uiMivtKSi I y 0. ILLINOIS LIBRARY AT URBANA CHAMPAIGN MATURAL HIST SURVEY ':^^'^y^^^-:ih: A Distribution and Abundance of Winter Populations of Bald Eagles in Illinois Stephen P. Havera and Glen W. Kruse Illinois Natural History Survey Biological Notes 129, February 1988 Cover Plioto: The bald easlc, Haluurlus kucocophalm. as photographed by Steven HoltyX'IREO. CONTENTS DESCRIPTION 3 DISTRIBUTION 3 POPULATION LEVELS 3 LIFE HISTORY 4 Reproduction 4 Mortality 5 LEGAL STATUS 5 BALD EAGLES IN ILLINOIS 5 Historical Records, 1200-1950 5 Recent Records 6 Statewide Winter Bald Eagle Counts 6 INHS Eagle Surveys 9 Upper Mississippi River Region 11 Upper Central Mississippi River Region 11 Lower Central Mississippi River Region 12 Lower Mississippi River Region 14 Upper Illinois River Region 14 Central Illinois River Region 15 Lower Illinois River Region 15 Central and Southern Lakes Region 17 Northeast Lakes Region 17 Surface-mined Lakes Region 17 Loxt'er Kaskaskia River Region 17 Other Inventories 17 DISCUSSION OF CENSUS DATA 18 NIGHT ROOSTS 20 RECENT BALD EAGLE NESTING 20 BALD EAGLE MORTALITY 21 SUMMARY 22 ACKNOWLEDGEMENTS 23 APPENDIX 24 LITERATURE CITED 28 Cilalniri: H.n.i.i, Si<|)Ikm I" . ,iiiii|>iil,inciiis oi b.ild (M^lcs in Illinois lllllinis N.lliu.ll IIIMOIN Sui\(\ Hic.lc>ni,,,l N.ilcs ]•>'.!. DISTRIBUTION AND ABUNDANCE OF WINTER POPULATIONS OF BALD EAGLES IN ILLINOIS Stephen P. Havera and Glen W. Kruse The bald eagle {Haliaeetus leucocephalus) is the only sea or fishing eagle that inhabits North America. Seven additional members of this genus occur in other parts of the world (Brown & Amadon 1968). Names sometimes applied to the bald eagle include American eagle, white-headed eagle, and Washington's eagle. Immature bald eagles have been called black eagles because of their dark color (Terres 1980). In recent years, the numbers of bald eagles in North America have rebounded frotn the critically low levels that resulted from loss of habitat, illegal shooting, and the widespread use of certain persistent pesticides. Illinois is currently an important winter area for bald eagles. DESCRIPTION Among the largest birds of North America, the bald eagle has an average body length of 34 to 43 in (85-109 cm) and a wingspan that is generally 6 to 7 ft (2.0—2.4 m). Robards (1967) reported wingspans of 78 to 93 in (1.98-2.36 m) for 39 eagles captured in Alaska. As in most raptors, female bald eagles are larger and heavier than males with the average weights of females ranging between 10 and 14 lb (4.5—6.4 kg) as compared with weights for males between 8 and 9 lb (3.6-4.1 kg) (Imler & Kalmbach 1955). Weights as high as 16.5 lb (7.5 kg) were reported by R()l)ar(ls (1967). Birds trapped in Minnesota weighed 8.() to 13.2 lb (3.9-6.0 kg) (Hennes 1985). Although different immature and subadult |)lu- mages of bald eagles occur, these plumages are mostly dark brown with irregular patches of white on the underside of the wings or elsewhere on the body (Clark 1983) (Fig. 1). For the purposes of this study, juvenile, immature, and subadult eagles were grouped into the immature age class because subtle differences in these plumages were diffuuit to discern during aerial inventories. The bill of an iiimiaiure is brown. the eyes are pale yellovv-giay, and ilu- feel .ut- lemon yellow. Immature bald eagles are distinguished from immature golden eagles by their larger heads, heavier bills, and unfeathered legs (Terres 1980). Thi.s paper is published by aiilhorily dI the Sialc i>l Illinois. It is a contribution of the .Section ol VVildlile Researc h o( the Natural History Survey. Dr. Stephen Havera is a Wildlilc K< oloj;ist at the Survey field station in Havana, (ilen Kruse. lornurly an .Assistant Research Biologist with the Survey, is now a I'rojed M.in.iK- Wikllilc Socifiy). A majority of eagles seen during the winter inventory was associated with the Mississippi River floodplain. Since 1973, nunil)ers of eagles tensused in Januaiv along the Illinois River usually ranged between 100 and 300, with a maximum of 383 observed in l'.)8(). Ihe percentage of all eagles censused by the I NHS during January that were immatures ranged between 16.5 and 46.5% (Table 3). In a majority of the years, the percentage of immature eagles varied between 30 and 37%. Southern (1963) cautioned, however, against using the proportion of immatures to adult eagles wintering in the Mississippi Valley bordering Illinois as a realistic estimate o( leprochu live suctess. Millsap (1986:436) re])orted that the jjercfiitage of imiTiature bald eagles observed on the midwintei sur- vey from 1979 to 1982 varied between 32.7 and 4 1 .2% . Sprunt (1965) reported that estimates of inimatuie bald eagle age ratios ranged between 21.(j and 26.5% from surveys (ondiKterl by the National AudulK)n Society from 1961 to 1963. The counts compiled by Elton Fawks were both aerial and ground (ounts (Table 3). F.ach count in- ( hided the Mississippi Ri\ei from I)ubu(|ue to "below St. Louis," the Illinois Rivei, and vaiying numbers and combinations of refuges and reservoirs. The Mis- sissippi River from (Irafton to C.aiio was added in 1976. The 1982 count also included the Fox. Wabash, Rock, and Ohio rivers. Fiom 1961 to 1978, the count was maove. but thev do not concen- tiatc- on .iic-.is neai large rivers where bald eagles are Illinois Natural History Survey Biological Notes No. 129 Southeast Alaska Northwest Interior Maine/ Nova Scotia I \\ Florida Breeding Area Represented by Bandings Fig. -4. -Major migraioi \ |)"l) Scrviii- 1986) Oilohii lliniii.jh M.mli li.iiul icmM-rics (IS. lisli .iii.l U ildlile most likely to occur. Areas covered in the Christmas bird counts are generally those surroimding the loca- tion of local Audubon Society chapters. Bald eagles observed on the Christmas bird counts ranged from 256 in 1974 to 767 in 1986 (Table 3). Differences in dates of counts, areas coveted, skills of observers, and weather conditions and the emphasis placed on counting eagles as opposed to other species create variability among the inventories (Table !5). Whereas counts for a single year vary widely, there are several instances when the changes between years are similar. Kxamiiiation of eagle numbers in Table 3 reveals that both the INHS and Fawks inventories show intreases from 1961 to 1962, decreases from 1962 through 1964, and increases fiom 1964 to 1965 and from 1977 tiirough 1979. Such similarities in trends suggest that patterns of changes in eagle nimi- bers in Illinois are real and demonstrate the impor- tance of more than one type of amnial count. Moving .3-year averages of the aiuuial winter February 1988 Havera and Kruse: Winter Populations of Bald Eagles Table 3. — Winter bald eagle counis (ondii couiu, 1957-1987. (1 \n the lllin()i> Natural Hisiorv Survey. KUon hawks, and the .Audubon Christmas bird INHS Fawk s'' Audubon''''- ^ear .\duli liiiinalure lolal .•\(luli Itninaturc L'nascd lolal .^duh Immature Unaged total 19.-) 7 359 19.59 207 1960 (i3 1961 142 30 172 365 51 1 417 1962 143 44 187 417 62 29 508 1963 108 64 172 331 91 7 429 1904 68 30 98 226 58 130 414 196-) ".■) 52 127" 420 160 44 624 1966 486 122 51 659 1967 485 175 28 688 196H 84 37 121'' 415 174 50 639 1969 391 261 3 655 1970 112 76 188 1971 100 40 140 406 159 11 576 1972 75 19 94 .562 302 31 895 158 95 29 282 1973 364 175 539 529 231 14 774 211 115 14 340 1974 398 153 551 413 128 8 549 140 113 3 2.56 1973 365 201 566 519 274 32 825 304 182 110 596 1976 489 270 759 459 214 33 706 287 269 6 562 1977 327 144 471 508 264 29 801 405 252 11 668 1978 369 166 535 69() 219 52 967 503 •247 11 761 1979 410 203 613 1,018 356 30 1.404 244 149 10 403 1980 514 288 802 931 4()(i 57 1 ,394 384 203 7 594 1981 454 304 758 228 153 5 386 1982 517 271 788 888 380 28 1 .296 211 128 3 342 1983 365 178 543 292 256 17 565 1984 613 317 930 316 218 3 537 198.5 404 247 651 420 235 12 667 1986 779 438 1.217 443 313 11 767 1987 368 201 569 ■'Kawks (1983). ''Struihcrs (1973. 1974, 1975, 1976, \'.r, 'Stroyls (1981, 1982, I9H3, 1984, 1985. ' Incomnlele count. 7, 1978, 1979, 1980). 1986, 1987). counts are presented in Figure 5. Each of the three inventories shows a general trend toward higher num- bers in recent years. As noted by Fawks ( 198S), weatli- er and other factors that interfere with ac( urate counts are of lesser importance if data interpretation is based on multi-year averages rather than one-year totals, (founts by the Illinois Audubon Society were probably also influented by changes in geographical coverage between years. INHS Eagle Surveys. The most extensive record o( bald eagle populations in Illinois was compiled by the INHS. Kagles were toiuited onte eat h year in I'C)?, H)59 to \9(]5, 1968, 1970, and 1971 (Table :<). Sinte I97'2, several counts were made each year in 1 1 regions of the slate (Table 4, Fig. 6) and provide de- tailed information. Riverine regions inventoried in- cluded the river pioj^er and accom])anying wetlands in the iloodplain. (Censuses conducted in the Missis- sippi River iloodplain included eagles seen in Iowa and Missouri. Kach region wasdi\idc-d into a muiiber of census areas for recording data. Individual .ueas inc iuded river segments def'med by pioiniiicni lanil- iti.irks, lakes, bays, marshes, sloughs, oi ollit-i wetland aieas. The census aieas included in each ic-gion ate listed in the Appendix. The numbei ol miles of river and the area of wetlands in each riverine region are presented in Table 5. The number of years in whic h aerial censuses were taken in a region and the niutiber of flights made each year were dependent upon availability of funds 1500 1200 1100 1000 o 800 "" 700 $ 600 O •" 500 <00 1- INHS ) - FAWKS i - AUDUBON 1977, 1986, I960 IMZ IM4 !««« IMS l«70 1972 1*74 1976 1978 IMO 1962 19*4 19*6 YEAR 1 liiii-vrai m<>\iii){ .ivei.igrs ol wiiiKi eagle ciHiiUs ill II- hoiii Illinois N.iim.il llislory Siirxcy data, lawks ( 1983), and linois .\iidulM.n So(ici\ (Siniiliers 1973, 1974, 1975. 1976. 1978. 1979. 1980; Siioyls 1981, 1982. 1983. 1984. 198.5, 1987). Illinois Natural History Survey Biological Notes No. 129 Table 4. — Regions aerially iinentoried for bald eagles by the INHS and the Irequeno, approximate period, and years of censuses in each region. Region Upper Mississippi Riyer (Dubuque— Interstate 80) Upper Central Mississippi Riyer (Moline— Keokuk) Lower Central Mississippi Riyer (Keokuk-Alton, lower 26 miles of Illinois Riyer) Lower Mississippi Riyer (St. Louis-Cairo) Upper Illinois River (Morris-Spring Valley) Central Illinois Riyer (Spring Valley-Lower Peoria Lake) Lower Illinois River (Pekin Lake-Naples) Surface-mined Lakes (Fulton, Knox counties) Northeast Lakes (natural lakes, cooling lakes, reservoirs in Cook, De Kalb, Grundy. Lake, La Salle. McHenry, Will counties) Central and Southern Lakes (Reservoirs: Carlyle, Rend, Shelbyville, Springfield. Cooling lakes: Baldwin, Clinton, Coffeen, Newton, Sangchris) Lower Kaskaskia River (Fayetteville-Chester) ■'Period lengths varied due to availability of llight tunds and on weather (Table 4). The number of aerial inven- tories, mean number of bald eagles observed per in- ventory, and maximum number of eagles seen on a single inventory for each census area for all census years are presented in the Appendix. The mean number of bald eagles sighted per inventory flight for all years was used as the indicator level of eagle Frequency of Inventories .•\pproxinialc Inventory Period'' Biweekly October- March 1980-1983 Weekly Sept.-Dec, Jan., Feb.-April I972-I986 Weekly Biweekly Sept-Dec. .Jan.. Feb.-April October- April 1972-1986 1972-1986 Biweekly Weekly Weekly Biweekly September- April Sept.- Dec. .Jan., Feb.- .April Sept.-Dec. .Jan., Feb.-April October- March 1976-1985 1972-1986 1972-1986 1980-1985 Biweekly Biweekly Biweekly September- .•\pril October- April October- April 1972-1986 1982-1986 Upper Cenlfal Mississippi "^ Sufface-mired Lower Central Mississippi Northeast Lakes Central and Southern Lakes use for a given area. The number of inventories and the mean number of eagles observed per inventorv were based on the period between the first sighting of a bald eagle in the fall of each year and the last eagle sighting the following spring, and not the entire census period from fall through spring. This proce- dure appropriately concentrated on the period when eagles were present in the various regions. The arrival and departure dates of bald eagles in Illinois vary each year as a result of differences in weather, freeze- ups, and other unpredictable phenomena of nature. Differences in the length of the inventory period and the number of years of censuses should be taken into consideration when making comparisons in values for the mean number of eagles coimted per inventory among regions (Table 4). For instance, the Upper Mississippi River Region was inventoried din- ing 1980-1983 as compared with 1972-1986 for the other Mississippi River regions. Bald eagles ha\e gen- erally increased in Illinois since 1972 (Table 3) and a mean value for the number seen per inventory in more recent years (1980-1983) mav pn)vide a differ- ent value than a mean calculated on data from 1972- Table 5. — River miles and wet and area inch ded in riverine census regions. R vcr Mile ,;j Wetland Area'' Region mi^ km- Upper Mississippi River 8.5.4 122.9 318.3 Upper Central Mississippi Ri\e i;n.:i 168.5 436.4 LowerC'entral Mississippi Rive 187.1 265.3 687.1 Lower Mississippi River 202.9 242.2 627.3 Upper Illinois River 57.0 24.6 63.7 Central Illinois River 57.8 82.4 213.4 Lower Illinois River l:l6.2 154.6 400.4 Lower Kaskaskia River :«).o 22.5 58.3 Tig. 6. -Regions ol llliiHiis inventoried lot bald e.igles In the lllii Natuial HiMnn Sui\<\. 1972-1987. •'U.S. .Army Corps of Fngineers (1978). Illinois Division of W atcr- wavs (I9(=i9). ''H.igen et al. (1977). Haveia (1985). February 1 988 Havera and Kruse: Winter Populations of Bald Eagles 1986. However, the mean number of eagles seen per inventory was the most logical method of presenting the data. Upper Mississippi River Region. The Upper Mis- sissippi River Region extends from the Illinois- Wisconsin border southward to approximately Rock Island, Illinois (Fig. 6). This region was surveyed from October through March during the three winters of 1980-1983 and substantial numbers of eagles were counted. The floodplain in this area provided attract- ive winter habitat for bald eagles (Fig. 7). Fig 7 - 1 vpual iivcniie habitat of the L'ppcr Mississi] gion. The seasonal pattern of eagle use of the Upper Mississippi River Region is shown in Figure 8. Annual peak population counts occurred between 2 February and 23 March. Substantial numbers of birds were still present when counts were terminated each spring. The greatest 1-day count was 213 eagles on 23 March 1983. Southern (1964) observed peak numbers of 268 and 225 eagles in this region on 2 February 1962 and on 5 February 1963, respectively. A total of 2,663 eagle sightings was recorded in this region; 34.99? of those were immatures. The proportion of immature ISO ■ 140- 130 - 70 ■ 60 ■ 50- iO ■ IMMATURES TOTAL birds ranged from 30.9 to 4 1 .69c on individual census flights. The mean count per inventory on individual cen- sus areas is shown in Figure 9. The area between the Wapsipinicon River and Princeton was the most heav- ily used in this region, with an average of 20.9 eagles sighted per inventory (Appendix). Other heavily used areas were the Bellevue to Green Island segment and the Fulton to Albany area. The Upper Mississippi River Region also includes the Oak Valley F.agle Ref- uge in Rock Island Clounty. The heavy use of this area by wintering bald eagles was documented by Klton Fawks through 25 years of observation and because of its importance, the refuge was purchased by the Illinois Department of Conservation (IDOC) in 1980. For the Upper Mississippi River Region as a whole, bald eagles occurred at an average density of 1.38 eagles per river mile or 0.96 eagle per square mile of wetland habitat from 1980-1983. These values are higher than those for any other region of the Missis- sippi and Illinois rivers; however, the Upper Missis- sippi River Region has been flown for fewer years than the other regions (Table 4, Fig. 6). Upper Central Mississippi River Region. The Upper Central Mississippi River Region stretches from ap- proximately Rock Island, Illinois, to Keokuk, Iowa (Fig. 6). Between 1972 and 1986, most areas within this region were surveyed from 120 to 130 times. Flights were made from September through De- cember in 1972 to 1985, during Januarv from 1973 to 1986, and from February through .April from 1980 to 1985. Lodi and Dam 16 15 29 12 26 lig. H.-Mcaii iiuinlur oi bald eagles (oiiiiliil |>< i Ippcr Mississipi Rivir Region. HIHO-HW:! Kig.O.-Mean luiinherof halrleagleMoiinteii pel invenloiv.iltensiis areas KJlhiii ihe I'ppei Mississippi Rlvei Regiriii. Htmt-Ht.S:V Illinois Natural History Survey Biological Notes No. 129 The level of eagle use in the Upper Central Missis- sippi River Region from fall through spring is shown in Figure 10. Few inventories were conducted during late January and early February. In each year except 1983, at least one inventory was conducted in the fall before any eagles were observed and those flights al- lowed an approximation of the arrival date of bald eagles in this region. Estimates of arrival dates of eagles observed from the air for 1 1 autumns indicated that arrival gradually became earlier. From 1972 to 1975 the first eagle sightings were in late November, but from 1976 to 1981 the first observations usually occurred in late October or early November. In 1982 and 1983 eagles were present in the Upper Central Mississippi River Region by late September. Spring data indicated that all eagles usually left this region by early April. Fig. 10. -Mean tuiiiiIki of b.ikl califs lounU'd per imciildi v in tlu- tapper Ck-nlr.il Mississippi Ri\ir Region, lT OCT NOV DEC JAN FEB MAR APR DATE Me.in luinibc'i nl b.ild e.igles loiniled pel in\enloi\ in the enli.il Mississi|)pi River ll h.iUI eagles iiMnilecl pel iruentiii\ M wilhin llie lower ( enli.il Mississippi Rivei Regiciii. 1 ig l.'i. Nciiilusesiward view of llie Ceil.ii (ilen K H-iiuixk (iiiiiiiiv. Illinois, and llie f1(M)d|)laiii l>cl10,00 Fig. 17.— Mean niniiljer ol bald eagles counted per inventory at census areas within the Lower Mississippi River Region, 1972-1986. or 0.20 per square mile of wetland habitat. These density values of bald eagles were the lowest of all the Mississippi Ri\er regions. Upper Illinois River Region. The L^pper Illinois River Region stretches from Morris, Illinois, to near Spring Valley, Illinois (Fig. 6). Most portions of this region were surveyed 59 times between September and .-Vpril from 1976 to 1985. This relativelv short, highly de- veloped segment of the Illinois River vallev seldom showed substantial use bv bald eagles. The seasonal pattern of eagle numbers in the Upper Illinois Ri\er Region is shown in Figure 18. The trend toward earlier observations of eagles in the fall during the years inventoried was also apparent in this legion. In the late 1970s, eagles were usually first seen during aerial inventories in the latter half of November, but in more recent years, the first observa- tions generalh occurred in mid-October. The earliest sighting ot bald eagles during any census in all regions (F"ig. 6) occurred in the LIpper Illinois Ri\er Region when four birds were seen between L'tica ami Mar- seilles on 10 September 1980. Spring departure ot eagles from the L'i)per Illinois Ri\er Region was gen- erall\ completed b\ earK .\piil. February 1988 Havera and Kruse: Winter Populations of Bald Eagles IS I 15 29 12 DEC JAN Fig. 1 8. -Mean numbci of bald eagles counted per invenloiv in ihe Upper Illinois River Region. 197(5-198.5. Seasonal patterns of hald eagle octiirrence in the Central Illinois Rivei Region are ilhistiated in Figure 20. No census tliglits were toiuhuied from mid- Peak population counts occurred between 3 Janu- ary and 9 March in this region with the highest count of 53 eagles reported on 20 January 1982. Generally, fewer than 25 eagles were seen on each census in this region. Immature birds consisted of to 50.0% of the eagles observed on individual inventories, and the av- erage proportion of immatures for all eagles seen was 31.8%. The mean number of bald eagles observed per inventory for individual areas is depicted in Figure 19. The downstream portion of this region, between La Salle, Illinois, and Spring Valley, Illinois, was gen- erally used the most by eagles (Appendix). The density of bald eagles in the Upper Illinois River Region av- eraged 0.20 per river mile or 0.47 per square mile of wetlands. Fig. 19. -Mean nnnilxi ol bald eagles (ounled [)er inventory at census areas within the L pper Illinois River Region. 1976-198.5. Central Illinois River lief^on. The Central Illinois River Region consists |)rimarily of the Peoria Naviga- tion Pool and extends from near .Spring Valley to Lower Peoria Lake (Fig. 6). This region was inven- toried approximately 125 times between 1972 aiif the La (iiange Pool of (he Illinois River and stretches lioin Pekin to Naples (Fig. <>). I his region has been more t loselv monitored for eagles than anv other with over 1.30 inventories l.iken liom 1972 to 1980. (^c'lisus lliglils \kv\v made fluiing Seplembei lliiough Deteiiibei from 1972 to 1985. during |.iiiuai V lioiii 197!^ I o 198(1. .tnd tlui ing Februarv through .\piil from 1980 to 19M5. Seasonal variation in eagle use ol this region is sliiivMi in Figure 23. No i ensus flights were tonclui led I loin mid- 1. Ill t I.I I V to l.ilc Icbi ti.iiv . I imesof f.illai i iv- linois Natural History Survey Biological Notes No. 129 <1.00 1.00-2.99 I 3.00-4.99 ) 5.00-9.99 Fig. 21. -Mean number of bald eagles tountecl per inventory at cen.sus areas within the Central Illinois River Region, 1972-1986. Fig. 23. -Mean number of bald eagles counted per Lower Illinois River Region. 1972-1986. to 50.09^ of tlie total eagles observed, and the overall proportion of immatiires was 34.79^ for all inven- tories. The mean number of eagles inventoried per cen- sus flight is illustrated in Figure 24. ,\reas with the most use by bald eagles intluded Chautauqua National Wildlife Refuge (Fig. 25), Crane Lake, Clear Lake, Cuba Island, and Jack Lake (Appendix). .\11 of these areas are privately owned except the 4,500-acre (1,822-ha) Chautauqua Refuge. al in late September and early October and spring dejiaiture by eailv \pv\\ were similar to those in the Cential Illinois Ri\er Region. Eagles arri\ed in the Lowei Illinois Ri\er Region approximately 1 week earlier than in the Cential Illinois River Region in 2 of the 15 years inventoried. Peak population counts have occurred from 2 De- cember until 18 March; the latter is also the date of the highest 1-day count of .345. The proportion of immatures seen on indixidual censuses ranged from Fig. 24. -Mean number ol b.iUl eagles i census are, IS uilliiii llie Lower Illinois Ri' • 100-299 % 3.00-4-99 innied per in\enior\ a er Region. 1972-1986. Fig. 22.-Fastern sh, into Senacliw'ine L.tk The estimated density of bald eagles in the Lower Illinois River Region averaged ().5(i per river mile or 0.49 per squaie mile of wetland habitat. The average density per scjuare mile of wetlands (0.49) in this re- gion was slighth higher than the value lor the ('entral and L'pj)ei Illinois River regions (0.47) and was also highei tlian the values for all of the Mississippi River February 1988 Havera and Kruse: Winter Populations of Bald Eagles Fig. 25.-Northcast\vai(l \icw ol the Chautauqua Lake area, an important winter site for bald eagle.s in the Lower Illinois River Re>;ion. regions (0.20-0.36), with the exception of the Upper Mississippi River Region (0.96). The average density of eagles per river mile (0.56) in the Lower Illinois River Region was slightly lower than the value for the Central Illinois River Region (0.67), higher than those values for the Upper Illinois (0.20), Lower Mississippi (0.24), and the Upper and Lower Central Mississippi regions (0.46 and 0.48, respectively), but substantially lower than the value (1.38) for the Upper Mississippi River Region. Central and Southern Lakes Region. A vast majority of bald eagles wintering in Illinois was found in the floodplains of the Illinois and Mississippi rivers (Figs. 6 and 26). Relatively small numbers of eagles were counted in other regions. Of the nine reservoirs and cooling lakes censused in central and southern Illinois (Fig. 6, Appendix), five (Sangchris, Carlyle, Rend, Baldwin, and Springfield) were inventoried on a reg- ular basis from October through April since 1972. However, only Carlyle and Rend lakes averaged more than one eagle per census (Fig. 26, Appendix). No eagles were sighted at Clinton Lake. Kagje use of these lakes in central and southern Illinois declined in recent years. Northeast Lakes Region. The natural lakes, reser- voirs, and cooling lakes in counties of northeastern Illinois (Fig. 6, Appendix) alsf) hosted few eagles. These lakes were censused from September through April since 1976. Lhe M( Henry Dam area in Mc Henry County was the only area that held an average of tnore than one eagle per census (Pig. 26, Appendix). Kaglc numbers were slighilv higher in the late 1970s in this region, but fell to near zero iti later years. Surffiir-wnirrI Litkc^ Rtgiini. Inveiitoi ies ol surface- mined lakes in Fulton and Knox (onniies in west- central Illinois (Fig. 6) reveali-d limited use bv bald eagles (Fig. 26). Inventories were conducted from Oc- tober tbiough Match frotn 1980 to I98.T. No eagles were sighted r)n most of the lakes in this r<-gion bul small numfjers were seen on a regular basis on the Central Illinois Light Company (CILCO) property (Appendix). Lower Kaskaskia River Region. The Lower Kaskaskia River Region (Fig. 6) included the lower 30 miles of the river from approximately Fayetleville to Chester on the Mississippi. This region was surveyed during October to April from 1982 to 1986 (Fig. 26). From 1982 to 1985 bald eagle use in this region was low with never more than one eagle sighted on a census; however, on 7 January 1986, nine eagles were ob- served (Appendix). Other Inventories. Other studies that monitored bald eagle populations over one or two winters in small geographic areas have been conducted in Il- linois. Such studies, because of the frequency of counts and the limited area involved, provided information on the abundance and use patterns of bald eagles that is not obtainable from periodic regional counts. Southern (1963) monitored eagle populations near the Savanna Army Depot in Jo Daviess County in the * •lOO • 1 00 2 99 • 3 00 4 99 • 5 00 9 99 • •10M liK 'Jli.~M<'.in niiinlH'i nl h.ilil ('.i|{k'> xiunlcd (mt invrnl(ii\ al (ensiis areas within all i enrolls. 1972-UINf). Illinois Natural History Survey Biological Notes Upper Mississippi River Region (Figs. 8 and 9) during the winterof 1961-1962. He reported that eagles were present in that area from before 9 December until 1 April. The peak population count occurred in this area on 2 February when 262 birds were counted along a 14-mile census route. A second study by Southern (1964) was done in the same area during the winter of 1 962-1 963. Counts were initiated earlier in the fall and the first eagles were sighted on 2 1 October. After a peak of 225 birds on 5 February, all eagles had left the area by 9 April. Through most of the winter, only about 10 to 12% of the bald eagle population in this area were imma- ture birds. Jonen (1973) studied bald eagles on a 6-mile reach of the Mississippi River extending downstream from Lock and Dam 19 at Keokuk (Figs. 12 and 13). In the winter of 1970-1971 he first noticed an immature bald eagle in this area on 3 October. The first adult eagle was sighted on 13 December and the peak popu- lation count for that winter was 49 (40 adults, 9 imma- tures) on 8 January. Jonen sighted the first eagle, an immature, during the 1971—1972 season on 18 Oc- tober. An adult eagle was first sighted on 29 November, and the largest niunber of eagles sighted was 66 (57 adults, 9 immatures) on 12 February. Two patterns noticed by Jonen (1973) were that immature eagles were present in the area earlier in the fall and later in the spring than adults and that the number of eagles present in his study area in- creased during periods of extremely cold weather. Fischer (1982) studied the same area described by Jonen (1973) and found that bald eagle use of this area had increased dramatically in the 7 years since Jonen's work. During the winter of 1978-1979, which was a winter of record-breaking low temperatures and heavy snowfall, Fischer (1982) monitored eagle num- bers from early December to early April. A peak count of 454 eagles (359 adults, 88 immatures, 7 not aged) occurred on 18 January 1979. The count far exceeded any recorded previously in the vicinity of Lock and Dam 1 9 and was not equaled in the following 6 years. Fischer (1982) also made inventories from late November 1979 until the end of March 1980. During a relatively mild winter, the peak number of eagles counted was 120 (89 adults, 31 iinmatures) on 18 Feb- ruary 1980. The peak for 1979-1980 was only 26.4% of the peak count for the previous winter. Harper (1983) studied eagle populations in the vicinity of Lock and Dam 24 at Clarksville, Missouri, in the Lower Central Mississippi River Region (Figs. 6 and 13). He conducted aerial and ground sinveys in 1980—1981 and ground counts in both foraging and night roosts during the winterof 1981-1982. Aer- ial counts conducted in 1980-1981 gave consistently lower counts than ground counts done simulta- neously. Daytime ground counts usually provided lower totals than counts done on the same dale at night roosts. I he highest count recorded during this study was 168 eagles (92 adults, 57 immatures, 19 unaged) observed at a night roost on 26 January 1982. Sabine and Klimstra ( 1 985) inventoried bald eagles in southern Illinois at Union County and Horseshoe Lake Conservation areas during 1979-1981 and at Crab Orchard National Wildlife Refuge (CONVVR) during 1980—1981. Eagles are probably attracted to these areas by the large numbers of wintering Canada geese because crippled and dead geese provide a read- ily available source of food for wintering eagles. Eagles arrived in these areas in late October and departed by early March. Peak population counts for eagles in 1979—1980 were 71 at Union County on 19 February and 78 at Horseshoe Lake on 8 February. During 1980-1981, peak populations of eagles occurred on 23 January when there were 79 at Union County and 77 at Horseshoe Lake. Approximately 76% and 65% of the peak populations of eagles were immatures at Union County and Horseshoe Lake, respectively. These values for immatures are much higher than those noted for areas along the Illinois and Mississippi rivers (Jonen 1973, Fischer 1982, Havera et al. 1984). CONWR was used by relatively few eagles as shown by a peak count of only 29 in mid-December 1980. Southern et al. (1985) conducted 12 statewide aer- ial surveys of bald eagles during the winter of 1984— 1985. This was the first attempt at 1-day statewide censuses. All previous studies included 1-day counts in relatively small areas (Jonen 1973, Sabine 1981) or counts over large areas that were completed in a period of days (I NHS aerial surveys). In addition to documenting changes in bald eagle populations in Illinois, which peaked at 989 on 8 Feb- ruary 1985, Southern et al. (1985) identified 13 essen- tial areas of winter habitat using criteria suggested by Crier et al. (1983). These included any area of 15 mi (24 km ) or less in length along a river that was used by 15 or more bald eagles for 2 weeks or longer. Southern et al. (1985) found that the most heavily used winter area was that portion of the Mississippi River near Lock and Dam 19 at Keokuk. In the Illinois River valley, the area used most by bald eagles was Pere Marquette State Pai k in Jersey County. .Ml areas found to be essential winter habitat by Southern et al. (1985) are shown in Figure 27. Small concentrations of bald eagles sometimes occur away from the floodplains of major rivers. For example, between 20 and 25 bald eagles were seen frequenting a wooded area of Pike Countv during the 1986-1987 winter. These eagles were feeding on live- stock carcasses and were roosting close to the food source and not in the Illinois or Mississippi river fioodplains. DISCUSSION OF CENSUS DATA Although a concentrated effort has been made to monitor the numbers of bald eagles wintering in II- February 1988 Havera and Kruse: Winter Populations of Bald Eagles 19 Clarksvllle (12) Horseshoe Lake (1 0) \© Fig. 27.-I.<)cali<)iis of essential bald eagle wiiilei li.iliiMi Kliriiilied by Southcin cl al. 1985. Numbers inditale iin|joilaiin- rankiiiH- linois, bald eagle census data for Illinois should be considered anestimateor index of actual pojjulalions. For a variety of identified and imdeteriTiined reasons, counts of bald eagles made on a specific area are subject to extreme short-term variations. This vari- ation is well demonstrated by data from Fischer (1982). In his study, two or three ground counts often were made on the same day, and those counts differed by as much as 35% or 150 eagles. However, variations in the numbers of eagles seen were not consistent. Numbers of eagles sometimes increased or decreased during the same day. I'he lack of sinuiltaneous counts over a wide area and the mobility of bald eagles make it diffic ult tf) determine whether daily flue luations in numbers were c aiised by lo( al movements or bv shifts in populations over larger geograjjhic aieas. Dunstan and Fawks (19HI) stated that food avail- ability is the primary reason (or the southwaid move- ment of bald eagles into Illinois during wintei. Keister et al. (1987) reported that the interaction of food avail- ability and proximity of loosting habilal appealed to be a primary factor in the behavior of wintering bald eagles. Structures such as locks and dams or activities that maintain open water in winter and, therefore, the availability of fish serve as an attractant to bald eagles, partic ularly if suitable daytime perches and night roost areas are nearby. As noted by Steenhof (1978), the abundance and distribution of wintering bald eagles have been greatly af fee ted by construe tion of dams and impoundments. Fischer (1982) found that the extent of ice cover on the Mississippi River influenced the local distribution of eagles. As the area of open water changed, the eagles shifted their pat- terns of use to take advantage of available forage. Harper (1983) reported similar shifts in bald eagle use patterns near IakV. and Dam 24. Jonen (1973) and Fischer (1982) reported a corre- lation between extremely cold temperatures and in- creasing numbers of bald eagles present on the Lock and Dam 19 study area. An increase in eagle numbers generally occurred a few days after the onset of a cold period. However, there was no apparent relationshi]) between population changes of eagles and low temper- atures over the large geographic areas sinveyed aer- ially by I NHS, a finding which suggests that the re- lationship between movement and temperature noted by Jonen (1973) and Fischer (1982) could be a local phenomenon. Millsap (1986) found that bald eagles appeared to winter farther south during unusually cold winters. Southern (1963, 1964) and others suggested that immature bald eagles move south earlier and move farther south than adults. Millsap (1986) found that the percentage of immatine bald eagles in the mid- winter survey increased at southern latitudes. Both jonen (1973) and Fischer (1982) reported sighting a higher proportion of innnatutes early in the winter than during midwinter. Fischer (1982) also noted an increase in the proportion of innnatures as spring approached. The largest proportions of immatures reported in Illinois were at wildlife areas in southern Illinois (Sabine & Klimstra 1985). These observations support the concept that immature bald eagles move soiuhward earliei and farther than adults. IXHS cen- sus data collected over lelalively large areas did not reveal such patterns of movements by immature bald eagles. The accuracy of various methods of counting eagles has been debated. Southern (19(")4) reported that simultaneous ground and aerial counts gave simi- lar results; he suggested that aerial counts validated the accuracy of ground counts. Harper (198.3) found that aerial surveys gave consistently lower counts th.in ground coinits in the same area. Havera et al. (1984) csiimaicd ihai 75 to 90'^ of the eagles present were counted during aerial survevs. Density of vegc-talion could easily affect the accii- lacy of observeis when counting eagles from the ground or the air. Kven inider good conditions, imma- ture bald eagles can be difficult to locate because of their moiilccl daik |)luTiiagc-. However, weather mav Illinois Natural History Survey Biological Notes have the greatest influence on the accuracy of bald eagle counts. Fog, rain, snow, and wind not only make it difficult for observers to see and age eagles but such conditions also may influence the activity of eagles, thereby making them less visible. Difficulties in obtaining accurate determinations of eagle numbers in Illinois do not eliminate the value of such counts. The counts can identify important winter areas and document trends in bald eagle popu- lations. The bald eagle inventories organized by Elton Fawks, for example, were prompted by concern over the low proportion of immatures seen in winter popu- lations during the 1950s (Fawks 1983). After quickly increasing from 12.3% in 1961 to 27.6% in 1965, the proportion of immatures sighted on Fawks' counts has fluctuated slightly around a value of 30%. As with any migratory species, suitable winter habitat is essen- tial to the survival of individuals and the subsequent reproductive success of the population. NIGHT ROOSTS Night roosts generally offered seclusion from human disturbance and protection from cold winds. At many of the roost sites, a windbreak was provided by the walls of a ravine or glen. On level areas, eagles selected trees for roosting that were protected by other trees (Fig. 28) (Fischer & Osterfeld 1981, Sabine & Klimstra 1985). Cotton woods (Populus deltoides) were the preferred roost tree in some areas, and it appeared that size and configuration of available trees were im- portant. Bald eagles generally selected trees that were taller than surrounding ones and had large, horizontal branches (Sabine & Klimstra 1985). Although bald eagles perch and forage along major stretches of the Illinois and Mississippi rivers, relatively few areas have been identified and reported as night roosts (Fig. 29). Other undiscovered or unre- ported night roosts undoubtedly exist along the Il- linois and Mississippi rivers. Dunstan and Fawks (1981) identified and described six night roosts as fol- lows: 1) The Savanna Army Depot and nearby islands include about 13,000 acres (5,260 ha) in Jo Daviess and Carroll counties. Four specific night roost sites have been identified on the area. There are also recent records of nesting attempts on Depot property. 2) The Oak Valley Eagle Refuge in Rock Island County is owned and managed by the NWF and IDOC. There are four wooded ravines on the refuge that were used by roosting eagles. 3) Cedar Glen Eagle Roost (Fig. 15) is located in Hancock County between Hamilton and Warsaw. The roost and surrounding eagle foraging areas have been purchased and protected by a cooperative effort of the Illinois Chapter of The Nature Conservancy, IDOC, and Western Illinois University. 4) Pere Marquette State Park is located near the confluence of the Illinois and Mississippi rivers in jer- sey County. The area around the park provides forag- Kig. 28.— Crystal fllen Eagle Roost in Hancock ("oiintv near Lock and Dam 19 on the Mississippi River. The sycamore {Plaluniis oc- inlniliilis) in the foreground was used by as many as 20 eagles nightly during January and February, 1979 (David L. Fischer, per- sonal communication 1986). Houses built nearby during the sum- mer of 1979 discouraged subsequent eagle use of this glen as a night roost (Photo by David L. Fischer). ing and resting habitat, but most night roosting occur- red in three hollows within the park. 5) Clarksville Island is located in the Mississippi River along Calhoun County. The island, owned bv The Nature Conservancy, was used by eagles through- out the winter. 6) Burlington, Craigel, and Otter islands are lo- cated in the Mississippi River along Henderson County and each has a night roost. A variety of habitats appropriate for daytime use by bald eagles is available in the area. Very few night roost areas along the Illinois River have been identified, mainly because of a lack of re- search. Robert Phelps, John Grigsby, and James Waycuilis (personal communications 1986) reported roosts in Putnam County near the Illinois River, at Rice Lake in Fulton County, and at Marshall County Conservation Area, respectively (Fig. 29). Additional night roosts likely occur in the vicinity of foraging areas such as Goose and Senachwine lakes. Peoria Lake, Chautauqua National Wildlife Refuge, and San- ganois Conservation Area. Roost areas were also iden- tified at Union County and Horseshoe Lake Conser- vation areas and at CONWR (Sabine & Klimstra 1985) (Fig. 29). RECENT BALD EAGLE NESTING Bald eagles have constructed seseral nests in Il- linois since 1973, and at least 16 eaglets fledged from these nests from 1978 to 1987 (Fable 6). The first rc( cut bald eagle nest in Illinois was built at COXWR in 1973. Eagles were seen at this nest again in 1974. Inil no eggs were laid in either of these \ears. .\ new nest was built at COXWR in 1979. One eaglet fledged fioin this nest in 1980 and two in 19S1. In 19S2. a second nest was constructed at COXWR. One nest February 1 988 Havera and Kruse: Winter Populations of Bald Eagles 21 Horseshoe Lake Fig. 29. -Locations ol bald eagle iiiglil rooMs idciuilicd bv Diiiislaii ami Kauksd 981). .Sabine and Klinistia (I9,S.')). Robeil l'liel|)s (per- sonal ( omnHMiicalion 19H(i). |obn (Irigsln (personal (onniuMMca- lion I'.I.SI'o. and |.inies W.ixcuilis (pcisonal i oMUniinic .ilion I'.ISli). was cl(.sti()\i(l l)\ liij^li winds, but the oiIr'i |)1(>(1iih-(1 two youn^ in 19X2, two in 1983, and one in 1981. This second nest was blown to the ground in 198') and ncj young were produced. Two nests were (on- strueled at CX)\VVR in the spring of 1987. Two young Hedged Iroin one nest, and one young fledged from liieoilui . 1 wo neslsweie also (oiisi I uded a! CONW'R in llie spring of 1987. Two voung fledged from one nesi and one young was killed in a sloini jusl before fledging fiom the oilier nesi. Allliougli (X)\\VR has lediilh been ihc inosi |)i(>- (hulive eagle nesliiig area in Illinois, the liisi sniiess- ful !iest was at Horseshoe Lake in Alexander Coiuii). Two yoiuig were produced from a single nesi ai Horseshoe Lake in 1978. NosubsecjuenI nesting a(li\ ■ ity was obseivt-d in that area until 1987 wlien a new nest was constructed but no Noung wcic flc'dged. Nesting activity has l)een seen at the Savanna Army Depot each yeai since 1978. A nest was constructed in 1978, enlaiged in 1979, blown down in 1980, and replaced by a new nest in 1981. Kggs were laid and incubated in 1982, but it is not known if the eggs hatched. The nest was again destroyed by wind in 198;^ and rebuilt in 1984. The first successful use of this nest was in 1985 when one eaglet fledged. Othei bald eagle nest construction has been seen at Union (louiity C.onsei vation Area in 1974 and 1980 and at Rend Lake in 1982 and 1983, but no young were pioduced from these nests. Although bald eagles are occasionalh' seen during the late spring and summer months in the Illinois River valley, the first documented nesting of bald eagles in central Illinois occurred in 198(5. A nest was established neai' the Illinois River in Pike County but was later abandoned. In 1987 a nest was built in Han- cock County; however, it was also abandoned. Table 6.— Bald eagle nesting activity in Illinois, 1973-1987. Year Number of Nests Number of Young Fledged 1973 1974 1975 1976 1977 1978 1979 1980 I98I 1982 1983 1984 1985 1986 1987 BALD EAGLE MORTALITY .\ total of 'M) bald eagles found dead in Illinois from 19()3to 1984 were submitted to federal laborato- ries for determination of cause of death. Lhese birds died from a variety of causes, many of which are re- lated to human activity (Table 7, NWHI. 1985). Like the nationwide data |)iesented earlier, these moitalilv figures should be used as an index lalhei than as an accurate measure of the causes of eagle deaths in a po|)ulation because it is clilTiciilt to collect a i.indom sample-. Shooting was the most common cause of de.ith of l)ald eagles in Illinois, accounting for 1 1 (.SO.O'r) of the deaihs (Table 7). At least two moic- eagles weie shot to death in 1985. .Six ac c iciental deaths of Ijald eagles resulted from traumatic injuries, such as collisions with wires or ve- hicles (Table 7). Other causes of eagle deaths, such as electrcK ulion. emac iaiion, and trapping, were of rela- tively minor importance in Illinois. .Six eagles also died from poisoning during this 22-year period in Illinois, and foiu of those six birds linois Natural History Survey Biological Notes Table 7.— Bald eagle mortality in Illinois, 1963-1984 (National Wildlife Health Laboratory 1985). Numberof Percent Cause of Death Mortalities ofTotal Gunshot 11 30.6 Trauma (excluding gunshot, electrocution, and trapping) 6 16.7 Poisoning 6 16.7 Lead 4 11.1 Other 2 5.6 Electrocution 3 8.2 Emaciation 1 2.8 Trapping 1 2.8 Other . ^ Decomposed, '' Open ' 8 22.2 "Death attributed to some cause other than those listed. ''Carcass too badly decomposed for diagnosis. ' No apparent cause of death could be determined from necropsy examination and ancillary laboratory analyses. were victims of secondary lead poisoning (Table 7). Waterfowl, especially crippled and dead birds, are a primary food item for bald eagles in some areas. Sev- eral species of waterfowl have been identified as food items for bald eagles in Illinois (Jonen 1973, Fischer 1982, Harper 1983, Sabine & Klimstra 1985). Feeding on waterfowl carcasses was usually accomplished by tearing the duck or goose apart with the beak while standing on the carcass (Fig. 30) (Sabine & Klimstra 1985). The growing problem of secondary lead poisoning in eagles resulted in the requirement that nontoxic (steel) shot be used for waterfowl hunting in 10 Illinois counties in 1985. The prohibition of lead shot was expanded to 22 counties for the 1 986 waterfowl season (U.S. Dept. of Interior 1986). IDOC modified the non- toxic shot zones for the 1986 waterfowl season to in- clude primarily those areas that are used extensively by waterfowl and bald eagles (W. Anderson, personal communication 1986). This modification resulted in the prohibition of the use of lead shot for sport hunt- ing of waterfowl in corridors along the Illinois and Mississippi rivers and in five entire counties, and in part of the Rend Lake area (Fig. 31). This corridor approach is supported by the findings of Havera and Kruse (1986) in their evaluation of the distribution of wintering bald eagles and waterfowl harvest in Illinois. Restrictions on the use of lead shot for waterfowl hunt- ing in Illinois are part of a growing nationwide effort to reduce the availability of lead to eagles by reducing the number of dead and crippled waterfowl with lead shot embedded in their flesh or present in their diges- tive systems. The USFWS is moving towards a total nationwide ban on the use of lead shot for the hunting of waterfowl by the 1991—1992 hunting season (USFWS 1986). SUMMARY Continental numbers of bald eagles have increased since eagles achieved legal protection and since the prohibition of certain pesticides that affected repro- ductive success of eagles. Wintering bald eagles are attracted to Illinois by the availability of food, primar- ily fish. Wintering eagles generally appeared in earlv to mid-October in the northern and central regions of Illinois and during late October in the southern regions. Eagles usually had departed Illinois by mid- April. Numbers of wintering bald eagles sighted in Illinois during the midwinter inventory have in- creased since its inception in 1957, and the highest number censused by INHS was 1,217 on 6— 7 January Fig. 30.-Iiiiiii.iiurc h.iM c.igle I IcvdiMH (in .1 <1<.M(1 Canada ijooso 1 (I'hdioln linioilu Sniallcv.Min- ' ncsoi.L Dcp.irlincnt ol Nalnr.il Rcsoums ). February 1988 Havera and Kruse: Winter Populations of Bald Eagles 23 1986. The proportion of immature eagles counted by INHS during the midwinter inventory typically ranged between 30 and 37%. The Illinois and Mississippi river floodplains and the Canada goose refuges in southern Illinois hosted a majority of bald eagles wintering in the state. The mean number of eagles seen per inventory was highest for the Upper Mississippi River Region with values ranging from 4. 1 to 20.9 eagles per flight at the various areas censused. Six of the 1 1 areas censused in the Upper Mississippi River Region had an average of more than 10 bald eagles per inventory. By compari- son, only four areas in the 10 other regions censused throughout Illinois, (the Keokuk- Meyer, Grand Tower— Cape Girardeau, and Cape Girardeau— Cairo stretches of the Mississippi River and the Flat and Swan lakes area near the confluence of the Illinois and Mississippi rivers) had values exceeding 10.0 for the mean number of eagles per inventory. The Upper Mississippi River Region was censused for just 3 years (1980-1983). and only the Kaskaskia River Region had fewer inventories. Values for the estimated density of bald eagles per river mile and per square mile of wetlands, respec- tively, for the inventory regions of the Mississippi and Illinois rivers were Upper Mississippi, 1.38 and 0.96; Upper Central Mississippi, 0.46 and 0.36; Lower Cen- tral Mississippi, 0.48 and 0.34; Lower Mississippi, 0.24 and 0.20; Upper Illinois, 0.20 and 0.47; Central Il- linois, 0.67 and 0.47; and Lower Illinois, 0.56 and 0.49. Fewer bald eagles were observed in the Surface- mined Lakes Region in west-central Illinois, the Northeast Lakes Region, the Central and Southern Lakes Region, and the Lower Kaskaskia River Region. Intensified enforcement of laws protecting bald eagles, the protection and management of habitats identified as feeding, roosting, and major eagle use areas, and the reduction of the availability of lead shot to eagles should result in an increase in the number of bald eagles that winter in Illinois. ACKNOWLEDGEMENTS We gratefully acknowledge the help of H. Kath- leen Belcher and Michelle M. Georgi in data tabulation and the contribution of Frank C. Bellrose and Robert D. Crompton, who conducted the aerial censuses of eagles. Funds for airplane rental were provided by the Illinois Department of Conservation; U.S. Army Corps of Engineers, St. Louis District; Illinois Power Company; Commonwealth Fdison; and Central Il- linois Public Service Company. Tabulation and analysis of some of the data and preparation of this report were supported in part by the Environmental Research Program of the Department of Energy and Natural Resources under C'ontrad Nos. EH-27 and EH-2, and we thank Elizabeth Johnson and Linda Vogt for their support. We also thank Dr. Glen C. Sanderson, Head of the Section of Wildlife Research; Drs. Richard and Jean Graber, and Dr. Christopher D. Burnett of the Illinois Natural History Survey; Dr. David L. Fischer, Brigham Young University; and Dr. James B. Elder, who is retired from the L'.S. Fish and Wildlife Service and lives in Minneapolis, Minnesota, for reviewing the manuscript. Assistance for the final manuscript was provided by Shirley McC'lellan, As- sociate lechnital Editor; Lloyd Le Mere, lechnical Illustrator; Molly Scott, Photographer; Patty Du/an, computerization support and typesetting; Eva Steger, typesetting; and Katie Roat. Illinois Natural History Survey. We thank Dr. Fischer for the use of some of his excellent photogi aphs. Fig. 3 1. -Illinois nontoxic shot zones for the 1986 waterfowl hiunling season. 24 linois Natural History Survey Biological Notes No 129 APPENDIX Aerial inventory areas by region, number of inventories, mean number of bald eagles counted per inventory, and maximum number of eagles censused during a single inventory at each area for all census years. (Number of years of inventories in each region is listed in Table 4.) Asterisks indicate areas located in the Illinois River floodplain but inventoried as part of the Lower Central Mi ssissippi River Region. Mean Number Maximum Number Number of of Eagles of Eagles Inventory Area Inventories^ Per Inventory" Censused Upper Mississippi River Region Dubuque-Galena River 22 6.5 25 Galena River-Bellevue 22 7.4 31 Bellevue-Green Island 22 16.4 39 Green Island-Savanna 22 11.8 28 Spring Lake Refuge 23 5.5 13 Elk River Refuge 22 4.1 18 Elk River— Fulton 23 11.3 23 Fulton-Albany 23 15.0 47 Albany-Wapsipinicon River 23 11.6 29 Wapsipinicon River-Princeton 23 20.9 68 Princeton-Interstate 80 23 7.4 28 Upper Central Mississippi River Region Andalusia Refuge 49 1.8 47 Moline-Muscatine 124 4.9 27 Muscatine-Louisa 125 5.1 30 Louisa Refuge 127 5.1 24 New Boston 127 1.2 19 Keithsburg Refuge 131 2.9 14 Keithsburg-Oquawka 131 5.6 28 Oquawka Refuge 127 2.0 13 Crystal Lake 127 0.5 11 Oquawka-Burlington 131 5.5 52 Turkey Slough 128 0.9 8 Burlington— Dallas City 132 5.6 29 Dallas City-Fort Madison 133 4.9 19 Fort Madison-Nauvoo 133 7.5 37 Arthur Refuge 129 <0.1 3 Nauvoo- Keokuk 133 6.7 35 Lower Central Mississippi River Region Keokuk-Meyer 130 11.7 90 Gardner Refuge 127 2.3 18 Meyer— Quincy 131 6.8 90 Quincy-Hannibal 125 5.0 57 Gardner Club 116 <0.1 3 Hannibal— Saverton 128 4.1 40 Shanks Refuge 126 2.9 18 Saverton-Louisiana 129 5.2 46 Delair Refuge 126 3.0 15 Sny Slough 127 0.8 39 Calumet Refuge 127 0.8 12 Duck Club 125 0.1 4 Cannon Refuge 126 2.8 13 Gilead 125 0.7 7 Batchtown Refuge 123 3.0 18 Batchtown Pool 123 6.3 52 Cuivre 124 2.1 16 February 1988 Havera and Kruse: Winter Populations of Bald Eagles 2S APPENDIX (continued) Inventory Area Mean Number Maximum Number Number of Eagles of Eagles of Inventories* Per Inventory" Censused 124 3.8 26 124 1.1 13 123 1.3 9 126 2.0 20 127 2.8 13 127 13.6 269 128 1.8 18 131 0.7 10 108 0.9 8 109 0.8 13 109 1.8 20 109 3.6 25 109 6.0 38 109 8.5 46 108 12.5 53 105 15.1 64 11 2.4 8 59 0.6 5 59 1.6 10 59 2.4 12 59 4.4 21 123 0.5 6 123 0.8 11 123 1.2 11 124 <0.1 4 125 6.6 26 125 4.8 20 125 0.9 6 125 0.7 8 125 2.4 12 125 1.5 15 125 0.1 3 125 <0.1 3 125 0.9 8 125 3.8 14 123 0.8 7 123 <0.1 1 124 3.8 19 125 2.3 23 125 6.0 24 122 1.3 10 128 2.1 9 137 3.9 34 108 0.4 7 1 22 0.8 6 120 1.8 17 Dardenne Marais Temps Clair Hager Grafton-Alton Gilbert Lake* Flat Lake, Swan Lake* Stump Lake* Godar Swamp* Lower Mississippi River Region Horseshoe Lake St. Louis-Kimmswick Kimmswick— Crystal City Crystal City— Ste. Genevieve Ste. Genevieve-Chester Chester-Grand Tower Grand Tower-Cape Girardeau Cape Girardeau— Cairo Upper Illinois River Region Morris-Seneca Seneca— Marseilles Marseilles-Utica Utica-La Salle La Salle-Spring Valley Central Illinois River Region Turner Lake DepueLake, Spring Lake Coleman Lake Bureau Ponds Goose Lake Senachwine Lake Siebold Lake Sawmill Lake Billsbach Lake Weis Lake Sparland Lake Wightman Lake Sawyer Slough Hitchcock Slough Round Pond Meadow Lake Rice P(jnd Goose Pond Upper Peoria Lake Lower Peoria Lake Lower Illinois River Region Pekiii Lake Poweiton Lake .Spring Lake Banner Marsh Duck Creek Cooling Lake Illinois Natural History Survey Biological Notes APPENDIX (continued) Mean Number Maximum Number Number of Eagles of Eagles Inventory Area of Inventories^ Per Inventory" Censused Rice Lake 134 2.2 11 Big Lake 134 3.0 14 Goose Lake 131 0.8 9 Clear Lake 134 6.0 28 Chautauqua Lake 135 9.4 35 Quiver Creek 136 4.1 23 Quiver Lake 110 0.7 10 Bath Lake 127 1.0 11 Grass Lake 133 1.6 10 Anderson Lake 134 1.3 9 Moscow Bay 135 0.9 9 Jack Lake 135 5.2 23 Snicarte Slough 135 0.6 7 Ingram Lake 135 1.8 13 Stewart Lake 132 0.9 11 Crane Lake 134 8.1 29 Cuba Island 133 5.5 40 Chain Lake 133 2.0 16 Sangamon Delta 132 4.3 33 Treadway Lake 132 1.1 12 Muscooten Bay 69 0.2 3 Big Prairie Lake 131 0.9 12 Meredosia Lake 133 4.9 25 Smith Lake 130 0.6 11 Central and Southern Lakes Region DeWitt County Clinton Lake 14 0.0 Sangamon County Lake Springfield 56 0.8 7 Christian County LakeSangchris 59 0.3 3 Montgomery County Cof feen Lake 41 0.4 5 Shelby and Moultrie counties Lake Shelbyville 31 0.4 3 Jasper County Newton Lake 26 0.6 6 Clinton and Fayette counties Carlyle Lake 56 2.2 9 Jefferson and Franklin counties Rend Lake 55 1.8 9 Randolph County Baldwin Lake 58 0.4 5 Northeast Lakes Region McHenry County McHenry Dam vicinity 31 1.4 7 Wonder Lake 28 0.3 3 Woodstock Ponds 28 0.0 Pistakee Bay 30 0.2 5 Lake County Fox Lake 30 0.3 3 February 1988 Havera and Kruse: Winter Populations of Bald Eagles 27 APPENDIX (continued) Inventory Area Number of Inventories ' Mean Number of F.agles Per Inventory'' Maximum Nuiulier of Eagles C'.ensused Grass Laice Antioch-Laice Villa DeKalb County Shabbona Lake Cook County Crabtree Nature Area Baker Lake vicinity McGinnis Slough vicinity McGinnis— Calumet Will County Des Plaines Wildlife Area Rendwick Lake vicinity Grundy County Dresden Lake Goose Lake Prairie Collins Lake Braidwood Lake La Salle County Seneca Lake Surface-mined Lakes Region Knox County Victoria vicinity Rapatee vicinity Peabody property Fulton County Norris vicinity Fairview vicinity Fin and Feather property CI LCO property Buckheart property Cebuhar property Justice property Metropolitan Sanitary District property Fulton County C^onservation Area Cuba vicinity Wee-Ma-Tuk property Red Barn property Giant Goose Ranch property Lower KashtLskia Kii \ving spring, and not llie entire (ensiis period troin f.ill through spring. Illinois Natural History Survey Biological Notes No. 129 LITERATURE CITED Anonymous. 1915. Bird-Lore's 15th Christmas census. Bird-Lore 17:22-48. Barnes, R.M. 1939. Law breaking mayor kills a bald eagle. Oologist 56(12):134-135. Bellrose, F.C., Jr. 1944. Bald eagles nesting in Illinois. Auk 61(3):467-468. Bonnell, C. 1935. The occurrence of the American bald eagle along the Ohio River in Illinois. Illinois State Academy of Science Transactions 28(2):249-250. Braun, C.E., F. Hamerstrom, T. Ray, and CM. White. 1975. Con- servation committee report on status of eagles. Wilson Bulletin 87:140-143. Brown, L. 1977. Eagles of the world. Universe Books, New York. 224 p. , and D. Amadon. 1968. Eagles, hawks, and falcons of the world. 2 volumes. McGraw-Hill, New York. 945 p. Cain, S.L. 1986. A new longevity record for the bald eagle. Journal of Field Ornithology 57(2): 173. Clark, W.S. 1 983. The field identification of North American eagles. American Birds 37(6):822-826. Dunstan, T.C., and E. Fawks. 1981. Documentation of bald eagle wintering areas and populations in Illinois. Unpublished report to Illinois Department of Conservation, Springfield. 60 p. Ekblaw, S.E. 1918. A record of the bald eagle from Champaign County, Illinois. Bird-Lore 20(6):421. Evans, D.L. 1982. Status reports on twelve raptors. U.S. Fish and Wildlife Service Special Scientific Report, Wildlife 238. Wash- ington, DC. 68 p. Fawks, E. 1983. Elton Fawks and bald eagles. Limited edition pub- lished by Elton Fawks, East Moline, Illinois. 208 p. Fischer, D.L. 1982. The seasonal abundance, habitat use, and forag- ing behavior of wintering bald eagles, Haliaeetus leucocephalus, in west-central Illinois. M.S. Thesis. Western Illinois University, Macomb. 178 p. , and D.J. Osterfeld. 1981. Influence of wind exposure on use of bald eagle loafing and night-roosting areas, with impor- tant implications for management. Pages 1 l-13m'r.N. Ingram, ed. 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Wilson Bulletin 62(4):210. High quality manuscripts dealing with any aspect of natural history will be considered for publication in one of the Illinois Natural History Survey series: Bulletin, Biological Notes, Circular, and Special Publication. Authors who arc not employees of the Survey arc required to pay printing costs. Manuscripts should follow the rccommcndatii)ns of the third edition of the Couniil of Biology Etlilors Style Manual except that journal names in the literature cited section are to be spelled in full. The Survey exjx-c is to publish only one or two manuscripts by non-.Survcy authors yearly. Send three copies of manuscripts to be considered lor publiiatinn lo Olliic ol ili<- (Ihicl. Illinois N.iiiii.il HiMorv Survey, 607 East Peabody Drive. Champaign, Illinois 61820. Illinois Natural History Survey 607 East Peabody Drive Champaign, Illinois 61820 Division of the Department of Energy and Natural Resources US ISSN ()O73-490X 64 184 — 3M — 2/88