THE UNIVERSITY OF ILLINOIS LIBRARY m 95.79 ^5o e^wrwfKi^: WAY 2 1946 MAY 18 1946 NOV 2 7 1946 ■M Z(j 1947 J1JW9 tgfet juN e-iesB MEMOIRS CAENEGIE MUSEUM, 1901-1904 VOL. I. W. J. HOLLAND, Edilor. J. B. HATCHER, Associate Editv PITTSBURGH. Published hy the Autiiokity of the Board ok Tuustbes of XHt carnegip: institute. J 1 -" ^ I PREFATORY NOTE. The first volume of the Memoirs of the C-arnegie Museum is brought to its con- clusion with the publication of Dr. William H. Ashmead's exceedingly important Monograph upon the Chalcidoidea, the descriptive portion of which is based upon the collections made in South America by Mr. Herbert H. Smith, now the property of the Carnegie Museum. No less valuable from a scientific standpoint tlian the concluding paper of the series embodied in this volume are the papt'is of Mr. Hatcher on Paleontology and the Osteological Monograph of Dr. Slmfeldt. It is with a feeling of satisfaction that the Editor contemplates the issue in final form of this volume, which marks the initiation in the city of Pittsburgli of an enterprise which is destined to give to it as the years go by an enduring reputation as a center of scientific culture. To the fame of Pittsburgh as the seat of some of the most Cyclopean industries of the age is being added reputation as a seat of learning. Under the cloud of smoke, which attests the industry of her inhabitants, and is the sign of her material prosperity, live men who find their pleasure in ex- ploring the wonders of the material universe, and the record of their discoveries and researches will from year to year be found in the Annals and the Memoii-s of the great Museum which the more tlian princely generosity of Mr. Andrew ( 'ar- negie has called into being. Here ai-e being brought together from all over the globe materials for study illustrating the different arts and sciences, and upon these collections is destined to be built up an extensive literature, which will add to the sum of human knowledge. Among the first fruits of this movement the following pages are found, and they are submitted to the attentive i-egard of students through- out theworld in the confident belief that those who are capable of con-ectly esti- mating the value of such publications will find in them l)otli pleasure and profit. W. J. Holland. TnK (jARNEfiiE Museum, Marcli 31, 1904. 5Kijj/o<]ocu.s lonf/ux :\Iar.sli 5 2. Dentary Bone of IHplmlonis Ioikjus 8 3. Section of Maxillary Bone of JUjJodocus loncjus 9 4. Atlas of Diplodocus lom/us, Side View 19 5. Atlas of Diplodocim lonr/m, Front View 19 6. Axis of DiplodocuH cwnef/ii, Seen from Right Side '20 7. Tenth, Eleventh, and Twelfth Cervicals of Diplodoem canuyil, Seen from Right Side. . 22 8. Posterior View of Eleventh Cervical of Dip/odocus carnegil .• 23 9. Inferior View of Sacrum and Ilia of Diplodocus carner/ii 30 10. Sacrum and Ilium of Diplodocus hiiffus Marsh, Seen from Below 33 11. Coossified Caudal Vertebrae of Diplodocua oartiefjfii 3(5 12. Superior View of Pair of Sternal Bones of Dlp/ndocns cdnurfii 40 13. Supposed Clavicle of IHplodocua carnaju 41 14. External View of Right Scapula and Coracoid of Dlplodocm carna/i'i 44 15. Proximal P]nd of Left Femur o{ Diplodocus cariKyii 4() K). Distal End of Left Femur of Diplodocus canuyii 4G 17. Oblique Internal Front View of I^eft Femur of f)ij>loilocus ciuimjii 47 18. Proximal End of Right Tibia and Fibula of Diplodocus carneijii 48 19. Distal End of Right Tibia and Fibula of Diplodocus carne/i,nii(s fr/itiux Scott. XX. Mounted Skeleton of J>aj,l,uni(s frlinus ^ott. XXI. Dorsal Aspect of the Pelvis of I'lotux Irvuikanfi ; Basal View of tlic Skull of Pborthon a-thrrem ; Superior Aspect of the Skull of Pharlhon faviroxtris; Skeleton of the Left Foot of /"'yrf/ff/rt (iqiii/a : Ventral View of the Sternum of /V(«;7/(o/i (r/Ar/riw ; Left Lateral Aspect of the Trunk Skeleton of P/metlion flavirostrlx. XXII. Right Lateral View of the Skull and Mandible of Sidu go.sui ; Basal View of Same Skull ; Basal View of the Skull of »S'«/« brrirdrri. Plate. XXIII. Superior A-spect of Skull of Said r/o-ssi ; Superior Aspect (jf Skull of Sii/,t hrvwderi ; Left Lateral View of Trunk Skeleton of Sulci ffossi. XXIV. Left hatera.] Yiew of SknW o( Plialncrocorax pelafflcun ro/jiistas ,• Left Lateral View o{ Phalacrocorax albiventris ; Left Lateral View of Skull of Fluikicroroid.r luilc ; Basal View of Skull of Phalacrocorax melnnoleucus ; Left Lateral View of Skull and Mandible of Phalacrocorax dilojylms ; Superior Aspect of Skull of Palacrocorax J). 7-ohuisttM ; Superior of Skull of Pluilacrocorax dilophus; Superior Aspect of Skull of Phahtcrocorax tnelanoleucus ; Superior Aspect of Skull of Phalacrocorax albiventris. XXV. Dorsal View of Trunk Skeleton of Phalacrocorax urile; Left Lateral View of Trunk Skeleton of Phalacrocorax urile. XXVI. Aiiconal Aspect of Right Carpo-metacarpus of Fregata aquila; Left Lateral View of Skull of Phalacrocorax mdanoleucus ; Auconal Aspect of Proximal Phalanx of the Medius Digit of the Manus of Right Pectoral Limb of PVef/afa aquila ; Basal Aspect of Skull of Phalacrocorax dilophus ; Basal Aspect of Skull of Phalacrocorax p. robustus; Basal Aspect of Skull of Phalacrocorax albivevtris; Dorsal Aspect of Sternum and Os Furcula of Phalacrocorax albiventris. XXVII. Anconal Aspect, Right LTlna, Pelecanus shurpei; Anterior Aspect, Left Femur, Pelecanus sharpei; Anterior Aspect of the Left Tibio-tarsus, Pelecanus sharpei; Palmar Aspect of Left Humerus, Pelecanus sharpei; Basal View of Skull of Pelecanus fuscu^-i ; Basal View of Skull of Pelecanus sharpei ; Left Lateral View of Skull of Pelecanus sharpei ; Anterior Aspect of Right Coracoid of Pelecanus sliarpei ; Anconal Aspect of Left Carpo-metacarpus, Pelecanus sharpei; Superior View of Skull of Pelecanus fuseus ; Superior View of Mandible of Pelecanus fuscus. XXVIII. Dorsal Aspect of Sternum and Coossified Os Furcula of Pelecanus sharpei; Palmar Aspect of Left Pectoral Limb of Phalacrocorax urile ; Inner Aspect of Right Pelvic Limb of Phalacrocorax urile. XXIX. Skull, Including Mandible, of Frajata (npiila (No. 1S,4S5, Coll. U. S. Nat. Mus.). XXX. Ventral Aspect of Pelvis of Pelecanus sharpei; Ventral Aspect of Sternum and Shoulder-girdle of Fretjuta aquila ; Left Lateral Aspect of Sternum of Fregata aquila. XXXI.-XXXIX. Figures of Xew Genera and Species of Chalcidoidca, described by William H. Ashmead. Publicatiom of the Carneyie Mmemu. Serial. No. 21. MEMOIRS OV THU CAENEGIE MUSEUM. Vol. I. No. 4. W. J. HOLLAND, Pji.D., Sc.D., LL.D., Kkitor. J. B. Hatchek, Ph.B., Associate P^ditok. CLASSIFICATION OF THE CHALCID FLIES OK THE SUPERFAMILY CHALCIDOIDEA, WITH DESCUIPTIONS OF NEW SPECIES IN THE CARNEGIE MUSEUM, COLLECTED IN SOUTH AMERICA BY HERBERT H. SMITH. By WILLIAM HARRIS A8HMEAD, A.M., Sc.D., ASSISTANT Curator, U. S. National Museum. PITTSBURGH. Published by the Authority of the Boarp of Trustef.s of thk CARNEGIE INSTITUTE. January, 1904. For sale by Messrs. Win. Wesley it .Son, 28 Elssex St., Strand, London, England ; Mes-irs. li. Friedhcnder n. .Sohn, 11 Carlstrasse, Berlin, N. W. 0, GeriuaQy ; and at the Carnegie Museum, Sohenley Park, Pittsburgh, Pa., U. S. / THIS VOLUME IS RESPECTFULLY DEDICATED TO DR. ('ARL W. VON DALLA TORRE THE EMINENT BOTANIST AND HYMENOPTERIST, PROFESSOE IN THE UNIVERSITY OF INNSBRUCK, WHO, IN THE COMPILATION OF "CATALOGUS HYMESOPTERORUM," HAS DONE SO MUCH FOR WORKING H YMENOPTERISTS THE AUTHOR Washington, D. C, January, 1904 TABLE OF CONTENTS. Dedication iii Errata and Corrigenda x Introduction 22") Part I. Classification of the Ciialcid Flies, or tlie Siiporfaniily ( 'liuloidoidea 220 Table of Families 22(i Family LX. Agaonidse 2.31 Table of Subfamilies 232 Subfamily I. Agaoninie 232 Table of Genera 233 Subfamily II. Sycopbagiuie 234 Table of Genera 23-5 Family LXI. Torymidte 23.') Table of Subfamilies 23t! Subfamily I. Idarniuje 237 Table of (ienera 237 Subfamily II. Toryminje 241 Table of Genera 241 Subfamily III. Monodontomeriiue 242 Table of Genera 242 Subfamily IV. Podagrioninai 244 Table of Genera 244 Subfamily V. Megastigmiua; 244 Table of Genera 244 Subfamily VI. Ormyriuse 245 Table of (jJenera 245 Family LXII. Chalcidid;e 246 Table of Subfamilies 246 Subfamily I. Leucospidina^ 246 Table of Genera 247 Subfamily IT. Chalcidince 247 Table of Tribes 248 Tribe I. Chalcidini 248 Table of Genera 248 Table II. Smicrini 250 Table of Genera 2.50 Tribe III. Chalcitellini 254 Table of Genera 254 Tribe IV. Haltichellini 254 Table of Genera 254 TABLE OF fONTENTS Tribe Y. Dirliinini 2')! Tabic of (Jencra 2o7 Family LXIII. Eurytomidfe 257 fable of Tribes 258 Tribe I. Axiraini 258 Table of Genera 258 Tribe II. Isosomiiii 259 Table of Genera 259 Tribe III. Eurytomini 260 Table of Genera 261 Tribe IV. Rileyini 263 Table of Genera 264 Table V. Decatomini 265 Table of Genera 265 Family LXIV. PerilampicLe 265 Table of Genera '. 266 Family LX V. Eucharida; 266 Table of Genera 267 Family LXVI. Miscogasteridie 270 Table of Subfamilies 271 Subfamily I. Pirenime 271 Table of Genera 271 Subfamily II. Tridyminae 272 Table of Tribes 273 Tribe I. Tridymini 273 Table of Genera 273 Tribe II. Metastenini 275 Table of Genera 275 Subfamily III. Miscogasterinse 276 Table of Tribes 277 Tribe I. Haltioopterini 277 Table of Genera 277 Tribe II. Miscogasterini 278 Table of Genera 278 Subfamily IV. Lelapina; 279 Table of Genera 279 Family LXVII. Cleonymida; 280 Table of Subfomilies 280 Subfamily I. Chalcedectiiife 281 Table of Genera 281 Subfamily II. Cleouyminae 282 Table of Genera 282 Subfamily III. Pelecinellinie 285 Subfamily IV. Golotrecbninie 285 Family LX VIII. Encyrtidse 286 fable of Subfamilies 286 TAI5LK OF CONTENTS VII Subfamily I. Eiipelmiiiip 287 Table of Tribes 287 Tribe I. Eupclmini 287 Table of Genera 2S7 Tribe II. Tauaostif^mini 2!n Table of Genera 201 Subfamily II. Eneyrtinre 2!l2 Table of Tribes 292 Tribe I. Ectromini 2!l2 Table of Genera 2!i;? Tribe II. Encyrtini 2!»7 Table of Genera 2f)7 Tribe III. Mirini 298 Table of Genera 298 Tribe IV. Arrhenopbagini 311 Table of Genera .'511 Subfamily III. Signiphorimc 311 Family LXIX. Pteromalidis 311 fable of Subfamilies 312 Subfamily I. Pteromaliua; 312 Table of Tribes 313 Tribe I. Metaponini -"i 1 3 Table of Genera 313 Tribe II. Raphitelini 31 ."> Table of Genera 315 Tribe III. Eutelini 317 Table of Genera 317 Tribe IV. Pteromalini 3.1 8 Table of Genera 318 Subfamily II. Merisinte 318 Table of Tribes 322 Tribe I. Roptrocerini 322 Table of Genera 323 Tribe II. Merisini 323 Table of Genera 323 Tribe III. Isoplatiui 324 Table of Genera 324 Subfamily III. Eunotinie 325 Table of Genera 325 Subfamily IV. Sphegigasterinaj 327 Table of Tribes 327 Tribe I. Asaphini 327 Table of Genera 327 Tribe II. Pachyneuriiii 329 Table of Genera 329 Tribe III. Sphegigasterini 330 TABLE OF CONTENTS Tal)le of Genera 330 Tribe lY. C'ratomini 332 Table of Genera 332 Subfamily lY. Spalangiina? 333 Table of Genera 3.')3 Subfamil)^ Y. Dipariuie 334 Table of Genera 334 Family LXX. Elasmidfe 335 Table of Genera 335 Family LXXI. Eulophiilie 336 Table of Subfamilies 337 Subfamily I. Entedoninte 337 Table of Tribes 337 Tribe I. Tetracampini 337 Table of Genera 338 Tribe II. Omphalini 338 Table of Genera 339 Tribe III. Entedonini 340 Table (if Genera 340 Tribe I Y. Pediobiini 343 Subfamily II. Apheliuina? 344 Table of Tribes .-US Tribe I. Aphelinini 345 Table of Gene-a 345 Tribe II. Pteroptricini 347 Table of Genera 347 Subfamily Til. Tetrastiehinte 347 Table of Tribes 347 Tribe I. Ceratoneurini 347 Tribe II. Tetrasticliini 347 Table of Genera 348 Subfamily lY. Elachertinte 350 Table of Tribes 351 Tribe I. Euplectrini 351 Table of Genera 351 Tribe II. Ophelinini 352 Table of Genera .352 Tribe III. Elachertini ; 354 Table or Genera 354 Subfamily Y. Eulophina; 355 Table of Tribes 356 Tribe I. i:uloi)hini 356 Table of (ienera 356 Tribe II. Hemiptarsenini 358 Table of Genera 358 Family LXXII. Tricliogrammidse 358 TABLK OF C'ONTI'.NTS IX Table of Subfamilies 'A'/J Subfamily I. Oligositina- .-JoO Table of Genera 859 Subfamily II. Tridiograuiuiiuiu ."560 Table of Genera .-JdO Family LXXIII. Mymaridte :•,(;] fable of Subfamilies :)r>-2 Subfamily I. Gonatoeerime '.U')'2 Table of Tribes :}62 Trii)e I. Ooctonini .'562 Table of Genera .'502 Tribe II. Gonatocerini ."'.02 Table of Genera .•;(;2 Subfamily II. Myniariiiie ."SO;! Table of Tribes :5(;:5 Tribe I. Anaphini :W-i Table of Genera 3f)3 Tribe II. Mymarini 'M'.] Table of ( Jenera '.W'.] Genera unknown to author and ikiI elassiiiid 3()4 Genera incorrectly placed with ( 'halcidoidea ;Ui4 Literature and Abbreviations 522 Bibliography of genera al])habftically arranged .')G5 Part II. South American C'lialcidoidca .'UM Family LX. Agaonidic ;3y4 Family I^XI. Torymida; 395 Family LXII. Ghalcididic. 402 Family LXIII. Kiirytomidio' 458 Family i^XIV. J'erilampidic 467 Family I^XV. Eucharidie 4(i7 Family I^X^'I. Miscogasterida' 474 Family LXVI I. C'leonymida-; 483 Family I.XVII I. Encyrtid.-r. 486 Family LXIX. Ptcromalidtu . 497 Family LXX. Elasmida; 502 Family LXXI. Eulophidse 503 Family LXXII. Trichogrammida- 521 Family LXXII I. Mymaridre 521 Literature 522 Special Index 533 ERRATA ET CORRIGENDA. Page 240, line 27, Otetewlla sliould read Otitcxdla. Page 254, line 30, Anacryptiia should read Aiuicri/jthiy. Page 201, line 43, Prodecohina sliould read ProihrdlmiKi. Page 2!t3, line 40, (type Pha;w,hx,-nH) should read /'liis. Page 365, line 1, (= Merjadtymm Dalmaii) shoidd read (= Mei/((spi/iifi AVestwooil). Page 371, line 13, (= KriechbaimencUa Dalla Torre) should read (= KriivlihdununHd). Page 373, line 31, Enarrjopdtc should read Euan/opcltc. Page 377, line 32, (type //. dicliroiis Thomson) sliould read (typo If. nh/rr Ashuua.l). Page 380, line 5, Li/iinwuon shoidd read Li/iiiiucdii. Page 389, line 15, (type .Va/ra Aslim.) should read mcllca. Page 400, line 19, P/rsl,,si;,/nii)diis should rrad Pledodigmodes. Page 496, line 7, I'dniici/r/iis />r(i.'. .")i2 (type .1. n'Kjrinpx Aslmi.). Aseiba Cameron ( = Cerchysius Westw. tcufe Howard), Biol. Centr.-Amer. Hym., I., ISSil, p. 1 Bruchobius a. sh mead, gen. nov., ante, p. 314 (type B. laticeps Ashm.). Neocatolacccs A.shmead, gen. nov., ante, p. 320 (type Catolaceus tylodernise Ashm.). Nkolei-AI'S Ashmead, Fauna Hawaiiensis, I., 1901, p. 312 (type N. huwaiien.sis Ashm.). Paekardiella Ashniead, u. ii. fbr I'teratomus Paekard, ante, p. 364 (type Pteratomus ptitiia Paek.). Tetraenemoidea Howard, Proe. U. S. Natl. Mus., XXI., 1 898, p. 2:'.:', (tyi)e T. australiei Howard). Westwoodella Ashmead, gen. nov., ante, j). 359 (type < Hiijoxita mbfaxciddi Westw.). Xanthoatomus Ashmead, gen. nov., ante p. 360 (ty])e A', alblpcn Ashm.). MEMOIRS OF THE OAENEGIE MUSEUM. VOL. I. NO. 4. CLASSIFICATION OF THE SUPEKFAMILY CHALCIDOIDEA. By Wm. Harris Ashmead, A. M. INTRODUCTION. About fourteen years ago Mr. Herbert H. Smith placed in ray hands for study part of his extensive collection of South American parasitic Hymenoptera, repre- senting the old families Proctotrypidfe, Cynipidse and Chalcididse. The collection was a valuable one and when I took it I confidently expected to find time to finish with it within two or three years ; but so much other material Avas thrust upon me for study that I found it a physical impossibility to do it justice in that length of time ; only a few short papers on it have been published. The work accomplished during these past fourteeen years has I think fully demonstrated that I have not been idle, ray knowledge of the groups, genera, and species of the Hymenoptera has greatly increased and broadened, and I now feel better able to do justice to the splendid col- lection received so many years ago. The Herbert H. Smith collection of insects, made in South America, repre- senting nearly all orders, has been acquired by the Carnegie Museum, Pitts- burgh, Pa., and much to my regret, the many fine species, except a set of dupli- cates in my hands, and which 1 had always hoped might be secured by the National Museum, must go to that Museum. Dr. W. J. Holland, the Director of the Carnegie Museum, desires now that I complete my work as soon as pos- sible and return the material in my possession. In acceding to this request I have stipulated that he publish my classification of the chalcid-flies, or the Super- family Chalcidoidea, as a part of this work, and it is through his liberality that I am able to give so many fine illustrations and present in a more attractive form my ideas on the cla.ssification of another great complex in the Order Hymenoptera. 225 226 MEMOIRS OF THE CARNEfJIE MUSEUM The Prodotrypoidea] and Cynipoxdca in the coUeetioia will be described in another paper. In this contribution I have restricted myself to elucidating the chalcid-flies and have divided it into two parts. The first part is devoted to a classification of the superfamily Chalcidoidea ; the second part deals with the South American Chalci- doidea. In this latter part, besides my report upon the Herbert H. Smith c(j1 lection, the descriptions of the new genera and species, is given a complete bibliographical catalogue of all the known South American species. PART I. CLASSIFICATION OF THE CHALCID-FLIES, OR THE SUPERFAMILY CHALCIDOIDEA. Among the ten great groups or superfamilies of the Hymenoptera, recognized by the author, there is none so large numerically, more important economically, or so difficult to study and classify as the superfamily Chalcidoidea or the Chalcid-flies. The species exist everywhere not by hundreds, but by thousands and millions, and they are probably of far greater importance, from an economic standpoint, than are the Ichneumonoidea or ichneumon-flies. Only a few of them, comparatively speaking, are in any sense injurious, a single minor group, the tribe Isosomini, in the family Eurytomidte, alone being injurious to vegetation. All the others, except the fig-insects forming the family Agaonidse, the Megastigminas, a subfamily in the Torymidse, and some few hyperparasitic genera in different families, being genuine parasites and beneficial to man. It is true, however, that some of the chalcid-flies destroy a few beneficial insects, but the vast majority of the known species destroy mostly the injurious species in the other orders, i. e., the Coleoptera, the Lepidoptera, the Diptera, the Rhynchota, etc.; they attack the eggs, the larvje, and the pupte, and in some cases even the imagoes of their hosts, and their value to many great industries of the world cannot easily be estimated. Who, for example, can estimate the value of the fig-insects to the fig industry of the United States ? Through the efforts of Dr. L. 0. Howard and Mr. W. T. Swingle, of the U. S. Department of Agriculture, the caprifier of the Smyrna fig, Blustophaga j)senes Linne {grossomm Gravenhorst), has been successfully introduced into the fig-orchards of California, and the experiments already made fully demonstrate the great value this little chalcid-fl}^ has to the fig industry. It is evidently destined to revolutionize fig-growing in the United States, making it exceedingly profitable, and, on account of the superiority of the American-grown ASHJIEAD : CLASSIFICATION OV THE CHALCID FLIES 227 Smyrna fig jx^lk-nized by these Chalcid-flies, will in time diminish the impor- tation of the Smyrna fig. The Me(jastiyin:une too, I suspect, are like the fig- insects, and will he found to be of great importance as pollenizers of various plants and trees. At present the known genera and species of the ( "halcidoidea are considerably less in number than are the Ichneumonoidea recently classified by the writer; but this is due simply to the fact that the literature on the subject, in various languages, is widely scattered, in many foreign journals, magazines, proceedings of learned societies, etc., and the minute size of most of the species and the difficulty of their study, have deterred entomologists from giving them any attention. Only a little over 5,000 species have been described. If we look back for a century and a half we find comparatively few who have given much attention to these " atoms of creation," and of these a few names only stand out conspicuously as students of this great complex. The study of the group began about one hundred and fifty years ago with Linnaeus in Sweden and with GeofiVoy in France. Linne, or Linnaeus, in his Systema Natura, tenth edition, published in 1758, described several species under his genera Ichneumon, Cynips, Sphex, etc. Others took up their study, and an interest in them Avas aroused in Sweden, Austria, Germany, France, Italy, and England. In Sweden, besides Ijinne, Swederus, Fabri- cius, Zetterstedt, Dalman, Boheman, Dahlbom and Thomson did much valuable work in the group ; in Germany, Klug, Nees von Esenbeck, Bouche, Ratzeburg, and Forster — the last mentioned, the greatest systematic worker in the group; in Austria, Reinhard and Mayr; in Russia, Motschulsky ; in France, Geotfroy, Latreille, Fons- colombe, Perty, Guerin, Giraud, Sichel, and Andre ; in Italy, Spinola and Rondani ; in England, Curtis, Haliday, Westwood, Walker, Saunders, Kirby, and Cameron ; and in America, Say, Fitch, Cresson, Walsh, Riley, Frovancher, Howard, and Ash- mead. A few others might be mentioned, but those specified are the ones whose nanies stand out prominently as adding materially to our knowledge of the group during the eighteenth and nineteenth centuries. For years I have been studying this great complex and in the following pages have attempted to define the families, subfamilies, tribes, and genera of the world. The work has been a gigantic and most laborious one, necessitating the microscopic examination of many th9usands of these minute creatures ; but if it shall be found that I have brought some order into their classification, that I shall excite an interest in the collecting of the many thousands still unknown, and that I shall stimulate others to study them, my time has not been wasted and I shall be amply repaid. 228 MEMOIRS OF THE CARNEGIE MUSEUM SuPERFAMiLY VII. CHALCIDOIDEA. 1758. Cynips, Linne (partim), Syst. nat., Ed. 10", I., p. 343. 1802-05. Diplolepaires, Familie soixante deuxieme (partim) Latreille, Hist. Nat. Crust, et Ins., XIII., p. 198. 1802-05. Cynipseres, Familie soixante-troisieme (partim), opus, cit., p. 211. 1809. DiploleparisB, Familiaquinta (partim) Latreille, Gen. Crust, et Ins., III., p. 15. 1809. Cynipsera, Familia sexta (partim) Latreille, opus, cit., supra, p. 21. 1811. Diploleparia, Familia (partim) Spinola, Ann. du Mus. Natur., XVIL, p. 138. 1S20. Pteromalini, Familia (partim) Dalman, Kongl. Svensk. Vet.-Akad. Hand!., p. 132. 1825. Chalcidites, Quatrieme Tribu (partim) Latreille, Fam. Natur. d. Regne anim., p. 446. 1830. Chalcida, Family, Leach, Edinb. Encycl., IX., p. 144. 1840. Chalcididfe, Family, Westwood, Intro. Mod. Class. Ins., II., p. 154. 1846. Chalcidites, Order, Walker, List Chalc. Brit. Museum, I., p. 1. 1856. Chalcidite, Forster, Hym. Stud., II., pp. 1-90. 1875. Fteromalidte, Familia, Thomson, Hym. Skand., IV., p. 3. 1877. Chalcidita, Tribe 4, Forster, Ueber den Syst. Werth d. Fliigelg. bei d. Hym., p. 19. 1886. Chalcididae, Family, Howard, Ent. Amer., I., p. 197. 1897. Chalcidoidea, Superfamily VII., Ashmead, Proc. Ent. Soc. Washington, IV., p. 243. TABLE OF FAMILIES. 1. Hiiid wings exceedingly uairow, linear, pedunculate at base ; ovipositor issuing from beneath just anterior to tip of abdomen ; antennae without a ring-joint, the scape rather small, short, compressed 12 I Hind wings never very narrow, nor linear, not pedunculate at base ; ovipositor issuing far anterior to the tip of abdomen ; antennse elbowed, with 1, 2, or 3 ring-joints, very rarely without, the scape large and rather long 2 2. Axillae triangularly produced or advanced forward into the basal region of the scapulie, their base or anterior margin on or in advance of an imaginary line drawn from tegula to tegula ; anterior tibial spur most frequently small or weak ; tarsi 3-4-jointed, rarely 5-jointed or heteromerous 10 Axillfe normal, or at least never produced forward into the basal region of the scapulae, their base or anterior margin straight and always back of an imaginary line drawn from tegula to tegula ; anterior tibial spur large and strong ; tarsi 5-jointed (rarely 4-jointed, or 3- or 4-jointed in some wingless males) 3 3. Head in ? oblong, with a deep, broad longitudinal furrow above, the occipital margin superiorly, usually with a small recurved tubercle or spine at its middle ; mandibles or palpi most frequently furnished with saw-like appendages ; anterior and posterior legs very stout, their tibiie very much shorter than their femora, the middle legs very slender, sometimes aborted ; hypoi>ygium very ASHMEAD : CLASSIFICATION OF THE CHALCII) FLIES 229 prominent, acute, cultriform or lanceolate ; ovipositor long, prominently exserted ; ^ always apterous, the head anteriorly with a deep triangular fovea, in which are placed the short 3-9-jointed antennae ; the abdomen in the c? 's broadly sessile, long and tubular, thickened at base, or broadened apically with a tubercle or filament at each apical angle Family LX. AgaoniDvE. Head rarely oblong and quite differently formed, never with a deep broad longitudinal furrow above, most frequently transverse, or subquadrate, the occipital margin never with a small recurved spine ; mandibles and palpi without saw-like appendages ; middle legs not especially slender, the anterior and posterior legs are often stout, but their tibia; are always longer, at least never shorter than their femora ; hypopygium rarely very prominent ; cf most freiiuently winged, rarely apterous ; in the latter case the abdomen is normal, not long and tubular. Mesopleura large, entire, without a femoral furrow, except occasionally in some males, the ^ mes'i&pisterirUm large, triangular, not extending to base of front coxie ; middle tibial spur saltatorial, most frequently long and stout, or dilated at base 8 Mesopleura always with a femoral furrow or impression, the inescpisterhhin variable, rarely large, except in the Cleonymidm, most frequently small, wedge-shaped, or linear and extend- ing to base of front coxa; ; if largeand triangular, either the auterior or the posterior femora are mucrh swollen ; middle tibial spur not saltatorial, usually short or weak, never very stout. Hind tibiie with 2 apical spurs, rarely with 1 only ; In the latter case the radius termi- nates in a large, rounded stigma, the ovipositor very long 4 Hind tibite with 1 apical spur ; ovipositor rarely long ; if long the stigma is small 9 Mandible%i falcate, usually with 1 or 2 teeth within ; thorax most frequently very gibbous, the scutellum usually very large, often abuormally developed, elevated and produced posteriorly, the axilliB connate, not distinctly separated from the surrounding surface and broadly united along their inner margins 6 Mandibles usually 3-4-dentate at apex, rarely falcate, with 1 or 2 teetli within ; thorax not, or very slightly, gibbous, the axilhe distinctly separate, their inner margins most fiecpiently widely separated, very rarely touching. Hind coxie rarely much larger than the anterior coxse, most frequently smaller or equal ; if much larger, the pronotum is elongate, mesepisternum large, the hind legs very long, the postmarginal vein very long ; ovipositor very rarely prominent. .' 5 Hind coxie very large and long, usually five or six times larger than the anterior cox;e. Hind coxaj subtriquetrous, or at least compressed into a shaii) ridge above ; hind femora rarely very much swollen, aud most frequently simple, rarely with one large tooth or den- ticulate beneath ; hind tibia usually straight ; abdomen most frequently subcompressed (more rarely depressed), with a long ovipositor ; if without au exserted ovipositor, the abdomen is conical or conic-ovate with a peculiar sculpture, the radius (stigmal vein) usually very short, the hind tibia; at apex normal Family LXI. Torymid^e. ^ Hind coxa; usually very long and subcylindrical ; hind femora always much swollen and most frequently armed with teeth beneath or finely serrated, rarely without teeth ; hind tibia; strongly curved and obliquely truncateiy produced at apex, so that the tarsi seem to be attached a little before tips ; abdomen of various shapes, most frequently conical or conic-ovate, more rarely globose, or oblong-oval, the ovipositor very rarely prominent ; radius variable, rarely very short Family LXII. Chalciuid-k. Pronotum rarely transverse quadrate, conical or coiiically produced anteriorly, or very short, trans- verse linear, aud very much narrowed medially, rarely as wide as the mesonotum 7 :.u.>.i. 230 MEMOIRS OF THE CARNEGIE MUSEUM Pronotum large, quadrate or transverse quadrate, never very short, if somewhat shortened always as wide as the mesonotum. Pronotum quadrate or subquadrate ; abdomen in $ not triangulated, either globose, ovate, conic- ovate or lanceolate and compressed or subcompressed, the hypopygium most frequently prominent plowshare-shaped ; second doreal segment never very large ; mandibles not strong, most fi-equently 4-dentate Family LXIII. Eueytomid^. Pronotum shorter, more transverse and as wide as the mesonotum ; abdomen in ? most frequently triangulated, or globose, the second and third segments occupying most of the dorsal surface, the following very short and more or less retracted within the third ; hypopygium not prominent ; mandibles 2- or 3-dentate at apex Family LXIV. Perilampid^. G. Second abdominal segment very large and most frequently enclosing the following ; coxse not large, subglobose, nearly equal ; all legs very slender ; radius scarcely developed, its stigma sessile or sub- sessile Family LXV. Eucharid.e. 7. Mesepisternum not large, triangular ; anterior femora never much swollen, the posterior femora also normal or only slightly swollen ; marginal vein in hind wings usually long, the costal cell not reach- ing to the booklets or spinulje and most frequently very narrow ; radius well developed. Family LXVI. Miscogasterid^. Mesepisternum large, triangular ; either the anterior or the posterior femora are much swollen and sometimes toothed, or both are swollen with the hind femora toothed ; if with slender legs, the hind legs are very long, their coxse long, cylindrical, while the radius (stigmal vein) in front wings is verj' short, with the postmarginal vein very long extending to the apex of the wing {Pelecinella). Family LXVII. Cleoxymid^. 8. Mesonotum either depressed, with more or less distinct parapsidal furrows, the scapulae longitudinally ridged, or convex or subconvex, entirely without furrows, rarely convex with distinct furrows ; axillie most frequently meeting at inner basal angles, rarely very widelj' separated. Family LXVIII. Encyrtid.e. 9. Mesonotum subconvex with incomplete or complete parapsidal furrows ; hind coxse rarely much larger than the front coxse ; axiilse separated, not meeting at inner basal angles ; mesepisternum usually small, wedge-shaped or triangular ; hind wings with a long marginal vein ; mandibles usually stout, 3- or 4-dentate at apex Family LXIX. Pterom alid^e. 10. Hind coxse normal ; mesopleura impressed 11 Hind coxse abnormally large and dilated, their femora flat or compressed ; tarsi very long ; mesopleura entire, not impressed ; marginal vein in front wings most frequently extraordinarily lengthened, the radius very short, scarcely dilated ; mesonotum without furrows Family LXX. Elasmid^. 11. Tarsi 4-5-jointed, rarely heteromerous ; anterior wings not short and broad, with the pubescence normal , marginal and radial veins normal ; postmarginal vein often wanting ; mesonotum with complete or incomplete furrows Family LXXI. Eulophid^e. Tarsi 3-jointed ; anterior wings short and broad, broadly rounded at apex with the pubescences most frequently arranged in rows, more rarely normally pubescent ; marginal and radial veins united in the form of a strongly curved line n Family LXXII. TrichogrammiDjE. 12. Pronotum usually large, rounded, or conically produced anteriorly ; wings always with a long marginal fringe, nearly veinless and always without a radius (stigmal vein), the marginal vein most frequently reduced to a mere dot ; antennae in 9 most frequently terminating in a distinct fusiform or egg-shaped solid club, more rarely with a 2-jointed club ; tarsi 4-5-jointed. Familv LXXIII. Mymarid-e. ASHMEAD : OLASSIFICATION OF THE CHALf'in FLIES 231 Family LX. AGAONIDyE. 1846. Agaonid*, Family G (partim) Walker, List Chalc. Brit. Museum, I., p. 23. 1856. Agaonoidaj, Familie (S. descrip.) Forster, Hym. Stud., II., p. 29. 1871. Agaonidfe, Family (partim) Walker, Notes on Chalc, Ft. IV., p. 58. 1867. Blastophagidte, Familia, Kirchnei-, Cat. Hym. Eur., p. 188. 1882. CynipidPB, Sycophagides, Division 1, Saunders, Trans. Ent. Soc. London, p. 20. 1897. Agaonidse, Family LX., Ashmead, Proc. Ent. Soc. Washington, IV., p. 243. This family is one of the most striking and remarkable of any in the super- family Chalcidoidea. It is based ui)on the genus Ayaotb Dalman, established in 1818, from a specimen taken in Sierra Leone, Africa. The species composing this family, on account of their habits, curious forms, and the diversity of structure in the sexes, were long a puzzle to the ablest and most astute of the European hymenopterologists, but it is now definitely settled that they form a component of this great complex. Sir Sidney Saunders, as late as 1883, placed them as a division with the family Cynipidts. In my opinion, however, they have little in common with the Cynipoidea, and I concur with Walker, Westwood, and Mayr, in believing them a component of this major group. Mr. Francis Walker, an Englishman, was the first to give the group family rank ; but, as is the case with most of his families, he never properly defined or character- ized it, and merely lumped together a miscellaneous lot of insects obtained from figs, and called them a family — the Agaonidie. His ideas of the family were extremely vague and indefinite, and he placed in it many foi-ms with which they had no relationship. In 1871, Walker, in speaking of them said: "The Agaonidse appear as yet chiefly in thi-ee aspects, and in three different regions. The first region is the Mauritius, where they have been discovered by the researches of Dr. Coquerel. The three species figured are said to be 'condemned to eternal darkness' in the central regions of figs. These figs are the fruit of Firna h rrposed female of Dorylaa; and thereby the multitudinous tiibc of ants wIiosl' (■coiiuniy is 232 MEMOIRS OF TITK CARNEGIE MUSEUM SO reinarka1)le, emerges from blind and radical ApocrypUc and Sijcocrupkc, the per- petual dwellers in the interior of figs." " But the affinity of these two genera to the Chalcidix is more evident and appears by several connecting links in the Agaonidx; and thus the near relation to the general ancestors of the thousands and perhaps tens of thousands of the Chalcidiaj species, the trilje being considered in unity, are cradled in figs." Our knowledge of fig-insects, within the past twenty-five years, has been very greatly augmented by the studies of Prof. John O. Westwood and Sir Sidney Saun- ders, of England, Dr. Gustav Mayr, of Vienna, Austria, Dr. Paul Mayer, of the Naples Station, Italy, and ray own studies on some Florida, Mexican and West Indies species, so that to-day sufficient forms are known in both sexes to enable me to segregate, define and place in their proper groups, the miscellaneous insects known as fig-insects. In this work I have restricted the Agaonidx to the caprifiers, or true fig-insects, chalcid-fiies that live in and pollenize, or fructify, fig-trees. The others, heretofore classified with them, belong elsewhere, in three or four different families, and are either inquilinous or genuine parasites. Some, the vast majority, belong to the Tonjmhhe, while others belong to the Chalcididx, 3Iiscogas- teridee, Pteromalidse, etc. All fig-trees, in a wild state, are dioecious and wherever fig-trees grow, principally in tropical and semi-tropical countries, there also will be found fig-insects, for these microscopic creatures are essential to their pollenization. Undoubtedly, judging from the great number of fig-trees known to botanists, many genera and hundreds of species still remain unknown to us. Among the genuine fig-insects, two well-marked subfamilies may be distin- guished, separable by the aid of the following table : TABLE OF SUBFAMILIES. Abdomen in 9 subcompressed, the ovipositor prominent, the mandibles with an appendage, usually ser- rate ; males apterous, the abdomen long, narrowed towards apex and curving beneath the thorax. Subfamily I. AGAONlNiE. Abdomen in ? subcompressed, the ovipositor prominent, the mandibles without an appendage, the palpi sometimes with an appendage ; otherwise similar to the Agaoninse ; males apterous, the abdomen broad- ened towards apex, narrowed towards the base, and with a tubercle or long filament at each apical angle Subfamily II. SycoPHAGiNiE. Subfamily I. Agaonin.e. 1883. Cynipidca3, Division I., Sycophagides (partim), Saunders, Trans. Ent. Soc. London, 1883, p. 20. ASHMEyVD : CLASSIFICATION OF THE CHALCIl) FLIES 233 The males in this subfamily are easily distinguished from those in the next by the shape of the abdomen, which is always long and tubular, narrowed toward apex and never broadened at apex as in the Si/cophagimv. It is also more flexible and is usually curved or bent forward under the thorax. The females are more difficult to separate and, althtnigh with practice one can detect a difference in cephalic characters, almost impossible to define, the only reli- able character to separate them from those in the other subfamily is tlie mandibular appendage, which is usually finely serrate. In tlie ^ijcophaijiinv the mandibles are always without an appendage. The known genera falling in this group may be easily recognized by the us(; of the following table : TABLE OF OENEKA. 1. Females, always winged 2 Males, always apterous 10 2. AnteniiiB with the third or fourth joint with a ilistinct process 3 AntennsB without a joint with a distinct proces 9 3. Front wings with the marginal, stigmal and postmarginal veins fully deveU)pcd, or at the most with the postmarginal vein absent 4 Front wings without marginal, stigmal and postmarginal veins. Head oblong, about li times as long as wide ; antennse 11-jointed, thickened toward ape.x. Eupristina Saunders (type E. masonii Saund.). 4 Postmarginal vein present 5 Postmarginal vein absent 7 5. Antennae 11 or 12-jointed 6 AntennfB 10-jointed, the last five joints enlarged Kradibia Saunders (type A', coiraitii Saunders). 6. Head not very long, quadrate or nearly so 8 Head oblong, or very long, from 2* to 3 times as long as wide, tlie facial channel narrow ; mandibles at apex bidentate ; antennjE 11-jointed Pleistodontes Saunders (type P. imperialis Saunders). 7. Head quadrate or nearly, only a little narrower in front than behind ; stigmal vein usually a little longer than the marginal Eisenia Ashmead gen. nov. (type E. mexicana Ashm.). 8. Seventh abdominal segment on each side with a .small rounded or at most oval spiracular fovea. Blastophagus Gravenhorst (type Cynipa pscnes Linn^). Seventh abdominal segment on each side witli an itoiniule, disk-sliaped, spiracular fovea. Ceratosolens Mayr (type C. appendiculalus Mayr). 9. Front wings with the marginal, stigmal and postmarginal veins wanting ; head trapezoidal, a little longer than wide ; antennae 11-jointed, not thickened towards apex, the joints smooth ; middle legs very minute or subobsolete Tetrapus Mayr (type T. americanus Mayr). Front wings with the marginal, stigmal and po.stmarginal veins present ; head oblong, 24 times as long as wide; antennje (?) 12-jointed, the scape large, triangularly dilated, the funicle slender, filiform, the club abruptly enlarged, 3-jointed ; mandibles 3-dentate ; thorax bidentate at each side. Agaon Dalman (type A. paradoxum Dalm.). 10. Head anteriorly ivith a large, deep, triangular fovea. Ba.sal part of antennse enclosed in a canal 11 Basal part of antenna; 7iot enclosed in a canal 12 234 MEMOIRS OF THE CARXEGIE MUSEUM 11. Front tarsi 2-jointed, the front and hind femora stout and more or less compressed ; eyes very small or wanting Ceratosolens Jlayr. 12. Front tarsi 2- to 5-jointed, not reposing in a sulcus at the tip of the front tibiae 13 Front tarsi 1-jointed, reposing in a sulcus at the tip of the front tibiie ; claws small, almo.st straight. Legs four, the middle pair aborted ; antennse 3-jointed, compressed ; head and thorax subquadrate. Tetrapus Mayr. 13. The usually large anteunal fovea terminates posteriorly in a narrow or broad longitudinal slit ; if it does not, then the transverse oval anteunal fovea does not extend to the middle of the head 14 14. Body somewhat narrow ; anteunse 3- to 6-jointed. Front tarsi 2 or 3-jointed 16 Front tarsi 5-jointed. Thorax in outline trapezoidal ; antennte 6-jointed, with 3 ring-joints. Pleistodontes Saunders. Thorax not so shaped, either bell-shaped or ellip.soidal, antennai 3- to 6-joiuted 15 15. Thorax not oblong, in outline either ellipsoidal or bell-shaped ; antennse 3- to 6-jointed. Thorax ellipsoidal, slightly contracted at the sides before the middle ; antennse 6-jointed, with with one ring-joint and a 2-jointed club Kradibia Saunders. Thorax bell-shaped; antennae 3- or 4-jointed, with one ring-joint Eupristina Saunders. 16. Head about twice as long as wide ; front tarsi 2-jointed ; antennje 6-jointed Ceratosolens Mayr. Head not or scarcely longer than wide, at the most not more than one and one half times as long as wide ; front tarsi 3-jointed ; antennae 3-jointed. Head a little wider than long ; tarsi of hind legs very long, more than twice longer than their tibiae Blastophaga Gravenhorst. Head usually a little longer than wide, never wider than long ; tarsi of hind legs short, not or only a little longer than tibiae Eisenia Ashmead, gen. nov. Subfamily II. Sycoph.agix.e. 1883. Cynipidse, Division II., Aploastomata, Saunders, Trans. Ent. Soc. London, 1883, p. 20. Tliis group was first correctly separated by Sir Sidney Saunders, who designated it as Division II., Aploastomata, in the family Cynipidie. It has, however, nothing to do Avith the Cynipidse, but forms a natural group of the genuine fig-insects. At present no species is known outside of the Asiatic and African faunae. The females belonging to the group are distinguished from those in the pre\ious subfamily principally by the absence of mandibular appendages. The head, how- ever, is also somewhat differently shaped, being flatter, with a broader frontal sul- cus and no hook-like tubercle on the middle of the occiput, while the a]>domen is more depressed and the ventral valve is not prominent. The males are more ea.sily recognized, being quite different from those in the Agaonhifc and the wingless males in the Torymvlie. The head is very long and narrow, while the abdomen is long^ sessile, and gradually broadened towards apex, each lateral apical angle being furnished with a tubercle, or a long, thread-like organ. asumead: classification of thk chalcid flip:s 235 The genus I'hitijscajxi Motschulsky I do not know, Imt, judging from Mot- schulsky's description, it seems to be one of these insects, and I liave included it in my table with the hope that it may be rediscovered, and its proper position in our system definitely settled. TABLE OF GENERA. 1 . Females 2 Males 4 2. Postmarginal vein obsolete or nearly ; maxillary palpi with rows of teeth along the under side 3 Postmarginal vein well developed, longer than Ihe stigmal vein ; maxillary jialpi without rows of teeth beneath. Antenna; 13-jointed, with 2 ring-joints, of which the second is rather large ; liind tibiie with rows of comb-like teeth at apex ; stigmal vein not short Sycophaga Westwood (type Cynips sycomori HasseUiuist). Antenna- 9-jointed {teste Motschulsky) ; hind tiljia' normal ; stigmal vein short. Platyscapa MotschuLsky (type P. frontalis Motsch.). 3. Antenme 11-jointed ; hind tibia' without rows of teeth at apex Crossogaster Mayr (type C. triform is Mayr). 4. Apterous 5 Winged. Postmarginal vein subobsolete ; head w'ith a large, deep triangular fovea anteriorly that extends posteriorly to the middle of the head ; antenme 11-jointed Crossogaster Mayr. 5. Head oblong, without a deep triangular fovea anteriorly ; antennfe 3-jointed 6 Head nearly quadrate, a little wider than long, with a deep, triangular fovea anteriorly. Antennae 8-jointed, the scape thick, depressed ; body broad Crossogaster Mayr. 6. Scape of antennae about as broad as long ; mandibles broad, 3-jointed ; tarsal joints 2-4 wider than long ; abdomen terminating in two long filaments, one on each apical angle... Sycophaga Westwood. Scape of antennai long, subclavate, more than twice longer than thick ; mandibles conical, without teeth ; tarsal joints 2-4, not wider than long ; abdomen not terminating in two long filaments. Apocrypta Coquerel (type A. perpleia Coquerel). Family LXI. TORYMID/E. 1833. Torymida', Family II. (partim) Walker, Ent. Mag., I., p. 115. 184G. Torymidie, Family I., Walker, List Chalc. Brit. Museum, I., p. 14. 1848. Torymidffi, Family I., Walker, opus, cit, II., p. 100. 1856. Torymoidaj, Familie XL, Forster, Hyra. Stud., II., pp. 19, 23 and 43. 1875. Torymina, Tribus (partim) Thomson, Hyni. Skand., IV., pp. 11 and 59. 1886. Toryminte, Subfamily (partim), Howard, Ent. Amer., I., p. 198. 1897. Torymidte, Family XLL, Ashmead, Proc. Ent. See. Washington, IV., p. 243. This is a very large and most interesting family, approaching nearest, on one side, to the genuine fig-insects (Agaonid*) ; on the other side exhibiting affinities that ally it to the Chulclduhv, and some forms in the Misnx/dstcridir and the I'Uru- malidie. 236 MEMOIRS OF THE CARNEGIE MUSETTM Many of tlie genera, especially in my subfamily Iduridnx, were included by Walker, Westwood and Mayr, among the Agaonidx, a position not tenable. The genus Podagrion Spinola, too, on account of the swollen and dentate hind femora was placed by Mayr, Howard and others with the Ghakididse. The swollen and dentate hind femora, although of great taxonomic importance, in themselves should not alone be depended upon to place genera and species. Many genera and species are now known with such femora that unquestionably belong to other families. In the family Cleonymidte there is a whole subfamily Avith such hind femora and it would be absurd and most unnatural to classify it with the Chahididx on that ac- count alone. In establishing families many characters must be carefully considered, weighed and analyzed, and we should not be led astray by superficial resemblances or b}' characters common to many groups. Six fairly well defined subfamilies have l)een recognized. TABLE OF SUBFAMILIES. 1. Mesotlioracic furrows not well defined, the scapulae therefore scarcely or indistinctly separated ; abdo- men in 9 conically pointed, the ovipositor not exserted 6 Mesotboracic furrows distinctly defined, the scapulae therefore well separated ; ovipositor always prominently exserted, most frequently very long 2 2. Hind tibise with only one apical spur 5 Hind tibiie with two apical spurs. Posterior margin of the mesepisternnm incised beyond the middle ; metepimeron curved, dilated above the apex ; posterior femora simple, neither armed with a tooth nor serrate 3 Posterior margin of the mesepisternnm entire ; posterior femora rarely simple, more or less swollen, serrate or armed with one or two teeth beneath, sometimes much swollen and armed with several teeth beneath 4 3. Stigmal vein always long ; abdomen usually more or less depressed ; if subcompressed the hyjiopygium is large and prominent ; males frequently apterous, the head usually oblong, with a triangular fovea anteriorly in which lie the antennae ; abdomen short, never tubularly lengthened. Subfamily I. Idarnin^. Stigmal vein very short, the knob of same being sessile or subsessile ; abdomen usually subcompressed, the hypopygium not prominent ; males most frequently winged ; if apterous, the head not oblong and without a triangular fovea anteriorly Subfamily II. Toeymin^. 4. Hind femora not much swollen, as long as their tibiae, beneath feebly serrate or armed with one or two teeth beyond the middle towards apex, their tibiaj straight ; 'fcietanotiinJ' usually with a median carina, coarsely rugulose or punctate ; stigmal vein not long, but still longer than in the Torijminx, oblique and clavate Subfamily III. Monodontomerin^. Hind femora much swollen and armed with from four to cli/lit teeth, their tibiae arcuate ; metanotum with a A-sluiped carina ; stigmal vein sliort, the knob subsessile, similar to the Torijmlnie. Subfamily IV. PoDAGRiONiNiE. ASHMEAD : CLASSIFICATION OF THE CHALCID FLIES 237 5. Stigmal vein in front wings terminating in a large rounded or dilated knob or stignia, the basal nervure distinct, straight Subfamily V. Megastigmin.k. 6. Abdomen in ? conic-ovate or produced at apex, in (^ oblong, usually with a peculiar sculpture in both sexes, the middle segments most frequently with transverse rows of pits or rounded punctures ; stigmal vein very short Subfamily VI. Ormyrin^. Subfamily I. iDARNiNiE. 1846. Agaoiiida;, Family 6 (partim), Walker, List Chalc. Brit. INliiseum, I., p. 23. 1897. Idarninaj, Subfamily I., Ashmead, Proc. Ent. Soc. Washington, IV., p. 235. All of the species composing this subfemily are found associated with the genuine fig-insects ; they are either inquilinous or genuine parasites, as is the case in the Cynipidx and the Etirjjtomuhe, and in at least another group in the Torymidaj, i. e., the ^ fejast igmimi'. The group comes nearest to the subfamily Toryminie, agreeing with it closely in cephalic and thoracic characters, and in having two apical spurs on the hind tibi* ; but here the resemblance ceases, and it is easily separated by the difference in vena- tion, shape of abdomen, etc. The males are most frequently apterous, the head being long or oblong, with a triangular fovea anteriorly in which lie the antennae, a character also found in the Agaonidse ; but the abdomen is short and never tubularly lengthened nor broadened at the apex, as in that family. The females agree somewhat with the Agaonidx in venation, but differ decidedly in cephalic, mandibular, and abdominal characters, the abdomen being either de- pressed, imthout a prominent ventral valve, or subcompressed with a prominent ventral valve, with a long ovipositor. From the Toryminx they are at once distin- guished by the long stigmal vein and the different shape of the abdomen. TABLE OF GEXERA. 1. Females 1 Males 16 2. Abdomen normal, the last two segments not tubular 2 Abdomen abnormal, the last two segments very narrow and produced into a kind of tube or tail, the ovipositor very long 3 3. Antennje 10-jointed. 4 AntenniB 13-jointed, with three ring-joints; mesonotum witli distinct parapsidal furrows. Seventh abdominal segment very long, longer than the preceding segments united ; scape of antennffi scarcely thrice the length of the pedicel Philotrypesis Forster. (type Cynips caricve Hasselq.). Seventh abdominal segment hardly as long as the preceding segments united ; scape of antennie long, about four times as long as the pedicel Sycoscaptella Westw. (type S. affinia Westw.). 4. Prothorax long, conical ; metathorax long ; legs short and .-itout, the front femora incrassated. Polanisa Walker (type/rfarnes transiens Walk.). 238 MKMOIKS OF THE f'ARNEGIE MUSEUM 5. Scutellum«o/ flat, usually convex or at least subconvex, and usually, but not always, without longitudinal grooved lines C Scutellum flat and broad, quadrate, with Hvo longitudinal grooved lines. Metallic ; flagellum filiform, the joints of the funicle at least twice as long as thick ; ocelli ob- tusely triangularly arranged Idarnes Walker (type I. carme Walker). Non-metallic ; flagellum subelavate, the joints of the flagellum not or scarcely longer than thick ; ocelli arranged nearly in a straight line Koebelea Ashmead, g. nov. (type A', auslraliensis Ashm.). 6. Ovipositor shorter than the body and sometimes not at all prominent ' 11 Ovipositor longer than the body ; funicle of antennse 5-jointed 7 7. Abdomen with the ventral segments normal, not widened downwards 8 Abdomen with the ventral segments wideJied downwards. Hypopygium very prominent, plowshare shaped ; antennre 12-jointed, with 2 ring-joints. Goniogaster JIayr (type G. variiconiis Mayr). 8. Mesothoracic furrows distinct, complete 9 Mesothoracic furrows not distinct, wanting posteriorly. AntenniB 11-jointed, with one ring-joint inserted much nearer to the front margin of the bead than to the middle ; stigmal vein long, nearly three fourths the length of the marginal, Sycoryctes Mayr (type S. patellaris Mayr). 9. Autennte 12-jointed or less 10 Antennte 13-jointed, with three ring-joints inserted at an equal distance between the middle and the front margin of the head ; stigmal vein less than one third the length of the marginal. Trichaulus Mayr (type T. versicolor Mayr). 10. Antennte 12-jointed with one ring-joint; stigmal vein about half the length of the marginal. Apocryptophagus Ashmead, g. nov. (tj'pe Chakis '? explorator Coquerel). Antennae 9-joinied (teste Motschulsky) ; abdomen elliptic, much depressed, the ovipositor twice the length of the body Platyneura Motschulsky (type P. testacea Motsch.). 11. Postmargiual vein distinct, always present 12 Postmarginal vein wanting or never well developed 14 12. Ovipositor hidden 15 Ovipositor always prominent or exserted 13 13. Antennse 12-jointed with two ring-joints, the funicle 5-jointed, inseited near the clypeus, the joints of the flagellum fluted ; body metallic ; ovipositor about as long as the thorax and abdomen united or a little longer, and thickened towards apex. Colyostichus Mayr (type ? C. longicaudis jMayr, rf Heteradrivm longipes Mayr). Antennie 11-jointed with one ring-joint, the funicle 5-jointed, the club 3-jointed, the joints of the funicle hardly longer than thick ; ovipositor scarcely as long as the abdomen, gradually thick- ened towards apex. Heterandrium Mayr (type c? S. biannulatum Mayr, $ Colyostichns brevicaudis Mayr). 14. Scutellum with two longitudinal grooved Hues, the axillse widely separated ; ovipositor a little shorter than the abdomen Sycophila Walker (type S. decaiomoides Walk.). Scutellum luithout grooved lines, the axilhe nearly uniting at base of scutellum ; ovipositor only slightly projecting Froggattia Ashmead, g. nov. (type F. polita Ashm.). 15. Mesothoracic' fui rows indistinct, never complete ; antennic 9-jointed. Micranisa Walker (type Idarnes pteromaloides Walk.), ASHMEAD : CLASSII'ICATION OF THE (HALCIU I'LIES 239 16. Tarsi 5-jointed 17 Tarsi 3- or 4-jointed 31 17. Apterous or with rudimentary wings 18 Winged 34 18. Antennfe inserted on the anterior margin of the head or far below its middle ; clypeus not extending to the middle of the inner margin of eyes 1 "J Antennae inserted on the middle of the face 30 19. Head anteriorly with a large, deep, triangular fovea from which originate the antennrc 20 Head anteriorly without such a fovea ; wings entirely absent, not represented by filaments 2.5 20. Wings represented by two pairs of thread-like filaments 21 Wings repre.sented by one pair of thread-like filaments or by bristles 22 21 Head oblong, not or scarcely wider than the thorax, the eyes minute ; antennas separated by a sharp carina ; hind tarsi with the first joint dilated Philotrypesis Forster. Head large, oblong, .slightly narrowed anteriorly, the liiiid niarfjiii simuite, the hind angles rounded, setose ; antennae 8- or 9-jointed with 1 ring-joint, the scape large, davate ; mandibles large, [lorrect, curved, acute and entire at apex, but within, 3-dentate. Tetranemopteryx A.shmead, g. nov. {tyite Sycoseajder J,-Kelosa Westw.). 22. Head oblong, the hind angles rounded, not acute 23 Head oblong-quadrate, the hind angles acutely produced, not rounded 24 23. Head large, oblong, broader than the thorax ; antenna; 10-joiuted, with 1 ring-joint inserted close together near the front margin of the head ; mandibles porrect, stout and curved, the tips acute, entire, the inner margin with a tooth behind the middle Sycoscapter Westwood (type iS'. insignis Westw.). Head oblong-quadrate, sinuate posteriorly ; antcnnte 8- or 'J-jointed, without a ring-joint, inserted close together near the mouth ; mandibles porrect, falcate, the tips bidentate, the inner margin with a tooth near the base Sycoscapteridea Ashmead, g. nov. (type Sycoscapter monilifer). 24. Head oblong, the sides anteriorly nearly straight, the front angles rounded, the hind angles acutely produced ; antenna; 8-joiuted (or 9-jointed with 1 ring-joint), inserted close together near the mouth ; mandibles small, falcate, entire; wings represented by short filaments. Sycoscapterella Ashmead, g. nov. (type Syam-aptcr anguliceps Westw.). 25. AntenuiE shorter, at the most 9-jointed, the joints closely united, inserted further from each other than to the sides of the head, most frequently with only 4 or .') free joints 26 Antennae longer, 10- or 11-jointed, with only the last three joints closely united, inserted near or not far away from the mouth border -~ 26. Head between the clypeus and the sides concave ; mandibles bidentate ; lal)iuni and paliii wanting, in place is a membranous tube; antenn;e coniposein'i-uiUy sliort ; inetatliorax declivous, not largely punctate. Torymus Dalman (tj-pe Ichneumon hedef/uarix Liunel. Prothorax short, the metathorax abruptly declining, largely punctate ; antenna; subclavate, the club lanceolate Torymoides Walker (type T. amabilis Walker). 8. Antenna; 13-jointed, with only one ring-joint 9 Antenna; 13-jointed, with livo ring-joints Lochites Forster. 9. Scutellum without a cross-furrow before apex 10 Scutellum with a cross-furrow before apex Syntomaspis Funster. 10. Mandibles 3-dentate ; costal cell wide 11 Mandibles 2-dentate ; costal cell narrow Callimomus Thomson. 11. Clypeus anteriorly normal, not produced 12 Clypeus anteriorly subi)roduced medially Lioterphus Thomson. 1 2. Thorax normal, the pronotum not especially long 13 Thoi-ax long, the pronotum elongate, conical Ecdamua Walker. 13. Pronotum not especially short, the metathorax not largely punctate Torymus Dalman. Pronotum short, the metathorax abruptly declivous, largely punctate Torymdides Walker. Subfamily III. Monodontomerin.'e. 1875. Toryrnides, Subtribus (partim) Thomson, Skancl. Hym., IV., p. 59. 1899. Monodontomerinse, Subfamily III., Ashm .Proc. Ent. Soc. Wash., IV., p. 247. In this group the hind femora are more swollen than in the previous groups, while the lower edge is serrate or armed with one or two teeth, thus showing an approach to the Podagrioninie, and through tliat subfamily to the Leucospidinic and the Cludcidinse. Some of the species are parasitic upon hymenopterous and dipterous gall-mak- ing insects; other.s, belonging to the genera Physothorax and Plesiostigma, have been bred from fig-insects, while species belonging to the genera Diamorus and Monodon- tomcrus are bred commonly from the nests of bees and wasps, and they are said to be parasites of these insects. In a single case, at least, I have positive evidence that Monodontomems was bred from the puparium of a tachinid-fly found in the nest with the bee. TABLE OF GENERA. 1. Females ; ovipositor i)rominently exserted 2 IMales 9 2 Hind femora beneath very finely denticulate and usually also with a single large tooth, rarely with 2 teth 7 Hind femora beneath smooth, with 1 or 2 large teeth, some distance from the apex 3 3. Scutellum with a cross furrow before the apex 4 Scutellum without a cross furrow before the apex. Abdomen shoi-t, not longer than the head and thorax united ; stigmal vein very short. Holaspis Mayr (type Torymus militaris Boheman). Abdomen elongate, much longer than the head and thorax united ; stigmal vein not short, oblique Websterellus Ashmead (type Iritici). ashmead: classification ov the ciiALcin ki,iks 243 4. Abdomen with the hind margin of the first dorsal segment (or its flap) incised at the middle 5 Abdomen with the hind margin of tlie first dorsal segment straight, not incised. Monodontomerus Westwood (tyi)e ilf. ohsrurtis Westw.). 5. Funicle with 6 joints ; head smooth or at most with sparse punctures (> Funicle with 7 joints ; head and thorax with large, thimble-like imnolures. Hind femora with a single tooth beneath towards apex Diamorus Walker (type 'Ihnjinii^armatus Boheman). Hind femora with two teeth beneath toward apex Physothorax JIayr (type Diamorus variabilis Mayr ?, Physothorax disciger Mayrcf). ti. Thorax delicately shagreencd ; hine Ifhnnimon dort^dix Fahricius. SUJ'.KAMILY Y]. OrmykiN.E. 185G. Orniyruichu, Ftuiiilio X., lAirstei-, Hyin. Stud., II., pp. 1!), 22 and 24. 1875. Ormyrides, Sul)ti-il»ii.s, Tliomson, Skuiid. Ilyni., IV., p. 100. 1899. Orinyi-ina% Suhfaiuily V., Ashmead, Proc. Ent. Soc. Wash., IV'., p. 247. Df. Aniold Frirstc'i- was the Hrst to sepai-ate tliis gfoup from other I'oiymids. It is a very distinct and compact subfamily, agreeing with the sul)family Toryniin;e in venation and in pleural and pedal characters, but otherwise it is totally different from it and the others. The females are easily recognized by the elongate, pointed, orconically produced abdomen, non-prominent ovipositor, and by its peculiar sculp- ture ; the males by the oblong oval shape of the abdomen, as well as by the sculpture. It shows some affinity with the KurytomvJiv, and particularly with my tril)e Kilei/ivi, in the shape of the pronotum, and in antennal and abdominal peculiarities. All are bred from hymenopterous and dipterous gall-insects, belonging princi- pally to the families Ciiitiphhf and QxlilonujUdn' ; oidy a few species have been described. Only three genera are known distinguished as follows : TABLE OF GENERA. 1. Males ; abdomen oblong, depressed 4 Females ; abdomen long, conically i)ointed and nioi-e or less eonipres.sed, especially towards the ajiex, the ovipositor never prominent 2 2. Abdomen without a. peculiar sculpture, normal ;? Abdomen with a peculiar sculpture, and some of the .segments wit h two rows of pits or dee]) punctures. No large, coarse punctures at base of middle abdominal segments ; antenuie with one ring- joint Monobseus Foreter (type M. cingulatus Foi-st.). With large, coarse punctures at base of middle abdominal segments ; antenme with two ring- .joints Ormyrus Westwood (type 0. puncliger Westw.). 3. Abdomen (inely iiunctate ; antenna; with three ring-joints. Tribaeus F6rster(type T. punctulaliis Forst.). 4. Abdomen without a peculiar sculpture 5 Abdomen with a peculiar sculpture. Antennffi with one ring-joint Monobaeus Fiirster. Antennte with two ring-joints Ormyrus Westwood. 5. Antennre with three ring-joints Tribaeus Fiirster. 246 memoirs of the carnegie museum Family LXII. CHALCIDID.E. 1830. Chalcidfe, Family (paiiim), Leach, Edinb. Encyc, IX., p. 144. 1840. ChalcidsE, Subfaniil}^ I. (partim), Westwood, Intro. Mod. Class. Ins., I., p. 10(5, Synop., p. 65. 1846. Chalcididfc, Family II. (partim), Walker, List Clialc. Brit. Mils., I., p. 2. 1850. Chalcidoida;, Familie II., Forster, Hym. Stud., II., pp. 18, 21 and 29. 1875. Chalcidina Tribus, Thomson, Hym. Skand., IV., i^p. 11, 12. 1886. Chalcidinai, Subfamily (partim), Howard, Ent. Amer , I., p. 197. 1897. Chalcididte, Family LXIL, Ashmead, Proc. Ent. Soc. Washington, IV., p. 245. 1900. Chalcididai, Family LXIL, Ashmead, Proc. U. S. National Museum, XXIII. , p. 202. This family, although allied to the Torymidx and to the EarytomidiV, is very distinct in many particulars ; by the usually very long, subcylindrical hind coxie, the greatly swollen hind femora, usually dentate or serrate beneath, by the strongly arcuate hind tibia3 which are usually obliquely truncately produced at apex, so that the tarsi appear to be attached a little before the tips, and by pro- notal and abdominal differences. The group is most extensively represented in South America, where many genera and many species have been discovered. Two subfamilies have been recognized, distinguishaljle as follows: TABLE OF SUBFAMILIES. Front wings longitiulinallj' folded ; ovipo.sitor curving upwards and bacliwards over the dorsum of the abdomen Subfamily I. Leucospidin.e. Front wings not folded ; ovipositor when prominent, straight, not curving over the dorsum of the abdomen Subfamily II. CuALClDlNiE. Subfamily I. LEUcosPiDiNiE. 1833. Leucopsida^ Family, Walker, Ent. Mag., II., p. 13. 1839. Leucospidaj, Family XV., Haliday, Hym. Synop., p. ii. 1846. Leucospidee, Family I., Walker, List Chalc. Brit. Museum, I., p. 1. 1856. Leucospoidte, Family I., Forster, Hym. Stud., II., pp. 18, 20 and 29. 1886. Leucospinte, Subfamily, Howard, Ent. Amer., I., p. 197. 1897. Leucospidinte, Subfamily I., Ashmead, Proc. Ent. Soc. Wash., IV., p. 235. Dr. von Dalla Torre, in his Catalogus Hymenopterorum, Vol. V., has incor- rectly credited this subfamily to Forster ; it should be credited to Walker, who desig- nated it as a family as earl 3'^ as 1833. The group is a natural one, and is very distinct from the other groups of the Chal- cididve, in haljits, in abdominal peculiarities — the ovipositor being curved forward ASHMEAD : CLASSIFICATION OF THE CIIALCin FLIES 247 over tlie dorsum of tlie abdomen, often reacliing to tlie scutelluin, and in haviiij^; the wings longitudinally folded as in the \'. Scape of antenna; normal ; body of abdomen in female fusiformly pointed or conic-ovate, the petiole very short, not or rarely longer than thick Spilochalcis Thomson (partim). Scape of antenna; usually long and extending much beyond the ocelli ; body i>f abdomen ovate, the petiole very long and slender, as long or nearly as the thorax .... Mischosmicra Ashmead, gen. nov. (type M. Kahlii Ashm.). 13. Metathorax normal or with one small tooth on each side of the petiole ; abdomen variable, subglobose, conic-ovate or fusiform, but rarely ending in a stylus ; antennae K! jointed. Spilochalcis Tliomson (partim). 14. Antenme 13- or U-jointed 15 Antennae 12-jointed. Metathorax po.steriorly y (type Smicra Iriicotelus Walk.). 18. Scutellum at apex usually ending in an emarginate or bidentate plate ; abdomen in female lanceolate or conically produced ; the Sth segment often long, styliform. Octosmicra Ashmead, gen. nov. (type O. lalicepf) A.shm.). 19. Hind femora armed with 6 large teeth or less 20 Hind femora armed with 7 large teeth, the seventh usually much reduced in size. Abdomen in female conically produced or fusiform, the 8th segment sometimes stj lifonn or pro- duced into a stylus, the petiole not short ; mandibles in 9 hi- in r^ tri-dentate. Heptasmicra Ashmead, gen. nov. (type Smicra oblitcrata Walk.). 20. Hind femora armed with 5 large teeth or less 21 Hind femora armed with 6 large teeth (the 6th tootli sometimes broad and at apex 3-deutate). Metathorax armed with two triangular teeth and a lamina or plate just behind ihe insertion of the hind wings, the latter extending slightly over the metapleura, the sculpture coarse ; teeth of hind femora short and stout Diplodontia Ashmead (type Smiera Carolina Ashm.). Metathorax usually, but not always, armed with two, not very prominent, teeth, but wiihoutthe lamina or plate behind the insertion of the hind wings, the seulpture alveolate ; teeth of hind femora large, long. Abdomen in female .subglobose or short ovate, the petiole always more than twice longer than thick Metadontia Ashmead (iype'»icm montana Ashm.). Abdomen in female lanceolate or fusiformly produced, longer than the head and thorax united, the petiole very short, net longer than thick. Hexasmicra Ashmead, gen. nov. (type Smic7-a transrersa Walk.). 21. Hind femora armed with 4 large teeth or less 22 Hind femora armed with 5 large teeth. Eyes very large, occupying nearly the whole sides of the head ; mandil)Ies broad, 3-denlate ; scutellum usually ending in an emarginate or bidentate plate ; metathorax alveolate. Pentasmicra Ashmead, gen. nov. (type P. brasUiensis Ashm.). 22. Hind femora with 4 large teeth ; metathorax usually with teeth or projections, rarely unarmed ; plate at apex of scutellum bidentate or broadly emarginate ; abdomen ovate or conic-ovate, rarely -ftisiform, the eighth segment in female never very long Tetrasmicra Ashmead, gen. nov (type Smicra concUata Walk.). Hind femoia with 3 large teeth Trismicra Ashmead, g. nov. (type S. contractu Walk.). 23. Middle tibiie without an apical sjiur 24 Middle tibise with an apical spur 25 ASIIMEAD ; 'LASSIl-ICATION OF THF CIIAI.cn) FI.IF.S 2~)^ 24. Antemue 12- or 13-joiiited ; tlionix l)l;uk ; liiiid fi'ii\(ini aniicil with iiiiiiiy minute teetli. .Smicra Sjiinohi Aiiti'iiiia; 14-joiuted ; thorax yellow or yellow niarkcd with black ; liiiul femora armed with (> teeth. Epitranus Walker. 25. Hind femora armed with one large tooth near base followed by many .small or minute teeth, from 10 to 20 or more 2G Hind femora armed with from H to 9 large teeth 30 26. Metathorax armed with from 2 to 4 teeth or projections 29 Metathorax unarmed. Scutelliim at apex unarmed, without an emarginate or bidcntate plate 27 Scutellum at apex armed with an emarginate or bidentate plate. 27. Abdominal petiole very long and slender, as long as the thorax or nearly 28 Abdominal petiole usually short and stout, or at least never very long nor very slender. Pronotum anteriorly rounded or sloping, not acute ; jjetiole carinate. Eustypiura Ashmead. Pronotum anteriorly acute ; petiole not carinate Spilcchalcis Thomson. 28. Scape of anteume abnormal, long and clavate, extending far al)ove the ocelli. Ceratosmicra .Vshmead. Scape of antennsE slender, cylindrical, not or only slight extending above the ocelli. Sayiella .Vshmead. 29. Metathorax bi-dentate ; a tooth on each side of the petiole. Scape of antennre normal ; abdominal petiole not very long nor slender. Spilcchalcis Thorns. ( partim). Scape of antenme usually long and slender, extending much beyond the ocelli ; abdominal petiole very long and slender, as long or nearly as long as the tlioi ax. Mischosmicra Ashmead. Metathorax quadridentate ; two teeth on each side of the petiole. Scutellum at apex, bidentate or with an emarginate plate Xanthomelanus Ashmead. Scutellum at apex normal, unarmed. Thorax wholly black ; scape of antenn;e normal, petiole of abdomen normal. Melanosmicra Ashmead. Thorax yellow or yellow marked with black ; scape of antenmc abnormal ; petiole of ab- domen very long. Scape long and clavate, excavate at apex beneath, extending far above the ocelli. Ceratosmicra .Vshmead. Scape slender, cylindrical and extending only slightly beyond the ocelli. Sayiella Ashmead. 30. Antenntc 13- or 14-)ointed 31 Antenmc 12-jointed : metatliorax iiuudii.le..l..l Thaumapus Kirby. 31. Antennae 13-jointed ■52 AntenniC 14-jointed. Hind femora armed with large or modcratesi/ed teeth Epinseus Kirby. 32. Hind femora armed with 8 large teeth or less 33 Hind femora armed with 9 moderately large teetli Enneasmicra Ashmead. 33. Hind femora armed with 7 large teeth or less 34 Hind femora armed with 8 large teeth. Mesonotum without distinct parapsidal furrow Protoceras Kirby. Mesonotum with distinct parapsidal furrows Octosmicra Ashmead. 34. Hind femora with 6 large teeth or less 35 Hind femora with 7 large teeth Heptasmicra Ashmead. 254 MKMOIKS OF THE CARNEGIE MUSEUM 35. Hind femora armed witli 5 large teeth or less 3G Hind femora armed with 6 large teeth. Metathorax armed with two triangular teeth or projections and a lamina or plate just behind the insertion of the hind wings that extends slightly over the mesopleura ; teeth of hind femora short and stout Diplodontia Ashmead. Metathorax sometimes armed with two teeth, sometimes unarmed but always without the lamina or plate. ? cf unknown Metadontia Ashmead. ? cf unknown Hexasmicra Ashmead. 36. Hind femora armed with 4 large teeth or less 37 Hind femora armed with 5 large teeth Pentasmicra Ashmead. 37. Hind femora ai-med with 4 large teeth Tetrasmicra Ashmead. Hind femora armed with 3 large teeth Trismicra Ashmead. Tribe III. Chalciiclliui. This tribe is proposed for a small group closely allied to the tribe HaltichcJIini, but is easily distinguished by the distinctly petiolate abdomen. The group is unknown to the American fauna, although some South American genera in the Smicrini come quite close to it; they are, however, easily separated b}'^ having the antennae inserted farther up on the face and not low down close to the mouth, as in this tiibe. TABLE OF GENERA. 1. Females 2 Males 4 2. Antennte 13-jointed 3 Antennae 11-jointed (the club indistinctly jointed). Hind femora with 7 or 8 large teeth Chalcitella West wood (type C. evanioides Westw ) 3. Hind tibise without a tooth outwardly near base, the hind femora armed with many very minute teeth. Arretocera Kirby (type Epitranus albipennis Walk.). Hind tibise with a tooth outwardly near base, the hind femora with one large tooth and then finely serrate beyond Anacryptua Kirby (type Epitranus impulsulor Walk.). 4. Characters as in females. Tribe IV. HaUichdlini. This tribe resembles most closely the tribe Chalcidini and is separated from it with dicuffilty, the difference in the insertion of the antennae alone being the only character tliat readily distinguishes the two tribes. In this trilje the antennse are inserted close to the mouth border, the scape being long, while the postmarginal vein is wanting, or rarely vei-y long as in the Chakidhd and the Sinicrini. TABLE OF GENERA. 1. Females 2 Males 17 ASHMEAI) : CLASSIKICA'IUOX OF TllK CllAI-CII) KI.IKS 255 2. Tliiul femora with one or more large teetli or projections beneatli, as well as heini; finely denticulate or serrate 3 Hind femora simple or at most very finely (lenticulate beneath, without a larf;e to- iin.sti rioily ; sul.costal vein ending in a knoU, tin- marginal and stignial veins not .li'Velop.-d Hybothorax Katzelmrg. Metathorax sliurt, ending in two long diveigent spines; marginal and stigmal veins present. Kriechbaumerella Dal la Torre. 30. Antenme 10-juinled (or 1 1-Jointed witli a ring-joint) Bippota Walker. TiMUF V. Dirhhih,;. Tltis tribe is distintiuislicd from all tliu utlan-s by biiviiig tlic head deu[)ly eniar- ginate and honied. The anteniue ure inserted close to the mouth as in the Haltichellhil. TABLIC OF GENERA. 1. Female 2 Males 4 2. Ovipositor not exserted ; head with two horns 3 Ovipositor strongly exserted, long; head with /oiir hums ; antennie l;;-Jointed. Hontalia t'anu'ron (lyi)e H. civrulea Cameron). 3. Metathorax strongly bidentate. AntenniE 12-jointed Dirhinus Dalnian (type D. excavatiis Dalman). Antennfe 13-jointed Eniaca KirUy (type Chnjsis hesperidum Rossi). 4. Head with hvo horns 5 Head with four horns. Metathorax without teeth ; antenme lij-jointed Hontalia Cameron. 5. Metathorax strongly bidentate. Antenme 12-jointed Dirhinus Dalman. Antennae 13-jointed Eniaca Kirby. Family L.XIIl. " EURYTOMID.E. 1830. Cynipsida Leaeh, Kdinb. Kncye., IX., p. 144. 1833. Eurytoniida^ Family 1., Walker, Ent. Mag., I., p. 12. 1840. Eurytomides, Subfamily 2, Westwood, Intro. Mod. Class. Ins., II., p. 166; Synop., p. ()(). 1816. Eurytoniid;e, Family 3, AValker, List Chalc. IJrit. Museum, i., p. 8. 1856. Eurytomoidte, Familie Xll., Fr.rster, Hym. Stud., ii., pi). 19, 23, 44. 1875. Eurytomina, Tribus, Thomson, Hym. Skand., IV., pp. 11, 25. 1886. Eurytominte, Subftimily, Howard, Ent. Amer., I., p. 19. 1897. EurytomidiP, Family LXIII, Ashmead, Troc. Ent. Soc. Wash., IV, p. 245. On account of the diversity of habits among the various groups composing this family it is one of the most interesting of all of the families in this great complex to study, except possibly the family Aguonidu. 258 .MEMOIRS OF THE CARNEGIE MUSEUM Some are phytophagous or gall-makers; others live in the nests of Vjees and wasps; others are parasitic upon gall-making Hymenoptera and Diptera; while still others are genuine parasites on Coleoptera and other insects, and a few, repre- senting my tribe RUeyini, destroj'^ the eggs of orthopterous insects. The family may be separated into five tribes as follows: .*^^ ... .>j^^ J'^^'^^^nP'*^' TABLE OF TRIBES. rj^^ orax seen from above short, not lousrer than the scutellum, usually d 1. Metathorax seen from above short, not longer than the scutellum, usually distinctly shorter, and some- what abruptly declivous 4 Metathorax seen from above long, always longer than the .scutellum and usually gradually sloping to apex (in a single case quadrate and squarely truncate behind) 2 2. Head not cornuted ; eyes oval or ovate, not round 3 Head cornuted, much wider than the thorax ; eyes round Tribe I. Aximiui. 3. Marginal vein slender, longer than the stigmal or rarely shorter, but never very stout or stigmated ; abdomen most frequently long in both sexes, in female conic-ovate or couically pointed ; antenuse dissimilar in the sexes, in females usually subclavate, in males with the joints of the funicle con- stricted or excised at apex with whorls of long haire Tribe H. Isosomini. 4. Marginal vein short and stout, stigmated, either quadrate or semicircular 6 Marginal vein usually slender, linear, rarely stout, but never stigmated, quadrate, or semicircular 5 5. AntennsB 10- to 12-jointed, with only one ring-joint, and dissimilar in the sexes, in females filiform or subclavate, in males with the funicle joints excised or petiolate at apex, with whorls of long haiis or with sparse long hairs Tribe III. Eurytomini. AntennsB 13-jointed, with two or three ring-joints, and veiy .similar in the sexes, the funicle joints not excised or pedicellate at apex, and without whorls of long hairs Tribe IV. Rileyiui. 6. Antennse at the most 11-jointed, with 1 ring-joint, the funicle 4-jointed and very similar in both sexes, filiform or subclavate, without whorls of long hairs ; abdomen short, globose, or oval, never very long or strongly compressed ; hind tibife with rigid bristles behind Tribe V. Decatomini. Tribe I. Aximlni. Mr. Peter Cameron, in Biol. Centr. Amer. Hyra., Vol. 1, p. Ill, established for Axima Walker and HontaUa Cameron, the subfamily A.rtiuhiiv. The two genera, however, have nothing in common, are quite distinct in structural characters and belong to different families. Mr. Cameron was evidently deceived in regard to their relationship by a superficial resemblance in the structure of the heads of these insects. Axima is clearly a eurytomid, as I first pointed out several years ago, while Hoiitiiliu is a chalcidid allied to Dirhhius, and must be placed in my tribe Dirhhiini. Axima is parasitic in the nests of the small carpenter bees, CeraUnUhc. TABLE OF GENERA. • 1. Females 2 Males 3 2. Marginal vein four or more times longer than the stigmal vein ; antennre ll-joiuted, the funicle 6- jointed ; abdomen long, lanceolate, compressed Axima Walker (type A. spin! fro its Walk.). ASnMEAD : CLASSIFrCATrOX OF TITR CirAT.rTD FLTF.S 259 Marginal vein hardly twice the length of the stiguial vein ; antenna' ll-j()iMtclaced within it, but always above it near the anterior margin of the vertex 8 Head with a deep frontal channel or antennal furrow, the front ocellus always placed at the apex of this furrow, never above it. Scape elongate, more than twice longer than the first joint of the funicle, the funicle joints rather long, the first about twice as long as thick ; abdomen compres.sed, not longer than the head and thorax united, ending in a conical point ; the dorsum, as seen from the side, is highly convexly elevated ; postmarginal vein variable, sometimes shorter than the marginal, but rarely very much longer; hind tibia' with rather .short, stifl!' bristles behind. Prodecatoma Ashmead, gen. nov. (type P. ftarescens Ashm.). 262 MEMOIRS OF THE CARNEGIE MI^SEUM 8. Marginal vein always distinctly longer than the stigmal vein, the postmarginal vein well developed, sometiines very long 9 Marginal vein short, not, or scarcely, longer than the stigmal vein, usually a little shorter, the post-mar- ginal vein rarely well developed, rarely as long as the stigmal (in only a single case is it very long) . . 13 9. Hind tibise with 2 apical spurs 10 Hind tibias with 1 apical spur. Funicle with the joints oval-moniliform ; eyes broadly oval. Phylloxeroxenus Ashmead (type Eurytoma phyllexerse Ashm.). 10. Postmarginal vein very long, fully twice as long (or even longer) as the stigmal vein 11 Postmarginal vein not much longer than the stigmal vein. Marginal vein scarcely longer than the stigmal ; antennae clavate, the club large, 3-jointed, the joints of the funicle moniliform ; abdomen with the fifth segment, as seen from the side, two or more times longer than wide ; head convex in front, the occiput deeply concave. Eurytcmocliaris Ashmead (type E. mimda Ashm.). Marginal vein very distinctly longer than the stigmal ; antennaj filiform or nearly, at most sub- clavate, not greatly thickened towards apex, the joints of the funicle oblong, cylindrical ; abdomen conic-ovate, subcompressed, the fifth segment, as seen from the side, shorter than wide Eurytoma Illiger (type E. planata Illig.). 11. Hind tibi;e normal, not dilated 12 Hind tibiie compressed, dilated. Head transverse, wider than the thorax, and thin antero-posteriorly, the eyes more or less rounded, prominent ; antennse inserted far above the middle of the face, the scape very long, reaching far above the ocelli and with a tooth or tubercle at apex beneath, the flagellum filiform with sparse hairs, the iunicle joints more than twice longer than thick ; abdomen very strongly compressed, the petiole long and slender. Eudoxinna Walker (type Sesxetra transversa Walk.). 12. Marginal vein at least one and a half times as long as the stigmal vein, the postmarginal vein not longer than the stigmal ; abdomen conic-ovate, longer than the thorax, the fifth segment nearly twice as long as the fourth ; antennse 11-jointed, the flagellum subclavate, the joints of the funicle submoniliform Xanthosoma Ashmead (type A', nigricornis Ashm.). Marginal vein not or scarcely longer than the stigmal vein and stout ; antennaj 11-jointed, the fla- gellum clavate or subclavate ; abdomen globose, or short ovate, shorter than the thorax, the segments subequal Systole Walker (type S. albipennis Walk.). 13. Abdomen short, subglobose, the fourth segment much the longest, enclosing the following. Systolodes Ashmead (type S. brevicornis Ashm.). Abdomen ovate, subcompressed (the tip sometimes produced into a stylus), the fourth and fifth seg- ments short, although a little longer than the others, and subequal Bruchophagus Ashmead (type B. boreal is Ashm.). 14. Head with a deep frontal furrow ; abdomen ovate, subcompressed, petiolate, the jietiolc usually not short, the fifth segment the longest, but not greatly longer than the fourth. Funicle 5-jointed, the joints moniliform or submonilifoiin, the first much shorter than the pedicel Decatomidea Ashmead (type D. xanthochroa Ashm.). 1-5. Non-metallic, smooth, delicately shagreened or umbilicate punctate, raiely finely punctate 10 Metallic green or blue, coarsely umbilicately punctate. Head in front quadricarinate, the carina along the orbits sometimes delicate ; funicle with the joints excised at apex above, with whorls of long hairs Chryseida i^pinola. ASIIMKAI) : CLASSIFICATION OF TIIF, CHALCII) FLIES 263 IG. Mcsoiiotum closuly, distinctly, uiiil)ilicately imnctiite 17 Mesoiiotuin iwl umbilifatcly inim-tate, litlKT smootli or nearly, or shaj^reuiii'd or finely, re-^nlarly sculp- tured 0(; 17. First joint of the llagellum long, as long as the scape or longer IS First joint of the flagellum not especially long, always much shorter than the scape IS) 18. Head with a deep antennal furrow, the front ocellus placed within tlie furrow. ? Known to me in ? sex only Bephrata Canici-on. ? Known to me in 9 sex only Aximogastra AshmeaXV., Ashmead, Proc. Ent. Soc. Washington, IV., p. 235. 1899. Eucharidse, Family LXV., Ashmead, loc. cit, p. 245. ASHMEAD : CLASSIFICATION OF THE CIIALCID FMKS 267 1900. EuclKiridiC, Family LXV., Aslunead, Proc. U. S. National Museum, XX III., p. 202. In this family are found some of the most singular looking and wonderfully shaped Chalcids known, the structure of the thorax, and particularly of the scutel- lum, being most wonderfully and curiously modified and developed ; and this de- velopment, in connection with the brilliant metallic green and blue colors of its members, makes the group the most striking and attractive of any in the Superfam- ily. Some of the species are now known to be parasitic upon ants and probalily the whole group attacks these insects. In temperate regions the family is poorly represented, l)ut in tropical counti-ics, where ants most abound and flourish in enor- mous colonies, these insects are not rare and seem to have reached a very highly specialized development. The known genera may l)e tabulated as follows : TABLE OF GENERA. 1. Females 2 Males -1 2. Antennw 13-jointed or less 3 Antennae 14-pointed or more. Scutellum simple; antennae 16-18-jointe sliiatc ; aiitcnii.i' 14-joinlc(l. Saccharissa Kii-liy (type Kucliiiri.^ ronlingcns Walker). 3. Scutellum simple, neither bidentate nor produced into long processes 4 Scutellum bidentate or produced posteriorly into long processes over the al«lomen 11 4. Antenna; moniliform -"J Antenna; not moniliform f> r>. Abdomen compressed, ascending Eucharis Lalreille (type Ci/nlps ndsccndcns Falir.). .'Vhdonicn neither compressed nor ascending. Hind tarsi with the first joint nuuh tliickened ; antenna- 1 1-jointed Tricoryna Kiihy (ty|ic Eucharis jello Walt.). Ilin.l tarsi with the lirst joint very hmg, l)at not thickened Metagea Kirby (type Eucharis Zaliitcs Walk.). f>. .Joints of antenna' not serrate, cylindrical ' •Joints of antenna; serrate or subdentate !' 7. .\ntenna' 13jointed, the joints short !^ .\ntenna' 1 1-jointed, the joints long. Thorax smooth, polislicd ; i)eti()le of abdomen abruptly enlarged at apex. Pseudometagea Ashmead (type Metagea schicarzii Ashni.). Thorax rugose : petiole of alxlonien normal, long and cylindrical .Psilogaster Blanchard (type P. ciiprciis lilanchard). 8. Thora.x not greatly elevated, similar to Chrysolampus in the l'erilanipida>, punctate and with complete panipsidal furrows ; mandildes long, acute at apex, the right mandible with two teeth within, the left with one tooth within Orasema Cameron (type O. siramineipes Cam.). 268 MEMOIRS OF THE CARNEGIE MUSEUM 9. Antennae 1 l-jointed 10 Antennic 13-jointed JlMpipallus Kirby (type Enchnrix vohisnK AValk.). 10. Scutellum rounded, not conically elevated posteriorly ; wings hyaline. Pseudochalcura Ashniead g. nov. (type Eucliai-is ylhbosa Provaneher). Scutellum subconically elevated posteriorly; wings with a substigmal cloud or fascia. Chalcura Kirby (type Eucharis deprivafa Walk.). 11. Scutellum with the processes verj' long, usually as long as the abdomen and sometimes very broad, forming a shield over the abdomen, or conically produced 13 Scutellum bidentate, the processes never very long. Metathorax armed with strong lateral projections or teeth 12 Metathorax unarmed, without teeth. A hump-like elevation above the metapleura Stilbula Spinola (type Ichneumon cynipiformis Rossi). No hump-like elevation above the metapleura ScMzaspidia We.stwood (type S. furcifera Westw.). 12. Metathoracic processes curving downwards LophyTOcera Cameron (type L. straminelpes Cam.). Metathoracic processes con.sisting of two diverging horizontal teeth Tetramelia Kirby (type Schizaspidia phigiafa Walk.). 13. Scutellum not conically produced 14 Scutellum conically produced over the abilomen. Head with a deep antennal furrow ; hind femora very broad ; abdomen subsessile, fusifoiTn, depressed Destefania Dalla Torre (type Sternodes Pasaieri De Stef ). 14. Seutellar processes long and slender, generally curving inward toward tips 18 Scutellar processes very broad and covering the entire abdomen. Thorax not pubescent, the apex of the scutellar processes simple, or cleft or notched 15 Thorax pubescent, the apex of the scutellar processes rounded and not sharply cleft, the notch extending two thirds the entire length 1" 15. Scutellar processes long, broad and contiguous, but very flat, the extremities rounded, subtruncate, or furnished with two rounded short .spines 16 Scutellar processes not so shaped. Scutellar processes very broad, triangular .... Thoracantha Latreille (type T. latreillei Gueriii). Seutellar processes long, contiguous and acutelj' pointed at tips, longitudinally striate. Uromelia Kirby (type Thoracantha striata Perty). 16. Mesonotum and scutellum medially impressed ; head almost as wide as the thorax ; antennae lO-jointed, the third joint as long as the scape, the following much wider than long Dicoelothorax Ashmead (type D. plftli/rerii-fi Ashm.). Mesonotum and .scutellum not so impressed, the scutellar processes having the basal portion as wide as the thorax, briefly compressed in the center, then dilated and at the apex furnished with two rounded short spines Laetocantha Shipp (type Thoracantha nasvn Walk.). 17. Antenmc 10-jointcd, the first funicle joint the longest, the following short Dilocantha Shipp (type TJioracantha flaricornh AValk.). 18. Head and eyes noi-mal, not tubereulate 19 Head and eyes tubereulate. Antenna; 12-jointcd Isomeralia Shipp (type Thonicnntha coronala Westw.). 19. Antenna; 11-jointed, the third joint not much longer than the fourth 20 ASHJtRAD : PLASSIFIfATION OF THE CHALCIP FIJES 269 Antenna? 10-jointe-jointed 29 Antennic 18-jointed Saccharissa Kirby. 2.3. Antenna; 10-13-jointed 24 Antenna; 22-jointcd Eucharissa Westwood. 24. Antenmo simjile, witliout branches 25 AntenniB ramose or with branches 28 25. Flagellar joints moniliform 2G Flagellar joints cylindrical, not moniliform 27 26. Abdomen compressed, ascending Eucharis Latreille. Abdomen neither compressed nor ascending. First joint of tarsi much thickened Tricoryna K irby . First joint of tarsi very long, slender Metagea Kirliy. 27. Antennse lO-U-jointed. Petiole of abdomen abruptly enlarged at apex ; thorax smooth Pseudometagea Ashmead. Petiole of abdomen normal, long, cylindrical ; thorax rugose Psilogaster Itlanchard. Antenna' 13-jointed, rather short ; thorax closely pun(;tate, the parapsidal furrows distinct : right man- dible with two teeth within, the left with one tooth within Orasema Cameron. 28. Scutellum spined, bidcntate or produced into long processes that extend over the abdomen 2i» Scutellum normal, unarmed. Antenna' with 4 branches ; wings hyaline Pseudochalcura Aslniiea. 59. 1875. Tridyminse Tribus, (partim) Thomson, Hjnn. Skand. p|). 12 and 192. 1886. Tridyrainte, Subfamily, (partim) Howard, Ent. Amer., I., p. 198. 1899. Tridyminse, Subfamily II. , Ashmead, Proc. Ent. Soc. Washington, IV., p. 247. ashmead: classification of the chalcii) flies 273 This subfamily seems to l)e a natural group of gall-iiilialiitiiig species, allied to the Pireniiuv, but easil}'^ separated by the structure of the anteniue. Two tribes may be distinguished : table of tribes. Mesonotum with complete, distinct fiiiiows Tribe I. Tridymini. Mesouotuui with incomplete furrows, iiulitati'd only anteriorly Tribe IL Metasteuini Tkikk 1. Tndiinunl. The complete mesonotal furrows distinguish this tribe. Most of the species fall- ing in this tribe are parasitic upon gall-making or gall-inhabiting Diptera, belonging to the famil}' Cecidorui/ildn: J\jtnarn.s Mayr, however, is a genus living parasitic- ally upon fig-insects in Urazil, ami one or two exotic genera attack other gall- inhabiting insects. TABLE OF (iENERA. 1. Females 2 Males 15 2. Antenna; insertef Walk., the flagellum long and slender, the fuuicle joints all long; left mandible 3-, the right 4-(lentate ; metathorax not short, without a median carina, the spiracles large, oval ; abdomen large, depressed, very much longer than the head and thorax united Stylophorella Ashmead, g. nov. (type .V. perplexa Ashm.). 5. Pronotum not distinctly separated 6 Pronotum distinctly separated 8 276 MEMOIRS OF THE CARNEGIE MUSEUM G. Funiclo filiform, the first joint large. Marginal vein slender, always longer than the stigmal vein ; abdomen ovate 7 Marginal vein thickened, not longer than the stigmal vein. Body short ; clypeus bidentate ; mandibles 3-dentate Xenocrepis Fiirster (type denorrcpis areiiicola Thorns.). 7. Abdomen with a yellow band at base ; metathora.x short, wilh a median carina and lateral folds ; cly- peus separated, anteriorly arcuate Dimaclius Thomson (type Pteromalus discolor Walk.). Abdomen inlhout a yellow band at base ; metathorax not .short, without a median carina ; clypeus separated, smooth, with a median tooth anteriorly. . Hemitrichus Thomson (type H. rufipes Thoms.). 8. Flagcllum with the first joint short 9 Flagellum with the first joint long, cylindrical 10 9. Antennae .short, clavate in both sexes, inserted a little below the middle of the face ; clypeus at apex truncate ; metathorax rather short, without lateral folds, tlie spiracles rounded.. ..Habritus Thomson (type Pteromalus brevicornis Ratzeburg). Antennte filiform, inserted on the middle of the face ; clypeus at apex with a median incision ; meta- thorax not long, without a median carina, spiracles large, nearly linear Dinarmus Thomson (type D. acutus Thoms.). 10. AntennfE filiform, insei-ted on the middle of the face, the funicle joints all longer than thick ; clypeus anteriorly with a median sinus ; metathorax not short, without lateral folds, median carina, or spir- acular sulci, the spiracles oval Arthrolysis Fiirster (type Pteromalus scabriculiis Nees). 11. Antennfe 12-jointed, with tivo ring-joints 12 Antennte 13-jointed, with two ring-joints 13 12. Marginal vein not thickened, longer than the stigmal vein ; metanotum .short, smooth but with a dis- tinct median carina, the spiracles small Metastenus Walker. Marginal vein thickened, not longer than the stigmal vein ; metanotum jjunctate, with a median carina Disema Forster. 13. Pronotum not distinctly separated 14 Pronotum distinctly separated 16 14. Marginal vein slender, always longer than the stigmal vein 1.5 Marginal vein thickened, not longer than the stigmal vein. Clypeus bidentate ; mandibles 3-dontate Xenocrepis Fiirster. 15. Metathorax .short, %vith a median carina Dimachus Thomson. Metathorax not short, without a median carina. Hemitrichus Thomson. 16. Antennse inserted on the middle of the face, long 17 Antennie inserted somewhat below the middle of the face Habritus Thomson. 17. Metathorax without a median carina, the lateral UtMs prfsoil, the spiracles large, nearly linear. Dinarmus Thomson. Metathorax without a median carina, the lateral folds absent, the spiracles oval. Arthrolysis Foister. Subfamily III. MISCOGASTERINtE. 1833. MiscogasteridjB, Family IV (partim), Walker, Ent. Mag., I., p. 370. 1835. Ormoceridffi, Family (partim). Walker, Eiit. Mag., I., p. 167. 1835. Pteromalidaj, Family (partim). Walker, Eiit. Mag., II., p. 286. ASHMEAD : CLASSIFICATION OF THE ('HALCID FLIES 277 1856. Miscogasteroida\ Family XIV (partiin), F(>rster, Tlym. Stud., II., pi>. 10. 24 and 59. 1875. Pteromalina, Tribus (partim), Thomson, Hym. Skand., IV., pp. 12 and 21G. 1885. Pteromalinse, Sul>family (partim), Howard, Ent. Amcr., I., p. 198; II., 188(;, p. .S3. 1885. Miscliogastrides, Suhtril.us Thomson, Hym. Skand., IV., pp. Hi and 219. 1880. Miscliogastrides, Tribe, Howard, Ent. Amer., II., p. 33. This su))family, although distinct, has aflfinities which ally rt with the fiimilies Perilaiiipi(hc and the Eucharkhi', and also with the Ptcromalidir, through the sub- family Sphc.s is from North American, and super- ficially resembles rhilnchi/ra Haliday, a genus in the Eari/toiuidu'. table of gekera. 1. Females - Males '5 2. Autennse 14-jointed with 2 ring-joints '^ Antenna 12- or 13-jointed '' 3. Winged ■* Apterous. Abdonu'n conically pointed, the petiole distinct, a little longer than thick, the second segment occupying hardly half its surface ; llagellum rather stout, subclavate, the joints fluted. Apterolelaps Aslimead (type A. nigricejin Ashni.). 4. Abdomen conically produced at apex, and usually ending in a prominent ovipositor; second segment large, occupying fully half the surface, the third to fifth very short, the sixth and seventh together conical, longer than half the length of the second. Lelaps Haliday (type Merostenus sodales Walk.). 5. Abdomen conically produced at apex, the second segment not much longer than the third and fourth united, the fifth longer than the fourth, the seventh conically ].roduced ; scutellum with a cross-furrow at its apical third Neolelaps Aslnnead (type .Y. hauaiicnaix Ashm.). 280 MKMOIKS OF THE CARNEGIE MUSEUM Abdomen subglobose or short oval, the second segment very large, occupying nearly the whole surface, the following very short, more or less retracted within the second ; scutellum with a cross-fun-ow vei-y near its apex Mesolelaps Ashmead (type M. cyaneivenlris Ashm.). 6. Petiole of abdomen long, the body small, spatulate ; antenna; very long, 14-jointed, longer than the whole body, the joints long, cylindrical, clothed with long, sparse haire Lelaps Haliday. Petiole of abdomen very short, the body oblong-oval, truncate at apex; antenna' not longer than the thorax, 13-jointed, the flagellum filiform, pubescent, the joints after the first about twice as long as thick Mesolelaps Ashmead. Family LXVII. CLEONYMID/E. 1837. Cleonyiiiidie, Family (partiin), Walker, Ent. Mag., J\'., p. 349. 184(3. Eupelmicke, Family 9 (partim), Walker, List (/hulcid. Brit. .Museum, I., p. 52. 185G. Cleonymoida-, Fauiilie XIV.; (partim), Forster, Hym. Stud., II., pp. 19, 24 and 46. 1875. Cleonymides, Sulitribus, Thomson, Hym. Skand., IV., p. 217. 1878. Cleonyniides, Subtribus, Thomson, Hym. Skand., V,, p. 3. 1886. Cleonymides, Triljo, Howard, Entora. Amer., II., pp. 33, 34. 1899. (Jleonymidaj, Family LXVL, Ashmead, Proc. Ent. Soc. Washington, IV., p. 200. An historical sketch of this family was given in my paper entitled : " On the Genera of the Cleonymida)," published in 1899. Unquestionably, the family comes nearest to the family Eaetjrtkhv, and forms a connecting link between it and some families previously treated, i. e., the ChalcididsR, Eurytomidm and the Miscogastendx ; some genera in the subfamily Chalcedectinfe especially being remarkably like some genuine Clinlcidi(hr. jMany males, too, are easily mistaken for genuine EajKlmiues and Ejicyrtlnefy. Pdechulla has some char- acters similar to the Eurytomidse, and the Turymidx. Coleotrechnus, placed in this fixmily, is unknown to me. It is placed here from the description alone, and may be a genuine Encyrtid, although nothing is said of a salta- torial middle tibial spur. TABLE OF SUBFAMILIES. 1. Mesonotal furrows not at all indicated 4 Mesonotal furrows more or less distinct 2 2. Abdomen longly petiolated 3 Abdomen sessile, or subpetiolate, never longly petiolate. Po.sterior femora much swollen and usually toothed or finely denticulate beneath, as in Chaleis, Smicra, etc. ; abdomen usually depressed, the ovipositor rai-cly exserted ; autennse at the most II. jointed Subfamily I. Chalcedectin^. Posterior femora not much swollen and very rarely toothed beneath, the anterior femora usually more or less enlarged, sometimes very much swollen and often excised or dentate beneath toward apex ; ovipositor often, but not always, exserted ; antenna; Il-I3-jointed. Subfamilv 11. CLEONYMlNiE. ASHMEAU: CLASSIEICATIO.N OF Till': CIIAhClD FLIE8 281 3. Body of abdomen strongly compressed, sword-shai^ed, ending in a long ovipositor, the basal sheaths very broad ; all legs slender, the hind pair very long, their coxic long, cylindrical ; front wings with the marginal and postmarginal veins very long, the latter extending to tip of wing, the stigmal vein very small, subsessile ; antennse U-jointed, inserted close to the mouth. .Subfamily III. Pelecinellin.'E. 4. Abdomen subsessile, compressed, conically produced toward apex ; front wings with the postmarginal and stigmal veins very short, the kiiub of the latter nearly sessile, rounded ; posterior tibiic compres.sed, the hind margin denticulate Subfamily IV. CoLOTRECHNiNyK. Subfamily 1. Ciialcedectin.e. 1889. Polychrommintc, Subfamily, Ashmead, Proc. Ent. Soc. Washington, I., ]>. 22(5. 1895. Chalcedectinaj, Subfamily, Ashmead, Proc. Ent. Soc. Washington, III., p. 230. 1898. Chalcodectinaj, Subfamily, Dalla Torre, (.'at. Hym., V., p. 186. 1899. Chalcedectinae, Subfamily I., Ashmead, Proc. Ent. Soc. Washington, IV., p. 201. The sessile abdomen and the greatly swollen and usually dentate or serrate hind femora, as in the Chalcididie, distinguish the family. It is not represented in Europe and has reached its greatest development in South America and Australia, where the species are evidently numerous. 1 have also seen an undescribed species from Africa, where it should be well represented. table of genera. 1. Females 2 Males 7 2. Hind femora much swollen, minutely denticulate beneath .5 Hind femora much swollen and armed beneath with large, distinct teeth, or at least with one large tooth. Hind femora armed with 1 to 3 large teeth ;', Hind femora armed with several teeth. Abdomen depressed, the fourth dorsal segment the largest, transversely striate or aeieulate- Chalcedectes Walker (type ('. mactiliconiis Walker). 3. Hind femora with one large tooth, with three or foui- smaller teeth beyond 4 Hind femora with three large teeth Cleptimorpha Walker (type C. binolata Walker) 4. Abdomen oblong-oval, subcompressed beneath, nearly baie, and ending in a long ovipositor ; head and thorax coarsely reticulately punctate ; flagelliun lung, slender, iililorm Chalcidiscelis Ashmead (type C. ko.-hclei Ashm.). Abdomen shorter than the thorax, compressed, the ovipositor luddeu ; mcsouotum nearly smooth, or transversely striate or wrinkled ; antenn;e 12-joiuted, the funicle 6-jointed, the joints transverse. Systolomorpha Ashmead (type S. thyridopterygls Ashm.). 5. Antennas 1 1-jointcd (> Antenn;e 10-jointcd Amotura Cameron (type A. unniilininiis Ciini.). 6. Abdomen ovate or conic-ovate. Abdomen not depressed, convex heiieatli, the fotiith dorsal segment longer than tlie third, the latter a little longer than the lil'tli ; melathoraeie spiracles reniform. Agamerion Ilaliday ' (type Miscof/aslcr gejo Walk.). Abdomen depressed. Hat above and beneath, llie third and fourth doreal segments short, united shorter than the liftli ; stigmal vein rather short : metathoracic spiracles oval. Euchrysia Westwood (type E. cleplidea Westw.). ' Kirby has incorrectly classified this f,'emis witli the ('!iiihiiliii:r. 282 MKMOrKS OK THE CARNF/ilE MUSEUM 7. Hind femora much swollen, minutely denticulate beneath 10 Hind femora much swollen and armed beneath with large, distinct teeth or at least with one large tooth. Hind femora armed with 1 to 3 large teeth 8 Hind femora armed with several teeth Chalcedectes Walker. 8. Hind femora with one large tooth 9 Hind femora with three large teeth Cleptimorpha Walker. 9. Head and thorax coarsely, reticulately punctate Chalcidiscelis Ashmead. Head and thorax not coarsely reticulately punctate, the niesonotum almost smooth. Systolomorpha Ashmead. 10. Antenna' 11-jointed 11 Antennaj 10-joiuted Amotura Cameron. 11. Metathoracic spiracles reniform Agamerion Haliday. Metathoracic spiracles oval Euchrysia Westw ood. Subfamily II. Clennymin;e. 1897. Cleonyminie, Subfamily II, Ashmead, Proc. Ent. Soc. Washington, iv, pp. 201, 202. To this group belong the better known genera of the family, namely Cleonipnus, Chdropachys, TrigunoJerus, Acroronnu.% etc., genei-a of world-wide distribution. It is the largest group of the family and includes many genera and species. Most of the species appear to attack only coleopterous larv;e and are of great economic importance, since they destroy beetles destructive to our fruit, shade and forest trees. TABLE OF GENERA. 1. Females 2 Males 23 2. Anterior femora more or less distinctly swollen, and never excised dentate beneath 18 Anterior femora iiuieb swollen, or e.vcised dentate beneath • 3 3. Pronotum not inueli narrowed and always wider than long 4 Pronotum nuuh lengthened and narrowed, longer than wide. Front femora greatly swollen, but not excised dentate beneatli ; abdunieii conieally produced, but not much longer than the head and thorax united. Heydenia Fiirster (type H. pretiosa Fiirster). Front femora much swollen, and excised dentate beneath ; abdomen elongate, conieally produced. Lycisca Spinola (type L. rciptoria Spinula). 4. Eyes bare 1 Eyes pubescent. Ovipositor very long 6 Ovipositor not prominent, at the most subexserted. Abdomen subrotund, oblong or conic-ovate, the sides rounded, not carinated 5 Abdomen conic-ovate, or conieally lengthened, the sides distinctly carinated. Epistenia Westwood (type E. ca-ndea Westw.). ASHMEAD : CLASSIFICATION OF Till', CIIA 1,(11) FLTKS 283 5. Labrum inconspicuous oi- hidden 7 Labrum conspicuous. Abdomen conic-ovate Cleonymus Latreillc (type Ichneumon depresxtis Fabr ). Abdomen sul>r()tnn 12. Marginal vein not thickened 18 Marginal vein thickened, shorter than the i)ostmarginal. Zapachia Fiirster (tyiie X. apilaplcni Filrst.). 1.3. Abdomen ovi'.l, hardly as long as the thorax, above dejiressed 14 Abdomen conical, longer than the thorax, the segments after the first, which is the longest, subequal. Front wings with one ti-ansverse band, the stigmal club large Acrocormus Fiirster (type A. semifasciatus Thorns.). Front wings without a band, the stigmal cluli small. Caudonia Walker (type C. agijlla Walk.). 14. Front wings with a large, broad fuscous band beneath, the marginal and stigmal veins having a tri- angular hyaline streak within from the marginal vein ; .stigmal vein long, subclavate, as long as the marginal : fiuiicle joints 3 to fi a little wider than long Brachycaudonia .V.shmead g. nov. (tvpe />'. callfnrnica Ashni.), 284 MEMOIRS OF THE CARNEGIE MUSEUM l"). Abdomen ovate, shorter tliaii the thorax, the segments subequal : midille tarsi incrassate. Notanisus Walker (type jY. versicolor Walk.). IG. Abdomen .sessile 17 Abdomen petiolate 22 17. Abdomen elongate, conieally produced or acuminate at apex ; postmarginal vein well developed. 18 Abdomen long, conic-ovate, ending in a prominent ovipositor, which is dilated into three broad, leaf- like expansions, like a propeller in a naphtha launch. Dinoura Ashmead (type D. auriventris Ashm.). 18. Hcutellum without a transverse grooved line before tip ; head triangular, narrowed anteriorly 21 Scutellum with a transverse grooved line before the tip ; head with rounded, convex cheeks. Pronotum transverse, not large 19 Pronotum large, almost quadrate Merostenus Walker (type M. phedyma Walk.). 19. Pronotum very short, visible from above as a fine transverse line 20 Pronotum transverse Trigonoderus Westwood (type T. prince})s Westw.). 20. Antenna} 13-joiuted ; middle tibite normal ; metanotum with a sharp median carina. Anoglyphis Forster (type A. nubilosa Fcirst.). Antennse 14-jointed ; middle tibite much lengthened. Macromesus Walker (type M. amphireius Walk.). 21. Middle tibia; not dilated at apex Platygerrhus Thomson (type P. gracilis Thorns.). Middle tibi;e dilated at apex, their tarsi broad at base Pegopus Forster (type Prosopon montanus Walk.). 22. Scutellum without a transverse grooved line before tip Photismus Thomson (type P. nubilosus Thorns.). 23. Front femora not, or less distinctly, swollen, and never excised dentate beneath 28 Front femora much swollen, and sometimes excised dentate beneath. Pronotum not much narrowed and always wider than long 24 Pronotum much narrowed and lengthened. Front femora much swollen but not excised dentate beueatli ; abdomen clavate. de- pressed Heydenia Forster. Front femora swollen and excised dentate beneath. Lycisca Spinola. 24. Eyes bare 27 Eyes pubescent. Abdomen not carinate along the sides 25 Abdomen carinate along the sides Epistenia Westwood. 25. Labrum inconspicuous or hidden 26 Labrum conspicuous. Metathorax with a median carina, the spiracles large, oblong or oval ; (Ingellum subclavate, densely hairy, the joints of the funicle wider than long Cleonymus Latreille. Metathorax without a median carina, the spiracles small, rounded ; flagellum long, filiform, densely hairy, the joints of the funicle long Micradelus Walker. 2fi. Front wings bifasciate or maculate ; marginal vein slender, about twice as long as the stigmal ; pro- notum not short ; metanotum with a median carina ; head lenticular, much wider than the thorax. Ptmobius Ashmead. 27. Front wings bifasciate ; marginal vein not longer than the stigmal, the latter long, strongly clavate ; pronotum short, transverse, slightly narrowed ; metanotum with a short median carina, the spiracles elliptic ; head transverse ; pedicel shorter than the first joint of the funicle. Cheiropachys Westwood. Front wings hyaline, not fasciate ; marginal vein longer than the stigmal, the latter not especially long, ending in a small knob: pronoUmi large, (juadrate. well-sei)arated Schizonotus Hatzeberg. ASIIMKAB : t'LASSIFICATroN OF THE CITALCin FI.TKS 285 28. Mesouotuni with the furrows complete 30 Mesouotum with incomplete furrows, iudicated only anteriorly. Marginal vein not thickened 29 Marginal vein thickened, shorter than the postnirtrgina] ZapacMa Fiirster. 29. Wing.s with a transverse band or fascia. Stigmal club very large Acrocormus Forster. Stigmal club small Brachycaudonia Ashmead. Wings hyaline, without a fascia. Stigmal club small Caudonia Walker. 30. Abdomen petiolate 34 Abdomen sessile. Scutelhun irilli a transverse grooved line before apex 31 Scutelhim irithind a tran.sverse grooved line before apex 32 31. Pronotum very sliort, visible from above as a fine transverse line 32 Pronotum not very short. Pronotum quadrate Merostenus Walker. Pronotum transverse Trigonoderus Westwood. 32. Metanotum not short, smooth and with a sharp median carina : antenna" 13-jointeil. Anoglyphis Filrster. Metanotum very .short, closely punctate ; antenna- U-.jointed Macromesus Walker. 32. Postmarginal vein well developed ^'5 Postmarginal vein scarcely longer than the short stigmal ve'ii with its knob ; scutclUuu with two par- allel dorsal grooved lines ; metanotum with a median carina : antenna' Dinoura Ashmead. 33. Middle tibi:e not dilated at apex Platygerrhus 'Ibomson. Middle tilme dilated at apex, the tarsi much thickened at base Pegopus Forster. 34. Scutellum without a transverse grooved line before the tip Photismus Thomson. hiUBFAMILY III. PFLKCINELLINiK. 1897. PelecinellinfP, Subfamily III., Ashmead, Proc. Ent. Soc.Washington,IV.,p. 201. This subfamily is at present represented by a single genus, Pelecinella Westwood. Three species have been described, all from Brazil, and these are the largest and in some respects the most striking looking ( 'halcid-Hies known. The affinities of the group were discussed in my paper, '• ( )ii tlie ( icnus Peleci- nella Westwood," published in 1897. Form very elongate ; pronotum verj' long, longer and narrower than the mesonotum, contracted in front ; head with a deep frontal furrow; abdomen long, compi-essed, lanceolate, ending in a long ovipositor, and lougly petiolate ; hind legs long, the coxa" long and cylindrical, the tibisB longer than the femora, very gradually widened toward apex and terminating in two spurs, the tai-si slender, as long as the feraoi-a, the first joint longer than Joints 2 oiniited ; stigmal vein very short, the marginal vein long, the postmarginal longer than the marginal Pelecinella Westwood (type Ji. phantasma Wcstw.). Sur.FAMir.V 1\'. CoLOTRECHNIN.i;. 1875. Colotrochnidos, Subtribus. Tliom.son, Ilym. 8kand., IV., p. 217. 1878. ColotrechnidL's, Sul)tril.u.s, Thomson, llym. Skand., \'., p. 40. 286 MEMOIRS OF THE CARXEGIE MUSEUM 1886. Colotrechiiides, Tribe, Howard, Ent. Amer., II., pp. 33 and 34. 1897. Colotreclmiiiie, Sul.faniily IV , Ashmcad. Proc. Ent. Soc. Washington, IV., p. 201. This group is unknown to me in nature. It is based upon Thomson's genus Colotrechnus, occurring in Sweden, who called it a subtribe. Thomson's description clearly indicates that it belongs to the family Cleonipnidic, although the absence of mesonotal furrows is strongly suggestive of the Encijriidir. Wings with the stigmal and postinarginal veins very sliort, the knob of the former rounded, subses.sile ; frontal depression long ; antennse 12-jointed, inserted below the middle of the face, the funicle .5-jointed ; hind tibise compressed, the bind margin denticulate . . Colotreclinus Thomson (type C. subenerulens Thorns. ). Family LXVIII. ENCYRTID.E. 1837. Encyrtidfe, Family (partim), Walker, Ent. Mag., IV., p. 430. 1840. Encyrtidfe, Subfamily 4, Westwood, Intro. Mod. Classif Ins., IF., p. 166; Synop., p. 66. 1846. Eupelmidfc, Family 9 (partim). Walker, List Chalc. Brit. Museum, I., p. 52. 1846. Encyrtidfe, Family (partim), Walkei-, List Chalc. Brit. Museum, I., p. 53. ,1848. PteromaIida3, Family V. (partim), Walker, List Chalc. Brit. Museum, II., p. 104. 1856. Eupelmoida?, Familie IV. (partim), Fiirster, Hym. Stud., II., pp. 18, 21 and 30. 1856. Encyrtoidaj, Famihe V. (partim), FGrster, Hym. Stud., II., pp. 18, 21 and 32. 1875. Encyrtina, Subtribus (partim), Thomson, Hym. Skand., IV., pp. 12 and 112. 1885. Encyrtinte, Subfamily (partim), Howard, Ent. Amer., I., pp. 198, 216. 1897. Encyrtida?, Family LXVIII., Ashmead, Proc. Ent. Soc. Washington, IV., p. 236. 1900. Encyrtidie, Family LXVIII., Ashmead, Proc. U. S. Nat. Museum, XXIL, p. 202. This family was very fully characterized and discussed in my paper entitled "On the Genera of the Subfamily Encyrtime," published in the Proceedings of the U. S. National Museum for 1900. It is unnecessary, therefore, to repeat the charac- terization of the group, since the classification here is practically identical, the only change being the establishment of a new tribe in the Eupclmiinr, and some new genera in the various tribes. TABLE OF SUBPAMILIE8. 1. Mesonotnm entire, convex or subconvex, the parapsidal furrows entirely absent 2 Mesonotum not entire, usually depressed oi- impressed, rarely convex, the parapsidal farrows distinct or at least more or less distinct, never entirely wanting ; marginal vein usually long. Subfamily I. Eupelmin.^. ASHMKAD : CLASSIFICATION OK THE CUALCID FLIES 287 2. Marginal vein rarely very long, ofttMi ii\iii(til'(inii. aiul always much sliorter than the submurginal or subcostal vein; stigmal vein usually sliort, laicly Imiu- ; scutel hi in never short or transversely linear ; middle tibiro without lateral spurs Subfamily II. Encyrtin.e. Marginal vein long, as long as the submarginal or subcostal vein ; scutellum very short, transversely linear ; middle tibia; with lateral spurs, the apical spur lobed Subfamily III. Signiphokin^. Subfamily I. Eui'Elmin.e. 1846. Eupelmidte, Family (pcartim), Walker, Ann. & Mag. Nat. Hist., XVI 1., p. 114. 1856. Eupelmoidiu, Familie 4, Forstor, Hym. Stud., II., pp. 18, 21 and 30. 1875. Eupelmina, Trihus, Thomson, Hym. Skand., IV., })]>. 11 and 102. 1886. Eupelminpe, Subfamily, Howard, Ent. Amer., I., p. 198. 1897. Eupelminfe, Subfamily 1., Aslimoad, J*roc. Ent. Soc. Washington, IV., p. 238. This subfamily is quite distinct from tlic other two subfamilies, an