581.526325 
 
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 Naturcl ^?^i5cry Survey 
 Libtary 
 
 BIOLOGICAL REPORT 88(26.4) 
 MAY 1988 
 
 
 NATIONAL LIST OF PLANT SPECIES 
 
 THAT OCCUR IN WETLANDS: 
 
 NORTH PLAINS (REGION 4) 
 
 Fish and Wildlife Service 
 
 In Cooperation with the National and 
 Regional Interagency Review Panels 
 
 U.S. Department of the Interior 
 
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Biological Report 88 (26.4) 
 
 May 1988 
 
 Supersedes WELUT-86/W13.04 
 
 NATIONAL LIST OF PLANT SPECIES 
 THAT OCCUR IN WETLANDS: 
 NORTH PLAINS (REGION 4) 
 
 by 
 
 Porter B. Reed, Jr. 
 
 National Ecology Research Center 
 
 U.S. Fish and Wildlife Service 
 
 Suite 101, Monroe Building 
 
 9720 Executive Center Drive 
 
 St. Petersburg, FL 33702 
 
 for 
 
 National Wetlands Inventory 
 U.S. Fish and Wildlife Service 
 
 In cooperation with the National and Regional 
 Interagency Review Panels: 
 
 U.S. Fish and Wildlife Service 
 
 U.S. Army Corps of Engineers 
 
 U.S. Environmental Protection Agency 
 
 o.S. Soil Conservation Service 
 
 U.S. Department of the Interior 
 
 Fish and Wildlife Service 
 
 Research and Development 
 
 Washington, DC 20240 
 
DEDICATION 
 
 The National List of Plant Species That Occur i_n Wetlands is respectfully 
 dedicated to Neil Hotchkiss and Francis M. Uhler, who made this work possible 
 by their contributions to wetland botany and ecology. These two scientists 
 focused their long and productive careers on the study of wetlands long before 
 our current emphasis on wetland systems began. They aided in producing the Fish 
 and Wildlife Service's first wetland classification system, developed initial 
 lists and guidebooks to the wetland plants of the United States, and produced 
 many studies on the ecology and management of wetlands and their value to 
 wildlife. It is my sincere pleasure and humble gratitude to acknowledge the 
 dependence of this current work on their long and untiring efforts. 
 
 Suggested citation: 
 
 Reed, P.B., Jr. 1988. National list of plant species that occur in wetlands 
 
 North Plains (Region 4). U.S. Fish Wildl. Serv. Biol. Rep. 88(26.4). 64 pp. 
 

 CONTENTS 
 
 Page 
 
 ACKNOWLEDGMENTS i v 
 
 INTRODUCTION 1 
 
 DEVELOPMENT OF THE CURRENT LIST 2 
 
 REVIEW PANELS AND REVIEW PROCESS 4 
 
 COMPOSITION OF THE LISTS 7 
 
 DIGITAL DATA BASE AVAILABILITY 13 
 
 REGIONAL INDICATORS 14 
 
 TRINOMIALS 38 
 
 SYNONYMY 40 
 
 LITERATURE CITED 57 
 
 APPENDIXES 
 
 A. Review Procedure 58 
 
 B. Instructions for Completing the Review Sheet for 
 
 Plant Species That Occur in Wetlands 59 
 
 C. Review Sheet for Plant Species That Occur in 
 
 Wetl ands 60 
 
 D. Interagency Review Panel Members and Regional 
 
 Reviewers 61 
 
 11 1 
 
ACKNOWLEDGMENTS 
 
 The National List of Plant Species That Occur in Wetlands (hereafter referred 
 to as the National List ) is the result of the collective efforts of a large 
 number of dedicated biologists. Special recognition and thanks are extended to 
 the many people who contributed to the successful completion of this project. 
 The National Wetlands Inventory, especially John Montanari and William Wilen, 
 provided the funding and administrative support necessary for the development 
 of the National List and coordination of the review effort. 
 
 The other Federal agencies, especially William Sipple, John Meagher, and Dave 
 Davis of the Environmental Protection Agency, Robert Pierce, Richard Macomber, 
 and Dana Sanders of the Corps of Engineers, and Billy Teels and Carl Thomas of 
 the Soil Conservation Service, provided much needed funding to aid development 
 of the Annotated National Wetland Plant Species Data Base and technical support 
 through their agency biologists' participation in the review effort. 
 
 Special recognition and appreciation is extended to Lewis Cowardin who conceived 
 the need for the National List , supervised the compilation of the initial draft 
 list, and provided invaluable advice and direction during the initial phase of 
 the project. 
 
 The contribution of the authors of the almost 300 regional and State floras and 
 regional wetland manuals used in compiling the Annotated National Wetland Plant 
 Species Data Base is gratefully acknowledged. 
 
 The development of the National List was greatly facilitated by the dedicated 
 staff who contributed over 50 person years to compile the Annotated National 
 Wetland Plant Species Data Base. This data base provided a solid information 
 base from which the National List was derived. This staff included Karen E. 
 Amidei, James G. Armstrong, Sheryl A. Brenner, Steven I. Candileri, Mark A. 
 Charneski, Diana Fry, Thomas B. Gunter, Lillian A. Gustafson, Iris A. Kendall, 
 Mary E. Klee, Annie L. Kosvanec, David R. Lindsey, Stephen Mortellaro, Kent A. 
 Moyer, Laura E. Pittman, Donald R. Richardson, Richard N. Rowse, Angela F. Salem, 
 Deana Ulmer, Sheri A. Ulrich, Sandra M. Upchurch, Diane Wallace, Debora L. 
 Wegner, August M. Wooten and Kevin R. Youngberg. This landmark effort to collect 
 much of the taxonomic and ecological information about wetland plants into a 
 textual computer data base was a protracted and extremely tedious task. It was 
 truly a pleasure to work with such a productive and conscientious group. 
 
 A special debt is owed to the regional ecologists who so generously gave of their 
 time and experience in reviewing the lists. Their review helped to refine the 
 information presented in the botanical manuals and in many cases provided the 
 only and often best description of the ecology of many plant species. The 
 
 iv 
 
Nationa l List would not be as accurate and complete if it were not for the 
 enormous amount of review so generously provided by the regional reviewers. 
 State distributions and common names provided by John Kartesz from unpublished 
 data bases allowed the production of accurate State lists and common name 
 assignments for almost all species. The high quality and completeness of the 
 Nationa l List is in large part due to the data provided by John Kartesz from the 
 Biota of North America Program. 
 
 The composition and pleasing format of the lists is largely due to the computer 
 skills of Jill Muhlenbruck and Gregor Auble of the National Ecology Research 
 Center. 
 
 The contributions of the many National and regional review panel members is 
 gratefully acknowledged. These biologists represented their agencies in a 
 technically competent and thoroughly professional manner. The strength of the 
 Regional lists and Indicator assignments is largely due to the diverse background 
 of the review panel members and a firm desire by all to make the process work. 
 The unswerving resolve and determination displayed by all the review panels to 
 complete each phase of the task during the marathon regional review panel 
 meetings was truly inspiring. Well over 200 weeks of staff time was contributed 
 by the review panel members. 
 
 It has been a pleasure to coordinate this effort and to work so closely with so 
 many dedicated and professional biologists. The credit and recognition for the 
 completion of this task applies equally to all who contributed so much. As 
 compiler, I must necessarily take the responsibility for any transcription errors 
 which may have occurred during the production of this report. The extensive use 
 of the National List has made the entire effort professionally satisfying for 
 all who have contributed. 
 
Digitized by the Internet Archive 
 
 in 2013 
 
 http://archive.org/details/nationallistofpl04reed 
 
INTRODUCTION 
 
 The National List of Plant Species That Occur in Wetlands (hereafter referred 
 to as the National List ) represents the combined efforts of many biologists over 
 the last decade to define the wetland flora of the United States. The National 
 List has undergone a number of revisions based on intensive review by regional 
 ecologists. National, regional, and State lists are being distributed to provide 
 users with the most current information. We welcome and encourage modification 
 and improvement of the National List . Refinement of the National List will occur 
 continually, reflecting increased knowledge in Indicator assignments, taxonomy, 
 and geographic distribution. We anticipate that further refinement of the 
 National List will lead to additional infra-specific and subregional Indicator 
 assignments. Review documents and procedures are included with the National List 
 to aid and encourage additional review (Appendix A). The U.S. Fish and Wildlife 
 Service initially developed the National List in order to provide an appendix 
 to the Classification of Wetlands and Deepwater Habitats of the United States 
 (Cowardin et al . 1979) to assist in the field identification of wetlands. Plant 
 species that occur in wetlands as used in the National List are defined as 
 species that have demonstrated an ability (presumably because of morphological 
 and/or physiological adaptations and/or reproductive strategies) to achieve 
 maturity and reproduce in an environment where all or portions of the soil within 
 the root zone become, periodically or continuously, saturated or inundated during 
 the growing season (adapted from Huffman 1981). The development of the National 
 List changed significantly when a cooperative review effort was established by 
 the major Federal agencies involved in wetland identification and management. 
 The utility of the National List goes far beyond a simple catalog of wetland 
 plants. The Fish and Wildlife Service, in cooperation with North Carolina State 
 University, has produced a weighted average procedure for using the wetland 
 Indicator assignments of individual species to assist in determining the 
 probability that a community is a wetland (Wentworth and Johnson 1986). This 
 procedure is used by the Soil Conservation Service to aid in the determination 
 of wetlands included under the conservation provisions of the Food Security Act 
 of 1985. The Fish and Wildlife Service, Army Corps of Engineers, Environmental 
 Protection Agency, and Soil Conservation Service use the National List to aid 
 in identifying wetlands falling under their various wetland program 
 responsibilities. Wetland identification manuals which incorporate the National 
 List have recently been produced by the Corps of Engineers (Environmental 
 Laboratory 1987) and the U.S. Environmental Protection Agency (Sipple 1988). 
 
DEVELOPMENT OF CURRENT LIST 
 
 The Fish and Wildlife Service recognized that accessory lists of hydrophytes 
 (plant species that occur in wetlands) and hydric soils would need to be 
 developed to apply the wetland classification system accurately and consistently 
 in the field. The scientific names of the plant species included in the major 
 wetland plant lists and manuals were collected and merged into a single 
 computerized list with those species on the National List of Scientific Plant 
 Names (U.S. Department of Agriculture 1971) that had emergent, floating, or 
 submergent life forms. This initial list of 1,626 species, completed in March 
 1976, was obviously incomplete, and was especially deficient in plant species 
 from the western United States and the Alaska, Caribbean, and Hawaii regions. 
 Dr. Donovan Correll, Fairchild Tropical Garden, Miami, Florida, reviewed this 
 initial list in 1977 and suggested many additional species for inclusion. Dr. 
 Correll 's additions were combined with the initial list, and a draft list of 
 4,235 species was developed in 1977. This draft list, although plagued by 
 problems of plant nomenclature and synonymy, was remarkably complete, considering 
 the small amount of time which had been spent on its development. 
 
 Review and refinement of this draft list has continued since 1977. Initial tasks 
 were to maintain and improve computer storage and retrieval of the draft list 
 information, align the listed species with a national taxonomic treatment, and 
 subdivide the species according to their fidelity to wetlands. The importance 
 of the development of an accurate National List of Plant Species That Occur in 
 Wetlands to the Federal community and the need to substantiate the occurrence 
 of these plant species in wetlands from the botanical literature led the Fish 
 and Wildlife Service to begin development of the Annotated National Wetland Plant 
 Species Data Base. This textual data base documents the taxonomy, distribution, 
 and ecology of each species based on a synthesis of almost 300 National and 
 regional wetland plant and botanical manuals representing the major State and 
 regional floras. Computer storage of the Annotated National Wetland Plant 
 Species Data Base allowed for the efficient maintenance of the initial National 
 List and creation of early draft regional subdivisions of the National List . 
 Data collection for the Annotated National Wetland Plant Species Data Base for 
 all plant species was completed in 1987, but incorporation of this information 
 into a single data base on the National Wetlands Inventory minicomputer remains 
 to be accomplished. The species data base is presently stored on a 
 microcomputer. Information on the data base and preliminary species listings 
 or data reports from the Annotated National Wetland Plant Species Data Base can 
 be obtained from the National Wetlands Inventory, St. Petersburg, Florida. 
 
 The Soil Conservation Service, through a contract to the Smithsonian Institution, 
 produced a revised National List of Scientific Plant Names in 1982 (U.S. 
 Department of Agriculture 1982.) This national treatment provided a standard 
 nomenclature for the National List of Plant Species That Occur j_n Wetlands , 
 
supplied a listing of synonyms linked to the accepted names, and updated the 
 regional distribution of each species. The Soil Conservation Service list was 
 selected as the taxonomic standard in order to facilitate the eventual 
 correlation of the National List of Plant Species That Occur in Wetlands with 
 the Hvdric Soils of the United States (U.S. Department of Agriculture 1987). 
 Copies of the National List of Scientific Plant Names (1982) are available from 
 the State offices of the Soil Conservation Service. 
 
 A wetland fidelity rating system was created during the initial development of 
 the Annotated National Wetland Plant Species Data Bas??. Early coding of verbatim 
 habitat from the botanical manuals for a wide variety of plant species indicated 
 that an obvious separation of obligate (restricted to wetlands) and facultative 
 (not restricted to wetlands) species could be made. Further refinement led to 
 subdivision of the facultative category into three subcategories, with a range 
 of percent occurrences in wetland versus nonwetland applied to each subcategory 
 to enhance user understanding and consistent application. 
 
 The ecological information obtained from the botanical manuals during data 
 collection for the development of the Annotated National Wetland Plant Species 
 Data Base led to the identification and addition to the National List of many 
 additional species for which at least one manual reported the species occurring 
 in an obvious wetland site. The National List had increased as a result of this 
 process to 5,244 species in 1982, 6,042 species in 1986, and presently is 
 composed of 6,728 plant species. 
 
REVIEW PANELS AND REVIEW PROCESS 
 
 The desire of the Federal agencies involved in wetland identification and 
 delineation for a Federal list of plant species that Occur in Wetlands led to 
 the suggestion by the Fish and Wildlife Service that a review process be 
 established similar to that developed to review the Hvdric Soils of the United 
 States (U.S. Department of Agriculture 1987). In early 1983, the Fish and 
 Wildlife Service formally requested that the Army Corps of Engineers, 
 Environmental Protection Agency, and Soil Conservation Service participate 
 cooperatively in an interagency review and development of a National List of 
 Plant Species that Occur in Wetlands. Each agency nominated staff wetland 
 ecologists with a strong background in wetland botany to a National Interagency 
 Review Panel. Two organizational meetings were held in the summer of 1983 to 
 determine the responsibilities and goals of the National and Regional Interagency 
 Review Panels and the structure of the review process. The four Federal agencies 
 also nominated staff wetland ecologists to represent them on each of the Regional 
 Review Panels. Selection and appointment of the Regional Interagency Review 
 Panel representatives was completed by each agency by the spring of 1984. All 
 four agencies have been represented on most Review Panels, with some change in 
 agency representatives occurring through the years. 
 
 Initial organizational meetings for all the conterminous United States Regional 
 Interagency Review Panels were held in 1984. The regional review process was 
 discussed, and review materials were developed. Potential reviewers, principally 
 field botanists and ecologists associated with State and Federal agencies and 
 universities, were identified for each region, and the responsibility for 
 contacting the potential regional reviewers was partitioned among the Review 
 Panel members. Regional reviewers were contacted during the summer and fall of 
 1984 to determine if they could review the list and return their review comments 
 by the winter of 1984-85. Regional reviewers were sent the most current copy 
 of the regional list (dated September 1982) during the summer and fall of 1984. 
 The Regional review lists contained, for many species, a tentative Indicator 
 assignment developed from the data collected for the Annotated National Wetland 
 Plant Species Data Base. All the Regional Review Panels met during the spring 
 of 1985 to consider the review provided by the regional ecologists. A total of 
 142 ecologists and botanists from across the country initially reviewed the 1982 
 lists. The number of reviewers varied from 10 to 30 per region. Regional 
 reviewers assigned a wetland Indicator to as many species as possible, based upon 
 their field experience, using Cowardin et al . (1979) for the definition of a 
 wetland. 
 
 The Regional Interagency Review Panels examined the Indicator assignments 
 suggested by each reviewer and any additional supporting documentation which 
 reviewers provided. Each of the Regional Review Panel members independently 
 synthesized the review received and developed a regional Indicator assignment 
 
for each species based upon all the regional review and information gathered 
 about the species. The tentative Indicator assignment developed from the 
 Annotated National Wetland Plant Species Data Base often was regarded as the 
 equivalent of a regional reviewer's assignment and was given the same status or 
 weight in Review Panel deliberations. The Regional Review Panel collectively 
 considered Indicator assignments for each species made by each agency panel 
 member and, with each agency having one vote, attempted to achieve unanimous 
 agreement on a Regional Indicator assignment. The plus (+) and minus (-) 
 designations, specifying respectively the higher or lower part of the frequency 
 range for a particular Indicator, were used by some Review Panels as a means of 
 achieving interagency agreement. The number of reviewers for each species varied 
 considerably, and each reviewer was generally given the same weight by a Review 
 Panel. The number of reviewers commenting on individual species varied, 
 depending on the distribution of the species across the region. Particular 
 species ranged from 20 review comments to only a single review, and in some cases 
 received no review. Review reflecting a wide range of suggested Indicators was 
 received for some species. This broad range of suggested Indicators for these 
 species was difficult to synthesize and blend into a single Indicator status. 
 These species were given an NA (no agreement) assignment if the Review Panel 
 could not reach a unanimous decision. Unreviewed species were assigned an NI 
 (no indicator) assignment if the Review Panel had little or no information on 
 which to base an Indicator status. 
 
 The National Review Panel met in July 1985 to review progress, to examine the 
 Indicator assignments for consistency across regions, and to develop a procedure 
 for the assignment of an Indicator to as many unassigned species as possible. 
 National, regional, and State lists of plant species that Occur in Wetlands were 
 produced in the spring of 1986, and were distributed widely. 
 
 The Regional Review Panels met during the summer and fall of 1986 to apply an 
 Indicator assignment to as many remaining unassigned species as possible. The 
 Review Panels principally relied on additional review received from former and 
 new reviewers, the habitat information recorded in the Annotated National Wetland 
 Plant Species Data Base, or examination of the habitat given in selected regional 
 manuals if the Species Data Base information was not available, to assign a 
 regional Indicator status. An asterisk (*) was assigned by the Regional Review 
 Panels to Indicators derived from limited ecological information. The asterisk 
 reflected a tentative assignment made with less confidence and data than the 
 other Indicator assignments. Usually no review was received from regional 
 ecologists for these asterisked species. A question mark (?) following a 
 National Indicator denoted a tentative Indicator assignment assigned by the 
 compiler and not confirmed by Regional Interagency Review Panel concurrence. 
 The 1986 National Wetland Plant List - Regional Indicator Compilation (Reed 1986) 
 reported the status of the review process. 
 
 The National Review Panel reexamined the review process and current list in 1986 
 and directed the Regional Panels to complete the review of the regional lists. 
 The Regional Review Panels met during the spring and summer of 1987 to complete 
 the initial assignment of as many unassigned species remaining on the regional 
 lists as possible. The Review Panels also considered species suggested for 
 addition by reviewers. Additional Regional Indicator assignments and changes 
 to previous assignments were made based on new review received from reviewers, 
 
ecological information from the Annotated National Wetland Plant Species Data 
 Base, or information from botanical manuals. 
 
 The task of the Regional Interagency Review Panels was to interpret and 
 synthesize reviewers' comments and the range of habitat descriptions given for 
 each species by the various authors of the botanical manuals into a single 
 wetland Indicator category for their regions. There was an overwhelming 
 similarity of independent Indicator assignments made by both the reviewers, based 
 on their field experience, and the Regional Review Panel members, based on the 
 habitat expressed in the botanical literature. This repeatability of Indicator 
 assignments derived in a variety of ways by ecologists with a wide variety of 
 backgrounds confirmed that the Indicators were both reproducible and defensible. 
 The Regional Review Panels were able to assign, with the highest degree of 
 confidence. Indicators to those species which had been reviewed by a number of 
 regional reviewers and also had a large literature base. The comparability of 
 the Indicator designations is also demonstrated by the large number of species 
 (6,114 species, 91% of the species assigned an Indicator) that were assigned only 
 a single Indicator or a narrow range of Indicators by the independently 
 functioning Regional Review Panels. An analysis of the National Indicator range 
 shows that 483 species (7%) were assigned an NI (no indicator), and 675 species 
 (10%) have a question mark following the Indicator. The question mark signifies 
 a tentative assignment. An analysis of the Regional Indicators for all regions 
 shows that an * was added to 729 Regional Indicators and an NA was assigned to 
 28 species. The National List of Plant Species that Occur in Wetlands represents 
 our progress to date and is provided both for current use and as a base for 
 future revisions. 
 
COMPOSITION OF THE LISTS 
 
 The National, regional, and State lists are reported in a variety of formats. 
 All of the lists are initially arranged alphabetically by scientific name 
 followed by a second list, also alphabetical by scientific name, of those species 
 with infra-specific (subspecies, variety, or form) Indicator assignments. A 
 third list, alphabetic by scientific name, provides a list of synonyms related 
 to the equivalent accepted name shown in the alphabetical scientific name list. 
 The State lists also contain a fourth compilation, following the synonymy report, 
 of the species found in each State, alphabetized by common name and subdivided 
 into eight separate lists by life form (trees, shrubs, vines, forbs, grasses, 
 grasslikes [sedges and rushes], ferns and allies, and aquatics). 
 
 Nomenclature and distribution follow the National List of Scientific Plant Names 
 (1982) except as modified by State distribution data from an unpublished plant 
 species geographic data base (Kartesz). 
 
 Epiphytic (e.g., mistletoes and some orchids) and nonrooted species (e.g., 
 dodder) were excluded from the National List because their roots were 
 respectively either never exposed to ground-level soil conditions or were not 
 in existence during all or part of the plant's life span. The current National 
 List contains only vascular plants, but a companion list of Bryophytes (mosses 
 and liverworts) that occur in wetlands is being developed to further define the 
 wetland flora of the United States. The Algae have also been excluded from the 
 current list. 
 
 Listings of the members of the National and appropriate Regional Interagency 
 Review Panels and the regional reviewers are included at the end of each list 
 (Appendix D) . Not all categories listed below are displayed in each National, 
 regional, or State report. The National alphabetical scientific name list is 
 reported by SCI-NAME, NAT-IND, RIIND, R2IND, R3IND, R4IND, R5IND, R6IND, R7IND, 
 R8IND, R9IND, POIND, RAIND, RCIND, RHIND, and REGION. The National trinomial 
 list is reported by SCI-NAME, TRINOM, RIIND, R2IND, R3IND, R4IND, R5IND, R6IND, 
 R7IND, R8IND, R9IND, POIND, RAIND, RCIND, RHIND, and REGION. The National, 
 regional, and State synonym lists are reported by SYNONYMY, SCI-NAME, AUTHOR, 
 and REGION. The regional and State alphabetical scientific name lists are 
 reported by SCI-NAME, AUTHOR, COMMON-NAME, RIND, NAT-IND, and HABIT. The 
 regional and State trinomial lists are reported by SCI-NAME, AUTHOR, TRINOM, 
 RIND, NAT-IND, and HABIT. The State alphabetical common-name lists are reported 
 within life forms (HABIT) by COMMON-NAME, RIND, NAT-IND, SYMBOL, SCI-NAME, and 
 HABIT. 
 
 The information in these lists is presented in the following categories. A brief 
 definition of the categories reported in the various lists is given below: 
 
SCI-NAME (Scientific Name) 
 
 The genus and species applied to the taxon by the National List of 
 Scientific Plant Names (1982). 
 
 SYMBOL 
 
 Symbol assigned in the National List of Scientific Plant Names (1982), 
 consisting of the first two letters of the genus name and the first two 
 letters of the specific epithet, with additional numbers added in numeric 
 sequence to the four-letter symbol to break ties. Tentative plant 
 symbols for species not in the National List of Scientific Plant Names 
 (1982) have been created by taking the first two letters of the genus and 
 specific epithet, adding a numeric tie breaker, if necessary, and ending 
 with a question mark. All species have a unique symbol. 
 
 AUTHOR 
 
 The author of the scientific name as cited by the National List of 
 Scientific Plant Names (1982). 
 
 SYNONYMY 
 
 Alternate scientific names applied to the species by major regional or 
 State floras. 
 
 TRINOM (Trinomial) 
 
 Varieties, subspecies, or forms which differ in Indicator assignment from 
 the species. 
 
 NAT-IND (National Range Of Indicators) 
 
 The National Indicators reflect the range of estimated probabilities 
 (expressedas a frequency of occurrence) of a species occurring in wetland 
 versus nonwetland across the entire distribution of the species. A 
 frequency, for example, of 67%-99% (Facultative Wetland) means that 67%-99% 
 of sample plots containing the species randomly selected across the range 
 of the species would be wetland. Aquestion mark (?) following an Indicator 
 denotes a tentative assignment based on the botanical literature and not 
 confirmed by regional review. When two indicators are given, they reflect 
 the range from the lowest to the highest frequency of occurrence in 
 wetlands across the regions in which the species is found. A positive (+) 
 or negative (-) sign was used with the Facultative Indicator categories 
 to more specifically define the regional frequency of occurrence in 
 wetlands. The positive sign indicates a frequency toward the higher end 
 of the category (more frequently found in wetlands), and a negative sign 
 
 8 
 
indicates a frequency toward the lower end of the category (less frequently 
 found in wetlands) . 
 
 Indicator Categories 
 
 Obligate Wetland (OBL). Occur almost always (estimated probability >99%) 
 under natural conditions in wetlands. 
 
 Facultative Wetland (FACW). Usually occur in wetlands (estimated 
 probability 67%-99%), but occasionally found in nonwetlands. 
 
 Facultative (FAC). Equally likely to occur in wetlands or nonwetlands 
 (estimated probability 34%-66%). 
 
 Facultative Upland (FACU). Usually occur in nonwetlands (estimated 
 probability 67%-99%), but occasionally found in wetlands (estimated 
 probability l%-33%). 
 
 Obligate Upland (UPL). Occur in wetlands in another region, but occur 
 almost always (estimated probability >99%) under natural conditions in 
 nonwetlands in the region specified. If a species does not occur in 
 wetlands in any region, it is not on the National List . 
 
 The wetland Indicator categories should not be equated to degrees of 
 wetness. Many obligate wetland species occur in permanently or 
 semipermanently flooded wetlands, but a number of obligates also occur 
 and some are restricted to wetlands which are only temporarily or 
 seasonally flooded. The facultative upland species include a diverse 
 collection of plants which range from weedy species adapted to exist in 
 a number of environmentally stressful or disturbed sites (including 
 wetlands) to species in which a portion of the gene pool (an ecotype) 
 always occurs in wetlands. Both the weedy and ecotype representatives of 
 the facultative upland category occur in seasonally and semipermanently 
 flooded wetlands. 
 
 R_IND (Regional Indicator) 
 
 The estimated probability (likelihood) of a species occurring in wetlands 
 versus nonwetlands in the region. Regional Indicators reflect the 
 unanimous agreement of the Regional Interagency Review Panel. If a 
 regional panel was not able to reach a unanimous decision on a species, 
 NA (no agreement) was recorded in the regional indicator (RIND) field. 
 An NI (no indicator) was recorded for those species for which insufficient 
 information was available to determine an indicator status. A 
 nonoccurrence (NO) designation indicates that the species does not occur 
 in that region. An asterisk (*) following a regional Indicator identifies 
 tentative assignments based on limited information from which to determine 
 the indicator status. In the listings for the States divided into two 
 regions (Montana, Wyoming, and Colorado), both regional Indicators are 
 reported. 
 
REGION 
 
 The distribution of the species expressed by the regional codes used in 
 the National List of Scientific Plant Names (1982). These code numbers 
 and regions are displayed in Figure 1. 
 
 REGION 
 
 CODE REGION 
 
 1 Northeast 
 
 2 Southeast 
 
 3 North Central 
 
 4 North Plains 
 
 5 Central Plains 
 
 6 South Plains 
 
 7 Southwest 
 
 8 Intermountain 
 
 9 Northwest 
 California 
 A Alaska 
 
 C Caribbean 
 
 Hawaii 
 
 STATE(S) IN REGION 
 
 ME,NH,VT,MA,CT,RI,WV,KY,NY,PA, 
 
 NJ,MD,DE,VA,OH 
 
 NC,SC,GA,FL,TN,AL,MS,LA,AR 
 
 MO,IA,MN,MI,WI,IL,IN 
 
 ND,SD,MT(Eastern) ,WY(Eastern) 
 
 NE,KS,CO(Eastern) 
 
 TX,OK 
 
 AZ,NM 
 
 NV,UT,CO{Western) 
 
 WA,OR, ID,MT(Western) ,WY{Western) 
 
 CA 
 
 AK 
 
 PR(Puerto Rico), VI(U.S. Virgin 
 
 Isls.)» CZ(Canal Zone), SQ (Swan 
 
 Isls.) 
 
 HI(Hawaiian Isls.), AQ(American 
 
 Samoa), GU(Guam), IQ(U.S. Misc. 
 
 Pacific Isls.), MQ(Midway Isls.), 
 
 TQ(Trust territories of the 
 
 Pacific Isls.), WQ(Wake Isl.), 
 
 YQ(Ryukyu Isls. Southern) 
 
 COMMON -NAME 
 
 HABIT 
 
 A popular name applied to the species. Common name selection generally 
 follows Common Names for North American Plants (Kartesz and Thieret, in 
 press), but some common names follow the current common name list 
 maintained by the Soil Conservation Service. 
 
 The plant characteristics and life forms assigned to each species in the 
 National List of Scientific Plant Names (1982) and by the Soil Conservation 
 Service. Family names are listed alphabetically under specific life forms 
 restricted to these families. 
 
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 11 
 

 CHARACTERISTIC 
 
 
 
 CHARACTERISTIC 
 
 SYMBOL 
 
 OR LIFE FORM 
 Annual 
 
 SYMBOL 
 G 
 
 OR LIFE FORM 
 
 A 
 
 Grass 
 
 B 
 
 Biennial 
 
 
 
 POACEAE 
 
 C 
 
 Clubmoss 
 
 LYCOPODIACEAE 
 SELAGINELLACEAE 
 
 GL 
 
 = 
 
 Grassl ike 
 CYPERACEAE 
 JUNCACEAE 
 
 E 
 
 Emergent 
 
 H 
 
 = 
 
 Partly woody 
 
 
 
 Epiphytic 
 
 HS 
 
 = 
 
 Half shrub 
 
 F 
 
 Forb 
 
 H2 
 
 = 
 
 Horsetail 
 
 / = 
 
 Floating 
 
 
 
 EQUISETACEAE 
 
 F3 = 
 
 Fern 
 
 I 
 
 = 
 
 Introduced 
 
 
 ADIANTACEAE 
 
 N 
 
 = 
 
 Native 
 
 
 ASPLENIACEAE 
 
 P 
 
 = 
 
 Perennial 
 
 
 BLECHNACEAE 
 
 + 
 
 = 
 
 Parasitic 
 
 
 CYATHEACEAE 
 
 P3 
 
 = 
 
 Pepperwort 
 
 
 DAVALLIACEAE 
 
 
 
 MARSILEACEAE 
 
 
 DENNSTAEDTIACEAE 
 
 Q 
 
 = 
 
 Quillwort 
 
 
 DRYOPTERIDACEAE 
 
 
 
 ISOETACEAE 
 
 
 GLEICHENIACEAE 
 
 S 
 
 = 
 
 Shrub 
 
 
 GRAMMITIDACEAE 
 
 - 
 
 = 
 
 Saprophytic 
 
 
 HYMENOPHYLLACEAE 
 
 z 
 
 = 
 
 Submerged 
 
 
 LOMARIOPSIDACEAE 
 
 $ 
 
 = 
 
 Succulent 
 
 
 MARATTIACEAE 
 
 T 
 
 = 
 
 Tree 
 
 
 OPHIOGLOSSACEAE 
 
 V 
 
 = 
 
 Herbaceous Vine 
 
 
 OSMUNDACEAE 
 
 w 
 
 = 
 
 Waterfern 
 
 
 PARKERIACEAE 
 
 
 
 AZOLLACEAE 
 
 
 POLYPODIACEAE 
 
 
 
 SALVINIACEAE 
 
 
 PSILOTACEAE 
 
 wv 
 
 = 
 
 Woody vine 
 
 
 PTERIDAC EAE 
 
 
 
 
 
 SCHIZAEACEAE 
 
 
 
 
 The HABIT symbols are combined to describe the life form of the 
 {e.g., ANG means annual native grass, IT means introduced tree). 
 
 species 
 
 12 
 
DIGITAL DATA BASE AVAILABILITY 
 
 The National List of Plant Species that Occur jn Wetlands is maintained in a 
 microcomputer data base. This data base was created to track and document the 
 decisions made by the Regional Interagency Review Panels and to facilitate 
 generation of National, regional and State reports. The data base is organized 
 into 26 fixed-length and 3 variable-length fields and contains information on 
 plant taxonomy, geography, and wetland Indicator status. We expect to make 
 regional subdivisions of the data base and a user's manual available on selected 
 magnetic media. An announcement of the format and ordering procedures will be 
 distributed widely when the data base is available in digital form. 
 
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LITERATURE CITED 
 
 Cowardin, L.M., V. Carter, F.C. Golet, and E.T. LaRoe. 1979. Classification 
 of wetlands and deepwater habitats of the United States. U.S. Fish Wildl. 
 Serv. FWS/OBS-79/31. 103 pp. 
 
 Environmental Laboratory. 1987. Corps of Engineers wetlands del ineation manual . 
 Technical Report 4-87-1. Corps of Engineers, Waterways Experiment Station, 
 Vicksburg, MS. 
 
 Huffman, R.T. 1981. Multiple-parameter approach to the field identification 
 and delineation of aquatic and wetland ecosystems. Technical Report EL-81. 
 U.S. Army Corps of Engineers, Waterways Experiment Station, Vicksburg, MS. 
 
 Kartesz, J.T. Unpublished plant species geographic data base. Biota of North 
 America program. North Carolina Botanical Garden, Chapel Hill. 
 
 Kartesz, J.T., and J. Thieret. Common names for North American plants. In 
 press. 
 
 Reed, P.B., Jr. 1986. 1986 National wetland plant list - regional indicator 
 compilation. U.S. Fish Wildl . Serv. WELUT 86/W17.01 . 121pp. 
 
 Sipple, W.S. 1988. Wetland identification and delineation manual. Volume I. 
 Rationale, wetland parameters, and overview of jurisdictional approach. 
 Volume II. Field methodology. U.S. Environmental Protection Agency, 
 Washington, DC. Vol. 1:30 pp.. Vol. 2:40 pp. 
 
 U.S. Department of Agriculture. 1971. National List of scientific plant names. 
 U.S. Soil Conservation Service, Lincoln, NE. 281 pp. 
 
 U.S. Department of Agriculture. 1982. National List of scientific plant names. 
 Vol. 1. List of plant names. Vol. 2. Synonymy. U.S. Soil Conservation 
 Service, SCS-TP-159, Washington, DC. Vol. 1:416 pp.. Vol. 2:438 pp. 
 
 U.S. Department of Agriculture. 1987. Hydric soils of the United States. Soil 
 Conservation Service, Washington, DC. np. 
 
 Wentworth, T.R., and G.P. Johnson. 1986. Use of vegetation for the designation 
 of wetlands. U.S. Fish Wildl. Serv., Washington DC. 107 pp. 
 
 57 
 
APPENDIX A. REVIEW PROCEDURE 
 
 The procedure for adding, deleting, or changing the Indicator status of plant 
 species on the National List is given below. 
 
 We welcome improvement of the National List and encourage the submission of 
 Indicator changes to misclassified, unclassified, or unlisted species. We also 
 encourage any additional suggestions for changes to any of the data contained 
 in the National List , especially improved information on distribution and 
 additional synonyms, or for improvement of the format of data presentation. 
 
 If certain species should be removed, added, or remain on the National List with 
 a revised Indicator status, gather the information to support your recommendation 
 and submit it on the enclosed review sheet to: 
 
 Porter B. Reed, Jr. 
 
 National Ecology Research Center 
 
 U.S. Fish & Wildlife Service 
 
 Monroe Building - Suite 101 
 
 9720 Executive Center Drive 
 
 St. Petersburg, FL 33702-2440 
 
 Proposed changes and supporting information will be presented to the appropriate 
 Regional Review Panel for consideration. The National Review Panel will review 
 all regional Indicator assignments for consistency and accuracy and serve as the 
 final arbitrator for the Regional Review Panels. 
 
 The National List will be updated as changes to the current list are made, and 
 revised editions will be printed as needed. Improvement of the National List 
 is dependent on a strong review by knowledgable botanists and ecologists from 
 across the country. We sincerely appreciate your help in completing this task. 
 
 58 
 
APPENDIX B. INSTRUCTIONS FOR COMPLETING THE REVIEW SHEET FOR PLANT SPECIES THAT 
 OCCUR IN WETLANDS 
 
 Reviewer: Reviewer's name 
 
 Address: Mailing address and phone number of reviewer 
 
 Species Symbol : The symbol given in the National List of Scientific Plant Names 
 
 (1982) 
 
 Scientific Name: Genus, species, and any additional infraspecific designations 
 
 as well as the author's name. 
 
 Geographic area 
 
 covered by reviewer: The area of your fieldwork and experience from which your 
 
 review is derived. Be as specific as possible with the 
 
 regional wetland plant list area. 
 
 Action: Check appropriate line. 
 
 Indicator: Complete appropriate lines. Deleted species will not have a 
 recommended Indicator. Added species will not have a current 
 Indicator. 
 
 Review Comments: 
 
 This is the primary section for explaining the rationale for any regional or 
 subregional changes, deletions, or additions to the list. Information on the 
 specific habitats, microsites, or soils that the species occurs in should be 
 provided. Use the back of the page if additional space is needed. 
 Site-specific information supporting review comments: 
 
 This section is for the description of the specific community (or communities) 
 in which the species is found and which support your review comments. Use 
 additional sheets for more community descriptions - staple all sheets for each 
 species together. The soils and hydrology sections are optional, but it will 
 be very useful if you can enter this information. Provide as much information 
 as possible on the composition of the plant communities, soils, and hydrology 
 that contain the species of concern. Include, where possible, community 
 composition, soils, and hydrology data on other sites, drier or wetter, that the 
 species also occurs in, which help support your suggested Indicator status. 
 Include site-specific information, with the locations well specified, to identify 
 the field sites where information has been gathered. Present quantitative data 
 to support your change in a form that can be readily reviewed. If possible, enter 
 the soil series and water regime as described in Cowardin et al . (1979). 
 
 59 
 
APPENDIX C. REVIEW SHEET FOR PLANT SPECIES THAT OCCUR IN WETLANDS 
 
 Rev i ewer 
 
 Address 
 
 Species symbol 
 
 Scientific name 
 
 Geographic area covered by reviewer 
 
 Actionlndicator 
 
 Addition 
 
 Change of Indicator Current _ 
 
 Deletion Recommended 
 
 REVIEW COMMENTS. 
 
 Including physiographic setting and additional data sources - cite specific 
 reference{s) , other than standard botanical texts, if used. 
 
 Site-specific information supporting review comments 
 
 Community composition and dominants: 
 Overstory 
 Shrub 
 
 Herbaceous 
 Soil 
 Hydrology 
 
 60 
 
APPENDIX D. INTERAGENCY REVIEW PANEL MEMBERS AND REGIONAL REVIEWERS 
 
 National Interagency Review Panel 
 
 U.S. Fish and Wildlife Service 
 
 Joseph Dowhan 
 
 U.S. Fish & Wildlife Service 
 Dept. of the Interior, Rm. 2553 
 Washington, DC 20240 
 
 Russell Kologiski 
 Dept. of Biology 
 Univ. of NC, Charlotte 
 Charlotte, NC 28223 
 
 Porter B. Reed, Jr. 
 National Wetlands Inventory 
 U.S. Fish and Wildlife Service 
 Monroe Bldg., Suite 101 
 9720 Executive Center Drive 
 St. Petersburg, FL 33702 
 
 Stephen S. Talbot 
 
 Refuge Division 
 
 U.S. Fish and Wildlife Service 
 
 1011 E. Tudor Road 
 
 Anchorage, AK 99503 
 
 U.S. Army Corps of Engineers 
 
 * Richard Macomber, Chief 
 Environmental Division 
 Board of Engineers for 
 
 Rivers and Harbors 
 U.S. Army Corps of Engineers 
 Kingman Building 
 Ft. Belvoir, VA 22060 
 
 Charles J. Newl ing 
 Waterways Experiment Station 
 U.S. Army Corps of Engineers 
 P.O. Box 631 
 Vicksburg, MS 39180 
 
 Robert Pierce 
 
 U.S. Army Corps of Engineers 
 
 CECW-OR 
 
 20 Mass. Ave., NW 
 
 Washington, DC 20314-1000 
 
 (* - Former Panel Members) 
 
 * Dana Sanders 
 Waterways Experiment Station 
 U.S. Army Corps of Engineers 
 P.O. Box 631 
 Vicksburg, MS 39180 
 
 Fred Weinmann, Chief 
 Estuarine Research Unit 
 U.S. Army Corps of Engineers 
 Seattle District 
 P.O. Box C-3755 
 Seattle, WA 98124 
 
 61 
 
U.S. Environmental Protection Agency 
 
 Charles Rhodes 
 
 U.S. Environ. Protect. Agcy. 
 
 Environmental Impact & Marine 
 
 Policy Branch 3ES40 
 841 Chestnut Bldg. 
 Philadelphia, PA 19107 
 
 Ralph Rogers 
 
 U.S. Environ. Protect. Agcy. 
 
 Oregon Operations Office 
 
 522 Southwest 5th Ave. 
 
 Yeon Building 
 
 Portland, OR 97204 
 
 William Sipple 
 
 Office of Wetlands Protection 
 
 A-104F 
 U.S. Environmental Pr. Agcy. 
 401 M Street, SW 
 Washington, DC 20460 
 
 U.S. Soil Conservation Service 
 
 * Robert S. Maclauchlan 
 
 Nat. Plant Materials Special. 
 USDA-Soil Conservation Service 
 P.O. Box 2890 
 Washington, DC 20013 
 
 * Donald Pendleton 
 
 National Range Conservationist 
 USDA-Soil Conservation Service 
 P.O. Box 2890 
 Washington, DC 20013 
 
 * Thomas N. Shiflet 
 
 Director of Ecological Sciences 
 USDA-Soil Conservation Service 
 P.O. Box 2890 
 Washington, DC 20013 
 
 Billy Teels, Nat'l Biologist 
 USDA-Soil Conservation Service 
 P.O. Box 2890 
 Washington, DC 20013 
 
 * Carl Thomas 
 National Biologist 
 USDA-Soil Conservation Service 
 P.O. Box 2890 
 Washington, DC 20013 
 
 62 
 
Regional Interagency Review Panel : 
 
 North Plains and 
 (Regions 4 & 5) 
 
 Central Plains 
 
 U.S. Fish and Wildlife Service 
 
 Chuck Ell iott 
 
 Regional Wetland Coordinator 
 
 U.S. Fish and Wildlife Service 
 
 Denver Federal Center 
 
 P.O. Box 25486 
 
 Denver, CO 80228 
 
 * Russell Kologiski 
 Dept. of Biology 
 Univ. of NC, Charlotte 
 Charlotte, NC 28223 
 
 U.S. Army Corps of Engineers 
 
 Mike Gilbert 
 
 U.S. Army Corps of Engineers 
 
 Regulatory Branch 
 
 P.O. Box 5 
 
 Omaha, NE 68101-0005 
 
 U.S. Environmental Protection Agency 
 
 Glen Yager 
 
 U.S. Environmental Prot. Agcy. 
 
 Federal Facilities Coordinator 
 
 Environment Review Branch 
 
 726 Minnesota Avenue 
 
 Kansas City, KS 66101 
 
 U.S. Soil Conservation Service 
 
 Rex Hamilton 
 
 USDA-Soil Conservation Service 
 100 Centennial Mall North 
 Federal Bldg., Rm. 345 
 Lincoln, NE 68508-3866 
 
 63 
 
Regional Reviewers : North Plains and Central Plains (Regions 4 & 5) 
 
 Will iam T. Barker 
 
 Dept. of Botany 
 
 North Dakota State University 
 
 Fargo, ND 58105 
 
 Gary E. Larson 
 Botany-Biology Dept. 
 Agriculture Hall 304 
 SD State University 
 Brookings, SD 57007 
 
 Ralph Brooks, Research Asst. 
 Kansas Biological Survey 
 2291 Irving Mill Rd., Campus W. 
 Lawrence, KS 66045 
 
 Bonnie Meidel 
 Natural Heritage Program 
 1424 West Century Avenue 
 Bismarck, ND 58501 
 
 David J. Cooper 
 
 Adjunct Assistant Professor 
 
 Environmental Sciences and 
 
 Engineering Ecology 
 Colorado School of Mines 
 Golden, CO 80401 
 
 Dave Ode 
 
 Natural Heritage Program 
 
 Div. of Parks and Recreation 
 
 Anderson Bldg. 
 
 445 East Capitol Ave. 
 
 Pierre, SD 57501 
 
 Paul Currier 
 
 Platte River Trust 
 
 2550 North Diers Avenue 
 
 Suite H 
 
 Grand Island, NE 68801 
 
 David Sutherland 
 Dept. of Biology 
 University of Nebraska 
 Omaha, NE 68132 
 
 Ron Hartmann 
 
 Rocky Mountain Herbarium 
 3165 University Station 
 University of Wyoming 
 Laramie, WY 82071 
 
 Richard Walter 
 
 3525 Terry Lake Road 
 
 Fort Collins, CO 80524 
 
 Hal Kantrud 
 
 U.S. Fish and Wildlife Service 
 
 Northern Prairie Wildlife 
 
 Research Center 
 P.O. Box 1747 
 Jamestown, ND 58401 
 
 Robert Kaul 
 
 School of Biological Sciences 
 University of Nebraska 
 Lincoln, NE 68588 
 
 64 
 
30372-101 
 
 REPORT DOCUMENTATION 
 PAGE 
 
 1. REPORT NO. 
 
 Biological Report 88(26.4) 
 
 4. TItIa and Subtltl* 
 
 National List of Plant Species That Occur in Wetlands: 
 North Plains (Region 4) 
 
 3. RaclDlant's Accattlon No. 
 
 5. Report Data 
 
 May 1988 
 
 7. Author(») 
 
 Porter B. Reed 
 
 8. Parformlng Organization Rapt. No. 
 
 Jr. 
 
 9. Parforming Organization Nama and Addraaa 
 
 National Ecology Research Center 
 U.S. Fish and Wildlife Service 
 Creekside One Bldg., 2627 Redwing Rd 
 Fort Collins, CO 80526-2899 
 
 10. Projact/Tatk/Worl( Unit No. 
 
 11. Contract(C) or Grant(G) No. 
 
 (C) 
 
 (G) 
 
 12. Sponsoring Organization Nama and Address 
 
 Department of the Interior 
 U.S. Fish and Wildlife Service 
 Research and Development 
 Washington, D.C. 20240 
 
 13. Typ« of Report & Period Covered 
 
 14. 
 
 IS. Supplementary Notes 
 
 le. Abstract (Limit: 200 words) 
 
 The National U 
 efforts of many 
 United States 
 in order to pro 
 Habitats of the 
 of wetlands. P 
 are defined as 
 reproduce in an 
 become, periodi 
 season. The 11 
 regulatory prog 
 
 St of Plant Species That Occu r in Wetlands represents the combined 
 biologists over the last decade to define the wetland flora of the 
 The U.S. Fish and Wildlife Service initially developed the list 
 vide an appendix to the Classification of Wetlands and Deepwater 
 United States (FWS/OBS 79/31) to assist in the field identification 
 lant species that occur in wetlands, as used in the National List , 
 species that have demonstrated an ability to achieve maturity and 
 environment where all or portions of the soil within the root zone 
 cally or continuously, saturated or inundated during the growing 
 st will facilitate the Implementation and management of the wetland 
 rams of the Federal Government and most State governments. 
 
 This list for the North Plains (Region 4) is a subset of the National List , 
 
 17. Oocumant Analysis a. Oascriptors 
 
 Plants 
 
 Plant ecology 
 
 b. Idantlfiars/Open-Endad Terms 
 
 Wetland plant species 
 North Plains (Region 4) 
 
 c. COSATI Field/Group 
 
 18. Availability Statement 
 
 Release unlimited 
 
 19. Security Class (This Report) 
 
 Unclassified 
 
 20. Security Class (This Page) 
 
 Unclassified 
 
 21. No. of Pages 
 
 64 
 
 22. Price 
 
 (See ANSI-Z39.18) 
 
 See Instructions on Reverie 
 
 (}P0 576-831/85234 
 
 OPTIONAL FORM 272 (4-77) 
 (Formerly NTlS-35) 
 Department of Commerce 
 
Hawaiian Islands ■' 
 
 Puerto Rico and 
 Virgin Islands 
 
 ^ 
 
 Headquarters, Research and Developnnenl 
 Washington, DC 
 
 ♦ Locations ol Regional Offices 
 
 REGION 1 
 
 Regional Director 
 
 U.S, Fish and Wildlife Service 
 
 Lloyd Five Hundred Building, Suite 1692 
 
 500 N.E. Multnomah Street 
 
 Portland, Oregon 97232 
 
 REGION 2 
 
 Regional Director 
 
 U,S, Fish and Wildlife Service 
 
 P,0, Box 1306 
 
 Albuquerque, New Mexico 87103 
 
 REGION 3 
 
 Regional Director 
 U.S. Fish and Wildlife Service 
 Federal Building, Fort Snelling 
 Twin Cities, Minnesota 55111 
 
 REGION 4 
 
 Regional Director 
 U.S. Fish and Wildlife Service 
 Richard B. Russell Building 
 75 Spring Street, S.W. 
 Atlanta, Georgia 30303 
 
 REGION 5 
 
 Regional Director 
 
 U.S. Fish and Wildlife Service 
 
 One Gateway Center 
 
 Newton Corner, Massachusetts 02158 
 
 REGION 6 
 
 Regional Director 
 
 U.S. Fish and Wildlife Service 
 
 P.O. Box 25486 
 
 Denver Federal Center 
 
 Denver, Colorado 80225 
 
 REGION 7 
 
 Regional Director 
 U.S. Fish and Wildlife Service 
 1011 E. Tudor Road 
 Anchorage, Alaska 99503 
 
UNIVERSITY OF ILLINOIS-URBANA 
 
 Knal t^'foF Vant spe'cTs that occu 
 
 TAKE 
 PRIDE IN 
 AMERICA 
 
 3 0112 017575975 
 
 Preserve Our Natural Resources 
 
 I .s. 
 
 HSH*WII.IM.IKt 
 SF.RVKK 
 
 DEPARTMENT OF THE INTERIOR 
 
 U.S. FISH AND WILDLIFE SERVICE 
 
 As the Nation's principal conservation agency, the Department of the Interior has respon- 
 sibility for most of our nationally owned public lands and natural resources. This includes 
 fostering the wisest use of our land and water resources, protecting our fish and wildlife, 
 preservmg thfrenvironmental and cultural values of our national parks and historical places, 
 and providing for the enjoyment of life through outdoor recreation. The Department as- 
 sesses our energy and mineral resources and works to assure that their development is in 
 the best interests of all our people. The Department also has a major responsibility for 
 American Indian reservation communities and for people who live in island territories under 
 U.S. administration.