch long, simple or furcate, margin irregularly lobed, mid-rib with two lateral rudimentary strands, perianth longer and narrower than in type or var. Wilsoniana. Spores light to dark brown. Elaters very light brown, very narrow, bi-spiral, 8-10 spiral turns. Hab. — In marshy, sandy situations on the coast. Very rare. I. Malahide, Br. B. Moore, D. McArdle. Dimensions. — Perianth 5*5 mm. x 1'5 mm., capsule 2*5 mm. x 1"1 mm., spores "05 mm., elaters '275 mm. x "01 mm., "2 mm. x '015 mm. Obs. — With reference to this variety or species Professor Tansley writes : " We have finished our investigation of the Pallavicmia-Si/m^jhi/of/yna-Hi/nieiiophi/fon group in reference to the strand of prosenchymatous cells. I was fortunate enough to get living- specimens of PaUavichna LyelUi from Professor Howe of ."Nev*' York as you suggested, and find that the strand is water- FALL A VICINIA . 437 conducting, though water passes through it much more slowly than is the case in mosses. In P. hibernica from Malahide, which Mr. McArdle sends me, we find two lateral strands of very thin walled but distinctly prosenchymatous cells which may be water conducting, but not having living material I cannot tell for certain. In larger plants from Lough Bray {Jun(/. hibernica, Hook. Brit. Jung, type) we can find no trace of such strands, neither can we in the specimens you sent from Coatham Marshes, Yorkshire, Wm. Ingram (var. Wihoniana). "... there is the occurrence in the Malahide plant of two distinct lateral veins in the mid-rib. The walls of the cells of these veins are distinctly lignified though very little if at all thickened, and in other respects, so far as can be seen from dried material, the strands are rudimentary compared with that of P. Lyellii and other species ot Pallavicijiia, Si/mpho(///jia, &c. I shall, of course, be glad to know if you find other characters in the Malahide plant distinguishing it from true P. hibernica, but even if not, the marked anatomical characters I find seem to me to justify its separation as a distinct species. In the absence of more conspicuous external characters I would suggest leptodesma as a specific name in allusion to the rudimentary character of the strands." For an account and figures of these strands see Tansley and Chick, " Notes on the Conducting Tissue System in Bryophyta," Annals of Botany, Vol. XV., March, 1901. Description of Plate CXCII. — Fig. 1. Plant natural size (Hooker). 2. Frond x ? (ditto). 3. Cross-section of midrib x 24 (Lough Bray, McArdle). 4. Portion of frond x 290 (var. Wiho7iiana), (Hayle, Curnow). 5. Portion of bract and perianth X ? (Hooker). 6. Portion of bract and pistillidia x 24 (Lough Bray, McArdle). 7. Capsule x ? (Hooker). 8. Portion of male frond X ? (Hooker). Description of Plate CXCIII. — Fig. 1. Plants natural size (var. leptodesnia), (Malahide, McArdle). 2, 3. Ditto (var. IFilsoniana), (Aberffraw, Wilson). 4. Plants slightly enlarged (ditto). 5. Fertile plant x 11 (var. leptodesma), (Malahide, McArdle). (5-8. Cross-sections of mid-rib x 24 (ditto). 9. Perianth x 11 (var. 438 HEPATIC^. Wihouiana), (Hayle, Curnow). 10. Pistillidium x 64 (Lough Bray, McArclle). 11. Spore x 290 (var. Wihoniana), (Hayle, Curnow). 12-14. Perigonial bracts x 24 (ditto), (Soiithport, Wild). 3. Pallavicinia Blyttii [Moerck), Lindh. Jangermania Bhjttil, Moerck Fl. Dan. x. 34, t. 2004 (1830). Gymnomitrion Blyttii, Hiiben. Hep. Germ. p. 44 (1834). Gordea Blyttii, Corda in Sturm Deutschl. Fl. (1835). J)i2domitrium Blyttii, Coixla in Sturm Fl. Germ. 11, p. 126 (1835). JHjjlolaena Blyttii, Nees Hep. Eur. Ill, p. 339 (1838). Thedenia Blyttii, Hartm. Skand. Fl. (1820-43). Blyttii Moeo'ckii, G.L.N. Syn. Hep. p. 474 (1844). Moerckia norvegica, Gottsch. in G. &, R. Hep. Eur. n. 295 (1863). Moerckia Blyttii, Gottsch. Ann. sc. nat. p. 83 (1864). Dilaena Blyttii, Dum. Hep. Eur. p. 138 (1874). Dilaena Blyttii, Dum. Hep. Eur. p. 138 (1874). Pallavicinia Blyttii, Lindb. Muse. Scand. p. 10 (1879) ; Kaalaas Leverm. Norge, p. 451 (1893). Galycularia BhjttH (Moerck) St. Sp. Hep. p. 360 (1900). Dioicous, loosely or closely ca.'spitose, medium size, pale green colour. Fronds oblong, simple or furcate, prostrate, some- what thick; lobes sub-membranaceous ; costate, mid-rib ver}^ prominent on the postical side ; a cross-section of frond shows 20-30 layers of thin (those below purple-walled) cells, gradually becoming thinner near the margin where it is one layer thick ; epidermic cells smaller than the inner, quadrate, large, walls firm, no trigones ; margin undulate, sinuate, crispate ; radiculose, rootlets numerous, close, reddish-brown. Female flowers antical ; bracts surrounding the base of the perianth, undulate, plicate, unequally laciniate. Perianth projecting very much beyond the bracts, oval-campanulate, upper portion plicate, slightly laterally compressed, composed of several layers of cells, mouth unequally laciniate-dentate. Pistillidia 5-8. Calyptra thick. Capsule oval, valves firm, bi-tri-stratose, lanceolate, pedicel very long. Spores dark brown, verruculose. Elaters very narrow, bi-spiral, paler than the spores. Male flowers antical, bracts imbricate, biseriate. PALLAVICINIA. 439 ])licate, obovate, simple or bilobed; antheridia largo, single, roundish, style short and thick, composed of two rows of cells. Paraphyses few. Dimensions. — Fronds 2 c. mm. long x 8* mm. broad ; bracts 1' mm. high x 2" mm. broad ; perianth 5' mm. long; pistillidia ■5 mm. X '075 mm.; calyptra 3' mm. x 2* mm.; valves of capsule 3- mm. x r mm. ; spores '035 mm. ; elaters '275 mm. x "01 mm. ; male frond 3' mm. broad x "7 mm. thick near the middle, maU- bracts "7 mm. x '05 mm., '7 mm. x -4 mm. ; antheridia '275 mm. x •25 mm. Hab. — On small bare vertical peatv banks on grassy slopes, in alpine regions. Extremely rare. Scotland, Jo/ut Sadler. 15. Locli-na-gar, Georf/c Stabler, July 1884. Ben Wyvis, George E. Davies, July 1888. Ben Lawers, in several localities, 3100 to 3400 ft., /S'. M. Macvicar, June 1901. Found on the Continent (Germany, Norway). Obs. — This is an extremely rare British species, being only found in a few alpine situations. From Pellia it is at once distinguished by its oval capsule, no other frondose species with which it is likely to be mistaken for, being found at such altitudes where it is met with. Description oe Plate CXCIV. — Fig. 1. Fertile plant natural size. 2. Ditto x ? 3. Portion of male frond x ? 4. Bract, explanate x ? 5. Young perianth x ? 6. Perianth x ? 7. Mouth of perianth, explanate x ? 8. Calyptra x ? ("Flora Danica"). Description oe Plate CXCV. — Fig. 1. Cross-section of male stem x 24 (Norway, Herb. Schimper). 2. Portion of frond X 290 (Ben Lawers, S. M. Macvicar). 3. Bracts x 24 (Loch-na-gar, G. Stabler). 4. Mouth of perianth, explanate x 16 (Norway, Herb. Schimper). 5-8. Male bracts x 24 (ditto). ed, 440 HEPATIC^. Genus 41. BLASIA, iMich. Blasia, Mich. Nov. PL Gen. p. 14 (1729); Linn. Fl. Suec. ed. l,p. 9:^3 p. 105 (1715), et Sp. PI. p. 1605 (1753). Janyermania, Hook. Brit. Jung. tt. 82-84 (181G). Plant frondose, nerved, rructifi cation produced in a rupture of the nerve. Involucre bladder-shaped, adnate to the apex of the frond. Perianth enclosed within the utricular involucre. Calyptra obovate. Peduncle emerging from the apex. Capsule oval-globose, 4-valved, coriaceous. Elaters bi-spiral, deciduous. Antheridia immersed in the thallus, covered with dentate scales. Gemma' globose, arising by a slender ascending tube from their large Hask-like receptacles which are immersed in the thallus. Blasia pusilla, Z. Blasia pusilla, Linn. Sp. pi. 1605 (1753). Jungermania Blasia, Hook. Brit. Jung. t. 82 & 84 (1816). Dioicous, in flat imbricating patches, medium size,pale yellowish or bluish-green in colour. Fronds prostrate, horizontal, costate, simple or with a single lateral branch, dichotomously divided, palmate with apices forked or rarely stellate ; lobes usually oblong or linear-obovate, apices always broader than the base, often erect, margin pinnatifid or shortly lobate, sinuous, undulate, incurved ; throuo-liout the centre of the frond and its divisions runs an evident and broad nerve, most prominent on the postical side ; texture fleshy, thinnest at the margin where it is membranaceous ; at the middle 8-14 cells thick, gradually becoming thinner to the margins where it is one cell thick ; cells somewhat indistinct, at the middle of the frond striate, narrow, elongate, near the margin medium size, oblong-quadrate or quadrate, marginal cells smaller, quadrate, very finely crenulate, walls thick and firm ; radiculose, rootlets numerous, proceeding from the postical side of the frond but (diiefly along the nerve, long, slender, hyaline. On the postical side of the frond are scattered small, hyaline, oval scales BLASIA. 441 resembling stipules vvliicli are unequcilly dentate. Involucre antical, oblong-lanceolate, inflated, semi-transparent, vanishing- early, imbedded in an oval cavity in the mid-rib of the fronds. Calyptra obovate, free. Capsule oval-globose, on a long pedicel, 4-valved, valves thick, dull reddish brown in colour, inner layer with imperfect spiral threads. Spores sub- spherical, greyish- brown, slightly granulate ; elaters very narrow, about \ the diameter of the spores, hyaline, bi-spiral. Male plant smaller, narrower and rarer ; androecia imbedded in the antical side of the frond ; antheridia elliptical, yellowish in colour, shortly stipitate. (xemmiparous ; gemma3 numerous, globose, issuing by a slender ascending tube from their flask-like receptacles, which are immersed in the frond. Fruits March and April, quickly dis- appearing. Dimensions. — Fronds ^ to 1 inch long, 4'-G' mm. wide; cells •03 mm. X '04 mm. ; spores "05 mm. diam. ; elaters '2 mm. long X '01 mm. broad. Hab. — On wet clayey banks in woods, or on moist heaths. Moderately rare. 1. Trembath, Chyanhal, Trungle Moor, Cornwall, if. Curnoic ; Madron Valley, near Carfury, and near Gurnard's Head, Cornwall, J'J. D. Marqaand. 3. Road leading from Langton Green to Speld- hurst Church, Fon^fcr, " Fl. Tunbr." 5. Banks of Severn, Seckley ; Arley Wood ; Dimmings Dale, Staffordshire, IF. Bagnall. 7. Cvvin Bychan, Merionethshire, Ttev. T. Salway. 8, 9. Marple, Cheshire, G. A, Holt; Ashley, Cheshire, W. H. P. ; Daisy Nook, Lane, G. A. Holt, Sailor's Shore, Lane, G. A. Holt, Holden Clough, Lane, G. A. Holt. 10. Near Claphara, i)r. Carrington ; Clay pool-side, near Sherburn, Br. F. A. Lees; Halifax, ./. Bolton ; Todmorden, John No well ; Pennant Clough, Hebden Bridge, A. Stansjielcl. 12. Low Borrow Bridge ; Foulshaw ; by river Lune, near Ingmire Hall, Westmorland, G. Stabler. 13. Damp road- sides, New Galloway, J. McAnclreir ; near Carronbridge, Charles Scott; Cluden Mill, Kirkcudbrightshire, /<7//^<:^s 6y7//r/-.s//r/y//-. 15. Den of Garrol, Aberdeenshire, /. Sim. IG. Moidart, West Inverness, S. M. Macvicar. 442 HEPATIC^. I. Castle Kelly Grlen, Dublin, Dr. TaijJor ; near Wooden Bridge, Wicklow, Br. D. Moore ; at the base of Brandon Moun- tain, Kerry, Dr. D. Moore. Found on the Continent, Northern Asia, and in North America. Obs. — A very distinct species, the only one of the genus and easily recognised, even when barren, by the remarkable flask-like receptacles for the gemmae and the beautiful hyaline serrated bracts on the postical side of the fronds. Description of Plate CXCVI.— Figs. 1 & 2. Plants natural isize. 3. Young frond x . 4. Portion of frond with young receptacles for the gemmae, the mouth not being yet opened- 5. Fertile plant x . 6. Portion of frond, postical view showing scales. 7. Cross-section of female frond x . 8. Capsule opened x. 9. Antheridium x. 10. Gemmae x (Hooker). G-enus 42. PELLIA, Raddi. Jungermcmia, Linn. Fl. Suec. 1 eel., p. 399 (174.")), et. Sp. PI. 1 ed. 2, 1135, n. 23 (17.33); Hook. Brit. Jung. t. 17 (ISlC). Fellia, Eaddi in Att. Soc. Modena, 1. Ci'.t (isilT). Pellia ejnphylla, Lindb. Hep. in Hib. p. 534 (1874) ; Limpr. in Colin Kr\ [)t. Scliles, 1, p. 328 (187G). Marsilia epijjhijlla, Lindb. Muse. 8cand. p. 10 (187!)). Monoicous, growing in spreading, flat patches, largish, deep green in colour. Fronds imbricating, and matted together by means of their rootlets, simple or irregularly once divided, segments oblong or broadly cuneate, margin entire near the base, near the apex irregularly incised, sinuate or sublobate, broadly nerved, near the middle 14-16 cells thick, gradually becoming thinner towards the margins; texture submembranaceous, cells largish to large, 4-, 5-, and 6-sided, elongate, w^alls thin, no trigones ; radiculose, rootlets brownish white, numerous, close, short. Involucre proceeding from the antical side of the frond, near the apex or more rarely the middle, subcylindrical, plicate, mouth slightly dilated, irregularly inciso-dentate. Calyptra ovate, when older oblong, tuberculate, exserted, carnose, composed of several layers of cells. Pedicel long, hyaline, 18 cells thick. Capsule spherical, pale greyish-brown, dividing into 4 equal, ovate valves, which soon recurve, outer layer composed of hexagonal cells, whose corners are thickened, inner layer with elongated cells containing numerous semi-spiral rings. Elater-bearers for some time persistent at the base of the capsule, 20 to 40, in a dense entangled mass, brownish colour, irregular in length, at the free end conical, obtuse, usually with 1 or 2 brown spiral threads, sometimes 3 or 4, rarely 5 or 6, at their free end, they are more or less uncinate, holding some of the true elaters and spores for a longer or shorter time back. Elatei's very long, slender, twisted, usually bi-spiral. Spores oval, greenish-yellow, finely muriculate. Male flowers situated on the antical surface of the frond and always confined to the nerve, in which the antheridia are imbedded irregularly in two rows. Fruits in Spring. 414 HEPATIC^. Dimensions. — Fronds ^ to 2 inches long, J to |- inch broad, cells "09 mm. x '045 mm., "09 mm. x '055 mm., 'll mm. x "05 mm., '09 mm. x '05 mm. ; pedicel 1 to 2 inches long ; calyptra 3" mm. high X r mm. broad; valves of capsule 175 mm. x 12 mm.; elater-bearers 1* mm long x '025 mm. broad; elaters '5 mm. long; spores '125 mm. x *1 ram. Hab. — Growing in large patches on banks and rocks in damp or wet shady situations, in ditches or springy places. Common. 1 to 18. I. Europe, North America. Obs. — A common species, which fruits plentifully in Spring. It is distinguished when fertile from other frondose species by its long delicate pedicel, which supports a rather large capsule, containing an extraordinary number of spores and elaters; Herr Jack has counted one containing no fewer than 4500 of the former and 5000 of the latter. When sterile, the large, submembranaceous, dark green, broadly nerved frond will distinguish it. For the characters separating it from Pellia caJycina (Tayl.), and Pellia Neesiana (G.), see notes under those species. Decriptions of Plate CXCVII. — Fig. 1. Fertile plants, natural size (Ekart, after Hooker). 2. Young fertile plant, magnified (ditto). 3. Cross-section of the same, more higldy magnified (ditto). 4. Portion of frond x 290(Llanfaelog, W. H. P.). 5. Capsule magnified (Ekart, after Hooker). 6. Opened capsule, showing elater-bearers x 10 (Jack). 7. Elaters and spores x 75 (ditto). 8. Elater-bearers x 7b (ditto). 9. Elater x 316 (ditto). PEL LI A. 445 2. Pellia Neesiana [Got/.sc/i.), Limpr. PelUa epijyhylla, B forma yeesiana, Gottsche in Hedwigia, p. (5i» (]H(i7). Pellia Xeesiana, Limpr. in Cohn Krypt. Fl. Schles. 1, p. 329 (1876) ; Jack, Flora, p. 81 (1895); Howe Hep. and Antho. of California, p. 7(5 (1890); Macvicar, Journ. of Bot. July (1900); Stephani Sp. Hep. p. :'.(;(; (1900). Marsilia N'eesii, Lindb. Muse. Scand. p. 10 (187'9). Pellia N'eesii, Limpr. Kaalaas, Leverm. Norge, p. l.")!', (189;)). Dioicous, growing in flat creeping layers, especially the fertile plants, medium to largish in size, not so robust as Pellia epijjlnjlla, dark green to reddish brown, older plants dark brown in colour. Fronds elongate, cuneate, somewhat sparingly dichotomously branched with oblong or linear segments, margin plane or frequently undulate, middle of frond 10-12 cells thick ; radiculose, rootlets numerous, brownish- white. Involucre antical, smooth, perfect, viz., forming a complete ring, cylindrical, longitudinally plicate, mouth truncate, entire or crenulate. Calyptra usually more or less exserted, sometimes included, thick, composed of 5 layers of cells at the base, at apex bistratose. Pedicel hyaline, delicate. Capsule spherical, pale brown, dividing into 4 equal, ovate valves, outer layer of cells nodulose, thickened at the angles, inner layer with numerous semi-spiral rings. Elaters and spores similar to those of Pellia epiphjlla. Male plants simple or bifid, never repeatedly innovantly furcate ; antheridia numerous, imbedded in the frond. Fruits in Spring and early Summer. Dimensions. — Fronds \ to 1|- in. long by 7 cm. to 1* cm. broad; involucre 1'2 mm. high; calyptra 1'3 mm. high; pedicel 3'5 cm. long ; capsule 1'5 mm. diameter. Hab. — In wet grassy ground, Moidart, West Inverness, S. M. Macvicar, 1900. Commoner than P. epijjlii/lla, above 3000 ft. on Ben Lawers, Perthshire ; fruit 3200 ft. S. M. Macvicar, 1901. Middle and North of Europe. North America. Obs. — Through the acuteness of Symers M. Macvicar, Esq., I am enabled to add with certainty this distinct species to my list. 446 HEPATIC^. In the "Journal of Botany," July 1900, lio published it as new to Britain, and gives some interesting notes how to dis- tinguisli it from the two other species of the genus, from which I take the following : " The fronds bear considerable resemblance to those of P. calycina, the species being un distinguishable with certainty when barren. The younger fronds are reddish-brown to dai k green, frequently wavy at the edges ; the older ones are dark brown, narrower, and flat. It also resembles calycina in bein^- dioicous, but the male plant is without the hammer-shaped furcate innovations of the latter. The most distinctive character between the two species lies in the structure of the interior layer of the capsule wall, caJyciyia being entirely without rings, while the present plant agrees with epijjliyJJa in being furnished with them. The involucre forms a complete ring, as in calycina, the cylinder varying in length, but never so long as is frequently seen in the latter. The calyptra is more or less exserted, sometimes only slightly beyond the involucre, but more commonly for some distance, and frequently as much as in e/jiphylla. It is usually included in calycina. " Pellia epijiliyUa differs in being monoicous, the antheridia being on the same frond as the female, and it has commonly much broader fronds, which are generally green. Tlie darker coloured narrower forms cannot be distinguished with certainty in the barren state from Neesiana. Besides the difference in the inflorescence. P. ejiiphylla also differs from the two other species in that its involucre does not form a complete ring, the antical portion being absent; and the mouth of the involucre looks towards the apex of the frond, while in the others it is vertical. The calyptra is always highly exserted. The interior layer of the capsule wall has more numerous rings than in Ncesianay PELL [A. U7 3. Pellia calycina {Taijl) Jangermania calycina, Tayl. in Mackay Fl. Hib. 11, 55 (l.s;)(i). Jungermania endivioifolia, Dicks. PI. crypt. Brit. fasc. iv. p. ID (l.SOl ) ? Jimgermania epiphylla, vax. farciyera, Hook. Brit. Jung. t. 47, f. 18 (1ru.sii>id<;s, Nees, Nat. Eur. Leb. I 1 1 , p. 4.'i<» (ls;;h), G. L. N. 8vii. Hep. p. r.i7 (!«»»). Ju'ccardia lauliijida, var. ainbrosioides (Nees), Lindb. Acta Soc. Sci. Fenn. M», r.ii(is7-.). Monoicous, loosely ca'spitose or croeping amongst mosses, small, dark green, turning dark brown, when young light green in colour. Stems prostrate, narrow, subjiinnate ; pinna- ascend- ing, irregular in length, sometimes attenuate ; cross-section bi-convex or piano-con ve.x ; margin minutely crenulate, composed of a single row of cells; texture rigid; exterior cells small, (juadrate, much smaller than the inner, 40 to GO round, walls dark, firm ; inner cells medium to rather large in size, 4-G-sided ; 4 to G cells thick near the middle. Female flowers lateral on the main stem ; bracts small, laciniate. Calyptra relatively very large, cylindrical, smooth. Androecia lateral, long, linear, in which are imbedded about G roundish-oval antheridia. I have never met with perfect fruit. Dimensions. — Fronds 1 inch long x 5' to 10" mm. broad, stem 75 mm. broad x "2 mm. thick ; cells interior 03 mm. X 05 mm., exterior 02 mm. ; male catkin 1 mm. long x 4 mm. broad. H.\H. — Amongst wet mosses or on shaded wet rocks. Some- what rare. 1. Mousehole Cave, Penzance, Cornwall, Miffen iS* dirnoic. 7. 'ryn-y-Groes, Merionethshire, //'. //. /^ . Ogwen Valley, Dr.Carrington ^- IW II. P. \). Hayfield, Derbyshire, G. A. UuU. 12. Grisedale, Burnt s ty Stabler ; Broad Gate Bog, Ings, Westmorland, G. Stabler. 16. Common on wet rockv i)anks. especially in ravines, Moidart, West Inverness. S. M. Marv'unr. \. Ventry ; Connor's Hill; O'Sullivan's Cascade, ifec. Dr. Carrinyhm and other?. Found on the C«»ntinent. Obs. — This has always been considered by previous writers as a variety of A. inuUifuln, but its usually tlark brown colour, rigid 454 HEPATIC jE. habit, narrow outline of frond, with ascending branches, margin of stem and branches usually minutely crenulate, are characters which I consider make it quite worthy of specific rank, and by them it can be easily distinguished from other British species. Description of Plate CCI. — Figs. 1-3. Plants natural size. 4. Frond x 11 (C. & P. n. 63). 5, 6. Cross-section of stem x 24 (Ogwen Valley, W. H. P.). 7, 8. Ditto x 24 (near Penzance, Mitten & Curnow). 9. Male catkin x 11 (C. & P. n. G3). 4. Aneura latifrons Lindb. Jungermania multifida, Schmid. Icon. pi. 11], pp. 213-21(5, excl. synon. et pp. (1797), Hook. Brit. Jung. p. 11), n. 75, pp. (1816). Riccardia multijida, Gr. & Benn. Nat. Arr. Br. PL p. G84, n. 1, pp. (1821). Aneura jxdmata, a major, Nees, Nat. Eur. Leb. Ill, p. 459 (1838), G. L. N. Syn. Hep. p. 498 (1840). Aneura latifrons, Lindb. Soc. F. Fl. Fenn. (1873). Riccardia latifrons, Lindb. Hep. Hibern. p. 513 (1875). Monoicous, broadly and densely csespitose, depressed, small, pale, bright or brownish-green in colour. Fronds suberect, irregularly subpinnate or sub-bipinnate, transverse section plano- convex, almost the whole breadth 4 or 5 ceils thick, the inner ones large, hyaline, exterior smaller and chlorophyllose, one cell thick at the entire margin; branches thin, plane, more or less fastigiate, broadly or oblong-cuneate or sublinear, transverse section linear, 3 cells thick near the middle ; radiculose, rootlets few or numerous, hyaline ; stoloniferous, stolons proceeding from the main stem and branches, radiculose, subterete and long, sometimes bearing inflorescence ; gemmiparous, gemmae roundish, composed of one or two cells at apex of the branches; texture opaque when dry, somewhat pellucid when living ; cells of the antical layer irregular, oblong-rhomboid, slightly or indistinctly thickened. Inflorescence lateral on the stem or branches, very often the $ and $ together, rarely $ alone. Perichsetial bracts few but large, broadly oval, irregularly lobate, fimbriate. Pistillidia 3-10, short, conical. Calyptra large, ascending, pyriform, clavate ANEURA. 455 or cylindrical, truncate, white, slightly verrucose, near the base 7 or 8 cells thick, above 3 or 4. Pedicel hyaline, somewhat thick. Capsule oval or elliptical, brown, dividing to the bast- into 4 valves, which are elliptical, when dry convolute, reflexed, composed of two layers of cells, inner small. Elaters short, some- what obtuse, unispiral, spiral threads brown. Spores globose, greenish-brown. Androecia narrowly oblong, base narrow and shortly stipitate, base and postical side slightly radiculose; anthe- ridia biseriate, globose, hyaline, very shortly stipitate. Fruits in Spring. Dimensions. — Fronds \ to h inch long, branches 4- mm. io 8* mm. long, stem 4* mm. broad x 2* mm. thick, branches of stem 4* mm. broad x 1* mm. thick; calyptra 2-3' mm. long x •75-1* mm. thick ; pedicel 4--7* mm. long; capsule '0 mm. x '4 mm.; valve '9 mm. x "3 mm.; spores •015-'0175 mm. ; elaters •225 mm. x •0175 mm. Hab. — On rotting trees, damp turf or in wet places. Rare. 9. Cotteril Ciough, Cheshire, IF. Wihou. 10. Goathland, M.B. Slater. 16. Moidart, West Inverness, ascending to 1400 it., S. M. Macvicar. I. Killarney, Dr. Carr'uifjton and others. Found on the Continent and in North America. Obs. — A species previously confounded either with A. niuUijida or A. jjahiKita until the late Prof. Lindberg recognised it as distinct from both. It is abundantly distinct from what was considered a variety of A. muJtijida [A. avibro.sioiclcs) and from A. palmed a ; from typical A. mult if da it is distinguished by its relatively thicker and rounder stems, with more irregular and broader branches, and the more tuberculate calyptra. Description of Plate CCII. — Fig. 1. Plant natural size (Hooker). 2. Frond magnified (ditto). 3. Portion of frond magnified (ditto). 4. Frond x 24 (Cr. k P. 493). 5. Cross- section of stem X 85 (ditto). 6. Cross-section of branch x 85 (ditto). 7. Calyptra x IG (Nr. Goathland, M. B. Slater). 8. Capsule x 24 (ditto). 456 HEPATIC^. 5. Aneura sinuata (Dids.), Bum. p.p. Lichenastrum chmncedryos mioltifidcp. divisura, Dill. Hist. Muse. pp. 511, t. 74, f. 14b (notf. 44a) (1741). Jungermania sinuata, Dicks. PI. crypt. Brit. fasc. '2, p. 16 (1790). Jungermania multifida, var. sinuata, Hook. Brit. Jung. t. 45, f. 2 (181G). Aneura sinuata, Dum. Comm. Bot. p. 115 (1822). Aneura pinnatifida, N. Carr. Irish Crypt, p. (1863), and of many others. Eiccardia multifida, var. major, Lindb. Hep. Hibern. (1875) ? Riccardia latifrons (Lindb.), var. sinuata (Dicks.), Lindb. Muse. Scand. p. 5 (1879) ; Kaalaas Leverm. Norge, p. 213 (1893). Monoicous, growing in flat spreading patches, medium size, pale green in colour. Fronds prostrate, imbricating, flat, irregu- larly divided, branches irregularly pinnate or bipinnate ; thin, the whole breadth almost equal in thickness ; cross-section linear ; cells oblong-rhomboid. Female flowers lateral; bracts very short. Calyptra oblong, clavate, smooth. Capsule oblong, dark brown ; spores pale brown, same size as the thickest portion of the elaters ; elaters unispiral, reddish-brown. Androecia lateral, situated near the female flower, oblong-oval ; antheridia oval, 8-10 in two rows, imbedded in the male catkin. Fruits in Spring. Dimensions. — Stems 1 to Ij inch long, 2* mm. broad, •2 mm. thick, branches '5 mm.-l'5 mm. broad; calyptra 3" mm. X "8 mm.; capsule 1" mm. x "6 mm.; spores "02 mm.; elaters •4 mm. X -02 mm. ; andrcecia "8 mm. x '4 mm. Hab.— On wet dripping perpendicular rocks in shady situa- tions. Rare. 7. Arthog, Dolgell}^ Tyn-y-Groes, Merionethshire, Dr. CarrlngioR §' JF. H. F. 12. Borrowdale, Cumberland, W.H.P. Isle of man, G. A. Holt. I. Killarney, Cromaglown, Tore Cascade, Dr. Carrington and others. Obs. — Distinguished from Aneura midtifida (L.) and Aneura latifrons Lindb. by its larger size, thinner fronds and smooth calyptra. It is a somewhat rare species, and, so far as I know, is only ANEUBA. 457 found on wet dripping perpendicular rocks, frequently growing with Juhula Hutclihu^icB. Dr. Scliiffner states that Aueura ^jiwiafifda, Nees, is quite a different species. Description of Plate CCIII. — Fig. 1. Plants nat. size. 2. Portion of frond magnified. (Isle oF Man, Holt). 3, 4. Cross- section of frond x IG (ditto). 5. Male catkin and young female flower X 24 (Artliog, W. H. P.). 6. Calyptra x IG (ditto). 7. Capsule x IG (ditto). G. Aneura pinguis (X.), Difiii. Marsilea media pingiois, pallide virens, Jlorihus nigricantibus. ad foliorum latera egredientihus, Mich. Nov. pi. gen. p. 5, tab. 4, f. 2 (1720). Lichenastrum ccqntidis oblong is ; juxta foliorum divisuras enascentibus, Dill. Hi.st. Muse. p. 509, tab. 74, fig. 42 (1741). Jungermania pinguis, Linn. Sp. pi. p. lloG (1753); Hook. JBrit. Jung. t. 4(! (181G). Aneura pinguis, Dum. Oomm. Bot, p. 115 (1822). Riccardius pinguis, Gray k Benn. Nat. Arr. Brit. PI. 1, p. 084 (1821). Dioicous, csespitose or creeping among mosses, of medium size, dark green in colour, turning pale yellowish-green when dry. Fronds decumbent, often imbricate, simple or lobate, oblong- sinuate, plane or concave, apex rounded and obtuse, margii: somewhat undulate ; texture succulent, fleshy, opaque, rigid when dry ; 6 to 8 cells thick near the middle, 1 to 2 near the margin, without nerve ; cells 4-, 5-sided, interior ones hyaline, walls firm, no trigones ; radiculose, rootlets few, minute, short, close, yellowish-white, some fronds rootless. Female flowers proceeding Irom the postical side near the margin ; bracts short, subhemis- pherical, fimbriate. Calyptra linear-oblong, cylindrical, smooth, nearly the same thickness throughout, straight or slightly waved and curved, mouth a little uneven ; pistillidia 7 or 8, small, ovate. Capsule oblong-oval, reddish-brown, striate or furrowed, 4-valvecl, inner layer of cells with 20 to 30 rows of spiral threads, base of capsule (apex of pedicel) composed of 8 x 8 large brown cells ; spores spherical, brown, muriculate, twice the diameter of the i58 HEPATIC^. broadest part of the elaters ; elaters pale brown, bispiral, attenu- ated at both ends. Male plant usually shorter and crisped at the margin, 4 to 16, roundish-oval, slightly pedicellate, antheridia imbedded in the antical side of the roundish, simple or bilobed processes. Fruits in Spring. Var. denfici'Iafa, N. Fronds smaller than the usual form, simple, linear, margin denticulate. Dimensions, — Fronds |- to Ih inch long, by 5" mm. to 10' mm. broad, "4 mm. to '5 mm. thick near the middle ; cells •05 mm. X "05 mm., "05 mm. x "06 mm. ; calyptra 5" mm. to 7' mm. long x 1*5 mm. broad ; pedicel 1 inch long; valves 1*2 mm. X '4 mm. ; spores "03 mm. ; elaters '25 x '015 mm. near the middle. Hab. — In damp places, ditches, bogs, or on wet rocks. Common. 1-3. 5,G. 7-12. 14-16. I. Europe, North America, Cuba. Yar. denticulata, N. Creeping amongst mosses, frequent in limestone districts. Obs. — A distinct species, and not likely to be mistaken for any of the other frondose ones ; distinguished by its thick, nerveless, tiesh}^ fronds, which are brittle when wet and have a greasy feel, like the leaves of Pinguicula vulgaris ; its usually larger size and smooth calyptra separate it from the other British species of the genus. Descriptiox of Plate CCIV. — Figs. 1-3. Plants, natural size (after Hooker). 4. Frond, postical aspect with involucres slightly magnified (ditto). 5. Cross-section of frond x 16 (Closter, Austin). 6. Cross-section of involucre magnified (after Hooker). 7. Cross-section of young calyptra, magnified (ditto). 8. Cal3^ptra, magnified (ditto). 9. Capsule, magnified (ditto). 10. Capsule opened, magnified (ditto). 11. Portion of male frond, magnified (ditto). 12. Male receptacle, magnified (ditto). METZGERIA. 459 Genus 44. METZGERIA H(i(/di. Jungermania, L. Sp. pi. ed. 1, p. o24 (17;57) ; Hook. I'lit. Jiu lt. t. !'.'>, .")() iind hb (181(0. Metzgeria, Raddi in Act. See. Sc. Modena, 18, p. ;M (1«18). Plants frondose, soft, pale green or yellow, rarely dark green, prostrate, stratified or creeping laxly amongst mosses. Fronds linear, diehotomously branclied, very rarely pinnate ; floriferous, branches postical, pilose or ciliate, especially on the costa on the postical side, sometimes on the whole of the postical surface, rarely on the antical. Pili or cilia solitary or 2-5. Costa very distinct, rarely terete, slightly compressed, cross-section oval, 4-11 cells in diameter, cortical cells larger and pellucid, inner narrow, subopaque. Pagina, viz. wings on both sides of tlie costa, 10-25 cells broad, composed of one layer, texture lax, pellucid, often decurved or undulate. Inflorescence dioicous, very rarely monoicous. Involucre ? (bracts) on short postical branches, emarginate or obcordate, sometimes reduced to a mere annulus around the base of the calyptra. Perianth absent. Pistillidia i'ew, 2-5-pairs, short, the sterile surrounding the base of the calyptra. Calyptra exserted, clavate or pyriform, fleshy, the whole or only the upper half pilose or villose. Pedicel short. Capsule oval, dividing to the base into 4 valves, composed of two layers, inner without annular or semi-annular fibres. Elater> long, unispiral, deciduous. Elater-bearers short, claviform, per- sistent at the apex of the valves. Spores minute, almost smooth. Involucre $ similar to the ? , globose, convolute ; antheridia about 4-pairs, globose, very shortly stipitate. Gemma? aggregated on the attenuate apices of the linear fronds, composed of one layer, margin fimbriated. 460 HEPATIC^. 1. Metzgeria pubescens [Schrcutk). Jioigermania pubescens, Schrank, Prim. fl. Germ. 1, p. 231, n. 860, excl. synon. (1792), Hook. Brit. Jung. p. 20, n. 79 (1816). Metzgeria pubescens, Radd. in Act. Soc. Modena, 18, p. 40 (1818). Dioicous, in flat patches, medium size, pale glaucous green in colour. Fronds closely imbricate, horizontal, alternately pinnate or sub-decompound, branches short, linear, of uniform width, flat, apices obtuse ; margins undulate, uniformly and densely villose on the antical and postical sides ; the hairs on the postical side longer, single, or many at the margin double or in threes, irregularly curved, without sucker-like branches at the apex ; midrib projecting on the antical and postical side, 8-12 cells tliick X 10-14 cells broad, showing scarcely any cortical cells ; texture delicate, cells smallish, 5-6 angled. Perichsetial bracts situated on the postical side of midrib near margin, densely villose, pistillidia 2-10. Androecia numerous, on the postical side of midrib ; antheridia 5-9, subglobose, shghtly stipitate. Only imperfect female flowers have yet been met with. Dimensions. — Fronds 1-1 j- inch long, '2. mm. broad ; cells •0225--0285 mm. Hab. — On rocks and stones, or creeping amongst mosses, rarely on trees, in limestone districts. Moderately common. 5. Eocks about Thor's Cave, Robert Gamer; Dove Dale, Staffordshire, J. E. Bagnall, W. H. P. 7, 8, 9. Easegill, near Leek ; Over Kellet, A. Wilson ; Silverdale, Lane, /. A. Wheldon- 10, 11, 12. On an old tree-stump, Barrow Field, near Kendal; Deep Grhyll, Mallerstang ; on shady rocks near the bridge, Kirkby Stephen ; on Stainmoor, near Barras ; in a shady ghyll at the village of Dent, Westmorland, G, Stabler. 13. Penton Linns, Charles Scott. 15. Ben Lawers, ascending to top, S. M. Macvicar. 16. Lismore, Argyll, S. M. Macvicar. I. Mountains near Belfast, Mr. Tenq^letoii ; on limestone rocks between Larne and Glenarm, and at Sillaghbraes, near Larne, Br. D. Moore, S. A. Stewart. METZaEllIA. 461 Found on the Continent, North Anioricu ; Simla, Himalayas. (Jbs. — The fronds, being densel}^ villose on the antical surface, distinguished the plant at once from the other species of the ^•('11 us. Description of Plate CCV. — Fig-. 1. Plants nat. size (Eng. Bot.). !2. Ditto (Hooker). 3. Portion of I'rond, antical view x (ditto). 4. Cross-section of portion of frond x 60 (Elfving). 5. Portion of frond, bearing antheridia x (Hooker). 6. Anthe- ridium x (ditto). 2. Metzgeria furcata (L.), Radd. Lichenastrum saxatile erectuvi, temufolium, furcatam, Dill. Cat. pi. Giss. p. 21 ;> (1718). Lichenastrum fur catum, Dill. Hist. Muse. p. h\o, ut synon. (1741). Jungennania /areata, L. Sp. pi, 1, ed. 2, p. lloG,n. 20, excl. synon. Mich. (1753). Metzyeria glabra, Rad. in Att. soc. Modena, 18, p. 45 (1818). Metzgeria /areata, Dum. Recueil, 1, p. 2(), min. p. (1835). Dioicous, coespitose, closely jDressed to surface, small; pale to yellowish-green in colour. Fronds imbricating, irregularly branched and furcate, sometimes subsimple or alternately sub- pinnate, linear, here and there narrower, plane or undulate, sometimes slightly convex, antical surface smooth, postical clothed with long white hairs, which are single, most being usually near the midrib and margin ; midrib prominent, projecting only on the postical side, a cross-section showing it to be oval, and about 10 cells in circumference; texture thin,membranaceou.^, one cell thick; cells medium to rather large, regularly 5- and G- angled, lax and chlorophyllose, walls thickened. Female inflores- cence produced from the midrib on the postical side, bracts two, very small, convex, roundish, nerveless, ciliate. Calyptra elongato-pyriform, carnose, beset with short rigid setie. Pedicel short. Capsule subglobose, brown, when ripe splitting to the base into 4 valves, walls composed of two layers of cells, the inner with semi-annular threads. Spores greenish-yellow, tetrahedral-roundish, finely granulate, broader than the elaters. 462 HEPATIC^. Elaters brown, unispiral. Aiidroecia proceeding from the mid- rib on the postical side of frond ; bracts ventricose, beset with numerous short hairs, enclosing 2-4 roundish antheridia, which are very shortly stipitate. Gemmiparous ; gemmae terminal on attenuated branches, discoid, oval or roundish, furnished later with hairs, Var. ani(/inosa. Hook. Minute, of a blue colour, changing to green. No other good characters are to be observed. Dimensions. — Fronds ^ to J inch long, '5 mm.-l" mm broad; cells "035 mm. --05 mm. Hab. — On trunks of trees, stones and rocks, in exposed or shady situations. Moderately common. 1 to 17. I. Europe, Africa, North and South America, Australasia. Obs. — The species belonging to this genus are easily recognised, even in their barren state, by their thin fronds with prominent midrib, which projects on the postical side, and the presence of long white hairs on their surface. The two good species separated from M. furcafa by the late Prof Lindberg are at once identified by the characters given with each species. Description of Plate CCVI. — Fig. 1. Plants nat. size (Eng. Bot. 1632). 2. Plant x (ditto). 3. Portion of frond, postical view ? X (Hooker). 4. Ditto $ (ditto). 5. Portion of frond X 60 (Elfving). 6. Cross-section of portion of frond x GO (ditto). 7. Calyptra x (Hooker). 8. Spores and elaters x (ditto). METZGEBIA. 463 3. Metzgeria conjugata, Lindh. Jungermania fiircata (non L.), Weiss. PI. crypt, fl. (Jott. p. ] OS (1770), and others. Jungermaniafurcata, var. elongata, Hook. Brit. Jung, in textu ad tab. hh et .')(> (pp. ?), (isis-k;). Metzgeria conjugaia, Lindb. in Act. See. Fl. Fenn. 10, p. 4',)."), n. 27 (1875), Schiffner in Engler k. Prantl, Pflanzenfamilien, 91 und !)2 Lief. p. 5^5 (with figures) (1.S03) ; McArdle in Proc. Royal Irish Academy, 3rd ser. vol. iv. No. 5 (with plate) (181)8). Monoicous, stratificato-c?espitose, medium size, pallid, pale g-reen to green in colour, when dry slightly polished, very pellucid. Fronds dichotomous, or in smaller forms irregularly branched and furcate or sub-bipinnate, branches short, linear, narrower in some parts, convex above, margins revolute, apices obtuse, usually broader, antical surface smooth, postical, with the margins and midribs pilose with rather long, straight, divaricate hairs, the hairs usually double and very frequently with sucker- like branches at their extremities, midrib about 20 cells in circumference, projecting on the postical side ; texture firm, cells largish, regularly 5- and 6-angled, lax. Female flowers situated on the midrib on the postical side ; bracts pilose, hairs straight and divaricate. Calyptra pyriform, clothed with numerous long divaricate hairs. Androecia very numerous, situated on the midrib, antheridia 3-10, shortly stipitate. Fruits in Spring and Autumn. Dimensions. — Fronds 1 to 2 inches long, 1* mm. -2* mm. broad ; cells '04 mm. -'065 mm. Hab. — On shaded rocks or trunks of living trees. Somewhat rare. 2. Ardingley Rocks, Sussex, G. E. Davic-s. 7. Llanberis, Carnarvonshire, IF. //. P. 9. Thrang End, Lane, ./. J. Whcldon. 10. Dent, G. Stabler; Ingleton, IF. West. 12. Kentmere Planta- tion ; Park Beck ; Grisedale ; Possett Ghyll ; Rayrigg "Wood, Windermere, Westmorland, G. Stabler ; Isle of Man, G, A. Holt. 13. In sub-alpine glens. New Galloway, /. J/-S'. M. Macvicar. 17a. Sutherlandshire, Br. R. K. Grecille. South of Ireland, Killarney, P)r. D. Moore, Prof. Li/idber//, Canon Petf, D. McArdle. Europe, Asia, North and South America, New Zealand, &c. Obs. — This is a much larger plant than M.fnrcata, of a paler yellow colour, with more elongate less furcate branches, which are remarkably convex, having fewer hairs on the postical surface, a cross- section showing about 2 near the margin and 3 on the midrib, the wings being otherwise smooth, the hairs are also curiously hooked-deflexed. Description of Plate CCVII. — Fig. 7. Plant natural size. 8. Portion of frond x CO (Elfving). 9. Cross-section of portion of frond x 60 (ditto). 1130 HEPATICJ^:. Suborder II. MAUCHANTlACE.E. Genus 45. MARCHANTIA, March. jiJ. ilarchantia, March, fil. in 3Iem. Acad. Paris. (17l;5)- Micheli Gen. (1720); Linn. Arc. Fronds flesh}-, rarely membranaceous, opaque, often ver\Mjroad, with a broad diffused midrib, densely radiculose, apex emarginate or dichotomous ; no postical branches; antical face reticulate or porose ; postical face with scales on each side, 2-5 seriate. Inflorescence dioicous, terminal. Peduncle arising from a sinus in the apex of the expanded forking frond, elongate, bi- (very rarely 3) canaliculate. Capitulam more or less convex, many lobed, radiating, rarely subentire, above smooth, below with narrow scales. Involucre with several (2-5) flowers. Perianth with apex 3-5-fid. Capsule globular, stipitate, exserted, pen- dulous, dehiscent by 4-8 laciniate segments, often revolute. Elaters long, slender, attenuate at each end, 2-, 3- (very rarely 1-) spiral. Spores medium-minute. Andra^cia pedunculate, peltate, radiate or lobed. Gemma? lenticular borne in a cup- shaped receptacle. Marchantia polymorpha, Linn. Sjj. PL 11, 1G03 (1753). Dioicous, c?espitose, medium to large in size, dull green on the antical side, on the postical brownish. Fronds closely imbri- cate, procumbent, dichotomoiisly branched, flattened, branches short, margin entire, sinuous, midrib broad, thick and very distinct; radiculose, rootlets numerous of two kinds, the stronger but simpler kind are the root hairs which penetrate the sub- stratum and perform the same functions as the true roots of higher plants ; the otliers are more delicate and have internal thickenings arranged spirally ; these delicate rhizoicls do not penetrate the sub-strnium but lie flat up to the postical surface MARCHANTIA. 467 of the thallus, covered by the scales ; texture between flesljy and membranaceous ; on the antical surface there are numerous stomafca, each situated in tiie centre of a diamond-sliaped area, the postical side beuig furnislied witli numerous brownish scales. Female receptacle produced from the g-rowing- points of the thallus, pedunculate, hemispherical, deeply divided to the base into G to 12, usually 9, linear decurved rays, coverinj^ as many involucres, wliich are united at the base and mixed with minute chafty scales. Peduncle 2-channelled. Involucre oblong-, open at the end and torn, enclosing an ovate quadrifid perianth. Calyptra obovate, persistent, fissured at the apex. C.^ipsule ovate or globular, exserted, pendulous, pale greenish brown, protruding a little beyond the perianth, opening into about 8 short re volute segments or teeth. Spores smooth ; elaters Inspiral, slender, attenuate at each end. Male receptacle greenish, peduncu- late, produced from tlie growing point of the thallus, peltate, fiat, horizontal, papillose, flesli}^ thin at the margin, crenately 8-lobed; antheridia flask-shaped, imbedded in the receptacle opening- by an orifice through the papillae. On the antical surface of the thallus are numerous hemispherical cups with laciniate margins contain- ing numerous roundish, lenticular (contracted near the middle) gemmte. Dimensions. — Fronds 1 to 4 inches long, -3 cm.-0"8 cm. wide ; peduncle 2*5 cm.-7'5 cm. high, 1 -5 mm. x r25 mm. diam. ; male peduncle 2*5 cm. high ; gemmio "Go mm. x 'bb mm. ; lacinitu of receptacle containing gemmae '2 mm. long. Hab.— By the sides of streams or in other damp and wet -situations, on earth, walls or stones. Frequent in greenhouses. Ver}^ common, except in the Highlands of Scotland. 1-17, 18b. I. Generally distributed over Europe, Northern Asia, North America, also found in Soutli Avnerica, Japan and Java. Obs. — One of the commonest of our frondose species and easily recognised. For man}' of my notes and drawings of this species I am indebted to a valuable paper contributed by my friend Mr. L. W. 468 HEPATIC^. Waechter to the Proceedings of the Manchester Microscopical Society, 1891. Description of Plate CCVIII. — Fig. 1. Female plants, natural size (Carrington). 2, Thallus with mature female inflorescence, two-thirds natural size, showing how the midrib of the thallus branches, and how the branches become the stalks of the inflorescence (Waechter). 3. Cross-section of the stalk of a female inflorescence x II (W. H. P.). 4. Vertical section of a young female receptacle when grown to about a quarter of an inch above the thallus with rows of pistillidia (Waechter). 5. Vertical section through an older female receptacle: youngest sporogonium (nearest the stalk) on the left ; oldest sporogonium very nearly ripe (Waechter). 6. Vertical section of female receptacle x ? (Bischoff). 7. Involucre, calyptra and capsule x ? (ditto). 8. Portion of an elater, highly magnified (Waechter). 9. Young thallus with immature male inflorescence ; the midrib in this case is not branched (ditto). 10. Vertical section through a male inflorescence, antheridium nearly ripe in the centre of the disc; youngest antheridia near the mar^^in of the disc (ditto). 11. Portion of cup x ? (ditto). 12. Gemma x 11 (W. H. P.). Genus 46. CONOCEPHALTJS, Necl^. Ifejyatica, Mich. Nov. pi. gen. p. 3 (1729); Lindb. Hep. utv. p. 5 (1877). Marchantia, L. Sp. pi. ed. p. 1138 (1753). C'onocepA«Z«s, Neck. Elem. Bot. Ill, p. 344 (1790); Dum. Comm. p. 115 (1822). Fegatella, Raddi, in Opusc. Scient. di Bologna, 11, p. 35G (1818) ; Nees, Eur. Leberm. iv. p. 170 (1838). Fronds large, dichotomous, cartilaginous, reticulate, with a narrow distinct nerve. Female flowers pedicellate, conical or mitriform, membranaceous. Involucres 5-8, tubular, 1 -fruited, suspended from the apex of the peduncle, coherent with the interior of the receptacle. Perianth wanting. Calyptra persistent, campanulate, 2-4-lobed at the apex. Capsule oblong-pyriform, dehiscing by 5-9 revolute segments, pedicellate. Spores muricu- CONOCEPHALUS. 469 Icite. Elaters short, thick, bispiral. Male flowers sessile Kear the apex of the frond, disciform or oval. Conocephalus conicus (^Xccl:), Bum. Ilepalica vulgaris, Mich. Nov. pi. gen. p. o (1729). Marchantia conica, L. Sp. pi, ed. 1, p. 1138 (1753). Conocephalus conicus, Dum. Coram, p. 115 (1822). Feyatella conica, Corda in Opiz Beitr. 1, p. Gl'J (1829). llepatica conica, Lindb. Hep. utv. p. 5 (1877). Conocephalmn conicum (L,), Necker, Steph. 8p. Hep. p. 141 (]90<»). Dioicous, closely csespitose in thick patches, large, pale green antically, often purple postically. Fronds aromatic, prostrate, dichotomous, segments oblong, obtuse or emarginate ; midrib distinct, about 20 cells thick near the middle. Epidermic cells large, 6-sided, thin-walled. Furnished on the postical side with numerous reniform or cordate scales of a purple colour and delicate texture. Hadiculose, rootlets proceeding from the mid- rib, long, white, intricately twisted. Spores dark brown to black, muriculate. Elaters short, thick, brown, bi-trispiral. Male receptacle arising from the midrib, antheridia about 20, white-opaque. Fruits in Spring. Dimensions. — Fronds 1 to 3 inches long, segments ^ inch broad, '5 mm. thick near the middle, "1 mm. near the margin; cells of postical la3^er '05 mm. diara. ; stomata on antical side •4 mm. X *3 mm. ; postical scales '4 mm. x "5 mm., pedicel 1 to 1 J inch long ; female receptacle G* mm. x 4'mm., 5* mm. x 4' mm. ; capsule 1"5 mm. diam. ; spores "OG mm. diam. ; elaters •5 mm. X "03 mm. ; male receptacle 3* mm. x 2* mm.; antheridia 1" mm. X '3 mm. Hab. — Damp shady banks or on wet rocks. Common. 1-18. I. Europe, Asia, Japan, North America. Obs. — Ptecognised from other frondose hepaticic by its distinctly reticulated epidermis, and the large whitish stomata which are easily seen by the naked eye. 470 HEPATIC.^. The plant has also a pleasant aromatic scent. Description of Plate CCIX. — Fig. 1. Sterile frond, natural size. 2,3. Male fronds, natural size. 4. Portion of cross-section of frond x 24. 5. Portion of frond, postical layer of cells x 85. 6, 7. Scales from postical side of frond x 24. 8, 9. Female receptacles, natural size. 10. Ditto, magnified (Bischoft). 11. More highly magnified (ditto). 12. Cross-section of same, magnified (ditto). 13. Caljptra [a) and capsule {b) magnified (BischoflP). 14. Capsule, explanate x. 15. Cross-section of male receptacle x 11. IG. Antheridium x IG. Genus 47. REBOULIA, Raddi. Marchcmtia, Linn. Sp. pi. 1 ed., 2, p. 1138; Fl. Lapp. (1753). Asterella, p.p. Beauv. in Encycl. Meth. Suppl. 1, p. 502 (1810). RehouUa, Raddi in Act. soc. Modena 11, p. 357 (1818), (nom. em. ISTees, 1846). Frondose, fronds rigid, very indistinctly porose, the midrib broad, strong and distinct. Female receptacle j^edunculate, conical, hemispherical or plane, 1-G- (usually 4-) lobed, lobes divided to about the middle, peduncle continuous with the frond, barbulate-palaceous beneath. Outer involucre 1 -fruited, coherent with the lobes, 2-valved. Perianth wanting. Calj-ptra oval, minute, lacerate, persistent at the base of the capsule. Capsule hidden by the valves of the involucre, greenish, subglobose, nearlj- sessile, rupturing at the apex by irregular narrow teeth, or by a fragmentary operculum. Spores tuberculate. Elaters moderately long, mostly bispiral. Inflorescence monoicous. Andrtecia sessile, lunate-disciform. Reboulia hemisphaerica, Bculdl. Marchantia hemtsphcprica, Linn. Sp. pi. p. IGOi (1753). Asterella hemispharica, p.p. Beauv. in Encjcl. Meth. Suppl. 1, p. 502 (181(1). Eeboidia hemisplia'rlca, Raddi in Act. soc. Modena 1], p. 357 (1818). Monoicous, ca^spitose, medium size, dark green above, purple below and at margin. Fronds rigid, simple, bilobed or dichoto- IIEBOULIA. 471 mously branched, sinus often deep and narrow: prostrate, Hat, costate, midrib broad and prominent, margin crenate, undulate ; epidermis smooth ; below squamose, scales dark purple, imbricate, quadrate-elongate ; radiculose, rootlets copious, slender, white, proceeding from the midrib. Female receptacle pedunculate, subrotund, afterwards hemispherical, deeply 4-5-lobed, proceeding from the antical side of frond, at the base of the sinus ; surround- ing the base of the female peduncle, also at the apex, there are long, white, silky, hair-like processes, 1 and 2 cells wide. Perianth wanting. Calyptra minute, lacerate, persistent at the base of the capsule. Capsule composed of several layers, the innermost without any spiral threads, globose, rupturing irregu- larly, teeth not revolute. Spores muriculate, pale yellow with hyaline border when young, brown when old and dr3\ Elaters very closely twisted, narrow. Antheridia narrow, numerous, immersed in the antical, sessile, crescent-shaped discs. Fruits in Spring and early Summer. Dimensions. — Fronds 1 to 2 inches long, 5* to (>• mm. broad; female peduncle 1 to 2 inches long; spores •! mm. diam. ; elaters '35 mm. long. Hab. — On dryish, exposed cracks oi" rock filled with earth. Somewhat rare. 1. Extremely rare, Paul Hill, Cornwall, //'. 6V/'yv/o/r. 2. Hamp- shire. 3. Sydenham, j\Iarf//ji ; lane leading from liustliall Com- mon to the High Rocks ; Lamberhurst Quarter, J e inter ; Boro' Green; Ightham ; Halstead, Kent, E. j]f. Jlolii/c-s. 4,5. Weaver Hill, Staffordshire, 7^. Gdnirr. 7. On rocks, Barmouth, Merionetli- shire, Rer. T. Sal toe//. Dr. Cdrrhujiou tS" // . H. P. 8. Castleton, Derbysliire, /. Uliiteliead. \). Dalton Crag, Lane, ./. A. Wheldan. 10. Ingleboro, iJr. Carr'nujtoii ; Dentdale, G. S fabler; Thornton Force, ir. West ; Settle, 7/. F. Parsons ; Malham Moor, V. l/'e.s/ ■ Ingleton, Jf. II. P. 11, 12. Among limestone rocks, Levens Park, Westmorland, G. Slahler. 13. Back of Kenmure Castle, New Galloway ; rocks near Grennan, Dairy; about Kirkcudbright, /. McAiidreir. 15, IG. Ptather common on exposed rocky banks, Moidart, West Inverness, S. J/. j\faevicar. 472 HEFATIG^. I. DunkeiTon, Br. Taylor ; Dingle Bay, i^y. Carrington ; lime- stone rocks near Cork and Fermoy, /. Carroll ; abundant on the walls of the bridge at Cong, Co. Galway ; Sillaghbraes, near Larne, Co. Antrim, Dr. D. Moore ; on sandy ground at the North Bull, near Dublin, D. Mc Anile ; Island of Eathlin, Co. Antrim, >S'. A. Steicart. Found on the Continent, Asia, North and South America, Java, Japan, Australia, New Zealand. Obs. — Much rarer than Preissia conim/dafa, with which it is frequently confounded. "Easily distinguished by the deeply divided 4 -5- fid receptacle, which is barbate beneath and at the base of the peduncle, and by the sessile antheridia, whereas the receptacle of Preissia cohtmutata is marked on the top with a cruciate crest and the ^ pedunculate. "^ — Dr. Carrington. Another character at once makes the difference certain — viz., the innermost layer of the walls of the capsule of P. cornmutata have in their cells reddish-brov/n spiral threads, whereas they are absent in this species. Description of Plate CCX. — Figs. 1-4. Fertile plants natural size. 5. Frond, postical view. 6. Plant with young (r/ c? ) and {h ?). 7. Cross-section of pedicel X 11 (W.H. P.). 8-13. Female receptacles. 14. Ditto, showing young capsule. 15,16,17. Cap- sule in different stages x ?. 18. Spores x ?. 19. Spore and elater, more highly magnified. 20. Cross-section of male recep- tacle x ? (Bischolf ). Genus 48. PREISSIA, Conla. Preissia, Corda in Opiz Naturalientausch (182D) ; Nees, Eur. Leberm. (1838). Frond obcordate, sparsely furcate, margin thin, increasing by joints from the apex, pores conspicuous, bearing the peduncle from a sinus of the terminal lobe. Gemma.^ wanting. Female receptacle pedunculate, hemispherical, 4-6-lobed, rays of the lobes shorter, rib-like, only free at the inferior part, librous-barbu- late underneatli. Perianth obconically campanulate, angular, PREISSIA. 473 unequally 4 or 5 parted. Calyptra persistent, rupturing obliquely at the apex. Capsule large, shortly pedicellate, dehiscing by 4 or 8 irregular revolute valves. Spores large, tuberculate. Elaters short, bispiral. Male receptacle pedunculate, peltate, repand, lobate. Preissia commutata [Lindenb.), Nees. Marchantia hemispharica , Linn. Fl. Suec. ii. 1052 (1745). Marchantia quadrata, Scop. Fl. Carn. t. 63, 1356 (1772). Marchantia commutata, Lindenb. Hep. Eur. p. 101 (1829). Marchantia androgyna, Tayl. in Trans. Linn. Soc. xvii. p. 380 (1835). Preissia commutata, Nees, Nat. Eur. Leber, iv. p. 117 (1838). Preissia hemisp/uerica, Cogn. in Bull. Soc. Bot. Belg. p. 49 (1872) ; Dum. Hep. Eur. p. 152 (1874). Chomiocarpon quadratus (Scop.), Lindb. Muse. Scand. (1879). Dioicous, closely ca?spitose, medium size, pale green to brownish-green antically, dark purple postically and at the margin. Fronds prostrate, oblong-sinuate, apices bilobed, porose, margin irregularly crenate, on the postical side squamose, scales purplish-black, oblong, acinaciforni. Female flowers pedunculate, peltate, angularly hemispherical or sub- quadrate, sub-entire or lacerate, above obtusely 4-costate, 2-4-lobed, loculi attached to the underside of the lobes, 1-3-fruited, de- hiscing irregularly. Perianth obconico-campanulate, 4-5-lobed. Peduncle below red, above pale green, bicanaliculate, channels on both sides with rootlet-threads. Calyptra rotundate, styled, persistent. Capsule globose, shortly pedicellate, reddish- brown or dark purple, dehiscing by revolute segments, cells of the inner wall with reddish-brown ring fibres. Spores brown, submuricate, roundish-tetrahedral. Elaters long, slender, bispiral. Male receptacle pedunculate ; peduncle very thick, obtusely angular and irregular in outline, outer walls dark purple, 1 or 2 inner layers purple, others hyaline, trigones large, walls thin, bicanaliculate, with rootlet threads in the channels; smaller than the female, often subsessile; peltate, margin scariose, thin, undulate, irregular but undivided ; 474 HEPATIC^E. antheiidia immersed, disposed in rays, oblong. At the base of the peduncle are a few ver}^ delicate white scales. Dimensions. — Fronds 1 to 2 inches long, -} to h inch broad, near the margin "5 mm. and at the middle l* mm. thick : spores -08 mm. diam. ; elaters '25 mm. long ; female pedicel 1 to 2 inches long; male pedicel |- to f inch long; 75 mm. x *5 mm. diam. ; male receptacle 2* mm. to 3' mm. diam. ; antheridia "4 mm. X -2 mm. H.VB. — On rocks in exposed or -shady situations. Frequent in sub-alpine localities. 6, 7. Barmouth, Merionethshire, Lr. Carrinr/fon 8^- IF. II. P. ; Llanberis, Carnarvonshire, J. Ca-'. Moore, Pro. Bo//. Ir'ixli Acad. p. G03 (1876). Kilhirney, Br. Carriu/jlon, and others. Found on the Continent (Pyrenees, l>r. Spruce, Prof. Sc/iiii//jcr, Italy), and in the Southern States of North America. Obs. — Our finest frondose species, and which seems to be almost peculiar to the South of Ireland, as only two other stations, where it has been sparingly collected, are known. When seen not likely to be confounded with any other of our frondose species. Dr. Spruce and Prof. Sehitfncr consider it a variety of the tropical Buuwrtlera hirs/ffa (Swartz), and Dr. Stephani in his Sp. Jlcp. merges it with that species. Description of Plate CCXIII. — Fig. 1. Portion of frond, natural size. 2-4. Female fronds, natural size (Tayl.), 5. Cross- section of frond x 11 (Killarney, S. 0. L.). 6, 7. Cilia x (U (ditto). 8. Cross-section of pedicel x (Ta}^.). 9. Calyptra x (Tayl). 10. Female receptacle x (ditto). 11. Ditto, reversed X (ditto). 12. Spores and elaters x (ditto). 13. Male frond, natural size (ditto). Genus 51. TARGIONIA, J//V//. Targionia, Mich. Nov. PI. Gen. (17;i;i). Frond furcate and continuous from the apex, conspicuously porose, squamulose beneatli. Distinct female receptacle absent, the involucre sessile springing from the apex of the frond, bivalved, 1-fruited. Pistillidia ;> to 4, of which one is perfected. Perianth absent. Calyptra thin, persistent, investing the capsule, at length vanishing at the apex. Style deciduous. Capsule solitary, shortly pedicellate, globose, rupturing irregularly. Spores globose, tuber- culate. Elaters bi-trispiral. Andnecia lateral, disciform, papillose, rising on a separate innovation from the antical costa. 480 HEPATIC^. Targionia hypophylla, L. Targionia hypophylla, Linn. Sp. pi. p. 1604 (1753). Targionia Michelii, Corda in Opiz Beiti\ 1, p. 649 (1829). Monoicous, growing in spreading patches, small, dark green in colour, with niore or less conspicuous whitish pores, margin and postical aspect chocolate-purple. Fronds imbricate, simple, obovate or obcuneate, plane, involute when dry, costate, thick along the middle, margin thinner, a cross-section showing the antical outer layer to be greenish, inner whitish, postical brownish purple ; clothed on the jDOstical side with numerous, densely imbricate, broadly lanceolate, chocolate-purple scales ; cells elongate, irregularly 4-, 5-, and 6-sided, with several, much smaller, roundish, hyaline ones containing minute, nucleate granules ; rootlets very numerous, hyaline. Perianth sessile, protruding postically from the apex of the frond, composed of two semi-roundish, slightly emarginate, dark purple bracts. Calyptra thin, persistent. Style deciduous. Capsule shortly pedicellate, when young pale brown, deep purple when ripe, oval-orbicular, protruding slightly or not at all beyond the perianth, bursting irregularly. Spores globose, dark brown, margin slightly paler, coarsely tuberculate. Elaters 2-4-spiral, pale brown, much narrower than the spores. Androecia lateral, disciform, papillose, rising on a separate innovation from the antical costa. Dimensions. — Fronds 8mm. to 10mm. long, by 4 mm. to 5 mm, broad, "75 mm. thick; scales '45 mm. x "15 mm. broad at the base; cells of scales '08 mm. x '03 mm., "05 mm. x 'OS mm., •05 mm. X "04 mm. ; small nucleate cells '02 mm. ; bracts 1*25 mm. X 1*5 mm. x 2" mm.; capsule 1"75 mm. x 1*4 mm.; spores "05 mm. diam. ; elaters *225 mm. x "015 mm. Hab. — On exposed warm dry or moist rocky banks. Rare. 1, 2. Hampshire. 5. Moist sunny red sandstone rocks at the entrance to Habberley Valley, Kidderminster, Worcester, Dr. F. A. Lees, March 1883; Dove Dale, Staffordshire, /. E. Bagnall SPHJEROCARPUS, 481 Barmouth, rocks above the town, and by the road to Harlech, Iiei\ T. Salwet/ ; Barmouth, on the Dolgelly Road, Br.Carriiiglon Sf ir. JI. P. 10. I. Cave Hill, Belfast, John Teuipleton ; on dry limestone rocks, Carrigaline, near Cork, /. Carroll ; on the warm basaltic rocks at Deerpark, Glenarm, Co. Antrim, Br. B. Moore. C. Bale de Saint Brelade, Aug. Martin. Found on the Continent, Central Asia, Africa, Australia, Tasmania, New Zealand, North and South America, &c. Obs. — As only one species of this peculiar genus has been found in Europe there is little difficulty in identifying it. From other genera it is distinguished by the postical sessile perianth, visible at the apex of the frond ; in habit it resembles the large Rlccia glaucescens Carr., but grows in larger patches with the fronds usually spreading in one direction, and when dry remarkably involute, showing the almost black postical scales. Description or Plate CCXIV. — Fig. 1. Plants natural size (Templeton). 2. Ditto, slightly enlarged. 3. Portion of frond, antical view x (ditto). 4. Ditto, postical view x (ditto). 5. Frond, postical view x 3 (Schiffner in Engl. & Prantl Pflanzf ). (). Portion of frond, antical view x (Templeton). 7. Cross- section of frond x 11 (Alliers, France, du Buysson). 8. Scale x (Schiffn. in Engl. & P. Pfl.). 9. Ditto x 64 (Alliers, du Buysson). 10. Portion of scale x 290 (ditto). 11. Bracts with ripe cap- sules X 3 (Schiffn, in Engl. & P. Pfl.). 12. Bracts and capsule X 16 (Kidderminster, F. A. Lees). 14. Bracts, explanate x 11 (ditto). 14. Male branch x 5 (Schiffn. in Engl. & P. Pfl.). 15. Longitudinal section of the same x 5 (ditto). Genus 52. SPHiEROCARPUS, MlcUclL Sph'i'rocarpus, Mich. Nov. PI. Gen. 4, t. 3 (1729); Dum. Comm. Bot., p. 78 (1822). 2\ir(/ionia, Dicks. PI. Crypt. Brit. fasc. 1, p. 8, n. 2 (178.J). Fronds small, orbicular, costate, epidermis indistinct. Perianths produced on the antical surface of the frond, aggregated, sessile, 2 H 482 HEP A TIC.E. pyriform or obtusely conical, perforated at the apex, 1 -fruited. Calyptra crowned with a deciduous style, closely investing the globose, indehiscent capsule. Spores globose, inuriculate, remain- ing in a coccus. No elaters. Antheridia in folliculose bodies on the surface of separate fronds. Sphsgrocarpus terrestris, Jficlt. Sphcerocarjms terrestris, Mich. Nov. pi. gen. p. 4, t. ;) {ll'l'd) ; Dill. Hist. muse. p. 53G, t. 78, f. 17 (1741); Smith, Eng. Bot. t. 297 (1790). Targionia Sj^hcerocarpus, Dicks. PI. crypt. Brit. fasc. 1, p. 8 (1785). Spha'rocarjms Jlichelii, Bell. Act. Taur. v. 2.j8 (1702) ; Cooke, Brit. Hep. f. 195 (1868). Sphcerocarjms Sphm-ocarjms (Dicks.), Howe, Hep. k, Anthoc. Calif, p. (iO (189!)). Dioicous, closely cjcspitose, small, pale green in colour, some- what glaucous. Fronds flat, orbicular, midrib 2 to 3 cells thick, indistinct, graduall}' disappearing towards the thin membranaceous margin, which is slightl}' and variously lobed, lobes short and rounded, concealed by the aggregated inflated perianths ; texture thin, reticulated; radiculose, rootlets long, single, hyaline, papillose within. Perianths aggregated on the frond, sessile, obovate or pyriform, when young longish or elliptical, variable in size, apex obtuse or sub-truncate, when 3'oung closed, afterwards opened, mouth small, rounded, texture semi-transparent, regularly 4-6-sicIed cells. Pistillidia 2-5, oblong. Calyptra very delicate, bistratose at the base, at the apex unistratose, crowned with a deciduous style, closely investing the capsule, and on its maturity disappearing. Capsule globose, included in the perianth, inde- hiscent, bursting irregularly, composed of a single laj^er of cells. Spores greenish-blaclv, angular, muriculate. No elaters. Antheridia spherical, shortly pedicellate, situated on the surface of separate fronds which are smaller than the others. Gemmiparous, gemmae dispersed on the frond, dark coloured, globose, elliptical or oval. Annual. DiAirNsioNs. — Thallus 'G mm. to 1'3 mm. diam, ; perianth 1"5 mm. io 2*5 mm. long. Hab. — On the ground, especially clover fields. Very rare. JIICCIA. 483 4. Near Yarmouth, Bawson Tamer ; Gcorr/e Fitt. 5. Sellach, near Ross, Herefordshire, 72^^?. A. Let/, 1872 ; B. Jf. IFatkifis, 1881. 8. On a stone in Collin's Lane, Coleman ; Stubble field, Twycross, Leicestershire, Hlo.ram. 10. Found on the Continent, North Africa, and in North America. Obs. — A very distinct species and the only known British one of the i^enus. Description of Plate CCXV.— Fig. 1. Plants natural size (W. H. P.). 2. Ditto X 2. 3, 4. Ditto x 4. 5. Perianths in different stages of growth with portion of frond and rootlets x . G. Portion of rootlet x . 7. Portion of frond with section of perianth («) showing pistillidia and (//) gemma. 8. Youn"- capsule X . i). Half-ripe capsule x. 10. Half-ripe spores x. 11. Perfect capsule x. 12. Portion of calyptra x. 13. Cross-section of capsule {(t) and a ripe spore {J)). 14. Gem- ma) X (Bischoff). Suborder III. lUCCIACLE. Genus 5:3. RICCIA, MicL Jliccia, Mich. PI. Gen. (1721)). Fronds generally disposed in rosettes, at first radiating from the centre, which often decays, segments bifid or dichotomous; antical plane, depressed or canaliculate ; postical usually convex, smooth or squamulose; margin either naked or ciliate ; epidermis usually distinct, without pores. Fruit immersed in the interior of the frond, rupturing antically, sessile. Involucre none. Calyptra delicate, coherent with the capsule, style-bearing, style persistent, protruding. Capsule spherical, sessile, indehiscent within the calyptra. Spores tetrahedral, alveolate or muriculate. Kootlets papillose within. 484 HEPATICJE. 1. Riccia glauca, Linn. liiccia ylaiica, Linn. Sp. pi. iCO.') (17");)), Monoicous, growing in orbicular patches, closely attached to the ground, small, glaucous green in colour, concolorous. Fronds bifid or dichotomous, substellate, lobes linear, obovately-linear or cuneate^ 8 to 12 cells thick near the middle, plane or channelled near the apex, punctate, apex emarginate or obtuse, margin thin, membranaceous, crenulate or smooth, postical side green ; radi- culose, rootlets plentiful ; texture fleshy ; cells hexagonal or sub- quadrate, walls thick, no trigones. Capsules subspherical in a sinorle or double row radiatincr from the centre of the frond and O C) runnino; alons: near the middle of each lobe, crowned with a style, 4 to 6 cells long, of a dark colour, which alone is protruded, and soon deciduous. Spores dark reddish-brow^n, almost black when ripe, angular-globose, reticulate, walls of convex side irregularly tuberculate, papilke narrowly conical, areolae regular, 5, 6-angled, 8-10 measuring the convex surface, other sides almost smooth ; when young with a narrow pellucid border, scarcely visible when ripe. Var. minima. Minute, green on both sides, fronds bifid, lobes linear, thin, 4 to 6 cells thick, plane, slightly channelled near apex. Dimensions. — Plants \ to J inch in diameter ; fronds 6' mm. X 2- mm., 7 mm. thick, 5' mm. x l" mm., "5 mm. thick ; capsule '6 mm. ; spores '09 mm. x '07 mm. Var. minima, \ inch in diameter; fronds 1-75 mm. x '5 mm., •3 mm. thick, 2*25 mm. x '6 mm., "3 mm. thick, 3*5 mm. x "7 mm., '3 mm. thick. Hab. — On moist ground, chiefly clayey fields and gardens. 1-16. I. Common; rarer in Scotland. Europe, Asia, Japan, North America. Var. minima. Probably generally distributed. 9. Stretford, Lane, C. J. Wild; Cheetham, Lane, C. J. Wild. 12. Fowlshaw Moss, Westmorland, G. Stabler, Rice I A, 485 Obs. — This is the commonest species of the genus, and is easily recognised by its glaucous green colour, the same on both sides. Some hepaticologists consider Rlccia minuiia L. to be the same as It. sorocarpa Bischoff, but as probably all small forms of Biccia would be included under this name, I prefer to follow Bischoff, as he was the first to correctly and fully describe and figure the species, and I retain the small form which cannot be separated as a species from R. (jlanca as var. riiiiiiina. This must not be confounded with the plant described and figured by Lindenberg, which is purple beneath (Lindenb. Mon. Eiccia, p. 427, t. XX (1836) ). DESCRIPTION OF Plate CCXVL — Figs. 1, 2. Plants natural size (Bischoff). 3, 4. Ditto magnified (ditto). 5. Ditto, var. major x 2 (ditto). 6. Ditto, var. minima, natural size (Fowlshaw Moss, G-. Stabler). 7. Ditto x 11 (ditto). 8. Ditto, var. major (Baden, Jack). 9, 10. Cross-section of fronds x (Bischofl'). 11-13. Ditto X IG (Fowlshaw Moss, G. Stabler). 14. Portion of rootlet X 200 (Hoft'meister). 15, Antheridium x (j4 (Fowlshaw Moss, Gr. Stabler). 16. Convex surface of spores, highly magnified (ditto). 2. B-iccia crystallina, Z. Lichen palusfris, Dill. Hist. muse. p. 5^)5, t. 78, f. 12 (1741). Riccia crystallina, Linn. Sp. pi. 1605 (17r)o). Monoicous, forming rosettes, closely attached to the ground, small, pale or yellowish green on both sides, crystalline. Fronds flat, circular or wedge-shaped, segments bilobed or furcate, obcordate, margin slightly crenate, sometimes slighth^ raised^ antical surface pitted, epidermic cells 6-sided ; cross-section shows frond to be of equal thickness and full of air-cavities ; radiculose, rootlets few, long, hyaline. Capsule embedded in the frond, spherical, brown ; style conical, projecting, covered with a hyaline or faintly purple calyptra. Spores roundish-angular, reddish- brown, reticulate, 6-8 lumen across each face, walls raised, 486 HEFATICjE, irregular. Antheridia embedded in the frond, cylindrical, hyaline ; style projecting, hyaline. Dimensions. — Fronds about \ inch long by about \ inch wide ; capsule '65 mm. diam. ; spores "08 mm. diam. Hab. — On very damp earth in fields, on wet banks, or especially by the margin of ponds. Very rare. 3. In a ditch near Deptford, Kent, Feiiccr, now extinct. 4. Helford, West Norfolk, Rer. E. N. Bloouifidd, Aug. 18S1. 5. Stoke-on-Trent, 11. Garner; Pottal Eeservoir, AVarwickshire, /. E. BagncdJ. 7. Dolgelly, Merionethshire, /. Tlalf^, 18G3 ; Aberffraw, Anglese}^ //'. IVUsoii. 8. Exposed mud banks of reservoirs at Cropstone and Thornton, Leicestershire, F. T. Matt, Aug. 1894. 9. Mere Mere, Cheshire, G. E. Hunt, 18G9. 13. Brownhall Orchard, Dumfries, /. Crn'u-k^hank ? Found on the Continent and in North America. Obs. — This rare and distinct species ma}' be distinguished from M. (jlauca by its larger size, paler and 3'ellower colour, spongy texture and its antical surface being pitted with, deep cavities. It has been conjectured that it might be a terrestrial form of Bicciocmyus nafan-s, but, in addition to its other distinguishing characters, the spores are nearly twice as large and are leticulate, not papillose. Description oe Plate CCXVII. — Figs. 1-4. Plants natural size. 5. Frond x . G. Cross-section of frond x . 7. Ditto. 8. Portion of frond x. 9. Eootlets x. 10. Calyptra x. 11. Capsule X. 12. Style x. 13. Spores x. 14. Portion of frond, showing styles of antheridia. 15. Styles of antheridia x '(Lindenberg). FdCCIA. 4.87 3. Riccia sorocarpa, Bi.v/iolf. Iiiccia sorocfW7;rt, Bischoff, Hep. Nov. Act. N. Car. xvii. p, 1053, t. 71, f. 11 (1.S3:.). Jiiccia minima, L. Schifin. in Engl, und Prantl Pflanzenf. 1)1 und 92 Lief. p. ir,(i8'J3). Iiiccia minima, L. p.p. Howe, Hep. it Antho. Calif, p. 2-\ (18119). Moiioicous, CiL'spitulose, small, pale green in colour, when dry albescent, concolorous. Fronds subradiate, bifurcate ; lobes oblong-linear, on cross-section boat-shaped, salcate, deeply so \vlien dr}', with margins erect ; margins thin, smooth, or with lew hyaline scales ; apex acute or obtuse ; texture firm, antical surface finely reticulate. Capsules embedded in the i'rond, roundish, dark brown. Spores reddish -brown, walls thin, finely tuberculate on the convex sides, projections very minute and numerous. Dimensions, — Fronds about 4" mm. long ; lobes "75 mm, broad x '7b mm. thick, 1" mm. x '7b mm., l"mm. x -G n)m. ; when dry '6 mm. broad x 1' mm. thick, '75 mm. x '[) mm., •8 mm, X "5 mm. ; spores -O'Jo mm., '08 mm. x '055 mm. Hab. — On thin rocky soil, or more rarely in cultivated fields in limestone districts. Hare. 1, Battery Hedge, near Penzance, Cornwall, //', Canioic, Sept. 1871). 5. Gt. Doward Hill, B. U. ll'afhiu.^, Sept. 1S70 ; Cheddar Cliff's, near lioss, Herefordshire, J3. Jf. llafki/hs. 7. Barmouth, Merionethshire, //'. IL P., Aug. 1877. b. Millers Dale, Derby- shire, IF. II. 1\, Jan. 1871). 12. On earth, among rocks, near Levens Church ; VVliitbarrow ; in sandy cultivated fields S.W. of Bridge End, Levens, Westmorland, (L Stdhlcr. I. In fissures and tops of old walls ; on an old wall near Dingle, Liiidbcr// ^- Moore, 1873; wall top by the roadside, leading from Dingle to Ventry, the Dingle side of the river near the Union, Leif S>- McArdlc, ls98. Found on the Continent and in North America. Obs. — From any form of Riccia (jluiica L. distinguished b\' its deeply sulcate lobes, when dry, being apparently thicker than 488 HEPATIC^. broad, and its remarkably firm texture ; it is also generally much paler in colour, and is almost peculiar to limestone districts. Description of Plate CCXVIIL— Figs. 1 and 2. Plants natural size. 3, 4. Ditto, slightly magnified. 5. Cross-section of frond with young capsule x . 6. Ditto with capsule x (Bischoff). 7-10. Cross-sections of fronds, when dry, x 31 (Cheddar, AVatkins). 11. Epidermis x. 12. Style x. 13. Young spores x. 14. Spores x (Biscbofi'). 4. Riccia bifurca, Jloff'm. Riccia hi/urea, Hoffmann, Deutschl. Fl. p. !):» (171)0) (excl. syn. Mich. tab. 57 fig. 4). Monoicous?, closely ctespitose, small; antical surface glaucous green, margins and postical surface of fronds purphsh-black in colour. Fronds crowded and frequently imbricated, firmly attached to the surface, tumid-flabellate or pedate ; lobes elliptic, contracted at each end or obovate, rarel}' oblong, cuspidate, retuse, bifurcate or obovate-emarmnate, marmn elevated, rounded, antical surface sphacelate ; texture solid, homogeneous, upper stratum columnar, intermediate portion darker green, epidermis of the postical surface purplish-brown. Rootlets numerous, either capillary and translucent or papillose within. Sporangia scattered irregularly over the disc of the frond, at length rupturing the epidermis. Spores conspicuous, dark brown, three-angled, sub- cristate, rounded and reticulate, muricate on the outer surface. Dimensions. — Fronds 375 mm. long x 2-5 mm. broad, 3" mm, X 1"25 mm., 5" mm. x 2'5 mm,, from "3 mm. to "5 mm. thick at the middle, near the margin '5 mm. to 7 mm. thick, spores •095 mm., '1 mm. diam. Hab. — On ledges of rocks where there is a thin layer of soil and mud-covered walls, chiefiy in limestone districts and near the sea. Rare. 1 . Penzance, JJ"^. Ciniiow. 7. Aberfii'aw, Anglesey, IF. IVihoUy 1830 ; Barmouth, Merionethshire, Br. Carrinyioii, 1887, on mud- RICCIA. 489 covered walls, IF. IL P. 1876. 12. On clamp limestone rocks, Levens Park, Westmorland, G. Sf abler, 1870. 13. Burnfoot Hill, New Galloway, ./. Mc Andrew. 15. Head of Glen Dole, Forfar, J.Sadler, 1872. Found on the Continent and in North America. Ohs. — The only other British species which is without cilia and has a purple margin is Riccia ir'up'eJla 1).C. ; this is smaller, with narrow linear lobes, dark green antical surface, the margin and postical surfiice of which are clothed with regularly imbricating scales. Riccia (;laucescem Carr. is a ciliate species, usually much larger and irregular in shape, but small forms where the cilia are not so evident require careful observation to discriminate from R. /jifi/rca. The description and drawings are partly taken from Dr. Carrington's " New British Hepaticoe " contributed to " Grevillea," Dec. 1874. Description oi<' PlxVTE CCXIX. — Figs. 1-3. Fronds natural size. 4-7. Fronds x ?. 8. Frond x ?. 9-14. Cross-sections of fronds x 16. 15. Cross-section of frond near apex x 16. 16-19. Spores x 250 (Dr. Carrington & W. H. P.). 5. Riccia nigrella, JJ.C. JRiccia niyrella, De Candolle, Fl. Fr. v., p. 19;] (1^05); Lindenb. Monog. Rice. p. 4(;7, I. xxix. f. 1 (18:50). Blccia lamellosa, Ralfs, MSS. ; Cooke, Brit. Hep. n. 1;}3 (1SG8). Exsicc. Husn. Hep. Gall. n. DC ; Carr. k Pears. Hep. Brit. Exsicc. n. G.'>, et 2!)(>. Dioicous, ca?spitose, minute, dark green in colour antically : clothed with dark purple, almost black, transverse, closely imbri- cated semi-circular scales postically or on the postical margin. Fronds stellate or sub-stellate, narrow, linear, narrowly canalicu- late, margins thickened, involute, strongly so wlien dry ; radicuiose, rootlets few, hyaline. Spores angular globose, dark brown, almost black, slightly verruculose. Dimensions. — Fronds 4* mm. x 1- mm. : 35' mm. x 75 mm., 490 HEPATIC.^. •3 mm. to '5 mm. thick; cells of scales '04 mm., -05 mm. ; spores •075 mm. x '05 mm. Hab. — On banks and mud-covered walls. 7. Barmouth, Merionethshire, ./. Balfs, Dr. Carrhir/fon, W. IF. P., and others. Very rare. Found on the Continent and in jN'orth America. Obs. — Avery rare and distinct species, the dark purple, almost black, semi-circular scales on the iuflexed margins easily distin- guishing- it. Herr Stephani considers the British species to be distinct from Biccia iiif/rclla D.C, and has published it in his " Sjj. Hep.'' as Biccia Bearsoni. It seems very ungracious to question his deter- mination when he does me such honour, but I have not been able to satisfy myself as to our British species being distinct from French specimens upon which De CandoHe founded his species. All French specimens I have had the opportunity of examining are dioicous like our British ; the specimens in my copy of Husnot's Hep. Gall. n. 96, are male plants. JStephani gives the size of the British, species as 10* mm. long, but 1 have examined a great number of specimens, and find them very regular in size, like the French and Italian, 3"5 mm. to 4" mm. long; the apices are all more or less rotundate, the channel varies in being more or less sulcate ; cross-sections of British specimens agree exactly with French ones. He also describes the spores as being different in size, but I find those of Italian plants to be similar to the British, '075 mm. x 05 mm. I wrote to Herr Stephani expressing my regret that I could not agree with his observations, and he wrote the following characteristic kindly letter : " You are perfectly at liberty to doubt my publication and to give expression to it. I am not touchy at all. The drawings I here give are copies of \w\ tbrmer examinations ; you see the form oi the Italian plant is altogether different from that of the English one, and thh alojie icoidd be sufficient, all other points being equal. I have examined a great number of plants, which you kindly sent years ago, and 1 cannot help differing from you." IIICCIA. 491 I give the drawings sent ; wluitever nuiy be the position of the Italian plant, most of my drawings agree exactly with the French ones. Description of Platk CCXX. — Fig. 1 . Plants natural size. 2, 3. Fronds, antical view, magnified. 4, 5. Ditto x Hi (J3ar- moiith, J. Kalis). 0-8. Cross-sections x 24 (ditto), i), 10. Ditto (Paimpol, Cotes dii Nord, France, Dr. Avice). 11-13. Ditto (Husn. Hep. Call. n. <}6). 14-lG. Ditto (C. t P. Hep. Brit. Exsicc. n. 290, Barmouth, W. H. P.). 17, 18. Ditto (Florence, Italy, Dr. Levier). 19, 20, 21. Cross-sections, apex, middle, adult, Bicciff Fear-soju, alter IStephani. 22, 23. Ditto, apex, middle, Biccia i/i//r('//fi, after Stephani. (). Riccia tumida, Lindenh. Tikc'ia tumida, Lindenb. Syn. Hep. Eur. p. ]2(1Sl^;)); Lindenb. Monogr. lliccia, p. 4r)'J. t. 27 (18;)(;). /'Iccia Jlichelii, Raddi, var. cili((ris, Levier, Bull. Herb. Bois.s. vol. ii. No. 4, p. 230 (1894). L'iccia Jlichelii, Raddi, Stepli. Sp. Hep. p. 1 1 (IDOO). Dioicous ?, caespitulose, crowded, minute, bicolorous, light grej'ish-green antically, margin and portion of postical sides deep purple, almost black, maddle yellowish-green. Fronds simple or bifid with segments again slightly divided in some instances, linear or linear-obovate, obtuse at apex, thick, canaliculate, margin thickened, when dry stroni>-lv inflexed : texture fleshy; ciliatc, cilia marginal, biseriate, subulate, hyaline, often swollen at the base ; radiculose, rootlets plentil'ul, long, of equal length. Capsule embedded in the middle of the IVonds, spherical, dark brown. Spores angular-globose, very dark brown, almost black, the plane sides with pale brown border, Jaintly reticulate, convex side irregularly tuberculate, spines shorter and blunter than in Ji. (/I a lie (I. Dimensions. — Fronds from 2- mm. to 4- mm. long x -5 mm. to 1" mm. broad, '3 mm. to -5 mm. thick ; cilia IG mm. to '25 mm. long X -05 mm. to -08 mm. broad at the base ; rootlets "015 mm. thick • capsule '35 mm. x '4 mm. ; spores '1 mm. x "075 mm. 492 HEPATIC^. Hab. — On miicl-covered walls, banks, and rocks. Very rare. 1. On rocks near Penzance, Cornwall, W. Cnrjww, 1870 (Herb. W. H. P.); Trebar- with - Strand, North Cornwall, E. M. Holmes, Mrs. Ma 31. Thulall. 6. Lantyssil, Cardigan- shh-e, W. Joshua, May 1877. 7. On mud-covered walls and banks, Barmouth, Merionethshire,//'^. //. P., April 187G; T.Bogers, Sept. 1877. Pound on the Continent and in North America. Obs. — A minute but very distinct species, recognised by its two colours, green and purple, and the two rows of cilia. Blcc'ia (/laiicescens Carr. is a larger species, more irregular in shape, margin somewhat acute, cilia in a single row. Riccia ciliafa Hoffm. is destitute of the purple scales, fronds plane, cilia much more abundant. The Biccia fnn/ida, var. Pearsoiil Carr. (Lond. Cat. 12 b. 1881) is a form found in damper situations, where the purple scales are not so evident, fronds thinner, and the cilia not so regularly biseriate ; it approaches B. ciliafa, but from close study of the plant in sifii (Barmouth) I have little doubt but that it is simply R. tiimida modified by its damper habitat. Dr. Levier reduces M. tumida to a variety (var. ciliaris) of Kiccia Michelii Paddi, and Herr Stephani refers it to that species ; all the specimens I have seen from different localities, and all I have seen growing for several years at Barmouth, are so regular in size, shape and in having the double row of cilia, that I follow Lindenberg, who described and figured our British form so accurately. Descriptiox of Plate CCXXI. — Pig. 1. Plants natural size. 2. Frond x ? (Lindenberg). 3-5. Fronds x 24 (Cardiganshire, Wm. Joshua). 6, 7. Cross-sections of fronds x ? (Lindenberg). «, 9. Ditto X 24 (Cardiganshire, Joshua). 10. Ditto x ? (ditto). 11-15. Cilia x 64 (ditto). 16. Cilium x 290 (ditto). Rice I A. ' 493 7. Riccia glaucescens, Carrhifjlon. Riccia glaucencens, Carr. in Carr. it Pears. Hep. Brit. Exsicc. fase. 1, n. GO (187><). liiccia Michelii, Raddi, Lindb. Muse. Scand. p. 2 (187!>). Riccia bifarca, Hoflm. Steph. 8p. Hep. p. 'M) (1 ",)()()). Dioicous, cicspitose, small, pale glaucous green antically, border somewhat paler, postical surface covered laterally by a delicate membrane or detached scales of a purple colour, best seen when the plant is dry and the border incurved, mid-line brownish. Fronds flabellate-furcate, sometimes crowded and imbricating each other ; segments linear-cuneate or battledore-shaped, concave, not canaliculate on the upper aspect, surrounded by a rather broad border, recurved and convex when moist, but inflexed when dry ; texture homogeneous, composed of large cells arranged in regular series, thickened along the mid-line of the lower surface, margins thinner, acute, fringed with small but strong translucent cilia, sometimes these are wanting or irregularly disposed, and the border simply crenulate ; radiculose, rootlets hyaline. Capsules few, occupying the hollow central channel near the base of the frond ; spores large, brown, muriculate, angular-convex, about 10 areolae across the convex side. Dimensions. — Fronds G" mm. to 12- mm. long, segments 1- mm to 3- mm. broad, -3 mm. to 7 mm. thick; cilia -l mm. x '05 mm. ; capsule "75 mm. ; spores "1 mm. Hab. — On damp banks or mud-covered rocks in exposed situations, usually near the coast. Eare. 1. Penzance, Cornwall, //^. (7^/r//o?r. 7. Barmouth, Merioneth- shire, i)>-. Carrhfffon, W. H. P.\ AberfFraw, Anglese}', TF. Wihon; Aber, Carnarvonshire, W. Wihon. 12. Leveus Park, Westmor- land, G. Stabler. 13. Portpatrick, /. McA/idrcw. 15. ^lontrose Links, A. Cruo.l/ ; Den of Fullarton, Forfar, ^1. Croall. Obs. — A large and characteristic species discovered by my late friend Dr. Carrington at Barmouth, where it grows in some abundance, remarkable for its glaucous antical surface, hence its name. The only other bicolourous ciliate species which has been found 494 HEP A TICJi. ill Britain \^ Riccia fumida Lindenb. which is veiy distinct from it, bein2^ a much smaller plant, very neat and regular in its shape, and possessing- a double rowof cilia; both species grow at Barmouth, and there is no difficulty in distinguishing them with the naked eye. Riccia Lesciirlana Aust., a North American species, approaches the nearest to it, but is thinner, not so purple in colour, with the margins more incurved and with more numerous cilia, which are often 2 and 3 rows deep ; it is also monoicous according to Prof. Howe. Riccia Bi.^cJioJfii Huben., a Continental species, is nearly allied to it, but in this the fronds are broader and shorter, generally bilobed, obovate or obcordate, channelled above, but thick and prominent beneath, surrounded with a broad membranaceous margin, fringed with large obtuse cilia. Herr Stephani in his ">S/y. Hep. " p. 14, refers Riccia (jJauccsceiis to Riccia hifiirca Hoffm., a non-ciliate species. Lindberg refers it to Riccia Michclii Raddi ; to the ciliate form of which Dr. Levier refers this species. The description and some of the observations are taken from Carr. & Pears. Hep. Brit. Exsicc. n. 6G, contributed by Dr. Carrington. Description or Plate CCXXII. — Fig. 1. Plants natural size. 2. Frond x 11 (Barmouth, W. H. P.). 3. Cross-section of Irond X 24 (ditto). 4-8. Ditto x 24 (Barmouth, Dr. Carring- ton). 9,10. Ditto X 16 (Aberffraw, W.Wilson). 11,12. Cilia X 25 (Barmouth, Dr. Carrington). 13. Style x 85 (ditto). 8. Riccia ciliata, Hoffm. Riccia ciliaia, Hoftmann, Deutschl. Fl. p. D.J (179G). Monoicous,'ca?spitulose, small, pale or yellowish-green in colour, concolorous. Fronds sub-orbicular or llabellate, somewhat thin, dichotomously or sub-stellately divided ; lobes linear or slightly cuneate, obtuse or sub-emarginate, towards the apex sub-canali- culate, margin slightly thickened and fringed with numerous long, PilCCIELLA. 495 subulate hyaline cilia, a little swollen at tlie base ; texture some- what delicate; epidermic cells punctate; radiculose, rootlets numerous, filiform, papillose within. Capsule embedded in the frond, globose, black; spores dark brown, almost black, tuber- culate, spines blunt. Dimensions. — Plants forming circular patches | to -J inch in diameter; lobes 3* mm. to 4" mm. long x 7 mm., "8 mm. and r mm. broad; cilia '075 mm. to 'I'lb mm. long x •01 mm. broad at the base, spores "09 mm. diam. Oi?s. — Distinguished from any of the other ciliate lUccioi by its green colour on the postical as well as antical surface of the fronds, and the numerous long cilia. Hab. — In damp places by ditch sides or on clayey fields. Yery rare. 1. On mud, Tavistock, Devon, ./. Ihilfn ^' //'. Ciirnow, July 1875. Found on the Continent and in North America. Description op Plate CCXXIII. — Fig. 1. Plants natural size. 2. Plant x . 3. Portion of lobe x . 4. Cross-section of lobe X . (). Cilia x (Lindenberg). Genus 54. RICCIELLA, Br am/. Lichenastrum aquaticuvi, Dill. Hist. muse. p. .")14, t. 7i, f. 47 (1741). Hiccia, Linn. Sp. pi. IGOO (175;',). Ricciella, Braun, Bot. Zeit. p. 7:)(; (1S21). Plants slender, linear, usually several times dichotomous, scales not present. Fruit globose, exserted, adnate to the postical surface of the frond. Ricciella fiuitans (/.), Bra/n/. Lichenastrum aquaticinti, Dill. Hist. muse. p. 514, t. 74, f. 47 (1741). Ricciajiuitans, Linn. Sp. pi. 1(50(5 (1753). Ricciella fiuitans, Braun, Bot. Zeit. p. 754 (1821). Dioicous, ca?spitose or spreading, not forming rosettes, medium size, yellowish-green in colour. Fronds dichotomously branched, segments narrow, linear, plane or slightly canaliculate, thin, con- 496 HEPATIC^. colorous, apex emarginate or obtuse ; radiculose when terrestrial only, rootlets hyaline. Female flowers adnate to the postical side of frond. Capsule globose, dark brown, coarsely reticu- lated, the areolae 5, 6 across the convex side, walls dark brown, thin, raised in a winged manner. Dimensions. — Fronds ^ to 2 inches long, 1" mm. wide, 'ii mm. thick; capsule '7b mm.; spore "1 mm. Hab. — Floating on stagnant ponds, or on mud at their margins. Somewhat rare. 2. Strathfieldsaye, Hampshire, i/"///, 1855. 3. \\.Q\\i,Forsfer. 4, 5, 8. Pond near Ashley ; pond between Donington and the Trent, Leicestershire, F. T. Mott. 9. Ashton Canal, h^nc,!. E. Smiderland; Moston, Lanc.,i?. Lees ; Reddish Canal, Lane, G. A. Holt ; Mere Mere, Cheshire, G. E. Hunt. 10. Hirst Courtney and Drax, H. F. Parsons ; pond near PoUington, T. Birh,Jnnr. ; near Goole, H. F. Parsons. 14. L Not unfrequent in ditches near the Shannon, Co. Limerick ; by the side of the Bann River, above Drogheda ; still ditches near Lough Neagh, where the canal joins the Lough at Lurgan, Dr. D. Moore. Found on the Continent, North America, &c. Obs. — The fertile terrestrial form of this species is considered by some writers to be a distinct species, but as it is only found on mud at the margins of ponds where the floating form is found, and where the ponds have become lower by dryness, I can only conclude that whatever slight changes it may have undergone are owing to the great difference in habitat to that of floating on water, where it is always barren ; the terrestrial form is much smaller, narrower, thicker, and more canaliculate. Description of Plate CCXXIV.^ — Fig. 1. Plants natural size (Bischoff). 2. Ditto (Lindenberg). 3. Frond x ? (ditto). 4, 5. Ditto (Bischoff). 6, 7- Cross-sections oF fronds x (Lindenberg). 8, 9. Ditto X 24 (Mere, Cheshire, Hunt). 10. Capsule x (Bischoff^). 11. Cross-section of frond showing capsule x (ditto). 12. Spore x 290 (Barmouth, Dr. Carrington). Rice IOC ARP US. 49 7 aenus 55. RICCIOCARPUS, Corda. Ricciocarpus, Corda in Opiz Nat. (1829). Riccia, Linn. Syst. 956. Plants obcordate, constantly dividing with growth, air-cavities abundant, postical surface naked or furnished with long dentate scales. Fruit immersed in the frond and not protruding on either surface, at length naked by the splitting of the central groove. Antheridia in a row in the middle of the frond. Ricciocarpus natans (Ziuh.), Corda. Riccia natans, Linn. Syst. Nat. ed. v. 12. p. 2 (1753). Ricciocarpus natans, Corda in Opiz Nat. p. (J51 (1829). Dioicous, floating on water or when terrestrial forming rosettes, small, pale or dark green above, jDurple below. Fronds simple, obcordate or obcuneate, broadly emarginate, channelled in the centre, furnished with long linear-lanceolate ribbon-like, flattened, purple, serrated membranaceous processes; epidermis with numerous uniform air-cavities beneath it ; texture spongy ; radiculose, root- lets few, very long, hyaline, smooth within. Capsules immersed in the frond about the middle in two rows. Spores brownish- black, angular, strongly papillose, papillae blunt. Dimensions. — Fronds J to |^ inch long, segments 3" mm. broad ; capsule '7b mm. diam. ; spores '05 mm. Hab. — Stagnant ponds, floating, or on damp margins of the same. Hare. 2. Hampshire,^///; Warner. -4,5. Near Eoss, Herefordshire, B. M. WatMns. 7, 8. Thurcaston, Leicestershire, i^. T.Motf. 10. Hebden Bridge, S. Gibson ; near Dringhouses, near York, JF. IFcsf. " Rare in Ireland. Abundant in a large boggy pool, about half-way between Drogheda and Navan, near the Railway, right- hand side going from Navan to Drogheda ; ditch by the side of the Shannon, near Portumna, Co. Gal way. Ditches near Passy, Co. Limerick, Dr. W. H. ILarccij'' — Dr. D. Moore. 2i 498 HEPATIC^E. Found on the Continent, North America, &c. Obs. — When floating the ribbon-like processes are long and abundant, sometimes they are only faintly tinged purj)le ; when growing on land, they frequently disappear, or are only few. Description of Plate CCXXV. — Fig. 1. Plant natural size {Lindenberg). 2,3. Ditto (BischofF). 4. Plant, slightly magnified (Bischoff). 5. Cross-section of frond, magnified (Lindenberg). 6. Cross-section of portion of frond, showing immersed capsules, magnified (Bischofi"). 7. Ditto, showing antheridia (ditto). 8. Epidermis, magnified (ditto). 9. Hair-like process, magnified (ditto). 10. Capsule, magnified (ditto). 11. Young spores, magnified (ditto). 12. Spores, magnified (ditto). Suborder IV. ANTHOCEUOTACE^E. Genus 56. ANTHOCEROS, Mich. Anthoceros, Mich. Gen. PI. (1729). Fronds dark green or blackish, often orbicular ; nearly plane or with centre depressed, radiately lobate ; lobes variously divided; costa very wide, confluent, confused with the pagina; texture lax, vesicular, with large chlorophyl granules, frequently glandularly thickened at the apex or in streaks along the middle so as to appear nerved. Inflorescence monoicous or dioicous. Involucre tubular. Capsule pedunculate, exserted, linear or cylindrical, very long, oblong, bivalved, furnished witli a columella. Elaters in a few species perfectly spiral, in most only undulate or sinuate, simple or branched, often geniculate, more or less heteromorphous, fibres wanting or indistinct. Spores papillose or smooth. Antheridia irregularly disposed on the antical surface of the frond. ANTHOGEROS. 499 1. Anthoceros laevis, L. Anthoceros kcvis, Linn. Sp. pi. 1 (!()G (17");3). Dioicous, CcTespitose, small to medium size, dark green. Fronds flat, smooth, without air-cavities ; lobate, lobes undulate, some- what orbicular, rounded, not laciniate ; texture somewhat fleshy, epidermic cells 4 times smaller than the inner, 3 to 4 cells thick near the middle of the frond ; radiculose, rootlets numerous, pale dirty brown. Calyptra trumpet-shaped, 1 to 2 cells thick near the middle, 3 to 4 near the base, mouth wide, repand, toothed, rarely scariose. Capsule brown, dark brown to black when dry ; spores yellowish-brown, flatfish, angular, almost smooth ; elaters rather short, geniculate, somewhat articulated. Andr^ecia irre- gularly dispersed over the face of the frond, oval; antheridia oval, pale green, slightly stipitate, about 6 in each cavity. Dimensions. — Fronds J to J inch long and broad, '2 mm. to *3 mm. thick ; calyptra 3* mm. long x 1- mm. broad, 2* mm. x 1" mm.; capsule f to 1 inch long; spores -05 mm. diam. ; androecia "3 mm. x '2 mm. Hab. — Wet fields, damp roadsides, by the sides of ditches, &c. Rare. 1. Stable Hobba, near Trereiff*e, Newlyn CUft; Trungle, Paul, Jr. C/fr//o/r. 3?, 5, 7?, 10, 12. In a cultivated field near Hever- sham Railway Station ; cultivated fields (sand and sandy-peat), Nether Levens and Foulshaw, Westmorland, G. Sfnhler. 17a. Caithness, Rev. B. LiUie. I. On a wet clay bank by the roadside, leading from Dingle to Ventry, left-hand side of the road about a mile and a half from Ventry, Lliidb. &f Moore, 1873. On a pathway in Burnham Wood, Ventry, Left §• McArdlc, 1898. Found on the Continent and in North America. Obs. — Distinguished from At/f//ocero-'> jjHricfat/f.s by its dioicous inflorescence, the smooth surface of fronds, no air-cavities, lobes rounded not laciniate, paler colour of the spores, which are not echinate. ^00 HEPATIC^. From A. Stableri, see notes under that species. Description of Plate CCXXVI. — Fig. 1. Fertile plant natural size. 2. Ditto x . 3. Cross-section of frond x 16. 4, 5. Calyptra x 11. 6. Cross-section of calyptra x 11. 7. Ripe capsule opened x . 8. Male plant natural size.. 9. Portion of male plant x 11. 2. Anthoceros punctatus, L. Anthoceros punctatus, Linn. Sp. pi. 1606 (1753). Monoicous, ca3Spitose, small to medium size, bright pale green- in colour. Fronds in layers, thin, subirabricate, spreading in a circular manner, centre depressed and concave, lobes suberect, somewhat dichotomous, oblong and obtuse, without any distinct midrib, if present confluent and confused with both sides of the frond, more or less deeply sinuate, pinnatifid or laciniate, crisped ; cross-section of frond shows about 2 to 3 layers of cells, rarely 5 to 6 ; texture delicate, glandular, reticulate ; cells lax, epidermic layer 4-, 5-, and 6-angled, chlorophyllose, chlorophyl granules large, angular ; walls thin ; stomata distinct ; small remote scales scattered over the frond ; radiculose, rootlets pale dirty brown, arising from base or centre of frond. Calyptra antical, lower half 4 to 6 cells thick, inner layer of cells very minute, others irregular in size with very thin walls, oblong-linear, Slightly repand, when young closed at the top, and later rup- tured by the protruding capsule, leaving an irregular lacerate opening, parts of which are torn aw^ay or remain adhering as scariose rudiments. Capsule pedunculate, exserted, very long, black, pod-iike, filiform, arising like blades of grass, tipped with a veil when young, bursting when ripe into 2 valves with a parallel partition. Spores black, strongly echinate, angular, on the convex side beset with blunt prickles, single or 2 and 3 together ; on the other two sides irregularly verrucose. Elaters brownish- black, short and broad, flattish, geniculate, variously contorted. ANTIIOCEUO^. 501 Androecia imbedded irregularly in the frond ; antheridia pale brown, oval, stipitate, stipes 4 cells long by 2 cells thick, 8- 1 in each cavity. Dimensions. — Fronds \ to \ inch long and broad, "2 mm. to '3 mm. thick at the middle ; capsule about 1 inch long ; spores •05 mm. ; elaters 'Vlh mm. x '015 mm. ; antheridia '125 mm. x '1 mm. Hab. — In damp fallow fields and on ditch banks, &c. Some- what rare. 1. West Cornwall, rather common, )V. Cnriiow ; Sidmouth, South Devon, Urn. FMa M. Tiiidall ; Somerset, T. Briffain. 3. Kent, DHIen'mx. 5. Little Fenton, Staffordshire, 11. Garner. 7. Very finely in Corsygedol Woods, Barmouth, Merionethshire, Rev. T. Salwei/ ; Dolgelly, Merionethshire, C. J. Wild. 8. Lough- borough, Leicestershire, Pulteneij; Ditch near Twycross, Leicester- shire, Roc. A. B/oxaii/. 10. 13. New Quay, Kirkcudbrightshire, /. CriAckxIiauk. IG. Hare ; at the side of ditches, Moidart, West Inverness, fruit Oct., /S'. M. ]\[acricar. 17a. Caithness, Rev. D. LilVu-. I. Frequent in the counties of Kerry and Cork, Dr. I). Moore. Mount Brandon, W. Vl/mu, 1829; Maghanabo Glen, Z). J/c., t. ix. (1847). GoTTSCHE, C. M.-^" Untersuchungen uber Haplomitrium Hookeri." Nv. E. Acta Acad. Caes. Leoj). Carol. Nat. Cur. xx. 267-398, t. xiii.-xx. (1843). " Uber die Fructification der Jungermannife Geocalycete." Acta Acad, Caes. Leop. Carol. Nat. Cnr. xxi. 419-466, t. xxx.-xxxii. (1845). " Keure Untersuchungen uber die Jungermannise Geocalycese." Ahhandl. des Natarw. Vereins in Hamburg, vii. 39-66, t. (1880), and numerous other contributions. GoTTSCHE, LiNDEXBERG, and Kees — " Synopsis Hepaticarum." 8vo, pp. 834, Hamburghi (1844-1«47). GoTTSCHE, C. M., et Eabexhorst, L. — " Hepatictv Europa? (exsiccatse)." Decas 1-66 (Nos. 1-660), Dresden (1862-1879). Gray, S, F. — " A Natural Arrangement of British Plants." Two vols. 8vo, London (1«21). Hoffmann, G. F. — " Deutschlands Flora Kryptogamie." Erlangen (179.")). Hooker, William Jackson — " British Jungermannite." Folio, 8S tab. London (1S16). " Flora Scotica." Two vols. 8vo, pp. x. 292, 297, London (1821). Hooker, W. J., and Taylor, T. — " Muscologia Britannica." Second ed. 8vo, pp. 272, 36 tab. London (1827). Hudson, William—" Flora Anglica." 8vo. pp. viii. 506, London (1762). HuBENER, J. W. p. — " Hepaticologia Germanica." 8vo, pp. 314, Mannheim (1834). HtJBENER, J. W. P., et Genth, C. F. F. — " Deutschlands Lebermoose exsiccata," HusNoT, Tranquille — " Hepaticologia Gallica." 8vo, pp. 102, 13 pi. Caen (18H1). " Hepaticai Gallise (exsiccatje)." Fascicules i.-ix. Nos. 1-231. Jack, Josef Bernard — " Die Lebermoose Badens. Bericht. der naturf. Gesellsch. zu Freiberg (1870). (Sep. pp. 92). " Hepaticee Europee." Bot. Zeituny, xxxv. 49-59, 65-74, 81-93, 97-111 (1877). (Review of Dumortier's work). " Die Europaischen Radula-Arten." Flora, Ixiv. 353-362, 385-400, t. vii., viii. (1881). (Sep. pp. 24). " Monographie der Lebermoosgattung Physiotum." Hedwiyia, xxv. 49-87, t. i.-x. (1886). Jenner, Edward — " New Localities for Jungermania Turneri." Phytoloyist, i. 718 (1844). Lett, H. W. — " Report on the Mosses, Hepatics, and Lichens of the Mourne Mountain District." Proc. Roy. Irish Acad. 3rd ser. i. 265-325 (1890). BLBLIOGEAPHY. 507 Libert, Marie Anna — " Sur un genre nouveau d'Hepatiques, Lejeunia." Ann. Gen. des Sci. Phys. (Bruxelles), vi. 372-374, t. 1)6 (182U). "Plantar Cryptogamica^ Ardennes (1830-37). LiGHTFOOT, John — " Flora Scotica." Two vols. 8vo, London (1777). LiMPRicHT, Karl Gustav — " Lebermoose, in Cohn : Kryptogamen- Flora von Schlesien" (1H77), and numerous other works. LiNDBERG, Sextus Otto — " Monographia Metzgeria^." 8vo, pp. 48, 2 pi. Hel- singfors (1877). " Hepaticee in Hibernia. Mense Julii (1.S73), lectee." Acta Soc. Sc. Feiin. X. 407-559 (1875), and numerous other works. LiNDBERG, S. O., et Lackstrom, E. F. — " Hepaticse Scandinavica^ exsiccata?." Fasc. 1, Nos. 1-25, Helsingfors (1874). LiNDENBERG, J. B. W. — " Synopsis Hepaticarum Europsearum." Nova Acta Acad. Caes. Leop. xiv. Suppl. pp. 126, 2 tab. Bonn (182'J). "Monographie der Riccien." Xova Acta Acad. Caes. Leoj). xviii. 361- 504, tab. xix.-xxxvii. (1836). LiNDENBERG, J. B. W., et GoTTSCHE, C. M. — " Species Hepaticarum." 4to, 07, tab. col. Bonn (1830-51). LiNN.EUS, Carl — "Species plantarum, etc." Two vols. 8vo, pp. 1200, Holmiai (1753). Macvicar, S. M. — " Hepaticse of Moidart." Journ. Bot. p. 348-356 (1899). " Pellia Neesiana Limpr. in Britain." Journ. Bot. p. 275 (1900). " New British Hepaticse." Journ. Bot. p. 36 (1901). " A Key to Bi-itish Hepaticae." Journ. Bot. p, 154-167 (1901). " Scapania crassiretis Bryhn in Britain." Journ. Bot. p. 210 (1901). Martius, K. F. p. von — " Flora cryptogamica Erlangensis." 8vo, pp. Ixxviii. 508, 6 tab. Nuremberg (1817). Massalongo, Caro — " Hepaticologia Veneta." 3 tav. Padova (1879). " Hepaticffi Italife-Venetfe exsiccatje." 4 fasc. Nos. 1-120 (1878-1881);^ and numerous other works. Massalongo, C, et Carestia, A. — "Epatiche delle Alpi Pennine." Nuooo Giorn. Bot. Ital. xiv. 212-258, t. x.-xiv. (1882); and other works. Mc Andrew, J. — " Radula voluta Tayl. in Scotland." Journ. Bot. xxvii. 51 (1889). McArdle, David — " Notes on some New or Rare Irish Hepaticfe." Proc. Pay. Duhlhi Soc. iii. 17-21, pi. v., vi. (1881). " Hepaticfe of Wicklow." Journ. of Bot. xxvii. 11, 12 (1889). " Hepaticaj of Lough Bray, County Wicklow." Journ. Bot. xxviii. 356- 361 (1890). McIvoR, W. C — " Hepaticfe Britannica?, or Pocket Herbarium of British Hepaticfe." Small 8vo, New Brentford (1847). MiCHELi, p. A. — " Nova plantarum genera." 4to, pp. 234, 108 tab. Florence (1729). Mitten, William — " A List of all the Mosses and Hepatica^ hitherto observed 508 HEPATIC^E, in Sussex." Ann. Nat. Hist. viii. 305-324, 3^vo, pp. xvi. 352, Leipzig (1848), &c. Raddi, G. — Jungermanniogi^aphia Etrusca." Mem. di Matem. e di Fisica della S'oc. Ital. delle Sci. (Modena), xviii. 14-56, 7 tab. (1820), &c. Reader, H. P. — "Monmouthshire Cryptogams." Journ. Bot. xx. 120 (1882). " The Hepaticje of Gloucestershire." Journ. Bot. xxii. 331, 332 (1885). RossETTi, C. — "Appunti di Epaticologia toscana." Xuovo Giwn. Bot. Ital. XX. 412, 413 (1888), &c. Roth, A. "VV. — " Tentamen Flora^ Germanicaj." Three vols. 8vo, Leipzig (1788- 1800). Salwey, T. — " A List of Certain Plants to be met with in the Neighbourhood of Barmouth, Dolgelley and Harlech." 12mo, pp. 40 (1863). ScHiFFNER, V, — " Observationes de quibusdam Hepaticis." Bot. Centralbl. xxvii. 207-211, 239-243, 1 Taf. ^886). " Hepaticje in Engler und Prantl." Pflanzenf. (1893-5), &c. BIBLIOGRAPHY. 500 ScuMiDEL, C. C. — " Icone.s Plantarum, Ac." Folio, pp. 11)7, .")0 tab. col. Nurem- bei- (1747), kc. SciiRADEK, H. A. — " Plantje Cryptogamicii' nova-, rariore.s aut minus cognitje." Jour II. far die Botanik, i. (!()-8(> (1801). ScHREBEK, J. C. D. — " Spicilegium Flora; Lipsica-.'' Svo, pp. 148, Leipzig (1771). ScHW.ECJKiciiEX, 0. F. — " Historia Muscoruiii hepaticarum Prodromu.s." 8vo, pp. o;>, 1 tab. Leipzig (1814), tfec. ScoPOLi, J. A,—" Flora Carnolica, etc." Second ed. 2 vols. 8vo, Vienna (1772). Scully, R. W. — " Hepatica^ found in Kerry, 1881)." Journ. J)oi. xwiii. 'JuO- 208 (18DU). Smith, J. E. — >See Sowerby, J . SowERBY, J. — " English Botany." Thirty-six vols. 8vo, 2592 tab. col. with text, London (170U-1814). Spuuce, Richard — " A List of Mosses and Hepaticae collected in Eskdule, Yorkshire." Phytoloyist, i. r)40-:»44 (1844). " On the Branch-bearing Leaves of Jungermannia juniperina Sw." I/u/to- logist,u. 85, 8G (1845). " A List of the Musci and llepatica; ot Yorkshire." Phi/tolofjiat, ii. 147-157 (1845). " The Musci and Hepatica3 of Teesdale." Trans. Hot. >Soc. Edin. ii. G5-89 (1846). " The Musci and Hepatica- of the Pyrenees." Ann. and May. Nat. Hist. 2nd ser. iii. Sl-lOG, 269-298, 858-380, 478-503; iv. 104-120, t. i.-iii. (1849); Trans. Bot. S'oc. Pdin. 108-216, t. i., ii., xiv. (1850). *' On Anomoclada, a New Genus of Hepaticse, and on its Allied Genera, Odontoschisma and Adelanthus." Journ. Bot. xiv. 129-136, 161-170, 193-208, 280-285, t. clxxviii., clxxix. (1876). (Sep. pp. 88). " Musci Pra?teriti sive de Muscis nonnuUis adhuc neglectis prsetervisis vel confusis, nunc recognitis." Journ. Bot. xix. 88-40 (1881). " On Marsupella Stableri, n. sp., and some Allied Species of European Hepaticae." Rev. Brijol. viii. 89-104 (1881). " On Cephalozia, its Sub-(4enera and some Allied Genera." 8vo, pp. 99, Malton (1882). " Hepatica? of the Amazon and of the Andes of Peru and Ecuador." Trans. Bot. > 302 Ajylozia — {continued) (jracillima, Dum. . hyalina, Dum. larida, Dum. pmnila, Dum. riparia, Dum. sphcerocarjxi, Dum. tristis, Dum. Aste7'ella. hemisphcerica, Beauv Bazzaxia, Gr. d- B. . Pearsoni {Stephani) triangularis {Schleich.) Lindb. tricrenata (Wahlenb.) trilobata (L.), Gr. cC- L Blasia, Afich. . pusilla, L. Blepharostoma, Dum. connivens, Dum. setacea, Dum. setiformis, Lindb. . trichophyllum {L.), JJum Blepharuzia, Dum. . ciliaris (Z.), Dam. lloffmanni, Dum. . pulcherrima(y/o//yM.),Z?« IVoodsii, Dum Blytiia. Lyellii, Endl. Blyttii. Moerckii, G. L, N. Calycularia. hlyttii (Mcerck). . Calyj)oyeia, Dill. aryuta, Mont, it Xees IVichomanis, Dum. Cephalozia, Dam. . au-aiia, Pearson albesceiis, Dum. 48 49 47 196 302 3G4 298 292 294 296 294 470 126 133 130 132 128 440 440 111 157 124 114 112 — 104 3.S 104 — 106 db. 39 106 — 102 — 432 — lliH ~ 438 — 1 :{4 — 139 — 13, 6 — 141 69 181 — 192 512 HEPATIC^. PLATE PAGE PLATE PAGE VOL.2 VOL. 1 VOL. 2 VOL. 1 Cephalozia — {continued) Clasmatocolea — {continuec I) alpestris, Cogn. 331 cuneifolia {Hook.), Spruce lOi 254 bicuspidata {L.), Bum. . 57 150 Codonia. catenulata {Hilben.) 54 144 Dumwtieri, Hiib. & connivens {Dicks.) . 60 157 Genth. . 423 curvifolia {Dicks.), Dum. 61 159 pusilla, Dum. 418 dentata {Eaddi), Lindb. 73 187 Ralfsii, Dum. 430 denudata {Xees), Sjn-uce 'o'o 174 Coleochila. divaricata {Smith), Dum. 67 177 anomala, Dum. 267 elachista {Jack), Lindb. . 71 184 cuneifolia. Spruce . 254 fluitans {Xees), Spruce . &?> 165 Taylori, Dum. 264 Francisci {Hook.), Dum. 62 103 CoxocEPHALUS, Xeck. 468 Helleri, Lindb. 351 conicus {Xeck.), Dum. .' 209 469 heterostipa, Carr. el Spruce 64 168 Cordea. hibernica, Spruce MS. . 59 155 Blyttii, Cord. — 438 islandica, Lindb. . — 194 Dilama. Jackii, Limpr. MS. 70 182 Blyttii, Dum. — 438 laxifolia, Dum. — 197 hibernica, Gr. & Benn. — 434 leucantha, Spruce . 12 186 Lyellii, Dum. 432 lunula?folia, Ditm. . 56 147 Diptlomitrion. media, Lindb. — 147 hibernicum. Cord. . 434 midtiflora. Spruce . — 147 Diplomitrium. obtusiloba, Lindb. . — 165 Blyttii, Cord. — 438 pallida, Sp'uce 55 146 Diplolcena. serriflora, Lindb, . — 144 Blyttii, Cord. — 438 Sphagni {Dicks.), Spruce 65 171 hibernica, Dum. — 434 stellulifera {Tayl. MS.) Turneri, Lindb. . 68 179 190 Lyellii, Nees — 1-430 \434 Cesia, Gr. ii' B. . 390 DiPLOPHYLLUM, Dum. 238 adusta (Nees), Lindb. . — 399 albicans (Z.), Dum. 97 239 alpina {Gottsch.), Lindb. . 172 391 Dicksoni {Hook.), Dum. 100 243 brevissima {Dum.) 176 399 Hellerianum, Dum. — 251 concinnata {Light/,), minutum, Dum. 353 G. cC- B. . . . 178 403 obtusifolium {Hook.), conferta, Limp'. . 175 397 Dum. 99 241 corallioides {Xees) 177 401 saxicolum, Dum. . 355 crassifolia {(Jarr.), Lindb. 174 395 taxifolium {Wahl.), Dum . 98 240 crenulata, Gottsche 179 407 DuMORTiERA, Xees 477 obtusa, Lindb. 179 405 hirsata (Swartz.), Stejyh. — 477 revoluta {Xees), Lindb. . 173 393 irrigua {Wils.), Xees 213 477 Chandonanthus, Mitt. — 113 Eremonotus, Lindb. . — 200 setiformis {Ehrh.), Mitt. 42 114 myriocarpus ( Carr.), Lind b. 78 201 Chiloscyphus, Dum. — 256 Eucalyx. polyanthos {L.), Dum. . 106 257 obovata, Lindb. — 366 Chomiocarpo7i. Fegatella. quadratus (Scop.), Lindb. — 473 conica, Corda — 469 Cincinmdus, Radd. . 134 FossoMBRONiA, Eaddi — 416 argutus, Dum. — 139 angulosa {Mich., Dicks.), Sprengelii, Dum. . 138 Raddi 185 421 Trichomanis, Dum. 136 angulosa, Raddi — 424 Clasmatocolea, Spruce — 254 ciespitiformis, De X. 187 424 INDEX. 513 PLATH I'AOB PI ATI PA OB VOL. 2 VOL. 1 VOL. •> VOL. I Fossomhroniii - (contlmted) HY(iROIJIELLA, Spruce i9(; coi'hulceformis, Trab. — 1:50 laxifolia {Ifoolc), Sprue ", t ( 197 cristata, JAndh. . ISI 120 ■aevicensis, Carr. . 3H9 Duiiiortieri {Ifilh. <( Jamksoxiella , Sprvce 2s(; aenth.), JAndh. . . ISC, 423 Carringtoni {Half.) J'oreoluta, J^indl). . — 423 Sjn'uce 121 287 Mittenii, y'iiid. 188 425 JuBULA, Dam. . 35 pusiUa {JJill., /,.), Bum Hutchinsiiu {Hook.) Lindh. . 183 418 Dum. 30 Frullania, Haddi . . 23 JUNGERMANIA, L. 288 dilatata {L.), Dum. 5 33 acuta, Lindl). 316 fragili folia, Tai/lor 3 2;) acuta, Wils. . 309 germana [Tai/lor) . 4 31 adunca, Dicks. — 99 Hutchinsiw; Nees . — 3(; a'qailoba, ,Schwaeg. 212 microphylla (GoUsche), P ears. 2 26 albescens. Hook. . — 192 Tamarisci (/>.), J)i(tn. 1 24 albicans, Linn. 239 Gymnant/ie. albicans, Nees 240 declpiexs, Mitt. * 204 ahjeriensis, Gott. . ;)09 Wilsoni, Tayl. 410 alpestris, Schleich. 142 331 Gymnocolea. anyulosa, Dicks. . 421 ajiiiis, Dum. , 300 anomala. Hook. 267 injlata, Dum. — 307 aquilegia, Tayl. 74 laxifoUa, Dum. — 197 asplenioides, L. 274 Gymnomitrion. attenuata, Lindb. . 342 Blyttii, Hiibeii. . Gymnomitrium. — 438 autumnalis, J)e ('. 129 1 3(13 1^305 adastum, Nees — 399 hadensis, Gott, 309 concinnatuvi, Cord. 403 barbata, Hook. 342 confertum, Limpr. . 397 barbata, Schreb. 148 344 corallioides, Nees . — 401 Bartlingii, Hampe — 209 crassifoliam, Carr. 395 Baueri, Mart. 1 59 cremdatum, Gottsche 407 (133 312 revolutum, Pliilib. — 393 bantriensis, Ifook. 'l34 314 liAKPANTiius, Xees . 259 i.3^ :;i(; Flotoviamts, Nees . _ 201 bicornis, Web. 35;; Flotovvii, Nees 108 2G1 bicrenata, Schinid. 139 324 scutatus, W. cf- .1/. . 107 259 bicuspidata, G. L. N. 144 IlejxUica. bicuspidata, Linn. — 150 conica, Lindl). — . 4(;9 bicuspidata, Nees . 153 .saxaiUis, Vaill. — 224 bidentata, Dum. . 247 vulgar in, Mich. — 4(;9 bide)/tata, Hook. . 312 Ilepaticoides. lilasKt, Hook. 440 albescens, Vaill. - 2:'. 9 niyttii, Moerck . 438 Hehherta, Gr. d- B. . _ 99 byssacea, Roth 177 adunca (Dicks.), Gr. <(■ /! :;i; !I9 calycina, Tayl. 447 llerpetiuni. calyjdrifolia, Hook. 64 repians, Nees - /117 |ll9 capitata, Jlook. catenulala, Hiiben. l;;s 144 trilohatitm, Nees . 128 ciliaris, L. 104 llyijropyla. cochlear if orm is, Weiss. . 91 irriyaa, Tayl. — 477 compacta, Iloth 2(tS - K 51' HEPATIC.-E. Jungermania — (continued) complanata. Hook. comjdanata, L. compressa, Hook. . concinnata, Lightf. conci7inata, Schleich. confervoides, Raddi connivens, Dicks. . connivens, Mart. . convexa, Scop. convoluta, Hiiben. corcyracea, Nees . Cordeana, Hiiben. cordifolia, Hook. . crenulata, Hooh. var. crenulata, Smith . culeccris, Wils. cuneifolia, Hook. . cu])7'essina, Swartz. citrta, Mart. . curvifolia, Dicks. . citspididigera, Nees decipiens, Hook. . deflexa, Mart. dentata, Raddi denudata, Nees Dicksoni, Hook. . distans, Schwein. . divaricata, Sm. Doniana, Hook. Donii, Hook. Dimiortieri, Lib. . elachista, Jack elongata, Hook. emarginata, Ehrh. endivif^folia, Dick; epiphylla., Hook. ejnphylla^ L. excisa, Dicks. excisa, Funck exigua, Tayl. exsecta, Schmid. Jissa, Scop. . flava, Swartz. Floerkii, Web. Flotoviana, Nees fluiians, Nees Francisci, Hook. Fi0ickii, Web. et Mohi furcata, L. PLATE \0L. 2 PAGE YOL.l _ 74 78 368 403 399 177 157 — 147 23(; 201 . 309 89 122 290 ]28 302 ]27 300 312 251 117 235 159 209 204 130 187 174 243 89 177 ir)7 361 361 269 184 - f463 ■\464 375 447 __ 447 443 322 379 282 145 337 135 49 348 261 165 163 379 — 461 Jungermania — {continued) furcata (non L.), Weiss, gelida, Tajjl. . Genthiana, Hiiben. Goepertiana, Hiiben. Goulardii, Husn. . gracilis, Schleich. . gracillima, Smith . hamatifolia, Hook. Hartmanni, Theden. Helleriana, Nees . heterophylla, Schrad . hihernica, Hook. , hibernica, Wils. Hqfinanni, Wallroth Hookeri, Lyell Hutchinsice, Hook. hyalina, Lyell hygrophylla^ Spruce incisa, Schrad. iitconspiciha, Raddi iiiilata, Hads. ivjiata, Nees . intermedia, Lindenb. interrupta, Nees ii-rigua, Nees islandica, Nees Taylori, Hook. julacea, L. julacea, Nees juniper ina, Hook. . Juratzk:mta, Lirapr. Kunzeana, Hiiben . laevigata, Dum. Lammersiana, Hiiben lanceolata, L. lunulcpfolia, Dum. . laxifolia. Hook. Leersii, Roth. lurida, Dum. lycopodioides, Wallr. Ljjellii, Hook. Lyoni, Tayl. . Mackaii, Hook. macrorhiza, Dicks. major, Mich. . 7nicroscopica, Tayl. minima, Mich, minuta, Crantz PLATE PAGB VOL. 2 VOL. 1 — 463 143 334 14; 15^ 36 364 312 335 61 307 — 165 — 322 — 269 — 230 — 194 — 264 — 94 — 97 — 99 — 97 151 349 144 131 — 80 — 153 — 284 — 147 — 197 — 106 — 298 (149 346 \150 348 — 432 146 339 — 40 153 'INDEX. 515 PLATE I'AOE VOL. 1 VOL.1 Jun^ei'inania — [GontiaauV) minatissima, Hook. 54 minutlssima, Sm. . Gl montana, Mart. _ 212 Micelleri, Nees ai4 multifida, Hook. . 45G multijida, Linn. 451 multifida, Schmidt 454 nana, Nees . 298 nemorosa, Hook, . - (225 \235 nemorosa, L. . 'Z2'2 nemorosa, Mich. 22-2 nevicensis, Csur. 3.S9 nivalis, Sw. . 97 obovata, Nees 3GG obtusa, Lindh. \M 318 ohtusifolia. Hook. . 241 orcadensis, Hook. . \k)b 357 ornithopodioides, With. . 219 ovata, ]3ickb. -_ 243 jKdmata, Hedw. . — 450 palusiris, Mich. — 284 Pearsoni, Spruce . 156 359 pingiiis, Linn. 457 planifolia, Hook. . 219 platyphylla, L. — 84 pohjanihos, Linn. . — 257 Porella, Dicks. 89 porphyrolenca, Nees 327 pubescens, Schrank — 4G0 jmlcherrima, Hofim. 106 pumihx, Wither. . 12;; 292 pmpurea, 8cop. — 91 jntsilla, Linn. — (418 "(421 quimjuedentata, Web. . 339 Ealfsii, Wils. — 430 reclusa, Tayl. 144. riparia, Tayl. 12i 294 repens, Mich, — 239 reptans, Dum. — 119 reptans, Hook. 117 7'esupiHata, Linn. . — 217 resupinata, Web. et Molir. 2< >8 rosacea, Corda — 233 rupestris, Schleich. 2(1! 1 saxicola, Schrad. . 154 3r»r» scalaris, Mart, 174 scalaris, Schrad, , 370 Schraderi, Ekart . 130 303 PLATK PAGE VOL. 1 VOL. I Juntrermania — {continued} Schraderi, Mart. . Schraderi, Nees scutattts, W. cfe M. Sehlmeyeri, Hiib. . serjryllifolia, Dicks. serpyllifolia. Hook. setacea, Web. setiformis, Ehrh. . siln-etUr, (lott. siuaaia, Dicks, socia, A^ees sphacelata, Gies. spluerocarpn, Ilook. Sphuijui, Difks. Spliaijid, Hook. spinulosa, Dicks. . spinulosa, Hook. . Sprenyelii, Mai't. . Starkii, Herb. Funck. Starkii, Nees stellnlifera, Tayl. MH. stipulacea, Hook. . suhaljiina, Nees subupicalis, Nees . subcompressa, Limpr, sudetica, Nees Tamarisci, L. taxi folia, Wahl. Taylori, Hook. terrestris, Mich. terrestris, Mich. te7-sa, Nees . Thuja, Dicks. tomentella, Ehrh. . triangularis, Schleichei Trichomanis, Dicks. trichophyUa, L. tricreitata, Wahlenb, trilobata, L. . tristis, Nees . turbinata, Raddi . Turneri, Hook. tyroletisis, Nees uUcina, Tayl. uliyinosa, Nees nliyinosa, Sw, umbrosa, Scluad. . undulata, Linn. ustulata, Hiiben. . 125 126 13: 516 HEPATICLE. PLATE PAGE PLATE PAGE VOL. 2 VOL. 1 TOL. 2 VOL.1 Jungervaiiniii~'(contim(sd) Lichenastrum -{continu,edj) ventricosa, Dicks. . ' UO) 141) 327 aqimticum. Dill. asplenii facie, Dill. — 495 274 verruculosa, Lindb. viiiculosa, Linn. — 351 413 arbor is vita' facie, Dill. (82 (84 Wallroihiana, Nees — 309 auricidatum, Dill. . — 217 Weheri, Mart. — 353 f219 J 222 [239 Wilsoni, Tayl. — 410 auriculatuin. Dill. . — Wilsoniana, Nees . — 309 Woodsii, Hook. — 102 cajntulis ohlongis, Dill. — 457 Kantia, Gr. ci' B. — 134 chama'dryos, Dill. . — 456 arguta {Mont. tH- 2\ ees). filicinum. Dill. — 108 Lindb. 53 139 furcatum, Dill. — 461 Sprengelii (Mart.) 52 138 imbricatum. Dill. . — 24 Trichomanis (L.), Gr ci- Ih •. 51 135 minus. Dill. . 33 Lejeunea, Libert calcarea, Lib. 16 39 58 minimum, Dill. — /327 \.418 calyptrifolia {Hook.), Dum. 20 64 multifidum, Dill. . — 119 diversiloba Spruce . 15 56 '128 echinata, Tayl. flava {Sioartz.) Nees 12 58 49 jnnnulis, Dill. . — 224 " 247 Holtii Spruce 18 51 250 hamatifolia {Hook.), Dum 9 43 scorpioides, Dill. . — 104 Mackaii {Hook.), Sj/reng. 7 40 trichoides. Dill. — 112 microscopica, Tayl. 19 63 Trichomanis, Dill. . — 257 minutissima {Sm.) 18 61 LlOCHLyENA, NeCS — 284 minutissima, Spruce — 54 lanceol.ata {L.), Nees 120 284 3IoUeri, Steph. — 42 LOPHOCOLEA, Dum. . — 245 Moorei, Lindb. — 49 bidentata (Z.), Dam. 101 247 ovata, Tayl. . s 42 bidentata, Nees — 249 patens, Lindb. 11 47 cuspidata, Limpr. . 102 249 Rossettiana, Massal. 17 60 heterophylla {Schrad.) 103 250 serpyllifolia {Dicks.), Lib. 10 45 Hookeriana, Nees . — 247 Taylori, Spruce — 61 spicata, Tayl. 104 252 ulicina, Tayl. U 54 Lophozia. Lepidozia, Dum. 116 acuta, Dum. . — 316 cupressina {Sw.) . 43 117 barbata, Dum. — 344 Pearsoni, Spruce . 45 121 bicrenata, Dum. — 324 pinnata, Dum. — 117 exsecta, Dum. -_ 337 reptans {I^.), Dum. 44 119 incisa, Dum. . — 335 setacea ( Web.), Mitt. 46 124 lycopodioides, Cogn. 346 tumidula, Tayl. — 117 Muelleri, Dum. 314 Widfsbergii, Lindb. — 121 scutata, Dum. — 259 Leptoscyphus. ventricosa, Dum. . 327 cuneifolia, Mitt. . — 254 LUNULARIA, Mich. . — 475 interruptus, Mitt. . — 269 cruciata (Z.), Dum. 212 476 Taylori, Mitt. 264 vulgaris, Mich. 476 Lichen jyalustris, Dill. — 485 Madotheca. Lichenastrum. • Thuja (Dicks.) — 82 aljnnitm, Dill. (91 platy])hylla, Dum. . — 84 ' 194 Porella, Nees — 89 INDEX. 517 ri,ATE PAGH VOL. 2 VOL. 1 Madotheca (continued) rividaris, Nees — 87 Madotheca, l)um. — 79 l(tm(/ata, Lindb. — «0 Marchantia, March, fil . — 46 G androcjyna, Tayl. . — 473 commutaia, Lindb. — 473 co7iica, L. . . . — 4g;j cruciata, L. . — 476 hemisphairica, L. . /470 \473 h'riyua, Wils. 477 polymorpha, L. •im 466 Marsilia. e7idivia'folia,{Dickii.), Lindb. — 447 ejn2)h)/Ua, Lindb. . 443 xVeesii, Lindb. 445 pingiiis, Mich. 457 Marsupella, Bum. . 375 conferta (Limpr.), Spruce — 397 emarginata {Ehrh.), Bum. ' j*'"' ' Funckii {Weh. et Mokr.), Dum. . . . 1 6(1 latifolia, Lindb. . . — nevicensis (Carr.), Kaal. olivacea, Spruce revohtta, Lindb. silvrettw, Dum. sparsifolia (Lindb.), Dum. sphacelata (Gies.), Duta. . Stableri, Spruce ustulata (Ililben.), Sprace Martinellia. cequiloha, Lindb. . convexa, Lindb. (jracilis, Lindb. oiemorosa, Gr. «fe B. undalata, Gr. cfe B. Mastiyohryum. J'earsoni, Stephani trilobatum, Nees . Mastkjophora, Nees . reptans, Nees Woodsii (Hook.), yees . Maurocenia. pasilla, Gr. & Ben. J71 107 169 165 170 I6.S orcadensis, Dum. comp'essa, Dum. scalaris, Dum. 375 I 379 389 389 380 393 372 384 377 386 382 •212 236 217 222 224 133 128 102 119 102 418 357 368 370 Metz(ieria. Ji'addi . conjugata, Lindb. . furcata (L.), Eaddi glabra, Dum. liamata, Lindb. linearis, Lindl). pubescens (Schrank) Mnium. Jissum, L. jungermania, L. 7Vicho7uanis (T>i\\.), L. Moerckia. Bhjttii, Gotts. hibernica, Gotts. . norregica, Gotts. . Muscoides. terrestre, Mich. terrestris, Mich. Mylia, Gr. (C- B. anoniala (Hook). G'r. d- B Taylori (Hook.)', Gr. tO B cuneifolia, Gr. & B. Nardia, Gr d: Ben. . Carringtoni, Balf. . comp-essa, Lindb. . compressa (Hook.), Gr. d Benn. geoscypha, Mass. . hcematosticta, Lindb. hyalina (Lyell), Carr. latifolia, Lindb. obovata (Nees), Carr. revoluta, Lindb. scalaris (Schrad.), Gr. ( Benn. silvrettae, Gotts. sparsifolia, Lindo. sphacelata (Gies.), Carr. Nowellia, Mitt. curvifolia, Mitt. . Odontochisma, Dum. . decipiens, Lindb. . denudatum, Dum. . Sphagni, Dum. Pallavicinia, Gr. d- Benn Blyttii (Moerck), Lindb. hibernica (/look.), Gr. d Benn. Lyii[Vn(/Iuok.),Gr.d-Be7in. PLATE PAGE VOL. 2 VOL. 1 — 459 463 461 461 464 464 460 207 206 207 110 109 J 60 158 159 161 162 (194) (1951 (192) 1193f 191 ] 35 91 135 438 434 438 119 128 263 267 264 254 363 2S7 287 368 372 372 364 389 366 393 370 372 384 377 141 159 (in 1^170 204 174 171 432 438 434 432 518 HEPATIC^. Pedinophyllum, Lindb. . interrviptum {Nees), Lindb . •pi/renaicitm, Lindb. Pellia, Raddi . calycina (Tai/l.) epiphylla (/>.), Lindb. Neesiana {Gotts.),Li7np Xeesii, Limpr. Petalophyllum, GoU. lamellatum, Lindb. Ralfsii (TF^7s.), Gott. Physiotium, Nees cochlear i forme, Nees Plagiochila, G. L. X. fiosa, 3RU. . asplenioides (Z.), Dwm. Bartlingii, M. et N. compacta, M. et N. corniculata, Dum. . carta, M. et N. decipiens, Dum. exigua, Tayl. interrupta, Dum. . pijrenaica, Lindb. . punctata, Tayl. rosacea, M. et JST. . spinulosa [Dicks.), Dum. Stableri, Dear son . tridenticulata, 7\tyl. tyrolensis, M. et N. uliginosa, M. et N. umbrosa, Mont. Dleuranthe. olivacea, Tayl. Dleiiroschisv la , Dum. demtdatum, Dum. . Donniana, Dum. . reptans, Dum. Sjihagni, Dum. trilobatum, Dum. . Pleuroolada, Spruce albescens {Hook.), Sjyruce albescens. Spruce islandica {Xees) Pleurozia, Dum. cochleariformis {Wei Dum. purpurea, Lindb. . PLATE PAGE VOL. 2 VOL. 1 — 269 269 442 447 443 445 445 421) 430 430 90 91 274 275 274 209 208 280 235 204 282 269 269 278 233 276 273 280 212 231 236 261 141 174 361 119 171 128 192 192 194 194 90 91 91 11 198 197 190 115 113) 114) 119 117 116 112 118 76 PLATE PAGE VOL. 2 VOL. 1 Porella {Dill^i . 79 Levigata {Schrad.), Lindb. 28 80 pinnata {Dill.), Lindb. . 32 89 platyphylla (Z.), Lindb. :'.0 84 rivularis {Xees), Lindb. . 31 87 Thuja {Dicks.), Moore . 29 >^i Preissia, Corda ATI commutata {Lindb.), Xees 211 473 hemisphwrica, Cogn. 473 Prionolobus, Hp>ruce . 189 Turneri {Hook.), Spruce . 74 190 Ptilidium. ciliare, Hamp. 104 Eadula, Dum. . W} a'qidloba, Dum. 212 aquilegia, Tayl. 25 74 aquilegia, Tayl. 76 asjilenioides, Dum. 274 Carringtonii, Jack 2(') 76 complanata (L.) 68 Gomplanata (L.) - — 70 complanata (//.), Dum. . 27 78 commutata, Gottsche — 68 cornicidata, Dum. . 280 carta, Dum. . — 235 decipiens, Dum. — 204 dentata, Dum. ___ 217 germana, Jack 23 70 Holtii, Spruce 24 72 Lindenbergiana, Gottsche 68 Lindbergii, Gottsche 22 68 nemorosa, Dum. — 222 resupinata, Dum. . — 208 uligi7iosa, Dum. . — 231 umbrosa, Dum. 236 undulata, Dum. 224 voluta, Tayl. . 21 67 Reboulia, Raddi 470 hemisphffirica, Rad. 210 470 Riccardia. latifrons, Lindb. — /454 1456 multifida, Gr. &, Benn. . — f451 \454 multifida (Nees), Lindb. — 453 midtifida, Lindb. . — 456 palmata, Carruth. . — 450 Riccardius. jnnguis, Gr. & Benn. — 457 RicciA, Mich. . — 483 bifurca, Hoffm. 219 488 TNBEX. 519 Riccia — (continuedC) bifurca, Hofim. ciliata, Iloffm. crystallina, L. Jluitans, h. . glauca, L. glaucesceiis, (Jarr. . lamellosa, Ralf.s. Michelii, Raddi minima, L. Schift". . natans, L. nigrella, B.C. sorocarpa, Bisch. . tumida, Lindevh. . RicciELLA, Braun. . fliiitans (L.), Brann. RicciocARPUS, Corda natans (Z.), Corda Saccogyna, Dum. viticulosa {Linn.), Dum Sa7'cosc)/phits. alphms, Gottsch. . anomalus, Jack capillaris, Limpr. . confertus, Limpr. . Ehrharti, Corda Funckii, Nees revolutus, Nees sparsifolius, Lindb. sphacelatus, Nees . ScALiA, Gr. (t- Benn. . Hookeri(Zye/^), Gr. c(-Bej ScAPANiA, Dam. JBquiloba (jSchway.), Dum. aspera, Bern. Bartlingii {Ilampe), JS Carestice, De. Not. compacta (Roth.), Ban convexa. Scop. carta, Carr. . curta {Mart.), Bam. dentata, Dum. intermedia, Hasnot irrigua {Xees), Dum. PLATE PAGE VOL. 2 VOL. 1 _ 49?, '. 22)\ 494 . 217 48:5 49:) . 2](; 484 222 493 489 491 487 497 220 489 . 218 487 . 221 491 . 495 224 495 — . 497 . 225 497 413 .* 182 413 „ 391 372 389 — 397 375 379 — 393 384 - — 377 -- 427 in. 180 427 207 m. 80 212 . .S4 214 es HI 209 20!) 80 208 . 236 233 . 95 235 217 . !)! 227 1)2 230 •OL. 2 88 H7 86 PAGE VOL.1 90 85 ;)4 82 96 93 89 tScapania — {continued) nemorosa (A.), J) am. nemorosa, Husnot . nimbosa, I'ayl. ornithopodioide.s {Dill, et With.). Peai-.s. . planifolia (Hook.), l^um. purpurascens {Hook.), Tayl MS ' re.supinata (//.), Dam. . rosacea {(.'orda), J) am. . subalpina {Nees), Dum. . ti/rolensis, G.L.N. . umbrosa (Schrad.), Bum. uliginosa {Sirartz.), Dum. undulata (//,), Dum. Schisma, Dum. . . • . adunca, Dum. Sendtnera, Nees adunca, Carr. juiiiperi'aa, Nees . Woodsii, G.L.N. Southbya. obovata, Dum. Sph^rocarpus, Mich. Michelii, Bell. terrestris, Mich. Sphaynoecetis, Nees . comm^inis, Nees Hilbtieriana, Rabenli Sphagni, Nees Bteetzia. L)/ellii,Jjehm. Targioxia, Mich. hypophylla {L.) Michelii, Corda . . — spharocarpiis, Dicks. . — Thedenia. Blyttii, Hartm. . . — Trkciiocolea {Dam.), Xees — tomentella {Ehrh.), Bum. 40 Tricholea. tomentella. Dum. . 215 214 222 227 220 219 219 225 217 233 211 2V2 236 231 224 99 99 99 99 99 102 366 481 482 482 ri4i \170 174 174 171 432 479 480 480 482 438 108 108 — 108 520 HEP A TIC.E. ERRATA. P, 11, col. 2, line 16, /or pistillidia read antheridia. P. 11, col. 2, line iW,for male read female. P. 24, line 22, for Monoicous read Dioicous, P. 45, line 29, /or Dioicous read Monoicous. P. 102, bottom line, /or smaller read larger. P. 157, line 18, /or flagelliferous read eflagelliferous. P. 214, line 24, delete Miiller. P. 221, line 18, /or smooth read minutely papillose. P. 339, above Jungermania Lyoni insert — Section 2. BARBATAE, Schiffti. Plants usually large. Leaves 3-5-fid. Stipules present, bifid, ciliate. Perianth oval, plicate almost to the base. P. 351, above Jungermania Helleriana insert — Section 3. P. 384, line 11, for sparsifolia read sparsifolius. P. 413, line 2 from the bottom, p. 414, lines 17 and 18, p. 4ir>, lines 1 and for perianth read, perigynium. Plate cxviii. for trideniculata read tridenticulata. Printed by Ballantvne, Hanson Cs^ Co. London <2^ Edinburgh