diomed uK 495 C24 671n 1889 THE LIBRARY OF THE UNIVERSITY OF CALIFORNIA LOS ANGELES NOTES ON THE PINKS OF WESTERN EUROPE BY FREDERIC N. WILLIAMS, F.L.S. Plants as wholes assume unlike attitudes towards their environments ; they have many ways of articulating their parts with one another ; tliey have many ways of adjusting their parts towards surrounding agencies. These are causes of special differentiations additional to tlwse general differentiations that result from increase of mass and increase of composition There is a parallel cause for tlie parallel differentiations exhibited by all aggregates, HEEBEET SPENCEK. LONDON : PRINTED BY WEST, NEWMAN AND CO., HATTON GARDEN, E.G. 1889. INTRODUCTION. THE grouping of the species in these 'Notes' is in accordance with that followed out in the author's ' Enumeratio Specierum Varietatumque Generis Dianthus.' Original references are given for species and varieties not mentioned in Nyman's ' Conspectus,' or .where his nomenclature is not followed. The matter relating to each species is arranged under the following heads : (1) A Latin diagnosis of the species, drawn up after examining and comparing original specimens with the best known descriptions, in which the chief essential and differential rather than the absolute specific characters are given ; (2) the countries of Western Europe (as included in the area under consideration) in which the species is igeiwus, with the vernacular names of the plant (if it has any) in these countries, and in which the following initials indicate the v countries named : jT S" E. = Great Britain. G. = Germany (small part of), i F. = France (most of). S. = Spain (Castilian, Catalan, ^ H. = Holland (and Luxemburg). and the Basque dialect). i B. = Belgium (Flemish). P. = Portugal. X s (3) the general geographical limits of the species; (4) pre-Linnean j references ; (5) derivation of the specific name ; (6) notes and jj remarks on the plant and its varieties. The measurements, in ^ centimetres and millimetres, refer to average specimens and to the 5 average length of their parts. Some justification may perhaps be. ^ needed of the ex cathedra manner in which these pages are written, ^ in the prevailing absence of synonymy, and references to authori- ^ ties, &c. ; were it not that they are rather a digest of results obtained by research, and not merely a compilation. It has not been^ thought desirable to burden the text with innumerable 4O1415 IV INTRODUCTION. synonyms, which are often no more than the record of the unnecessary labours, as well as the mistakes, of previous observers ; and which, in some systematic works, tend to reduce much of the text to a dry and unreadable index. BRENTFORD, September, 1889. NOTES ON THE PINKS OF WESTERN EUROPE. THE genus Dianthus was founded by Linnaeus, and many of his species are described in the first edition of the ' Species Plantarum,' published in 1753. The name is a most appropriate one, and is one of Linnaeus' happiest selections in his generic titles. It is derived from dios, anthos, the divine flower, in allusion to the singular beauty and fragrance of some of the species. A second derivation is di, anthos (double flower) ; but this is only a concession to the prejudice of horticulturists, who have degraded so many of the "divine flowers" of Nature to the rank of emasculated garden monstrosities. Caryophyllus was the pseudogeneric term used for many of the species described by the pre-Linnean botanists, Ruellius (' De natura stirpium,' 1536), Dodoens (1554), Tabernsemontanus (1588), J. Bauhin (1591), Grisley (1661), Cupani (1696), Tournefort (1694), and Seguier (' Plantse Veronenses,' 1745-1754). The limits of the geographical range of the genus are : N. On the coast of the Kara Sea, in Arctic Russia, lat. 70 N. E. Alaska, in N.W. America, long. 160 W. S. Cape of Good Hope, lat. 34 S. W. Corunna, at N.W. extremity of Spain, long. 8 W. In Europe the genus is distributed throughout its whole extent, but is absent, however, from Iceland and Ireland ; its extreme limits being : N. On the coast of the Kara Sea, in Arctic Russia, lat. 70 N. D. alpinus. E. Baku, in Russian Armenia, lat. 50 E. D. prolifer. S. The island of Crete, lat. 35 S. D. sphadoticus. W. Corunna, at N.W. extremity of Spain, long. 8 W. D. mons- pessulanus. The area of "Western Europe" to which these notes refer includes The United Kingdom, Holland, Belgium, France, ex- cluding that part E. of the Rhone (.Dauphine, Provence, and Savoy), the part of the German Empire W. of the Rhine, Portugal, and Spain. The only special work bearing upon the subject is ' Observa- tions sur les Di. attenuatus, as the authors of the ' Flore de France ' have done. It is somewhat variable in the comparative size of its different parts, in the relative length of the leaves, stems, shoots, and branches of the rootstock. On the dry calcareous soil of the southern part of Saragossa is found a dwarfed form with small flowers. 21. D. csesius Sin. Caespitosus, glaber, glaucus. Caules 22 centim., simplices v. superne furcati strict! 4-angulares. Folia 3-nervia patentia, radicalia 50 mm. obtusa plana, caulinia 36 mm. acuta carinata, vagina folii diam. sequante. Flores speciosi odorati. Bractese 4 obovatae mucronatao. Calyx dentibus ovato-lanceolatis acutis. Petala contigua, lamina obovato-cuneata, = unguis. E. (Cheddar or Mountain P.). F. ((E. bleuatre). G. (Graug- riine N.). B. (Blauwachtige A.). NOTES ON THE PINKS OP WESTERN EUROPE. 23 Area of distribution, 96. N. Somerset, 51. 8. Dauphiny, 45. E. Pilsen, in Bohemia, 13. W. Somerset, 3. Armeriae species flore in summo caule singulari (Ray, 1670). Tunica rupestris folio caasio molli flore carneo (Dillenius, 1732). Ray seems to base his name on the unbranched unifloral stems ; Dillenius selects its glaucous habit, the textur.e of its leaves divergent from the stem, the colour of the flowers, and its rocky mountain station. Referred to by Hudson as " D. glaucus " (1762). This is par excellence the English Pink. It is not a Linnean species, as stated in Sowerby's ' English Botany ' (3rd edition), but was described by Smith in his < English Botany,' tab. 62 (1792), and also referred to two years later (1794) in his Synopsis of the genus in the ' Transactions of the Linnean Society.' The specific name CCBSIUS refers to the grey-green appearance produced by the deposit of bloom on the leaves, and is used in the same sense as "glaucus" and "pruinosus" of many authors, though the last would rather indicate a more frosted, heavier deposit of white bloom. Its geographical distribution is extremely limited, and the only British locality for the plant is the Cheddar Cliffs, consisting of abrupt limestone precipices in Somerset. This solitary station should therefore be jealously guarded, being, as it is, the northern as well as the western limit of the plant in Europe. It is not very plentiful, and fortunately not very accessible, even at this one station, where it grows at a height of 50 ft. among jagged rocks ; and unless visitors forbear to take away specimens, it will soon become extinct as a native British plant. If they require mementoes, they might wreak their cacoethes prehendi on Diantkus deltoides, which grows hard by. In the east of France and Switzerland it is essentially a mountain pink, and reaches an altitude of 1500-1800 metres : often found in association with Kaxifraya aizoides, S. hypnoides, and Cerastium alpinum. When it is found at a lower elevation, it is from the seeds being washed down by rains, &c. In North Italy it is called "garofano appannato." The rootstock has long, slender divisions, producing ascending, short, barren shoots, which give the plant a caespitose habit, and many erect or ascending unbranched flowering stems ; the bracts are membranous, with an herbaceous apex. The flowers are 1 in. in diameter, with the teeth of the petals one-eighth to one-sixth the length of the lamina, and, as Smith says, " delightfully fragrant, of a delicate, pale rose-colour." It may be noticed that the highest pair of leaves are adpresscd to the stem, and much shorter, thus approaching the bracts in character. Subsectio 2. HEMISYRHIX. Bractese atting. % calycis longi- tudinem. 22. D. deltoides L. Caespitosus, pubescens. Caules 25 centim. furcati tenues. Folia plana patentia flaccida 3-nervia, 24 NOTES ON THE PINKS OF WESTERN EUROPE. radicalia oblanceolata obtusa, caulinia lineari-lanceolata acuta, suprema squamiformia fere scariosa, vagina folii diam. gequante. Flores breviter pedunculati odorati. Bracteae 2 (rarius 4) ovatse acuminatse. Calyx gracilis, dentibus lanceolatis acumi- natis. Petala non contigua, lamina obovata rosea v. purpu- rascente, = % unguis. var. glaucus L. var. microlepis Pet. E. (Maiden P.). F. (OB. deltoide). H. B. (Drylioekigc A.). G. (Delta-blumige N.). S. (Clavellina deltoidea). P. Area of distribution, 6027. N. Norway, 64. E. Japan, 142. S. Bengal, 23. W. Scotland (Inverness), 5. Pre-Linnean references. Earliest record in 1581, in which year Lobel published his ' Kruydtboek,' and gave a drawing of it : not mentioned by Dodoens, either in the ' Kruydeboeck ' or 'Pemptades.' It is referred to again, sixteen years later, in Gerarde's ' Herball,' where he says : " There is a wild creeping pink which groweth in our pastures near about London and in other places, but especially in the great field next to Deptford by the path side as you go from Redriffe to Greenwich, which hath many small tender leaves shorter than any other of the wild pinks, set upon little tender stalks, which lie flat upon the ground, taking hold of the same in sundry places, whereby it greatly increaseth, whereupon grow little reddish flowers. The root is small, tough, and long lasting." Armeria sylvestris (Lobel, 1581). Caryophyllus simplex supinus latifolius (Caspar Bauhin, 159G). Caryophyllus sylvestris (Clusius, 1598). Caryophyllus minor repens nostras (Hay, 1670). Armerise species flore summo caule singular! (Petiver, 1702). Tunica ramosior (Dillenius, 1732). Other pre-Linnean botanists also refer to the plant by one of these names. The species is so named from the resemblance of the marking of the lamina of the petals to the Greek letter delta. An old English name for it is " Sop-in-Wine." It has a very wide area of distribution. The most northern record for the plant is Levanger, on the Trondhjem fiord, in Norway, lat. 63 52', vide A. Blytt's ' Norges Flora,' p. 1073. The southern limit is Bengal. Franchet and Savatier record its eastern limit in Japan, in whose Flora it had also been previously described by Siebold and Miquel. The western limit is in the west of Scotland : in Watson's herbarium at Kew there is a specimen (dated 1833) from Cromwell's Mount, Campbelltown, in Inverness. It grows in all the countries of Europe, with the exception of Turkey and Greece ; though it is not recorded from Sardinia, or from Ireland. This pretty pink is found on grassy slopes, road-sides, and dry pastures : it flourishes on soils of primitive or volcanic origin, e. 1570; Gerarde, 1597; Grisley, 1661). Betonica coronaria squammosa sylvestris (Jean Bauhin, 1591). Caryophyllus sylvestris prolifer (Caspar Bauhin, 1596 ; Zanni- chelli, 1712). Caryophyllus floribus capitatis aggregatis, squamis calycinis ovatis obtusis muticis tubum superantibus (J. G. Gmelin, 1751). Caryophyllus aridus (Moench, 1794). Tunica prolifera (Scopoli, 1760). Pre-Linnean references. First record as a British plant (in Middlesex) by How in 1650, who calls it "Childing Sweet- william." Mentioned in Lobel's ' Kruydtboeck,' Gerarde's ' Herball," Grisley's ' Viridarium,' C. Bauhin's ' Theatrum Botanicum,' J. Bau- hin's ' Hist, universalis Plantarum,' Gmelin's ' Flora Sibirica,' Seguier's ' Plant Veronenses,' and Wulff's ' Flora Borussica.' Called "strigoli" by the early Italian botanists. Introduced by the colonists, and imperfectly naturalized in North America. As to its specific name, Asa Gray defines prolifer, as bearing progeny in the way of offshoots. 46 NOTES ON THE PINKS OF WESTERN EUROPE. In the ' Flora Berolinensis,' the genus " Kohlrauschia " wag invented for this species, and the following points of generic distri- bution were relied upon by Kunth for its separation from the Linnean genus : Flores capitato-congesti rarissime solitarii sessiles involucre universal! scarioso cincti. Bracteolse 2 scariosae convolutse. Calyx pentagono-tubulosus 15-costatus, inter angulos fere omnino scarioso-membranaceus, apice attenuate. Corolla caryophyllacea ; petala retusa lamina suberecta ungue longa lineolataque. Capsula maturescens lateraliter fissus usque ad basin, oblongo-fusiforinis. Seguier's figure wrongly depicts 'the bracts as acute ; in the living specimens they are most distinctly mucronate, like the bracteiform leaves, which usually inclose the flower-head as an involucre con- sisting of three pairs. There is no perennial rootstock, or barren leafy shoots ; and each stem or fork terminates in a small ovate- fusiform head of flowers. Dr. Boswell-Syme, in the 3rd edition of the 'English Botany,' says that in all the specimens he has had an opportunity of dissecting, he has been able to find only one bract to the calyx, but they may not have been perfect specimens. The heads of flowers are about 18 mm. long, and the average individual flowers about 9 mm. in diam. The calyx is very narrow, and faintly ribbed. In England it grows in sandy places and gravelly pastures in four counties only Berks, Hants, Sussex, Norfolk, and in Jersey. In France it occurs on the light sandy soil of road-sides and dry places, and on the limestone along the Doubs and in the Vosges. In Spain, Boissier found specimens at an elevation of 600-1600 metres. The authority for the northern limit of the species is that of the more recent German Floras ; for the southern limit, the ' Catalogue of the Native Plants of the Canary Isles'; for the western limit, in the 'Botany of the Voyage of the Herald' (1852); and for the eastern limit, E. F. Hohenacker in 'Enum. pi. in it. per prov. Taliisch' (1838). In his ' Florule du Tarn' (1864), Martrin-Donos notes a local form, scarcely of the nature of a variety, in which the stems are very slender, and forked almost from the base with emarginate or nearly entire petals ; growing on sandy banks, and also on old walls. 55. D. velutinus Guss. Velutinus. Caules simplices strict! graciles, supra angulares, infra teretes. Folia plana, radicalia lanceolato-linearia recurva, caulinia linearia stricta, sursum decrescentes, vagina folii diam. duplo longiore : phylla involucri epunctulata mucronulata ad florum apices. Iii- florescentia capitulis 5 8 florum roseorum pedicellatorum inodororum. Bractese phyllis similes. Calyx dentibus ob- ovatis obtusis. Petala contigua, lamina barbulata, = unguis. Semina tuberculata. F. ((E. veloute). S. P. NOTES ON THE PINKS OF WESTERN EUROPE. 47 Geographical limits, 660. N. Crimea, 46. E. Armenia, 47. 8. Algeria, 84. W. N. Portugal, 8. First described by Gussone in 1825, and so named from the glandular hairs with which the stem is covered. Often confused with the preceding species, from which, however, it is very distinct. Besides the general differences which are specified above, may be mentioned the greater proportionate breadth of the radical leaves to the upper ones of the stem, the laxer inflorescence, and the almost bifid character of the petals. It grows on the dry pastures of mountains, often at high altitudes, e.g., on Sierra Nevada, near Trevolez, at 1650 metres. In the ' Flore de France' (1848), it is only recorded from Corsica, but in the Essay of Timbal-Lagrave it is mentioned as occurring in the depts. of Pyrenees Orientales and Aude. RATIO OF THE SPECIES OF WESTERN EUROPE TO THE WHOLE NUMBER OF SPECIES KNOWN: 55 species, i. e., 24 per cent. Spain. 43 species, i. e., 18'61 per cent. France. 33 species, i. e., 14'29 per cent. Portugal. 13 species, i. e., 5*6 per cent. Germany. 7 species, i.e., 3 per cent. Belgium. 5 species, i. e., 2*2 per cent. Holland. 5 species, i. e., 2'2 per cent. England. 4 species, i. e., T731 per cent. ERRATUM, On p. 22, line 11, for 'rosea infra pallida," read "rosei infra pallidl,' UNIVERSITY OF CALIFORNIA LIBRARY Los Angeles This book is DUE on the last date stamped below. Form L9-25m-9,'55(B4283s4)444 UNIVERSITY OF CALIFOR1S AT A 000467597 1