Digitized by the Internet Archive in 2017 with funding from University of Illinois Urbana-Champaign Alternates https://archive.org/details/historymethodsof00mars_0 '■'M 1 HISTORY AND METHODS PALiEOMOLOGICAL DISCOVERY. AlSf ADDRESS, DELIVERED BEFORE THE AMERICAN ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE, AT SARATOGA, N. Y., AUGUST 28, 1879, Professor 0. C. MARSH, President, 12320 MAHSH (Othlel C.) History and Methods of Palaeontological Discovery. An Address delivered before the American Association for the Advancement of Science at Saratoga, N.Y. August 28, 1873* Dp. 50# &oy# 8vo, orig. wrappers. [N.p.d.] Presentation copy from the author. Scholarly resum£ of the whole history of palaeontology from Xenophanes of Colophon [500 B.C.] to modern times. HISTORY AND METHODS OP PALEONTOLOGICAL DISCOVERY. AE ADDRESS, DELIVERED BEFORE THE AMERICAN ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE, AT SARATOGA, N. Y., AUGUST 28, 1879, BY Professor 0. C. MARSH, President, 9J?an mag tie lebenbtgen SBtrfungen ber 9?atur im ©anjen unb ©ro£en uberfefyen, ober man mag bie UeberMeibfel ifjrer entflo^enen ©eifter jergltebern: fte bleibt immer gletd), immer rneljr bemunberungSmurbtg.—© o e tl) e. . hM HISTORY AND METHODS OF PALEONTOLOGICAL DISCOVERY. “S Co N 55 h I In the rapid progress of knowledge, we are constantly brought face to face with the question, What is Life ? The answer is not yet, hut a thousand earnest seekers after truth seem to be slowly approaching a solution. This question gives a new interest to every department of science that relates to life in any form, and the history of life offers a most sug¬ gestive held for research. One line of investigation lies through embryology, and here the advance is most encourag¬ ing. Another promising path leads back through the life history of the globe, and in this direction we may hope for increasing light, as a reward for patient work. The plants and animals now living on the earth interest alike the savage and the savant, and hence have been carefully observed in every age of human history. The life of the remote past, however, is preserved only in scanty records, buried in the earth, and therefore readily escapes attention. For these reasons, the study of ancient life is one of the latest of modern sciences, and among the most difficult. In view of the great advances which this department of knowledge has made within the last decade, especially in this country, I have thought it fitting to the present occasion to review briefly its development, and have chosen for my subject this even¬ ing, The History and Methods of Palaeontological Discovery. 4 In the short time now at my command, I can only attempt to present a rapid sketch of the principal steps in the progress of this science. The literature of the subject, especially in connection with the discussions it provoked, is voluminous, and an outline of the history itself must suffice for my present purpose. In looking over the records of Palaeontology, its history may conveniently be divided into four periods, well marked by prominent features, but, like all stages of intellectual growth, without definite boundaries. The first period, dating back to the time when men first noticed fossil remains in the rocks, and queried as to their nature, is of special historic interest. The most prominent characteristic of this period was, a long and bitter contest as to the nature of fossil remains. Were they mere “ sports of Hature,” or had they once been endowed with life ? Simple as this problem now seems, centuries passed before the wise men of that time were agreed upon its solution. Sea shells in the solid rock on the tops of mountains early attracted the attention of the ancients, and the learned men among them seem to have appreciated in some instances their true character, and given rational explanations of their presence. The philosopher Zenophanes, of Colophon, who lived about 500 B. C., mentions the remains of fishes and other animals in the stone quarries near Syracuse; the impression of an anchovy in the rock of Paros, and various marine fossils at other places. His conclusion from these facts was, that the surface of the earth had once been in a soft condition at the bottom of the sea; and thus the objects mentioned were entombed. Herodotus, half a century later, speaks of marine shells on the hills of Egypt, and over the Libyan desert, and he inferred therefrom that the sea had once covered that whole region. Empedocles, of Agrigentum (450 B. C.), believed 5 40 that the many hippopotamus bones found in Sicily were remains of human giants, in comparison with which the pres¬ ent race were as children. Here, he thought, was a battle held between the gods and the Titans, and the bones belonged to the slain. Pythagoras (582 B. C.) had already anticipated one conclusion of modern geology, if the following statement, attributed to him by Ovid, was his own :* Vidi ego quod fuerat solidissima tellus, Esse fretum : vidi factas ex sequore terras ; Et procul a pelago conchae jacuere marinse. Aristotle (384-322 B. C.) was not only aware of the exist¬ ence of fossils in the rocks, but has also placed on record saga¬ cious views as to the changes in the earth’s surface necessary to account for them. In the second book of his Meteorics, he says : “ The changes of the earth are so slow in comparison to the duration of our lives, that they are overlooked ; and the migrations of people after great catastrophes and their removal to other regions, cause the event to be forgotten.” Again, in the same work, he says: “As time never fails, and the universe is eternal, neither the Tanais, nor the Nile, can have flowed for ever. The places where they rise were once dry, and there is a limit to their operations : but there is none to time. So of all other rivers ; they spring up, and they perish ; and the sea also continually deserts some lands and invades others. The same tracts, therefore, of the earth are not, some always sea, and others always continents, but everything changes in the course of time.” Aristotle’s views on the subject of spontaneous generation were less sound, and his doctrines on this subject exerted a powerful influence for the succeeding twenty centuries. In the long discussion that followed concerning the nature of fossil remains, Aristotle’s views were paramount. He believed that animals could originate from moist earth or the slime of rivers, and this seemed to the people of that period a much * Metamorphoses, Liber XV, 262. 6 simpler way of accounting for the remains of animals in the rocks than the marvelous changes of sea and land otherwise required to explain their presence. Aristotle’s opinion was in accordance with the Biblical account of the creation of Man out of the dust of the earth, and hence more readily obtained credence. Theophrastus, a pupil of Aristotle, alludes to fossil fishes found near Heraclea, in Pontus, and in Paphlagonia, and says : “ They were either developed from fish spawn left behind in the earth, or gone astray from rivers or the sea into cavities of the earth, where they had become petrified.” In treating of fossil ivory and bones, the same writer supposed them to be produced by a certain plastic virtue latent in the earth. To this same cause, as we shall see, many later authors attributed the origin of all fossil remains. Previous to this, Anaximander, the Miletian philosopher, who was born about 610 years before Christ, had expressed essentially the same view. According to both Plutarch and Censorinus, Anaximander taught that fishes, or animals very like fishes, sprang from heated water and earth, and from these animals came the human race; a statement which can hardly be considered as anticipating the modern idea of evolution, as some authors have imagined. The Bomans added but little to the knowledge possessed by the Greeks in regard to fossil remains. Pliny (23-79 A. H), however, seems to have examined such objects with interest, and in his renowned work on Natural History gave names to several forms. He doubtless borrowed largely from Theo¬ phrastus, who wrote about three hundred years before. Among the objects named by Pliny were, “ Bucardia , like to an ox’s heart“ Brontia , resembling the head of a tortoise, supposed to fall in thunder storms;” “ Glossojotra, similar to a human tongue, which does not grow in the earth, but falls from heaven while the moon is eclipsed “ the Horn of Ammon , possessing, with a golden color, the figure of a ram’s horn e 7 Ceraunia and Ombria , supposed to be thunderbolts; Ostra- cites , resembling the oyster shell; Sjpongites , having the form of sponge ; Phycites , similar to sea-weed or rushes. He also mentions stones resembling the teeth of hippopotamus; and says that Theophrastus speaks of fossil ivory, both black and white, of bones born in the earth, and of stones bearing the figure of bones. Tertullian (160 A. D.) mentions instances of the remains of sea animals on the mountains, far from the sea, but uses them as a proof of the general deluge recorded in Scripture. During the next thirteen or fourteen centuries, fossil remains of animals and plants seemed to have attracted so little atten¬ tion, that few references are made to them by the writers of this period. During these ages of darkness, all departments of knowledge suffered alike, and feeble repetitions of ideas de¬ rived from the ancients seem to have been about the only contributions of that period to Natural Science. Albert the Great (1205-1280 A. D.), the most learned man of his time, mentions that a branch of a tree was found, on which was a bird’s nest containing birds, the whole being solid stone. He accounted for this strange phenomenon by the ms formativa of Aristotle, an occult force, which, according to the prevalent notions of the time, was capable of forming most of the extraordinary objects discovered in the earth. Alexander db Alexandro , of Naples, states that he saw, in the mountains of Calabria, a considerable distance from the sea, a variegated hard marble, in which many sea shells but little changed were heaped, forming one mass with the marble. With the beginning of the sixteenth century, a great impetus was given to the investigation of organic fossils, especially in Italy, where this study really began. The discovery of fossil shells, which abound in this region, now attracted great atten¬ tion, and a fierce discussion soon arose as to the true nature of these and other remains. The ideas of Aristotle in regard to 8 spontaneous generation, and especially his view of the hidden forces of the earth, which he claimed had power to produce such remains, now for the first time were seriously ques¬ tioned, although it was not till nearly two centuries later that these doctrines lost their dominant influence. Leonardo da Vinci, the renowned painter and philosopher, who was horn in 1452, strongly opposed the commonly accepted opinions as to the origin of organized fossils. He claimed that the fossil shells under discussion were what they seemed, and had once lived at the bottom of the sea. “You tell me,” he says, “ that Nature and the influence of the stars have formed these shells in the mountains; then show me a place in the mountains where the stars at the present day make shelly forms of different ages, and of different species in the same place.” Again, he says, “In what manner can such a cause account for the petrifactions in the same place of various leaves, sea¬ weeds, and marine crabs ?” In 1517, excavations in the vicinity of Verona brought to light many curious petrifactions, which led to much specu¬ lation as to their nature and origin. Among the various authors who wrote on this subject was Fracastoro, who declared that the fossils once belonged to living animals, which had lived and multiplied where found. He ridiculed the prevailing ideas that the plastic force of the ancients could fashion stones into organic forms. Some writers claimed that these shells had been left by Noah’s flood, hut to this idea Fracastoro offered a mass of evidence, which would now seem conclusive, hut which then only aroused hitter hostility. That inundation, he said, was too transient; it consisted mainly of fresh water ; and if it had transported shells to great distances, must have scattered them over the surface, not buried them in the interior of mountains. Conrad Gesner (1516-1565), whose history of animals has been considered the basis of modern zoology, published at Zurich in 1565 a small hut important work entitled u De omni rerum fossilium genere .” It contained a catalogue of the 9 collection of fossils made by John Kentmann. This is the oldest catalogue of fossils witli which I am acquainted. George Agricola (1494-1555) was, according to Cuvier, the first mineralogist who appeared after the revival of learning in Europe. In his great work, “De lie Metallica ,” published in 1546, he mentions various fossil remains, and says they were produced by a certain “ materia jpinguis ,” or fatty matter, set in fermentation by heat. Some years later, Bauhin published a descriptive catalogue of the fossils he had collected in the neighborhood of Boll, in Wurtemberg.* Andrew Mattioli, a distinguished botanist, adopted Agricola’s notion as to the origin of organized fossils, but admitted that shells and bones might be turned into stone by being permeated by a “lapidifying juice.” Ealloppio, the eminent professor of anatomy at Padua, believed that fossil shells were generated by fermentation where they were found ; and that the tusks of elephants, dug up near Apulia, were merely earthy concretions. Mercati, in 1574, published figures of the fossil shells preserved in the Museum of the Vatican, but expressed the opinion that they were only stones, that owed their peculiar shapes to the heavenly bodies. Olivi, of Cremona, described the fossils in the Museum at Yerona, and considered them all “ sports of nature.” Palissy, a French author, in 1580, opposed these views, and is said to have been the first to assert in Paris that fossil shells and fishes had once belonged to marine animals. Fabio Colonna appears to have first pointed out that some of the fossil shells found in Italy w T ere marine, and some terrestrial. Another peculiar theory discussed in the sixteenth century deserves mention. This was the vegetation theory, especially advocated by Tournefort and Camerarius, both eminent as botanists. These writers believed that the seeds of minerals and fossils were diffused throughout the sea and the earth, and were developed into their peculiar forms by the regular incre¬ ment of their particles, similar to the formation of crystals. * Historia novi et admirabilis Fontis Balneique Bollensis , in Ducaiu Wirtem- bergico. Montbeliard , 1598. 9 “ How could the Cornu Ammonis ,” Tournefort asked, “ which is constantly in the figure of a volute, he formed without a seed containing the same structure in the small, as in the larger forms ? Who moulded it so artfully, and where are the moulds ? ” The stalactites which formed in caverns in various parts of the world were also supposed to be proofs of this vegetative growth. Still another theory has been held at various times, and is not yet entirely forgotten, namely: that the Creator made fossil animals and plants just as they are found in the rocks, in pursuance of a plan beyond our comprehension. This theory has never prevailed among those familiar with scientific facts, and hence needs here no further consideration. An interest in fossil remains arose in England later than on the continent; but when attention was directed to them, the first opinions as to their origin were not less fanciful and erroneous than those to which we have already referred. Dr. Plot, in his “Natural History of Oxfordshire,” published in 1677, considered the origin of fossil shells and fishes to be due to a “ plastic virtue, latent in the earth,” as Theophrastus had suggested long before. Lhwyd, in his “LithojpJiylacii Britan- nici Ichnograjphia ,” published at Oxford in 1699, gives a cata¬ logue of English fossils contained in the Ashmolean Museum. He opposed the vis jplastioa theory, and expressed the ojfinion that the spawn of fishes and other marine animals had been raised with the vapors from the sea, conveyed inland by clouds, and deposited by rain, had permeated into the interior of the earth, and thus produced the fossil remains we find in the rocks. About this time several important works were pub¬ lished in England by Dr. Martin Lister, which did much to infuse, a true knowledge of fossil remains. He gave figures of recent shells side by side with some of the fossil forms, so that the resemblance became at once apparent. The fossil species of shells he called “turbinated and bivalve stones,” and adds, “ either these were terriginous, or, if otherwise, the animals which they so exactly represent have become extinct.” 11 During tlie seventeenth century there was a considerable advance in the study of fossil remains. The discussions in regard to the nature and origin of these objects, had called attention to them, and many collections were now made, espe¬ cially in Italy, and also in Germany, where a strong interest in this subject had been aroused. Catalogues of these collec¬ tions were not unfrequently published, and some of them were illustrated with such accurate figures, that many of the species can now he readily recognized. In this century, too, an important step in advance was made by the collection and description of fossils from particular localities and regions, in distinction from general collections of curiosities. Casper Schwenkfeld, in 1600, published a catalogue of the fossils discovered in Silesia ; in 1622, a detailed description of the renowned Museum of Calceolarius, of Yerona, appeared; and in 1642, a catalogue of Besler’s collection. Wormius’s cata¬ logue was published in 1652 ; Spener’s in 1663 ; and Septala’s in 1666. A description of the Museum of the King of Den¬ mark was issued in 1669 ; Cottorp’s catalogue in 1614; and that of the renowned Kirscher in 1618. Dr. Grew gave an account in 1681 of the specimens in the Museum of Gresham’s College in England; and in 1695, Petiver of London published a cata¬ logue of his very extensive collection. A catalogue by Fred. Lauchmund, on the fossils of Hildesliein, appeared in 1669, and the fossils of Switzerland were described by John Jacob Wagner in 1689. Among similar works, were the dissertations of Gyer, at Frankfort, and Alhertus, at Leipsic. Steno, a Dane, who had been professor of anatomy at Padua, published, in 1669, one of the most important works of this period.* He entered earnestly into the controversy as to the origin of fossil remains, and by dissecting a shark from the Mediterranean, proved that its teeth were identical with some found fossil in Tuscany. He also compared the fossil shells found in Italy with existing species, and pointed out their resemblance. In the same work, Steno expressed some very * Be solido intra solidum naturaliter contento. important views in regard to the different kinds of strata, and their origin, and first placed on record the important fact that the oldest rocks contain no fossils. Scilla, the Sicilian painter, published in 1670 a work on the fossils of Calabria, well illustrated. He is very severe against those who doubted the organic origin of fossils, but is inclined to consider them relics of the Mosaic deluge. Another instance of the power of the lusus natural theory, even at the close of the seventeenth century, deserves mention. In the year 1696, the skeleton of a fossil elephant was dug up at Tonna, near Gotha, in Germany, and was described by William Ernest Tentzel, a teacher in the Gotha Gymnasium. He declared the bones to be the remains of an animal that had lived long before. The Medical Faculty in Gotha, however, considered the subject, and decided officially that this specimen was only a freak of nature. Beside the authors I have mentioned, there were many others who wrote about fossil remains before the close of the seventeenth century, and took part in the general discussion as to their nature and origin. During the progress of this con¬ troversy the most fantastic theories were broached and stoutly defended, and although refuted from time to time by a few clear-headed men, continually sprang up anew, in the same or modified forms. The influence of Aristotle’s views of equivo¬ cal generation, and especially the scholastic tendency to dispu¬ tation, so prevalent during the middle ages, had contributed largely to the retardation of progress, and yet a real advance in knowledge had been made. The long contest in regard to the nature of fossil remains was essentially over, for the more intelligent opinion' at the time now acknowledged that these objects were not mere “ sports of nature,” but had once been endowed with life. At this point, therefore, the first period in the history of Palaeontology,* as I have indicated it, may appropriately end. It is true that later still, the old exploded errors about the plastic force and fermentation were from time to time revived, 13 as they have been almost to the present day; bnt learned men, with few exceptions, no longer seriously questioned that fossils were real organisms, as the ancients had once believed. The many collections of fossils that had been brought together, and the illustrated works that had been published about them, were a foundation for greater progress, and, with the eighteenth cen¬ tury, the second period in the history of Palaeontology began. The main characteristic of this period was the general belief, that fossil remains were deposited by the Mosaic deluge. W e have seen that this view had already been advanced, but it was not till the beginning of the eighteenth century that it became the prevailing view. This doctrine was strongly opposed by some courageous men, and the discussion on the subject soon became even more bitter than the previous one, as to the nature of fossils. In this diluvial discussion theologians and laymen alike took part. For nearly a century the former had it all their own way, for the general public, then as now, believed what they were taught. Noah’s flood was thought to have been universal, and was the only general catastrophe of which the people of that day had any knowledge or conception. The scholars among them were of course familiar with the accounts of Deucalion and his ark, in a previous deluge, as we are to-day with similar traditions held by various races of men. The firm belief that the earth and all it contains was created in six days ; that all life on the globe was destroyed by the deluge, except alone what Noah saved; and that the earth and its inhabitants were to be destroyed by fire, was the foun¬ dation on which all knowledge of the earth was based. With such fixed opinions, the fossil remains of animals and plants were naturally regarded as relics left by the flood described in Holy Writ. The dominant nature of this belief is seen in nearly all the literature in regard to fossils published at this time, and some of the works which then appeared have become famous on this account. 14 In 1710, David Biittner published a volume entitled “Rudera Diluvii Testes .” He strongly opposed Lhwyd’s explanation of the origin of fossils, and referred these objects directly to the Flood. The most renowned work, however, of this time, was published at Zurich, in 1726, by Scheuchzer, a physician and naturalist, and professor in the University of Altorf. It bore the title “Homo Dilumi Testis P The specimen upon which this work was based was found at Oeningen, and was regarded as the skeleton of a child destroyed by the Deluge. The author recognized in this remarkable fossil, not merely the skeleton, but also portions of the muscles, the liver, and the brain. The same author was fortunate enough to discover, subsequently, near Altorf, two fossil vertebrae, which he at once referred to that “ Accursed race destroyed by the Flood!” These, also, he carefully described and figured in his “Physiea Sacra,” published at Ulm in 1731. Engravings of both were subsequently given in the “ Copper-Bible.” Cuvier afterward examined these interesting relics, and pronounced the skeleton of the supposed child to be the remains of a gigantic Salamander, and the two vertebrae to be those of an Ichthyosaurus! Another famous book appeared in Germany in the same year in which Scheuclizer’s first volume was published. The author was John Bartholomew Adam Beringer, professor at the Uni¬ versity of Wurtzburg, and his great work* indirectly had an important influence upon the investigation of fossil remains. The history of the work is instructive, if only as an indication of the state of knowledge at that date. Professor Beringer, in accordance with views of his time, had taught his pupils that fossil remains, or “ figured stones,” as they were called, were mere “ sports of nature.” Some of his fun-loving students reasoned among themselves, “ If Nature can make figured stones in sport, why cannot we ?” Accordingly, from the soft * Lithographia Wirceburgensis, ducentis lapidum figuratorum, a potiori , insecti- formium, prodigiosis imaginibus exornaia. Wirceburgi, 1726. Edit. II. Franco- furti et Lipsiae. limestone in the neighboring hills, they carved out figures of marvelous and fantastic forms, and buried them at the localities where the learned Professor was accustomed to dig for his fossil treasures. His delight at the discovery of these strange forms encouraged further production, and taxed the ingenuity of these youthful imitators of Nature’s secret processes. At last Beringer had a large and unique collection of forms, new to him, and to science, which he determined to publish to the world. After long and patient study, his work appeared, in Latin, dedicated to the reigning prince of the country, and illustrated with twenty-one folio plates. Soon after the book was published, the deception practiced upon the credulous Professor became known; and, in place of the glory he ex¬ pected from his great undertaking, he received only ridicule and disgrace. He at once endeavored to repurchase and destroy the volumes already issued, and succeeded so far that few copies of the first edition remain. His small fortune, which had been seriously impaired in bringing out his grand work, was exhausted in the effort to regain what was already issued, as the price rapidly advanced in proportion as fewer copies remained; and, mortified at the failure of his life’s work, he died in poverty. It is said that some of his family, dissatisfied with the misfortune brought upon them by this disgrace and the loss of their patrimony, used a remaining copy for the production of a second edition, which met with a large sale, sufficient to repair the previous loss, and restore the family fortune. This work of Beringer, in the end, exerted an excellent influence upon the dawning science of fossil remains. Observers became more cautious in announcing supposed discoveries, and careful study of natural objects gradually replaced vague hypotheses. The above works, however, are hardly fair examples of the literature on fossils during this part of the eighteenth century. Scheuchzer had previously published his well-known “ Com¬ plaint and Vindication of the Fishes,” illustrated with good plates. Moro, in his work on “ Marine Bodies which are 16 found in tlie Mountains,” 1740, showed the effects of volcanic action in elevating strata, and causing faults. Yallisneri had studied with care the marine deposits of Italy. Donati, in 1750, had investigated the Adriatic, and ascertained by sound¬ ings that shells and corals were being imbedded in the deposits there, just as they were found in the rocks. John Gesner’s dissertation, u De Petrificatis ,” published at Leyden in 1758, was a valuable contribution to the science. He enumerated the various kinds of fossils, and the different conditions in which they are found petrified, and stated that some of them, like those at Oeningen, resembled the shells, fishes, and plants of the neighboring region, while others, such as Ammonites and Belemnites, were either unknown species, or those found only in distant seas. He discusses the structure of the earth at length, and speculates as to the causes of changes in sea and land. He estimates that, at the observed rate of reces¬ sion of the ocean, to allow the Appenines, whose summits are filled with marine shells, to reach their present height, would have taken about eighty thousand years, a period more than “ ten times greater than the age of the universe.” He accord¬ ingly refers the change to the direct command of the Deity, as related by Moses, that, “ The waters should be gathered together in one place, and the dry land appear.” Voltaire (1694-1778), discussed geological questions and the nature of fossils in several of his works, but his published opinions are far from consistent. He ridiculed effectively and justly the cosmogonists of his day, and showed, also, that he knew the true nature of organic remains. Finding, however, that theologians used these objects to confirm the Scriptural account of the deluge, he changed his views, and accounted for fossil shells found in the Alps, by suggesting that they were Eastern species, dropped by the pilgrims on their return from the Holy Land ! Buffon, in 1749, published his important work on Natural History, and included in it his “ Theory of the Earth,” in which he discussed, with much ability, many points in Geology. 6 I 17 Soon after the hook was published, he received an official letter from the Faculty of Theology in Paris, stating that fourteen propositions in his works were reprehensible, and contrary to the creed of the church. The first objectionable proposition was as follows : “ The waters of the sea have produced the mountains and valleys of the land,—the waters of the heavens reducing all to a level, will at last deliver the whole land over to the sea, and the sea successively prevailing over the land, will leave dry new continents like those we inhabit.” Buffon was politely requested by the college to recant, and having no particular desire to he a martyr to science, submitted the following declaration, which he was required to publish in his next work: “ I declare that I had no intention to contra¬ dict the text of Scripture ; that I believe most firmly all therein related about the creation, both as to order of time and matter of fact; and I abandon everything in my hook respect¬ ing the formation of the earth, and, generally, all which may he contrary to the narration of Moses.” This single instance will suffice to indicate one great obsta¬ cle to the advancement of science, even up to the middle of the eighteenth century. Another important work appeared in France about this time, Bourguet’s “Traite des Petrifactions ,” published in 1758, which is well illustrated with faithful plates. In England, a discourse on earthquakes, by Dr. Robert Flooke, was published in 1705. This author held some views in advance of his time, and main¬ tained that figured stones were “ really the several bodies they represent or the moldings of them petrified, and not, as some have imagined, a lusus naturaz , sporting herself in the needless formation of useless things.” He anticipates one important conclusion from fossils, when he states that “ though it must he very difficult to read them and to raise a chronology out of them, and to state the intervals of time wherein such or such catastrophes and mutations have happened, yet it is not im¬ possible.” He also states that fossil turtles, and such large Ammonites as are found in Portland, seem to have been the 3 productions of liotter countries, and lienee it is necessary to suppose that England once lay under the sea within the torrid zone. He seems to have suspected that some of the fossils of England belonged to extinct species, but thought they might possibly be found living in the bottom of distant oceans. Dr. Woodward’s “Natural History of the Fossils of Eng¬ land” appeared in 1729. This work was based on a systematic collection of fossils which he had brought together, and which he subsequently bequeathed to the University of Cambridge, where it is still preserved, with his arrangement carefully retained. This descriptive part of this work is interesting, but his conclusions are made to coincide strictly with the Scriptural account of the creation and deluge. He had previously stated, in another work, that he believed, “ the whole terrestrial globe to have been taken to pieces and dissolved at the flood, and the strata to have settled down from this promiscuous mass.” In support of this view, he stated that, “Marine bodies are lodged in the strata according to the order bf their gravity, the heavier shells in stones, the lighter in chalk, and so of the rest.”* The most important w T ork on fossils published in Germany at this time, was that of George Wolfgang Knorr, which was continued after his death by Walcli. This work consisted of four folio volumes, with many plates, and was printed at Nuremberg, 1755-73. A large number of fossils were accu¬ rately figured and described, and the work is one of permanent value.f A French translation of this work appeared in 1767-78. Burton’s “Oryctographie de Bruxelles ,” 1784, con¬ tains figures and descriptions of fossils found in Belgium. Abraham Gottlieb Werner (1750-1817), Professor of Min¬ eralogy at Freyberg, did much to advance the science of Geology, and indirectly, that of fossils. He first indicated the relations of the main formations to each other, and, according to his pupil, Professor Jameson, first made the highly important * Essay towards a Natural History of the Earth. 1695. f Lapid.es ex celeberr. viror. sententia diluvii universalis testes, quos in ordines ac species distribuit, suis coloribus expremit , etc. 272 Tab. 1755-73. 19 observation “that different formations can be discriminated by tbe petrifactions they contain.” Moreover, “ that the petri¬ factions contained in the oldest rocks are very different from any of the species of the present time ; that the newer the formation, the more do the remains approach in form to the organic beings of the present creation.” Unfortunately, Werner published little, and his doctrines were mainly dis¬ seminated by his enthusiastic pupils. The great contest between the Yulcanists and the Neptunists started at this time, mainly through Werner, whose doctrines led to the controversy. The comparative merits of tire and water, as agencies in the formation of certain rocks, were dis¬ cussed with a heat and acrimony characteristic of the subject and the time. Werner believed in the aqueous theory, while the igneous theory was especially advocated by Hutton of Edin¬ burgh, and his illustrator, Playfair. This discussion resulted in the advancement of descriptive geology, but the study of fossils gained little thereby. The “ Protogaea ” of Leibnitz, the great mathematician, pub¬ lished in 1749, about thirty years after his death, was a work of much merit. This author supposed that the earth had gradually cooled from a state of igneous fusion, and was subse¬ quently covered with water. The subsidence of the lower part of the earth, the deposits of sedimentary strata from inunda¬ tions, and their induration, as well as other changes, followed. All this, he supposed to have been accomplished in a period of six natural days. In the same work Leibnitz shows that he had examined fossils with considerable care. Linnaeus (1707-1778), the famous Swedish botanist, and the founder of the modern system of nomenclature in Natural History, confined his attention almost entirely to the living forms. Although he was familiar with the literature of fossil remains, and had collected them himself, he did not include them in his system of plants and animals, but kept them sepa¬ rate, with the minerals; hence he did little directly to advance this branch of science. 20 During tlie last quarter of the eighteenth century, the belief that fossil remains were deposited by the Deluge sensibly declined, and the dawn of a new era gradually appeared. Let us pause for a moment here, and see what real progress had been made; what foundation had been laid on which to estab¬ lish a science of fossil remains. The true nature of these objects had now been clearly deter¬ mined. They were the remains of animals and plants. Most of them certainly were not the relics of the Mosaic Deluge, but had been deposited long before, part in fresh water and part in the sea. Some indicated a mild climate, and some the tropics. That any of these were extinct species, was as yet only suspected. Large collections of fossils had now been made, and valuable catalogues, well illustrated, had been pub¬ lished. Something was known, too, of the geological position of fossils. Steno, long before, had observed that the lowest rocks were without life. Lehmann had shown that above these primitive rocks, and derived from them, were the sec¬ ondary strata, full of the records of life; and above these were alluvial deposits, which he referred to local floods, and the Deluge of Hoah. Rouelle, Fuchsel, and Odoardi had shed new light on this subject. Werner had distinguished the tran¬ sition rocks, containing fossil remains, between the primitive and the secondary, while everything above the chalk he grouped together, as the “overflowed land.” Werner, as we have seen, had done more than this, if we give him the credit his pupils claim for him. ILe had found that the formations he examined contained each its own peculiar fossils, and that from the older to the newer there was a gradual approach to recent forms. William Smith had worked out the same thing in England, and should equally divide the honor , of this important discovery. The greatest advance, however, up to this time, was that men now preferred to observe , rather than to believe , and facts were held in greater esteem than vague speculations. With this preparation for future progress, the second period in the history of Palaeontology, as I have divided it, may appropri¬ ately be considered at an end. 21 Tims far, I have said nothing in regard to one branch of my subject, the rnethods of Palaeontological research, for up to this time, of method there was none. We have seen that those of the ancients who noticed marine shells in the solid rock, called them such, and concluded that they had been left there by the sea. The discovery of fossils led directly to theories of how the earth was formed. Here the progress was slow. Subterranean spirits were supposed to guard faithfully the mysteries of the earth; while above the earth, Authority guarded with still greater power the secrets men in advance of their age sought to know. The dominant idea of the first sixteen centuries of the present era was, that the universe was made for Man. This was the great obstacle to the correct determination of the position of the earth in the universe, and, later, of the age of the earth. The contest of Astronomy against authority was long and severe, but the victory was at last with science. The contest of Geology against the same power followed, and continued almost to our day. The result, is still the same. In the early stages of this contest, there was no strife, for science was benumbed by the embrace of super¬ stition and creed, and little could be done till that was cast off. In a superstitious age, when every natural event is referred to a supernatural cause, science cannot live; and often as the sacred fire may be kindled by courageous far-seeing souls, will it be quenched by the dense mists of ignorance around it. Scarcely less fatal to the growth of science is the age of Authority, as the past proves too well. With freedom of thought, came definite knowledge, and certain progress;—but tw T o thousand years was long to wait. With the opening of the present century, began a new era in Palaeontology, which we may here distinguish as the third period in its history. This branch of knowledge became now a science. Method replaced disorder, and systematic study superseded casual observation. For the next half century the advance w T as continuous, and rapid. One characteristic of this 22 period was, the accurate determination of fossils by compar¬ ison with living forms. This will separate it from the two former epochs. Another distinctive feature of this period was the general belief that, every species , recent and extinct , was a separate creation. At the very beginning of the epoch we are now to consider, three names stand out in bold relief: Cuvier, Lamarck, and William Smith. To these men, the science of palaeontology owes its origin. Cuvier and Lamarck, in France, had all the power which great talent, education, and station could give; William Smith, an English surveyor, was without culture or influence. The last years of the eighteenth century had been spent by each of these men in preparation for his chosen work, and the results were now given to the world. Cuvier laid the foundation of the palaeontology of Yertebrate animals; Lamarck, of the Invertebrates ; and Smith established the prin¬ ciples of Stratigraphical Palaeontology. The investigator of fossils to-day seldom needs to consult earlier authors of the science. George Cuvier (1769-1832), the most famous naturalist of his time, was led to the study of extinct animals by ascertaining that the remains of fossil elephants he examined were extinct species. “ This idea,” he says later, “ which I announced to the Institute in the month of January 1796, opened to me views entirely new respecting the theory of the earth, and determined me to devote myself to the long researches and to the assiduous labors which have now occupied me for twenty- five years.”* It is interesting to note here that in this first investigation of fossil vertebrates, Cuvier employed the same method that gave him such important results in his later researches. Remains # of elephants had been known to Europe for centuries, and many authors, from Pliny down to the contemporaries of Cuvier, had written about them. Some had regarded the bones as those of human giants, and those who recognized what they were consid¬ ered them remains of the elephants imported by Hannibal or * Ossemens Fossiles , Second Edition, Vol. I, p. 1*78. the Romans. Cuvier, however, compared the fossils directly with the bones of existing elephants, and proved them to be distinct. The fact that these remains belonged to extinct species was of great importance. In the case of fossil shells, it was difficult to say that any particular form was not living in a distant ocean; but the two species of existing elephants, the Indian and the African, were w T ell known, and there w r as hardly a possibility that another living one would be found. It is important to bear in mind, too, that Cuvier’s preparation for the study of the remains of animals was far in advance of any of his predecessors. He had devoted himself for years to careful dissections in the various classes of the animal kingdom, and was really the founder of comparative anatomy, as we now understand it. Cuvier investigated the different groups of the whole kingdom with care, and proposed a new classiiication founded on the plan of structure, which in its main features is the one in use to-day. The first volume of his Comparative Anatomy appeared in 1800, and the work was completed in five volumes in 1805. Previous to Cuvier, the only general catalogue of animals was contained in Linnaeus’ “ Systerna Natures?' In this work, as we have seen, fossil remains were placed with the Minerals, not in their appropriate places among the animals and plants. Cuvier enriched the animal kingdom by the introduction of fossil forms among the living, bringing all together into one comprehensive system. His great work, “ Le Regne Animal ,” appeared in four volumes, in 1817, and with its two subsequent editions remains the foundation of modern zoology. Cuvier’s classic work on vertebrate fossils—“ Recherehes sur les Ossemens Fossiles ,” in four volumes, appeared in 1812-13. Of this work, it is but just to say that it could only have been written by a man of genius, profound knowledge, the greatest industry, and with the most favorable opportunities. The introduction to this work was the famous “Discourse on the Revolutions of the Surface of the Globe,” which has perhaps been as widely read as any other scientific essay. The 24 discovery of fossil bones in the gypsum quarries of Paris, by the workmen, who considered them human remains; the care¬ ful study of these relics by Cuvier, and his restorations from them of strange beasts that had lived long before, is a story with which you are all familiar. Cuvier was the first to prove that the earth had been inhabited by a succession of different series of animals, and he believed that those of each period were peculiar to the age in which they lived. In looking over his work after a lapse of three-quarters of a century, we can now see that Cuvier was wrong on some important points, and failed to realize the direction in which science was rapidly tending. With all his knowledge of the earth, he could not free himself from tradition, and believed in the universality and power of the Mosaic deluge. Again, he refused to admit the evidence brought forward by his distin¬ guished colleagues against the permanence of species, and used all his great influence to crush out the doctrine of evolution, then first proposed. Cuvier’s definition of a species, the domi¬ nant one for half a century, was as follows: “ A species comprehends all the individuals which descend from each other, or from a common parentage, and those which resemble them as much as they do each other.” The law of “ Correlation of Structures,” as laid down by Cuvier, has been more widely accepted than almost any thing else that bears his name; and yet, although founded in truth, and useful within certain limits, it would certainly lead to serious error if applied widely in the way he proposed. In his Discourse, he sums this law as follows: “ A claw, a shoulder blade, a condyle, a leg or arm bone, or any other bone separately considered, enables us to discover the description of teeth to which they have belonged; so also reciprocally we may determine the form of the other bones from the teeth. Thus, commencing our investigation by a careful survey of any one bone by itself, a person who is sufficiently master of the laws of organic structure, may, as it were, reconstruct the whole animal to which that bone had belonged.” We know to-day that unknown extinct animals cannot be restored from a single tooth or claw, unless they are very similar to forms already known. Had Cuvier, himself, applied his methods to many forms from the early Tertiary or older formations, he would have failed. If, for instance, he had had before him the disconnected fragments of an Eocene Tillodont, he would undoubtedly have referred a molar tooth to one of his Pachyderms ; an incisor tooth to a Rodent; and a claw bone to a Carnivore. The tooth of a Hesperornis would have given him no possible hint of the rest of the skeleton, nor its swim¬ ming feet the slightest clue to the ostrich-like sternum or skull. And yet, the earnest belief in his own methods led Cuvier to some of his most important discoveries. Jean Lamarck (1744-1829), the philosopher and naturalist, a colleague of Cuvier, was a learned botanist before he became a zoologist. His researches on the invertebrate fossils of the Paris Basin, although less striking, were not less important than those of Cuvier on the vertebrates; while the conclu¬ sions he derived from them form the basis of modern biology. Lamarck’s method of investigation was the same, essentially, as that used by Cuvier, namely: a direct comparison of fossils with living forms. In this way, he soon ascertained that the fossil shells imbedded in the strata beneath Paris were, many of them, extinct species, and those of different strata differed from each other. His first memoir on this subject appeared in 1802,* and, with his later works, effected a revolution in concliology. His “ System of Invertebrate Animals” appeared the year before, and his famous “Philosophie Zoologique ,” in 1809. In these two works, Lamarck first announced the princi¬ ples of Evolution. In the first volume of his “ Natural History of Invertebrate Animals,”f he gave his theory in detail; and to-day one can only read with astonishment his far-reaching anticipations of modern science. These views were strongly * Memoires sur les fossiles des environs de Paris. 1802-6. \ Histoire naturelle des animaux sans vertebres. 1 vols. Paris, 1815-1822. 2d edition. 11 vols. 1835-1845. 26 supported by Geoffroy Saint-Hilaire, but bitterly opposed by Cuvier; and tbeir great contest on this subject is well known. In looking back from this point of view, the philosophical breadth of Lamarck’s conclusions, in comparison with those of Cuvier, is clearly evident. The invertebrates on which Lamarck worked offered less striking evidence of change than the various animals investigated by Cuvier; yet they led Lamarck directly to Evolution, while Cuvier ignored what Avas before him on this point, and rejected the proof offered by others. Both pursued the same methods, and had an abun¬ dance of material on which to work, yet the facts observed induced Cuvier to believe in catastrophes; and Lamarck, in the uniform course of nature. Cuvier declared species to be permanent; Lamarck, that they were descended from others. Both men stand in the first rank in science; but Lamarck was the prophetic genius, half a century in advance of his time. While the Paris Basin was yielding such important results for Paleontology, its geological structure was being worked out with great care. The results appeared in a volume by Cuvier and Alex. Brongniart, chiefly the work of the latter, published in 1808.* This was the first systematic investigation of Tertiary strata. Three years later, the work was issued in a more extended form. The separate formations were here carefully distinguished by their fossils, the true importance of which for this purpose being distinctly recognized. This advance was not accepted without some opposition, and it is an interesting fact that Jameson, who claimed for Werner the theory here put in practice, rejected its application, and wrote as follows: “ To Cuvier and Brongniart we are indebted for much valuable information in their description of the country around Paris, but we must protest against the use they have made of fossil organic remains in their gcognostical descrip¬ tions and investigations.’^ * Essai sur la Geographie Mineralogique des environs de Paris. 4to, 1808. f Translation of Cuvier’s Discourse. Note K. (B.), p. 103, 1817. 27 William Smith (1769-1839), “ the father of English Geology,” had previously published a “Tabular View of the British Strata.” He appears to have arrived independently at essentially the same view as Werner in regard to the relative position of strat¬ ified rocks. lie had determined that the order of succession was constant, and that the different formations might be identi¬ fied at distant points by the fossils they contained. In his later works, “Strata identified by Organized Fossils,” pub¬ lished in 1816-20, and “ Stratigrapliical System of Organized Fossils,” 1817, he gave to the world results of many years of careful investigations on the Secondary formations of England. In the latter work, he speaks of the success of his method in determining strata by their fossils, as follows : “ My original method of tracirlg the strata by the organized fossils imbedded therein, is thus reduced to a science not difficult to learn. Ever since the first written account of this discovery was circulated in 1799, it has been closely investigated by my scientific acquaintances in the vicinity of Bath, some of whom search the quarries of different Strata in that district, with as much cer¬ tainty of finding the characteristic Fossils of the respective rocks, as if they were on the shelves of their cabinets.” The systematic study of fossils now attracted attention in England, also, and was prosecuted with considerable zeal, although with less important results than in France. An ex¬ tensive work on this subject, by James Parkinson, entitled “ Organic Remains of a Former World,” was begun in 1804, and completed in three volumes in 1811. A second edition appeared in 1833. This work was far in advance of previous publications in England, and, being well illustrated, did much to make the collection and study of fossils popular. The belief in the geological effects of the Deluge had not yet lost its power, although restricted now to the later deposits; for Park¬ inson in his later edition wrote as follows: “Why the earth was at first so constituted that the deluge should be rendered necessary—why the earth could not have been at first stored with all those substances, and endued with all those properties which seemed to have proceeded from the deluge—why so many beings were created, as it appears, for the purpose of being destroyed — are questions which I presume not to answer.” William Buckland (1784-1856), published in 1823 his cele¬ brated “Reliquiae Diluviance ,” in which he gave the results of his own observations in regard to the animal remains found in the caves, fissures and alluvial gravels of England. The facts presented are of great value, and the work was long a model for similar researches. Buckland’s conclusions were, that none of the human remains discovered in the caves were as old as the extinct mammals found with them, and that the Deluge was universal. In speaking of fossil bones found in the Himalaya mountains, he says: “ The occurrence of these bones at such an enormous elevation in the region of eternal snow, and consequently in a spot now unfrequented by such animals as the horse and deer, can, I think, be explained only by sup¬ posing them to be of antediluvian origin, and that the carcasses of the animals were drifted to their present place, and lodged in sand, by the diluvial waters.” The foundation of the “ Geological Society of London,” in 1807, marks an important point in the history of palaeontology. To carefully collect materials for future generalizations, was the object in view, and this organization gradually became the centre in Great Britain for those interested in geological science. The society was incorporated in 1826, and has since been the leading organization in Europe for the advancement of the sciences within its field. The Geological Society of France, established at Paris in 1832, and the German Geologi¬ cal Society, founded at Berlin in 1848, have likewise contribu¬ ted largely to geological investigations in these countries, and to some extent in other parts of the world. In the jmblications of these three societies, the student of palaeontology will find a mine of valuable materials for his work. The systematic study of fossil Plants may be said to date from the publication of Adolphe Brongniart’s “Prodrome” 29 in 1828.* This was very soon followed by his larger work, “Ilistoire des vegetaux fos sites” issued in 1828-48. Brong- niart pursued the same method as Cuvier and Lamarck, viz: the comparison of fossils with living forms, and his results were of great importance. In his “ Tableau des genres vegetaux fossiles ,” etc., published in Paris in 1849, he gives the classifica¬ tion and distribution of the genera of fossil plants, and traces out the historical progression of vegetable life on the globe, as he had done to a great extent in his previous works. He shows that the cryptogamic forms prevailed in the primary formations; the conifers and cycads in the secondary, and the higher forms in the Tertiary, while four fifths of living plants are exogens. In England, Lindley and Hutton published, in 1831-37, a valuable wmrk in three volumes, entitled, “ Fossil Flora of Great Britain.” This work was illustrated by many accurate plates, in which the plants of the coal formation were especially represented. Henry Witham also published two works in 1831 and 1833, in which he treated especially of the internal struc¬ ture of fossil plants. “ Antediluvian Phytology,” by Artis, was published in London in 1838. Bowerbank’s “ History of the Fossil Fruits and Seeds of the London Clay,” appeared in 1843. Hooker’s memoir “On the Yegetation of the Carbon¬ iferous Period as compared with that of the present day,” pub¬ lished in 1848, was an important contribution to the science. Bunbury, Williamson, and others, also published various papers on fossil plants. This branch of Palseontology, how¬ ever, attracted much less attention in England, than on the Continent. In Germany, the study of fossil plants dates back to the beginning of the century. Yon Schlotheim, a pupil of Wer¬ ner, published in 1804 an illustrated volume on this subject. A more important work was that of Count Sternberg, issued in 1820-38, and illustrated with excellent plates. Cotta in 1832 published a book with the title, “Die Dendrolithen ,” in * Prodrome d'une histoire des vegetaux fossiles. 8vo. Paris, 1828. 30 wliicli lie gave the results of his investigations on the inner structure of fossil plants. Yon Gutbier in 1835, and Germar in 1844-53, described and figured the plants of two important localities in Germany. Corda’s “ Beitrage zur Flora der Vorwelt ,” issued at Prague, in 1845, was essentially a continu¬ ation of the work of Sternberg. Unger’s “ Chloris protogcea” 1841-45, “ Genera et species plantarum fossilium” 1850, and his larger work published in 1852, are all standard authorities. In the latter, the theory of descent is applied to the vegetable world. Schimper and Mougeot’s “Monograph on the fossil plants of the Vosges,” 1845, was well illustrated, and contained noteworthy results. Goppert, in 1836, published a valuable memoir entitled, “ Systerna Filicum Fossilium ,” in which he made known the results of his study of fossil ferns. In the same year, this botanist began a series of experiments, in which he attempted to imitate the process of fossilization, as found in nature. lie steeped various animal and vegetable substances in waters hold¬ ing, some calcareous, others siliceous, and others metallic matter in solution. After a slow saturation, the substances were dried, and exposed to heat until the organic matters were burned. In this way Goppert successfully imitated various processes of petrifaction, and explained many things in regard to fossils that had previously been in question. His discovery of the remains of plants throughout the interior of coal did much to clear up the doubts about the formation of that substance. In 1841, Goppert published an important work in which he com¬ pared the genera of fossil plants with those now living. In 1852, another extensive work by this author appeared, entitled, u Fossile Flora des Uebergangs- Gebirges ” Andrse, Braun, D linker, Ettirisgliausen, Geinitz, and Golden- berg, all made notable contributions to fossil Botany in Germany, during the period we are now considering. The systematic study of invertebrate fossils, so admirably begun by Lamarck, was continued actively in France. The 31 Tertiary shells of the Seine valley were further investigated hy Def ranee, and especially hy Deshayes, whose great work on this subject was begun in 1824* DesMoulin’s essay on Sjpher- ulites in 1826, Blainville’s memoir on Belemnites in 1827, Ferussac’s various memoirs on land and fresh water fossil shells, were valuable additions to the subject. A later work of great importance was D’Orbigny’s Paleontologie Frangaise , 1840-44, which described the mollusca and radiates in detail, according to formations. The other publications of this author are both numerous and valuable. Brongniart and Desmarest’s “Histoire naturelle des Crustaces Fossiles ,” published in 1822, is a pioneer work on this subject. Michelins’ memoir on the fossil corals of France, 1841-46, was another important contri¬ bution to palaeontology. Agassiz’s works on fossil Echinoderms and Mollusks are valuable contributions to the science. The works of d’Archiac, Coquand, Cotteau, Desor, Edwards, Ilaime, and De Verneuil, are likewise of permanent value. In Italy, Bellardi, Merian, Michellotti, Pliillipi, Zigno, and others, contributed important results to Palaeontology. In Belgium, Bosquet, Nyst, Koninck, Byckholt, Yan Ben- eden, and others, have all aided materially in the progress of the science. In England, also, invertebrate fossils were studied with care, and continued progress was made. Sowerby’s “ Mineral Concli- ology of Great Britain,” in six volumes, a systematic work of great value, was published in 1812-30, and soon after was trans¬ lated into French and German. Its figures of fossil shells are excellent, and it is still a standard work. Miller’s “ Natural History of the Crinoidea,” published at Bristol, in 1821, and Austin’s later monograph, are valuable for reference. Brown’s “ Fossil Conchology of Britain and Ireland” appeared in 1839, and Brodie’s History of the Fossil Insects of England, in 1845. Phillips’ illustration of the geology of Yorkshire, 1829-36, and his work on the Palseozoic fossils of Cornwall, Devonshire, and West Somerset, 1843, contained a great deal of original matter * Description des coquilles fossiles des environs de Paris. 3 vols. Paris, 1824-37. 32 in regard to fossil remains. Morris’ “ Catalogue of British Fossils,” issued in 1843, and the later edition in 1854, is most useful to the working paleontologist. The memoirs of David¬ son on the Brachiopoda, Edwards, Forbes, Morris, Lycett, Sharpe, and Wood on other Mollusca, Wright on the Echino- derms, Salter on Crustacea, Busk on Polyzoa, Jones on the Entomostraca, and Duncan and Lonsdale on Corals, are of especial value. King’s volume on Permian fossils, Mantell’s various memoirs, Dixon’s work on the fossils of Sussex, 1850, and McCoy’s works on Palaeozoic fossils, all deserve honorable mention. Sedgwick, Murchison, and Lyell, although their greatest services were in systematic geology, each contributed important results to the kindred science of palaeontology during the period we are reviewing. In Germany, Schlotheim’s treatise, “ Die Petrifactenkunde ,” published at Gotha in 1820, did much to promote a general interest in fossils. By far the most important work issued on this subject was the “ Petrifacta Germanica ,” by Goldfuss, in three folio volumes, 1826 to 1844, which has lost little of its value. Bronn’s “ Geschichte der Natur ,” 1841-46, was a work of great labor, and one of the most useful in the litera¬ ture of this period. The author gave a list of all the known fossil species, with full references, and also their distribution through the various formations. This gave exact data on which to base generalizations, hitherto of comparatively little value. Among other early works of interest in this department may be mentioned, Dalman’s memoir on Trilobites , 1828, and Bur- meister’s on the same subject, 1843. Giebel’s well known “Fauna der Vorwelt ,” 1847-1856, gave lists of all the fossils described up to that time, and hence is a very useful work. The “ Letheea Geognostica ” by Bronn, 1834-38, and the second edition by Bronn and Boemer, 1846-56, is a compre¬ hensive general treatise on Paleontology, and the most valuable work of the kind yet published. The researches of Ehrenberg, in regard to the lowest forms • of animals and plants, threw much light on various points in 33 *) Palaeontology, and showed the origin of extensive deposits, the nature of which had before been in doubt. Yon Buch, Barrande, Beyrich, Berendt, Dunker, Geinitz, Heer, Hornes, Klipstein, Yon Munster, Beuss, Roemer, Sandberger, Suess, Yon Hagenow, Yon Hauer, Zeiten, and many others, all aided in the advancement of this branch of science. Angelin, His- inger, and Nilsson, in Scandinavia; Abich, DeWaldheim, Eich- wald, Keyserling, Kutorga, Nordmann, Pander, Rouillier, and Yolborth, in Russia*; and Pusch in Poland, published important results on fossil invertebrates. The impetus given by Cuvier to the study of vertebrate fossils extended over Europe, and great efforts were made to continue discoveries in the direction he had so admirably pointed out. Louis Agassiz (1807-73), a pupil of Cuvier, and long an honored member of this association, attained eminence in the study of ancient as well as of recent life. His great work on Fossil Fishes* deserves to rank next to Cuvier’s “Ossemens FossilesP The latter contained mainly fossil mammals and reptiles, while the fishes were left without a historian till Agassiz began his investigations. His studies had admirably fitted him for the task, and his industry brought together a vast array of facts bearing on the subject. The value of this grand work consists not only in its faithful descriptions and plates, but also in the more profound results it contained. Agassiz first showed that there is a correspondence between the succession of fishes in the rocks, and their embryonal development. This is now thought to be one of the strongest points in favor of evolution, although its discoverer interpreted the facts as bearing the other way. Pander’s memoirs on the fossil fishes of Russia form a worthy supplement to Agassiz’s classic work. Brandt’s publi¬ cations are likewise of great value; and those of Lund, in Sweden, have an especial interest to Americans, in consequence of his researches in the caves of Brazil. * Recherches sur les Poissons fossiles, 1833-45. 5 34 Croizet and Jobert’s u Pecker ches sur les ossemens fossiles du department du Puy-de-Dome” published in 1828, contained valuable results in regard to fossil mammals. Geoffroy St. Hilaire’s researches on fossil Reptiles, published in 1831, were an important advance. He Serres and He Christol’s explora¬ tions in the caverns in the South of France, published between 1829 and 1839, were of much value. Schmerling’s researches in the caverns of Belgium, published in 1833-36, were especially important on account of the discovery of human remains min¬ gled with those of extinct animals. Heslongchamp’s memoirs on fossil reptiles, 1835, are still of great interest. Pictet’s general treatise on palaeontology was a valuable addition to the literature, and has done much to encourage the study of fossils,* HeBlainville, in his grand work, “ Osteographies issued in 1839-56, brought together the remains of living and extinct vertebrates, forming a series of the greatest value for study. Aymard and Pomel’s contributions to vertebrate Palae¬ ontology are both of value. Gervais and Lartet added much to our knowledge of the subject, and Bravard and Hebert’s memoirs are well known. The brilliant discoveries of Cuvier in the Paris Basin, excited great interest in England, and when it was found that the same Tertiary strata existed in the south of England, care¬ ful search was made for vertebrate fossils. Remains of some of the same genera described by Cuvier were soon discovered, and other extinct animals new to science were found in various parts of the kingdom. Konig, to whom we owe the name Ichthyosa%irus , and Conybeare, who gave the generic designation Plesiosaurus , and also Mosasaurus , were among the earliest writers in England on fossil reptiles. The discov¬ ery of these three extinct types, and the discussion as to their nature, forms a most interesting chapter in the annals of Palseontology. The discovery of the Iguanodon , by Mantell, and the Megalosaurus , by Buckland, excited still higher * li Traite elementaire de paleontologies etc*., Geneve. 4 vols. 1844 -46. Second Edition. Paris, 1853-55. 35 interest. These great reptiles differed much more widely from living forms than the mammals described by Cuvier, and the period in which they lived soon became known as the “ age of Reptiles.” The subsequent researches of these authors added largely to the existing knowledge of various extinct forms, and their writings did much to arouse public interest in the subject. Richard Owen, a pupil of Cuvier, followed, and brought to bear upon the subject an extensive knowledge of comparative anatomy, and a wide acquaintance with existing forms. His contributions have enriched almost every department of palae¬ ontology, and of extinct vertebrates especially, he has been, since Cuvier, the chief historian. The fossil reptiles of England, he has systematically described, as well as those of South Africa, The extinct Strutliious birds of Hew Zealand, he has made known to science, and accurately described in extended memoirs. His researches on the fossil mammals of Great Britain, the extinct Edentates of South America, and the ancient Marsupials of Australia, each forms an important chapter in the history of our science. The personal researches of Falconer and Cautley in the Sewalik Hills of India brought to light a marvelous vertebrate fauna of Pliocene age. The remains thus secured were made known in their great work, “Fauna Antiqua Sivalensis ,” published at London in 1845. The important contributions of Egerton to our knowledge of fossil fishes, and Jardine’s well known work, “Ichnology of Annandale,” also belong to this period. The study of vertebrate fossils in Germany was pros¬ ecuted with much success during the present period. Blumen- bach, the ethnologist, in several publications between 1803 and 1814, recorded valuable observations on this subject. In 1812, Sommerring gave an excellent figure of a pterodac¬ tyl e, which he named and described. Goldfuss’ researches on the fossil vertebrates from the Caves of Germany, published in 1820-23, made known the more important facts of that 36 interesting fauna. His later publications on extinct Amphibians and Reptiles were also noteworthy. Jager’s investigations on the extinct vertebrate fauna of Wurtemberg, published between 1824 and 1839, were an important advance. To Plieninger’s researches in the same region, 1834-44, we owe the discovery of the first Triassic mammal (. Microlestes ), as well as important information in regard to Labyrinthodonts. Kaup’s researches on fossil mammals, 1832-41, brought to light many interesting forms, and to him we are indebted for the generic name Dinotherium , and excellent descriptions of the remains then known. Count Munster’s “ Beitrage zur Petrifacten-Kunde ,” pub¬ lished 1843-46, contained several valuable papers on fossil vertebrates; and the separate papers by the same author are of interest. Andreas Wagner wrote on Pterosaurians in 1837, and later gave the first description of fossil mammals of the Tertiary of Greece, 1837-40. Johannes Muller published an important illustrated work on the Zeuglodonts, in 1849, and various notable memoirs; and Quenstedt, interesting descrip¬ tions of fossil reptiles, as well as other papers of much value. Putimeyer’s suggestive memoirs are widely known. Hermann von Meyer’s contributions to vertebrate palaeon¬ tology are by far the most important published in Germany during the period we are now considering. From 1830, his investigations on this subject were continuous for nearly forty years, and his various publications are all of value. His u Beitrage zur Petrifactenkunde ,” 1831-33, contains a series of valuable memoirs. His “ Palceologica” issued in 1832, includes a synopsis of the fossil vertebrates then known, with much original matter. His great work, “ Zur Fauna der Vorwelt ,” 1845-60, includes a series of monographs invaluable to the student of vertebrate palaeontology. This work, as well as his other chief publications, was illustrated with admirable plates from his own drawings. Other memoirs by this author will be found in the “ Paloeontograjpliica ,” of which he was one of the editors. In the many volumes of this publication, which began 37 in 1851, and is still continued, will be found much to interest the investigator in any branch of palaeontology. The “ Palseontographical Society of London,” established in 1847, has also issued a series of volumes containing valuable memoirs in various branches of Palaeontology. These two publications together are a storehouse of knowledge in regard to extinct forms of animal and vegetable life. It may be interesting here to note briefly the use of general terms in Palaeontology, as the gradual progress of the science was indicated to some extent in its terminology. At first, and for a long time, the name “fossil” was appropriately used for objects dug from the earth, both minerals and organic remains. The term “ Oryctology,” having essentially the same meaning, was also used for this branch of study. For a long period, too, the termination ites a stone) was applied to fossils to distinguish them from the corresponding living forms; as, for instance, “ Ostraeites” used by Pliny. At a later date, the general name “figured stones” (lapides figurati ) was exten¬ sively used ; and less frequently, “ Deluge stones ” ( lajpides diluviani). The term “ organized fossils ” was used to distin¬ guish fossils from minerals, when the real difference became known, although the name “Reliquiae” was sometimes em¬ ployed. The term “petrifactions” (. Petrificata ) was defined by John Gesner in his work on fossils in 1758, and was afterwards extensively used. Palaeontology is comparatively a modern term, having come into use only within the last half century. It was introduced about 1830, and soon was generally adopted in France and England; but in Germany it met with less favor, though used to some extent. It would be interesting, too, did time permit, to trace the various opinions and superstitions, held at different times, in regard to some of the more common fossils, for example, the Ammonite, or the Belemnite. Of their supposed celestial origin; of their use as medicine by the ancients, and in the 38 East to-day; of tlieir marvellous power as charms, among the Romans, and still among the American Indians. It would he instructive, also, to compare the various views expressed by stu¬ dents in science, concerning some of the stranger extinct forms, for instance, the Nummulites, among Protozoa; the Rudistes, among Mollusks; or the Mosasaurus, among Reptiles. Dissim¬ ilar as such views were, they indicate in many cases gropings after truth,—natural steps in the increase of knowledge. The third period in the history of Palaeontology, which, as I have said, began with Cuvier and Lamarck at the beginning of the present century, forms a natural epoch extending through six decades. The definite characteristics of this period, as stated, were dominant during all this time, and the progress of Palaeon¬ tology was commensurate with that of intelligence and culture. For the first half of this period, the marvelous discoveries in the Paris Basin excited astonishment, and absorbed attention; but the real significance and value of the facts made known by Cuvier, Lamarck, and William Smith, were not appreciated. There was still a strong tendency to regard fossils merely as interesting objects of natural history, as in the previous period, and not as the key to profounder problems in the earth’s his¬ tory. Many prominent geologists were still endeavoring to identify formations in different countries by their mineral characters, rather than by the fossils imbedded in them. Such names as “Old Red Sandstone,” and “Hew Red Sandstone, were given in accordance with this opinion. Humboldt, for example, attempted to compare the formations of South Amer¬ ica and Europe by their mineral features, and doubted the value of fossils for this purpose. In 1823, he wrote as follows: “ Are we justified in concluding that all formations are char¬ acterized by particular species ? that the fossil shells of the chalk, the muschelkalk, the Jura limestone, and the Alpine limestones, are all different ? I think this would be pushing the induction much too far.”* Jameson still thought minerals * Essai geognostique sur legisement des Roches, p. 41. 39 more important than fossils for characterizing formations; while Bakewell, later yet, defines Palaeontology as comprising u Fossil Zoology and Fossil Botany, a knowledge of which may appear to the student as having little connection with Geology.” During the later half of the third period, greater progress was made, and before its close Geology was thoroughly estab¬ lished as a science. Let us consider for a moment what had really been accomplished up to this time. It had now been proved beyond question that portions at least of the Earth’s surface had been covered many times by the sea, with alternations of fresh water and of land ; that the strata thus deposited were formed in succession, the lowest of the series being the oldest; that a distinct succession of animals and plants had inhabited the earth during the different geological periods; and that the order of succession found in one part of the earth was essentially the same in all. More than 30,000 new species of extinct animals and plants had now been described. It had been found, too, that from the oldest formations to the most recent, there had been an advance in the grade of life, both animal and vegetable, the oldest forms being -among the simplest, and the higher forms successively making their appearance. It had now become clearly evident, moreover, that the fossils from the older formations were all extinct species, and that only in the most recent deposits were there remains of forms still living. The equally important fact had been established, that in several groups of both animals and plants, the extinct forms were vastly more numerous than the living; while several orders of fossil animals had no representatives in modern times. Human remains had been found mingled with those of extinct, animals, but the association was regarded as an accidental one by the authorities in science; and the very recent appearance of Man on the earth was not seriously questioned. Another important conclusion reached, mainly through the labors of Lyell, was, that the earth had not been subjected in the past to sudden and violent revolutions; but the great changes wrought 40 liad been gradual, differing in no essential respect from those still in progress. Strangely enough, the corollary to this propo¬ sition, that Life, too, had been continuous on the earth, formed at that date no part of the common stock of knowledge. In the physical world, the great law of “ Correlation of forces ” had been announced, and widely accepted ; but in the organic world, the dogma of the miraculous creation of each separate species still held sway, almost as completely as when Linnaeus declared: “ There are as many different species as there were different forms created in the beginning by the Infinite Being.” But the dawn of a new era was already breaking, and the third period of palaeontology we may consider now at an end. Just twenty years ago, science had reached a point when the belief in “special creations” was undermined by well estab¬ lished facts, slowly accumulated. The time was ripe. Many naturalists were working at the problem, convinced that Evo¬ lution was the key to the present and the past. But how had Nature brought this change about ? While others pondered, Darwin spoke the magic word — “ Natural Selection ,” and a new epoch in science began. The fourth period in the history of Palaeontology dates from this time, and is the period of to-day. One of the main char¬ acteristics of this epoch is the belief that all life , living and extinct , has been evolved from simple forms . Another prom¬ inent feature is the accepted fact of the great antiquity of the human race. These are quite sufficient to distinguish this period sharply from those that preceded it. The publication of Charles Darwin’s work on the u Origin of Species,” November, 1859, at once aroused attention, and started a revolution which has already in the short space of two decades changed the whole course of scientific thought. The theory of “Natural Selection,” or as Spencer has hap¬ pily termed it, the “ Survival of the Fittest,” had been worked out independently by Wallace, who justly shares the honor of 41 tlie discovery. We have seen that the theory of Evolution was proposed and advocated by Lamarck, but he was before his time. The anonymous author of the “ Vestiges of Creation,” which appeared in 1844, advocated a somewhat similar theory which attracted much attention, but the belief that species were immutable was not sensibly affected until Darwin’s work appeared. The difference between Lamarck and Darwin is essentially this : Lamarck proposed the theory of Evolution ; Darwin changed this into a doctrine, which is now guiding the investi¬ gations in all departments of biology. Lamarck failed to real¬ ize the importance of time, and the interaction of life on life. Darwin, by combining these influences with those also suggested by Lamarck, has shown how the existing forms on the earth may have been derived from those of the past. This revolution has influenced Palaeontology as extensively as any other department of science, and hence the new period we are discussing. In the last epoch, species were repre¬ sented independently, by parallel lines; in the present period, they are indicated by dependent, branching lines. The former was the analytic, the latter is the synthetic, period. To-day, the animals and plants now living are believed to be genetically connected with those of the distant past; and the palaeontologist no longer deems species of the first importance, but seeks for relationships and genealogies, connecting the past with the present. Working in this spirit, and with such a method, the advance during the last decade has been great, and is an earnest of what is yet to come. The progress of Palaeontology in Great Britain during the present period has been great, and the general interest in the science much extended. The views of Darwin soon found acceptance here. Next to his discovery of “Natural Selection,” Darwin was fortunate in having so able and bold an expounder as Huxley; who was one of the first to adopt his theory, and give it a vigorous support. Huxley’s masterly researches have 6 42 been of great benefit to all departments of Biology, and bis contributions to Palaeontology are invaluable. Among the latter, his original investigations on the relations of Birds and Beptiles are especially noteworthy. His various memoirs on extinct Beptiles, Amphibians and Fishes, belong to the perma¬ nent literature of the subject. The important researches of Owen on the fossil vertebrates have been continued to the present time. He has added largely to his previous publications on the British fossil Beptiles, Birds, and Mammals; the extinct reptiles of South Africa, and the Post-Tertiary birds of Hew Zealand. His description of the Archaeopteryx near the begin¬ ning of the period was a most ’welcome contribution. The investigations of Egerton on Fossil Fishes have likewise been continued with important results. Busk, Dawkins, Flower and Sanford have made valuable contributions to the history of fossil Mammals. Bell, Gunther, Hulke, Lankester, Powrie, Miall, and Seely, have made notable additions to our knowledge of Beptiles, Amphibians, and Fishes. Among Invertebrates, the Crustacea have been especially studied by Jones, Salter, and Woodward. Davidson, Etheridge, Lycett, Morris, Phillips, Wood, and Wright have continued their researches on the Mollusks; Duncan, Hicholson and others have investigated the extinct Corals; and Binney, Carruthers, and Williamson, the Fossil Plants. Mumerous other important contributions have been made to the science in Great Britain during the present period. On the Continent, the advance in Palaeontology has, during the last two decades, been equally great. In France, Gervais continued his memoirs on extinct vertebrates nearly to the present date; while Gaudry has published several volumes on the subject that are models for all students of the science. His work on the fossil animals of Greece is a perfect monograph of its kind, and his later publications are all of importance. Lartet’s various works are of permanent value, and his applica¬ tion of Palaeontology to Archaeology brought notable results. The volume of Alphonse Milne-Edwards on fossil Crustacea 43 was a fit supplement to Brongniart and Desmarest’s well known work ; wliile his grand memoir on fossil Birds deserves to rank with the classic volumes of Cuvier. Duvernoy, Filhol, Hebert, Sauvage and others have also published interesting results on fossil vertebrates. Van Beneden’s researches on the fossil vertebrates of Bel¬ gium have produced results of great value. Pictet, Riitimeyer, and Wedersheim in Switzerland, Bianconi, Forsytli-Major, and Sismonda in Italy, and Nodot in Spain, have likewise published important memoirs. The extinct vertebrates have been studied in Germany by Von Meyer, Carus, Fraas, Giebel, Haeckel, Haase, Ilensel, Kayser, Kner, Ludwig, Peters, Portis, Maack, Salenka, Zittel, and many others; in Holland by Winkler, in Denmark by Reinhardt; and in Russia by Brandt and Kowalewsky. The fossil invertebrates have been investigated with care by D’Archiac, D’Orbigny, Bayle, Fromentel, Oustalet, and others in France; Desor, Loriol and Roux in Switzerland; Cappellini, Massalongo, Michellotti, Meneghini, and Sismonda in Italy, Barrande, Benecke, Beyrich, Dames, Dorn, Ehlers, Geinitz, Giebel, Giimbel, Feistmantel, Hagen, von Hauer, von Heyden, von Fritsch, Laube, Oppel, Quenstedt, Roemer, Schliiter, Suess, Speyer, and Zittel in Germany. The fossil Plants have been studied in these countries by Massalongo, Saporta, Zigno, Fiedler, Goldenberg, Gehler, Heer, Goeppert, Ludwig, Schim- per, Schenk, and many others. Among the recent researches in Palaeontology in other regions may be mentioned those of Blanford, Feistmantel, Lydekker, and Stoliczka, in India; Haast and Hector in New Zealand, and Krefft and McCoy in Australia; all of whom have published valuable results. Of the progress of palaeontology in America, I have thus far said nothing, and I need now say but little, as many of you are doubtless familiar with its main features. During the first and second periods in the history of palaeontology, as I have 44 defined them, America, for most excellent reasons, took no part. In tlie present century, during the tliird period, appear the names of Bigsby, Green, Morton, Mitchell, Rafinesque, Say, and Troost, all of whom deserve mention. More recently, the researches of Conrad, Dana, Deane, DeKay, Emmons, Gibbes, Hitchcock, Holmes, Lea, McChesney, Owen, Redfield, Rogers, Shumard, Swallow, and many others, have enlarged our knowledge of the fossils of this country. The contributions of James Hall to the Invertebrate Palaeon¬ tology of this country form the basis of our present knowledge of the subject. The extensive labors of Meek in the same department are likewise entitled to great credit, and will form an important chapter in the history of the science. The memoirs of Billings, Gabb, Scudder, White, and Whitfield are numerous and important ; and the publications of Derby, Hartt, James, Miller, Shaler, Rathburn, and Winchell, are also of value. To Dawson, Lesquereux, and Hewberry, we mainly owe our present knowledge of the fossil plants of this country. The foundation of our vertebrate Palaeontology was laid by Leidy, whose contributions have enriched nearly every depart¬ ment of the subject. The numerous publications of Cope are well known. Agassiz, Allen, Baird, Dawson, Deane, DeKay, Emmons, Gibbes, Harlan, Hitchcock, Jefferson, Lea, LeConte, He wherry, Redfield, St. John, Warren, • Whitney, Worthen, Wyman, and others, have all added to our knowledge of American fossil vertebrates. The chief results in this depart¬ ment of our subject, I have already laid before you on a previous occasion, and hence need not dwell upon them here. In this rapid sketch of the history of Palaeontology, I have thought it best to speak of the earlier periods more in detail, as they are less generally known, and especially as they indicate the growth of the science, and the obstacles it had to surmount. With the present work in palaeontology, moreover, you are all more or less familiar, as the results are now part of the current literature. To assign every important discovery to its author, 45 would have led me far beyond my present plan. I have only endeavored to indicate the growth of the science by citing the more prominent works that mark its progress, or illustrate the prevailing opinions and state of knowledge at the time they were written. In considering what has been accomplished, directly or indi¬ rectly, it is well to bear in mind that without palaeontology there would have been no science of geology. The latter- science originated from the study of fossils, and not the reverse, as generally supposed. Palaeontology, therefore, is not a mere branch of geology, but the foundation on which that science mainly rests. This fact is a sufficient excuse, if one were want¬ ing, for noting the early opinions in regard to the changes of the earth’s surface, as these changes were first studied to explain the position of fossils. The investigation of the latter first led to theories of the earth’s formation, and thus to geology. When speculation replaced observation, fossils were discarded, and for a time the mineral characters of strata were thought to be the key to their position and age. For some time after this, geologists, as we have seen, apologized for using fossils to deter¬ mine formations, but for the last half century their value for this purpose has been fully recognized. The services which Palaeontology has rendered to Botany and Zoology are less easy to estimate, but are very extensive. The classification of these sciences has been rendered much more complete by the intercalation of many intermediate forms. The probable origin of various living species has been indicated by the genealogies suggested by extinct types; while our knowledge of the geographical distribution of animals and plants at the present day has been greatly improved by the facts brought out in regard to the former distribution of life on the globe. Among the vast number of new species which have been added are the representatives of a number of new orders entirely un¬ known among living forms. The distribution of these extinct orders, among the different classes, is interesting, as they 46 are mainly confined to the higher groups. Among the fossil Plants, no new orders have yet been found. There are none known among the Protozoa, or the Mollusca. The Padiates have been enriched by the extinct orders of Blastoidea, Cystidea, and Edrioasterida; and the Crustaceans by the Eurypterida and Trilobita. Among the Vertebrates, no extinct order of fossil Fishes has yet been found; but the Amphibians have been enlarged by the important order Labyrinthodonta. The great¬ est additions have been among the Peptiles, where the majority of the orders are extinct. Here we have at the present date the Iehthyosauria, Sauranodontia, Plesiosauria and Mosasauria, among the marine forms; the Pterosauria, including the Pteranodontia, containing the flying forms ; and the Dinosauria, including the Sauropoda—the giants among reptiles; likewise the Dicynodontia, and probably the Theriodontia, among the terrestrial forms. Although but few fossil Birds have been found below the Tertiary, we have already among the Mesozoic forms three new orders: the Saururse, represented by Archce- opteryx / the Odontotormse, with Ichthyornis as the type; and the Odontolcse, based upon Ilesperornis / all of these orders being included in the sub-class Odontornithes, or toothed birds. Among Mammals, the new groups regarded as orders are the Toxodontia, and the Dinocerata, among the Ungulates; and the Tillodontia, including strange Eocene Mammals whose exact affinities are yet to be determined. Among the important results in vertebrate palaeontology, are the genealogies, made out with considerable probability, for various existing animals. Many of the larger mammals have been traced back through allied forms in a closely connected series to early Tertiary times. In several cases the series are so complete that there can be little doubt that the line of descent has been established. The Evolution of the horse, for example, is to-day demonstrated by the specimens now known. The demonstration in one case stands for all. The evidence in favor of the genealogy of the horse now rests on the same 47 foundation as the proof that any fossil bone once formed part of the skeleton of a living animal. A special creation of a single bone is as probable as the special creation of a single species. The method of the palaeontologist in the investigation of the one, is the method for the other. The only choice lies between natural derivation and supernatural creation. For such reasons it is now regarded among the active workers in science as a waste of time to discuss the truth of Evolution. The battle on this point has been fought, and won. The geographical distribution of animals and plants, as well as their migrations, has received much new light from Palae¬ ontology. The fossils found in some natural divisions of the earth are related so closely to the forms now living there, that a genetic connection between them can hardly be doubted. The extinct Marsupials of Australia, and the Edentates of South America, are well known examples. The Pliocene hip¬ popotami of Asia and the South of Europe point directly to migrations from Africa. Other similar examples are numerous. The fossil plants of the Arctic region prove the existence of a climate there far milder than at present, and recent researches at least render more probable the suggestion, made long ago by Buffon, in his “ Epochs of Nature,” that life began in the polar regions, and by successive migrations from them the conti¬ nents were peopled. The great services which Comparative Anatomy rendered to Palaeontology at the hands of Cuvier,’ Agassiz, Owen, and others, have been amply repaid. The solution of some of the most difficult problems in Anatomy has received scarcely less aid from the extinct forms discovered, than from Embryology; and the two lines of research supplement each other. Our present knowledge of the vertebrate skull, the limb-arches, and the limbs, has been much enlarged by researches in Palaeontol¬ ogy. On the other hand, the recent labors of Gegenbaur, Huxley, Parker, Balfour, and Thacker, will make clear many obscure points in ancient Life. 48 One of the important results of recent palaeontological research, is the law of brain-growth, found to exist among extinct mammals, and to some extent in other vertebrates. According to this law, as I have briefly stated it elsewhere: “ All Tertiary mammals had small brains. There was, also, a gradual increase in the size of the brain during this period. This increase was confined mainly to the cerebral hemispheres, or higher portions of the brain. In some groups, the convolu¬ tions of the brain have gradually become more complicated. In some, the cerebellum and the olfactory lobes have even diminished in size.” More recent researches render it probable that the same general law of brain-growth holds good for birds and reptiles from the Mesozoic to the present time. The Cre¬ taceous birds, that have been investigated with reference to this point, had brains only about one-tliird as large in proportion as those nearest allied among living species. The Dinosaurs from our Western Jurassic follow the same law, and had brain cav¬ ities vastly smaller than any existing reptiles. Many other facts point in the same direction, and indicate that the general law will hold good for all extinct vertebrates. Palaeontology has rendered great service to the more recent science of Archaeology. At the beginning of the present period, a re-examination of the evidence in regard to the antiquity of the human race was going on, and important results were soon attained. Evidence in favor of the presence of man on the earth at a period far earlier than the accepted chronology of six thousand years would imply, had been gradually accumulating; but had been rejected from time to time by the highest authori¬ ties. In 1823, Cuvier, Brongniart, and Buckland, and later, Lyell, refused to admit that human relics, and the bones of extinct animals found with them, were of the same geological age, although experienced geologists, such as Bone and others, had been convinced by collecting them. Christol, Serres, and Tournal, in France, and Schmerling in Belgium, had found human remains in caves, associated closely with those of various extinct mammals, and other similar facts were on record. 49 Boucher de Perthes, in 1841, began to collect stone imple¬ ments in the gravels of the valley of the Somme, and, in 1847, published the first volume of his “Antiquites Celtiques .” In this work, he described the specimens he had found, and asserted their great antiquity. The facts as presented, however, were not generally accepted. Twelve years later, Falconer, Evans, and Prestwich examined the same localities with care, became convinced, and the results were published in 1859 and 1860. About the same time Gaudry, Hebert, and Desnoyers, also ex¬ plored the same valley, and announced that the stone imple¬ ments there wfcre as ancient as the mammoth and rhinoceros found with them. Explorations in the Swiss lakes and in the Danish shell heaps added new testimony hearing in the same direction. In 1863, appeared Lyell’s work on the “ Geo¬ logical Evidences of the Antiquity of Man,” in which facts were brought together from various parts of the world, proving beyond question the great age of the human race. The additional proof since brought to light has been exten¬ sive, and is still rapidly increasing. The Quaternary age of man is now generally accepted. Attempts have recently been made to approximate in years the time of man’s first ajDpear- ance on the earth. One high authority has estimated the antiquity of man merely to the last glacial epoch of Europe as 250,000 years; and those best qualified to judge, would, I think, regard this as a fair estimate. Important evidence has likewise been adduced of man’s existence in the Tertiary, both in Europe and America. The evidence to-day is in favor of the presence of man in the Pliocene of this country. The proof offered on this point by Professor J.. D. Whitney, in his recent work,* is so strong, and his careful, conscientious method of investigation so well known, that his conclusions seem irresistible. Whether the Pliocene strata he has explored so fully on the Pacific coast corresponds strictly with the deposits which bear this name in Europe, may be a question requiring further consideration. * Auriferous Gravels of the Sierra Nevada of California. 18t9. 1 50 At present, tlie known facts indicate that the American beds containing human remains, and works of man, are as old as the Pliocene of Europe. The existence of Man in the Tertiary period seems now fairly established. In looking back over the history of Palaeontology, much seems to have been accomplished; and yet the work has hut just begun. A small fraction only of the earth’s surface has been examined, and two large continents are waiting to be explored. The “ imperfection of the geological record,” so often cited by friends and foes, still remains, although much improved; but the future is full of promise. In filling out this record, America, I believe, will do her full share, and thus aid in the solution of the great problems now before us. I have endeavored to define clearly the different periods in the history of Palaeontology. If I may venture, in conclu¬ sion, to characterize the present period in all departments of science, its main feature would be a belief in universal laws. The reign of Law, first recognized in the physical world, has now been extended to Life, as well. In return, Life has given to inanimate nature the key to her profounder mysteries— Evolution, which embraces the universe. What is to be the main characteristic of the next period 1 ~No one now can tell. But if we are permitted to continue in imagination the rapidly converging lines of research pursued to-day, they seem to meet at the point where organic and inorganic nature become one. That this point will yet be reached, I cannot doubt. i Tuttle, Morehouse & Taylor, Printers, New Haven, Conn.